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<document id="03D316DAFAB4741A8B60A859FC35055C" ID-DOI="10.5281/zenodo.278442" ID-GBIF-Dataset="6b9505e0-ac6e-4baa-9066-6f729d38e9e2" ID-ISSN="1175-5326" ID-Zenodo-Dep="278442" IM.bibliography_approvedBy="felipe" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1460341493798" checkinUser="plazi" docAuthor="Hershler, Robert, Liu, Hsiu-Ping &amp; Landye, Jerry" docDate="2011" docId="03E687C5BF7B8300FF2E12BDFCF11D7B" docLanguage="en" docName="zt03001p032.pdf" docOrigin="Zootaxa 3001" docStyle="DocumentStyle:890A69B780ED73D6DB8551B71C8AC79E.4:Zootaxa.2009-2012.journal_article" docStyleId="890A69B780ED73D6DB8551B71C8AC79E" docStyleName="Zootaxa.2009-2012.journal_article" docStyleVersion="4" docTitle="Tryonia angosturae Hershler, Liu &amp; Landye, 2011, sp. nov." docType="treatment" docVersion="8" lastPageNumber="15" masterDocId="FFDFFFBDBF77830EFFB9171AFFA4194F" masterDocTitle="New species and records of springsnails (Caenogastropoda: Cochliopidae: Tryonia) from the Chihuahuan Desert (Mexico and United States), an imperiled biodiversity hotspot" masterLastPageNumber="32" masterPageNumber="1" pageNumber="13" updateTime="1698261240372" updateUser="plazi">
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<mods:title id="DCB7B02D671AA22F8FE6BDB8A1880319">New species and records of springsnails (Caenogastropoda: Cochliopidae: Tryonia) from the Chihuahuan Desert (Mexico and United States), an imperiled biodiversity hotspot</mods:title>
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<mods:namePart id="05A5A70F306CD380D988261E30E4FADF">Hershler, Robert</mods:namePart>
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<mods:namePart id="76347243AB3FB6E9E4FE24810C1D943F">Liu, Hsiu-Ping</mods:namePart>
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<mods:namePart id="AE60869FF7C47A58C9C933301B33B82F">Landye, Jerry</mods:namePart>
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<mods:date id="A2C8C3BAA1FCDE0B7D3C56AB03431624">2011</mods:date>
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<treatment id="03E687C5BF7B8300FF2E12BDFCF11D7B" ID-DOI="http://doi.org/10.5281/zenodo.5629195" ID-GBIF-Taxon="119406533" ID-Zenodo-Dep="5629195" LSID="urn:lsid:plazi:treatment:03E687C5BF7B8300FF2E12BDFCF11D7B" httpUri="http://treatment.plazi.org/id/03E687C5BF7B8300FF2E12BDFCF11D7B" lastPageId="14" lastPageNumber="15" pageId="12" pageNumber="13">
<subSubSection id="C3556558BF7B8302FF2E12BDFDD71FFA" pageId="12" pageNumber="13" type="nomenclature">
<paragraph id="8BF036D3BF7B8302FF2E12BDFE4D1C8E" blockId="12.[151,489,1447,1506]" box="[151,489,1447,1473]" pageId="12" pageNumber="13">
<heading id="D0B881BFBF7B8302FF2E12BDFE4D1C8E" bold="true" box="[151,489,1447,1473]" fontSize="11" level="1" pageId="12" pageNumber="13" reason="1">
<emphasis id="B93BEAC1BF7B8302FF2E12BDFE4D1C8E" bold="true" box="[151,489,1447,1473]" pageId="12" pageNumber="13">
<taxonomicName id="4C4F4D50BF7B8302FF2E12BDFE201C8E" box="[151,388,1447,1473]" class="Gastropoda" family="Hydrobiidae" genus="Tryonia" kingdom="Animalia" order="Littorinimorpha" pageId="12" pageNumber="13" phylum="Mollusca" rank="species" species="angosturae" status="sp. nov.">
<emphasis id="B93BEAC1BF7B8302FF2E12BDFE201C8E" bold="true" box="[151,388,1447,1473]" italics="true" pageId="12" pageNumber="13">Tryonia angosturae</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A20857BABF7B8302FE3512BDFE4D1C8E" box="[396,489,1447,1473]" pageId="12" pageNumber="13" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
<paragraph id="8BF036D3BF7B8302FF2E12D0FE771CAD" blockId="12.[151,489,1447,1506]" box="[151,467,1482,1506]" pageId="12" pageNumber="13">
(
<figureCitation id="13742A56BF7B8302FF2612D0FF411CAD" box="[159,229,1482,1506]" captionStart="FIGURE 5" captionStartId="10.[151,244,1857,1878]" captionTargetBox="[241,1348,158,1832]" captionTargetId="figure@10.[229,1363,145,1846]" captionTargetPageId="10" captionText="FIGURE 5. Penes. A. T. peregrina sp. nov., USNM 1153677. B. T. zaragozae sp. nov., USNM 1153679. C. T. ovata sp. nov., USNM 1153676. D. T. angosturae sp. nov., USNM 874174. E. T. molinae sp. nov., USNM 1153674. F. T. contrerasi sp. nov., USNM 1153670. G. T. taylori sp. nov., USNM 874047. H. T. taylori sp. nov., USNM 873252. I. T. metcalfi sp. nov., USNM 1153672. J. T. chuviscarae sp. nov., USNM 1001756. K. T. minckleyi sp. nov., USNM 1153673. L. T. julimesensis sp. nov., USNM 1153671. M. T. allendae sp. nov., USNM 1153667. N. T. oasiensis sp. nov., USNM 1153675. O. T. seemani, USNM 874125. P. T. seemani, USNM 874123. Scale bars = 250 µm. Db = distal bulb, Pa = papilla, Pd = penial duct, St = stylet." httpUri="https://zenodo.org/record/278447/files/figure.png" pageId="12" pageNumber="13">Figs 5</figureCitation>
D, 7CE, HI, M
<collectingCountry id="F3587643BF7B8302FE0E12D0FE6F1CAD" box="[439,459,1482,1506]" name="Iceland" pageId="12" pageNumber="13">O</collectingCountry>
)
</paragraph>
<paragraph id="8BF036D3BF7B8302FF2E1109FCAF1F67" blockId="12.[151,779,1554,1576]" box="[151,779,1554,1576]" pageId="12" pageNumber="13">
<treatmentCitationGroup id="AB5F11FDBF7B8302FF2E1109FCAF1F67" box="[151,779,1554,1576]" pageId="12" pageNumber="13">
<taxonomicName id="4C4F4D50BF7B8302FF2E1109FF411F67" box="[151,229,1555,1576]" class="Gastropoda" family="Hydrobiidae" genus="Tryonia" kingdom="Animalia" order="Littorinimorpha" pageId="12" pageNumber="13" phylum="Mollusca" rank="genus" status="sp. nov.">
<emphasis id="B93BEAC1BF7B8302FF2E1109FF411F67" box="[151,229,1555,1576]" italics="true" pageId="12" pageNumber="13">Tryonia</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A20857BABF7B8302FF551109FE861F67" box="[236,290,1555,1576]" pageId="12" pageNumber="13" rank="species">n. sp.</taxonomicNameLabel>
3.—
<treatmentCitation id="0AEE10C2BF7B8302FEE81108FDFA1F67" author="Hershler" box="[337,606,1554,1576]" page="1757" pageId="12" pageNumber="13" year="2005">
<bibRefCitation id="EFDE4B22BF7B8302FEE81108FD851F67" author="Hershler" box="[337,545,1554,1576]" pageId="12" pageNumber="13" refString="Hershler, R., Mulvey, M. &amp; Liu, H. - P. (2005) Genetic variation in the Desert springsnail (Tryonia porrecta): implications for reproductive mode and dispersal. Molecular Ecology, 14, 1755 - 1765." type="journal article" year="2005">
Hershler
<emphasis id="B93BEAC1BF7B8302FE0B1109FE7A1F67" box="[434,478,1554,1576]" italics="true" pageId="12" pageNumber="13">et al</emphasis>
. 2005
</bibRefCitation>
: 1757
</treatmentCitation>
(COI sequence).
</treatmentCitationGroup>
</paragraph>
<paragraph id="8BF036D3BF7B8302FF2E114FFDD71FFA" blockId="12.[151,1437,1621,1970]" pageId="12" pageNumber="13">
<emphasis id="B93BEAC1BF7B8302FF2E114FFF7E1F22" bold="true" box="[151,218,1621,1645]" pageId="12" pageNumber="13">
<typeStatus id="54F48871BF7B8302FF2E114FFF7E1F22" box="[151,218,1621,1645]" pageId="12" pageNumber="13">Types</typeStatus>
</emphasis>
.
<typeStatus id="54F48871BF7B8302FF48114FFEF81F22" box="[241,348,1621,1645]" pageId="12" pageNumber="13" type="holotype">Holotype</typeStatus>
, USNM 873259, Ojos de Arrey, western-most head spring, south of Galeana, Chihuahua,
<geoCoordinate id="EE7B5014BF7B8302FF2E1163FE8F1FDE" box="[151,299,1657,1681]" direction="north" orientation="latitude" pageId="12" pageNumber="13" precision="1" value="30.05786">30º3'28.3&quot; N</geoCoordinate>
,
<geoCoordinate id="EE7B5014BF7B8302FE811163FE491FDE" box="[312,493,1657,1681]" direction="west" orientation="longitude" pageId="12" pageNumber="13" precision="1" value="-107.590645">107º35'26.3&quot; W</geoCoordinate>
, leg. J.J.L.
<emphasis id="B93BEAC1BF7B8302FDC81160FD001FDE" box="[625,676,1657,1681]" italics="true" pageId="12" pageNumber="13">et al</emphasis>
.,
<date id="FFF11013BF7B8302FD021163FCEC1FDE" box="[699,840,1657,1681]" pageId="12" pageNumber="13" value="1971-08-25">25/viii/1971</date>
.
<typeStatus id="54F48871BF7B8302FCED1160FC651FDD" box="[852,961,1658,1682]" pageId="12" pageNumber="13" type="paratype">Paratypes</typeStatus>
(from same lot), USNM 873261, USNM 873285, USNM 905267, USNM 1153668.
</paragraph>
</subSubSection>
<subSubSection id="C3556558BF7B8302FF7E11D8FAA91F95" box="[199,1293,1729,1754]" pageId="12" pageNumber="13" type="etymology">
<paragraph id="8BF036D3BF7B8302FF7E11D8FAA91F95" blockId="12.[151,1437,1621,1970]" box="[199,1293,1729,1754]" pageId="12" pageNumber="13">
<emphasis id="B93BEAC1BF7B8302FF7E11D8FEEA1F95" bold="true" box="[199,334,1730,1754]" pageId="12" pageNumber="13">Etymology.</emphasis>
The species name is a geographic epithet referring to a nearby village, La
<taxonomicName id="4C4F4D50BF7B8302FB2911D8FAAD1F95" box="[1168,1289,1730,1754]" class="Gastropoda" family="Hydrobiidae" genus="Tryonia" kingdom="Animalia" order="Littorinimorpha" pageId="12" pageNumber="15" phylum="Mollusca" rank="species" species="angostura">Angostura</taxonomicName>
.
</paragraph>
</subSubSection>
<subSubSection id="C3556558BF7B8302FF7E11FFFC3D1EFE" pageId="12" pageNumber="13" type="materials_examined">
<paragraph id="8BF036D3BF7B8302FF7E11FFFC3D1EFE" blockId="12.[151,1437,1621,1970]" pageId="12" pageNumber="13">
<emphasis id="B93BEAC1BF7B8302FF7E11FFFE021FB1" bold="true" box="[199,422,1765,1790]" pageId="12" pageNumber="13">Referred material</emphasis>
. CHIHUAHUA.
<collectionCode id="ED5EAE16BF7B8302FDC211FFFD681FB2" LSID="urn:lsid:biocol.org:col:34871" box="[635,716,1765,1789]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34871" name="Smithsonian Institution, National Museum of Natural History" pageId="12" pageNumber="13">USNM</collectionCode>
873268, topotypes, leg. J.J.L.,
<date id="FFF11013BF7B8302FBEB11FFFB6B1FB2" box="[1106,1231,1765,1789]" pageId="12" pageNumber="13" value="1973-04-14">14/iv/1973</date>
.
<collectionCode id="ED5EAE16BF7B8302FB5811FFFA961FB2" LSID="urn:lsid:biocol.org:col:34871" box="[1249,1330,1765,1789]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34871" name="Smithsonian Institution, National Museum of Natural History" pageId="12" pageNumber="13">USNM</collectionCode>
874250,
<collectionCode id="ED5EAE16BF7B8302FF2E1013FF4C1E6E" LSID="urn:lsid:biocol.org:col:34871" box="[151,232,1801,1825]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34871" name="Smithsonian Institution, National Museum of Natural History" pageId="12" pageNumber="13">USNM</collectionCode>
874174,
<emphasis id="B93BEAC1BF7B8302FEED1013FE241E6E" box="[340,384,1801,1825]" italics="true" pageId="12" pageNumber="13">ibid</emphasis>
., leg. J.J.L. and P. Hines,
<date id="FFF11013BF7B8302FD151013FC8D1E6E" box="[684,809,1801,1825]" pageId="12" pageNumber="13" value="1990-09-16">16/ix/1990</date>
.
<collectionCode id="ED5EAE16BF7B8302FC8C1013FC221E6E" LSID="urn:lsid:biocol.org:col:34871" box="[821,902,1801,1825]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34871" name="Smithsonian Institution, National Museum of Natural History" pageId="12" pageNumber="13">USNM</collectionCode>
854943,
<emphasis id="B93BEAC1BF7B8302FC4A1013FBBB1E6E" box="[1011,1055,1801,1825]" italics="true" pageId="12" pageNumber="13">ibid</emphasis>
., leg. R.H. and J.J.L.,
<date id="FFF11013BF7B8302FA9B1013FA3D1E6E" box="[1314,1433,1801,1825]" pageId="12" pageNumber="13" value="1998-12-07">7/xii/1998</date>
.
<collectionCode id="ED5EAE16BF7B8302FF2E1037FF4C1E0A" LSID="urn:lsid:biocol.org:col:34871" box="[151,232,1837,1861]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34871" name="Smithsonian Institution, National Museum of Natural History" pageId="12" pageNumber="13">USNM</collectionCode>
873293, canal outflow from Ojos de Arrey at Hwy 10,
<geoCoordinate id="EE7B5014BF7B8302FCE71037FC7F1E0A" box="[862,987,1837,1861]" direction="north" orientation="latitude" pageId="12" pageNumber="13" precision="15" value="30.069723">30º4'11&quot; N</geoCoordinate>
,
<geoCoordinate id="EE7B5014BF7B8302FC5E1037FB211E0A" box="[999,1157,1837,1861]" direction="west" orientation="longitude" pageId="12" pageNumber="13" precision="15" value="-107.604164">107º36'15&quot; W</geoCoordinate>
, leg. J.J.L.,
<date id="FFF11013BF7B8302FAB51037FA3D1E0A" box="[1292,1433,1837,1861]" pageId="12" pageNumber="13" value="1971-08-25">25/viii/1971</date>
. BellMNH 21128, Oja del Rey (sic), leg. D.W. Taylor,
<date id="FFF11013BF7B8302FD57104BFCFA1E26" box="[750,862,1873,1897]" pageId="12" pageNumber="13" value="1969-09-03">3/ix/1969</date>
. BellMNH 21175, east Ojo del Rey (sic), northwest end of Cerro
<taxonomicName id="4C4F4D50BF7B8302FE95106CFE001EC1" box="[300,420,1910,1934]" class="Gastropoda" family="Hydrobiidae" genus="Tryonia" kingdom="Animalia" order="Littorinimorpha" pageId="12" pageNumber="15" phylum="Mollusca" rank="species" species="angostura">Angostura</taxonomicName>
, leg. D.W. Taylor and A. Metcalf,
<date id="FFF11013BF7B8302FC91106FFC331EC2" box="[808,919,1909,1933]" pageId="12" pageNumber="13" value="1969-09-03">3/ix/1969</date>
. BellMNH 21174, Ojo del Torreon, northwest end of Cerro
<taxonomicName id="4C4F4D50BF7B8302FE921080FE071EFD" box="[299,419,1946,1970]" class="Gastropoda" family="Hydrobiidae" genus="Tryonia" kingdom="Animalia" order="Littorinimorpha" pageId="12" pageNumber="15" phylum="Mollusca" rank="species" species="angostura">Angostura</taxonomicName>
, leg. D.W. Taylor and A. Metcalf,
<date id="FFF11013BF7B8302FC9F1083FC311EFE" box="[806,917,1945,1969]" pageId="12" pageNumber="13" value="1969-09-03">3/ix/1969</date>
.
</paragraph>
</subSubSection>
<caption id="DF30665BBF7A8303FF2E1035FEC41EBB" httpUri="https://zenodo.org/record/278449/files/figure.png" pageId="13" pageNumber="14" targetBox="[188,1404,151,1821]" targetPageId="13">
<paragraph id="8BF036D3BF7A8303FF2E1035FEC41EBB" blockId="13.[151,1436,1839,2036]" pageId="13" pageNumber="14">
<emphasis id="B93BEAC1BF7A8303FF2E1035FEA31E0B" bold="true" box="[151,263,1839,1860]" pageId="13" pageNumber="14">FIGURE 7</emphasis>
.
<taxonomicName id="4C4F4D50BF7A8303FEAD1035FE3E1E0B" box="[276,410,1839,1860]" class="Gastropoda" family="Hydrobiidae" genus="Tryonia" kingdom="Animalia" order="Littorinimorpha" pageId="13" pageNumber="14" phylum="Mollusca" rank="species" species="ovata" status="sp. nov.">
<emphasis id="B93BEAC1BF7A8303FEAD1035FE3E1E0B" box="[276,410,1839,1860]" italics="true" pageId="13" pageNumber="14">Tryonia ovata</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A20857BABF7A8303FE1B1035FE4B1E0B" box="[418,495,1839,1860]" pageId="13" pageNumber="14" rank="species">
<emphasis id="B93BEAC1BF7A8303FE1B1035FE4D1E0B" bold="true" box="[418,489,1839,1860]" pageId="13" pageNumber="14">sp. nov</emphasis>
.
</taxonomicNameLabel>
and
<taxonomicName id="4C4F4D50BF7A8303FD991035FD791E0B" box="[544,733,1839,1860]" class="Gastropoda" family="Hydrobiidae" genus="Tryonia" kingdom="Animalia" order="Littorinimorpha" pageId="13" pageNumber="14" phylum="Mollusca" rank="species" species="angosturae" status="sp. nov.">
<emphasis id="B93BEAC1BF7A8303FD991035FD791E0B" box="[544,733,1839,1860]" italics="true" pageId="13" pageNumber="14">Tryonia angosturae</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A20857BABF7A8303FD5D1035FC951E0B" box="[740,817,1839,1860]" pageId="13" pageNumber="14" rank="species">
<emphasis id="B93BEAC1BF7A8303FD5D1035FC8F1E0B" bold="true" box="[740,811,1839,1860]" pageId="13" pageNumber="14">sp. nov</emphasis>
.
</taxonomicNameLabel>
A. Holotype,
<taxonomicName id="4C4F4D50BF7A8303FC041035FBAC1E0B" box="[957,1032,1839,1860]" class="Gastropoda" family="Hydrobiidae" genus="Tryonia" kingdom="Animalia" order="Littorinimorpha" pageId="13" pageNumber="14" phylum="Mollusca" rank="species" species="ovata">
<emphasis id="B93BEAC1BF7A8303FC041035FBAC1E0B" box="[957,1032,1839,1860]" italics="true" pageId="13" pageNumber="14">T. ovata</emphasis>
</taxonomicName>
, USNM 873264. B. Male shell,
<taxonomicName id="4C4F4D50BF7A8303FAF11035FA321E0B" box="[1352,1430,1839,1860]" class="Gastropoda" family="Hydrobiidae" genus="Tryonia" kingdom="Animalia" order="Littorinimorpha" pageId="13" pageNumber="14" phylum="Mollusca" rank="species" species="ovata">
<emphasis id="B93BEAC1BF7A8303FAF11035FA321E0B" box="[1352,1430,1839,1860]" italics="true" pageId="13" pageNumber="14">T. ovata</emphasis>
</taxonomicName>
, UTEP 977. C. Holotype,
<taxonomicName id="4C4F4D50BF7A8303FE301057FDA91E2D" box="[393,525,1869,1890]" class="Gastropoda" family="Hydrobiidae" genus="Tryonia" kingdom="Animalia" order="Littorinimorpha" pageId="13" pageNumber="14" phylum="Mollusca" rank="species" species="angosturae">
<emphasis id="B93BEAC1BF7A8303FE301057FDA91E2D" box="[393,525,1869,1890]" italics="true" pageId="13" pageNumber="14">T. angosturae</emphasis>
</taxonomicName>
, USNM 873259. DE. Shells,
<taxonomicName id="4C4F4D50BF7A8303FC8F1057FC241E2D" box="[822,896,1869,1890]" class="Gastropoda" family="Hydrobiidae" genus="Tryonia" kingdom="Animalia" order="Littorinimorpha" pageId="13" pageNumber="14" phylum="Mollusca" rank="species" species="ovata">
<emphasis id="B93BEAC1BF7A8303FC8F1057FC241E2D" box="[822,896,1869,1890]" italics="true" pageId="13" pageNumber="14">T. ovata</emphasis>
</taxonomicName>
, USNM 1153668 (E is a male). FG. Operculum (outer, inner sides),
<taxonomicName id="4C4F4D50BF7A8303FEAB1070FEFB1E30" box="[274,351,1898,1919]" class="Gastropoda" family="Hydrobiidae" genus="Tryonia" kingdom="Animalia" order="Littorinimorpha" pageId="13" pageNumber="14" phylum="Mollusca" rank="species" species="ovata">
<emphasis id="B93BEAC1BF7A8303FEAB1070FEFB1E30" box="[274,351,1898,1919]" italics="true" pageId="13" pageNumber="14">T. ovata</emphasis>
</taxonomicName>
, USNM 1153676. HI. Operculum (outer, inner sides),
<taxonomicName id="4C4F4D50BF7A8303FCC01070FC591E30" box="[889,1021,1898,1919]" class="Gastropoda" family="Hydrobiidae" genus="Tryonia" kingdom="Animalia" order="Littorinimorpha" pageId="13" pageNumber="14" phylum="Mollusca" rank="species" species="angosturae">
<emphasis id="B93BEAC1BF7A8303FCC01070FC591E30" box="[889,1021,1898,1919]" italics="true" pageId="13" pageNumber="14">T. angosturae</emphasis>
</taxonomicName>
, USNM 1153688. J. Portion of radular ribbon,
<taxonomicName id="4C4F4D50BF7A8303FF70109DFEBD1ED3" box="[201,281,1927,1948]" class="Gastropoda" family="Hydrobiidae" genus="Tryonia" kingdom="Animalia" order="Littorinimorpha" pageId="13" pageNumber="14" phylum="Mollusca" rank="species" species="ovata">
<emphasis id="B93BEAC1BF7A8303FF70109DFEBD1ED3" box="[201,281,1927,1948]" italics="true" pageId="13" pageNumber="14">T. ovata</emphasis>
</taxonomicName>
, USNM 1153676. K. Central radular teeth,
<taxonomicName id="4C4F4D50BF7A8303FD69109DFCBB1ED3" box="[720,799,1927,1948]" class="Gastropoda" family="Hydrobiidae" genus="Tryonia" kingdom="Animalia" order="Littorinimorpha" pageId="13" pageNumber="14" phylum="Mollusca" rank="species" species="ovata">
<emphasis id="B93BEAC1BF7A8303FD69109DFCBB1ED3" box="[720,799,1927,1948]" italics="true" pageId="13" pageNumber="14">T. ovata</emphasis>
</taxonomicName>
, USNM 1153676. L. Lateral and inner marginal teeth,
<taxonomicName id="4C4F4D50BF7A8303FAFF109DFA321ED3" box="[1350,1430,1927,1948]" class="Gastropoda" family="Hydrobiidae" genus="Tryonia" kingdom="Animalia" order="Littorinimorpha" pageId="13" pageNumber="14" phylum="Mollusca" rank="species" species="ovata">
<emphasis id="B93BEAC1BF7A8303FAFF109DFA321ED3" box="[1350,1430,1927,1948]" italics="true" pageId="13" pageNumber="14">T. ovata</emphasis>
</taxonomicName>
, USNM 1153676. M. Portion of radular ribbon,
<taxonomicName id="4C4F4D50BF7A8303FDC110BFFCA41EF5" box="[632,768,1957,1978]" class="Gastropoda" family="Hydrobiidae" genus="Tryonia" kingdom="Animalia" order="Littorinimorpha" pageId="13" pageNumber="14" phylum="Mollusca" rank="species" species="angosturae">
<emphasis id="B93BEAC1BF7A8303FDC110BFFCA41EF5" box="[632,768,1957,1978]" italics="true" pageId="13" pageNumber="14">T. angosturae</emphasis>
</taxonomicName>
, USNM 1153688. N. Central radular teeth,
<taxonomicName id="4C4F4D50BF7A8303FB0610BFFAE31EF5" box="[1215,1351,1957,1978]" class="Gastropoda" family="Hydrobiidae" genus="Tryonia" kingdom="Animalia" order="Littorinimorpha" pageId="13" pageNumber="14" phylum="Mollusca" rank="species" species="angosturae">
<emphasis id="B93BEAC1BF7A8303FB0610BFFAE31EF5" box="[1215,1351,1957,1978]" italics="true" pageId="13" pageNumber="14">T. angosturae</emphasis>
</taxonomicName>
, USNM 1153688. O. Lateral and inner marginal teeth,
<taxonomicName id="4C4F4D50BF7A8303FDEB10D8FD711E98" box="[594,725,1986,2007]" class="Gastropoda" family="Hydrobiidae" genus="Tryonia" kingdom="Animalia" order="Littorinimorpha" pageId="13" pageNumber="14" phylum="Mollusca" rank="species" species="angosturae">
<emphasis id="B93BEAC1BF7A8303FDEB10D8FD711E98" box="[594,725,1986,2007]" italics="true" pageId="13" pageNumber="14">T. angosturae</emphasis>
</taxonomicName>
, USNM 115368. Scale bars AE = 1.0 mm, FI = 250 µm, J, M = 25 µm, KL, MO = 10 µm.
</paragraph>
</caption>
<subSubSection id="C3556558BF798300FF7E178DFD1F19B8" pageId="14" pageNumber="15" type="diagnosis">
<paragraph id="8BF036D3BF798300FF7E178DFD1F19B8" blockId="14.[151,1437,151,1076]" pageId="14" pageNumber="15">
<emphasis id="B93BEAC1BF798300FF7E178DFE9E19FF" bold="true" box="[199,314,151,176]" pageId="14" pageNumber="15">Diagnosis</emphasis>
. Shell medium-sized, conic; penis having two distal papillae on the inner edge and a basal papilla on the outer edge. Contrasted with
<taxonomicName id="4C4F4D50BF798300FE4517A7FDF1199B" box="[508,597,189,212]" class="Gastropoda" family="Hydrobiidae" genus="Tryonia" kingdom="Animalia" order="Littorinimorpha" pageId="14" pageNumber="15" phylum="Mollusca" rank="species" species="ovata">
<emphasis id="B93BEAC1BF798300FE4517A7FDF1199B" box="[508,597,189,212]" italics="true" pageId="14" pageNumber="15">T. ovata</emphasis>
</taxonomicName>
above. Differentiated from other Río Santa Maria basin congeners by having a basal papilla on the outer edge of the penis.
</paragraph>
</subSubSection>
<subSubSection id="C3556558BF798300FF7E1619FE081A88" pageId="14" pageNumber="15" type="description">
<paragraph id="8BF036D3BF798300FF7E1619FE6C18E3" blockId="14.[151,1437,151,1076]" pageId="14" pageNumber="15">
<emphasis id="B93BEAC1BF798300FF7E1619FEF51853" bold="true" box="[199,337,259,284]" pageId="14" pageNumber="15">Description</emphasis>
. Shell (
<figureCitation id="13742A56BF798300FE10161EFE4F1853" box="[425,491,260,284]" captionStart="FIGURE 7" captionStartId="13.[151,244,1839,1860]" captionTargetBox="[188,1404,151,1821]" captionTargetId="figure@13.[188,1404,145,1821]" captionTargetPageId="13" captionText="FIGURE 7. Tryonia ovata sp. nov. and Tryonia angosturae sp. nov. A. Holotype, T. ovata, USNM 873264. B. Male shell, T. ovata, UTEP 977. C. Holotype, T. angosturae, USNM 873259. D E. Shells, T. ovata, USNM 1153668 (E is a male). F G. Operculum (outer, inner sides), T. ovata, USNM 1153676. H I. Operculum (outer, inner sides), T. angosturae, USNM 1153688. J. Portion of radular ribbon, T. ovata, USNM 1153676. K. Central radular teeth, T. ovata, USNM 1153676. L. Lateral and inner marginal teeth, T. ovata, USNM 1153676. M. Portion of radular ribbon, T. angosturae, USNM 1153688. N. Central radular teeth, T. angosturae, USNM 1153688. O. Lateral and inner marginal teeth, T. angosturae, USNM 115368. Scale bars A E = 1.0 mm, F I = 250 µm, J, M = 25 µm, K L, M O = 10 µm." httpUri="https://zenodo.org/record/278449/files/figure.png" pageId="14" pageNumber="15">Fig. 7</figureCitation>
CE) up to
<quantity id="4CB79B36BF798300FDD6161EFD611853" box="[623,709,260,284]" metricMagnitude="-3" metricUnit="m" metricValue="4.9" pageId="14" pageNumber="15" unit="mm" value="4.9">4.9 mm</quantity>
tall, large females having 5.08.0 whorls, spire height 85110% width of shell, male shells smaller than those of females. Teleoconch whorls weak to medium convex, evenly rounded, sutures impressed. Aperture weakly angled adapically, parietal lip usually complete, usually adnate, rarely very slightly disjunct, umbilicus absent. Outer lip thin, weakly prosocline. Sculpture of strong growth lines. Periostracum brown or tan.
</paragraph>
<paragraph id="8BF036D3BF798300FF7E16ADFA881B58" blockId="14.[151,1437,151,1076]" pageId="14" pageNumber="15">
<emphasis id="B93BEAC1BF798300FF7E16ADFE6D189F" bold="true" box="[199,457,439,464]" pageId="14" pageNumber="15">Shell measurements (</emphasis>
mean in parentheses): height
<quantity id="4CB79B36BF798300FCA016ADFC1A1880" box="[793,958,439,463]" metricMagnitude="-3" metricUnit="m" metricValue="3.025" metricValueMax="3.36" metricValueMin="2.69" pageId="14" pageNumber="15" unit="mm" value="3.025" valueMax="3.36" valueMin="2.69">2.693.36 mm</quantity>
(3.01), width
<quantity id="4CB79B36BF798300FBDA16ADFAAC1880" box="[1123,1288,439,463]" metricMagnitude="-3" metricUnit="m" metricValue="1.27" metricValueMax="1.38" metricValueMin="1.16" pageId="14" pageNumber="15" unit="mm" value="1.27" valueMax="1.38" valueMin="1.16">1.161.38 mm</quantity>
(1.25), body whorl height
<quantity id="4CB79B36BF798300FE9416C6FE7418BB" box="[301,464,476,500]" metricMagnitude="-3" metricUnit="m" metricValue="1.545" metricValueMax="1.68" metricValueMin="1.41" pageId="14" pageNumber="15" unit="mm" value="1.545" valueMax="1.68" valueMin="1.41">1.411.68 mm</quantity>
(1.55), body whorl width
<quantity id="4CB79B36BF798300FD4016C6FC3918BB" box="[761,925,476,500]" metricMagnitude="-3" metricUnit="m" metricValue="1.29" metricValueMax="1.45" metricValueMin="1.13" pageId="14" pageNumber="15" unit="mm" value="1.29" valueMax="1.45" valueMin="1.13">1.131.45 mm</quantity>
(1.26), aperture height
<quantity id="4CB79B36BF798300FB1E16C6FAEE18BB" box="[1191,1354,476,500]" metricMagnitude="-4" metricUnit="m" metricValue="8.25" metricValueMax="8.9" metricValueMin="7.6" pageId="14" pageNumber="15" unit="mm" value="0.825" valueMax="0.89" valueMin="0.76">0.760.89 mm</quantity>
(0.82), aperture width
<quantity id="4CB79B36BF798300FEF916E5FE471B58" box="[320,483,511,535]" metricMagnitude="-4" metricUnit="m" metricValue="7.35" metricValueMax="8.2" metricValueMin="6.5" pageId="14" pageNumber="15" unit="mm" value="0.735" valueMax="0.82" valueMin="0.65">0.650.82 mm</quantity>
(0.72), total number of whorls 5.256.75 (5.92) (USNM 1153668,
<emphasis id="B93BEAC1BF798300FB6D151BFB461B57" box="[1236,1250,513,536]" italics="true" pageId="14" pageNumber="15">n</emphasis>
= 15).
</paragraph>
<paragraph id="8BF036D3BF798300FF7E153EFC751B10" blockId="14.[151,1437,151,1076]" pageId="14" pageNumber="15">
<emphasis id="B93BEAC1BF798300FF7E153EFDA71B73" bold="true" box="[199,515,547,572]" pageId="14" pageNumber="15">
Measurements of
<typeStatus id="54F48871BF798300FE25153EFDA71B73" box="[412,515,548,572]" pageId="14" pageNumber="15" type="holotype">holotype</typeStatus>
</emphasis>
: height
<quantity id="4CB79B36BF798300FDD9153EFD631B73" box="[608,711,548,572]" metricMagnitude="-3" metricUnit="m" metricValue="2.76" pageId="14" pageNumber="15" unit="mm" value="2.76">2.76 mm</quantity>
, width
<quantity id="4CB79B36BF798300FCA0153EFC241B73" box="[793,896,548,572]" metricMagnitude="-3" metricUnit="m" metricValue="1.25" pageId="14" pageNumber="15" unit="mm" value="1.25">1.25 mm</quantity>
, body whorl height
<quantity id="4CB79B36BF798300FBD8153EFB6D1B73" box="[1121,1225,548,572]" metricMagnitude="-3" metricUnit="m" metricValue="1.49" pageId="14" pageNumber="15" unit="mm" value="1.49">1.49 mm</quantity>
, body whorl width
<quantity id="4CB79B36BF798300FF2E155DFF591B10" box="[151,253,583,607]" metricMagnitude="-3" metricUnit="m" metricValue="1.19" pageId="14" pageNumber="15" unit="mm" value="1.19">1.19 mm</quantity>
, aperture height
<quantity id="4CB79B36BF798300FE01155DFDBA1B10" box="[440,542,583,607]" metricMagnitude="-4" metricUnit="m" metricValue="8.2" pageId="14" pageNumber="15" unit="mm" value="0.82">0.82 mm</quantity>
, aperture width
<quantity id="4CB79B36BF798300FD6A155DFC9D1B10" box="[723,825,583,607]" metricMagnitude="-4" metricUnit="m" metricValue="6.8" pageId="14" pageNumber="15" unit="mm" value="0.68">0.68 mm</quantity>
, 5.75 whorls.
</paragraph>
<paragraph id="8BF036D3BF798300FF7E1576FB131A78" blockId="14.[151,1437,151,1076]" pageId="14" pageNumber="15">
Edges of last 0.5 operculum whorl sometimes weakly frilled on outer side (
<figureCitation id="13742A56BF798300FBA41576FBC61BCB" box="[1053,1122,620,644]" captionStart="FIGURE 7" captionStartId="13.[151,244,1839,1860]" captionTargetBox="[188,1404,151,1821]" captionTargetId="figure@13.[188,1404,145,1821]" captionTargetPageId="13" captionText="FIGURE 7. Tryonia ovata sp. nov. and Tryonia angosturae sp. nov. A. Holotype, T. ovata, USNM 873264. B. Male shell, T. ovata, UTEP 977. C. Holotype, T. angosturae, USNM 873259. D E. Shells, T. ovata, USNM 1153668 (E is a male). F G. Operculum (outer, inner sides), T. ovata, USNM 1153676. H I. Operculum (outer, inner sides), T. angosturae, USNM 1153688. J. Portion of radular ribbon, T. ovata, USNM 1153676. K. Central radular teeth, T. ovata, USNM 1153676. L. Lateral and inner marginal teeth, T. ovata, USNM 1153676. M. Portion of radular ribbon, T. angosturae, USNM 1153688. N. Central radular teeth, T. angosturae, USNM 1153688. O. Lateral and inner marginal teeth, T. angosturae, USNM 115368. Scale bars A E = 1.0 mm, F I = 250 µm, J, M = 25 µm, K L, M O = 10 µm." httpUri="https://zenodo.org/record/278449/files/figure.png" pageId="14" pageNumber="15">Fig. 7</figureCitation>
H); inner side of operculum smooth (
<figureCitation id="13742A56BF798300FF411595FE981BE8" box="[248,316,655,679]" captionStart="FIGURE 7" captionStartId="13.[151,244,1839,1860]" captionTargetBox="[188,1404,151,1821]" captionTargetId="figure@13.[188,1404,145,1821]" captionTargetPageId="13" captionText="FIGURE 7. Tryonia ovata sp. nov. and Tryonia angosturae sp. nov. A. Holotype, T. ovata, USNM 873264. B. Male shell, T. ovata, UTEP 977. C. Holotype, T. angosturae, USNM 873259. D E. Shells, T. ovata, USNM 1153668 (E is a male). F G. Operculum (outer, inner sides), T. ovata, USNM 1153676. H I. Operculum (outer, inner sides), T. angosturae, USNM 1153688. J. Portion of radular ribbon, T. ovata, USNM 1153676. K. Central radular teeth, T. ovata, USNM 1153676. L. Lateral and inner marginal teeth, T. ovata, USNM 1153676. M. Portion of radular ribbon, T. angosturae, USNM 1153688. N. Central radular teeth, T. angosturae, USNM 1153688. O. Lateral and inner marginal teeth, T. angosturae, USNM 115368. Scale bars A E = 1.0 mm, F I = 250 µm, J, M = 25 µm, K L, M O = 10 µm." httpUri="https://zenodo.org/record/278449/files/figure.png" pageId="14" pageNumber="15">
Fig.
<date id="FFF11013BF798300FE951595FE981BE8" box="[300,316,655,679]" pageId="14" pageNumber="15">7</date>
</figureCitation>
I). Radula (
<figureCitation id="13742A56BF798300FE011595FE5E1BE8" box="[440,506,655,679]" captionStart="FIGURE 7" captionStartId="13.[151,244,1839,1860]" captionTargetBox="[188,1404,151,1821]" captionTargetId="figure@13.[188,1404,145,1821]" captionTargetPageId="13" captionText="FIGURE 7. Tryonia ovata sp. nov. and Tryonia angosturae sp. nov. A. Holotype, T. ovata, USNM 873264. B. Male shell, T. ovata, UTEP 977. C. Holotype, T. angosturae, USNM 873259. D E. Shells, T. ovata, USNM 1153668 (E is a male). F G. Operculum (outer, inner sides), T. ovata, USNM 1153676. H I. Operculum (outer, inner sides), T. angosturae, USNM 1153688. J. Portion of radular ribbon, T. ovata, USNM 1153676. K. Central radular teeth, T. ovata, USNM 1153676. L. Lateral and inner marginal teeth, T. ovata, USNM 1153676. M. Portion of radular ribbon, T. angosturae, USNM 1153688. N. Central radular teeth, T. angosturae, USNM 1153688. O. Lateral and inner marginal teeth, T. angosturae, USNM 115368. Scale bars A E = 1.0 mm, F I = 250 µm, J, M = 25 µm, K L, M O = 10 µm." httpUri="https://zenodo.org/record/278449/files/figure.png" pageId="14" pageNumber="15">Fig. 7</figureCitation>
M
<collectingCountry id="F3587643BF798300FD9C1595FD9E1BE8" box="[549,570,655,679]" name="Iceland" pageId="14" pageNumber="15">O</collectingCountry>
): dorsal edge of central radular teeth weakly convex, basal tongue rounded Vshaped, median cusps distally pointed, parallel-sided proximally, lateral cusps foursix, basal cusps onethree (innermost larger) (
<figureCitation id="13742A56BF798300FECC15CDFE1F1BA0" box="[373,443,727,751]" captionStart="FIGURE 7" captionStartId="13.[151,244,1839,1860]" captionTargetBox="[188,1404,151,1821]" captionTargetId="figure@13.[188,1404,145,1821]" captionTargetPageId="13" captionText="FIGURE 7. Tryonia ovata sp. nov. and Tryonia angosturae sp. nov. A. Holotype, T. ovata, USNM 873264. B. Male shell, T. ovata, UTEP 977. C. Holotype, T. angosturae, USNM 873259. D E. Shells, T. ovata, USNM 1153668 (E is a male). F G. Operculum (outer, inner sides), T. ovata, USNM 1153676. H I. Operculum (outer, inner sides), T. angosturae, USNM 1153688. J. Portion of radular ribbon, T. ovata, USNM 1153676. K. Central radular teeth, T. ovata, USNM 1153676. L. Lateral and inner marginal teeth, T. ovata, USNM 1153676. M. Portion of radular ribbon, T. angosturae, USNM 1153688. N. Central radular teeth, T. angosturae, USNM 1153688. O. Lateral and inner marginal teeth, T. angosturae, USNM 115368. Scale bars A E = 1.0 mm, F I = 250 µm, J, M = 25 µm, K L, M O = 10 µm." httpUri="https://zenodo.org/record/278449/files/figure.png" pageId="14" pageNumber="15">Fig. 7</figureCitation>
N). Lateral teeth having twothree cusps on inner and threefour cusps on outer sides, length of outer wing about 180% width of cutting edge, central cusp pointed (
<figureCitation id="13742A56BF798300FC4315E6FB981A5B" box="[1018,1084,764,788]" captionStart="FIGURE 7" captionStartId="13.[151,244,1839,1860]" captionTargetBox="[188,1404,151,1821]" captionTargetId="figure@13.[188,1404,145,1821]" captionTargetPageId="13" captionText="FIGURE 7. Tryonia ovata sp. nov. and Tryonia angosturae sp. nov. A. Holotype, T. ovata, USNM 873264. B. Male shell, T. ovata, UTEP 977. C. Holotype, T. angosturae, USNM 873259. D E. Shells, T. ovata, USNM 1153668 (E is a male). F G. Operculum (outer, inner sides), T. ovata, USNM 1153676. H I. Operculum (outer, inner sides), T. angosturae, USNM 1153688. J. Portion of radular ribbon, T. ovata, USNM 1153676. K. Central radular teeth, T. ovata, USNM 1153676. L. Lateral and inner marginal teeth, T. ovata, USNM 1153676. M. Portion of radular ribbon, T. angosturae, USNM 1153688. N. Central radular teeth, T. angosturae, USNM 1153688. O. Lateral and inner marginal teeth, T. angosturae, USNM 115368. Scale bars A E = 1.0 mm, F I = 250 µm, J, M = 25 µm, K L, M O = 10 µm." httpUri="https://zenodo.org/record/278449/files/figure.png" pageId="14" pageNumber="15">Fig. 7</figureCitation>
<collectingCountry id="F3587643BF798300FB8515E6FBF71A5B" box="[1084,1107,764,788]" name="Iceland" pageId="14" pageNumber="15">O</collectingCountry>
). Inner marginal teeth having 1419 cusps, outer marginal teeth having 2738 cusps. Radula data are from USNM 1153668.
</paragraph>
<paragraph id="8BF036D3BF798300FF7E145EFE081A88" blockId="14.[151,1437,151,1076]" pageId="14" pageNumber="15">
Animal darkly pigmented. Penis (
<figureCitation id="13742A56BF798300FDFC145EFD231A13" box="[581,647,836,860]" captionStart="FIGURE 5" captionStartId="10.[151,244,1857,1878]" captionTargetBox="[241,1348,158,1832]" captionTargetId="figure@10.[229,1363,145,1846]" captionTargetPageId="10" captionText="FIGURE 5. Penes. A. T. peregrina sp. nov., USNM 1153677. B. T. zaragozae sp. nov., USNM 1153679. C. T. ovata sp. nov., USNM 1153676. D. T. angosturae sp. nov., USNM 874174. E. T. molinae sp. nov., USNM 1153674. F. T. contrerasi sp. nov., USNM 1153670. G. T. taylori sp. nov., USNM 874047. H. T. taylori sp. nov., USNM 873252. I. T. metcalfi sp. nov., USNM 1153672. J. T. chuviscarae sp. nov., USNM 1001756. K. T. minckleyi sp. nov., USNM 1153673. L. T. julimesensis sp. nov., USNM 1153671. M. T. allendae sp. nov., USNM 1153667. N. T. oasiensis sp. nov., USNM 1153675. O. T. seemani, USNM 874125. P. T. seemani, USNM 874123. Scale bars = 250 µm. Db = distal bulb, Pa = papilla, Pd = penial duct, St = stylet." httpUri="https://zenodo.org/record/278447/files/figure.png" pageId="14" pageNumber="15">Fig. 5</figureCitation>
D) having two distal papillae on the inner edge and a basal papilla on the outer edge (all
<specimenCount id="9D49FD5ABF798300FED6147DFDAD1A30" box="[367,521,871,895]" pageId="14" pageNumber="15" type="generic">36 specimens</specimenCount>
from three samples). Distal bulb of penis expanded laterally on inner side, dark brown; stylet small. Penial duct undulating along most of length. Penial data are from USNM 873293, USNM 874174, USNM 905267.
</paragraph>
</subSubSection>
<subSubSection id="C3556558BF798300FF7E14C9FCF11D7B" pageId="14" pageNumber="15" type="distribution">
<paragraph id="8BF036D3BF798300FF7E14C9FCF11D7B" blockId="14.[151,1437,151,1076]" pageId="14" pageNumber="15">
<emphasis id="B93BEAC1BF798300FF7E14C9FE551AA3" bold="true" box="[199,497,979,1004]" pageId="14" pageNumber="15">Distribution and habitat.</emphasis>
<taxonomicName id="4C4F4D50BF798300FE4114CFFD6C1AA3" box="[504,712,981,1004]" class="Gastropoda" family="Hydrobiidae" genus="Tryonia" kingdom="Animalia" order="Littorinimorpha" pageId="14" pageNumber="15" phylum="Mollusca" rank="species" species="angostura">
<emphasis id="B93BEAC1BF798300FE4114CFFD6C1AA3" box="[504,712,981,1004]" italics="true" pageId="14" pageNumber="15">Tryonia angostura</emphasis>
</taxonomicName>
is endemic to the
<typeStatus id="54F48871BF798300FC2D14CFFC601AA2" box="[916,964,981,1005]" pageId="14" pageNumber="15">type</typeStatus>
locality area in the Río Santa Maria basin (
<figureCitation id="13742A56BF798300FF2614EDFF451D40" box="[159,225,1015,1039]" captionStart="FIGURE 1" captionStartId="2.[151,244,1795,1816]" captionTargetBox="[179,1402,157,1789]" captionTargetId="figure@2.[179,1423,125,1806]" captionTargetPageId="2" captionText="FIGURE 1. Map showing the distribution of Tryonia species in the Chihuahuan Desert. 1. Tryonia hertleini, Las Palomas. 2. Tryonia peregrina sp. nov., Ojo Vareleño, northwest of Casas Grandes. 3. Tryonia peregrina sp. nov., Yepomera. 4. Tryonia zaragozae sp. nov., northeast of Ignacio Zaragoza. 5. Tryonia angostura sp. nov., T. ovata sp. nov., south of Galeana. 6. Tryonia molinae sp. nov., El Molino. 7. Tryonia taylori sp. nov., vicinity of Rancho Ojo Caliente. 8. Tryonia contrerasi sp. nov., vicinity of Rancho Nuevo. 9. Tryonia chuviscarae sp. nov., T. minckleyi sp. nov., Balneario de San Diego de Alcala. 10. Tryonia julimesensis sp. nov., south of Julimes. 11. Tryonia allendae sp. nov., southwest of Talamantes. 12. Tryonia metcalfi sp. nov., La Cienega, south of Candelaria. 13. Tryonia oasiensis sp. nov., Caroline Spring, Oasis Ranch. 14. Tryonia cheatumi, Phantom Lake Spring. 15. Tryonia circumstriata, Diamond Y Draw. 16. Tryonia seemani, Ojo de Agua del Obispo, Durango City. 17. Tryonia seemani, Nombre de Dios. 18. Tryonia seemani, Amado Nervo." httpUri="https://zenodo.org/record/278443/files/figure.png" pageId="14" pageNumber="15">Fig. 1</figureCitation>
, locality 5). Habitat as for
<taxonomicName id="4C4F4D50BF798300FDB514E3FDC11D5F" box="[524,613,1017,1040]" class="Gastropoda" family="Hydrobiidae" genus="Tryonia" kingdom="Animalia" order="Littorinimorpha" pageId="14" pageNumber="15" phylum="Mollusca" rank="species" species="ovata">
<emphasis id="B93BEAC1BF798300FDB514E3FDC11D5F" box="[524,613,1017,1040]" italics="true" pageId="14" pageNumber="15">T. ovata</emphasis>
</taxonomicName>
(described above).
<taxonomicName id="4C4F4D50BF798300FCFB14E3FBB51D5F" box="[834,1041,1017,1040]" class="Gastropoda" family="Hydrobiidae" genus="Tryonia" kingdom="Animalia" order="Littorinimorpha" pageId="14" pageNumber="15" phylum="Mollusca" rank="species" species="angostura">
<emphasis id="B93BEAC1BF798300FCFB14E3FBB51D5F" box="[834,1041,1017,1040]" italics="true" pageId="14" pageNumber="15">Tryonia angostura</emphasis>
</taxonomicName>
has also been collected in the main spring canal (from Ojos de Arrey) at the Highway 10 crossing.
</paragraph>
</subSubSection>
</treatment>
</document>