217 lines
32 KiB
XML
217 lines
32 KiB
XML
<document id="993EC5A5B3B23E0F8DB7A917659AE503" ID-DOI="10.5281/zenodo.278442" ID-GBIF-Dataset="6b9505e0-ac6e-4baa-9066-6f729d38e9e2" ID-ISSN="1175-5326" ID-Zenodo-Dep="278442" IM.bibliography_approvedBy="felipe" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1460341493798" checkinUser="plazi" docAuthor="Hershler, Robert, Liu, Hsiu-Ping & Landye, Jerry" docDate="2011" docId="03E687C5BF6E8317FF2E178DFE761C3E" docLanguage="en" docName="zt03001p032.pdf" docOrigin="Zootaxa 3001" docStyle="DocumentStyle:890A69B780ED73D6DB8551B71C8AC79E.4:Zootaxa.2009-2012.journal_article" docStyleId="890A69B780ED73D6DB8551B71C8AC79E" docStyleName="Zootaxa.2009-2012.journal_article" docStyleVersion="4" docTitle="Tryonia allendae Hershler, Liu & Landye, 2011, sp. nov." docType="treatment" docVersion="8" lastPageNumber="26" masterDocId="FFDFFFBDBF77830EFFB9171AFFA4194F" masterDocTitle="New species and records of springsnails (Caenogastropoda: Cochliopidae: Tryonia) from the Chihuahuan Desert (Mexico and United States), an imperiled biodiversity hotspot" masterLastPageNumber="32" masterPageNumber="1" pageNumber="26" updateTime="1698261240372" updateUser="plazi">
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<mods:titleInfo id="2C8773B50FED87031D9035DF62397295">
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<mods:title id="13F0D32F7E69EF9836EDEB43917CC18C">New species and records of springsnails (Caenogastropoda: Cochliopidae: Tryonia) from the Chihuahuan Desert (Mexico and United States), an imperiled biodiversity hotspot</mods:title>
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<mods:name id="C22137C71F034360181DFE502406A1AA" type="personal">
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<mods:namePart id="E246638FAFFC6F3ED1991582BB5A9821">Hershler, Robert</mods:namePart>
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<mods:roleTerm id="E3A8B99EDAFCB7E483A50299D56CF910">Author</mods:roleTerm>
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<mods:namePart id="95F6DEBC9D9D39F827698A1EF3229DD1">Liu, Hsiu-Ping</mods:namePart>
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<mods:roleTerm id="28661FFD3382AB16D7B971CA097B474B">Author</mods:roleTerm>
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<mods:namePart id="6BD7498C55D47AD49B58E71AC4634B1D">Landye, Jerry</mods:namePart>
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<mods:typeOfResource id="11C1E9426586019908942E7C2EC27373">text</mods:typeOfResource>
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<mods:title id="9FD27E46B529F19E7678B4A77E5588F9">Zootaxa</mods:title>
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<mods:part id="80801AE6B2354DABA09AEB70B06415FC">
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<mods:date id="5FB478DC04726F518D36394AE0C58060">2011</mods:date>
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<mods:number id="5939AAF7DB0BF682FFEB3699CD5D0C92">3001</mods:number>
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<mods:classification id="CC2860C65BB9F6AE19DEDED8E197BC09">journal article</mods:classification>
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<mods:identifier id="4B9643BC64AA210DEA0E221A14DD4570" type="DOI">10.5281/zenodo.278442</mods:identifier>
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<mods:identifier id="5A71A9188C5E4223CF8DF813A550E4EA" type="GBIF-Dataset">6b9505e0-ac6e-4baa-9066-6f729d38e9e2</mods:identifier>
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<mods:identifier id="99832FD4A3BB1F90BE2BD30DBCC45814" type="ISSN">1175-5326</mods:identifier>
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<treatment id="03E687C5BF6E8317FF2E178DFE761C3E" ID-DOI="http://doi.org/10.5281/zenodo.5629213" ID-GBIF-Taxon="119406526" ID-Zenodo-Dep="5629213" LSID="urn:lsid:plazi:treatment:03E687C5BF6E8317FF2E178DFE761C3E" httpUri="http://treatment.plazi.org/id/03E687C5BF6E8317FF2E178DFE761C3E" lastPageNumber="26" pageId="25" pageNumber="26">
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<subSubSection id="C3556558BF6E8317FF2E178DFC5818CE" pageId="25" pageNumber="26" type="nomenclature">
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<paragraph id="8BF036D3BF6E8317FF2E178DFE6C19FE" blockId="25.[151,487,151,210]" box="[151,456,151,177]" pageId="25" pageNumber="26">
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<heading id="D0B881BFBF6E8317FF2E178DFE6C19FE" bold="true" box="[151,456,151,177]" fontSize="11" level="1" pageId="25" pageNumber="26" reason="1">
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<emphasis id="B93BEAC1BF6E8317FF2E178DFE6C19FE" bold="true" box="[151,456,151,177]" pageId="25" pageNumber="26">
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<taxonomicName id="4C4F4D50BF6E8317FF2E178DFEC019FE" box="[151,356,151,177]" class="Gastropoda" family="Hydrobiidae" genus="Tryonia" kingdom="Animalia" order="Littorinimorpha" pageId="25" pageNumber="26" phylum="Mollusca" rank="species" species="allendae" status="sp. nov.">
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<emphasis id="B93BEAC1BF6E8317FF2E178DFEC019FE" bold="true" box="[151,356,151,177]" italics="true" pageId="25" pageNumber="26">Tryonia allendae</emphasis>
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</taxonomicName>
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<taxonomicNameLabel id="A20857BABF6E8317FED2178DFE6C19FE" box="[363,456,151,177]" pageId="25" pageNumber="26" rank="species">sp. nov.</taxonomicNameLabel>
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</emphasis>
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</heading>
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</paragraph>
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<paragraph id="8BF036D3BF6E8317FF2E17A0FE43199D" blockId="25.[151,487,151,210]" box="[151,487,186,210]" pageId="25" pageNumber="26">
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(
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<figureCitation id="13742A56BF6E8317FF2617A0FF40199D" box="[159,228,186,210]" captionStart="FIGURE 5" captionStartId="10.[151,244,1857,1878]" captionTargetBox="[241,1348,158,1832]" captionTargetId="figure@10.[229,1363,145,1846]" captionTargetPageId="10" captionText="FIGURE 5. Penes. A. T. peregrina sp. nov., USNM 1153677. B. T. zaragozae sp. nov., USNM 1153679. C. T. ovata sp. nov., USNM 1153676. D. T. angosturae sp. nov., USNM 874174. E. T. molinae sp. nov., USNM 1153674. F. T. contrerasi sp. nov., USNM 1153670. G. T. taylori sp. nov., USNM 874047. H. T. taylori sp. nov., USNM 873252. I. T. metcalfi sp. nov., USNM 1153672. J. T. chuviscarae sp. nov., USNM 1001756. K. T. minckleyi sp. nov., USNM 1153673. L. T. julimesensis sp. nov., USNM 1153671. M. T. allendae sp. nov., USNM 1153667. N. T. oasiensis sp. nov., USNM 1153675. O. T. seemani, USNM 874125. P. T. seemani, USNM 874123. Scale bars = 250 µm. Db = distal bulb, Pa = papilla, Pd = penial duct, St = stylet." httpUri="https://zenodo.org/record/278447/files/figure.png" pageId="25" pageNumber="26">Figs 5</figureCitation>
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M, 11D–E, H–I, M–
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<collectingCountry id="F3587643BF6E8317FE7217A0FE7B199D" box="[459,479,186,210]" name="Iceland" pageId="25" pageNumber="26">O</collectingCountry>
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)
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</paragraph>
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<paragraph id="8BF036D3BF6E8317FF2E1619FCAF1857" blockId="25.[151,779,258,280]" box="[151,779,258,280]" pageId="25" pageNumber="26">
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<treatmentCitationGroup id="AB5F11FDBF6E8317FF2E1619FCAF1857" box="[151,779,258,280]" pageId="25" pageNumber="26">
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<taxonomicName id="4C4F4D50BF6E8317FF2E1619FF411857" box="[151,229,259,280]" class="Gastropoda" family="Hydrobiidae" genus="Tryonia" kingdom="Animalia" order="Littorinimorpha" pageId="25" pageNumber="26" phylum="Mollusca" rank="genus" status="sp. nov.">
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<emphasis id="B93BEAC1BF6E8317FF2E1619FF411857" box="[151,229,259,280]" italics="true" pageId="25" pageNumber="26">Tryonia</emphasis>
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</taxonomicName>
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<taxonomicNameLabel id="A20857BABF6E8317FF551619FE861857" box="[236,290,259,280]" pageId="25" pageNumber="26" rank="species">n. sp.</taxonomicNameLabel>
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4.—
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<treatmentCitation id="0AEE10C2BF6E8317FEEB1618FDFA1857" author="Hershler" box="[338,606,258,280]" page="1757" pageId="25" pageNumber="26" year="2005">
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<bibRefCitation id="EFDE4B22BF6E8317FEEB1618FD851857" author="Hershler" box="[338,545,258,280]" pageId="25" pageNumber="26" refString="Hershler, R., Mulvey, M. & Liu, H. - P. (2005) Genetic variation in the Desert springsnail (Tryonia porrecta): implications for reproductive mode and dispersal. Molecular Ecology, 14, 1755 - 1765." type="journal article" year="2005">
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Hershler
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<emphasis id="B93BEAC1BF6E8317FE0B1619FE7A1857" box="[434,478,258,280]" italics="true" pageId="25" pageNumber="26">et al</emphasis>
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. 2005
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</bibRefCitation>
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: 1757
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</treatmentCitation>
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(COI sequence).
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</treatmentCitationGroup>
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</paragraph>
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<paragraph id="8BF036D3BF6E8317FF2E165FFC5818CE" blockId="25.[151,1436,325,1393]" pageId="25" pageNumber="26">
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<typeStatus id="54F48871BF6E8317FF2E165FFF7E1812" box="[151,218,325,349]" pageId="25" pageNumber="26">
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<emphasis id="B93BEAC1BF6E8317FF2E165FFF7E1812" bold="true" box="[151,218,325,349]" pageId="25" pageNumber="26">Types</emphasis>
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</typeStatus>
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.
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<typeStatus id="54F48871BF6E8317FF55165FFEF31812" box="[236,343,325,349]" pageId="25" pageNumber="26" type="holotype">Holotype</typeStatus>
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, USNM 892115, El Ojito, southwest of Talamantes, Chihuahua,
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<geoCoordinate id="EE7B5014BF6E8317FB8E165FFB7C1812" box="[1079,1240,325,349]" direction="north" orientation="latitude" pageId="25" pageNumber="26" precision="1" value="26.9035">26º54'12.6" N</geoCoordinate>
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,
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<geoCoordinate id="EE7B5014BF6E8317FB5C165FFA3C1812" box="[1253,1432,325,349]" direction="west" orientation="longitude" pageId="25" pageNumber="26" precision="1" value="-105.4555">105º27'19.8" W</geoCoordinate>
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, leg. R.H. and J.J.L.,
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<date id="FFF11013BF6E8317FEC51673FDA518CE" box="[380,513,361,385]" pageId="25" pageNumber="26" value="1998-12-10">10/xii/1998</date>
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.
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<typeStatus id="54F48871BF6E8317FDB21670FDDC18CD" box="[523,632,362,386]" pageId="25" pageNumber="26" type="paratype">Paratypes</typeStatus>
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(from same lot), USNM 1153667.
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</paragraph>
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</subSubSection>
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<subSubSection id="C3556558BF6E8317FF7E1697FA3418EA" box="[199,1424,397,422]" pageId="25" pageNumber="26" type="etymology">
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<paragraph id="8BF036D3BF6E8317FF7E1697FA3418EA" blockId="25.[151,1436,325,1393]" box="[199,1424,397,422]" pageId="25" pageNumber="26">
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<emphasis id="B93BEAC1BF6E8317FF7E1697FEE218EA" bold="true" box="[199,326,397,421]" pageId="25" pageNumber="26">Etymology</emphasis>
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. A geographic epithet referring to the municipality of Allende in which this species is distributed.
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</paragraph>
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</subSubSection>
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<subSubSection id="C3556558BF6E8317FF7E16AAFE3D1B7A" pageId="25" pageNumber="26" type="diagnosis">
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<paragraph id="8BF036D3BF6E8317FF7E16AAFE3D1B7A" blockId="25.[151,1436,325,1393]" pageId="25" pageNumber="26">
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<emphasis id="B93BEAC1BF6E8317FF7E16AAFE9E1886" bold="true" box="[199,314,432,457]" pageId="25" pageNumber="26">Diagnosis</emphasis>
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. Shell medium-sized, ovate-conic or conic; penis having two distal papillae on the inner edge and a basal papilla on the outer edge. Contrasted with conchologically similar
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<taxonomicName id="4C4F4D50BF6E8317FC7F16CCFBEF18A2" box="[966,1099,469,493]" class="Gastropoda" family="Hydrobiidae" genus="Tryonia" kingdom="Animalia" order="Littorinimorpha" pageId="25" pageNumber="26" phylum="Mollusca" rank="species" species="minckleyi">
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<emphasis id="B93BEAC1BF6E8317FC7F16CCFBEF18A2" box="[966,1099,469,493]" italics="true" pageId="25" pageNumber="26">T. minckleyi</emphasis>
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</taxonomicName>
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above. Differentiated from
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<taxonomicName id="4C4F4D50BF6E8317FA3E16CCFEBA1B5E" class="Gastropoda" family="Hydrobiidae" genus="Tryonia" kingdom="Animalia" order="Littorinimorpha" pageId="25" pageNumber="26" phylum="Mollusca" rank="species" species="chuviscarae">
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<emphasis id="B93BEAC1BF6E8317FA3E16CCFEBA1B5E" italics="true" pageId="25" pageNumber="26">T. chuviscarae</emphasis>
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</taxonomicName>
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and
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<taxonomicName id="4C4F4D50BF6E8317FEE016E0FDA41B5E" box="[345,512,505,529]" class="Gastropoda" family="Hydrobiidae" genus="Tryonia" kingdom="Animalia" order="Littorinimorpha" pageId="25" pageNumber="26" phylum="Mollusca" rank="species" species="julimesensis">
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<emphasis id="B93BEAC1BF6E8317FEE016E0FDA41B5E" box="[345,512,505,529]" italics="true" pageId="25" pageNumber="26">T. julimesensis</emphasis>
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</taxonomicName>
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, which are also distributed in the Río Conchos basin, in having a more strongly undulating penial duct.
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</paragraph>
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</subSubSection>
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<subSubSection id="C3556558BF6E8317FF7E155AFBAF1DAE" pageId="25" pageNumber="26" type="description">
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<paragraph id="8BF036D3BF6E8317FF7E155AFDC91BA6" blockId="25.[151,1436,325,1393]" pageId="25" pageNumber="26">
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<emphasis id="B93BEAC1BF6E8317FF7E155AFEF51B16" bold="true" box="[199,337,576,601]" pageId="25" pageNumber="26">Description</emphasis>
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. Shell (
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<figureCitation id="13742A56BF6E8317FE0D155BFDA31B16" box="[436,519,577,601]" captionStart="FIGURE 11" captionStartId="24.[151,244,1770,1791]" captionTargetBox="[151,1436,151,1743]" captionTargetId="figure@24.[151,1436,151,1743]" captionTargetPageId="24" captionText="FIGURE 11. Tryonia julimesensis sp. nov. and Tryonia allendae sp. nov. A. Holotype, T. julimesensis, USNM 874130. B – C. Male shells, T. julimesensis, USNM 1153671. D. Holotype, T. allendae, USNM 892115. E. Shell, T. allendae, USNM 1153667. F – G. Operculum (outer, inner sides), T. julimesensis, USNM 1153671. H – I. Operculum (outer, inner sides), T. allendae, USNM 1153671. J. Portion of radular ribbon, T. julimesensis, USNM 1153671. K. Central radular teeth, T. julimesensis, USNM 1153671. L. Lateral tooth, T. julimesensis, USNM 1153671. M. Portion of radular ribbon, T. allendae, USNM 1153667. N. Central radular teeth, T. allendae, USNM 1153667. R. Lateral teeth, T. allendae, USNM 1153667. Scale bars A – E = 1.0 mm; F – I = 250 µm; J, K – L, N – O = 10 µm; M = 25 µm." httpUri="https://zenodo.org/record/278453/files/figure.png" pageId="25" pageNumber="26">Fig. 11</figureCitation>
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D–E) up to
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<quantity id="4CB79B36BF6E8317FD19155BFD581B16" box="[672,764,577,601]" metricMagnitude="-3" metricUnit="m" metricValue="5.8" pageId="25" pageNumber="26" unit="mm" value="5.8">5.8 mm</quantity>
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tall, large females having 5.0–5.75 whorls, spire height 110–140% width of shell, male shells smaller than those of females. Teleoconch whorls weak to medium convex, evenly rounded, sutures impressed. Aperture angled adapically, parietal lip complete, usually adnate, rarely slightly disjunct, umbilicus narrow or absent. Outer lip thin, orthocline. Sculpture of strong growth lines, weak spiral thread also sometimes present. Periostracum tan.
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</paragraph>
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<paragraph id="8BF036D3BF6E8317FF7E15EFFA881A1A" blockId="25.[151,1436,325,1393]" pageId="25" pageNumber="26">
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||
<emphasis id="B93BEAC1BF6E8317FF7E15EFFE6D1A41" bold="true" box="[199,457,757,782]" pageId="25" pageNumber="26">Shell measurements (</emphasis>
|
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mean in parentheses): height
|
||
<quantity id="4CB79B36BF6E8317FCA015EFFC1A1A42" box="[793,958,757,781]" metricMagnitude="-3" metricUnit="m" metricValue="3.175" metricValueMax="3.66" metricValueMin="2.69" pageId="25" pageNumber="26" unit="mm" value="3.175" valueMax="3.66" valueMin="2.69">2.69–3.66 mm</quantity>
|
||
(2.92), width
|
||
<quantity id="4CB79B36BF6E8317FBDA15EFFAAC1A42" box="[1123,1288,757,781]" metricMagnitude="-3" metricUnit="m" metricValue="1.58" metricValueMax="1.76" metricValueMin="1.4" pageId="25" pageNumber="26" unit="mm" value="1.58" valueMax="1.76" valueMin="1.4">1.40–1.76 mm</quantity>
|
||
(1.53), body whorl height
|
||
<quantity id="4CB79B36BF6E8317FE941403FE741A7E" box="[301,464,793,817]" metricMagnitude="-3" metricUnit="m" metricValue="1.92" metricValueMax="2.2" metricValueMin="1.64" pageId="25" pageNumber="26" unit="mm" value="1.92" valueMax="2.2" valueMin="1.64">1.64–2.20 mm</quantity>
|
||
(1.82), body whorl width
|
||
<quantity id="4CB79B36BF6E8317FD401403FC391A7E" box="[761,925,793,817]" metricMagnitude="-3" metricUnit="m" metricValue="1.45" metricValueMax="1.67" metricValueMin="1.23" pageId="25" pageNumber="26" unit="mm" value="1.45" valueMax="1.67" valueMin="1.23">1.23–1.67 mm</quantity>
|
||
(1.43), aperture height
|
||
<quantity id="4CB79B36BF6E8317FB1E1403FAEE1A7E" box="[1191,1354,793,817]" metricMagnitude="-3" metricUnit="m" metricValue="1.08" metricValueMax="1.22" metricValueMin="0.94" pageId="25" pageNumber="26" unit="mm" value="1.08" valueMax="1.22" valueMin="0.94">0.94–1.22 mm</quantity>
|
||
(1.06), aperture width
|
||
<quantity id="4CB79B36BF6E8317FEF91427FE471A1A" box="[320,483,829,853]" metricMagnitude="-4" metricUnit="m" metricValue="9.399999999999999" metricValueMax="10.6" metricValueMin="8.2" pageId="25" pageNumber="26" unit="mm" value="0.94" valueMax="1.06" valueMin="0.82">0.82–1.06 mm</quantity>
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||
(0.89), total number of whorls 5.00–5.75 (5.35) (USNM 1153667,
|
||
<emphasis id="B93BEAC1BF6E8317FB6D1424FB461A1A" box="[1236,1250,830,853]" italics="true" pageId="25" pageNumber="26">n</emphasis>
|
||
= 15).
|
||
</paragraph>
|
||
<paragraph id="8BF036D3BF6E8317FF7E1478FC671AD2" blockId="25.[151,1436,325,1393]" pageId="25" pageNumber="26">
|
||
<emphasis id="B93BEAC1BF6E8317FF7E1478FDA71A35" bold="true" box="[199,515,865,890]" pageId="25" pageNumber="26">
|
||
Measurements of
|
||
<typeStatus id="54F48871BF6E8317FE251478FDA71A35" box="[412,515,866,890]" pageId="25" pageNumber="26" type="holotype">holotype</typeStatus>
|
||
</emphasis>
|
||
: height
|
||
<quantity id="4CB79B36BF6E8317FDD9147BFD631A36" box="[608,711,865,889]" metricMagnitude="-3" metricUnit="m" metricValue="2.91" pageId="25" pageNumber="26" unit="mm" value="2.91">2.91 mm</quantity>
|
||
, width
|
||
<quantity id="4CB79B36BF6E8317FCA0147BFC241A36" box="[793,896,865,889]" metricMagnitude="-3" metricUnit="m" metricValue="1.44" pageId="25" pageNumber="26" unit="mm" value="1.44">1.44 mm</quantity>
|
||
, body whorl height
|
||
<quantity id="4CB79B36BF6E8317FBD8147BFB6D1A36" box="[1121,1225,865,889]" metricMagnitude="-3" metricUnit="m" metricValue="1.72" pageId="25" pageNumber="26" unit="mm" value="1.72">1.72 mm</quantity>
|
||
, body whorl width
|
||
<quantity id="4CB79B36BF6E8317FF2E149FFF591AD2" box="[151,253,901,925]" metricMagnitude="-3" metricUnit="m" metricValue="1.35" pageId="25" pageNumber="26" unit="mm" value="1.35">1.35 mm</quantity>
|
||
, aperture height
|
||
<quantity id="4CB79B36BF6E8317FE01149FFDBA1AD2" box="[440,542,901,925]" metricMagnitude="-4" metricUnit="m" metricValue="9.7" pageId="25" pageNumber="26" unit="mm" value="0.97">0.97 mm</quantity>
|
||
, aperture width
|
||
<quantity id="4CB79B36BF6E8317FD6A149FFC9D1AD2" box="[723,825,901,925]" metricMagnitude="-4" metricUnit="m" metricValue="8.299999999999999" pageId="25" pageNumber="26" unit="mm" value="0.83">0.83 mm</quantity>
|
||
, 5.5 whorls.
|
||
</paragraph>
|
||
<paragraph id="8BF036D3BF6E8317FF7E14B3FC171D3A" blockId="25.[151,1436,325,1393]" pageId="25" pageNumber="26">
|
||
Outer side of operculum smooth (
|
||
<figureCitation id="13742A56BF6E8317FDF114B3FD3C1A8E" box="[584,664,937,961]" captionStart="FIGURE 11" captionStartId="24.[151,244,1770,1791]" captionTargetBox="[151,1436,151,1743]" captionTargetId="figure@24.[151,1436,151,1743]" captionTargetPageId="24" captionText="FIGURE 11. Tryonia julimesensis sp. nov. and Tryonia allendae sp. nov. A. Holotype, T. julimesensis, USNM 874130. B – C. Male shells, T. julimesensis, USNM 1153671. D. Holotype, T. allendae, USNM 892115. E. Shell, T. allendae, USNM 1153667. F – G. Operculum (outer, inner sides), T. julimesensis, USNM 1153671. H – I. Operculum (outer, inner sides), T. allendae, USNM 1153671. J. Portion of radular ribbon, T. julimesensis, USNM 1153671. K. Central radular teeth, T. julimesensis, USNM 1153671. L. Lateral tooth, T. julimesensis, USNM 1153671. M. Portion of radular ribbon, T. allendae, USNM 1153667. N. Central radular teeth, T. allendae, USNM 1153667. R. Lateral teeth, T. allendae, USNM 1153667. Scale bars A – E = 1.0 mm; F – I = 250 µm; J, K – L, N – O = 10 µm; M = 25 µm." httpUri="https://zenodo.org/record/278453/files/figure.png" pageId="25" pageNumber="26">Fig. 11</figureCitation>
|
||
H), inner side smooth or with edges of last 0.5 whorl weakly frilled (
|
||
<figureCitation id="13742A56BF6E8317FF2614D7FF541AAA" box="[159,240,973,997]" captionStart="FIGURE 11" captionStartId="24.[151,244,1770,1791]" captionTargetBox="[151,1436,151,1743]" captionTargetId="figure@24.[151,1436,151,1743]" captionTargetPageId="24" captionText="FIGURE 11. Tryonia julimesensis sp. nov. and Tryonia allendae sp. nov. A. Holotype, T. julimesensis, USNM 874130. B – C. Male shells, T. julimesensis, USNM 1153671. D. Holotype, T. allendae, USNM 892115. E. Shell, T. allendae, USNM 1153667. F – G. Operculum (outer, inner sides), T. julimesensis, USNM 1153671. H – I. Operculum (outer, inner sides), T. allendae, USNM 1153671. J. Portion of radular ribbon, T. julimesensis, USNM 1153671. K. Central radular teeth, T. julimesensis, USNM 1153671. L. Lateral tooth, T. julimesensis, USNM 1153671. M. Portion of radular ribbon, T. allendae, USNM 1153667. N. Central radular teeth, T. allendae, USNM 1153667. R. Lateral teeth, T. allendae, USNM 1153667. Scale bars A – E = 1.0 mm; F – I = 250 µm; J, K – L, N – O = 10 µm; M = 25 µm." httpUri="https://zenodo.org/record/278453/files/figure.png" pageId="25" pageNumber="26">
|
||
Fig.
|
||
<date id="FFF11013BF6E8317FF6D14D7FF541AAA" box="[212,240,973,997]" pageId="25" pageNumber="26" value="1911-01">11</date>
|
||
</figureCitation>
|
||
I). Radula (
|
||
<figureCitation id="13742A56BF6E8317FED614D7FE181AAA" box="[367,444,973,997]" captionStart="FIGURE 11" captionStartId="24.[151,244,1770,1791]" captionTargetBox="[151,1436,151,1743]" captionTargetId="figure@24.[151,1436,151,1743]" captionTargetPageId="24" captionText="FIGURE 11. Tryonia julimesensis sp. nov. and Tryonia allendae sp. nov. A. Holotype, T. julimesensis, USNM 874130. B – C. Male shells, T. julimesensis, USNM 1153671. D. Holotype, T. allendae, USNM 892115. E. Shell, T. allendae, USNM 1153667. F – G. Operculum (outer, inner sides), T. julimesensis, USNM 1153671. H – I. Operculum (outer, inner sides), T. allendae, USNM 1153671. J. Portion of radular ribbon, T. julimesensis, USNM 1153671. K. Central radular teeth, T. julimesensis, USNM 1153671. L. Lateral tooth, T. julimesensis, USNM 1153671. M. Portion of radular ribbon, T. allendae, USNM 1153667. N. Central radular teeth, T. allendae, USNM 1153667. R. Lateral teeth, T. allendae, USNM 1153667. Scale bars A – E = 1.0 mm; F – I = 250 µm; J, K – L, N – O = 10 µm; M = 25 µm." httpUri="https://zenodo.org/record/278453/files/figure.png" pageId="25" pageNumber="26">Fig. 11</figureCitation>
|
||
M-O): dorsal edge of central radular teeth concave, basal tongue rounded, median cusps distally pointed, parallel-sided proximally, lateral cusps four–six, basal cusps one–two (innermost larger) (
|
||
<figureCitation id="13742A56BF6E8317FAD714EBFF161D62" captionStart="FIGURE 11" captionStartId="24.[151,244,1770,1791]" captionTargetBox="[151,1436,151,1743]" captionTargetId="figure@24.[151,1436,151,1743]" captionTargetPageId="24" captionText="FIGURE 11. Tryonia julimesensis sp. nov. and Tryonia allendae sp. nov. A. Holotype, T. julimesensis, USNM 874130. B – C. Male shells, T. julimesensis, USNM 1153671. D. Holotype, T. allendae, USNM 892115. E. Shell, T. allendae, USNM 1153667. F – G. Operculum (outer, inner sides), T. julimesensis, USNM 1153671. H – I. Operculum (outer, inner sides), T. allendae, USNM 1153671. J. Portion of radular ribbon, T. julimesensis, USNM 1153671. K. Central radular teeth, T. julimesensis, USNM 1153671. L. Lateral tooth, T. julimesensis, USNM 1153671. M. Portion of radular ribbon, T. allendae, USNM 1153667. N. Central radular teeth, T. allendae, USNM 1153667. R. Lateral teeth, T. allendae, USNM 1153667. Scale bars A – E = 1.0 mm; F – I = 250 µm; J, K – L, N – O = 10 µm; M = 25 µm." httpUri="https://zenodo.org/record/278453/files/figure.png" pageId="25" pageNumber="26">Fig. 11</figureCitation>
|
||
N). Lateral teeth having two–three cusps on inner and three–four cusps on outer sides, length of outer wing about 150% width of cutting edge, central cusp pointed (
|
||
<figureCitation id="13742A56BF6E8317FD681323FC861D1E" box="[721,802,1081,1105]" captionStart="FIGURE 11" captionStartId="24.[151,244,1770,1791]" captionTargetBox="[151,1436,151,1743]" captionTargetId="figure@24.[151,1436,151,1743]" captionTargetPageId="24" captionText="FIGURE 11. Tryonia julimesensis sp. nov. and Tryonia allendae sp. nov. A. Holotype, T. julimesensis, USNM 874130. B – C. Male shells, T. julimesensis, USNM 1153671. D. Holotype, T. allendae, USNM 892115. E. Shell, T. allendae, USNM 1153667. F – G. Operculum (outer, inner sides), T. julimesensis, USNM 1153671. H – I. Operculum (outer, inner sides), T. allendae, USNM 1153671. J. Portion of radular ribbon, T. julimesensis, USNM 1153671. K. Central radular teeth, T. julimesensis, USNM 1153671. L. Lateral tooth, T. julimesensis, USNM 1153671. M. Portion of radular ribbon, T. allendae, USNM 1153667. N. Central radular teeth, T. allendae, USNM 1153667. R. Lateral teeth, T. allendae, USNM 1153667. Scale bars A – E = 1.0 mm; F – I = 250 µm; J, K – L, N – O = 10 µm; M = 25 µm." httpUri="https://zenodo.org/record/278453/files/figure.png" pageId="25" pageNumber="26">Fig. 11</figureCitation>
|
||
<collectingCountry id="F3587643BF6E8317FC9B1323FC9D1D1E" box="[802,825,1081,1105]" name="Iceland" pageId="25" pageNumber="26">O</collectingCountry>
|
||
). Inner marginal teeth having 20–26 cusps, outer marginal teeth having 22–40 cusps. Radula data are from USNM 1153677.
|
||
</paragraph>
|
||
<paragraph id="8BF036D3BF6E8317FF7E139BFBAF1DAE" blockId="25.[151,1436,325,1393]" pageId="25" pageNumber="26">
|
||
Animal lightly pigmented. Penis (
|
||
<figureCitation id="13742A56BF6E8317FD86139BFD241DD6" box="[575,640,1153,1177]" captionStart="FIGURE 5" captionStartId="10.[151,244,1857,1878]" captionTargetBox="[241,1348,158,1832]" captionTargetId="figure@10.[229,1363,145,1846]" captionTargetPageId="10" captionText="FIGURE 5. Penes. A. T. peregrina sp. nov., USNM 1153677. B. T. zaragozae sp. nov., USNM 1153679. C. T. ovata sp. nov., USNM 1153676. D. T. angosturae sp. nov., USNM 874174. E. T. molinae sp. nov., USNM 1153674. F. T. contrerasi sp. nov., USNM 1153670. G. T. taylori sp. nov., USNM 874047. H. T. taylori sp. nov., USNM 873252. I. T. metcalfi sp. nov., USNM 1153672. J. T. chuviscarae sp. nov., USNM 1001756. K. T. minckleyi sp. nov., USNM 1153673. L. T. julimesensis sp. nov., USNM 1153671. M. T. allendae sp. nov., USNM 1153667. N. T. oasiensis sp. nov., USNM 1153675. O. T. seemani, USNM 874125. P. T. seemani, USNM 874123. Scale bars = 250 µm. Db = distal bulb, Pa = papilla, Pd = penial duct, St = stylet." httpUri="https://zenodo.org/record/278447/files/figure.png" pageId="25" pageNumber="26">Fig. 5</figureCitation>
|
||
M) having two distal papillae on inner edge and a basal papilla on outer edge (30 of
|
||
<specimenCount id="9D49FD5ABF6E8317FEA713BFFE181DF2" box="[286,444,1189,1213]" pageId="25" pageNumber="26" type="generic">30 specimens</specimenCount>
|
||
). Distal bulb of penis expanded laterally on inner side, black. Penial duct strongly undulating along most of length; stylet small. Penial data are from USNM 1153667.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C3556558BF6E8317FF7E13F7FE761C3E" pageId="25" pageNumber="26" type="distribution">
|
||
<paragraph id="8BF036D3BF6E8317FF7E13F7FE761C3E" blockId="25.[151,1436,325,1393]" pageId="25" pageNumber="26">
|
||
<emphasis id="B93BEAC1BF6E8317FF7E13F7FE4E1C49" bold="true" box="[199,490,1261,1286]" pageId="25" pageNumber="26">Distribution and habitat</emphasis>
|
||
. Endemic to a single site (El Ojito) in an upper reach of the Río Conchos watershed (
|
||
<figureCitation id="13742A56BF6E8317FF26120BFF401C66" box="[159,228,1297,1321]" captionStart="FIGURE 1" captionStartId="2.[151,244,1795,1816]" captionTargetBox="[179,1402,157,1789]" captionTargetId="figure@2.[179,1423,125,1806]" captionTargetPageId="2" captionText="FIGURE 1. Map showing the distribution of Tryonia species in the Chihuahuan Desert. 1. Tryonia hertleini, Las Palomas. 2. Tryonia peregrina sp. nov., Ojo Vareleño, northwest of Casas Grandes. 3. Tryonia peregrina sp. nov., Yepomera. 4. Tryonia zaragozae sp. nov., northeast of Ignacio Zaragoza. 5. Tryonia angostura sp. nov., T. ovata sp. nov., south of Galeana. 6. Tryonia molinae sp. nov., El Molino. 7. Tryonia taylori sp. nov., vicinity of Rancho Ojo Caliente. 8. Tryonia contrerasi sp. nov., vicinity of Rancho Nuevo. 9. Tryonia chuviscarae sp. nov., T. minckleyi sp. nov., Balneario de San Diego de Alcala. 10. Tryonia julimesensis sp. nov., south of Julimes. 11. Tryonia allendae sp. nov., southwest of Talamantes. 12. Tryonia metcalfi sp. nov., La Cienega, south of Candelaria. 13. Tryonia oasiensis sp. nov., Caroline Spring, Oasis Ranch. 14. Tryonia cheatumi, Phantom Lake Spring. 15. Tryonia circumstriata, Diamond Y Draw. 16. Tryonia seemani, Ojo de Agua del Obispo, Durango City. 17. Tryonia seemani, Nombre de Dios. 18. Tryonia seemani, Amado Nervo." httpUri="https://zenodo.org/record/278443/files/figure.png" pageId="25" pageNumber="26">Fig. 1</figureCitation>
|
||
, locality 11). The springs at this locality have been impounded with a concrete dam for recreational purposes (
|
||
<bibRefCitation id="EFDE4B22BF6E8317FF5D122FFE6E1C02" author="Edwards" box="[228,458,1333,1357]" pageId="25" pageNumber="26" refString="Edwards, R. J., Garrett, G. P. & Marsh-Matthews, E. (2003) Fish assemblages of the northern Chihuahuan Desert: status and conservation. In: Garrett, G. P. & Allan, N. L. (Eds), Aquatic Fauna of the Northern Chihuahuan Desert. Museum of Texas Tech University Special Publications 46, Lubbock, pp. 75 - 89." type="journal article" year="2003">
|
||
Edwards
|
||
<emphasis id="B93BEAC1BF6E8317FEF7122CFEDB1C02" box="[334,383,1333,1357]" italics="true" pageId="25" pageNumber="26">et al</emphasis>
|
||
. 2003
|
||
</bibRefCitation>
|
||
).
|
||
<taxonomicName id="4C4F4D50BF6E8317FE64122CFD381C02" box="[477,668,1333,1357]" class="Gastropoda" family="Hydrobiidae" genus="Tryonia" kingdom="Animalia" order="Littorinimorpha" pageId="25" pageNumber="26" phylum="Mollusca" rank="species" species="allendae">
|
||
<emphasis id="B93BEAC1BF6E8317FE64122CFD381C02" box="[477,668,1333,1357]" italics="true" pageId="25" pageNumber="26">Tryonia allendae</emphasis>
|
||
</taxonomicName>
|
||
was abundant (on rootlets and dead leaves) in the outflow (25°C) of the impoundment (
|
||
<figureCitation id="13742A56BF6E8317FED21243FE0A1C3E" box="[363,430,1369,1393]" captionStart="FIGURE 6" captionStartId="11.[151,244,1710,1731]" captionTargetBox="[221,1371,156,1678]" captionTargetId="figure@11.[212,1381,149,1685]" captionTargetPageId="11" captionText="FIGURE 6. Photographs of habitats of species treated in this paper. A. Ojo Vareleño, Chihuahua, type locality of T. peregrina sp. nov. (photograph taken on 16 / ix / 1990). B. Main spring in Yepomera, Chihuahua, T. peregrina habitat (18 / ix / 1990). C. Springs northeast of Ignacio Zaragoza, Chihuaua, type locality of T. zaragozae sp. nov. (8 / xii / 1998). D. Western-most head spring, Ojos de Arrey, Chihuahua, type locality of T. angosturae sp. nov. and T. ovata sp. nov. (7 / xii / 1998). E. Spring in northern part of El Molino, Chihuahua, type locality of T. molinae sp. nov. (26 / iv / 2010). F. Ojo del Apache, Chihuahua, type locality of T. contrerasi sp. nov. (14 / xii / 1998). G. Ojo Carbonero, Chihuahua, T. contrerasi habitat (2 / iv / 1991). H. Outflow of Ojo Caliente, Chihuahua, type locality of T. taylori sp. nov. (7 / xii / 1998). I – J. El Ojo de Servin, Chihuahua, T. taylori habitat (27 - viii- 1971, 7 / xii / 1998, respectively). K. Seeps in La Cienega, Texas, type locality of T. metcalfi sp. nov. (6 / xii / 2001). L – M. Balneario de San Diego de Alcala, Chihuahua, type locality of T. chuviscarae sp. nov. and T. minckleyi sp. nov. (4 / xii / 2001). N. Springs southeast of Julimes, Chihuahua, type locality of T. julimesensis sp. nov. (5 / iv / 1991). O. El Ojito (outflow), southwest of Talamantes, Chihuahua, type locality of T. allendae sp. nov. (10 / xii / 1998). P. Caroline Spring, Oasis Ranch, Texas, type locality of T. oasiensis sp. nov. (30 / iii / 2009). Q. Spring in Nombre de Dios, Durango, T. seemani habitat (2 / ix / 1971). R. Spring in Amado Nervo, Durango, T. seemani habitat (7 / iv / 1991)." httpUri="https://zenodo.org/record/278448/files/figure.png" pageId="25" pageNumber="26">Fig. 6</figureCitation>
|
||
<collectingCountry id="F3587643BF6E8317FE171243FE611C3E" box="[430,453,1369,1393]" name="Iceland" pageId="25" pageNumber="26">O</collectingCountry>
|
||
).
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |