157 lines
19 KiB
XML
157 lines
19 KiB
XML
<document id="AAD731B27D973D0EE7737FDECA12D526" ID-CLB-Dataset="5657" ID-DOI="10.11646/zootaxa.1407.1.3" ID-GBIF-Dataset="280b385e-c15a-4f01-8306-919d8f0101c4" ID-ISSN="11755334" ID-Zenodo-Dep="5076809" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1625624403707" checkinUser="felipe" docAuthor="Johanson, Kjell Arne & Malm, Tobias" docDate="2007" docId="03C4D450FFA46A142FD1FF2DBD2B342B" docLanguage="en" docName="zt01407p022.pdf" docOrigin="Zootaxa 1407 (1)" docSource="https://www.biotaxa.org/Zootaxa/article/view/zootaxa.1407.1.3" docStyle="DocumentStyle:6581673A57F01A3145754A1E615EDFF0.4:Zootaxa.2007-2008.journal_article" docStyleId="6581673A57F01A3145754A1E615EDFF0" docStyleName="Zootaxa.2007-2008.journal_article" docStyleVersion="4" docTitle="Helicopsyche laoensis Johanson & Malm 2007, new species" docType="treatment" docVersion="5" lastPageNumber="15" masterDocId="FFFDAC28FFA66A162F46FF9ABE473151" masterDocTitle="Three new Helicopsyche from Laos (Trichoptera: Helicopsychidae)" masterLastPageNumber="22" masterPageNumber="13" pageNumber="15" updateTime="1699025939101" updateUser="ExternalLinkService" zenodo-license-document="CLOSED">
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<mods:titleInfo id="057F3190B0A702D331695B0C9C9AAEB4">
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<mods:title id="A0728732266C3C2911849DB7E8F062DC">Three new Helicopsyche from Laos (Trichoptera: Helicopsychidae)</mods:title>
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<mods:name id="7A9E1ECB9E3443243B9763A75F89FE1F" type="personal">
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<mods:namePart id="C3769EE97AD0B16B61BA9F59D8FB6BE0">Johanson, Kjell Arne</mods:namePart>
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<mods:roleTerm id="01A274564CD843C5FAC1E70718F1C016">Author</mods:roleTerm>
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</mods:role>
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<mods:namePart id="7FE0749FA73A96411F7F05F3AAE3C20A">Malm, Tobias</mods:namePart>
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<mods:typeOfResource id="E54312D5C08FF9F7762115E3EDCB4F64">text</mods:typeOfResource>
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<mods:title id="95BDFFB08B8616B3C0A50A5A550A451F">Zootaxa</mods:title>
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</mods:titleInfo>
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<mods:part id="38EEDBF2BA1A5D9475B3E98615952B59">
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<mods:date id="52D1AE42159B1C5538A7E69C93CD0E1C">2007</mods:date>
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<mods:number id="A469919C0AD5299A4F84A53D76E49F96">2007-02-15</mods:number>
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<mods:detail id="732BBEDA15F4850DE1F95E6E1DED6BF6" type="volume">
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<mods:number id="19F07CE8F395A2C7B720E01B2FF0716B">1407</mods:number>
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<mods:detail id="9990A447C905F4A6B5FD6EEE9CD9FF1A" type="issue">
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<mods:number id="8F12A0C6AC294E2E4E24040C29350B4C">1</mods:number>
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<mods:url id="636DAA2E61964A414B32E90DCA7EA5D0">https://www.biotaxa.org/Zootaxa/article/view/zootaxa.1407.1.3</mods:url>
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<mods:classification id="488CAD8E7B2025C9F8BA23536A5AF20C">journal article</mods:classification>
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<mods:identifier id="73B018E708D24256A577343210B5B718" type="CLB-Dataset">5657</mods:identifier>
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<mods:identifier id="0E82C85E3ADCBB8ED739B72BE524AE0B" type="DOI">10.11646/zootaxa.1407.1.3</mods:identifier>
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<treatment id="03C4D450FFA46A142FD1FF2DBD2B342B" ID-DOI="http://doi.org/10.5281/zenodo.5076829" ID-GBIF-Taxon="183399642" ID-Zenodo-Dep="5076829" LSID="urn:lsid:plazi:treatment:03C4D450FFA46A142FD1FF2DBD2B342B" httpUri="http://treatment.plazi.org/id/03C4D450FFA46A142FD1FF2DBD2B342B" lastPageNumber="15" pageId="2" pageNumber="15">
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<subSubSection id="C37736CDFFA46A142FD1FF2DBC783180" box="[151,575,183,209]" pageId="2" pageNumber="15" type="nomenclature">
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<paragraph id="8BD26546FFA46A142FD1FF2DBC783180" blockId="2.[151,575,183,244]" box="[151,575,183,209]" pageId="2" pageNumber="15">
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<heading id="D09AD22AFFA46A142FD1FF2DBC783180" bold="true" box="[151,575,183,209]" fontSize="11" level="1" pageId="2" pageNumber="15" reason="1">
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<emphasis id="B919B954FFA46A142FD1FF2DBC783180" bold="true" box="[151,575,183,209]" pageId="2" pageNumber="15">
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<taxonomicName id="4C6D1EC5FFA46A142FD1FF2DBFE73180" authority="Johanson & Malm, 2007" authorityName="Johanson & Malm" authorityYear="2007" box="[151,416,183,209]" class="Insecta" family="Helicopsychidae" genus="Helicopsyche" kingdom="Animalia" order="Trichoptera" pageId="2" pageNumber="15" phylum="Arthropoda" rank="species" species="laoensis" status="sp. nov.">
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<emphasis id="B919B954FFA46A142FD1FF2DBFE73180" bold="true" box="[151,416,183,209]" italics="true" pageId="2" pageNumber="15">Helicopsyche laoensis</emphasis>
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</taxonomicName>
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,
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<taxonomicNameLabel id="A22A042FFFA46A142EF6FF2DBC783180" box="[432,575,183,209]" pageId="2" pageNumber="15" rank="species">new species</taxonomicNameLabel>
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</emphasis>
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||
</heading>
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</paragraph>
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||
</subSubSection>
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||
<subSubSection id="C37736CDFFA46A142FD1FF40BDB6358E" pageId="2" pageNumber="15" type="description">
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<paragraph id="8BD26546FFA46A142FD1FF40BF7731A5" blockId="2.[151,575,183,244]" box="[151,304,218,244]" pageId="2" pageNumber="15">
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<figureCitation id="135679C3FFA46A142FD1FF40BEAF31A5" box="[151,232,218,244]" captionStart="FIGURES 2–4" captionStartId="3.[151,269,1958,1982]" captionTargetBox="[396,1228,245,1927]" captionTargetId="figure-54@3.[374,1287,168,2018]" captionTargetPageId="3" captionText="FIGURES 2–4. Distal part of forewings of the new Helicopsyche species. 2 — H. laoensis, new species; 3 — H. vongsombathi, new species; 4 — H. pathoumthongi, new species." figureDoi="http://doi.org/10.5281/zenodo.5076815" httpUri="https://zenodo.org/record/5076815/files/figure.png" pageId="2" pageNumber="15">Figs. 2</figureCitation>
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,
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<figureCitation id="135679C3FFA46A142FB3FF40BF7731A5" box="[245,304,218,244]" captionStart="FIGURES 5–12" captionStartId="4.[151,269,1697,1721]" captionTargetBox="[167,1382,245,1639]" captionTargetId="figure-153@4.[151,1436,194,1676]" captionTargetPageId="4" captionText="FIGURES 5–12. Helicopsyche laoensis, new species, holotype. 5 — right wings; 6 — abdominal sternite VI, lateral view; 7 — process on abdominal sternite VI, ventral view; 8 — genitalia, lateral view; 9 — genitalia, dorsal view; 10 — genitalia, ventral view; 11 — phallus, lateral view; 12 — phallus, ventral view." figureDoi="http://doi.org/10.5281/zenodo.5076817" httpUri="https://zenodo.org/record/5076817/files/figure.png" pageId="2" pageNumber="15">5–12</figureCitation>
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</paragraph>
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<paragraph id="8BD26546FFA46A142FD1FEBABADC30D5" blockId="2.[151,1437,288,1402]" pageId="2" pageNumber="15">
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<taxonomicName id="4C6D1EC5FFA46A142FD1FEBABFDB3068" authority="Johanson & Malm, 2007" authorityName="Johanson & Malm" authorityYear="2007" box="[151,412,288,313]" class="Insecta" family="Helicopsychidae" genus="Helicopsyche" kingdom="Animalia" order="Trichoptera" pageId="2" pageNumber="15" phylum="Arthropoda" rank="species" species="laoensis" status="sp. nov.">
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<emphasis id="B919B954FFA46A142FD1FEBABFDB3068" box="[151,412,288,313]" italics="true" pageId="2" pageNumber="15">Helicopsyche laoensis</emphasis>
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</taxonomicName>
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,
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<taxonomicNameLabel id="A22A042FFFA46A142EEDFEBABC7C306B" box="[427,571,288,314]" pageId="2" pageNumber="15" rank="species">new species</taxonomicNameLabel>
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, has unique male genitalia consisting of a long, slender primary branch of the gonocoxite posteriorly curved at its apex, in lateral view a nearly rectangular segment IX, and a pair of lateral processes on segment X basally attached to segment X shortly after the mid-length.
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</paragraph>
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<paragraph id="8BD26546FFA46A142F81FE0ABEEA33CB" blockId="2.[151,1437,288,1402]" pageId="2" pageNumber="15">
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Male. Cephalic setal warts small, oval; interantennal setal warts absent. Maxillary and labial palps with brushes of setae or sensillae on dorsal face; labial palps with distal segment very short, about 1/4 the length of segment 2; maxillary palps 2-segmented, with distal segment slightly longer than proximal segment and slightly shorter than eye diameter. Antennal scape as long as eye diameter. Tibial spurs 1, 2, 4. Forewing brownish grey with characteristic pattern of small, pale spots scattered over wing surface (
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<figureCitation id="135679C3FFA46A142B98FDAABB6E331B" box="[1246,1321,560,586]" captionStart="FIGURES 2–4" captionStartId="3.[151,269,1958,1982]" captionTargetBox="[396,1228,245,1927]" captionTargetId="figure-54@3.[374,1287,168,2018]" captionTargetPageId="3" captionText="FIGURES 2–4. Distal part of forewings of the new Helicopsyche species. 2 — H. laoensis, new species; 3 — H. vongsombathi, new species; 4 — H. pathoumthongi, new species." figureDoi="http://doi.org/10.5281/zenodo.5076815" httpUri="https://zenodo.org/record/5076815/files/figure.png" pageId="2" pageNumber="15">Fig. 2</figureCitation>
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), without modified setae along R5, length
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<quantity id="4C95C8A3FFA46A142D51FDC2BC323323" box="[535,629,600,626]" metricMagnitude="-3" metricUnit="m" metricValue="4.1" pageId="2" pageNumber="15" unit="mm" value="4.1">4.1 mm</quantity>
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; hind wing length
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||
<quantity id="4C95C8A3FFA46A142C14FDC2BDE83323" box="[850,943,600,626]" metricMagnitude="-3" metricUnit="m" metricValue="3.1" pageId="2" pageNumber="15" unit="mm" value="3.1">3.1 mm</quantity>
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, with 14 hamuli; wing venation as in
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<figureCitation id="135679C3FFA46A142A2BFDC2BEEF33CB" captionStart="FIGURES 5–12" captionStartId="4.[151,269,1697,1721]" captionTargetBox="[167,1382,245,1639]" captionTargetId="figure-153@4.[151,1436,194,1676]" captionTargetPageId="4" captionText="FIGURES 5–12. Helicopsyche laoensis, new species, holotype. 5 — right wings; 6 — abdominal sternite VI, lateral view; 7 — process on abdominal sternite VI, ventral view; 8 — genitalia, lateral view; 9 — genitalia, dorsal view; 10 — genitalia, ventral view; 11 — phallus, lateral view; 12 — phallus, ventral view." figureDoi="http://doi.org/10.5281/zenodo.5076817" httpUri="https://zenodo.org/record/5076817/files/figure.png" pageId="2" pageNumber="15">Fig. 5</figureCitation>
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||
.
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||
</paragraph>
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||
<paragraph id="8BD26546FFA46A142F81FD32BDB6358E" blockId="2.[151,1437,288,1402]" pageId="2" pageNumber="15">
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||
Male abdomen and genitalia (
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||
<figureCitation id="135679C3FFA46A142D73FD32BCFC3393" box="[565,699,680,706]" captionStart="FIGURES 5–12" captionStartId="4.[151,269,1697,1721]" captionTargetBox="[167,1382,245,1639]" captionTargetId="figure-153@4.[151,1436,194,1676]" captionTargetPageId="4" captionText="FIGURES 5–12. Helicopsyche laoensis, new species, holotype. 5 — right wings; 6 — abdominal sternite VI, lateral view; 7 — process on abdominal sternite VI, ventral view; 8 — genitalia, lateral view; 9 — genitalia, dorsal view; 10 — genitalia, ventral view; 11 — phallus, lateral view; 12 — phallus, ventral view." figureDoi="http://doi.org/10.5281/zenodo.5076817" httpUri="https://zenodo.org/record/5076817/files/figure.png" pageId="2" pageNumber="15">Figs. 6–12</figureCitation>
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). Sternal process VI short, oriented posteroventrally, tubular along its length, densely covered by long microtrichiae. Segment IX nearly rectangular in lateral view, anterodorsal corner nearly right-angled, anteroventral margin nearly straight; without sub-marginal anterior apodeme; horizontal lateral apodeme present; dorsal margin nearly straight; inner margin U-shaped in dorsal view, shallowly concave in ventral view; posterior margin weakly convex in ventral view. Segment X straight, parallel-sided in lateral view; with 8 small megasetae confined to apex; 1 pair of lateral knife-shaped processes originating shortly after mid-length, separated from segment X at base, running posteriorly in parallel with lateral margin of segment X along their lengths. Superior appendage moderately long, tubular, nearly straight. Gonocoxite primary branch slightly broadening distally in lateral view, bent posteriorly before apex; secondary branch with single apical seta, short, completely hidden by primary branch; basal branch absent; gonocoxite basal plate short, ending before anterior margin of segment IX, anterior part slightly tapering towards rounded apex; in ventral view, anterior part sharply narrowing apically. Phallus without processes, more or less uniformly curved ventrally along its length, with strongly undulating ventral margin; basal half about equally wide, posterior half nearly 3x wider than anterior half; in ventral view, anterior half about as wide as in lateral view; posterior half about 3x broader than anterior part.
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</paragraph>
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</subSubSection>
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<subSubSection id="C37736CDFFA46A142F81FB71BD2B342B" pageId="2" pageNumber="15" type="materials_examined">
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<paragraph id="8BD26546FFA46A142F81FB71BB703402" blockId="2.[151,1437,288,1402]" pageId="2" pageNumber="15">
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<materialsCitation id="3B056F1BFFA46A142F81FB71BC3C3402" ID-GBIF-Occurrence="3314719304" collectingDate="2005-04-30" collectionCode="NBCA, NRM" country="Laos" county="Nam Ha" elevation="749" latitude="21.152" location="Niam Leung" longLatPrecision="75" longitude="101.355" municipality="Lakkhammai Village" pageId="2" pageNumber="15" specimenCount="1" stateProvince="Louangnamtha" typeStatus="holotype">
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<emphasis id="B919B954FFA46A142F81FB71BD243454" bold="true" box="[199,867,1259,1285]" pageId="2" pageNumber="15">
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<typeStatus id="54D6DBE4FFA46A142F81FB71BF7E3454" box="[199,313,1259,1285]" pageId="2" pageNumber="15" type="holotype">Holotype</typeStatus>
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male:
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<collectingCountry id="F37A25D6FFA46A142EDEFB71BFAF3454" box="[408,488,1259,1285]" name="Laos" pageId="2" pageNumber="15">LAOS</collectingCountry>
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PDR:
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<collectingRegion id="49A9ABA4FFA46A142D0EFB71BD193454" box="[584,862,1259,1285]" country="Laos" name="Louangnamtha" pageId="2" pageNumber="15">Luang Namtha Prov.</collectingRegion>
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||
:
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</emphasis>
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<collectingCounty id="62B31DCAFFA46A142C36FB76BD9F3457" box="[880,984,1260,1286]" pageId="2" pageNumber="15">Nam Ha</collectingCounty>
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||
<collectionCode id="ED7CFD83FFA46A142CA3FB76BA7E3457" box="[997,1081,1260,1286]" pageId="2" pageNumber="15">NBCA</collectionCode>
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||
,
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||
<collectingMunicipality id="6BB6FF3CFFA46A142B0DFB76BB0E3457" box="[1099,1353,1260,1286]" pageId="2" pageNumber="15">Lakkhammai Village</collectingMunicipality>
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||
,
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||
<location id="8EB2339DFFA46A142A1DFB76BEA5347D" LSID="urn:lsid:plazi:treatment:03C4D450FFA46A142FD1FF2DBD2B342B:8EB2339DFFA46A142A1DFB76BEA5347D" country="Laos" county="Nam Ha" latitude="21.152" longLatPrecision="75" longitude="101.355" municipality="Lakkhammai Village" name="Niam Leung" pageId="2" pageNumber="15" stateProvince="Louangnamtha">Niam Leung</location>
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stream,
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<collectingMethod id="522C1D51FFA46A142E03FA88BFF5347D" box="[325,434,1298,1324]" pageId="2" pageNumber="15">light trap</collectingMethod>
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||
, loc. 29,
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||
<geoCoordinate id="EE590381FFA46A142D5DFA88BCD7347D" box="[539,656,1298,1324]" degrees="21.152" direction="north" orientation="latitude" pageId="2" pageNumber="15" precision="55" value="21.152">21.152°N</geoCoordinate>
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||
,
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||
<geoCoordinate id="EE590381FFA46A142DDDFA88BD50347D" box="[667,791,1298,1324]" degrees="101.355" direction="east" orientation="longitude" pageId="2" pageNumber="15" precision="55" value="101.355">101.355°E</geoCoordinate>
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||
[UTM 47Q 0744602, 2339873], el.
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||
<quantity id="4C95C8A3FFA46A142B87FA88BB4A347D" box="[1217,1293,1298,1324]" metricMagnitude="2" metricUnit="m" metricValue="7.49" pageId="2" pageNumber="15" unit="m" value="749.0">
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||
<elevation id="00408275FFA46A142B87FA88BB4A347D" box="[1217,1293,1298,1324]" metricMagnitude="2" metricUnit="m" metricValue="7.49" pageId="2" pageNumber="15" unit="m" value="749.0">749 m</elevation>
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||
</quantity>
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||
,
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||
<date id="FFD34386FFA46A142A5FFA88BBDF347D" box="[1305,1432,1298,1324]" pageId="2" pageNumber="15" value="2005-04-30">
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||
<collectingDate id="EF97BA6EFFA46A142A5FFA88BBDF347D" box="[1305,1432,1298,1324]" pageId="2" pageNumber="15" value="2005-04-30">30.iv.2005</collectingDate>
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||
</date>
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, Jönsson, Malm, Viklund (
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<collectionCode id="ED7CFD83FFA46A142E83FAA3BC4B3402" box="[453,524,1337,1363]" country="Sweden" httpUri="http://grbio.org/cool/4m0z-xq14" name="Swedish Museum of Natural History - Zoological Collections" pageId="2" pageNumber="15" type="Museum">NRM</collectionCode>
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||
, alcohol)
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||
</materialsCitation>
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||
.
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<materialsCitation id="3B056F1BFFA46A142DC0FAA2BB743402" ID-GBIF-Occurrence="3314719306" box="[646,1331,1336,1363]" collectingDate="2005-04-30" collectionCode="NBCA, NRM" country="Laos" county="Nam Ha" elevation="749" latitude="21.152" location="Niam Leung" longLatPrecision="75" longitude="101.355" municipality="Lakkhammai Village" pageId="2" pageNumber="15" specimenCount="2" specimenCount-female="1" specimenCount-male="1" stateProvince="Louangnamtha" typeStatus="paratype">
|
||
<emphasis id="B919B954FFA46A142DC0FAA2BD4A3403" bold="true" box="[646,781,1336,1362]" pageId="2" pageNumber="15">
|
||
<typeStatus id="54D6DBE4FFA46A142DC0FAA2BD4F3403" box="[646,776,1336,1362]" pageId="2" pageNumber="15" type="paratype">Paratypes</typeStatus>
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||
:
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</emphasis>
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same data as for holotype —
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<specimenCount id="9D6BAECFFFA46A142B2AFAA3BAF93402" box="[1132,1214,1337,1363]" pageId="2" pageNumber="15" type="male">1 male</specimenCount>
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||
,
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||
<specimenCount id="9D6BAECFFFA46A142B8FFAA3BB743402" box="[1225,1331,1337,1363]" pageId="2" pageNumber="15" type="female">1 female</specimenCount>
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||
</materialsCitation>
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||
.
|
||
</paragraph>
|
||
<paragraph id="8BD26546FFA46A142FD1FAFABD2B342B" blockId="2.[151,1437,288,1402]" box="[151,876,1376,1402]" pageId="2" pageNumber="15">
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Etymology.
|
||
<taxonomicName id="4C6D1EC5FFA46A142E60FAFABFC13428" authorityName="Johanson & Malm" authorityYear="2007" box="[294,390,1376,1401]" class="Insecta" family="Helicopsychidae" genus="Helicopsyche" kingdom="Animalia" order="Trichoptera" pageId="2" pageNumber="15" phylum="Arthropoda" rank="species" species="laoensis">
|
||
<emphasis id="B919B954FFA46A142E60FAFABFC13428" box="[294,390,1376,1401]" italics="true" pageId="2" pageNumber="15">laoensis</emphasis>
|
||
</taxonomicName>
|
||
, named after the
|
||
<typeStatus id="54D6DBE4FFA46A142D14FAFABCC3342B" box="[594,644,1376,1402]" pageId="2" pageNumber="15">type</typeStatus>
|
||
locality
|
||
<collectingCountry id="F37A25D6FFA46A142DADFAFABD63342B" box="[747,804,1376,1402]" name="Laos" pageId="2" pageNumber="15">Laos</collectingCountry>
|
||
PDR.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |