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<document id="4CF97908B037319E072EEB7E3FA9F1BB" ID-CLB-Dataset="27348" ID-DOI="10.11646/zootaxa.4581.1.1" ID-GBIF-Dataset="5b259853-01f6-4307-8bef-58c98ec73bd3" ID-ISSN="1175-5326" ID-Zenodo-Dep="2633842" ID-ZooBank="20A9776D-AE5F-41BC-A35B-0C5E42EDFE48" IM.bibliography_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1554804587469" checkinUser="plazi" docAuthor="Winterton, Shaun L., Martins, Caleb Califre, Makarkin, Vladimir, Ardila-Camacho, Adrian &amp; Wang, Yongjie" docDate="2019" docId="03C47176FFC38D757AD20239FF4D9390" docLanguage="en" docName="zootaxa.4581.1.1.pdf.imd" docOrigin="Zootaxa 4581 (1)" docStyle="DocumentStyle:8B0D3ECF822058C8413568C103B59429.6:Zootaxa.2001-2006.monograph" docStyleId="8B0D3ECF822058C8413568C103B59429" docStyleName="Zootaxa.2001-2006.monograph" docStyleVersion="6" docTitle="Thaumatosmylus Kruger 1913" docType="treatment" docVersion="16" lastPageNumber="66" masterDocId="FFFD090EFF838D347A450740FFA0923D" masterDocTitle="Lance lacewings of the world (Neuroptera: Archeosmylidae, Osmylidae, Saucrosmylidae): review of living and fossil genera" masterLastPageNumber="99" masterPageNumber="1" pageNumber="65" updateTime="1698719428342" updateUser="ExternalLinkService">
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<mods:title id="FF87B61B622D7811355420F2CF75EBDB">Lance lacewings of the world (Neuroptera: Archeosmylidae, Osmylidae, Saucrosmylidae): review of living and fossil genera</mods:title>
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<mods:namePart id="D94F0D333B7348FEE3C301AC7FDEABCB">Winterton, Shaun L.</mods:namePart>
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<mods:namePart id="433609055446AA35C2B09D266A2D1350">Martins, Caleb Califre</mods:namePart>
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<mods:namePart id="330CBA83C33B169931D07EB703FAE376">Makarkin, Vladimir</mods:namePart>
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<mods:namePart id="FE87B0CC7CC4E358AB7F06CDB1ADADC9">Ardila-Camacho, Adrian</mods:namePart>
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<treatment id="03C47176FFC38D757AD20239FF4D9390" ID-DOI="http://doi.org/10.5281/zenodo.5631489" ID-GBIF-Taxon="156146951" ID-Zenodo-Dep="5631489" LSID="urn:lsid:plazi:treatment:03C47176FFC38D757AD20239FF4D9390" httpUri="http://treatment.plazi.org/id/03C47176FFC38D757AD20239FF4D9390" lastPageId="65" lastPageNumber="66" pageId="64" pageNumber="65">
<subSubSection id="C37793EBFFC38D747AD20239FD8097AE" box="[151,544,1401,1427]" pageId="64" pageNumber="65" type="nomenclature">
<paragraph id="8BD2C060FFC38D747AD20239FD8097AE" blockId="64.[151,544,1401,1461]" box="[151,544,1401,1427]" pageId="64" pageNumber="65">
<heading id="D09A770CFFC38D747AD20239FD8097AE" bold="true" box="[151,544,1401,1427]" fontSize="11" level="1" pageId="64" pageNumber="65" reason="1">
<taxonomicName id="4C6DBBE3FFC38D747AD20239FD8097AE" ID-CoL="646LN" ID-ENA="1935915" authority="Kruger, 1913 a" authorityName="Kruger" authorityYear="1913" box="[151,544,1401,1427]" class="Insecta" family="Osmylidae" genus="Thaumatosmylus" kingdom="Animalia" order="Neuroptera" pageId="64" pageNumber="65" phylum="Arthropoda" rank="genus">
<emphasis id="B9191C72FFC38D747AD20239FD8097AE" bold="true" box="[151,544,1401,1427]" pageId="64" pageNumber="65">
<emphasis id="B9191C72FFC38D747AD20239FEC797AE" bold="true" box="[151,359,1401,1427]" italics="true" pageId="64" pageNumber="65">Thaumatosmylus</emphasis>
<bibRefCitation id="EFFCBD91FFC38D747B2B0239FD8097AE" author="Kruger" box="[366,544,1401,1427]" firstAuthor="Kruger" pageId="64" pageNumber="65" pagination="3 - 123" refId="ref49715" refString="Kruger, L. (1913 a) Osmylidae. Beitrage zu einer Monographie der Neuropteren-Familie der Osmyliden. II. Charakteristik der Familie, Unterfamilien und Gattungen auf Grund des Geaders. Stettiner Entomologische Zeitung, 74, 3 - 123." type="journal article" year="1913 a">Krüger, 1913a</bibRefCitation>
</emphasis>
</taxonomicName>
</heading>
</paragraph>
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<subSubSection id="C37793EBFFC38D747AD202DCFBD897C7" pageId="64" pageNumber="65" type="description">
<paragraph id="8BD2C060FFC38D747AD202DCFE839789" blockId="64.[151,544,1401,1461]" box="[151,291,1436,1461]" pageId="64" pageNumber="65">
(
<figureCitation id="1356DCE5FFC38D747ADA02DCFEBB9789" box="[159,283,1436,1461]" captionStart-0="FIGURE 43" captionStart-1="FIGURE 44" captionStartId-0="65.[151,250,1938,1960]" captionStartId-1="66.[151,250,1011,1033]" captionTargetBox-0="[176,1412,949,1924]" captionTargetBox-1="[475,1111,193,990]" captionTargetId-0="figure@65.[176,1412,949,1924]" captionTargetId-1="figure@66.[475,1111,193,990]" captionTargetPageId-0="65" captionTargetPageId-1="66" captionText-0="FIGURE 43. Spilosmylinae. Thaumatosmylus delicatus Banks." captionText-1="FIGURE 44. Spilosmylinae. Thaumatosmylus hainanus Yang (photo: Shaun L. Winterton)." httpUri-0="https://zenodo.org/record/2633929/files/figure.png" httpUri-1="https://zenodo.org/record/2633931/files/figure.png" pageId="64" pageNumber="65">Fig. 4344</figureCitation>
)
</paragraph>
<paragraph id="8BD2C060FFC38D747AD202A4FBD897C7" blockId="64.[151,1144,1508,1530]" box="[151,1144,1508,1530]" pageId="64" pageNumber="65">
(=
<taxonomicName id="4C6DBBE3FFC38D747AF102A4FE5797C7" authority="Kruger, 1913 a: 91" authorityName="Kruger" authorityPageNumber="91" authorityYear="1913" box="[180,503,1508,1530]" class="Insecta" family="Osmylidae" genus="Glenosmylus" kingdom="Animalia" order="Neuroptera" pageId="64" pageNumber="65" phylum="Arthropoda" rank="genus">
<emphasis id="B9191C72FFC38D747AF102A4FE9797C7" box="[180,311,1508,1530]" italics="true" pageId="64" pageNumber="65">Glenosmylus</emphasis>
<bibRefCitation id="EFFCBD91FFC38D747B7902A5FE7297C7" author="Kruger" box="[316,466,1508,1530]" firstAuthor="Kruger" pageId="64" pageNumber="65" pagination="3 - 123" refId="ref49715" refString="Kruger, L. (1913 a) Osmylidae. Beitrage zu einer Monographie der Neuropteren-Familie der Osmyliden. II. Charakteristik der Familie, Unterfamilien und Gattungen auf Grund des Geaders. Stettiner Entomologische Zeitung, 74, 3 - 123." type="journal article" year="1913 a">Krüger, 1913a</bibRefCitation>
: 91
</taxonomicName>
.
<typeStatus id="54D67EC2FFC38D74784402A5FD9397C7" box="[513,563,1509,1530]" pageId="64" pageNumber="65">Type</typeStatus>
species.
<taxonomicName id="4C6DBBE3FFC38D7478D502A4FB8597C7" authority="Kruger, 1913 a: 91" authorityName="Kruger" authorityPageNumber="91" authorityYear="1913" box="[656,1061,1508,1530]" class="Insecta" family="Osmylidae" genus="Glenosmylus" kingdom="Animalia" order="Neuroptera" pageId="64" pageNumber="65" phylum="Arthropoda" rank="species" species="elegans">
<emphasis id="B9191C72FFC38D7478D502A4FCC697C7" box="[656,870,1508,1530]" italics="true" pageId="64" pageNumber="65">Glenosmylus elegans</emphasis>
<bibRefCitation id="EFFCBD91FFC38D74792902A5FBA197C7" author="Kruger" box="[876,1025,1508,1530]" firstAuthor="Kruger" pageId="64" pageNumber="65" pagination="3 - 123" refId="ref49715" refString="Kruger, L. (1913 a) Osmylidae. Beitrage zu einer Monographie der Neuropteren-Familie der Osmyliden. II. Charakteristik der Familie, Unterfamilien und Gattungen auf Grund des Geaders. Stettiner Entomologische Zeitung, 74, 3 - 123." type="journal article" year="1913 a">Krüger, 1913a</bibRefCitation>
: 91
</taxonomicName>
)
<emphasis id="B9191C72FFC38D747E7602A4FBD897C7" bold="true" box="[1075,1144,1508,1530]" pageId="64" pageNumber="65">syn. n.</emphasis>
</paragraph>
</subSubSection>
<subSubSection id="C37793EBFFC38D747AD20161FC49940A" box="[151,1001,1569,1591]" pageId="64" pageNumber="65" type="materials_examined">
<paragraph id="8BD2C060FFC38D747AD20161FC49940A" blockId="64.[151,1001,1569,1591]" box="[151,1001,1569,1591]" pageId="64" pageNumber="65">
<emphasis id="B9191C72FFC38D747AD20161FE82940A" bold="true" box="[151,290,1569,1591]" pageId="64" pageNumber="65">
<typeStatus id="54D67EC2FFC38D747AD20161FF6C940A" box="[151,204,1569,1591]" pageId="64" pageNumber="65">Type</typeStatus>
species.
</emphasis>
<taxonomicName id="4C6DBBE3FFC38D747B6C0161FD41940A" authority="Gerstaecker, 1894: 168" authorityName="Gerstaecker" authorityPageNumber="168" authorityYear="1894" box="[297,737,1569,1591]" class="Insecta" family="Osmylidae" genus="Osmylus" kingdom="Animalia" order="Neuroptera" pageId="64" pageNumber="65" phylum="Arthropoda" rank="species" species="diaphanus">
<emphasis id="B9191C72FFC38D747B6C0161FE51940A" box="[297,497,1569,1591]" italics="true" pageId="64" pageNumber="65">Osmylus diaphanus</emphasis>
<bibRefCitation id="EFFCBD91FFC38D747BB30161FD11940A" author="Gerstaecker" box="[502,689,1569,1591]" firstAuthor="Gerstaecker" pageId="64" pageNumber="65" refId="ref48773" refString="Gerstaecker, C. E. A. (1894) [1893] Ueber neue und weniger gekannte Neuropteren aus der familie Megaloptera Burm. Mittheilungen aus dem Naturwissenschaftlichen Verein fur Neu-Vorpommern und Rugen, 25, 93 - 73." type="book" year="1894">Gerstaecker, 1894</bibRefCitation>
: 168
</taxonomicName>
(by original designation).
</paragraph>
</subSubSection>
<subSubSection id="C37793EBFFC38D747AD20124FB3195FC" pageId="64" pageNumber="65" type="diagnosis">
<paragraph id="8BD2C060FFC38D747AD20124FB3195FC" blockId="64.[151,1437,1636,1985]" pageId="64" pageNumber="65">
<emphasis id="B9191C72FFC38D747AD20124FEB19440" bold="true" box="[151,273,1636,1661]" pageId="64" pageNumber="65">Diagnosis.</emphasis>
Ocelli present; wings broad, ovoid to sub-triangular with expanded posterior margin at mid-length of wings; wings hyaline with sparse to dense patterning especially in FW; FW costal area relatively broad along entire length; subcostal veinlets simple and densely arranged; stem of RP short, RP with 11-15 branches, slightly sinuous distally; crossveins numerous and semi-regularly arranged in basal half of wings, more regular distally with outer gradate series distinct from other crossveins while inner gradate series (when present) is often not distinct from basal crossveins; end-twigging regular and close to wing margin in radial and medial fields of both wings, endtwigging less extensive in cubital field; M vein forked before of the origin of t RP1; FW with 24
<emphasis id="B9191C72FFC38D747EAD007DFA809569" box="[1256,1312,1853,1876]" italics="true" pageId="64" pageNumber="65">m-cu</emphasis>
crossveins before fork of M; FW lacking embossed bullae, and fenestrate areas absent in infuscate areas of wing; male genitalia with ectoproct with one or two pairs of rounded lobes; gonarcus and entoprocessus narrow and joined by extensive membrane area; female spermatheca shape variable, often with only a few lobes.
</paragraph>
</subSubSection>
<subSubSection id="C37793EBFFC28D757A8207D7FF4D9390" pageId="65" pageNumber="66" type="discussion">
<paragraph id="8BD2C060FFC28D757A8207D7FF4D9390" blockId="65.[151,1436,151,429]" pageId="65" pageNumber="66">
<emphasis id="B9191C72FFC28D757A8207D7FEE8928D" bold="true" box="[199,328,151,176]" pageId="65" pageNumber="66">Comments</emphasis>
.
<taxonomicName id="4C6DBBE3FFC28D757B1307D7FDB4928D" box="[342,532,151,176]" class="Insecta" family="Osmylidae" genus="Thaumatosmylus" kingdom="Animalia" order="Neuroptera" pageId="65" pageNumber="66" phylum="Arthropoda" rank="genus">
<emphasis id="B9191C72FFC28D757B1307D7FDB4928D" box="[342,532,151,176]" italics="true" pageId="65" pageNumber="66">Thaumatosmylus</emphasis>
</taxonomicName>
is distinguished by the presence of 24
<emphasis id="B9191C72FFC28D75799E07D9FBB3928D" box="[987,1043,153,176]" italics="true" pageId="65" pageNumber="66">m-cu</emphasis>
crossveins before the fork of M in the forewing. Other characteristics include the lack of fenestrate spots in wing markings and lack of embossed bullae. Species from continental regions have relatively narrower wings, while those found on islands (e.g.,
<collectingCountry id="F37A80F0FFC28D757AD20644FF5E9320" box="[151,254,260,285]" name="Malaysia" pageId="65" pageNumber="66">Malaysia</collectingCountry>
and
<collectingCountry id="F37A80F0FFC28D757B710644FE039320" box="[308,419,260,285]" name="Indonesia" pageId="65" pageNumber="66">Indonesia</collectingCountry>
) tend to have broader wings.
<taxonomicName id="4C6DBBE3FFC28D7578AB0644FB919320" authority="Kruger" authorityName="Kruger" authorityYear="1913" box="[750,1073,260,285]" class="Insecta" family="Osmylidae" genus="Glenosmylus" kingdom="Animalia" order="Neuroptera" pageId="65" pageNumber="66" phylum="Arthropoda" rank="species" species="elegans">
<emphasis id="B9191C72FFC28D7578AB0644FC7A9320" box="[750,986,260,285]" italics="true" pageId="65" pageNumber="66">Glenosmylus elegans</emphasis>
Krüger
</taxonomicName>
is moved to this genus based on the crossveins in the M-Cu area while
<taxonomicName id="4C6DBBE3FFC28D7578350668FD53937D" authorityName=", Esben-Petersen" authorityYear="1914" box="[624,755,295,320]" class="Insecta" family="Osmylidae" genus="Glenosmylus" kingdom="Animalia" order="Neuroptera" pageId="65" pageNumber="66" phylum="Arthropoda" rank="species" species="kruegeri">
<emphasis id="B9191C72FFC28D7578350668FD219302" box="[624,641,296,319]" italics="true" pageId="65" pageNumber="66">G</emphasis>
.
<emphasis id="B9191C72FFC28D7578D10667FD53937D" box="[660,755,295,320]" italics="true" pageId="65" pageNumber="66">kruegeri</emphasis>
</taxonomicName>
is transferred to
<taxonomicName id="4C6DBBE3FFC28D75798C0667FBCE937D" box="[969,1134,295,320]" class="Insecta" family="Osmylidae" genus="Thyridosmylus" kingdom="Animalia" order="Neuroptera" pageId="65" pageNumber="66" phylum="Arthropoda" rank="genus">
<emphasis id="B9191C72FFC28D75798C0667FBCE937D" box="[969,1134,295,320]" italics="true" pageId="65" pageNumber="66">Thyridosmylus</emphasis>
</taxonomicName>
(see below).
<taxonomicName id="4C6DBBE3FFC28D757F530667FEC19358" authority="(Banks)" authorityName="Banks" baseAuthorityName="Banks" baseAuthorityYear="1931" class="Insecta" family="Osmylidae" genus="Spilosmylus" kingdom="Animalia" order="Neuroptera" pageId="65" pageNumber="66" phylum="Arthropoda" rank="species" species="fraternus">
<emphasis id="B9191C72FFC28D757F530667FF5D9359" italics="true" pageId="65" pageNumber="66">Spilosmylus fraternus</emphasis>
(Banks)
</taxonomicName>
and
<taxonomicName id="4C6DBBE3FFC28D757BDA060DFD089358" authority="(Walker)" authorityName="Walker" baseAuthorityName="Walker" baseAuthorityYear="1853" box="[415,680,332,357]" class="Insecta" family="Osmylidae" genus="Spilosmylus" kingdom="Animalia" order="Neuroptera" pageId="65" pageNumber="66" phylum="Arthropoda" rank="species" species="conspersus">
<emphasis id="B9191C72FFC28D757BDA060DFD9B9359" box="[415,571,333,356]" italics="true" pageId="65" pageNumber="66">S. conspersus</emphasis>
(Walker)
</taxonomicName>
are transferred from
<taxonomicName id="4C6DBBE3FFC28D7579E0060CFB8B9358" box="[933,1067,332,357]" class="Insecta" family="Osmylidae" genus="Spilosmylus" kingdom="Animalia" order="Neuroptera" pageId="65" pageNumber="66" phylum="Arthropoda" rank="genus">
<emphasis id="B9191C72FFC28D7579E0060CFB8B9358" box="[933,1067,332,357]" italics="true" pageId="65" pageNumber="66">Spilosmylus</emphasis>
</taxonomicName>
since they have more than one forewing m-cu crossvein before the fork of MA-MP and lack an embossed spot along the posterior forewing margin.
</paragraph>
</subSubSection>
</treatment>
</document>