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<mods:title id="6E2E5424ACCB4D4F4F5DB966E03FCFC5">An illustrated guide to lady beetles (Coleoptera: Coccinellidae) of the Indian Subcontinent. Part 1. Tribe Coccinellini</mods:title>
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<paragraph id="8BD1F074143D4652BDDF3244FDEBFBC8" blockId="101.[151,550,1080,1143]" box="[151,550,1080,1107]" pageId="101" pageNumber="102">
<heading id="D0994718143D4652BDDF3244FDEBFBC8" bold="true" box="[151,550,1080,1107]" fontSize="11" level="2" pageId="101" pageNumber="102" reason="4">
<taxonomicName id="4C6E8BF7143D4652BDDF3244FDEBFBC8" authority="(Hope)" baseAuthorityName="Hope" box="[151,550,1080,1107]" class="Insecta" family="Coccinellidae" genus="Coelophora" kingdom="Animalia" order="Coleoptera" pageId="101" pageNumber="102" phylum="Arthropoda" rank="species" species="circumvelata">
<emphasis id="B91A2C66143D4652BDDF3244FDEBFBC8" bold="true" box="[151,550,1080,1107]" pageId="101" pageNumber="102">
<emphasis id="B91A2C66143D4652BDDF3244FE05FBC8" bold="true" box="[151,456,1080,1107]" italics="true" pageId="101" pageNumber="102">Coelophora circumvelata</emphasis>
(Hope)
</emphasis>
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</heading>
</paragraph>
</subSubSection>
<subSubSection id="C374A3FF143D4652BDDF3221FF34FBEC" box="[151,249,1117,1143]" pageId="101" pageNumber="102" type="description">
<paragraph id="8BD1F074143D4652BDDF3221FF34FBEC" blockId="101.[151,550,1080,1143]" box="[151,249,1117,1143]" pageId="101" pageNumber="102">
(
<figureCitation id="1355ECF1143D4652BDD73221FF3CFBEC" box="[159,241,1117,1143]" captionStart="FIGURE 72" captionStartId="108.[151,250,1943,1967]" captionTargetBox="[265,1316,182,1918]" captionTargetId="figure-20@108.[245,1342,181,1920]" captionTargetPageId="108" captionText="FIGURE 72. Coelophora circumvelata (Mulsant): a. Type, dorsal view (BMNH); b, d. adult, dorsal view; c. adult, lateral view; e. adult, frontal view; f, g. female genitalia; h. spermatheca (enlarged)." figureDoi="http://doi.org/10.5281/zenodo.8261737" httpUri="https://zenodo.org/record/8261737/files/figure.png" pageId="101" pageNumber="102">Fig. 72</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="C374A3FF143D4652BDDF32DDFBF4FAEA" pageId="101" pageNumber="102" type="reference_group">
<paragraph id="8BD1F074143D4652BDDF32DDFE7AFB43" blockId="101.[151,1436,1185,1394]" pageId="101" pageNumber="102">
<treatmentCitationGroup id="AB7ED75A143D4652BDDF32DDFA51FB22" box="[151,1436,1185,1210]" pageId="101" pageNumber="102">
<treatmentCitation id="0ACFD665143D4652BDDF32DDFE3DFB21" author="Hope, F. W." box="[151,496,1185,1210]" page="31" pageId="101" pageNumber="102" year="1831">
<taxonomicName id="4C6E8BF7143D4652BDDF32DDFE3DFB21" authority="Hope, 1831: 31" authorityName="Hope" authorityPageNumber="31" authorityYear="1831" box="[151,496,1185,1210]" class="Insecta" family="Coccinellidae" genus="Coccinella" kingdom="Animalia" order="Coleoptera" pageId="101" pageNumber="102" phylum="Arthropoda" rank="species" species="cincta">
<emphasis id="B91A2C66143D4652BDDF32DDFE87FB22" box="[151,330,1185,1209]" italics="true" pageId="101" pageNumber="102">Coccinella cincta</emphasis>
<bibRefCitation id="EFFF8D85143D4652BC1932DEFE3DFB21" author="Hope, F. W." box="[337,496,1186,1210]" pageId="101" pageNumber="102" pagination="21 - 32" refId="ref113751" refString="Hope, F. W. (1831) s. n. In: Gray, J. E., Synopsis of the new species of Nepaul insects in the collection of Major General Hardwicke. The Zoological Miscellany, London, pp. 21 - 32." type="book chapter" year="1831">Hope, 1831: 31</bibRefCitation>
</taxonomicName>
</treatmentCitation>
(preoccupied) (
<typeStatus id="54D54ED6143D4652BFDA32DEFD36FB21" box="[658,763,1186,1210]" pageId="101" pageNumber="102" type="lectotype">Lectotype</typeStatus>
, HEC, OMNH; Type locality:
<collectingCountry id="F379B0E4143D4652B97E32DDFBB8FB22" box="[1078,1141,1185,1209]" name="Nepal" pageId="101" pageNumber="102">Nepal</collectingCountry>
).—Booth &amp;
<treatmentCitation id="0ACFD665143D4652B9B332DEFA51FB22" author="Pope, R. D." box="[1275,1436,1185,1210]" page="350" pageId="101" pageNumber="102" year="1989">
<bibRefCitation id="EFFF8D85143D4652B9B332DEFA51FB22" author="Pope, R. D." box="[1275,1436,1185,1210]" pageId="101" pageNumber="102" pagination="633 - 735" refId="ref118886" refString="Pope, R. D. (1989) A revision of the Australian Coccinellidae (Coleoptera). Part I. Subfamily Coccinellinae. Invertebrate Taxonomy, 3, 633 - 735. https: // doi. org / 10.1071 / IT 9880633" type="journal article" year="1989">Pope 1989: 350</bibRefCitation>
</treatmentCitation>
</treatmentCitationGroup>
(
<typeStatus id="54D54ED6143D4652BD8632BCFEE0FB43" box="[206,301,1216,1240]" pageId="101" pageNumber="102" type="lectotype">lectotype</typeStatus>
designation).
</paragraph>
<paragraph id="8BD1F074143D4652BDDF329CFE20FB6C" blockId="101.[151,1436,1185,1394]" box="[151,493,1247,1271]" pageId="101" pageNumber="102">
<taxonomicName id="4C6E8BF7143D4652BDDF329CFE24FB6C" authority=": Crotch 1874: 149" authorityName="Crotch" authorityPageNumber="149" authorityYear="1874" box="[151,489,1247,1271]" class="Insecta" family="Coccinellidae" genus="Lemnia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="101" pageNumber="102" phylum="Arthropoda" rank="species" species="cincta">
<emphasis id="B91A2C66143D4652BDDF329CFEE5FB6C" box="[151,296,1248,1271]" italics="true" pageId="101" pageNumber="102">Lemnia cincta</emphasis>
: Crotch 1874: 149
</taxonomicName>
.
</paragraph>
<paragraph id="8BD1F074143D4652BDDF3282FDDDFAAF" blockId="101.[151,1436,1185,1394]" pageId="101" pageNumber="102">
<taxonomicName id="4C6E8BF7143D4652BDDF3282FDC3FA8E" authority="Mulsant, 1850" authorityName="Mulsant" authorityYear="1850" box="[151,526,1277,1301]" class="Insecta" family="Coccinellidae" genus="Lemnia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="101" pageNumber="102" phylum="Arthropoda" rank="species" species="circumvelata">
<emphasis id="B91A2C66143D4652BDDF3282FEA3FA8E" box="[151,366,1277,1301]" italics="true" pageId="101" pageNumber="102">Lemnia circumvelata</emphasis>
<bibRefCitation id="EFFF8D85143D4652BC3E3281FDC3FA8E" author="Mulsant, E." box="[374,526,1277,1301]" pageId="101" pageNumber="102" pagination="1 - 1104" refId="ref117842" refString="Mulsant, E. (1850) Species des Coleopteres Trimeres Securipalpes. Annales des Sciences Physiques et Naturelles, d'Agriculture et d'Industrie, publiees par la Societe nationale d'Agriculture, etc., de Lyon, Deuxieme Serie, 2, 1 - 1104. https: // doi. org / 10.5962 / bhl. title. 8953" type="journal article" year="1850">Mulsant, 1850</bibRefCitation>
</taxonomicName>
; 387, 388 (replacement name); 1866: 255;
<bibRefCitation id="EFFF8D85143D4652BE843281FB72FA8D" author="Korschefsky, R." box="[972,1215,1277,1302]" pageId="101" pageNumber="102" refId="ref116133" refString="Korschefsky, R. (1932) Coleopterorum Catalogus. Pars 120. Coccinellidae II. W. Junk, Berlin, 435 pp." type="book" year="1932">Korschefsky 1932: 291</bibRefCitation>
;
<bibRefCitation id="EFFF8D85143D4652B9833281FA51FA8E" author="Iablokoff-Khnzorian, S. M." box="[1227,1436,1277,1301]" pageId="101" pageNumber="102" refId="ref114042" refString="Iablokoff-Khnzorian, S. M. (1982) Les Coccinelles Coleopteres-Coccinellidae. Tribu Coccinellini des regions Palearctique et Orientale. Boubee, Paris, 568 pp. [in French]" type="book" year="1982">Iablokoff-Khnzorian</bibRefCitation>
1982: 220;
<bibRefCitation id="EFFF8D85143D4652BC733361FDC1FAAF" author="Ren, S. X. &amp; Wang, X. M. &amp; Pang, H. &amp; Peng, Z. Q. &amp; Zeng, T." box="[315,524,1308,1332]" pageId="101" pageNumber="102" refId="ref120760" refString="Ren, S. X., Wang, X. M., Pang, H., Peng, Z. Q. &amp; Zeng, T. (2009) Colored pictorial handbook of ladybird beetles in China. Science Press, Beijing, 336 pp. [in Chinese]" type="book" year="2009">
Ren
<emphasis id="B91A2C66143D4652BC223361FE50FAAF" box="[362,413,1308,1332]" italics="true" pageId="101" pageNumber="102">et al.</emphasis>
2009: 206
</bibRefCitation>
.
</paragraph>
<paragraph id="8BD1F074143D4652BDDF3340FC9BFAC8" blockId="101.[151,1436,1185,1394]" box="[151,854,1339,1363]" pageId="101" pageNumber="102">
<taxonomicName id="4C6E8BF7143D4652BDDF3340FC9FFAC8" authority=": Iablokoff-Khnzorian 1984: 210" authorityName="Iablokoff-Khnzorian" authorityPageNumber="210" authorityYear="1984" box="[151,850,1339,1363]" class="Insecta" family="Coccinellidae" genus="Lemnia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="101" pageNumber="102" phylum="Arthropoda" rank="species" species="circumvelata" subGenus="Phrynocaria">
<emphasis id="B91A2C66143D4652BDDF3340FF2EFAC8" box="[151,227,1340,1363]" italics="true" pageId="101" pageNumber="102">Lemnia</emphasis>
(
<emphasis id="B91A2C66143D4652BDBA3347FEBFFAC8" box="[242,370,1339,1363]" italics="true" pageId="101" pageNumber="102">Phrynocaria</emphasis>
)
<emphasis id="B91A2C66143D4652BCC93347FDC9FAC8" box="[385,516,1339,1363]" italics="true" pageId="101" pageNumber="102">circumvelata</emphasis>
:
<bibRefCitation id="EFFF8D85143D4652BF593347FC9FFAC8" author="Iablokoff-Khnzorian, S. M." box="[529,850,1339,1363]" pageId="101" pageNumber="102" pagination="203 - 220" refId="ref114074" refString="Iablokoff-Khnzorian, S. M. (1984) Notes sur la tribu des Coccinellini (Coleoptera: Coccinellidae). Nouvelle Revue d'Entomologie, New Series, 1 (2), 203 - 220." type="journal article" year="1984">Iablokoff-Khnzorian 1984: 210</bibRefCitation>
</taxonomicName>
.
</paragraph>
<paragraph id="8BD1F074143D4652BDDF3325FBF4FAEA" blockId="101.[151,1436,1185,1394]" box="[151,1081,1369,1394]" pageId="101" pageNumber="102">
<taxonomicName id="4C6E8BF7143D4652BDDF3325FD65FAEA" authority=": Booth &amp; Pope 1989: 350" authorityName=": Booth &amp; Pope" authorityPageNumber="350" authorityYear="1989" box="[151,680,1369,1394]" class="Insecta" family="Coccinellidae" genus="Coelophora" kingdom="Animalia" order="Coleoptera" pageId="101" pageNumber="102" phylum="Arthropoda" rank="species" species="circumvelata">
<emphasis id="B91A2C66143D4652BDDF3325FE57FAEA" box="[151,410,1369,1393]" italics="true" pageId="101" pageNumber="102">Coelophora circumvelata</emphasis>
: Booth &amp;
<bibRefCitation id="EFFF8D85143D4652BF4E3326FD65FAEA" author="Pope, R. D." box="[518,680,1369,1394]" pageId="101" pageNumber="102" pagination="633 - 735" refId="ref118886" refString="Pope, R. D. (1989) A revision of the Australian Coccinellidae (Coleoptera). Part I. Subfamily Coccinellinae. Invertebrate Taxonomy, 3, 633 - 735. https: // doi. org / 10.1071 / IT 9880633" type="journal article" year="1989">Pope 1989: 350</bibRefCitation>
</taxonomicName>
;
<bibRefCitation id="EFFF8D85143D4652BFFA3325FCB6FAE9" author="Poorani, J." box="[690,891,1369,1394]" pageId="101" pageNumber="102" pagination="307 - 383" refId="ref118931" refString="Poorani, J. (2002 a) An annotated checklist of the Coccinellidae (excluding Epilachninae) of the Indian subregion. Oriental Insects, 36, 307 - 383. https: // doi. org / 10.1080 / 00305316.2002.10417335" type="journal article" year="2002">Poorani 2002a: 328</bibRefCitation>
;
<bibRefCitation id="EFFF8D85143D4652BECE3326FBF9FAEA" author="Kovar, I." box="[902,1076,1369,1394]" pageId="101" pageNumber="102" pagination="1807" refId="ref116365" refString="Kovar, I. (2007) Family Coccinellidae Latreille, 1807. In: L ˆ bl, I. &amp; Smetana, A. (Eds.), Catalogue of Palaearctic Coleoptera. Vol. 4. Apollo Books, Stenstrup, pp. 568 - 631." type="book chapter" year="2007">Kovář 2007: 614</bibRefCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="C374A3FF143D4652BDDF33E3FAD6F9F5" pageId="101" pageNumber="102" type="diagnosis">
<paragraph id="8BD1F074143D4652BDDF33E3FAD6F9F5" blockId="101.[151,1437,1439,2042]" pageId="101" pageNumber="102">
<emphasis id="B91A2C66143D4652BDDF33E3FEDFFA22" bold="true" box="[151,274,1439,1465]" pageId="101" pageNumber="102">Diagnosis.</emphasis>
Length: 6.00
<quantity id="4C965D91143D4652BCF233E3FDEFFA21" box="[442,546,1439,1466]" metricMagnitude="-3" metricUnit="m" metricValue="6.5" pageId="101" pageNumber="102" unit="mm" value="6.5">6.50 mm</quantity>
. Form almost circular, dorsum convex and glabrous. Head yellow, eyes large, narrowly separated and apically strongly divergent; pronotum yellow with a large black macula occupying most of the disc; scutellar shield black; elytra yellow, lateral margins with a black border, about 1/4th of width of elytron (
<figureCitation id="1355ECF1143D4652B8273394FF13F9BD" captionStart="FIGURE 72" captionStartId="108.[151,250,1943,1967]" captionTargetBox="[265,1316,182,1918]" captionTargetId="figure-20@108.[245,1342,181,1920]" captionTargetPageId="108" captionText="FIGURE 72. Coelophora circumvelata (Mulsant): a. Type, dorsal view (BMNH); b, d. adult, dorsal view; c. adult, lateral view; e. adult, frontal view; f, g. female genitalia; h. spermatheca (enlarged)." figureDoi="http://doi.org/10.5281/zenodo.8261737" httpUri="https://zenodo.org/record/8261737/files/figure.png" pageId="101" pageNumber="102">Fig. 72ac</figureCitation>
). Ventral side more or less yellowish except metaventrite and abdominal ventrite 1 medially darker. Female genitalia (
<figureCitation id="1355ECF1143D4652BC4E304CFEACF9D1" box="[262,353,1584,1610]" captionStart="FIGURE 72" captionStartId="108.[151,250,1943,1967]" captionTargetBox="[265,1316,182,1918]" captionTargetId="figure-20@108.[245,1342,181,1920]" captionTargetPageId="108" captionText="FIGURE 72. Coelophora circumvelata (Mulsant): a. Type, dorsal view (BMNH); b, d. adult, dorsal view; c. adult, lateral view; e. adult, frontal view; f, g. female genitalia; h. spermatheca (enlarged)." figureDoi="http://doi.org/10.5281/zenodo.8261737" httpUri="https://zenodo.org/record/8261737/files/figure.png" pageId="101" pageNumber="102">Fig. 72f</figureCitation>
) and spermatheca (
<figureCitation id="1355ECF1143D4652BF70304CFD5AF9D1" box="[568,663,1584,1610]" captionStart="FIGURE 72" captionStartId="108.[151,250,1943,1967]" captionTargetBox="[265,1316,182,1918]" captionTargetId="figure-20@108.[245,1342,181,1920]" captionTargetPageId="108" captionText="FIGURE 72. Coelophora circumvelata (Mulsant): a. Type, dorsal view (BMNH); b, d. adult, dorsal view; c. adult, lateral view; e. adult, frontal view; f, g. female genitalia; h. spermatheca (enlarged)." figureDoi="http://doi.org/10.5281/zenodo.8261737" httpUri="https://zenodo.org/record/8261737/files/figure.png" pageId="101" pageNumber="102">Fig. 72h</figureCitation>
) as illustrated. Rarely dorsum fully melanic (
<figureCitation id="1355ECF1143D4652B9C5304CFAC8F9D1" box="[1165,1285,1584,1610]" captionStart="FIGURE 72" captionStartId="108.[151,250,1943,1967]" captionTargetBox="[265,1316,182,1918]" captionTargetId="figure-20@108.[245,1342,181,1920]" captionTargetPageId="108" captionText="FIGURE 72. Coelophora circumvelata (Mulsant): a. Type, dorsal view (BMNH); b, d. adult, dorsal view; c. adult, lateral view; e. adult, frontal view; f, g. female genitalia; h. spermatheca (enlarged)." figureDoi="http://doi.org/10.5281/zenodo.8261737" httpUri="https://zenodo.org/record/8261737/files/figure.png" pageId="101" pageNumber="102">Fig. 72d, e</figureCitation>
) except inner ocular margins of head and anterolateral areas of pronotum yellowish, with similar genitalia (
<figureCitation id="1355ECF1143D4652B9E43028FAC2F9F5" box="[1196,1295,1620,1646]" captionStart="FIGURE 72" captionStartId="108.[151,250,1943,1967]" captionTargetBox="[265,1316,182,1918]" captionTargetId="figure-20@108.[245,1342,181,1920]" captionTargetPageId="108" captionText="FIGURE 72. Coelophora circumvelata (Mulsant): a. Type, dorsal view (BMNH); b, d. adult, dorsal view; c. adult, lateral view; e. adult, frontal view; f, g. female genitalia; h. spermatheca (enlarged)." figureDoi="http://doi.org/10.5281/zenodo.8261737" httpUri="https://zenodo.org/record/8261737/files/figure.png" pageId="101" pageNumber="102">
Fig.
<quantity id="4C965D91143D4652B9A93028FAC2F9F5" box="[1249,1295,1620,1646]" metricMagnitude="-2" metricUnit="kg" metricValue="7.2" pageId="101" pageNumber="102" unit="g" value="72.0">72g</quantity>
</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="C374A3FF143D4652BD8F300BFB5AF909" box="[199,1175,1655,1682]" pageId="101" pageNumber="102" type="distribution">
<paragraph id="8BD1F074143D4652BD8F300BFB5AF909" blockId="101.[151,1437,1439,2042]" box="[199,1175,1655,1682]" pageId="101" pageNumber="102">
<emphasis id="B91A2C66143D4652BD8F300BFEACF90A" bold="true" box="[199,353,1655,1681]" pageId="101" pageNumber="102">Distribution.</emphasis>
<collectingCountry id="F379B0E4143D4652BC2F3004FE68F909" box="[359,421,1656,1682]" name="India" pageId="101" pageNumber="102">India</collectingCountry>
: Northeastern region (
<collectingRegion id="49AA3E96143D4652BFD63004FD22F909" box="[670,751,1656,1682]" country="India" name="Assam" pageId="101" pageNumber="102">Assam</collectingRegion>
,
<collectingRegion id="49AA3E96143D4652BFB13004FCB6F909" box="[761,891,1656,1682]" country="India" name="Meghalaya" pageId="101" pageNumber="102">Meghalaya</collectingRegion>
);
<collectingCountry id="F379B0E4143D4652BEC63004FC19F909" box="[910,980,1656,1682]" name="Nepal" pageId="101" pageNumber="102">Nepal</collectingCountry>
; The
<collectingCountry id="F379B0E4143D4652B95A3004FB59F909" box="[1042,1172,1656,1682]" name="Philippines" pageId="101" pageNumber="102">Philippines</collectingCountry>
.
</paragraph>
</subSubSection>
<subSubSection id="C374A3FF143D4652BD8F30E0FB91F861" pageId="101" pageNumber="102" type="discussion">
<paragraph id="8BD1F074143D4652BD8F30E0FB91F861" blockId="101.[151,1437,1439,2042]" pageId="101" pageNumber="102">
<emphasis id="B91A2C66143D4652BD8F30E0FEDCF92D" bold="true" box="[199,273,1692,1718]" pageId="101" pageNumber="102">Notes.</emphasis>
<taxonomicName id="4C6E8BF7143D4652BC5230E0FDF7F92D" authorityName=": Booth &amp; Pope" authorityYear="1989" box="[282,570,1692,1718]" class="Insecta" family="Coccinellidae" genus="Coelophora" kingdom="Animalia" order="Coleoptera" pageId="101" pageNumber="102" phylum="Arthropoda" rank="species" species="circumvelata">
<emphasis id="B91A2C66143D4652BC5230E0FDF7F92D" box="[282,570,1692,1718]" italics="true" pageId="101" pageNumber="102">Coelophora circumvelata</emphasis>
</taxonomicName>
is of uncertain generic status and
<bibRefCitation id="EFFF8D85143D4652BE8E30E0FB79F92D" author="Poorani, J. &amp; Sankararaman, H. &amp; Anusree, S. S." box="[966,1204,1692,1718]" pageId="101" pageNumber="102" pagination="117 - 133" refId="ref119636" refString="Poorani, J., Sankararaman, H. &amp; Anusree, S. S. (2021) A review of Indian species of Phrynocaria Timberlake (Coleoptera: Coccinellidae) with notes on synonymy and a new species from South India, hitherto misidentified as Coelophora circumusta (Mulsant). Zootaxa, 4926 (1), 117 - 133. https: // doi. org / 10.11646 / zootaxa. 4926.1.8" type="journal article" year="2021">
Poorani
<emphasis id="B91A2C66143D4652B96E30E0FBACF92D" box="[1062,1121,1692,1718]" italics="true" pageId="101" pageNumber="102">et al.</emphasis>
(2021)
</bibRefCitation>
opined that it could be a synonym of
<taxonomicName id="4C6E8BF7143D4652BC2930BCFE10F941" baseAuthorityName="Fabricius" baseAuthorityYear="1792" box="[353,477,1728,1754]" class="Insecta" family="Coccinellidae" genus="Phrynocaria" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="101" pageNumber="102" phylum="Arthropoda" rank="species" species="unicolor">
<emphasis id="B91A2C66143D4652BC2930BCFE10F941" box="[353,477,1728,1754]" italics="true" pageId="101" pageNumber="102">P. unicolor</emphasis>
</taxonomicName>
because it is similar to
<taxonomicName id="4C6E8BF7143D4652BFBB30BCFB8CF941" authority="Weise, 1892" authorityName="Weise" authorityYear="1892" box="[755,1089,1728,1754]" pageId="101" pageNumber="102" rank="variety" variety="cinctipennis">
var.
<emphasis id="B91A2C66143D4652BE6D30BCFC61F942" box="[805,940,1728,1753]" italics="true" pageId="101" pageNumber="102">cinctipennis</emphasis>
<bibRefCitation id="EFFF8D85143D4652BEFD30BCFB8CF941" author="Weise, J." box="[949,1089,1728,1754]" pageId="101" pageNumber="102" pagination="16 - 30" refId="ref124175" refString="Weise, J. (1892) Les Coccinellides du Chota - Nagpore. Annales de la Societe Entomologique du Belgique, 36, 16 - 30." type="journal article" year="1892">Weise, 1892</bibRefCitation>
</taxonomicName>
of
<taxonomicName id="4C6E8BF7143D4652B92330BCFB25F941" baseAuthorityName="Fabricius" baseAuthorityYear="1792" box="[1131,1256,1728,1754]" class="Insecta" family="Coccinellidae" genus="Phrynocaria" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="101" pageNumber="102" phylum="Arthropoda" rank="species" species="unicolor">
<emphasis id="B91A2C66143D4652B92330BCFB25F941" box="[1131,1256,1728,1754]" italics="true" pageId="101" pageNumber="102">P. unicolor</emphasis>
</taxonomicName>
. Booth &amp;
<bibRefCitation id="EFFF8D85143D4652B82C30BCFF2FF965" author="Pope, R. D." pageId="101" pageNumber="102" pagination="633 - 735" refId="ref118886" refString="Pope, R. D. (1989) A revision of the Australian Coccinellidae (Coleoptera). Part I. Subfamily Coccinellinae. Invertebrate Taxonomy, 3, 633 - 735. https: // doi. org / 10.1071 / IT 9880633" type="journal article" year="1989">Pope (1989)</bibRefCitation>
designated a
<typeStatus id="54D54ED6143D4652BC333098FE2FF965" box="[379,482,1764,1790]" pageId="101" pageNumber="102" type="lectotype">lectotype</typeStatus>
for
<taxonomicName id="4C6E8BF7143D4652BF583098FD0FF965" authorityName=": Booth &amp; Pope" authorityYear="1989" box="[528,706,1764,1790]" class="Insecta" family="Coccinellidae" genus="Coelophora" kingdom="Animalia" order="Coleoptera" pageId="101" pageNumber="102" phylum="Arthropoda" rank="species" species="circumvelata">
<emphasis id="B91A2C66143D4652BF583098FD0FF965" box="[528,706,1764,1790]" italics="true" pageId="101" pageNumber="102">C. circumvelata</emphasis>
</taxonomicName>
and Hopes type specimen of
<taxonomicName id="4C6E8BF7143D4652B95D3098FB0AF965" authorityName=": Booth &amp; Pope" authorityYear="1989" box="[1045,1223,1764,1790]" class="Insecta" family="Coccinellidae" genus="Coelophora" kingdom="Animalia" order="Coleoptera" pageId="101" pageNumber="102" phylum="Arthropoda" rank="species" species="circumvelata">
<emphasis id="B91A2C66143D4652B95D3098FB0AF965" box="[1045,1223,1764,1790]" italics="true" pageId="101" pageNumber="102">C. circumvelata</emphasis>
</taxonomicName>
(
<figureCitation id="1355ECF1143D4652B99E3098FAFBF965" box="[1238,1334,1764,1790]" captionStart="FIGURE 72" captionStartId="108.[151,250,1943,1967]" captionTargetBox="[265,1316,182,1918]" captionTargetId="figure-20@108.[245,1342,181,1920]" captionTargetPageId="108" captionText="FIGURE 72. Coelophora circumvelata (Mulsant): a. Type, dorsal view (BMNH); b, d. adult, dorsal view; c. adult, lateral view; e. adult, frontal view; f, g. female genitalia; h. spermatheca (enlarged)." figureDoi="http://doi.org/10.5281/zenodo.8261737" httpUri="https://zenodo.org/record/8261737/files/figure.png" pageId="101" pageNumber="102">Fig. 72a</figureCitation>
, BMNH, examined) looks like a
<taxonomicName id="4C6E8BF7143D4652BCE93174FDE2F8B9" box="[417,559,1800,1826]" class="Insecta" family="Coccinellidae" genus="Phrynocaria" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="101" pageNumber="102" phylum="Arthropoda" rank="genus">
<emphasis id="B91A2C66143D4652BCE93174FDE2F8B9" box="[417,559,1800,1826]" italics="true" pageId="101" pageNumber="102">Phrynocaria</emphasis>
</taxonomicName>
but could not be dissected. Similar variants of
<taxonomicName id="4C6E8BF7143D4652B9013174FB08F8B9" baseAuthorityName="Fabricius" baseAuthorityYear="1792" box="[1097,1221,1800,1826]" class="Insecta" family="Coccinellidae" genus="Phrynocaria" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="101" pageNumber="102" phylum="Arthropoda" rank="species" species="unicolor">
<emphasis id="B91A2C66143D4652B9013174FB08F8B9" box="[1097,1221,1800,1826]" italics="true" pageId="101" pageNumber="102">P. unicolor</emphasis>
</taxonomicName>
from eastern
<collectingCountry id="F379B0E4143D4652B82B3174FA50F8B9" box="[1379,1437,1800,1826]" name="India" pageId="101" pageNumber="102">India</collectingCountry>
have been examined. There is also a rare variant of
<taxonomicName id="4C6E8BF7143D4652BFB43150FC5BF8DE" baseAuthorityName="Mulsant" baseAuthorityYear="1850" box="[764,918,1836,1861]" class="Insecta" family="Coccinellidae" genus="Phrynocaria" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="101" pageNumber="102" phylum="Arthropoda" rank="species" species="circumusta">
<emphasis id="B91A2C66143D4652BFB43150FC5BF8DE" box="[764,918,1836,1861]" italics="true" pageId="101" pageNumber="102">P. circumusta</emphasis>
</taxonomicName>
with black lateral borders of elytra as in
<taxonomicName id="4C6E8BF7143D4652B8CB3150FEE5F8F1" authorityName=": Booth &amp; Pope" authorityYear="1989" class="Insecta" family="Coccinellidae" genus="Coelophora" kingdom="Animalia" order="Coleoptera" pageId="101" pageNumber="102" phylum="Arthropoda" rank="species" species="circumvelata">
<emphasis id="B91A2C66143D4652B8CB3150FEE5F8F1" italics="true" pageId="101" pageNumber="102">C. circumvelata</emphasis>
</taxonomicName>
. It is included here in
<taxonomicName id="4C6E8BF7143D4652BF55312CFD6EF8F1" box="[541,675,1872,1898]" class="Insecta" family="Coccinellidae" genus="Coelophora" kingdom="Animalia" order="Coleoptera" pageId="101" pageNumber="102" phylum="Arthropoda" rank="genus">
<emphasis id="B91A2C66143D4652BF55312CFD6EF8F1" box="[541,675,1872,1898]" italics="true" pageId="101" pageNumber="102">Coelophora</emphasis>
</taxonomicName>
following Booth &amp;
<bibRefCitation id="EFFF8D85143D4652BECA312CFBC6F8F1" author="Pope, R. D." box="[898,1035,1872,1898]" pageId="101" pageNumber="102" pagination="633 - 735" refId="ref118886" refString="Pope, R. D. (1989) A revision of the Australian Coccinellidae (Coleoptera). Part I. Subfamily Coccinellinae. Invertebrate Taxonomy, 3, 633 - 735. https: // doi. org / 10.1071 / IT 9880633" type="journal article" year="1989">Pope (1989)</bibRefCitation>
and the male genitalia illustrated by
<bibRefCitation id="EFFF8D85143D4652BDDF3108FE9BF815" author="Ren, S. X. &amp; Wang, X. M. &amp; Pang, H. &amp; Peng, Z. Q. &amp; Zeng, T." box="[151,342,1908,1934]" pageId="101" pageNumber="102" refId="ref120760" refString="Ren, S. X., Wang, X. M., Pang, H., Peng, Z. Q. &amp; Zeng, T. (2009) Colored pictorial handbook of ladybird beetles in China. Science Press, Beijing, 336 pp. [in Chinese]" type="book" year="2009">
Ren
<emphasis id="B91A2C66143D4652BD843108FEC9F815" box="[204,260,1908,1934]" italics="true" pageId="101" pageNumber="102">et al.</emphasis>
(2009)
</bibRefCitation>
also appear to be characteristic of
<taxonomicName id="4C6E8BF7143D4652BF963108FCA9F815" box="[734,868,1908,1934]" class="Insecta" family="Coccinellidae" genus="Coelophora" kingdom="Animalia" order="Coleoptera" pageId="101" pageNumber="102" phylum="Arthropoda" rank="genus">
<emphasis id="B91A2C66143D4652BF963108FCA9F815" box="[734,868,1908,1934]" italics="true" pageId="101" pageNumber="102">Coelophora</emphasis>
</taxonomicName>
, with the penis composed of two distinct sclerites. A female closely matching
<taxonomicName id="4C6E8BF7143D4652BC8531E4FD4DF829" authorityName=": Booth &amp; Pope" authorityYear="1989" box="[461,640,1944,1970]" class="Insecta" family="Coccinellidae" genus="Coelophora" kingdom="Animalia" order="Coleoptera" pageId="101" pageNumber="102" phylum="Arthropoda" rank="species" species="circumvelata">
<emphasis id="B91A2C66143D4652BC8531E4FD4DF829" box="[461,640,1944,1970]" italics="true" pageId="101" pageNumber="102">C. circumvelata</emphasis>
</taxonomicName>
(
<figureCitation id="1355ECF1143D4652BFD831E4FCC7F829" box="[656,778,1944,1970]" captionStart="FIGURE 72" captionStartId="108.[151,250,1943,1967]" captionTargetBox="[265,1316,182,1918]" captionTargetId="figure-20@108.[245,1342,181,1920]" captionTargetPageId="108" captionText="FIGURE 72. Coelophora circumvelata (Mulsant): a. Type, dorsal view (BMNH); b, d. adult, dorsal view; c. adult, lateral view; e. adult, frontal view; f, g. female genitalia; h. spermatheca (enlarged)." figureDoi="http://doi.org/10.5281/zenodo.8261737" httpUri="https://zenodo.org/record/8261737/files/figure.png" pageId="101" pageNumber="102">Fig. 72ac</figureCitation>
) and an almost fully melanic variant conspecific with this (
<figureCitation id="1355ECF1143D4652BDD731C0FED5F84D" box="[159,280,1980,2006]" captionStart="FIGURE 72" captionStartId="108.[151,250,1943,1967]" captionTargetBox="[265,1316,182,1918]" captionTargetId="figure-20@108.[245,1342,181,1920]" captionTargetPageId="108" captionText="FIGURE 72. Coelophora circumvelata (Mulsant): a. Type, dorsal view (BMNH); b, d. adult, dorsal view; c. adult, lateral view; e. adult, frontal view; f, g. female genitalia; h. spermatheca (enlarged)." figureDoi="http://doi.org/10.5281/zenodo.8261737" httpUri="https://zenodo.org/record/8261737/files/figure.png" pageId="101" pageNumber="102">Fig. 72d, e</figureCitation>
) were examined from
<collectingRegion id="49AA3E96143D4652BF5A31C0FDACF84D" box="[530,609,1980,2006]" country="India" name="Assam" pageId="101" pageNumber="102">Assam</collectingRegion>
, north-eastern
<collectingCountry id="F379B0E4143D4652BE4E31C0FC8DF84D" box="[774,832,1980,2006]" name="India" pageId="101" pageNumber="102">India</collectingCountry>
and both are illustrated here. Their genitalia also were almost identical (
<figureCitation id="1355ECF1143D4652BC11319CFE1CF861" box="[345,465,2016,2042]" captionStart="FIGURE 72" captionStartId="108.[151,250,1943,1967]" captionTargetBox="[265,1316,182,1918]" captionTargetId="figure-20@108.[245,1342,181,1920]" captionTargetPageId="108" captionText="FIGURE 72. Coelophora circumvelata (Mulsant): a. Type, dorsal view (BMNH); b, d. adult, dorsal view; c. adult, lateral view; e. adult, frontal view; f, g. female genitalia; h. spermatheca (enlarged)." figureDoi="http://doi.org/10.5281/zenodo.8261737" httpUri="https://zenodo.org/record/8261737/files/figure.png" pageId="101" pageNumber="102">Fig. 72f, g</figureCitation>
). See Booth &amp;
<bibRefCitation id="EFFF8D85143D4652BF35319CFCCAF861" author="Pope, R. D." box="[637,775,2016,2042]" pageId="101" pageNumber="102" pagination="633 - 735" refId="ref118886" refString="Pope, R. D. (1989) A revision of the Australian Coccinellidae (Coleoptera). Part I. Subfamily Coccinellinae. Invertebrate Taxonomy, 3, 633 - 735. https: // doi. org / 10.1071 / IT 9880633" type="journal article" year="1989">Pope (1989)</bibRefCitation>
for notes on its nomenclature.
</paragraph>
</subSubSection>
</treatment>
</document>