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<mods:title id="C0F4E078821ACC054363A2CCF77E956C">An illustrated guide to lady beetles (Coleoptera: Coccinellidae) of the Indian Subcontinent. Part 1. Tribe Coccinellini</mods:title>
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<taxonomicName id="4C6E8BF71431465EBDDF3741FDF1FECC" authority="(Fabricius)" baseAuthorityName="Fabricius" baseAuthorityYear="1775" box="[151,572,317,344]" class="Insecta" family="Coccinellidae" genus="Coelophora" kingdom="Animalia" order="Coleoptera" pageId="105" pageNumber="106" phylum="Arthropoda" rank="species" species="inaequalis">
<emphasis id="B91A2C661431465EBDDF3741FDF1FECC" bold="true" box="[151,572,317,344]" pageId="105" pageNumber="106">
<emphasis id="B91A2C661431465EBDDF3741FE67FEC3" bold="true" box="[151,426,317,344]" italics="true" pageId="105" pageNumber="106">Coelophora inaequalis</emphasis>
(Fabricius)
</emphasis>
</taxonomicName>
</heading>
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(
<figureCitation id="1355ECF11431465EBDD7371DFF3CFEE0" box="[159,241,353,379]" captionStart="FIGURE 70" captionStartId="105.[151,250,1774,1798]" captionTargetBox="[153,1433,556,1747]" captionTargetId="figure-105@105.[151,1436,555,1750]" captionTargetPageId="105" captionText="FIGURE 70. Coelophora inaequalis (Fabricius): a. adult (India); b. adult (Malaysia); c. abdomen; d, e. spermatheca; f, g. male genitalia: f. tegmen, ventral view; g. penis." figureDoi="http://doi.org/10.5281/zenodo.8261733" httpUri="https://zenodo.org/record/8261733/files/figure.png" pageId="105" pageNumber="106">Fig. 70</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="C374A3FF1431465EBDDF37E3FC2AFD88" pageId="105" pageNumber="106" type="reference_group">
<paragraph id="8BD1F0741431465EBDDF37E3FB75FE2C" blockId="105.[151,1436,415,531]" box="[151,1208,415,440]" pageId="105" pageNumber="106">
<taxonomicName id="4C6E8BF71431465EBDDF37E3FDF2FE2C" authority="Fabricius, 1775: 80" authorityName="Fabricius" authorityPageNumber="80" authorityYear="1775" box="[151,575,415,439]" class="Insecta" family="Coccinellidae" genus="Coccinella" kingdom="Animalia" order="Coleoptera" pageId="105" pageNumber="106" phylum="Arthropoda" rank="species" species="inaequalis">
<emphasis id="B91A2C661431465EBDDF37E3FEB8FE2C" box="[151,373,415,439]" italics="true" pageId="105" pageNumber="106">Coccinella inaequalis</emphasis>
<materialsCitation id="3B06FA291431465EBC3337E3FDF2FE2C" ID-GBIF-Occurrence="4399052307" box="[379,575,415,439]" collectionCode="BMNH" pageId="105" pageNumber="106" specimenCount="1" typeStatus="lectotype">Fabricius, 1775: 80</materialsCitation>
</taxonomicName>
(
<typeStatus id="54D54ED61431465EBF0537E3FD79FE2C" box="[589,692,415,439]" pageId="105" pageNumber="106" type="lectotype">Lectotype</typeStatus>
female, BMNH; Type locality: Nova Hollandia).
</paragraph>
<paragraph id="8BD1F0741431465EBDDF37C2FE2AFE6F" blockId="105.[151,1436,415,531]" pageId="105" pageNumber="106">
<treatmentCitationGroup id="AB7ED75A1431465EBDDF37C2FE2AFE6F" pageId="105" pageNumber="106">
<treatmentCitation id="0ACFD6651431465EBDDF37C2FD9AFE4D" author="Mulsant, E." box="[151,599,446,470]" page="404" pageId="105" pageNumber="106" year="1850">
<taxonomicName id="4C6E8BF71431465EBDDF37C2FD9AFE4D" authority=": Mulsant, 1850: 404" authorityName="Mulsant" authorityPageNumber="404" authorityYear="1850" box="[151,599,446,470]" class="Insecta" family="Coccinellidae" genus="Coelophora" kingdom="Animalia" order="Coleoptera" pageId="105" pageNumber="106" phylum="Arthropoda" rank="species" species="inaequalis">
<emphasis id="B91A2C661431465EBDDF37C2FE4CFE4D" box="[151,385,446,470]" italics="true" pageId="105" pageNumber="106">Coelophora inaequalis</emphasis>
:
<bibRefCitation id="EFFF8D851431465EBCC637C2FD9AFE4D" author="Mulsant, E." box="[398,599,446,470]" pageId="105" pageNumber="106" pagination="1 - 1104" refId="ref117842" refString="Mulsant, E. (1850) Species des Coleopteres Trimeres Securipalpes. Annales des Sciences Physiques et Naturelles, d'Agriculture et d'Industrie, publiees par la Societe nationale d'Agriculture, etc., de Lyon, Deuxieme Serie, 2, 1 - 1104. https: // doi. org / 10.5962 / bhl. title. 8953" type="journal article" year="1850">Mulsant, 1850: 404</bibRefCitation>
</taxonomicName>
</treatmentCitation>
; Crotch 1874: 153;
<treatmentCitation id="0ACFD6651431465EBE6937C2FBDCFE4D" author="Korschefsky, R." box="[801,1041,446,470]" page="292" pageId="105" pageNumber="106" year="1932">
<bibRefCitation id="EFFF8D851431465EBE6937C2FBDCFE4D" author="Korschefsky, R." box="[801,1041,446,470]" pageId="105" pageNumber="106" refId="ref116133" refString="Korschefsky, R. (1932) Coleopterorum Catalogus. Pars 120. Coccinellidae II. W. Junk, Berlin, 435 pp." type="book" year="1932">Korschefsky 1932: 292</bibRefCitation>
</treatmentCitation>
;
<treatmentCitation id="0ACFD6651431465EB95337C2FB09FE4D" author="Pope, R. D." box="[1051,1220,446,470]" page="665" pageId="105" pageNumber="106" year="1989">
<bibRefCitation id="EFFF8D851431465EB95337C2FB09FE4D" author="Pope, R. D." box="[1051,1220,446,470]" pageId="105" pageNumber="106" pagination="633 - 735" refId="ref118886" refString="Pope, R. D. (1989) A revision of the Australian Coccinellidae (Coleoptera). Part I. Subfamily Coccinellinae. Invertebrate Taxonomy, 3, 633 - 735. https: // doi. org / 10.1071 / IT 9880633" type="journal article" year="1989">Pope, 1989: 665</bibRefCitation>
</treatmentCitation>
;
<treatmentCitation id="0ACFD6651431465EB98637C2FA55FE4D" author="Poorani, J." box="[1230,1432,446,470]" page="328" pageId="105" pageNumber="106" year="2002">
<bibRefCitation id="EFFF8D851431465EB98637C2FA55FE4D" author="Poorani, J." box="[1230,1432,446,470]" pageId="105" pageNumber="106" pagination="307 - 383" refId="ref118931" refString="Poorani, J. (2002 a) An annotated checklist of the Coccinellidae (excluding Epilachninae) of the Indian subregion. Oriental Insects, 36, 307 - 383. https: // doi. org / 10.1080 / 00305316.2002.10417335" type="journal article" year="2002">Poorani 2002a: 328</bibRefCitation>
</treatmentCitation>
;
<treatmentCitation id="0ACFD6651431465EBD8F37A2FE2EFE6F" author="Slipinski, A. &amp; Li, J. &amp; Pang, H." box="[199,483,476,503]" page="36" pageId="105" pageNumber="106" year="2020">
<bibRefCitation id="EFFF8D851431465EBD8F37A2FE2EFE6F" author="Slipinski, A. &amp; Li, J. &amp; Pang, H." box="[199,483,476,503]" pageId="105" pageNumber="106" refId="ref122780" refString="Slipinski, A., Li, J. &amp; Pang, H. (2020) Ladybird Beetles of the Australo-Pacific Region Coleoptera: Coccinellidae: Coccinellini. CSIRO Publishing, Clayton, Victoria, 240 pp. https: // doi. org / 10.1071 / 9781486303885" type="book" year="2020">
Ślipiński
<emphasis id="B91A2C661431465EBC6037A1FE99FE6F" box="[296,340,476,500]" italics="true" pageId="105" pageNumber="106">et al</emphasis>
. 2020: 3637
</bibRefCitation>
</treatmentCitation>
.
</treatmentCitationGroup>
</paragraph>
<paragraph id="8BD1F0741431465EBDDF3787FC2AFD88" blockId="105.[151,1436,415,531]" box="[151,999,507,531]" pageId="105" pageNumber="106">
<treatmentCitationGroup id="AB7ED75A1431465EBDDF3787FC2AFD88" box="[151,999,507,531]" pageId="105" pageNumber="106">
<treatmentCitation id="0ACFD6651431465EBDDF3787FDD7FD88" author="Mulsant, E." box="[151,538,507,531]" page="393" pageId="105" pageNumber="106" year="1850">
<taxonomicName id="4C6E8BF71431465EBDDF3787FDD7FD88" authority="Mulsant, 1850: 393" authorityName="Mulsant" authorityPageNumber="393" authorityYear="1850" box="[151,538,507,531]" class="Insecta" family="Coccinellidae" genus="Coelophora" kingdom="Animalia" order="Coleoptera" pageId="105" pageNumber="106" phylum="Arthropoda" rank="species" species="vidua">
<emphasis id="B91A2C661431465EBDDF3787FE82FD88" box="[151,335,507,531]" italics="true" pageId="105" pageNumber="106">Coelophora vidua</emphasis>
<bibRefCitation id="EFFF8D851431465EBC1D3787FDD7FD88" author="Mulsant, E." box="[341,538,507,531]" pageId="105" pageNumber="106" pagination="1 - 1104" refId="ref117842" refString="Mulsant, E. (1850) Species des Coleopteres Trimeres Securipalpes. Annales des Sciences Physiques et Naturelles, d'Agriculture et d'Industrie, publiees par la Societe nationale d'Agriculture, etc., de Lyon, Deuxieme Serie, 2, 1 - 1104. https: // doi. org / 10.5962 / bhl. title. 8953" type="journal article" year="1850">Mulsant, 1850: 393</bibRefCitation>
</taxonomicName>
</treatmentCitation>
.—Synonymized by
<treatmentCitation id="0ACFD6651431465EBFA43787FC2EFD88" author="Poorani, J. &amp; Slipinski, A. &amp; Booth, R. G." box="[748,995,507,531]" page="592" pageId="105" pageNumber="106" year="2008">
<bibRefCitation id="EFFF8D851431465EBFA43787FC2EFD88" author="Poorani, J. &amp; Slipinski, A. &amp; Booth, R. G." box="[748,995,507,531]" pageId="105" pageNumber="106" pagination="579 - 594" refId="ref119452" refString="Poorani, J., Slipinski, A. &amp; Booth, R. G. (2008) A revision of Synona Pope (Coleoptera: Coccinellidae: Coccinellini). Annales Zoologici, 58, 579 - 594. https: // doi. org / 10.3161 / 000345408 X 364427" type="journal article" year="2008">
Poorani
<emphasis id="B91A2C661431465EBE093780FCA3FD88" box="[833,878,507,531]" italics="true" pageId="105" pageNumber="106">et al</emphasis>
. 2008: 592
</bibRefCitation>
</treatmentCitation>
.
</treatmentCitationGroup>
</paragraph>
</subSubSection>
<caption id="DF11A0FC1431465EBDDF3092FD87F8B1" ID-DOI="http://doi.org/10.5281/zenodo.8261733" ID-Zenodo-Dep="8261733" httpUri="https://zenodo.org/record/8261733/files/figure.png" pageId="105" pageNumber="106" startId="105.[151,250,1774,1798]" targetBox="[153,1433,556,1747]" targetPageId="105" targetType="figure">
<paragraph id="8BD1F0741431465EBDDF3092FD87F8B1" blockId="105.[151,1436,1774,1834]" pageId="105" pageNumber="106">
<emphasis id="B91A2C661431465EBDDF3092FEEDF89D" bold="true" box="[151,288,1774,1798]" pageId="105" pageNumber="106">FIGURE 70.</emphasis>
<taxonomicName id="4C6E8BF71431465EBC6D3092FD4AF89D" authority="(Fabricius)" baseAuthorityName="Fabricius" baseAuthorityYear="1775" box="[293,647,1774,1798]" class="Insecta" family="Coccinellidae" genus="Coelophora" kingdom="Animalia" order="Coleoptera" pageId="105" pageNumber="106" phylum="Arthropoda" rank="species" species="inaequalis">
<emphasis id="B91A2C661431465EBC6D3092FDC2F89D" box="[293,527,1774,1798]" italics="true" pageId="105" pageNumber="106">Coelophora inaequalis</emphasis>
(Fabricius)
</taxonomicName>
: a. adult (India); b. adult (Malaysia); c. abdomen; d, e. spermatheca; f, g. male genitalia: f. tegmen, ventral view; g. penis.
</paragraph>
</caption>
<subSubSection id="C374A3FF1431465EBDDF3113FC91F86E" pageId="105" pageNumber="106" type="diagnosis">
<paragraph id="8BD1F0741431465EBDDF3113FC91F86E" blockId="105.[151,1437,1903,2037]" pageId="105" pageNumber="106">
<emphasis id="B91A2C661431465EBDDF3113FEDFF812" bold="true" box="[151,274,1903,1929]" pageId="105" pageNumber="106">Diagnosis.</emphasis>
Length: 4.306.00 mm; width:
<quantity id="4C965D911431465EBF383113FCD8F812" box="[624,789,1903,1930]" metricMagnitude="-3" metricUnit="m" metricValue="4.5" metricValueMax="5.5" metricValueMin="3.5" pageId="105" pageNumber="106" unit="mm" value="4.5" valueMax="5.5" valueMin="3.5">3.505.50 mm</quantity>
. Form broad, almost circular, convex and glabrous. Dorsal colour pattern highly variable, the form illustrated here (
<figureCitation id="1355ECF11431465EBE5C31EFFCBEF836" box="[788,883,1939,1965]" captionStart="FIGURE 70" captionStartId="105.[151,250,1774,1798]" captionTargetBox="[153,1433,556,1747]" captionTargetId="figure-105@105.[151,1436,555,1750]" captionTargetPageId="105" captionText="FIGURE 70. Coelophora inaequalis (Fabricius): a. adult (India); b. adult (Malaysia); c. abdomen; d, e. spermatheca; f, g. male genitalia: f. tegmen, ventral view; g. penis." figureDoi="http://doi.org/10.5281/zenodo.8261733" httpUri="https://zenodo.org/record/8261733/files/figure.png" pageId="105" pageNumber="106">Fig. 70a</figureCitation>
) superficially resembles
<taxonomicName id="4C6E8BF71431465EB9C631EFFAD6F836" box="[1166,1307,1939,1965]" class="Insecta" family="Coccinellidae" genus="Coelophora" kingdom="Animalia" order="Coleoptera" pageId="105" pageNumber="106" phylum="Arthropoda" rank="species" species="bissellata">
<emphasis id="B91A2C661431465EB9C631EFFAD6F836" box="[1166,1307,1939,1965]" italics="true" pageId="105" pageNumber="106">C. bissellata</emphasis>
</taxonomicName>
with a pair of black spots on pronotum and nine black spots on elytra. Abdominal postcoxal line incomplete (
<figureCitation id="1355ECF11431465EB9A531CBFA82F84A" box="[1261,1359,1975,2001]" captionStart="FIGURE 70" captionStartId="105.[151,250,1774,1798]" captionTargetBox="[153,1433,556,1747]" captionTargetId="figure-105@105.[151,1436,555,1750]" captionTargetPageId="105" captionText="FIGURE 70. Coelophora inaequalis (Fabricius): a. adult (India); b. adult (Malaysia); c. abdomen; d, e. spermatheca; f, g. male genitalia: f. tegmen, ventral view; g. penis." figureDoi="http://doi.org/10.5281/zenodo.8261733" httpUri="https://zenodo.org/record/8261733/files/figure.png" pageId="105" pageNumber="106">Fig. 70c</figureCitation>
). Male genitalia (
<figureCitation id="1355ECF11431465EBC4F31A7FEB3F86E" box="[263,382,2011,2037]" captionStart="FIGURE 70" captionStartId="105.[151,250,1774,1798]" captionTargetBox="[153,1433,556,1747]" captionTargetId="figure-105@105.[151,1436,555,1750]" captionTargetPageId="105" captionText="FIGURE 70. Coelophora inaequalis (Fabricius): a. adult (India); b. adult (Malaysia); c. abdomen; d, e. spermatheca; f, g. male genitalia: f. tegmen, ventral view; g. penis." figureDoi="http://doi.org/10.5281/zenodo.8261733" httpUri="https://zenodo.org/record/8261733/files/figure.png" pageId="105" pageNumber="106">Fig. 70f, g</figureCitation>
) and spermatheca (
<figureCitation id="1355ECF11431465EBF1031A7FD1CF86E" box="[600,721,2011,2037]" captionStart="FIGURE 70" captionStartId="105.[151,250,1774,1798]" captionTargetBox="[153,1433,556,1747]" captionTargetId="figure-105@105.[151,1436,555,1750]" captionTargetPageId="105" captionText="FIGURE 70. Coelophora inaequalis (Fabricius): a. adult (India); b. adult (Malaysia); c. abdomen; d, e. spermatheca; f, g. male genitalia: f. tegmen, ventral view; g. penis." figureDoi="http://doi.org/10.5281/zenodo.8261733" httpUri="https://zenodo.org/record/8261733/files/figure.png" pageId="105" pageNumber="106">Fig. 70d, e</figureCitation>
) diagnostic.
</paragraph>
</subSubSection>
<subSubSection id="C374A3FF1432465DBD8F36EAFED2FF62" pageId="106" pageNumber="107" type="distribution">
<paragraph id="8BD1F0741432465DBD8F36EAFED2FF62" blockId="106.[151,1437,150,321]" pageId="106" pageNumber="107">
<emphasis id="B91A2C661432465DBD8F36EAFEACFF2B" bold="true" box="[199,353,150,176]" pageId="106" pageNumber="107">Distribution.</emphasis>
<collectingCountry id="F379B0E41432465DBC2F36EBFE6CFF2A" box="[359,417,151,177]" name="India" pageId="106" pageNumber="107">India</collectingCountry>
(Andamans);
<collectingCountry id="F379B0E41432465DBF0836EBFD7FFF2A" box="[576,690,151,177]" name="Sri Lanka" pageId="106" pageNumber="107">Sri Lanka</collectingCountry>
; Southeast Asia (
<collectingCountry id="F379B0E41432465DBE3836EBFC15FF2A" box="[880,984,151,177]" name="Thailand" pageId="106" pageNumber="107">Thailand</collectingCountry>
,
<collectingCountry id="F379B0E41432465DBEAB36EBFB81FF2A" box="[995,1100,151,177]" name="Malaysia" pageId="106" pageNumber="107">Malaysia</collectingCountry>
,
<collectingCountry id="F379B0E41432465DB91F36EBFB0AFF2A" box="[1111,1223,151,177]" name="Indonesia" pageId="106" pageNumber="107">Indonesia</collectingCountry>
,
<collectingCountry id="F379B0E41432465DB99A36EBFA98FF2A" box="[1234,1365,151,177]" name="Philippines" pageId="106" pageNumber="107">Philippines</collectingCountry>
); New
<collectingCountry id="F379B0E41432465DBDDF36C7FF26FF4E" box="[151,235,187,213]" name="Guinea-Bissau" pageId="106" pageNumber="107">Guinea</collectingCountry>
;
<collectingCountry id="F379B0E41432465DBDB136C7FE60FF4E" box="[249,429,187,213]" name="New Caledonia" pageId="106" pageNumber="107">New Caledonia</collectingCountry>
;
<collectingCountry id="F379B0E41432465DBCF336C7FDF0FF4E" box="[443,573,187,213]" name="Micronesia" pageId="106" pageNumber="107">Micronesia</collectingCountry>
;
<collectingCountry id="F379B0E41432465DBF0136C7FD7EFF4E" box="[585,691,187,213]" name="Australia" pageId="106" pageNumber="107">Australia</collectingCountry>
; Introduced in
<collectingCountry id="F379B0E41432465DBE2B36C7FC32FF4E" box="[867,1023,187,213]" name="New Zealand" pageId="106" pageNumber="107">New Zealand</collectingCountry>
, Hawaii and Florida (
<bibRefCitation id="EFFF8D851432465DB9B336C7FA5AFF4E" author="Gordon, R. D." box="[1275,1431,186,213]" pageId="106" pageNumber="107" pagination="1 - 912" refId="ref113006" refString="Gordon, R. D. (1985) The Coccinellidae of America, north of Mexico. Journal of the New York Entomological Society, 93, 1 - 912." type="journal article" year="1985">Gordon 1985</bibRefCitation>
;
<bibRefCitation id="EFFF8D851432465DBDDF36A3FEDFFF62" author="Pope, R. D." box="[151,274,223,249]" pageId="106" pageNumber="107" pagination="633 - 735" refId="ref118886" refString="Pope, R. D. (1989) A revision of the Australian Coccinellidae (Coleoptera). Part I. Subfamily Coccinellinae. Invertebrate Taxonomy, 3, 633 - 735. https: // doi. org / 10.1071 / IT 9880633" type="journal article" year="1989">Pope 1989</bibRefCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="C374A3FF1432465DBD8F377FFB0CFEDA" pageId="106" pageNumber="107" type="discussion">
<paragraph id="8BD1F0741432465DBD8F377FFB0CFEDA" blockId="106.[151,1437,150,321]" pageId="106" pageNumber="107">
<emphasis id="B91A2C661432465DBD8F377FFEDCFE86" bold="true" box="[199,273,259,285]" pageId="106" pageNumber="107">Notes.</emphasis>
<bibRefCitation id="EFFF8D851432465DBC51377FFE69FE86" author="Pope, R. D." box="[281,420,259,285]" pageId="106" pageNumber="107" pagination="633 - 735" refId="ref118886" refString="Pope, R. D. (1989) A revision of the Australian Coccinellidae (Coleoptera). Part I. Subfamily Coccinellinae. Invertebrate Taxonomy, 3, 633 - 735. https: // doi. org / 10.1071 / IT 9880633" type="journal article" year="1989">Pope (1989)</bibRefCitation>
and
<bibRefCitation id="EFFF8D851432465DBC953778FD1BFE86" author="Slipinski, A. &amp; Li, J. &amp; Pang, H." box="[477,726,259,287]" pageId="106" pageNumber="107" refId="ref122780" refString="Slipinski, A., Li, J. &amp; Pang, H. (2020) Ladybird Beetles of the Australo-Pacific Region Coleoptera: Coccinellidae: Coccinellini. CSIRO Publishing, Clayton, Victoria, 240 pp. https: // doi. org / 10.1071 / 9781486303885" type="book" year="2020">
Ślipiński
<emphasis id="B91A2C661432465DBF01377FFD4EFE86" box="[585,643,259,285]" italics="true" pageId="106" pageNumber="107">et al.</emphasis>
(2020)
</bibRefCitation>
described and illustrated this species in detail. It is not known from the Indian mainland and the specimens studied were collected from the Andaman Islands.
</paragraph>
</subSubSection>
</treatment>
</document>