269 lines
37 KiB
XML
269 lines
37 KiB
XML
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<mods:title id="4884B912309799255D1E26A32A91DDE9">An illustrated guide to lady beetles (Coleoptera: Coccinellidae) of the Indian Subcontinent. Part 1. Tribe Coccinellini</mods:title>
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<mods:title id="7D672EC810B934FC032DA20BF4CE3DD7">Zootaxa</mods:title>
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<mods:date id="ECA2198B26ADBF905F2FE52D5825345A">2023</mods:date>
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<treatment id="03C741621423464ABDDF353FFB2DFE12" ID-DOI="http://doi.org/10.5281/zenodo.8261572" ID-GBIF-Taxon="212450066" ID-Zenodo-Dep="8261572" LSID="urn:lsid:plazi:treatment:03C741621423464ABDDF353FFB2DFE12" httpUri="http://treatment.plazi.org/id/03C741621423464ABDDF353FFB2DFE12" lastPageId="125" lastPageNumber="126" pageId="123" pageNumber="124">
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<subSubSection id="C374A3FF1423464CBDDF353FFD4BFCC6" box="[151,646,834,861]" pageId="123" pageNumber="124" type="nomenclature">
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<paragraph id="8BD1F0741423464CBDDF353FFD4BFCC6" blockId="123.[151,646,834,897]" box="[151,646,834,861]" pageId="123" pageNumber="124">
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<heading id="D09947181423464CBDDF353FFD4BFCC6" bold="true" box="[151,646,834,861]" fontSize="11" level="2" pageId="123" pageNumber="124" reason="4">
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<emphasis id="B91A2C661423464CBDDF353FFD4BFCC6" bold="true" box="[151,646,834,861]" pageId="123" pageNumber="124">
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<taxonomicName id="4C6E8BF71423464CBDDF353FFDFAFCC7" authority="Poorani" authorityName="Poorani" authorityYear="2023" box="[151,567,834,861]" class="Insecta" family="Coccinellidae" genus="Harmonia" kingdom="Animalia" order="Coleoptera" pageId="123" pageNumber="124" phylum="Arthropoda" rank="species" species="andamanensis" status="sp. nov.">
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<emphasis id="B91A2C661423464CBDDF353FFE07FCC6" bold="true" box="[151,458,834,861]" italics="true" pageId="123" pageNumber="124">Harmonia andamanensis</emphasis>
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Poorani
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</taxonomicName>
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,
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<taxonomicNameLabel id="A229911D1423464CBF0C353FFD4BFCC6" box="[580,646,835,861]" pageId="123" pageNumber="124" rank="species">sp. n.</taxonomicNameLabel>
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</emphasis>
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</heading>
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</paragraph>
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</subSubSection>
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<subSubSection id="C374A3FF1423464CBDDF351BFEF8FC1A" box="[151,309,870,897]" pageId="123" pageNumber="124" type="description">
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<paragraph id="8BD1F0741423464CBDDF351BFEF8FC1A" blockId="123.[151,646,834,897]" box="[151,309,870,897]" pageId="123" pageNumber="124">
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(
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<figureCitation id="1355ECF11423464CBDD7351BFF3BFC1A" box="[159,246,870,897]" captionStart="FIGURE 85" captionStartId="124.[151,250,1943,1967]" captionTargetBox="[176,1408,191,1903]" captionTargetId="figure-20@124.[166,1422,181,1919]" captionTargetPageId="124" captionText="FIGURE 85. Harmonia andamanensis Poorani, sp. n.: a. adult, dorsal view; b. adult, lateral view; c. adult, frontal view; d. abdomen, female; e. abdominal postcoxal line; f. female genitalia; g. spermatheca." figureDoi="http://doi.org/10.5281/zenodo.8261781" httpUri="https://zenodo.org/record/8261781/files/figure.png" pageId="123" pageNumber="124">Figs 85</figureCitation>
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,
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<figureCitation id="1355ECF11423464CBC4A351BFEE0FC1A" box="[258,301,871,897]" captionStart="FIGURE 199" captionStartId="275.[151,250,1943,1967]" captionTargetBox="[223,1377,194,1916]" captionTargetId="figure-15@275.[203,1385,181,1920]" captionTargetPageId="275" captionText="FIGURE 199. Harmonia andamanensis sp. n.: a. adult female, dorsal view; b. adult male, dorsal view; c. apical abdominal ventrites, male; d. apical abdominal ventrites, female; e. spermatheca; f–i. male genitalia: f. tegmen, lateral view; g. tegmen, ventral view; h. penis; i. penis apex." figureDoi="http://doi.org/10.5281/zenodo.8262209" httpUri="https://zenodo.org/record/8262209/files/figure.png" pageId="123" pageNumber="124">199</figureCitation>
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)
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</paragraph>
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</subSubSection>
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<subSubSection id="C374A3FF1423464CBDDF35D2FEB7FA96" pageId="123" pageNumber="124" type="diagnosis">
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<paragraph id="8BD1F0741423464CBDDF35D2FEB7FA96" blockId="123.[151,1437,942,2013]" pageId="123" pageNumber="124">
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<emphasis id="B91A2C661423464CBDDF35D2FEDFFC53" bold="true" box="[151,274,942,968]" pageId="123" pageNumber="124">Diagnosis.</emphasis>
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<taxonomicName id="4C6E8BF71423464CBC5435D3FDF4FC52" authority="POORANI, 2023" authorityName="Poorani" authorityYear="2023" box="[284,569,943,969]" class="Insecta" family="Coccinellidae" genus="Harmonia" kingdom="Animalia" order="Coleoptera" pageId="123" pageNumber="124" phylum="Arthropoda" rank="species" species="andamanensis" status="sp. nov.">
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<emphasis id="B91A2C661423464CBC5435D3FDF4FC52" box="[284,569,943,969]" italics="true" pageId="123" pageNumber="124">Harmonia andamanensis</emphasis>
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</taxonomicName>
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<taxonomicNameLabel id="A229911D1423464CBF0B35D3FD4FFC52" box="[579,642,943,969]" pageId="123" pageNumber="124" rank="species">sp. n.</taxonomicNameLabel>
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can be separated from its Indian congeners by its dull straw yellow body with very pale elytral maculae and the male genitalia. It has a superficial similarity to old specimens of
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<taxonomicName id="4C6E8BF71423464CBDDF358BFE99FB8A" baseAuthorityName="Fabricius" baseAuthorityYear="1781" box="[151,340,1015,1041]" class="Insecta" family="Coccinellidae" genus="Harmonia" kingdom="Animalia" order="Coleoptera" pageId="123" pageNumber="124" phylum="Arthropoda" rank="species" species="octomaculata">
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<emphasis id="B91A2C661423464CBDDF358BFE99FB8A" box="[151,340,1015,1041]" italics="true" pageId="123" pageNumber="124">H. octomaculata</emphasis>
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</taxonomicName>
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but can be separated by the very pale elytral maculae. The male genitalia of
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<taxonomicName id="4C6E8BF71423464CB99F358BFA51FB8A" authority="POORANI, 2023" authorityName="Poorani" authorityYear="2023" box="[1239,1436,1015,1041]" class="Insecta" family="Coccinellidae" genus="Harmonia" kingdom="Animalia" order="Coleoptera" pageId="123" pageNumber="124" phylum="Arthropoda" rank="species" species="andamanensis" status="sp. nov.">
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<emphasis id="B91A2C661423464CB99F358BFA51FB8A" box="[1239,1436,1015,1041]" italics="true" pageId="123" pageNumber="124">H. andamanensis</emphasis>
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</taxonomicName>
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||
<taxonomicNameLabel id="A229911D1423464CBDDF3267FF14FBAE" box="[151,217,1051,1077]" pageId="123" pageNumber="124" rank="species">sp. n.</taxonomicNameLabel>
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, particularly the penis guide in ventral view (
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<figureCitation id="1355ECF11423464CBFB03267FCA7FBAE" box="[760,874,1051,1077]" captionStart="FIGURE 199" captionStartId="275.[151,250,1943,1967]" captionTargetBox="[223,1377,194,1916]" captionTargetId="figure-15@275.[203,1385,181,1920]" captionTargetPageId="275" captionText="FIGURE 199. Harmonia andamanensis sp. n.: a. adult female, dorsal view; b. adult male, dorsal view; c. apical abdominal ventrites, male; d. apical abdominal ventrites, female; e. spermatheca; f–i. male genitalia: f. tegmen, lateral view; g. tegmen, ventral view; h. penis; i. penis apex." figureDoi="http://doi.org/10.5281/zenodo.8262209" httpUri="https://zenodo.org/record/8262209/files/figure.png" pageId="123" pageNumber="124">
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Fig.
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<quantity id="4C965D911423464CBE793267FCA7FBAE" box="[817,874,1051,1077]" metricMagnitude="-1" metricUnit="kg" metricValue="1.9899999999999998" pageId="123" pageNumber="124" unit="g" value="199.0">199g</quantity>
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</figureCitation>
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||
) is similar to that of
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<taxonomicName id="4C6E8BF71423464CB9263267FE80FBC2" authority="Bielawski, 1968" authorityName="Bielawski" authorityYear="1968" class="Insecta" family="Coccinellidae" genus="Harmonia" kingdom="Animalia" order="Coleoptera" pageId="123" pageNumber="124" phylum="Arthropoda" rank="species" species="nigromarginata">
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<emphasis id="B91A2C661423464CB9263267FA51FBAF" box="[1134,1436,1051,1076]" italics="true" pageId="123" pageNumber="124">Harmonia nigromarginata</emphasis>
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Bielawski, 1968
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</taxonomicName>
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and
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<taxonomicName id="4C6E8BF71423464CBCCB3243FC8CFBC2" authority="Korschefsky, 1943" authorityName="Korschefsky" authorityYear="1943" box="[387,833,1087,1113]" class="Insecta" family="Coccinellidae" genus="Harmonia" kingdom="Animalia" order="Coleoptera" pageId="123" pageNumber="124" phylum="Arthropoda" rank="species" species="novaehebridensis">
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<emphasis id="B91A2C661423464CBCCB3243FDA4FBC2" box="[387,617,1087,1113]" italics="true" pageId="123" pageNumber="124">H. novaehebridensis</emphasis>
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||
<bibRefCitation id="EFFF8D851423464CBF383243FC8CFBC2" author="Korschefsky, R." box="[624,833,1087,1113]" pageId="123" pageNumber="124" pagination="51 - 52" refId="ref116292" refString="Korschefsky, R. (1943) Ueber Coccinelliden von den Neuen Hebriden aus dem National-Museum, Paris. Entomologische Blatter, 39, 51 - 52." type="journal article" year="1943">Korschefsky, 1943</bibRefCitation>
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</taxonomicName>
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(both Australo-Pacific species, illustrated in
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<bibRefCitation id="EFFF8D851423464CB871323CFED9FBE6" author="Slipinski, A. & Li, J. & Pang, H." pageId="123" pageNumber="124" refId="ref122780" refString="Slipinski, A., Li, J. & Pang, H. (2020) Ladybird Beetles of the Australo-Pacific Region Coleoptera: Coccinellidae: Coccinellini. CSIRO Publishing, Clayton, Victoria, 240 pp. https: // doi. org / 10.1071 / 9781486303885" type="book" year="2020">
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Ślipiński
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<emphasis id="B91A2C661423464CBDDF321FFF07FBE6" box="[151,202,1123,1149]" italics="true" pageId="123" pageNumber="124">et al</emphasis>
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. 2020
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</bibRefCitation>
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) but the apical stem of penis guide is distinctly shorter, wider and angulate in
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<taxonomicName id="4C6E8BF71423464CB9D0321FFAA6FBE7" authorityName="Bielawski" authorityYear="1968" box="[1176,1387,1123,1148]" class="Insecta" family="Coccinellidae" genus="Harmonia" kingdom="Animalia" order="Coleoptera" pageId="123" pageNumber="124" phylum="Arthropoda" rank="species" species="nigromarginata">
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<emphasis id="B91A2C661423464CB9D0321FFAA6FBE7" box="[1176,1387,1123,1148]" italics="true" pageId="123" pageNumber="124">H. nigromarginata</emphasis>
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</taxonomicName>
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and distinctly more elongate and angulate in
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<emphasis id="B91A2C661423464CBF2532FBFC93FB3A" box="[621,862,1159,1185]" italics="true" pageId="123" pageNumber="124">
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<taxonomicName id="4C6E8BF71423464CBF2532FBFC97FB3A" authorityName="Korschefsky" authorityYear="1943" box="[621,858,1159,1185]" class="Insecta" family="Coccinellidae" genus="Harmonia" kingdom="Animalia" order="Coleoptera" pageId="123" pageNumber="124" phylum="Arthropoda" rank="species" species="novaehebridensis">H. novaehebridensis</taxonomicName>
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.
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</emphasis>
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Besides, the infundibulum in the female genitalia of
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<taxonomicName id="4C6E8BF71423464CBDFF32D7FE6DFB5E" authorityName="Korschefsky" authorityYear="1943" box="[183,416,1195,1221]" class="Insecta" family="Coccinellidae" genus="Harmonia" kingdom="Animalia" order="Coleoptera" pageId="123" pageNumber="124" phylum="Arthropoda" rank="species" species="novaehebridensis">
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||
<emphasis id="B91A2C661423464CBDFF32D7FE6DFB5E" box="[183,416,1195,1221]" italics="true" pageId="123" pageNumber="124">H. novaehebridensis</emphasis>
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</taxonomicName>
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is distinctly longer, narrower and curved and the spermatheca is differently shaped in
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<taxonomicName id="4C6E8BF71423464CB8C932D7FE8BFB73" authorityName="Bielawski" authorityYear="1968" class="Insecta" family="Coccinellidae" genus="Harmonia" kingdom="Animalia" order="Coleoptera" pageId="123" pageNumber="124" phylum="Arthropoda" rank="species" species="nigromarginata">
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<emphasis id="B91A2C661423464CB8C932D7FE8BFB73" italics="true" pageId="123" pageNumber="124">H. nigromarginata</emphasis>
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</taxonomicName>
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. The elytral pattern in these two Australo-Pacific species is also distinctly different from that of
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<taxonomicName id="4C6E8BF71423464CB8C932B3FEFAFA96" authority="POORANI, 2023" authorityName="Poorani" authorityYear="2023" class="Insecta" family="Coccinellidae" genus="Harmonia" kingdom="Animalia" order="Coleoptera" pageId="123" pageNumber="124" phylum="Arthropoda" rank="species" species="andamanensis" status="sp. nov.">
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<emphasis id="B91A2C661423464CB8C932B3FEFAFA96" italics="true" pageId="123" pageNumber="124">H. andamanensis</emphasis>
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</taxonomicName>
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||
<taxonomicNameLabel id="A229911D1423464CBC76328FFEB7FA96" box="[318,378,1267,1293]" pageId="123" pageNumber="124" rank="species">sp. n.</taxonomicNameLabel>
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</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C374A3FF1423464CBD8F336AFD62F8EA" pageId="123" pageNumber="124" type="description">
|
||
<paragraph id="8BD1F0741423464CBD8F336AFD62F8EA" blockId="123.[151,1437,942,2013]" pageId="123" pageNumber="124">
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||
<emphasis id="B91A2C661423464CBD8F336AFE94FAAB" bold="true" box="[199,345,1302,1328]" pageId="123" pageNumber="124">Description.</emphasis>
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Length:
|
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<quantity id="4C965D911423464CBC89336BFDE7FAAA" box="[449,554,1303,1329]" metricMagnitude="-3" metricUnit="m" metricValue="4.2" pageId="123" pageNumber="124" unit="mm" value="4.2">4.20 mm</quantity>
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||
; width:
|
||
<quantity id="4C965D911423464CBFCF336AFD3DFAAA" box="[647,752,1302,1329]" metricMagnitude="-3" metricUnit="m" metricValue="3.5" pageId="123" pageNumber="124" unit="mm" value="3.5">3.50 mm</quantity>
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||
. Form elongate oval, dorsum strongly convex and glabrous. Ground colour in museum specimens dull straw yellow to yellow, pronotum with a roughly M-shaped median macula, often indistinct to obsolete, elytral pattern as illustrated, each elytron consisting of two dorsal and two lateral maculae enclosing a larger, roughly comma-shaped macula (
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<figureCitation id="1355ECF11423464CBEE433FFFBF8FA06" box="[940,1077,1410,1437]" captionStart="FIGURE 85" captionStartId="124.[151,250,1943,1967]" captionTargetBox="[176,1408,191,1903]" captionTargetId="figure-20@124.[166,1422,181,1919]" captionTargetPageId="124" captionText="FIGURE 85. Harmonia andamanensis Poorani, sp. n.: a. adult, dorsal view; b. adult, lateral view; c. adult, frontal view; d. abdomen, female; e. abdominal postcoxal line; f. female genitalia; g. spermatheca." figureDoi="http://doi.org/10.5281/zenodo.8261781" httpUri="https://zenodo.org/record/8261781/files/figure.png" pageId="123" pageNumber="124">Figs 85a, b</figureCitation>
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||
;
|
||
<figureCitation id="1355ECF11423464CB90D33FFFB53FA06" box="[1093,1182,1411,1437]" captionStart="FIGURE 199" captionStartId="275.[151,250,1943,1967]" captionTargetBox="[223,1377,194,1916]" captionTargetId="figure-15@275.[203,1385,181,1920]" captionTargetPageId="275" captionText="FIGURE 199. Harmonia andamanensis sp. n.: a. adult female, dorsal view; b. adult male, dorsal view; c. apical abdominal ventrites, male; d. apical abdominal ventrites, female; e. spermatheca; f–i. male genitalia: f. tegmen, lateral view; g. tegmen, ventral view; h. penis; i. penis apex." figureDoi="http://doi.org/10.5281/zenodo.8262209" httpUri="https://zenodo.org/record/8262209/files/figure.png" pageId="123" pageNumber="124">199a, b</figureCitation>
|
||
). Ventral side yellow. Head with inner margins of eyes strongly divergent; interspaces between punctures with reticulate microsculpture. Antenna compact, terminal antennomere greatly transverse. Pronotum densely punctate, punctures separated by 1–3 diameters, interspaces with reticulate microsculpture. Elytral punctures on disc finer, more widely separated and more shallowly impressed than those on pronotum, lateral sides with coarser and more deeply impressed punctures; interspaces somewhat alutaceous to smooth. Prosternal carinae subparallel, reaching beyond anterior margin of coxal cavity. Mesoventrite anteriorly very shallowly emarginate. Tarsal claws appendiculate. Abdominal postcoxal lines (
|
||
<figureCitation id="1355ECF11423464CBD923003FEFBF902" box="[218,310,1662,1689]" captionStart="FIGURE 85" captionStartId="124.[151,250,1943,1967]" captionTargetBox="[176,1408,191,1903]" captionTargetId="figure-20@124.[166,1422,181,1919]" captionTargetPageId="124" captionText="FIGURE 85. Harmonia andamanensis Poorani, sp. n.: a. adult, dorsal view; b. adult, lateral view; c. adult, frontal view; d. abdomen, female; e. abdominal postcoxal line; f. female genitalia; g. spermatheca." figureDoi="http://doi.org/10.5281/zenodo.8261781" httpUri="https://zenodo.org/record/8261781/files/figure.png" pageId="123" pageNumber="124">Fig. 85e</figureCitation>
|
||
) incomplete, recurved and apically merged with a strongly curved, angulate associated line. Ventrite 5 and 6 posteriorly gently emarginate in male (
|
||
<figureCitation id="1355ECF11423464CBFCE30DFFD3EF926" box="[646,755,1699,1725]" captionStart="FIGURE 199" captionStartId="275.[151,250,1943,1967]" captionTargetBox="[223,1377,194,1916]" captionTargetId="figure-15@275.[203,1385,181,1920]" captionTargetPageId="275" captionText="FIGURE 199. Harmonia andamanensis sp. n.: a. adult female, dorsal view; b. adult male, dorsal view; c. apical abdominal ventrites, male; d. apical abdominal ventrites, female; e. spermatheca; f–i. male genitalia: f. tegmen, lateral view; g. tegmen, ventral view; h. penis; i. penis apex." figureDoi="http://doi.org/10.5281/zenodo.8262209" httpUri="https://zenodo.org/record/8262209/files/figure.png" pageId="123" pageNumber="124">Fig. 199c</figureCitation>
|
||
), distinctly arcuate in female (
|
||
<figureCitation id="1355ECF11423464CB90830DFFB62F926" box="[1088,1199,1699,1725]" captionStart="FIGURE 199" captionStartId="275.[151,250,1943,1967]" captionTargetBox="[223,1377,194,1916]" captionTargetId="figure-15@275.[203,1385,181,1920]" captionTargetPageId="275" captionText="FIGURE 199. Harmonia andamanensis sp. n.: a. adult female, dorsal view; b. adult male, dorsal view; c. apical abdominal ventrites, male; d. apical abdominal ventrites, female; e. spermatheca; f–i. male genitalia: f. tegmen, lateral view; g. tegmen, ventral view; h. penis; i. penis apex." figureDoi="http://doi.org/10.5281/zenodo.8262209" httpUri="https://zenodo.org/record/8262209/files/figure.png" pageId="123" pageNumber="124">Fig. 199d</figureCitation>
|
||
). Male genitalia (
|
||
<figureCitation id="1355ECF11423464CB82730DFFF2FF97A" captionStart="FIGURE 199" captionStartId="275.[151,250,1943,1967]" captionTargetBox="[223,1377,194,1916]" captionTargetId="figure-15@275.[203,1385,181,1920]" captionTargetPageId="275" captionText="FIGURE 199. Harmonia andamanensis sp. n.: a. adult female, dorsal view; b. adult male, dorsal view; c. apical abdominal ventrites, male; d. apical abdominal ventrites, female; e. spermatheca; f–i. male genitalia: f. tegmen, lateral view; g. tegmen, ventral view; h. penis; i. penis apex." figureDoi="http://doi.org/10.5281/zenodo.8262209" httpUri="https://zenodo.org/record/8262209/files/figure.png" pageId="123" pageNumber="124">Fig. 199f–i</figureCitation>
|
||
) as illustrated, penis guide in lateral view (
|
||
<figureCitation id="1355ECF11423464CBF8130BBFCFEF97A" box="[713,819,1735,1761]" captionStart="FIGURE 199" captionStartId="275.[151,250,1943,1967]" captionTargetBox="[223,1377,194,1916]" captionTargetId="figure-15@275.[203,1385,181,1920]" captionTargetPageId="275" captionText="FIGURE 199. Harmonia andamanensis sp. n.: a. adult female, dorsal view; b. adult male, dorsal view; c. apical abdominal ventrites, male; d. apical abdominal ventrites, female; e. spermatheca; f–i. male genitalia: f. tegmen, lateral view; g. tegmen, ventral view; h. penis; i. penis apex." figureDoi="http://doi.org/10.5281/zenodo.8262209" httpUri="https://zenodo.org/record/8262209/files/figure.png" pageId="123" pageNumber="124">Fig. 199f</figureCitation>
|
||
) shorter than parameres, parameres basally widest and strongly curved medially, densely setose; penis guide in ventral view (
|
||
<figureCitation id="1355ECF11423464CBEE23097FBDAF89E" box="[938,1047,1771,1797]" captionStart="FIGURE 199" captionStartId="275.[151,250,1943,1967]" captionTargetBox="[223,1377,194,1916]" captionTargetId="figure-15@275.[203,1385,181,1920]" captionTargetPageId="275" captionText="FIGURE 199. Harmonia andamanensis sp. n.: a. adult female, dorsal view; b. adult male, dorsal view; c. apical abdominal ventrites, male; d. apical abdominal ventrites, female; e. spermatheca; f–i. male genitalia: f. tegmen, lateral view; g. tegmen, ventral view; h. penis; i. penis apex." figureDoi="http://doi.org/10.5281/zenodo.8262209" httpUri="https://zenodo.org/record/8262209/files/figure.png" pageId="123" pageNumber="124">
|
||
Fig.
|
||
<quantity id="4C965D911423464CBE963097FBDAF89E" box="[990,1047,1771,1797]" metricMagnitude="-1" metricUnit="kg" metricValue="1.9899999999999998" pageId="123" pageNumber="124" unit="g" value="199.0">199g</quantity>
|
||
</figureCitation>
|
||
) distinctly shorter than parameres, roughly goblet shaped, gradually narrowed from the base to a short stem, posteriorly rounded; penis (
|
||
<figureCitation id="1355ECF11423464CB86D3173FA59F8B2" box="[1317,1428,1807,1833]" captionStart="FIGURE 199" captionStartId="275.[151,250,1943,1967]" captionTargetBox="[223,1377,194,1916]" captionTargetId="figure-15@275.[203,1385,181,1920]" captionTargetPageId="275" captionText="FIGURE 199. Harmonia andamanensis sp. n.: a. adult female, dorsal view; b. adult male, dorsal view; c. apical abdominal ventrites, male; d. apical abdominal ventrites, female; e. spermatheca; f–i. male genitalia: f. tegmen, lateral view; g. tegmen, ventral view; h. penis; i. penis apex." figureDoi="http://doi.org/10.5281/zenodo.8262209" httpUri="https://zenodo.org/record/8262209/files/figure.png" pageId="123" pageNumber="124">Fig. 199h</figureCitation>
|
||
) strongly curved with a distinct capsule, penis apex (
|
||
<figureCitation id="1355ECF11423464CBFAE314FFC82F8D6" box="[742,847,1843,1869]" captionStart="FIGURE 199" captionStartId="275.[151,250,1943,1967]" captionTargetBox="[223,1377,194,1916]" captionTargetId="figure-15@275.[203,1385,181,1920]" captionTargetPageId="275" captionText="FIGURE 199. Harmonia andamanensis sp. n.: a. adult female, dorsal view; b. adult male, dorsal view; c. apical abdominal ventrites, male; d. apical abdominal ventrites, female; e. spermatheca; f–i. male genitalia: f. tegmen, lateral view; g. tegmen, ventral view; h. penis; i. penis apex." figureDoi="http://doi.org/10.5281/zenodo.8262209" httpUri="https://zenodo.org/record/8262209/files/figure.png" pageId="123" pageNumber="124">Fig. 199i</figureCitation>
|
||
) modified as illustrated. Female genitalia (
|
||
<figureCitation id="1355ECF11423464CB870314FFA59F8D6" box="[1336,1428,1842,1869]" captionStart="FIGURE 85" captionStartId="124.[151,250,1943,1967]" captionTargetBox="[176,1408,191,1903]" captionTargetId="figure-20@124.[166,1422,181,1919]" captionTargetPageId="124" captionText="FIGURE 85. Harmonia andamanensis Poorani, sp. n.: a. adult, dorsal view; b. adult, lateral view; c. adult, frontal view; d. abdomen, female; e. abdominal postcoxal line; f. female genitalia; g. spermatheca." figureDoi="http://doi.org/10.5281/zenodo.8261781" httpUri="https://zenodo.org/record/8261781/files/figure.png" pageId="123" pageNumber="124">Fig. 85f</figureCitation>
|
||
) and spermatheca (
|
||
<figureCitation id="1355ECF11423464CBC29312BFE0AF8EA" box="[353,455,1878,1905]" captionStart="FIGURE 85" captionStartId="124.[151,250,1943,1967]" captionTargetBox="[176,1408,191,1903]" captionTargetId="figure-20@124.[166,1422,181,1919]" captionTargetPageId="124" captionText="FIGURE 85. Harmonia andamanensis Poorani, sp. n.: a. adult, dorsal view; b. adult, lateral view; c. adult, frontal view; d. abdomen, female; e. abdominal postcoxal line; f. female genitalia; g. spermatheca." figureDoi="http://doi.org/10.5281/zenodo.8261781" httpUri="https://zenodo.org/record/8261781/files/figure.png" pageId="123" pageNumber="124">
|
||
Figs
|
||
<quantity id="4C965D911423464CBCD2312AFE0AF8EA" box="[410,455,1878,1905]" metricMagnitude="-2" metricUnit="kg" metricValue="8.5" pageId="123" pageNumber="124" unit="g" value="85.0">85g</quantity>
|
||
</figureCitation>
|
||
,
|
||
<figureCitation id="1355ECF11423464CBC9A312BFDC7F8EA" box="[466,522,1879,1905]" captionStart="FIGURE 199" captionStartId="275.[151,250,1943,1967]" captionTargetBox="[223,1377,194,1916]" captionTargetId="figure-15@275.[203,1385,181,1920]" captionTargetPageId="275" captionText="FIGURE 199. Harmonia andamanensis sp. n.: a. adult female, dorsal view; b. adult male, dorsal view; c. apical abdominal ventrites, male; d. apical abdominal ventrites, female; e. spermatheca; f–i. male genitalia: f. tegmen, lateral view; g. tegmen, ventral view; h. penis; i. penis apex." figureDoi="http://doi.org/10.5281/zenodo.8262209" httpUri="https://zenodo.org/record/8262209/files/figure.png" pageId="123" pageNumber="124">199e</figureCitation>
|
||
) as illustrated.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C374A3FF1423464CBD8F3106FADEF846" pageId="123" pageNumber="124" type="materials_examined">
|
||
<paragraph id="8BD1F0741423464CBD8F3106FADEF846" blockId="123.[151,1437,942,2013]" pageId="123" pageNumber="124">
|
||
<emphasis id="B91A2C661423464CBD8F3106FE7DF80F" bold="true" box="[199,432,1914,1940]" pageId="123" pageNumber="124">Material examined.</emphasis>
|
||
<materialsCitation id="3B06FA291423464CBCFF3107FBE9F822" ID-GBIF-Occurrence="4399052301" collectingDate="2014-04-23" collectionCode="NCBS" country="India" latitude="13.248306" location="North Andaman" longLatPrecision="1" longitude="92.97708" pageId="123" pageNumber="124" specimenCount="1" typeStatus="holotype">
|
||
<typeStatus id="54D54ED61423464CBCFF3107FDEFF80E" box="[439,546,1915,1941]" pageId="123" pageNumber="124" type="holotype">Holotype</typeStatus>
|
||
, female: “
|
||
<collectingCountry id="F379B0E41423464CBFD93107FD2AF80E" box="[657,743,1915,1941]" name="India" pageId="123" pageNumber="124">INDIA</collectingCountry>
|
||
: Andaman & Nicobar,
|
||
<location id="8EB1A6AF1423464CBEAD3107FB50F80E" LSID="urn:lsid:plazi:treatment:03C741621423464ABDDF353FFB2DFE12:8EB1A6AF1423464CBEAD3107FB50F80E" box="[997,1181,1915,1941]" country="India" latitude="13.248306" longLatPrecision="1" longitude="92.97708" name="North Andaman" pageId="123" pageNumber="124">North Andaman</location>
|
||
: Diglipur,
|
||
<geoCoordinate id="EE5A96B31423464CB85A3106FF66F822" degrees="13" direction="north" minutes="14" orientation="latitude" pageId="123" pageNumber="124" precision="1" seconds="53.9" value="13.248306">13°14’53.9” N</geoCoordinate>
|
||
<geoCoordinate id="EE5A96B31423464CBDFF31E2FEADF822" box="[183,352,1950,1977]" degrees="92" direction="east" minutes="58" orientation="longitude" pageId="123" pageNumber="124" precision="1" seconds="37.5" value="92.97708">92°58’37.5” E</geoCoordinate>
|
||
, 15 mts,
|
||
<date id="FFD0D6B41423464CBC9C31E3FD80F822" box="[468,589,1951,1977]" pageId="123" pageNumber="124" value="2014-04-23">
|
||
<collectingDate id="EF942F5C1423464CBC9C31E3FD80F822" box="[468,589,1951,1977]" pageId="123" pageNumber="124" value="2014-04-23">23.iv.2014</collectingDate>
|
||
</date>
|
||
,
|
||
<collectingMethod id="522F88631423464CBF1531E3FD19F822" box="[605,724,1951,1977]" pageId="123" pageNumber="124">Light trap</collectingMethod>
|
||
, Yeshwanth, H.M.” (
|
||
<collectionCode id="ED7F68B11423464CBE8531E3FBD1F822" box="[973,1052,1951,1977]" pageId="123" pageNumber="124">NCBS</collectionCode>
|
||
)
|
||
</materialsCitation>
|
||
;
|
||
<materialsCitation id="3B06FA291423464CB97C31E3FAC2F846" ID-GBIF-Occurrence="4399052367" collectingDate="2001-01-05" collectionCode="NBAIR" collectorName="S. Andaman" country="India" location="One" pageId="123" pageNumber="124" specimenCount="1" typeStatus="paratype">
|
||
<typeStatus id="54D54ED61423464CB97C31E3FB68F822" box="[1076,1189,1951,1977]" pageId="123" pageNumber="124" type="paratype">Paratypes</typeStatus>
|
||
:
|
||
<location id="8EB1A6AF1423464CB9FD31E3FB29F822" LSID="urn:lsid:plazi:treatment:03C741621423464ABDDF353FFB2DFE12:8EB1A6AF1423464CB9FD31E3FB29F822" box="[1205,1252,1951,1977]" country="India" name="One" pageId="123" pageNumber="124">One</location>
|
||
female, “South Andaman Before 1999” (
|
||
<collectionCode id="ED7F68B11423464CBCFA31BFFDDDF846" box="[434,528,1987,2013]" pageId="123" pageNumber="124">NBAIR</collectionCode>
|
||
); One male, “
|
||
<collectorName id="269B95A21423464CBFE131BFFCF8F846" box="[681,821,1987,2013]" pageId="123" pageNumber="124">S. Andaman</collectorName>
|
||
, Chidyatapu, Canopy,
|
||
<date id="FFD0D6B41423464CB97A31BEFB43F846" box="[1074,1166,1986,2013]" pageId="123" pageNumber="124" value="2001-01-05">
|
||
<collectingDate id="EF942F5C1423464CB97A31BEFB43F846" box="[1074,1166,1986,2013]" pageId="123" pageNumber="124" value="2001-01-05">5.i.2001</collectingDate>
|
||
</date>
|
||
” (
|
||
<collectionCode id="ED7F68B11423464CB9E131BFFACAF846" box="[1193,1287,1987,2013]" pageId="123" pageNumber="124">NBAIR</collectionCode>
|
||
)
|
||
</materialsCitation>
|
||
.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<caption id="DF11A0FC1424464BBDDF31EBFC16F848" ID-DOI="http://doi.org/10.5281/zenodo.8261781" ID-Zenodo-Dep="8261781" httpUri="https://zenodo.org/record/8261781/files/figure.png" pageId="124" pageNumber="125" startId="124.[151,250,1943,1967]" targetBox="[176,1408,191,1903]" targetPageId="124" targetType="figure">
|
||
<paragraph id="8BD1F0741424464BBDDF31EBFC16F848" blockId="124.[151,1436,1943,2003]" pageId="124" pageNumber="125">
|
||
<emphasis id="B91A2C661424464BBDDF31EBFEEFF834" bold="true" box="[151,290,1943,1967]" pageId="124" pageNumber="125">FIGURE 85.</emphasis>
|
||
<taxonomicName id="4C6E8BF71424464BBC6231E4FD48F834" authority="Poorani" authorityName="Poorani" authorityYear="2023" box="[298,645,1943,1967]" class="Insecta" family="Coccinellidae" genus="Harmonia" kingdom="Animalia" order="Coleoptera" pageId="124" pageNumber="125" phylum="Arthropoda" rank="species" species="andamanensis" status="sp. nov.">
|
||
<emphasis id="B91A2C661424464BBC6231E4FDE1F834" box="[298,556,1943,1967]" italics="true" pageId="124" pageNumber="125">Harmonia andamanensis</emphasis>
|
||
Poorani
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicNameLabel id="A229911D1424464BBFDA31EBFD00F835" box="[658,717,1943,1966]" pageId="124" pageNumber="125" rank="species">sp. n.</taxonomicNameLabel>
|
||
: a. adult, dorsal view; b. adult, lateral view; c. adult, frontal view; d. abdomen, female; e. abdominal postcoxal line; f. female genitalia; g. spermatheca.
|
||
</paragraph>
|
||
</caption>
|
||
<subSubSection id="C374A3FF1425464ABD8F36EBFB96FF2A" box="[199,1115,151,177]" pageId="125" pageNumber="126" type="etymology">
|
||
<paragraph id="8BD1F0741425464ABD8F36EBFB96FF2A" blockId="125.[151,1437,151,393]" box="[199,1115,151,177]" pageId="125" pageNumber="126">
|
||
<emphasis id="B91A2C661425464ABD8F36EBFE83FF2A" bold="true" box="[199,334,151,177]" pageId="125" pageNumber="126">Etymology.</emphasis>
|
||
The specific epithet is in reference to its
|
||
<typeStatus id="54D54ED61425464ABE5436EBFC81FF2A" box="[796,844,151,177]" pageId="125" pageNumber="126">type</typeStatus>
|
||
locality, the Andamans.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C374A3FF1425464ABD8F36C6FCCDFF4E" box="[199,768,186,213]" pageId="125" pageNumber="126" type="distribution">
|
||
<paragraph id="8BD1F0741425464ABD8F36C6FCCDFF4E" blockId="125.[151,1437,151,393]" box="[199,768,186,213]" pageId="125" pageNumber="126">
|
||
<emphasis id="B91A2C661425464ABD8F36C6FEACFF4F" bold="true" box="[199,353,186,212]" pageId="125" pageNumber="126">Distribution.</emphasis>
|
||
<collectingCountry id="F379B0E41425464ABC2F36C7FE68FF4E" box="[359,421,187,213]" name="India" pageId="125" pageNumber="126">India</collectingCountry>
|
||
: Endemic to Andaman Islands.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C374A3FF1425464ABD8F36A2FD31FF62" box="[199,764,222,249]" pageId="125" pageNumber="126" type="biology_ecology">
|
||
<paragraph id="8BD1F0741425464ABD8F36A2FD31FF62" blockId="125.[151,1437,151,393]" box="[199,764,222,249]" pageId="125" pageNumber="126">
|
||
<emphasis id="B91A2C661425464ABD8F36A2FE67FF63" bold="true" box="[199,426,222,248]" pageId="125" pageNumber="126">Associated habitat.</emphasis>
|
||
Attracted to light (label data).
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C374A3FF1425464ABD8F377FFB2DFE12" pageId="125" pageNumber="126" type="discussion">
|
||
<paragraph id="8BD1F0741425464ABD8F377FFB2DFE12" blockId="125.[151,1437,151,393]" pageId="125" pageNumber="126">
|
||
<emphasis id="B91A2C661425464ABD8F377FFEDCFE86" bold="true" box="[199,273,259,285]" pageId="125" pageNumber="126">Notes.</emphasis>
|
||
This appears to be the species mentioned as
|
||
<taxonomicName id="4C6E8BF71425464ABE56377FFC76FE86" box="[798,955,259,285]" class="Insecta" family="Coccinellidae" genus="Harmonia" kingdom="Animalia" order="Coleoptera" pageId="125" pageNumber="126" phylum="Arthropoda" rank="species" species="undetermined">
|
||
<emphasis id="B91A2C661425464ABE56377FFC5CFE87" box="[798,913,259,284]" italics="true" pageId="125" pageNumber="126">Harmonia</emphasis>
|
||
sp.
|
||
</taxonomicName>
|
||
from South Andaman by
|
||
<bibRefCitation id="EFFF8D851425464AB9A1377FFE97FEDA" author="Veenakumari, K. & Mohanraj, P." pageId="125" pageNumber="126" pagination="232 - 235" refId="ref123804" refString="Veenakumari, K. & Mohanraj, P. (2007) Additions to the coccinellid fauna of the Andaman Islands and the biology of the endemic Chilocorus coelosimilis Kapur 1966 (Coleoptera: Coccinellidae). Journal of the Bombay Natural History Society, 104 (2), 232 - 235." type="journal article" year="2007">Veenakumari & Mohanraj (2007)</bibRefCitation>
|
||
. It was identified by Booth (1993) as ‘’an undescribed species that is probably endemic to these islands”. The description given here is based on
|
||
<specimenCount id="9D683BFD1425464ABFF43737FC88FEFE" box="[700,837,331,357]" pageId="125" pageNumber="126" type="female">two females</specimenCount>
|
||
and
|
||
<specimenCount id="9D683BFD1425464ABE353737FC29FEFE" box="[893,996,331,357]" pageId="125" pageNumber="126" type="male">one male</specimenCount>
|
||
collected from Andamans. It is unique and does not match any of the Australo-Pacific and Indian species of
|
||
<taxonomicName id="4C6E8BF71425464ABED73713FBDFFE13" authorityName="Mulsant" authorityYear="1846" box="[927,1042,367,392]" class="Insecta" family="Coccinellidae" genus="Harmonia" kingdom="Animalia" order="Coleoptera" pageId="125" pageNumber="126" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B91A2C661425464ABED73713FBDFFE13" box="[927,1042,367,392]" italics="true" pageId="125" pageNumber="126">Harmonia</emphasis>
|
||
</taxonomicName>
|
||
known at present.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |