treatments-xml/data/03/E6/20/03E62055FF9E0004FF7BFB22FF1F358B.xml

<document id="36217024E7C9EA78290E8B50F88F4F31" ID-DOI="10.11646/zootaxa.5323.2.1" ID-ISSN="1175-5326" ID-Zenodo-Dep="8204101" ID-ZooBank="0ABC4CFF-9D2A-449A-B7C2-CFE16DF16004" IM.bibliography_approvedBy="felipe" IM.illustrations_approvedBy="felipe" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" IM.treatmentCitations_approvedBy="felipe" IM.treatments_approvedBy="felipe" checkinTime="1690887012857" checkinUser="plazi" docAuthor="Soares, Matheus M. M., Runyon, Justin B., Capellari, Renato S. &amp; Ale-Rocha, Rosaly" docDate="2023" docId="03E62055FF9E0004FF7BFB22FF1F358B" docLanguage="en" docName="zootaxa.5323.2.1.pdf" docOrigin="Zootaxa 5323 (2)" docSource="http://dx.doi.org/10.11646/zootaxa.5323.2.1" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Paraclius brooksi Soares, Runyon &amp; Capellari 2023, sp. nov." docType="treatment" docVersion="3" lastPageNumber="169" masterDocId="FFDF582DFF910016FFECFFBDFFDB3247" masterDocTitle="Review of Paraclius arcuatus species-group (Diptera, Dolichopodidae) with description of two new species" masterLastPageNumber="182" masterPageNumber="151" pageNumber="166" updateTime="1691081218152" updateUser="valdenar" zenodo-license-document="CLOSED">
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<mods:title id="45D4270EC95285569DBE8210A672DEAE">Review of Paraclius arcuatus species-group (Diptera, Dolichopodidae) with description of two new species</mods:title>
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<mods:namePart id="B14216210D4824D8822EBF685CE5BC9B">Soares, Matheus M. M.</mods:namePart>
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<mods:affiliation id="29D527721F7609EE1B12185CC23DDF89">Graduate Program in Entomology (PPG-Ent), Instituto Nacional de Pesquisas da Amazônia (INPA). &amp; matheusmmsoares @ gmail. com; https: // orcid. org / 0000 - 0002 - 2355 - 1441</mods:affiliation>
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<mods:namePart id="7EFE90E4F32715DFACB4C4BEA35ABE5E">Runyon, Justin B.</mods:namePart>
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<mods:affiliation id="524424F3870E5D44B1FA3346B228C447">Rocky Mountain Research Station, USDA Forest Service, 1648 S. 7 th Avenue, Bozeman, Montana 59717, USA. &amp; justin. runyon @ usda. gov; https: // orcid. org / 0000 - 0002 - 0271 - 0511</mods:affiliation>
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<mods:namePart id="B51F561861ECBA0C73A416EF7810E6AD">Capellari, Renato S.</mods:namePart>
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<mods:affiliation id="54C5595EA647C379CAF676526E09E20F">Instituto Federal do Triângulo Mineiro, Campus Uberaba, Rua João Batista Ribeiro 4000, Distrito Industrial II, Uberaba, Minas Gerais 38064 - 790, Brazil. &amp; rscapellari @ gmail. com; https: // orcid. org / 0000 - 0002 - 8410 - 9235</mods:affiliation>
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<mods:namePart id="0F7BEBB96683F6CF0152502290D58FFD">Ale-Rocha, Rosaly</mods:namePart>
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<mods:affiliation id="B35D0310CE4732BB897DC28F92F83C76">Coordenação de Biodiversidade (COBIO, INPA), Av. André Araújo, 2936, Petrópolis, CEP 69067 - 375, Manaus, Amazonas, Brazil. &amp; alerocha @ inpa. gov. br; https: // orcid. org / 0000 - 0001 - 9874 - 9770</mods:affiliation>
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<treatment id="03E62055FF9E0004FF7BFB22FF1F358B" ID-DOI="http://doi.org/10.5281/zenodo.8204045" ID-Zenodo-Dep="8204045" LSID="urn:lsid:plazi:treatment:03E62055FF9E0004FF7BFB22FF1F358B" httpUri="http://treatment.plazi.org/id/03E62055FF9E0004FF7BFB22FF1F358B" lastPageId="18" lastPageNumber="169" pageId="15" pageNumber="166">
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<paragraph id="8BF09143FF9E0019FF7BFB22FCE236FD" blockId="15.[151,825,1183,1246]" box="[151,825,1183,1210]" pageId="15" pageNumber="166">
<heading id="D0B8262FFF9E0019FF7BFB22FCE236FD" bold="true" box="[151,825,1183,1210]" fontSize="11" level="1" pageId="15" pageNumber="166" reason="1">
<emphasis id="B93B4D51FF9E0019FF7BFB22FCE236FD" bold="true" box="[151,825,1183,1210]" pageId="15" pageNumber="166">
<taxonomicName id="4C4FEAC0FF9E0019FF7BFB22FD0F36FE" authority="Soares, Runyon &amp; Capellari" authorityName="Soares, Runyon &amp; Capellari" authorityYear="2023" box="[151,724,1183,1210]" class="Insecta" family="Dolichopodidae" genus="Paraclius" kingdom="Animalia" order="Diptera" pageId="15" pageNumber="166" phylum="Arthropoda" rank="species" species="brooksi" status="sp. nov.">
<emphasis id="B93B4D51FF9E0019FF7BFB22FEB736FD" bold="true" box="[151,364,1183,1210]" italics="true" pageId="15" pageNumber="166">Paraclius brooksi</emphasis>
Soares, Runyon &amp; Capellari
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<taxonomicNameLabel id="A208F02AFF9E0019FD30FB1DFCE236FD" box="[732,825,1184,1210]" pageId="15" pageNumber="166" rank="species">sp. nov.</taxonomicNameLabel>
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<subSubSection id="C355C2C8FF9E0019FF7BFB7EFE8B3699" box="[151,336,1219,1246]" pageId="15" pageNumber="166" type="description">
<paragraph id="8BF09143FF9E0019FF7BFB7EFE8B3699" blockId="15.[151,825,1183,1246]" box="[151,336,1219,1246]" pageId="15" pageNumber="166">
(
<figureCitation id="13748DC6FF9E0019FF73FB7EFEC43699" box="[159,287,1219,1246]" captionStart-0="FIGURE 11" captionStart-1="FIGURE12" captionStart-2="FIGURE 13" captionStartId-0="17.[151,250,1597,1621]" captionStartId-1="18.[151,250,1647,1671]" captionStartId-2="19.[151,250,655,679]" captionTargetBox-0="[151,1436,181,1574]" captionTargetBox-1="[151,1436,237,1624]" captionTargetBox-2="[151,1436,181,632]" captionTargetId-0="figure-18@17.[151,1436,181,1574]" captionTargetId-1="figure-42@18.[151,1436,237,1624]" captionTargetId-2="figure-18@19.[151,1436,181,632]" captionTargetPageId-0="17" captionTargetPageId-1="18" captionTargetPageId-2="19" captionText-0="FIGURE 11. Paraclius brooksi sp. nov., A–E male holotype (CNC), F male paratype (CNC). A. Habitus, lateral view; B. Thorax, dorsal view; C–D. Head anterior and lateral views, respectively; E. Abdomen and hypopygium, lateral view; F. Left wing." captionText-1="FIGURE12.Paraclius brooksisp.nov., male paratype (CNC).A. Abdomen, ventralview;B. Cerci, dorsalview;C. Hypopygium, left lateral view; D. Surstyli and apical edge of epandrium; E. Internal appendages; F. Hypandrium and phallus, ventral view; G. Tergite 7, dorsal view. Abreviations: a apv s = apical apicoventral epandrial seta; b apv s = basal apicoventral epandrial seta; cerc = cercus; dsur = dorsal lobe of surstylus; ej apod = ejaculatory apodeme; epand = epandrium; hypd = hypandrium; hypd pr = hypandrial process; pgt = postgonite; ph = phallus; ph pr = phallus process; prct = proctiger; sp = sperm pump; st = sternite; tg = tergite; vsur = ventral lobe of surstylus." captionText-2="FIGURE 13. Paraclius brooksi sp. nov, identified female from Mexico (CNC). A. Habitus, lateral view; B. Head, anterior view; C. Right antenna, outer view." figureDoi-0="http://doi.org/10.5281/zenodo.8204135" figureDoi-1="http://doi.org/10.5281/zenodo.8204139" figureDoi-2="http://doi.org/10.5281/zenodo.8204141" httpUri-0="https://zenodo.org/record/8204135/files/figure.png" httpUri-1="https://zenodo.org/record/8204139/files/figure.png" httpUri-2="https://zenodo.org/record/8204141/files/figure.png" pageId="15" pageNumber="166">Figs 11–13</figureCitation>
, 
<figureCitation id="13748DC6FF9E0019FEC7FB79FE933699" box="[299,328,1220,1246]" captionStart="FIGURE 23" captionStartId="29.[151,250,1033,1057]" captionTargetBox="[151,1436,237,1009]" captionTargetId="figure-36@29.[151,1436,237,1009]" captionTargetPageId="29" captionText="FIGURE 23. Known geographic distribution of Paraclius angusticauda Van Duzee, 1933, P. angustipennis Van Duzee, 1929, P. brooksi sp. nov., P. elongatus Van Duzee, 1930 and P. xibun sp. nov." figureDoi="http://doi.org/10.5281/zenodo.8204175" httpUri="https://zenodo.org/record/8204175/files/figure.png" pageId="15" pageNumber="166">23</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="C355C2C8FF9E0019FF7BFAB6FD0D342D" pageId="15" pageNumber="166" type="diagnosis">
<paragraph id="8BF09143FF9E0019FF7BFAB6FD0D342D" blockId="15.[151,1437,1291,2038]" pageId="15" pageNumber="166">
<emphasis id="B93B4D51FF9E0019FF7BFAB6FEB43762" bold="true" box="[151,367,1291,1317]" pageId="15" pageNumber="166">Diagnosis (males).</emphasis>
Postpedicel subrectangular, pointed dorsally at apex (
<figureCitation id="13748DC6FF9E0019FC2AFAB6FBF73761" box="[966,1068,1291,1318]" captionStart="FIGURE 11" captionStartId="17.[151,250,1597,1621]" captionTargetBox="[151,1436,181,1574]" captionTargetId="figure-18@17.[151,1436,181,1574]" captionTargetPageId="17" captionText="FIGURE 11. Paraclius brooksi sp. nov., A–E male holotype (CNC), F male paratype (CNC). A. Habitus, lateral view; B. Thorax, dorsal view; C–D. Head anterior and lateral views, respectively; E. Abdomen and hypopygium, lateral view; F. Left wing." figureDoi="http://doi.org/10.5281/zenodo.8204135" httpUri="https://zenodo.org/record/8204135/files/figure.png" pageId="15" pageNumber="166">Fig. 11D</figureCitation>
). Anepisternum with yellow spot above coxa I (
<figureCitation id="13748DC6FF9E0019FED5FA92FE7A370D" box="[313,417,1327,1354]" captionStart="FIGURE 11" captionStartId="17.[151,250,1597,1621]" captionTargetBox="[151,1436,181,1574]" captionTargetId="figure-18@17.[151,1436,181,1574]" captionTargetPageId="17" captionText="FIGURE 11. Paraclius brooksi sp. nov., A–E male holotype (CNC), F male paratype (CNC). A. Habitus, lateral view; B. Thorax, dorsal view; C–D. Head anterior and lateral views, respectively; E. Abdomen and hypopygium, lateral view; F. Left wing." figureDoi="http://doi.org/10.5281/zenodo.8204135" httpUri="https://zenodo.org/record/8204135/files/figure.png" pageId="15" pageNumber="166">Fig. 11A</figureCitation>
). Legs mostly yellow, except extreme base of coxa I, basal 2/3 of coxa II and basal 1/3 of coxa III, tarsi I and II from apex of tarsomere 1, apex of dorsal and posterior surfaces of femur III, apical 1/3 of tibia III and all of tarsus III brown (
<figureCitation id="13748DC6FF9E0019FDDCFACAFD4237D5" box="[560,665,1399,1426]" captionStart="FIGURE 11" captionStartId="17.[151,250,1597,1621]" captionTargetBox="[151,1436,181,1574]" captionTargetId="figure-18@17.[151,1436,181,1574]" captionTargetPageId="17" captionText="FIGURE 11. Paraclius brooksi sp. nov., A–E male holotype (CNC), F male paratype (CNC). A. Habitus, lateral view; B. Thorax, dorsal view; C–D. Head anterior and lateral views, respectively; E. Abdomen and hypopygium, lateral view; F. Left wing." figureDoi="http://doi.org/10.5281/zenodo.8204135" httpUri="https://zenodo.org/record/8204135/files/figure.png" pageId="15" pageNumber="166">Fig. 11A</figureCitation>
). Wing brownish; M 
<subScript id="17CB9306FF9E0019FC68FA37FC5737DF" attach="left" box="[900,908,1418,1432]" fontSize="6" pageId="15" pageNumber="166">1</subScript>
strongly bowed towards R, weakly concave posteriorly; CuAx ratio: 0.9 (
<figureCitation id="13748DC6FF9E0019FE0EFA26FD9D37F1" box="[482,582,1435,1462]" captionStart="FIGURE 11" captionStartId="17.[151,250,1597,1621]" captionTargetBox="[151,1436,181,1574]" captionTargetId="figure-18@17.[151,1436,181,1574]" captionTargetPageId="17" captionText="FIGURE 11. Paraclius brooksi sp. nov., A–E male holotype (CNC), F male paratype (CNC). A. Habitus, lateral view; B. Thorax, dorsal view; C–D. Head anterior and lateral views, respectively; E. Abdomen and hypopygium, lateral view; F. Left wing." figureDoi="http://doi.org/10.5281/zenodo.8204135" httpUri="https://zenodo.org/record/8204135/files/figure.png" pageId="15" pageNumber="166">Fig. 11F</figureCitation>
). Male tergite 6 with 4–5 strong lateral setae (
<figureCitation id="13748DC6FF9E0019FBBEFA26FB6D37F1" box="[1106,1206,1435,1462]" captionStart="FIGURE 11" captionStartId="17.[151,250,1597,1621]" captionTargetBox="[151,1436,181,1574]" captionTargetId="figure-18@17.[151,1436,181,1574]" captionTargetPageId="17" captionText="FIGURE 11. Paraclius brooksi sp. nov., A–E male holotype (CNC), F male paratype (CNC). A. Habitus, lateral view; B. Thorax, dorsal view; C–D. Head anterior and lateral views, respectively; E. Abdomen and hypopygium, lateral view; F. Left wing." figureDoi="http://doi.org/10.5281/zenodo.8204135" httpUri="https://zenodo.org/record/8204135/files/figure.png" pageId="15" pageNumber="166">Fig. 11E</figureCitation>
). Basal apicoventral epandrial seta about 2X longer than apical seta, and slightly shorter than ventral lobe of surstylus (
<figureCitation id="13748DC6FF9E0019FAEFFA02FA4B379D" box="[1283,1424,1471,1498]" captionStart="FIGURE12" captionStartId="18.[151,250,1647,1671]" captionTargetBox="[151,1436,237,1624]" captionTargetId="figure-42@18.[151,1436,237,1624]" captionTargetPageId="18" captionText="FIGURE12.Paraclius brooksisp.nov., male paratype (CNC).A. Abdomen, ventralview;B. Cerci, dorsalview;C. Hypopygium, left lateral view; D. Surstyli and apical edge of epandrium; E. Internal appendages; F. Hypandrium and phallus, ventral view; G. Tergite 7, dorsal view. Abreviations: a apv s = apical apicoventral epandrial seta; b apv s = basal apicoventral epandrial seta; cerc = cercus; dsur = dorsal lobe of surstylus; ej apod = ejaculatory apodeme; epand = epandrium; hypd = hypandrium; hypd pr = hypandrial process; pgt = postgonite; ph = phallus; ph pr = phallus process; prct = proctiger; sp = sperm pump; st = sternite; tg = tergite; vsur = ventral lobe of surstylus." figureDoi="http://doi.org/10.5281/zenodo.8204139" httpUri="https://zenodo.org/record/8204139/files/figure.png" pageId="15" pageNumber="166">Fig. 12C, D</figureCitation>
). Ventral lobe of surstylus long and narrow, somewhat L-shaped, strongly curved ventrally at apex (
<figureCitation id="13748DC6FF9E0019FAEEFA5EFA4B37B9" box="[1282,1424,1507,1534]" captionStart="FIGURE12" captionStartId="18.[151,250,1647,1671]" captionTargetBox="[151,1436,237,1624]" captionTargetId="figure-42@18.[151,1436,237,1624]" captionTargetPageId="18" captionText="FIGURE12.Paraclius brooksisp.nov., male paratype (CNC).A. Abdomen, ventralview;B. Cerci, dorsalview;C. Hypopygium, left lateral view; D. Surstyli and apical edge of epandrium; E. Internal appendages; F. Hypandrium and phallus, ventral view; G. Tergite 7, dorsal view. Abreviations: a apv s = apical apicoventral epandrial seta; b apv s = basal apicoventral epandrial seta; cerc = cercus; dsur = dorsal lobe of surstylus; ej apod = ejaculatory apodeme; epand = epandrium; hypd = hypandrium; hypd pr = hypandrial process; pgt = postgonite; ph = phallus; ph pr = phallus process; prct = proctiger; sp = sperm pump; st = sternite; tg = tergite; vsur = ventral lobe of surstylus." figureDoi="http://doi.org/10.5281/zenodo.8204139" httpUri="https://zenodo.org/record/8204139/files/figure.png" pageId="15" pageNumber="166">Fig. 12C, D</figureCitation>
). Dorsal lobe of surstylus short and wide, somewhat subrectangular, narrowing at apical 1/3, with 1 short and strong seta at middle of dorsal edge (
<figureCitation id="13748DC6FF9E0019FE1CF996FD813401" box="[496,602,1579,1606]" captionStart="FIGURE12" captionStartId="18.[151,250,1647,1671]" captionTargetBox="[151,1436,237,1624]" captionTargetId="figure-42@18.[151,1436,237,1624]" captionTargetPageId="18" captionText="FIGURE12.Paraclius brooksisp.nov., male paratype (CNC).A. Abdomen, ventralview;B. Cerci, dorsalview;C. Hypopygium, left lateral view; D. Surstyli and apical edge of epandrium; E. Internal appendages; F. Hypandrium and phallus, ventral view; G. Tergite 7, dorsal view. Abreviations: a apv s = apical apicoventral epandrial seta; b apv s = basal apicoventral epandrial seta; cerc = cercus; dsur = dorsal lobe of surstylus; ej apod = ejaculatory apodeme; epand = epandrium; hypd = hypandrium; hypd pr = hypandrial process; pgt = postgonite; ph = phallus; ph pr = phallus process; prct = proctiger; sp = sperm pump; st = sternite; tg = tergite; vsur = ventral lobe of surstylus." figureDoi="http://doi.org/10.5281/zenodo.8204139" httpUri="https://zenodo.org/record/8204139/files/figure.png" pageId="15" pageNumber="166">Fig. 12D</figureCitation>
). Cercus oval, about 3X longer than wide; brownish, except base yellow; about 1/2 as long as epandrium (
<figureCitation id="13748DC6FF9E0019FDEFF9F2FDB1342D" box="[515,618,1615,1642]" captionStart="FIGURE 11" captionStartId="17.[151,250,1597,1621]" captionTargetBox="[151,1436,181,1574]" captionTargetId="figure-18@17.[151,1436,181,1574]" captionTargetPageId="17" captionText="FIGURE 11. Paraclius brooksi sp. nov., A–E male holotype (CNC), F male paratype (CNC). A. Habitus, lateral view; B. Thorax, dorsal view; C–D. Head anterior and lateral views, respectively; E. Abdomen and hypopygium, lateral view; F. Left wing." figureDoi="http://doi.org/10.5281/zenodo.8204135" httpUri="https://zenodo.org/record/8204135/files/figure.png" pageId="15" pageNumber="166">Figs 11E</figureCitation>
, 
<figureCitation id="13748DC6FF9E0019FD9AF9EDFD12342D" box="[630,713,1616,1642]" captionStart="FIGURE12" captionStartId="18.[151,250,1647,1671]" captionTargetBox="[151,1436,237,1624]" captionTargetId="figure-42@18.[151,1436,237,1624]" captionTargetPageId="18" captionText="FIGURE12.Paraclius brooksisp.nov., male paratype (CNC).A. Abdomen, ventralview;B. Cerci, dorsalview;C. Hypopygium, left lateral view; D. Surstyli and apical edge of epandrium; E. Internal appendages; F. Hypandrium and phallus, ventral view; G. Tergite 7, dorsal view. Abreviations: a apv s = apical apicoventral epandrial seta; b apv s = basal apicoventral epandrial seta; cerc = cercus; dsur = dorsal lobe of surstylus; ej apod = ejaculatory apodeme; epand = epandrium; hypd = hypandrium; hypd pr = hypandrial process; pgt = postgonite; ph = phallus; ph pr = phallus process; prct = proctiger; sp = sperm pump; st = sternite; tg = tergite; vsur = ventral lobe of surstylus." figureDoi="http://doi.org/10.5281/zenodo.8204139" httpUri="https://zenodo.org/record/8204139/files/figure.png" pageId="15" pageNumber="166">12B, C</figureCitation>
).
</paragraph>
</subSubSection>
<paragraph id="8BF09143FF9E0019FB53FA37FB0237DF" blockId="15.[151,1437,1291,2038]" box="[1215,1241,1418,1432]" pageId="15" pageNumber="166">
<subScript id="17CB9306FF9E0019FB53FA37FB0237DF" attach="left" box="[1215,1241,1418,1432]" fontSize="6" pageId="15" pageNumber="166">4+5</subScript>
</paragraph>
<subSubSection id="C355C2C8FF9E0006FF2BF9CEFDC9344E" lastPageId="16" lastPageNumber="167" pageId="15" pageNumber="166" type="description">
<paragraph id="8BF09143FF9E0006FF2BF9CEFDC9344E" blockId="15.[151,1437,1291,2038]" lastBlockId="16.[151,1437,151,1977]" lastPageId="16" lastPageNumber="167" pageId="15" pageNumber="166">
<emphasis id="B93B4D51FF9E0019FF2BF9CEFE7E34C9" bold="true" box="[199,421,1651,1678]" pageId="15" pageNumber="166">Description. Male.</emphasis>
Body length: 
<quantity id="4CB73CA6FF9E0019FDA5F9C9FD0F34C9" box="[585,724,1652,1678]" metricMagnitude="-3" metricUnit="m" metricValue="2.8499999999999996" metricValueMax="2.9" metricValueMin="2.8" pageId="15" pageNumber="166" unit="mm" value="2.8499999999999996" valueMax="2.9" valueMin="2.8">2.8–2.9 mm</quantity>
; wing length: 
<quantity id="4CB73CA6FF9E0019FC94F9CEFBD934C9" box="[888,1026,1651,1678]" metricMagnitude="-3" metricUnit="m" metricValue="2.7" metricValueMax="2.8" metricValueMin="2.6" pageId="15" pageNumber="166" unit="mm" value="2.7" valueMax="2.8" valueMin="2.6">2.6–2.8 mm</quantity>
, width: 
<quantity id="4CB73CA6FF9E0019FB8CF9C9FB6C34C9" box="[1120,1207,1652,1678]" metricMagnitude="-4" metricUnit="m" metricValue="8.0" pageId="15" pageNumber="166" unit="mm" value="0.8">0.8 mm</quantity>
(n = 4). 
<emphasis id="B93B4D51FF9E0019FAF2F9C9FA8534C9" bold="true" box="[1310,1374,1652,1678]" pageId="15" pageNumber="166">Head</emphasis>
(
<figureCitation id="13748DC6FF9E0019FA83F9CEFF3634F5" captionStart="FIGURE 11" captionStartId="17.[151,250,1597,1621]" captionTargetBox="[151,1436,181,1574]" captionTargetId="figure-18@17.[151,1436,181,1574]" captionTargetPageId="17" captionText="FIGURE 11. Paraclius brooksi sp. nov., A–E male holotype (CNC), F male paratype (CNC). A. Habitus, lateral view; B. Thorax, dorsal view; C–D. Head anterior and lateral views, respectively; E. Abdomen and hypopygium, lateral view; F. Left wing." figureDoi="http://doi.org/10.5281/zenodo.8204135" httpUri="https://zenodo.org/record/8204135/files/figure.png" pageId="15" pageNumber="166">Fig. 11A–D</figureCitation>
). Similar to 
<taxonomicName id="4C4FEAC0FF9E0019FE65F925FDD434F6" authorityName=": Loew" authorityYear="1864" box="[393,527,1688,1714]" class="Insecta" family="Dolichopodidae" genus="Paraclius" kingdom="Animalia" order="Diptera" pageId="15" pageNumber="166" phylum="Arthropoda" rank="species" species="arcuatus">
<emphasis id="B93B4D51FF9E0019FE65F925FE4134F6" box="[393,410,1688,1713]" italics="true" pageId="15" pageNumber="166">P</emphasis>
. 
<emphasis id="B93B4D51FF9E0019FE42F925FDD434F6" box="[430,527,1688,1713]" italics="true" pageId="15" pageNumber="166">arcuatus</emphasis>
</taxonomicName>
, except: Upper-most 5 postocular setae black, remaining setae white. Frons subrectangular 2.3X wider than high, metallic green, with weak bluish reflections. Face ground color pale brown, but obscured by dense silvery pruinosity, except short rectangular area below base of antennae metallic green with weak bluish reflections. Face gradually narrowing below, slightly wider than mid ocellus at narrowest point. Antenna dark brown; scape conical, dorsal surface covered with short black setae, 2–3 longer setae at outer surface; postpedicel subrectangular, pointed dorsally at apex, 1.7X longer than wide; arista-like stylus dorsal, arising at base of postpedicel, length about 4/6 of eye height, two-segmented, first segment short and arched, slightly overlapping apex of postpedicel, covered by short pubescence, second segment long, about 2X longer than first segment, covered by short microtrichia. 
<emphasis id="B93B4D51FF9E0019FE78F805FE363595" bold="true" box="[404,493,1976,2002]" pageId="15" pageNumber="166">Thorax</emphasis>
(
<figureCitation id="13748DC6FF9E0019FE11F80AFD5D3595" box="[509,646,1975,2002]" captionStart="FIGURE 11" captionStartId="17.[151,250,1597,1621]" captionTargetBox="[151,1436,181,1574]" captionTargetId="figure-18@17.[151,1436,181,1574]" captionTargetPageId="17" captionText="FIGURE 11. Paraclius brooksi sp. nov., A–E male holotype (CNC), F male paratype (CNC). A. Habitus, lateral view; B. Thorax, dorsal view; C–D. Head anterior and lateral views, respectively; E. Abdomen and hypopygium, lateral view; F. Left wing." figureDoi="http://doi.org/10.5281/zenodo.8204135" httpUri="https://zenodo.org/record/8204135/files/figure.png" pageId="15" pageNumber="166">Fig. 11A, B</figureCitation>
). Mesonotum mostly metallic green, with narrow acrostichal coppery stripe and weak bluish reflections. Metepimeron dark gray. 
<emphasis id="B93B4D51FF9E0019FCA5F866FC1235B2" box="[841,969,2011,2037]" italics="true" pageId="15" pageNumber="166">Chaetotaxy</emphasis>
: Scutellum with 1 pair of strong medial scutellars and 1 pair of smaller setae laterad, about 1/6 as long as medial scutellars; upper-surface of proepisternum with sparse fine setulae and 2 more conspicuous setae in front of anterior spiracle. 
<emphasis id="B93B4D51FF810006FBBAFF07FB4C3293" bold="true" box="[1110,1175,186,212]" pageId="16" pageNumber="167">Wing</emphasis>
(
<figureCitation id="13748DC6FF810006FB44FF06FAD63292" box="[1192,1293,187,213]" captionStart="FIGURE 11" captionStartId="17.[151,250,1597,1621]" captionTargetBox="[151,1436,181,1574]" captionTargetId="figure-18@17.[151,1436,181,1574]" captionTargetPageId="17" captionText="FIGURE 11. Paraclius brooksi sp. nov., A–E male holotype (CNC), F male paratype (CNC). A. Habitus, lateral view; B. Thorax, dorsal view; C–D. Head anterior and lateral views, respectively; E. Abdomen and hypopygium, lateral view; F. Left wing." figureDoi="http://doi.org/10.5281/zenodo.8204135" httpUri="https://zenodo.org/record/8204135/files/figure.png" pageId="16" pageNumber="167">Fig. 11F</figureCitation>
). Membrane brownish; M 
<subScript id="17CB9306FF810006FEC7FF4CFEE832B8" attach="left" box="[299,307,241,255]" fontSize="6" pageId="16" pageNumber="167">1</subScript>
strongly bowed towards R 
<subScript id="17CB9306FF810006FD82FF4CFD5332B8" attach="left" box="[622,648,241,255]" fontSize="6" pageId="16" pageNumber="167">4+5</subScript>
and weakly concave posteriorly; CuAx ratio: 0.9. 
<emphasis id="B93B4D51FF810006FB36FF62FAC932BE" bold="true" box="[1242,1298,223,249]" pageId="16" pageNumber="167">Legs</emphasis>
(
<figureCitation id="13748DC6FF810006FAC9FF62FA4B32BE" box="[1317,1424,223,249]" captionStart="FIGURE 11" captionStartId="17.[151,250,1597,1621]" captionTargetBox="[151,1436,181,1574]" captionTargetId="figure-18@17.[151,1436,181,1574]" captionTargetPageId="17" captionText="FIGURE 11. Paraclius brooksi sp. nov., A–E male holotype (CNC), F male paratype (CNC). A. Habitus, lateral view; B. Thorax, dorsal view; C–D. Head anterior and lateral views, respectively; E. Abdomen and hypopygium, lateral view; F. Left wing." figureDoi="http://doi.org/10.5281/zenodo.8204135" httpUri="https://zenodo.org/record/8204135/files/figure.png" pageId="16" pageNumber="167">Fig. 11A</figureCitation>
). Mostly yellow, except extreme base of coxa I, basal 2/3 of coxa II and basal 1/3 of coxa III, tarsi I and II from apex of tarsomere 1, apex of dorsal and posterior surfaces of femur III, apical 1/3 of tibia III and entirely tarsus III brown. 
<emphasis id="B93B4D51FF810006FF7BFEF6FF3B3322" bold="true" box="[151,224,331,357]" pageId="16" pageNumber="167">Leg I.</emphasis>
Podomere ratios: 27, 27, 11/4/3/3/3. Anterior surface of coxa I covered with silvery pruinosity on apical 1/2, apical edge with 4–5 strong black setae. Femur I with anteroventral row ending in 2–3 long preapical setae, 1 preapical and 1 apical long posteroventral setae. Ventral surface of tarsus I from apex of It 
<subScript id="17CB9306FF810006FB56FE18FB1933F4" attach="left" box="[1210,1218,421,435]" fontSize="6" pageId="16" pageNumber="167">1</subScript>
with weak whitish pilosity (MSSC). 
<emphasis id="B93B4D51FF810006FE8FFE0AFE6D3396" bold="true" box="[355,438,439,465]" pageId="16" pageNumber="167">Leg II.</emphasis>
Podomere ratios: 40, 40, 20/12/10/7/5. Femur II with anteroventral row of setae from base to apical 5/6 (about 0.4X as long as width of femur at broadest point), ending in 2–3 long preapical setae, 1 posteroventral more conspicuous seta at apical 1/3, and 1 strong anterior preapical seta. 
<emphasis id="B93B4D51FF810006FBB4FE42FB69305E" bold="true" box="[1112,1202,511,537]" pageId="16" pageNumber="167">Leg III.</emphasis>
Podomere ratios: 42, 50, 14/20/15/10/7. Femur III laterally compressed, about 1.5X wider than femur II at broadest point. Femur III with dorsal row of more erect setae (about 1/3 as long as width of femur) from base to apical 4/6, with ventral row of short setae (about 1/4 as long as width of femur), from base to apical 5/6, ending in 3 more conspicuous preapical setae, 1 short posteroventral seta and 1 strong anterior preapical seta. 
<emphasis id="B93B4D51FF810006FCD3FD33FC6F30EF" bold="true" box="[831,948,654,680]" pageId="16" pageNumber="167">Abdomen</emphasis>
(
<figureCitation id="13748DC6FF810006FC2DFD32FB9330EE" box="[961,1096,655,681]" captionStart="FIGURE 11" captionStartId="17.[151,250,1597,1621]" captionTargetBox="[151,1436,181,1574]" captionTargetId="figure-18@17.[151,1436,181,1574]" captionTargetPageId="17" captionText="FIGURE 11. Paraclius brooksi sp. nov., A–E male holotype (CNC), F male paratype (CNC). A. Habitus, lateral view; B. Thorax, dorsal view; C–D. Head anterior and lateral views, respectively; E. Abdomen and hypopygium, lateral view; F. Left wing." figureDoi="http://doi.org/10.5281/zenodo.8204135" httpUri="https://zenodo.org/record/8204135/files/figure.png" pageId="16" pageNumber="167">Figs 11A, E</figureCitation>
, 
<figureCitation id="13748DC6FF810006FBB8FD32FB7230EE" box="[1108,1193,655,681]" captionStart="FIGURE12" captionStartId="18.[151,250,1647,1671]" captionTargetBox="[151,1436,237,1624]" captionTargetId="figure-42@18.[151,1436,237,1624]" captionTargetPageId="18" captionText="FIGURE12.Paraclius brooksisp.nov., male paratype (CNC).A. Abdomen, ventralview;B. Cerci, dorsalview;C. Hypopygium, left lateral view; D. Surstyli and apical edge of epandrium; E. Internal appendages; F. Hypandrium and phallus, ventral view; G. Tergite 7, dorsal view. Abreviations: a apv s = apical apicoventral epandrial seta; b apv s = basal apicoventral epandrial seta; cerc = cercus; dsur = dorsal lobe of surstylus; ej apod = ejaculatory apodeme; epand = epandrium; hypd = hypandrium; hypd pr = hypandrial process; pgt = postgonite; ph = phallus; ph pr = phallus process; prct = proctiger; sp = sperm pump; st = sternite; tg = tergite; vsur = ventral lobe of surstylus." figureDoi="http://doi.org/10.5281/zenodo.8204139" httpUri="https://zenodo.org/record/8204139/files/figure.png" pageId="16" pageNumber="167">12A, G</figureCitation>
). Metallic green, with greenish and bluish reflections. Posterior margin of tergite 5 with long setae, but not overlapping posterior margin of tergite 6. Lateral margin of tergite 6 with 4 long and strong setae, apicalmost seta longer than tergite 6. Tergite 7 covered with weak pruinosity, with narrow and deep concavity at posterior margin, reaching 2/3 of tergite. Sternite 5 U-shaped anteriorly, with two narrow 0-shaped sclerites posteriorly. Sternite 8 subovoid, setose on posterior half. 
<emphasis id="B93B4D51FF810006FF7BFCFFFEE8311B" bold="true" box="[151,307,834,860]" pageId="16" pageNumber="167">Hypopygium</emphasis>
(
<figureCitation id="13748DC6FF810006FEACFCFEFE7C311A" box="[320,423,835,861]" captionStart="FIGURE 11" captionStartId="17.[151,250,1597,1621]" captionTargetBox="[151,1436,181,1574]" captionTargetId="figure-18@17.[151,1436,181,1574]" captionTargetPageId="17" captionText="FIGURE 11. Paraclius brooksi sp. nov., A–E male holotype (CNC), F male paratype (CNC). A. Habitus, lateral view; B. Thorax, dorsal view; C–D. Head anterior and lateral views, respectively; E. Abdomen and hypopygium, lateral view; F. Left wing." figureDoi="http://doi.org/10.5281/zenodo.8204135" httpUri="https://zenodo.org/record/8204135/files/figure.png" pageId="16" pageNumber="167">Figs 11E</figureCitation>
, 
<figureCitation id="13748DC6FF810006FE5DFCFEFDDA311A" box="[433,513,835,861]" captionStart="FIGURE12" captionStartId="18.[151,250,1647,1671]" captionTargetBox="[151,1436,237,1624]" captionTargetId="figure-42@18.[151,1436,237,1624]" captionTargetPageId="18" captionText="FIGURE12.Paraclius brooksisp.nov., male paratype (CNC).A. Abdomen, ventralview;B. Cerci, dorsalview;C. Hypopygium, left lateral view; D. Surstyli and apical edge of epandrium; E. Internal appendages; F. Hypandrium and phallus, ventral view; G. Tergite 7, dorsal view. Abreviations: a apv s = apical apicoventral epandrial seta; b apv s = basal apicoventral epandrial seta; cerc = cercus; dsur = dorsal lobe of surstylus; ej apod = ejaculatory apodeme; epand = epandrium; hypd = hypandrium; hypd pr = hypandrial process; pgt = postgonite; ph = phallus; ph pr = phallus process; prct = proctiger; sp = sperm pump; st = sternite; tg = tergite; vsur = ventral lobe of surstylus." figureDoi="http://doi.org/10.5281/zenodo.8204139" httpUri="https://zenodo.org/record/8204139/files/figure.png" pageId="16" pageNumber="167">12B–F</figureCitation>
). Epandrium long and narrow, about 2.8X longer than high, with acute apicoventral lateral ridge (
<figureCitation id="13748DC6FF810006FEDCFCDAFE6631C6" box="[304,445,871,897]" captionStart="FIGURE12" captionStartId="18.[151,250,1647,1671]" captionTargetBox="[151,1436,237,1624]" captionTargetId="figure-42@18.[151,1436,237,1624]" captionTargetPageId="18" captionText="FIGURE12.Paraclius brooksisp.nov., male paratype (CNC).A. Abdomen, ventralview;B. Cerci, dorsalview;C. Hypopygium, left lateral view; D. Surstyli and apical edge of epandrium; E. Internal appendages; F. Hypandrium and phallus, ventral view; G. Tergite 7, dorsal view. Abreviations: a apv s = apical apicoventral epandrial seta; b apv s = basal apicoventral epandrial seta; cerc = cercus; dsur = dorsal lobe of surstylus; ej apod = ejaculatory apodeme; epand = epandrium; hypd = hypandrium; hypd pr = hypandrial process; pgt = postgonite; ph = phallus; ph pr = phallus process; prct = proctiger; sp = sperm pump; st = sternite; tg = tergite; vsur = ventral lobe of surstylus." figureDoi="http://doi.org/10.5281/zenodo.8204139" httpUri="https://zenodo.org/record/8204139/files/figure.png" pageId="16" pageNumber="167">Fig. 12C, D</figureCitation>
). Basal apicoventral seta about 2X longer than apical seta, slightly shorter than ventral lobe of surstylus (
<figureCitation id="13748DC6FF810006FEB2FC36FE3D31E2" box="[350,486,907,933]" captionStart="FIGURE12" captionStartId="18.[151,250,1647,1671]" captionTargetBox="[151,1436,237,1624]" captionTargetId="figure-42@18.[151,1436,237,1624]" captionTargetPageId="18" captionText="FIGURE12.Paraclius brooksisp.nov., male paratype (CNC).A. Abdomen, ventralview;B. Cerci, dorsalview;C. Hypopygium, left lateral view; D. Surstyli and apical edge of epandrium; E. Internal appendages; F. Hypandrium and phallus, ventral view; G. Tergite 7, dorsal view. Abreviations: a apv s = apical apicoventral epandrial seta; b apv s = basal apicoventral epandrial seta; cerc = cercus; dsur = dorsal lobe of surstylus; ej apod = ejaculatory apodeme; epand = epandrium; hypd = hypandrium; hypd pr = hypandrial process; pgt = postgonite; ph = phallus; ph pr = phallus process; prct = proctiger; sp = sperm pump; st = sternite; tg = tergite; vsur = ventral lobe of surstylus." figureDoi="http://doi.org/10.5281/zenodo.8204139" httpUri="https://zenodo.org/record/8204139/files/figure.png" pageId="16" pageNumber="167">Fig. 12C, D</figureCitation>
). Ventral lobe of surstylus long, somewhat L-shaped, strong curved ventrally at apex, with short and stout modified apical seta (
<figureCitation id="13748DC6FF810006FD85FC12FD2A318E" box="[617,753,943,969]" captionStart="FIGURE12" captionStartId="18.[151,250,1647,1671]" captionTargetBox="[151,1436,237,1624]" captionTargetId="figure-42@18.[151,1436,237,1624]" captionTargetPageId="18" captionText="FIGURE12.Paraclius brooksisp.nov., male paratype (CNC).A. Abdomen, ventralview;B. Cerci, dorsalview;C. Hypopygium, left lateral view; D. Surstyli and apical edge of epandrium; E. Internal appendages; F. Hypandrium and phallus, ventral view; G. Tergite 7, dorsal view. Abreviations: a apv s = apical apicoventral epandrial seta; b apv s = basal apicoventral epandrial seta; cerc = cercus; dsur = dorsal lobe of surstylus; ej apod = ejaculatory apodeme; epand = epandrium; hypd = hypandrium; hypd pr = hypandrial process; pgt = postgonite; ph = phallus; ph pr = phallus process; prct = proctiger; sp = sperm pump; st = sternite; tg = tergite; vsur = ventral lobe of surstylus." figureDoi="http://doi.org/10.5281/zenodo.8204139" httpUri="https://zenodo.org/record/8204139/files/figure.png" pageId="16" pageNumber="167">Fig. 12C, D</figureCitation>
). Dorsal lobe of surstylus short and wide, narrowing at apical 1/3, about 1/3 as long as ventral lobe of surstylus, with 1 short and strong seta at middle of dorsal edge, 1 slender seta at middle of inner surface, 1 short and slender seta at apical 1/3 of dorsal edge and 1 long and slender seta at apical 1/3 of outer edge (
<figureCitation id="13748DC6FF810006FE59FBA6FDC43672" box="[437,543,1051,1077]" captionStart="FIGURE12" captionStartId="18.[151,250,1647,1671]" captionTargetBox="[151,1436,237,1624]" captionTargetId="figure-42@18.[151,1436,237,1624]" captionTargetPageId="18" captionText="FIGURE12.Paraclius brooksisp.nov., male paratype (CNC).A. Abdomen, ventralview;B. Cerci, dorsalview;C. Hypopygium, left lateral view; D. Surstyli and apical edge of epandrium; E. Internal appendages; F. Hypandrium and phallus, ventral view; G. Tergite 7, dorsal view. Abreviations: a apv s = apical apicoventral epandrial seta; b apv s = basal apicoventral epandrial seta; cerc = cercus; dsur = dorsal lobe of surstylus; ej apod = ejaculatory apodeme; epand = epandrium; hypd = hypandrium; hypd pr = hypandrial process; pgt = postgonite; ph = phallus; ph pr = phallus process; prct = proctiger; sp = sperm pump; st = sternite; tg = tergite; vsur = ventral lobe of surstylus." figureDoi="http://doi.org/10.5281/zenodo.8204139" httpUri="https://zenodo.org/record/8204139/files/figure.png" pageId="16" pageNumber="167">Fig. 12D</figureCitation>
). Postgonite well sclerotized, short and slightly curved ventrally (
<figureCitation id="13748DC6FF810006FAEBFBA6FA4B3672" box="[1287,1424,1051,1077]" captionStart="FIGURE12" captionStartId="18.[151,250,1647,1671]" captionTargetBox="[151,1436,237,1624]" captionTargetId="figure-42@18.[151,1436,237,1624]" captionTargetPageId="18" captionText="FIGURE12.Paraclius brooksisp.nov., male paratype (CNC).A. Abdomen, ventralview;B. Cerci, dorsalview;C. Hypopygium, left lateral view; D. Surstyli and apical edge of epandrium; E. Internal appendages; F. Hypandrium and phallus, ventral view; G. Tergite 7, dorsal view. Abreviations: a apv s = apical apicoventral epandrial seta; b apv s = basal apicoventral epandrial seta; cerc = cercus; dsur = dorsal lobe of surstylus; ej apod = ejaculatory apodeme; epand = epandrium; hypd = hypandrium; hypd pr = hypandrial process; pgt = postgonite; ph = phallus; ph pr = phallus process; prct = proctiger; sp = sperm pump; st = sternite; tg = tergite; vsur = ventral lobe of surstylus." figureDoi="http://doi.org/10.5281/zenodo.8204139" httpUri="https://zenodo.org/record/8204139/files/figure.png" pageId="16" pageNumber="167">Fig. 12C, E</figureCitation>
). Proctiger plain, weakly sclerotized (
<figureCitation id="13748DC6FF810006FDDBFB82FD7B361E" box="[567,672,1087,1113]" captionStart="FIGURE12" captionStartId="18.[151,250,1647,1671]" captionTargetBox="[151,1436,237,1624]" captionTargetId="figure-42@18.[151,1436,237,1624]" captionTargetPageId="18" captionText="FIGURE12.Paraclius brooksisp.nov., male paratype (CNC).A. Abdomen, ventralview;B. Cerci, dorsalview;C. Hypopygium, left lateral view; D. Surstyli and apical edge of epandrium; E. Internal appendages; F. Hypandrium and phallus, ventral view; G. Tergite 7, dorsal view. Abreviations: a apv s = apical apicoventral epandrial seta; b apv s = basal apicoventral epandrial seta; cerc = cercus; dsur = dorsal lobe of surstylus; ej apod = ejaculatory apodeme; epand = epandrium; hypd = hypandrium; hypd pr = hypandrial process; pgt = postgonite; ph = phallus; ph pr = phallus process; prct = proctiger; sp = sperm pump; st = sternite; tg = tergite; vsur = ventral lobe of surstylus." figureDoi="http://doi.org/10.5281/zenodo.8204139" httpUri="https://zenodo.org/record/8204139/files/figure.png" pageId="16" pageNumber="167">Fig. 12E</figureCitation>
). Hypandrium trough-shaped, with weak connection to epandrium basiventrally, free laterally, apex bifurcate, weakly sclerotized and asymmetrical, left lateral margin with 1 short dentiform preapical projection, and right lateral margin with minute dentiform projection, base of hypandrium projecting up inside epandrial capsule, cradling phallus (
<figureCitation id="13748DC6FF810006FCFFFB16FC6D3682" box="[787,950,1195,1221]" captionStart="FIGURE12" captionStartId="18.[151,250,1647,1671]" captionTargetBox="[151,1436,237,1624]" captionTargetId="figure-42@18.[151,1436,237,1624]" captionTargetPageId="18" captionText="FIGURE12.Paraclius brooksisp.nov., male paratype (CNC).A. Abdomen, ventralview;B. Cerci, dorsalview;C. Hypopygium, left lateral view; D. Surstyli and apical edge of epandrium; E. Internal appendages; F. Hypandrium and phallus, ventral view; G. Tergite 7, dorsal view. Abreviations: a apv s = apical apicoventral epandrial seta; b apv s = basal apicoventral epandrial seta; cerc = cercus; dsur = dorsal lobe of surstylus; ej apod = ejaculatory apodeme; epand = epandrium; hypd = hypandrium; hypd pr = hypandrial process; pgt = postgonite; ph = phallus; ph pr = phallus process; prct = proctiger; sp = sperm pump; st = sternite; tg = tergite; vsur = ventral lobe of surstylus." figureDoi="http://doi.org/10.5281/zenodo.8204139" httpUri="https://zenodo.org/record/8204139/files/figure.png" pageId="16" pageNumber="167">Fig. 12C, E, F</figureCitation>
). Phallus entirely divided into two strongly sclerotized narrow arms, narrowing at apex, right arm with 1 thumb-shaped projection (
<figureCitation id="13748DC6FF810006FB6CFB72FADD36AE" box="[1152,1286,1231,1257]" captionStart="FIGURE12" captionStartId="18.[151,250,1647,1671]" captionTargetBox="[151,1436,237,1624]" captionTargetId="figure-42@18.[151,1436,237,1624]" captionTargetPageId="18" captionText="FIGURE12.Paraclius brooksisp.nov., male paratype (CNC).A. Abdomen, ventralview;B. Cerci, dorsalview;C. Hypopygium, left lateral view; D. Surstyli and apical edge of epandrium; E. Internal appendages; F. Hypandrium and phallus, ventral view; G. Tergite 7, dorsal view. Abreviations: a apv s = apical apicoventral epandrial seta; b apv s = basal apicoventral epandrial seta; cerc = cercus; dsur = dorsal lobe of surstylus; ej apod = ejaculatory apodeme; epand = epandrium; hypd = hypandrium; hypd pr = hypandrial process; pgt = postgonite; ph = phallus; ph pr = phallus process; prct = proctiger; sp = sperm pump; st = sternite; tg = tergite; vsur = ventral lobe of surstylus." figureDoi="http://doi.org/10.5281/zenodo.8204139" httpUri="https://zenodo.org/record/8204139/files/figure.png" pageId="16" pageNumber="167">Fig. 12E, F</figureCitation>
). Ejaculatory apodeme long and narrow, about as long as 1/2 of epandrium, laterally compressed and weakly sclerotized (
<figureCitation id="13748DC6FF810006FA83FB4EFF133776" captionStart="FIGURE12" captionStartId="18.[151,250,1647,1671]" captionTargetBox="[151,1436,237,1624]" captionTargetId="figure-42@18.[151,1436,237,1624]" captionTargetPageId="18" captionText="FIGURE12.Paraclius brooksisp.nov., male paratype (CNC).A. Abdomen, ventralview;B. Cerci, dorsalview;C. Hypopygium, left lateral view; D. Surstyli and apical edge of epandrium; E. Internal appendages; F. Hypandrium and phallus, ventral view; G. Tergite 7, dorsal view. Abreviations: a apv s = apical apicoventral epandrial seta; b apv s = basal apicoventral epandrial seta; cerc = cercus; dsur = dorsal lobe of surstylus; ej apod = ejaculatory apodeme; epand = epandrium; hypd = hypandrium; hypd pr = hypandrial process; pgt = postgonite; ph = phallus; ph pr = phallus process; prct = proctiger; sp = sperm pump; st = sternite; tg = tergite; vsur = ventral lobe of surstylus." figureDoi="http://doi.org/10.5281/zenodo.8204139" httpUri="https://zenodo.org/record/8204139/files/figure.png" pageId="16" pageNumber="167">Fig. 12E</figureCitation>
). Sperm pump short and wide, somewhat C-shaped, about 1/2 as long as ejaculatory apodeme (
<figureCitation id="13748DC6FF810006FACBFAAAFA4B3776" box="[1319,1424,1303,1329]" captionStart="FIGURE12" captionStartId="18.[151,250,1647,1671]" captionTargetBox="[151,1436,237,1624]" captionTargetId="figure-42@18.[151,1436,237,1624]" captionTargetPageId="18" captionText="FIGURE12.Paraclius brooksisp.nov., male paratype (CNC).A. Abdomen, ventralview;B. Cerci, dorsalview;C. Hypopygium, left lateral view; D. Surstyli and apical edge of epandrium; E. Internal appendages; F. Hypandrium and phallus, ventral view; G. Tergite 7, dorsal view. Abreviations: a apv s = apical apicoventral epandrial seta; b apv s = basal apicoventral epandrial seta; cerc = cercus; dsur = dorsal lobe of surstylus; ej apod = ejaculatory apodeme; epand = epandrium; hypd = hypandrium; hypd pr = hypandrial process; pgt = postgonite; ph = phallus; ph pr = phallus process; prct = proctiger; sp = sperm pump; st = sternite; tg = tergite; vsur = ventral lobe of surstylus." figureDoi="http://doi.org/10.5281/zenodo.8204139" httpUri="https://zenodo.org/record/8204139/files/figure.png" pageId="16" pageNumber="167">Fig. 12E</figureCitation>
). Cercus long and oval, about 3X longer than wide, and about 1/2 as long as epandrium, inner and outer surfaces covered by short and sparse setae, ventral edge with long setae, remarkably long on basal 1/3 (
<figureCitation id="13748DC6FF810006FB25FAE2FAEB373E" box="[1225,1328,1375,1401]" captionStart="FIGURE12" captionStartId="18.[151,250,1647,1671]" captionTargetBox="[151,1436,237,1624]" captionTargetId="figure-42@18.[151,1436,237,1624]" captionTargetPageId="18" captionText="FIGURE12.Paraclius brooksisp.nov., male paratype (CNC).A. Abdomen, ventralview;B. Cerci, dorsalview;C. Hypopygium, left lateral view; D. Surstyli and apical edge of epandrium; E. Internal appendages; F. Hypandrium and phallus, ventral view; G. Tergite 7, dorsal view. Abreviations: a apv s = apical apicoventral epandrial seta; b apv s = basal apicoventral epandrial seta; cerc = cercus; dsur = dorsal lobe of surstylus; ej apod = ejaculatory apodeme; epand = epandrium; hypd = hypandrium; hypd pr = hypandrial process; pgt = postgonite; ph = phallus; ph pr = phallus process; prct = proctiger; sp = sperm pump; st = sternite; tg = tergite; vsur = ventral lobe of surstylus." figureDoi="http://doi.org/10.5281/zenodo.8204139" httpUri="https://zenodo.org/record/8204139/files/figure.png" pageId="16" pageNumber="167">Fig. 12B</figureCitation>
). 
<emphasis id="B93B4D51FF810006FAA9FAE2FA47373E" bold="true" box="[1349,1436,1375,1401]" pageId="16" pageNumber="167">Female</emphasis>
(
<figureCitation id="13748DC6FF810006FF73FA3EFF2F37DA" box="[159,244,1411,1437]" captionStart="FIGURE 13" captionStartId="19.[151,250,655,679]" captionTargetBox="[151,1436,181,632]" captionTargetId="figure-18@19.[151,1436,181,632]" captionTargetPageId="19" captionText="FIGURE 13. Paraclius brooksi sp. nov, identified female from Mexico (CNC). A. Habitus, lateral view; B. Head, anterior view; C. Right antenna, outer view." figureDoi="http://doi.org/10.5281/zenodo.8204141" httpUri="https://zenodo.org/record/8204141/files/figure.png" pageId="16" pageNumber="167">Fig. 13</figureCitation>
). Body length: 
<quantity id="4CB73CA6FF810006FE4FFA3EFE2637DA" box="[419,509,1411,1437]" metricMagnitude="-3" metricUnit="m" metricValue="3.2" pageId="16" pageNumber="167" unit="mm" value="3.2">3.2 mm</quantity>
; wing length: 
<quantity id="4CB73CA6FF810006FD72FA3EFD2E37DA" box="[670,757,1411,1437]" metricMagnitude="-3" metricUnit="m" metricValue="2.6" pageId="16" pageNumber="167" unit="mm" value="2.6">2.6 mm</quantity>
(n = 1). Similar to male, except as noted: Postpedicel short, about as long as wide, somewhat subtriangular (
<figureCitation id="13748DC6FF810006FD58FA1AFCC13786" box="[692,794,1447,1473]" captionStart="FIGURE 13" captionStartId="19.[151,250,655,679]" captionTargetBox="[151,1436,181,632]" captionTargetId="figure-18@19.[151,1436,181,632]" captionTargetPageId="19" captionText="FIGURE 13. Paraclius brooksi sp. nov, identified female from Mexico (CNC). A. Habitus, lateral view; B. Head, anterior view; C. Right antenna, outer view." figureDoi="http://doi.org/10.5281/zenodo.8204141" httpUri="https://zenodo.org/record/8204141/files/figure.png" pageId="16" pageNumber="167">Fig. 13C</figureCitation>
); face and clypeus wide, both wider than ocellar tubercle (
<figureCitation id="13748DC6FF810006FF73FA76FEDD37A2" box="[159,262,1483,1509]" captionStart="FIGURE 13" captionStartId="19.[151,250,655,679]" captionTargetBox="[151,1436,181,632]" captionTargetId="figure-18@19.[151,1436,181,632]" captionTargetPageId="19" captionText="FIGURE 13. Paraclius brooksi sp. nov, identified female from Mexico (CNC). A. Habitus, lateral view; B. Head, anterior view; C. Right antenna, outer view." figureDoi="http://doi.org/10.5281/zenodo.8204141" httpUri="https://zenodo.org/record/8204141/files/figure.png" pageId="16" pageNumber="167">Fig. 13B</figureCitation>
). Terminalia as in 
<taxonomicName id="4C4FEAC0FF810006FE3EFA76FD8937A3" authorityName=": Loew" authorityYear="1864" box="[466,594,1483,1509]" class="Insecta" family="Dolichopodidae" genus="Paraclius" kingdom="Animalia" order="Diptera" pageId="16" pageNumber="167" phylum="Arthropoda" rank="species" species="arcuatus">
<emphasis id="B93B4D51FF810006FE3EFA76FE3837A3" box="[466,483,1483,1508]" italics="true" pageId="16" pageNumber="167">P</emphasis>
. 
<emphasis id="B93B4D51FF810006FE1DFA76FD8937A3" box="[497,594,1483,1508]" italics="true" pageId="16" pageNumber="167">arcuatus</emphasis>
</taxonomicName>
, except as noted: tergite 10 divided medially into hemitergites each bearing 4 spines and 1 minute lateral seta.
</paragraph>
</subSubSection>
<subSubSection id="C355C2C8FF810006FF2BF9AEFE5135FE" pageId="16" pageNumber="167" type="materials_examined">
<paragraph id="8BF09143FF810006FF2BF9AEFD8534A6" blockId="16.[151,1437,151,1977]" pageId="16" pageNumber="167">
<emphasis id="B93B4D51FF810006FF2BF9AEFEAD346B" bold="true" box="[199,374,1554,1581]" pageId="16" pageNumber="167">Type material.</emphasis>
<materialsCitation id="3B279B1EFF810006FE92F9AFFDBC3432" collectingDate="1969-06-04" collectionCode="V, CNC" country="Mexico" elevation="900" latitude="15.199111" location="Chiapas" longLatPrecision="1" longitude="-92.46717" pageId="16" pageNumber="167" specimenCount="2" stateProvince="Chiapas" typeStatus="holotype">
<emphasis id="B93B4D51FF810006FE92F9AFFDC6346B" bold="true" box="[382,541,1554,1580]" pageId="16" pageNumber="167">
<typeStatus id="54F42FE1FF810006FE92F9AFFDC6346B" box="[382,541,1554,1580]" pageId="16" pageNumber="167">HOLOTYPE</typeStatus>
</emphasis>
♁ (
<figureCitation id="13748DC6FF810006FDA6F9AEFD41346A" box="[586,666,1555,1581]" captionStart="FIGURE 11" captionStartId="17.[151,250,1597,1621]" captionTargetBox="[151,1436,181,1574]" captionTargetId="figure-18@17.[151,1436,181,1574]" captionTargetPageId="17" captionText="FIGURE 11. Paraclius brooksi sp. nov., A–E male holotype (CNC), F male paratype (CNC). A. Habitus, lateral view; B. Thorax, dorsal view; C–D. Head anterior and lateral views, respectively; E. Abdomen and hypopygium, lateral view; F. Left wing." figureDoi="http://doi.org/10.5281/zenodo.8204135" httpUri="https://zenodo.org/record/8204135/files/figure.png" pageId="16" pageNumber="167">Fig. 11</figureCitation>
) labelled: “ 
<collectingCountry id="F358D1D3FF810006FCF2F9AEFCA2346A" box="[798,889,1555,1581]" name="Mexico" pageId="16" pageNumber="167">Mexico</collectingCountry>
, Chis. [
<collectingRegion id="498B5FA1FF810006FC21F9AEFBF6346A" box="[973,1069,1555,1581]" country="Mexico" name="Chiapas" pageId="16" pageNumber="167">Chiapas</collectingRegion>
] 20–25 | mi. N Huixtla 3000’ [= 
<quantity id="4CB73CA6FF810006FF7BF98AFF393416" box="[151,226,1591,1617]" metricMagnitude="2" metricUnit="m" metricValue="9.0" pageId="16" pageNumber="167" unit="m" value="900.0">
<elevation id="00627670FF810006FF7BF98AFF393416" box="[151,226,1591,1617]" metricMagnitude="2" metricUnit="m" metricValue="9.0" pageId="16" pageNumber="167" unit="m" value="900.0">900 m</elevation>
</quantity>
] [ca 
<geoCoordinate id="EE7BF784FF810006FEF1F985FE6C3414" box="[285,439,1592,1619]" degrees="15" direction="north" minutes="11" orientation="latitude" pageId="16" pageNumber="167" precision="1" seconds="56.8" value="15.199111">15°11′56.8″N</geoCoordinate>
<geoCoordinate id="EE7BF784FF810006FE2CF985FDB83414" box="[448,611,1592,1619]" degrees="92" direction="west" minutes="28" orientation="longitude" pageId="16" pageNumber="167" precision="1" seconds="01.8" value="-92.46717">92°28′01.8″W</geoCoordinate>
] | 
<date id="FFF1B783FF810006FD6DF98AFD313416" box="[641,746,1591,1617]" pageId="16" pageNumber="167" value="1969-06-04">
<collectingDate id="EFB54E6BFF810006FD6DF98AFD313416" box="[641,746,1591,1617]" pageId="16" pageNumber="167" value="1969-06-04">4.vi.1969</collectingDate>
</date>
| B. 
<collectionCode id="ED5E0986FF810006FCF5F98AFCEB3416" box="[793,816,1591,1617]" country="Canada" lsid="urn:lsid:biocol.org:col:13946" name="Royal British Columbia Museum - Herbarium" pageId="16" pageNumber="167" type="Museum">V</collectionCode>
. Peterson” “HOLOTYPE | 
<taxonomicName id="4C4FEAC0FF810006FB86F98AFAE83416" authorityName="Soares, Runyon &amp; Capellari" authorityYear="2023" box="[1130,1331,1591,1617]" class="Insecta" family="Dolichopodidae" genus="Paraclius" kingdom="Animalia" order="Diptera" pageId="16" pageNumber="167" phylum="Arthropoda" rank="species" species="brooksi">
<emphasis id="B93B4D51FF810006FB86F98AFAE83416" box="[1130,1331,1591,1617]" italics="true" pageId="16" pageNumber="167">Paraclius brooksi</emphasis>
</taxonomicName>
| Soares, Runyon &amp; Capellari” [red label] (
<collectionCode id="ED5E0986FF810006FDCCF9E6FD843432" box="[544,607,1627,1653]" country="Canada" httpUri="http://grbio.org/cool/y2kv-9w3k" name="Canadian National Collection of Insects, Arachnids, and Nematodes" pageId="16" pageNumber="167">CNC</collectionCode>
)
</materialsCitation>
. 
<typeStatus id="54F42FE1FF810006FD99F9E6FD063432" box="[629,733,1627,1653]" pageId="16" pageNumber="167">Holotype</typeStatus>
condition: good, not dissected. 
<materialsCitation id="3B279B1EFF810006FBBDF9E7FF2134FA" collectingDate="1969-06-03" collectionCode="NMNH" country="Mexico" elevation="900" latitude="15.199111" location="Chiapas" longLatPrecision="1" longitude="-92.46717" pageId="16" pageNumber="167" specimenCount="1" stateProvince="Chiapas" typeStatus="paratype">
<emphasis id="B93B4D51FF810006FBBDF9E7FB243433" bold="true" box="[1105,1279,1626,1652]" pageId="16" pageNumber="167">
<typeStatus id="54F42FE1FF810006FBBDF9E7FB223433" box="[1105,1273,1626,1652]" pageId="16" pageNumber="167">PARATYPES</typeStatus>
:
</emphasis>
same data as holotype, except: 
<date id="FFF1B783FF810006FE8EF9C2FE1734DE" box="[354,460,1663,1689]" pageId="16" pageNumber="167" value="1969-06-03">
<collectingDate id="EFB54E6BFF810006FE8EF9C2FE1734DE" box="[354,460,1663,1689]" pageId="16" pageNumber="167" value="1969-06-03">3.vi.1969</collectingDate>
</date>
(1 ♁ with head and terminalia dissected, and left wing mounted between cover slips, 
<collectionCode id="ED5E0986FF810006FF7BF91EFF2934FA" box="[151,242,1699,1725]" country="USA" httpUri="http://grbio.org/cool/142r-0w94" name="Smithsonian Institution, National Museum of Natural History" pageId="16" pageNumber="167" type="Museum">NMNH</collectionCode>
)
</materialsCitation>
; 
<materialsCitation id="3B279B1EFF810006FEE5F91EFAC934FA" box="[265,1298,1699,1727]" collectingDate="1963-05-14" collectionCode="INPA" country="Mexico" latitude="17.792973" location="Valle Nacional" longLatPrecision="1" longitude="-96.26186" pageId="16" pageNumber="167" specimenCount="1" stateProvince="Oaxaca" typeStatus="paratype">
<collectingRegion id="498B5FA1FF810006FEE5F91EFEBB34FA" box="[265,352,1699,1725]" country="Mexico" name="Oaxaca" pageId="16" pageNumber="167">Oaxaca</collectingRegion>
, above 
<location id="8E90C798FF810006FE51F91EFDBC34FA" LSID="urn:lsid:plazi:treatment:03E62055FF9E0004FF7BFB22FF1F358B:8E90C798FF810006FE51F91EFDBC34FA" box="[445,615,1699,1725]" country="Mexico" latitude="17.792973" longLatPrecision="1" longitude="-96.26186" name="Valle Nacional" pageId="16" pageNumber="167" stateProvince="Oaxaca">Valle Nacional</location>
[ca 
<geoCoordinate id="EE7BF784FF810006FD4DF919FCE134F8" box="[673,826,1700,1727]" degrees="17" direction="north" minutes="47" orientation="latitude" pageId="16" pageNumber="167" precision="1" seconds="34.7" value="17.792973">17°47′34.7″N</geoCoordinate>
<geoCoordinate id="EE7BF784FF810006FCAAF919FC3734F8" box="[838,1004,1700,1727]" degrees="96" direction="west" minutes="15" orientation="longitude" pageId="16" pageNumber="167" precision="1" seconds="42.7" value="-96.26186">96°15′42.7″W</geoCoordinate>
], 
<date id="FFF1B783FF810006FBEEF91EFBAA34FA" box="[1026,1137,1699,1725]" pageId="16" pageNumber="167" value="1963-05-14">
<collectingDate id="EFB54E6BFF810006FBEEF91EFBAA34FA" box="[1026,1137,1699,1725]" pageId="16" pageNumber="167" value="1963-05-14">14.v.1963</collectingDate>
</date>
(1 ♁, 
<collectionCode id="ED5E0986FF810006FB2AF91EFAD134FA" box="[1222,1290,1699,1725]" country="Brazil" httpUri="http://grbio.org/cool/rm88-499z" name="Instituto Nacional de Pesquisas da Amazonia" pageId="16" pageNumber="167">INPA</collectionCode>
)
</materialsCitation>
; 
<materialsCitation id="3B279B1EFF810006FACEF91EFD8134A6" collectingDate="1963-05-21" collectionCode="CNC" country="Mexico" latitude="17.792973" location="Valle Nacional" longLatPrecision="1" longitude="-96.26186" pageId="16" pageNumber="167" specimenCount="1" stateProvince="Oaxaca" typeStatus="paratype">
same data, except: 
<date id="FFF1B783FF810006FF03F97AFE8534A6" box="[239,350,1735,1761]" pageId="16" pageNumber="167" value="1963-05-21">
<collectingDate id="EFB54E6BFF810006FF03F97AFE8534A6" box="[239,350,1735,1761]" pageId="16" pageNumber="167" value="1963-05-21">21.v.1963</collectingDate>
</date>
(1 ♁ dissected, 
<collectionCode id="ED5E0986FF810006FDFFF97AFD8A34A6" box="[531,593,1735,1761]" country="Canada" httpUri="http://grbio.org/cool/y2kv-9w3k" name="Canadian National Collection of Insects, Arachnids, and Nematodes" pageId="16" pageNumber="167">CNC</collectionCode>
)
</materialsCitation>
.
</paragraph>
<paragraph id="8BF09143FF810006FF2BF957FE5135FE" blockId="16.[151,1437,151,1977]" pageId="16" pageNumber="167">
<emphasis id="B93B4D51FF810006FF2BF957FD303543" bold="true" box="[199,747,1770,1796]" pageId="16" pageNumber="167">
Additional material examined. 
<collectingCountry id="F358D1D3FF810006FDB8F957FD3D3543" box="[596,742,1770,1796]" name="Grenada" pageId="16" pageNumber="167">GRENADA</collectingCountry>
:
</emphasis>
W. Indies 190–66, 172, Mt. Maitland [ca 
<geoCoordinate id="EE7BF784FF810006FAEEF951FA473540" box="[1282,1436,1772,1799]" degrees="12" direction="north" minutes="02" orientation="latitude" pageId="16" pageNumber="167" precision="1" seconds="50.3" value="12.047306">12°02′50.3″N</geoCoordinate>
<geoCoordinate id="EE7BF784FF810006FF7BF8ADFEE6356C" box="[151,317,1808,1835]" degrees="61" direction="west" minutes="42" orientation="longitude" pageId="16" pageNumber="167" precision="1" seconds="48.9" value="-61.713585">61°42′48.9″W</geoCoordinate>
], (Leeward side), W.I., H. H. Smith, 
<taxonomicName id="4C4FEAC0FF810006FD0DF8B2FBC9356E" authority="(Loew)" baseAuthorityName="Loew" baseAuthorityYear="1861" box="[737,1042,1807,1833]" class="Insecta" family="Dolichopodidae" genus="Paraclius" kingdom="Animalia" order="Diptera" pageId="16" pageNumber="167" phylum="Arthropoda" rank="species" species="arcuatus">
<emphasis id="B93B4D51FF810006FD0DF8B2FC6C356F" box="[737,951,1807,1833]" italics="true" pageId="16" pageNumber="167">Paraclius arcuatus</emphasis>
(Loew)
</taxonomicName>
det. Fred C. Harmston, Collection of the California Academy of Sciences, San Francisco, Calif., Collection of Calif. Acad. of Sci. 1994 Gift Fred C. Harmston, 
<taxonomicName id="4C4FEAC0FF810006FEF9F8EAFDF93536" authority="Lw." authorityName=": Loew" authorityYear="1864" box="[277,546,1879,1905]" class="Insecta" family="Dolichopodidae" genus="Paraclius" kingdom="Animalia" order="Diptera" pageId="16" pageNumber="167" phylum="Arthropoda" rank="species" species="arcuatus">
<emphasis id="B93B4D51FF810006FEF9F8EAFE373537" box="[277,492,1879,1905]" italics="true" pageId="16" pageNumber="167">Paraclius arcuatus</emphasis>
Lw.
</taxonomicName>
, 
<taxonomicName id="4C4FEAC0FF810006FDC2F8EAFD013536" box="[558,730,1879,1905]" class="Insecta" family="Dolichopodidae" genus="Paraclius" kingdom="Animalia" order="Diptera" pageId="16" pageNumber="167" phylum="Arthropoda" rank="species" species="undefined-1">
<emphasis id="B93B4D51FF810006FDC2F8EAFD403536" box="[558,667,1879,1905]" italics="true" pageId="16" pageNumber="167">Paraclius</emphasis>
sp. 1
</taxonomicName>
(nr 
<taxonomicName id="4C4FEAC0FF810006FCE0F8EAFCB63537" authorityName=": Loew" authorityYear="1864" box="[780,877,1879,1904]" class="Insecta" family="Dolichopodidae" genus="Paraclius" kingdom="Animalia" order="Diptera" pageId="16" pageNumber="167" phylum="Arthropoda" rank="species" species="arcuatus">
<emphasis id="B93B4D51FF810006FCE0F8EAFCB63537" box="[780,877,1879,1904]" italics="true" pageId="16" pageNumber="167">arcuatus</emphasis>
</taxonomicName>
) det. S.E. Brooks, 2004 (
<specimenCount id="9D495ACAFF810006FB78F8EAFB653536" box="[1172,1214,1879,1905]" pageId="16" pageNumber="167" type="female">1 ♀</specimenCount>
, 
<collectionCode id="ED5E0986FF810006FB21F8EAFAD33536" box="[1229,1288,1879,1905]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:15690" lsid="urn:lsid:biocol.org:col:34700" name="California Academy of Sciences" pageId="16" pageNumber="167" type="Museum">CAS</collectionCode>
). 
<emphasis id="B93B4D51FF810006FAF1F8EBFA473537" bold="true" box="[1309,1436,1878,1904]" pageId="16" pageNumber="167">
<collectingCountry id="F358D1D3FF810006FAF1F8EBFA433537" box="[1309,1432,1878,1904]" name="Mexico" pageId="16" pageNumber="167">MEXICO</collectingCountry>
:
</emphasis>
Chis. [
<collectingRegion id="498B5FA1FF810006FF33F8C6FEE435D2" box="[223,319,1915,1941]" country="Mexico" name="Chiapas" pageId="16" pageNumber="167">Chiapas</collectingRegion>
] 20–25, mi. N Huixtla 3000’ [= 900 meters high] [ca 
<geoCoordinate id="EE7BF784FF810006FC76F8C1FBE835D0" box="[922,1075,1916,1943]" degrees="15" direction="north" minutes="11" orientation="latitude" pageId="16" pageNumber="167" precision="1" seconds="56.8" value="15.199111">15°11′56.8″N</geoCoordinate>
<geoCoordinate id="EE7BF784FF810006FBD7F8C1FB3A35D0" box="[1083,1249,1916,1943]" degrees="92" direction="west" minutes="28" orientation="longitude" pageId="16" pageNumber="167" precision="1" seconds="01.8" value="-92.46717">92°28′01.8″W</geoCoordinate>
], 
<date id="FFF1B783FF810006FB1FF8C6FA8435D2" box="[1267,1375,1915,1941]" pageId="16" pageNumber="167" value="1969-06-02">2.vi.1969</date>
, B. 
<collectionCode id="ED5E0986FF810006FA6EF8C6FA4235D2" box="[1410,1433,1915,1941]" country="Canada" lsid="urn:lsid:biocol.org:col:13946" name="Royal British Columbia Museum - Herbarium" pageId="16" pageNumber="167" type="Museum">V</collectionCode>
. Peterson (
<specimenCount id="9D495ACAFF810006FEE6F822FEE935FE" box="[266,306,1951,1977]" pageId="16" pageNumber="167" type="female">1 ♀</specimenCount>
, 
<collectionCode id="ED5E0986FF810006FEACF822FEA535FE" box="[320,382,1951,1977]" country="Canada" httpUri="http://grbio.org/cool/y2kv-9w3k" name="Canadian National Collection of Insects, Arachnids, and Nematodes" pageId="16" pageNumber="167">CNC</collectionCode>
).
</paragraph>
</subSubSection>
<caption id="DF30C1CBFF800007FF7BF980FF0A34D9" ID-DOI="http://doi.org/10.5281/zenodo.8204135" ID-Zenodo-Dep="8204135" httpUri="https://zenodo.org/record/8204135/files/figure.png" pageId="17" pageNumber="168" startId="17.[151,250,1597,1621]" targetBox="[151,1436,181,1574]" targetPageId="17" targetType="figure">
<paragraph id="8BF09143FF800007FF7BF980FF0A34D9" blockId="17.[151,1436,1597,1694]" pageId="17" pageNumber="168">
<emphasis id="B93B4D51FF800007FF7BF980FEFA3412" bold="true" box="[151,289,1597,1621]" pageId="17" pageNumber="168">FIGURE 11.</emphasis>
<taxonomicName id="4C4FEAC0FF800007FEC6F983FE3B3411" authority="Soares &amp; Runyon &amp; Capellari &amp; Ale-Rocha, 2023" authorityName="Soares &amp; Runyon &amp; Capellari &amp; Ale-Rocha" authorityYear="2023" box="[298,480,1598,1622]" class="Insecta" family="Dolichopodidae" genus="Paraclius" kingdom="Animalia" order="Diptera" pageId="17" pageNumber="168" phylum="Arthropoda" rank="species" species="brooksi" status="sp. nov.">
<emphasis id="B93B4D51FF800007FEC6F983FE3B3411" box="[298,480,1598,1622]" italics="true" pageId="17" pageNumber="168">Paraclius brooksi</emphasis>
</taxonomicName>
<emphasis id="B93B4D51FF800007FE05F983FDE73411" bold="true" box="[489,572,1598,1622]" pageId="17" pageNumber="168">
<taxonomicNameLabel id="A208F02AFF800007FE05F983FDE73411" box="[489,572,1598,1622]" pageId="17" pageNumber="168" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
, 
<emphasis id="B93B4D51FF800007FDA0F980FDA73412" bold="true" box="[588,636,1597,1621]" pageId="17" pageNumber="168">A–E</emphasis>
male holotype (CNC), 
<emphasis id="B93B4D51FF800007FC99F983FC5F3411" bold="true" box="[885,900,1598,1622]" pageId="17" pageNumber="168">F</emphasis>
male paratype (CNC). 
<emphasis id="B93B4D51FF800007FB90F980FB4E3412" bold="true" box="[1148,1173,1597,1621]" pageId="17" pageNumber="168">A.</emphasis>
Habitus, lateral view; 
<emphasis id="B93B4D51FF800007FA69F983FA473411" bold="true" box="[1413,1436,1598,1622]" pageId="17" pageNumber="168">B.</emphasis>
Thorax, dorsal view; 
<emphasis id="B93B4D51FF800007FE98F9DCFE77343E" bold="true" box="[372,428,1633,1657]" pageId="17" pageNumber="168">C–D.</emphasis>
Head anterior and lateral views, respectively; 
<emphasis id="B93B4D51FF800007FC60F9DFFC7F343D" bold="true" box="[908,932,1634,1658]" pageId="17" pageNumber="168">E.</emphasis>
Abdomen and hypopygium, lateral view; 
<emphasis id="B93B4D51FF800007FABBF9DFFAB1343D" bold="true" box="[1367,1386,1634,1658]" pageId="17" pageNumber="168">F.</emphasis>
Left wing.
</paragraph>
</caption>
<subSubSection id="C355C2C8FF800007FF2BF94DFEBB35F9" pageId="17" pageNumber="168" type="discussion">
<paragraph id="8BF09143FF800007FF2BF94DFEBB35F9" blockId="17.[151,1436,1776,2018]" pageId="17" pageNumber="168">
<emphasis id="B93B4D51FF800007FF2BF94DFEE0354D" bold="true" box="[199,315,1776,1802]" pageId="17" pageNumber="168">Remarks.</emphasis>
<taxonomicName id="4C4FEAC0FF800007FEA9F94DFDD5354D" authority="Soares &amp; Runyon &amp; Capellari &amp; Ale-Rocha, 2023" authorityName="Soares &amp; Runyon &amp; Capellari &amp; Ale-Rocha" authorityYear="2023" box="[325,526,1776,1802]" class="Insecta" family="Dolichopodidae" genus="Paraclius" kingdom="Animalia" order="Diptera" pageId="17" pageNumber="168" phylum="Arthropoda" rank="species" species="brooksi" status="sp. nov.">
<emphasis id="B93B4D51FF800007FEA9F94DFDD5354D" box="[325,526,1776,1802]" italics="true" pageId="17" pageNumber="168">Paraclius brooksi</emphasis>
</taxonomicName>
<emphasis id="B93B4D51FF800007FDFBF94DFDA8354D" bold="true" box="[535,627,1776,1802]" pageId="17" pageNumber="168">
<taxonomicNameLabel id="A208F02AFF800007FDFBF94DFDA8354D" box="[535,627,1776,1802]" pageId="17" pageNumber="168" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
possesses mostly yellow legs and male tergite 6 with long and strong setae, similar to 
<taxonomicName id="4C4FEAC0FF800007FEBEF8A8FDD13569" authorityName="Van Duzee" authorityYear="1933" box="[338,522,1812,1838]" class="Insecta" family="Dolichopodidae" genus="Paraclius" kingdom="Animalia" order="Diptera" pageId="17" pageNumber="168" phylum="Arthropoda" rank="species" species="angusticauda">
<emphasis id="B93B4D51FF800007FEBEF8A8FEB83569" box="[338,355,1813,1838]" italics="true" pageId="17" pageNumber="168">P</emphasis>
. 
<emphasis id="B93B4D51FF800007FE9FF8A9FDD13569" box="[371,522,1812,1838]" italics="true" pageId="17" pageNumber="168">angusticauda</emphasis>
</taxonomicName>
and 
<taxonomicName id="4C4FEAC0FF800007FDAEF8A8FD0B3569" authorityName="Van Duzee" authorityYear="1930" box="[578,720,1812,1838]" class="Insecta" family="Dolichopodidae" genus="Paraclius" kingdom="Animalia" order="Diptera" pageId="17" pageNumber="168" phylum="Arthropoda" rank="species" species="elongatus">
<emphasis id="B93B4D51FF800007FDAEF8A8FD883569" box="[578,595,1813,1838]" italics="true" pageId="17" pageNumber="168">P</emphasis>
. 
<emphasis id="B93B4D51FF800007FD8FF8A9FD0B3569" box="[611,720,1812,1838]" italics="true" pageId="17" pageNumber="168">elongatus</emphasis>
</taxonomicName>
, but can be differentiated by the brownish wing (
<figureCitation id="13748DC6FF800007FAEEF8A9FABF3569" box="[1282,1380,1812,1838]" captionStart="FIGURE 11" captionStartId="17.[151,250,1597,1621]" captionTargetBox="[151,1436,181,1574]" captionTargetId="figure-18@17.[151,1436,181,1574]" captionTargetPageId="17" captionText="FIGURE 11. Paraclius brooksi sp. nov., A–E male holotype (CNC), F male paratype (CNC). A. Habitus, lateral view; B. Thorax, dorsal view; C–D. Head anterior and lateral views, respectively; E. Abdomen and hypopygium, lateral view; F. Left wing." figureDoi="http://doi.org/10.5281/zenodo.8204135" httpUri="https://zenodo.org/record/8204135/files/figure.png" pageId="17" pageNumber="168">Fig. 11F</figureCitation>
) and long basal apicoventral epandrial seta which is about 2X longer than apical seta (
<figureCitation id="13748DC6FF800007FBD8F885FB403515" box="[1076,1179,1848,1874]" captionStart="FIGURE12" captionStartId="18.[151,250,1647,1671]" captionTargetBox="[151,1436,237,1624]" captionTargetId="figure-42@18.[151,1436,237,1624]" captionTargetPageId="18" captionText="FIGURE12.Paraclius brooksisp.nov., male paratype (CNC).A. Abdomen, ventralview;B. Cerci, dorsalview;C. Hypopygium, left lateral view; D. Surstyli and apical edge of epandrium; E. Internal appendages; F. Hypandrium and phallus, ventral view; G. Tergite 7, dorsal view. Abreviations: a apv s = apical apicoventral epandrial seta; b apv s = basal apicoventral epandrial seta; cerc = cercus; dsur = dorsal lobe of surstylus; ej apod = ejaculatory apodeme; epand = epandrium; hypd = hypandrium; hypd pr = hypandrial process; pgt = postgonite; ph = phallus; ph pr = phallus process; prct = proctiger; sp = sperm pump; st = sternite; tg = tergite; vsur = ventral lobe of surstylus." figureDoi="http://doi.org/10.5281/zenodo.8204139" httpUri="https://zenodo.org/record/8204139/files/figure.png" pageId="17" pageNumber="168">Fig. 12D</figureCitation>
) (wing mostly grayish (
<figureCitation id="13748DC6FF800007FF73F8E1FF313531" box="[159,234,1884,1910]" captionStart="FIGURE 1" captionStartId="4.[151,250,1581,1605]" captionTargetBox="[151,1436,306,1555]" captionTargetId="figure-87@4.[151,1436,306,1555]" captionTargetPageId="4" captionText="FIGURE 1. Paraclius angusticauda Van Duzee, 1933, male holotype (AMNH). A–B. Habitus, lateral and dorsal views respectively; C. Labels; D–E. Head, anterior and lateral views, respectively; F–H. Hypopygium, lateral view; I. Right wing. Abbreviations: a apv s = apical apicoventral epandrial seta; b apv s = basal apicoventral epandrial seta; cerc = cercus; dm-m = discal medial crossvein; M1 = first branch of media; M4 = fourth branch of media; R 4+5 = third branch of radius. Images originally lacked scale bars. Photographs by Agnieszka Pierwola, AMNH. " figureDoi="http://doi.org/10.5281/zenodo.8204105" httpUri="https://zenodo.org/record/8204105/files/figure.png" pageId="17" pageNumber="168">Fig. 1I</figureCitation>
) and basal apicoventral epandrial seta as long as apical seta in 
<taxonomicName id="4C4FEAC0FF800007FC72F8E0FB883531" authorityName="Van Duzee" authorityYear="1933" box="[926,1107,1884,1910]" class="Insecta" family="Dolichopodidae" genus="Paraclius" kingdom="Animalia" order="Diptera" pageId="17" pageNumber="168" phylum="Arthropoda" rank="species" species="angusticauda">
<emphasis id="B93B4D51FF800007FC72F8E0FC743531" box="[926,943,1885,1910]" italics="true" pageId="17" pageNumber="168">P</emphasis>
. 
<emphasis id="B93B4D51FF800007FC50F8E1FB883531" box="[956,1107,1884,1910]" italics="true" pageId="17" pageNumber="168">angusticauda</emphasis>
</taxonomicName>
(
<figureCitation id="13748DC6FF800007FB8EF8E1FB6C3531" box="[1122,1207,1884,1910]" captionStart="FIGURE 1" captionStartId="4.[151,250,1581,1605]" captionTargetBox="[151,1436,306,1555]" captionTargetId="figure-87@4.[151,1436,306,1555]" captionTargetPageId="4" captionText="FIGURE 1. Paraclius angusticauda Van Duzee, 1933, male holotype (AMNH). A–B. Habitus, lateral and dorsal views respectively; C. Labels; D–E. Head, anterior and lateral views, respectively; F–H. Hypopygium, lateral view; I. Right wing. Abbreviations: a apv s = apical apicoventral epandrial seta; b apv s = basal apicoventral epandrial seta; cerc = cercus; dm-m = discal medial crossvein; M1 = first branch of media; M4 = fourth branch of media; R 4+5 = third branch of radius. Images originally lacked scale bars. Photographs by Agnieszka Pierwola, AMNH. " figureDoi="http://doi.org/10.5281/zenodo.8204105" httpUri="https://zenodo.org/record/8204105/files/figure.png" pageId="17" pageNumber="168">Fig. 1G</figureCitation>
)). 
<taxonomicName id="4C4FEAC0FF800007FB3BF8E1FA473531" authority="Soares &amp; Runyon &amp; Capellari &amp; Ale-Rocha, 2023" authorityName="Soares &amp; Runyon &amp; Capellari &amp; Ale-Rocha" authorityYear="2023" box="[1239,1436,1884,1910]" class="Insecta" family="Dolichopodidae" genus="Paraclius" kingdom="Animalia" order="Diptera" pageId="17" pageNumber="168" phylum="Arthropoda" rank="species" species="brooksi" status="sp. nov.">
<emphasis id="B93B4D51FF800007FB3BF8E1FA473531" box="[1239,1436,1884,1910]" italics="true" pageId="17" pageNumber="168">Paraclius brooksi</emphasis>
</taxonomicName>
<emphasis id="B93B4D51FF800007FF7BF83DFF2935DD" bold="true" box="[151,242,1920,1946]" pageId="17" pageNumber="168">
<taxonomicNameLabel id="A208F02AFF800007FF7BF83DFF2935DD" box="[151,242,1920,1946]" pageId="17" pageNumber="168" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
was previously mentioned by Aldrich (1902) as a “light variety” of 
<taxonomicName id="4C4FEAC0FF800007FC17F83CFBA735DD" authorityName=": Loew" authorityYear="1864" box="[1019,1148,1920,1946]" class="Insecta" family="Dolichopodidae" genus="Paraclius" kingdom="Animalia" order="Diptera" pageId="17" pageNumber="168" phylum="Arthropoda" rank="species" species="arcuatus">
<emphasis id="B93B4D51FF800007FC17F83CFBD735DD" box="[1019,1036,1921,1946]" italics="true" pageId="17" pageNumber="168">P</emphasis>
. 
<emphasis id="B93B4D51FF800007FBF7F83CFBA735DD" box="[1051,1148,1921,1946]" italics="true" pageId="17" pageNumber="168">arcuatus</emphasis>
</taxonomicName>
and by 
<bibRefCitation id="EFDEECB2FF800007FB35F83DFAA635DC" author="Brooks, S. E." box="[1241,1405,1920,1947]" pageId="17" pageNumber="168" pagination="1 - 158" refId="ref21534" refString="Brooks, S. E. (2005) Systematics and phylogeny of Dolichopodinae (Diptera: Dolichopodidae). Zootaxa, 857 (1), 1 - 158. https: // doi. org / 10.11646 / zootaxa. 857.1.1" type="journal article" year="2005">Brooks (2005)</bibRefCitation>
as “ 
<taxonomicName id="4C4FEAC0FF800007FF48F819FE9035F9" box="[164,331,1956,1982]" class="Insecta" family="Dolichopodidae" genus="Paraclius" kingdom="Animalia" order="Diptera" pageId="17" pageNumber="168" phylum="Arthropoda" rank="species" species="undefined-1">
<emphasis id="B93B4D51FF800007FF48F819FECA35F9" box="[164,273,1956,1982]" italics="true" pageId="17" pageNumber="168">Paraclius</emphasis>
sp. 1
</taxonomicName>
”.
</paragraph>
</subSubSection>
<subSubSection id="C355C2C8FF800004FF2BF875FEB63292" lastPageId="18" lastPageNumber="169" pageId="17" pageNumber="168" type="etymology">
<paragraph id="8BF09143FF800004FF2BF875FEB63292" blockId="17.[151,1436,1776,2018]" lastBlockId="18.[151,1436,151,213]" lastPageId="18" lastPageNumber="169" pageId="17" pageNumber="168">
<emphasis id="B93B4D51FF800007FF2BF875FE9535A5" bold="true" box="[199,334,1992,2018]" pageId="17" pageNumber="168">Etymology.</emphasis>
Named after Scott Brooks (CNC), who has contributed substantially to the study of 
<taxonomicName id="4C4FEAC0FF800007FB05F875FA4735A5" box="[1257,1436,1992,2018]" class="Insecta" family="Dolichopodidae" higherTaxonomySource="GBIF" kingdom="Animalia" order="Diptera" pageId="17" pageNumber="168" phylum="Arthropoda" rank="subFamily" subFamily="Dolichopodinae">Dolichopodinae</taxonomicName>
worldwide, and for his willingness to separate and prepare specimens for loan, including those used in the description of the new species.
</paragraph>
</subSubSection>
<caption id="DF30C1CBFF830004FF7BF9D2FD8E357C" ID-DOI="http://doi.org/10.5281/zenodo.8204139" ID-Zenodo-Dep="8204139" httpUri="https://zenodo.org/record/8204139/files/figure.png" pageId="18" pageNumber="169" startId="18.[151,250,1647,1671]" targetBox="[151,1436,237,1624]" targetPageId="18" targetType="figure">
<paragraph id="8BF09143FF830004FF7BF9D2FD8E357C" blockId="18.[151,1437,1647,1852]" pageId="18" pageNumber="169">
<emphasis id="B93B4D51FF830004FF7BF9D2FEC634C0" bold="true" box="[151,285,1647,1671]" pageId="18" pageNumber="169">FIGURE 12.</emphasis>
<taxonomicName id="4C4FEAC0FF830004FECCF9D2FE0934C0" authority="Soares &amp; Runyon &amp; Capellari &amp; Ale-Rocha, 2023" authorityName="Soares &amp; Runyon &amp; Capellari &amp; Ale-Rocha" authorityYear="2023" box="[288,466,1647,1671]" class="Insecta" family="Dolichopodidae" genus="Paraclius" kingdom="Animalia" order="Diptera" pageId="18" pageNumber="169" phylum="Arthropoda" rank="species" species="brooksi" status="sp. nov.">
<emphasis id="B93B4D51FF830004FECCF9D2FE0934C0" box="[288,466,1647,1671]" italics="true" pageId="18" pageNumber="169">Paraclius brooksi</emphasis>
</taxonomicName>
<emphasis id="B93B4D51FF830004FE39F9D2FDF834C0" bold="true" box="[469,547,1647,1671]" pageId="18" pageNumber="169">
<taxonomicNameLabel id="A208F02AFF830004FE39F9D2FDF834C0" box="[469,547,1647,1671]" pageId="18" pageNumber="169" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
, male paratype (CNC). 
<emphasis id="B93B4D51FF830004FCE0F9D2FCFF34C0" bold="true" box="[780,804,1647,1671]" pageId="18" pageNumber="169">A.</emphasis>
Abdomen, ventral view; 
<emphasis id="B93B4D51FF830004FBF7F9D2FBE934C0" bold="true" box="[1051,1074,1647,1671]" pageId="18" pageNumber="169">B.</emphasis>
Cerci, dorsal view; 
<emphasis id="B93B4D51FF830004FB18F9D2FAD734C0" bold="true" box="[1268,1292,1647,1671]" pageId="18" pageNumber="169">C.</emphasis>
Hypopygium, left lateral view; 
<emphasis id="B93B4D51FF830004FEAAF92EFE8434EC" bold="true" box="[326,351,1683,1707]" pageId="18" pageNumber="169">D.</emphasis>
Surstyli and apical edge of epandrium; 
<emphasis id="B93B4D51FF830004FD15F92EFCCA34EC" bold="true" box="[761,785,1683,1707]" pageId="18" pageNumber="169">E.</emphasis>
Internal appendages; 
<emphasis id="B93B4D51FF830004FC1EF92EFBDE34EC" bold="true" box="[1010,1029,1683,1707]" pageId="18" pageNumber="169">F.</emphasis>
Hypandrium and phallus, ventral view; 
<emphasis id="B93B4D51FF830004FF7BF90AFF6A3488" bold="true" box="[151,177,1719,1743]" pageId="18" pageNumber="169">G.</emphasis>
Tergite 7, dorsal view. Abreviations: a apv s = apical apicoventral epandrial seta; b apv s = basal apicoventral epandrial seta; cerc = cercus; dsur = dorsal lobe of surstylus; ej apod = ejaculatory apodeme; epand = epandrium; hypd = hypandrium; hypd pr = hypandrial process; pgt = postgonite; ph = phallus; ph pr = phallus process; prct = proctiger; sp = sperm pump; st = sternite; tg = tergite; vsur = ventral lobe of surstylus.
</paragraph>
</caption>
<subSubSection id="C355C2C8FF830004FF2BF830FF1F358B" pageId="18" pageNumber="169" type="distribution">
<paragraph id="8BF09143FF830004FF2BF830FA4635E0" blockId="18.[151,1437,1933,1996]" box="[199,1437,1933,1960]" pageId="18" pageNumber="169">
<emphasis id="B93B4D51FF830004FF2BF830FEBA35E0" bold="true" box="[199,353,1933,1959]" pageId="18" pageNumber="169">Distribution.</emphasis>
<taxonomicName id="4C4FEAC0FF830004FE8BF830FDF635E0" authority="Soares &amp; Runyon &amp; Capellari &amp; Ale-Rocha, 2023" authorityName="Soares &amp; Runyon &amp; Capellari &amp; Ale-Rocha" authorityYear="2023" box="[359,557,1933,1959]" class="Insecta" family="Dolichopodidae" genus="Paraclius" kingdom="Animalia" order="Diptera" pageId="18" pageNumber="169" phylum="Arthropoda" rank="species" species="brooksi" status="sp. nov.">
<emphasis id="B93B4D51FF830004FE8BF830FDF635E0" box="[359,557,1933,1959]" italics="true" pageId="18" pageNumber="169">Paraclius brooksi</emphasis>
</taxonomicName>
<emphasis id="B93B4D51FF830004FDDFF833FD5735EF" bold="true" box="[563,652,1934,1960]" pageId="18" pageNumber="169">
<taxonomicNameLabel id="A208F02AFF830004FDDFF833FD5735EF" box="[563,652,1934,1960]" pageId="18" pageNumber="169" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
is known to occur in 
<collectingCountry id="F358D1D3FF830004FC97F830FC0135E0" box="[891,986,1933,1959]" name="Grenada" pageId="18" pageNumber="169">Grenada</collectingCountry>
and 
<collectingCountry id="F358D1D3FF830004FBE3F830FBBE35E0" box="[1039,1125,1933,1959]" name="Mexico" pageId="18" pageNumber="169">Mexico</collectingCountry>
(
<collectingRegion id="498B5FA1FF830004FB9EF830FB1535E0" box="[1138,1230,1933,1959]" country="Mexico" name="Chiapas" pageId="18" pageNumber="169">Chiapas</collectingRegion>
and 
<collectingRegion id="498B5FA1FF830004FAEFF833FA8235EF" box="[1283,1369,1934,1960]" country="Mexico" name="Oaxaca" pageId="18" pageNumber="169">Oaxaca</collectingRegion>
) (
<figureCitation id="13748DC6FF830004FA83F830FA4635E0" box="[1391,1437,1933,1959]" captionStart="FIGURE 23" captionStartId="29.[151,250,1033,1057]" captionTargetBox="[151,1436,237,1009]" captionTargetId="figure-36@29.[151,1436,237,1009]" captionTargetPageId="29" captionText="FIGURE 23. Known geographic distribution of Paraclius angusticauda Van Duzee, 1933, P. angustipennis Van Duzee, 1929, P. brooksi sp. nov., P. elongatus Van Duzee, 1930 and P. xibun sp. nov." figureDoi="http://doi.org/10.5281/zenodo.8204175" httpUri="https://zenodo.org/record/8204175/files/figure.png" pageId="18" pageNumber="169">Fig.</figureCitation>
</paragraph>
<paragraph id="8BF09143FF830004FF7BF80FFF1F358B" blockId="18.[151,1437,1933,1996]" box="[151,196,1970,1996]" pageId="18" pageNumber="169">23).</paragraph>
</subSubSection>
</treatment>
</document>