228 lines
25 KiB
XML
228 lines
25 KiB
XML
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<mods:title id="9398F538946DCA94CEE59579EA35EE96">Lance lacewings of the world (Neuroptera: Archeosmylidae, Osmylidae, Saucrosmylidae): review of living and fossil genera</mods:title>
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<mods:namePart id="94F331757DCCE0A72E206F4AB0F415B6">Winterton, Shaun L.</mods:namePart>
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<mods:namePart id="F86BF71311F53AD3ECF7357C50B04030">Martins, Caleb Califre</mods:namePart>
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<mods:namePart id="AA3C0F3D7DE7C75282F5E4E81F969BE0">Makarkin, Vladimir</mods:namePart>
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<mods:namePart id="9667605E002489E922F9272A0B973573">Ardila-Camacho, Adrian</mods:namePart>
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<mods:namePart id="02DA0E0987E883954E004E9DB33E6939">Wang, Yongjie</mods:namePart>
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<mods:title id="469B31D479645B4292EB7B64F38F24F6">Zootaxa</mods:title>
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<mods:date id="B136BBDDD8454DFF4FB50BC05ABA9D8E">2019</mods:date>
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<mods:number id="B7E29E8982FB8596C37AB250CBC18961">4581</mods:number>
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<treatment id="03C47176FFB98D0E7AD20454FE5495F7" ID-GBIF-Taxon="156146949" LSID="urn:lsid:plazi:treatment:03C47176FFB98D0E7AD20454FE5495F7" httpUri="http://treatment.plazi.org/id/03C47176FFB98D0E7AD20454FE5495F7" lastPageNumber="59" pageId="58" pageNumber="59">
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<subSubSection id="C37793EBFFB98D0E7AD20454FDE69113" box="[151,582,788,814]" pageId="58" pageNumber="59" type="nomenclature">
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<paragraph id="8BD2C060FFB98D0E7AD20454FDE69113" blockId="58.[151,582,788,814]" box="[151,582,788,814]" pageId="58" pageNumber="59">
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<heading id="D09A770CFFB98D0E7AD20454FDE69113" bold="true" box="[151,582,788,814]" fontSize="11" level="1" pageId="58" pageNumber="59" reason="1">
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<taxonomicName id="4C6DBBE3FFB98D0E7AD20454FDE69113" authority="Kruger, 1913 a" authorityName="Kruger" authorityYear="1913" box="[151,582,788,814]" class="Insecta" family="Osmylidae" kingdom="Animalia" order="Neuroptera" pageId="58" pageNumber="59" phylum="Arthropoda" rank="subFamily" subFamily="Spilosmylinae">
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<emphasis id="B9191C72FFB98D0E7AD20454FDE69113" bold="true" box="[151,582,788,814]" pageId="58" pageNumber="59">
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SPILOSMYLINAE
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<bibRefCitation id="EFFCBD91FFB98D0E7BD10454FDE69113" author="Kruger" box="[404,582,788,814]" firstAuthor="Kruger" pageId="58" pageNumber="59" pagination="3 - 123" refId="ref49715" refString="Kruger, L. (1913 a) Osmylidae. Beitrage zu einer Monographie der Neuropteren-Familie der Osmyliden. II. Charakteristik der Familie, Unterfamilien und Gattungen auf Grund des Geaders. Stettiner Entomologische Zeitung, 74, 3 - 123." type="journal article" year="1913 a">Krüger, 1913a</bibRefCitation>
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</emphasis>
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</taxonomicName>
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</heading>
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</paragraph>
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</subSubSection>
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<subSubSection id="C37793EBFFB98D0E7AD2041AFDE0914D" box="[151,576,858,880]" pageId="58" pageNumber="59" type="materials_examined">
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<paragraph id="8BD2C060FFB98D0E7AD2041AFDE0914D" blockId="58.[151,576,858,880]" box="[151,576,858,880]" pageId="58" pageNumber="59">
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<emphasis id="B9191C72FFB98D0E7AD2041AFEB6914D" bold="true" box="[151,278,858,880]" pageId="58" pageNumber="59">
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<typeStatus id="54D67EC2FFB98D0E7AD2041AFF6C914D" box="[151,204,858,880]" pageId="58" pageNumber="59">Type</typeStatus>
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genus.
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</emphasis>
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<taxonomicName id="4C6DBBE3FFB98D0E7B59041AFDE0914D" authority="Kolbe, 1897: 34" authorityName="Kolbe" authorityPageNumber="34" authorityYear="1897" box="[284,576,858,880]" class="Insecta" family="Osmylidae" genus="Spilosmylus" kingdom="Animalia" order="Neuroptera" pageId="58" pageNumber="59" phylum="Arthropoda" rank="genus">
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<emphasis id="B9191C72FFB98D0E7B59041AFE35914D" box="[284,405,858,880]" italics="true" pageId="58" pageNumber="59">Spilosmylus</emphasis>
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<bibRefCitation id="EFFCBD91FFB98D0E7BD9041AFDBC914D" author="Kolbe" box="[412,540,858,880]" firstAuthor="Kolbe" pageId="58" pageNumber="59" pagination="1 - 42" refId="ref49598" refString="Kolbe, H. J. (1897) Neuropteren. Die Netzflugler. In: Mobius, K. A. (Ed.), Die Thierwelt [Deutsch-] Ost-Afrikas und der Nachbargebiete. Band IV (Wirbellose Thiere). Dietrich Reimer, Berlin, pp. 1 - 42." type="book chapter" year="1897">Kolbe, 1897</bibRefCitation>
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: 34
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</taxonomicName>
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</paragraph>
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</subSubSection>
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<subSubSection id="C37793EBFFB98D0E7AD204DDFF4397B7" pageId="58" pageNumber="59" type="diagnosis">
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<paragraph id="8BD2C060FFB98D0E7AD204DDFF4397B7" blockId="58.[151,1437,925,1994]" pageId="58" pageNumber="59">
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<emphasis id="B9191C72FFB98D0E7AD204DDFEAA918B" bold="true" box="[151,266,925,950]" pageId="58" pageNumber="59">Diagnosis</emphasis>
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. Antenna shorter than FW; ocelli sometimes with median (or all) ocelli absent; pronotum length subequal to width; female forecoxa without processes; wing shape variable, including ovoid, sub-triangular, elongate or slightly falcate; costal area relatively narrow, subcostal veinlets simple (rarely forked) and without interlinking crossveins, densely spaced; trichosors present along entire wing margin; wings typically hyaline with dark pattern (especially FW) as mottling, bands, or fenestrations, sometimes with embossed spot present in FW CuP branches; FW with single sc-r crossvein basally; RP with 8-16 branches that may be slightly sinuous distally, 1–3 distinct gradate series in both wings, often dense crossveins in basal part of wing; FW M fork close to level with origin of RP1; MP base usually with a spur vein; HW medial area not expanded; CuA usually with more branches in HW than in FW; HW CuP short, simple, sometimes forked near wing margin; male abdominal tergites 8 and 9 as separate sclerites without scent glands, gonarcus narrow, arch-like and lacking setae, entoprocesses narrow and curved, mediuncus with a basal sclerotized arm, parameres present, fused medially as a single sclerite; female genitalia with sternite 8 as a small, knob-like process, located posteriorly proximal to gonocoxite 9, gonapophyses and gonocoxites 9 closely associated or appearing fused, spermathecal duct greatly elongate and coiled.
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</paragraph>
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</subSubSection>
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<subSubSection id="C37793EBFFB98D0E7A8202D5FE5495F7" pageId="58" pageNumber="59" type="discussion">
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<paragraph id="8BD2C060FFB98D0E7A8202D5FA2495BC" blockId="58.[151,1437,925,1994]" pageId="58" pageNumber="59">
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<emphasis id="B9191C72FFB98D0E7A8202D5FEE89793" bold="true" box="[199,328,1429,1454]" pageId="58" pageNumber="59">Comments</emphasis>
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.
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<taxonomicName id="4C6DBBE3FFB98D0E7B1002D5FE559793" authorityName="Kruger" authorityYear="1913" box="[341,501,1429,1454]" class="Insecta" family="Osmylidae" kingdom="Animalia" order="Neuroptera" pageId="58" pageNumber="59" phylum="Arthropoda" rank="subFamily" subFamily="Spilosmylinae">Spilosmylinae</taxonomicName>
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comprise three extant and two extinct genera widely distributed throughout tropical and subtropical parts of the Old World, ranging from southern Africa and
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<collectingCountry id="F37A80F0FFB98D0E7E5602F9FB3C97EF" box="[1043,1180,1465,1490]" name="Madagascar" pageId="58" pageNumber="59">Madagascar</collectingCountry>
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, throughout Asia and Australasia; while being found in
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<collectingCountry id="F37A80F0FFB98D0E7854029DFDD897CB" box="[529,632,1501,1526]" name="Australia" pageId="58" pageNumber="59">Australia</collectingCountry>
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the subfamily is restricted to northern tropical regions.
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<taxonomicName id="4C6DBBE3FFB98D0E7EA7029DFA2297CB" authorityName="Kruger" authorityYear="1913" box="[1250,1410,1501,1526]" class="Insecta" family="Osmylidae" kingdom="Animalia" order="Neuroptera" pageId="58" pageNumber="59" phylum="Arthropoda" rank="subFamily" subFamily="Spilosmylinae">Spilosmylinae</taxonomicName>
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is by far the most species rich subfamily with 118 species, the bulk of which being placed in
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<taxonomicName id="4C6DBBE3FFB98D0E7E9D0141FAFD9427" box="[1240,1373,1537,1562]" class="Insecta" family="Osmylidae" genus="Spilosmylus" kingdom="Animalia" order="Neuroptera" pageId="58" pageNumber="59" phylum="Arthropoda" rank="genus">
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<emphasis id="B9191C72FFB98D0E7E9D0141FAFD9427" box="[1240,1373,1537,1562]" italics="true" pageId="58" pageNumber="59">Spilosmylus</emphasis>
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</taxonomicName>
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. The distinction of
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<taxonomicName id="4C6DBBE3FFB98D0E7B710165FE1A9403" box="[308,442,1573,1598]" class="Insecta" family="Osmylidae" genus="Spilosmylus" kingdom="Animalia" order="Neuroptera" pageId="58" pageNumber="59" phylum="Arthropoda" rank="genus">
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<emphasis id="B9191C72FFB98D0E7B710165FE1A9403" box="[308,442,1573,1598]" italics="true" pageId="58" pageNumber="59">Spilosmylus</emphasis>
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</taxonomicName>
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from
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<taxonomicName id="4C6DBBE3FFB98D0E78440165FD1E9403" box="[513,702,1573,1598]" class="Insecta" family="Osmylidae" genus="Thaumatosmylus" kingdom="Animalia" order="Neuroptera" pageId="58" pageNumber="59" phylum="Arthropoda" rank="genus">
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<emphasis id="B9191C72FFB98D0E78440165FD1E9403" box="[513,702,1573,1598]" italics="true" pageId="58" pageNumber="59">Thaumatosmylus</emphasis>
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</taxonomicName>
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and
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<taxonomicName id="4C6DBBE3FFB98D0E78B20165FC3C9403" box="[759,924,1573,1598]" class="Insecta" family="Osmylidae" genus="Thyridosmylus" kingdom="Animalia" order="Neuroptera" pageId="58" pageNumber="59" phylum="Arthropoda" rank="genus">
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<emphasis id="B9191C72FFB98D0E78B20165FC3C9403" box="[759,924,1573,1598]" italics="true" pageId="58" pageNumber="59">Thyridosmylus</emphasis>
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</taxonomicName>
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is poorly defined and lacks characters which diagnose any one of these genera from the other two consistently across all species. Diagnostic characters presently used are themselves highly variable with multiple exceptions and species exhibiting apparent intermediate conditions. In their phylogenetic analysis,
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<bibRefCitation id="EFFCBD91FFB98D0E783E01D1FC229497" author="Winterton" box="[635,898,1681,1706]" firstAuthor="Winterton" pageId="58" pageNumber="59" pagination="555 - 574" refId="ref55173" refString="Winterton, S. L., Zhao, J., Garzon-Orduna, I., Wang, Y. & Liu, Z. (2017) Phylogeny of lance lacewings (Neuroptera: Osmylidae). Systematic Entomology, 42, 555 - 574. https: // doi. org / 10.1111 / syen. 12231" type="journal article" year="2017">
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Winterton
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<emphasis id="B9191C72FFB98D0E78B101D2FC8F9497" box="[756,815,1681,1706]" italics="true" pageId="58" pageNumber="59">et al.</emphasis>
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(2017)
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</bibRefCitation>
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recovered
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<taxonomicName id="4C6DBBE3FFB98D0E7E4701D1FB609497" box="[1026,1216,1681,1706]" class="Insecta" family="Osmylidae" genus="Thaumatosmylus" kingdom="Animalia" order="Neuroptera" pageId="58" pageNumber="59" phylum="Arthropoda" rank="genus">
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<emphasis id="B9191C72FFB98D0E7E4701D1FB609497" box="[1026,1216,1681,1706]" italics="true" pageId="58" pageNumber="59">Thaumatosmylus</emphasis>
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</taxonomicName>
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as sister to a clade containing
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<taxonomicName id="4C6DBBE3FFB98D0E7B5901F5FE0194F3" box="[284,417,1717,1742]" class="Insecta" family="Osmylidae" genus="Spilosmylus" kingdom="Animalia" order="Neuroptera" pageId="58" pageNumber="59" phylum="Arthropoda" rank="genus">
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||
<emphasis id="B9191C72FFB98D0E7B5901F5FE0194F3" box="[284,417,1717,1742]" italics="true" pageId="58" pageNumber="59">Spilosmylus</emphasis>
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</taxonomicName>
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and
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<taxonomicName id="4C6DBBE3FFB98D0E7BAF01F5FD2F94F3" box="[490,655,1717,1742]" class="Insecta" family="Osmylidae" genus="Thyridosmylus" kingdom="Animalia" order="Neuroptera" pageId="58" pageNumber="59" phylum="Arthropoda" rank="genus">
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<emphasis id="B9191C72FFB98D0E7BAF01F5FD2F94F3" box="[490,655,1717,1742]" italics="true" pageId="58" pageNumber="59">Thyridosmylus</emphasis>
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</taxonomicName>
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, with all three genera as monophyletic. Traditionally placed in
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<taxonomicName id="4C6DBBE3FFB98D0E7AD20199FE9794CF" authorityName="Kruger" authorityYear="1913" box="[151,311,1753,1778]" class="Insecta" family="Osmylidae" kingdom="Animalia" order="Neuroptera" pageId="58" pageNumber="59" phylum="Arthropoda" rank="subFamily" subFamily="Spilosmylinae">Spilosmylinae</taxonomicName>
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(e.g.,
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<bibRefCitation id="EFFCBD91FFB98D0E7BC2019AFD8394CF" author="Kruger" box="[391,547,1754,1778]" firstAuthor="Kruger" pageId="58" pageNumber="59" pagination="3 - 123" refId="ref49715" refString="Kruger, L. (1913 a) Osmylidae. Beitrage zu einer Monographie der Neuropteren-Familie der Osmyliden. II. Charakteristik der Familie, Unterfamilien und Gattungen auf Grund des Geaders. Stettiner Entomologische Zeitung, 74, 3 - 123." type="journal article" year="1913">Krüger, 1913</bibRefCitation>
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),
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<taxonomicName id="4C6DBBE3FFB98D0E7879019AFD2F94CC" box="[572,655,1754,1777]" class="Insecta" family="Osmylidae" genus="Lysmus" kingdom="Animalia" order="Neuroptera" pageId="58" pageNumber="59" phylum="Arthropoda" rank="genus">
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<emphasis id="B9191C72FFB98D0E7879019AFD2F94CC" box="[572,655,1754,1777]" italics="true" pageId="58" pageNumber="59">Lysmus</emphasis>
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</taxonomicName>
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was instead recovered in
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<taxonomicName id="4C6DBBE3FFB98D0E798D0199FBCA94CF" authorityName="Kruger" authorityYear="1913" box="[968,1130,1753,1778]" class="Insecta" family="Osmylidae" kingdom="Animalia" order="Neuroptera" pageId="58" pageNumber="59" phylum="Arthropoda" rank="subFamily" subFamily="Protosmylinae">Protosmylinae</taxonomicName>
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as sister to
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<taxonomicName id="4C6DBBE3FFB98D0E7F440199FA3594CF" box="[1281,1429,1753,1778]" class="Insecta" family="Osmylidae" genus="Gryposmylus" kingdom="Animalia" order="Neuroptera" pageId="58" pageNumber="59" phylum="Arthropoda" rank="genus">
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||
<emphasis id="B9191C72FFB98D0E7F440199FA3594CF" box="[1281,1429,1753,1778]" italics="true" pageId="58" pageNumber="59">Gryposmylus</emphasis>
|
||
</taxonomicName>
|
||
.
|
||
<taxonomicName id="4C6DBBE3FFB98D0E7AD201BDFE97952B" authorityName="Kruger" authorityYear="1913" box="[151,311,1789,1814]" class="Insecta" family="Osmylidae" kingdom="Animalia" order="Neuroptera" pageId="58" pageNumber="59" phylum="Arthropoda" rank="subFamily" subFamily="Spilosmylinae">Spilosmylinae</taxonomicName>
|
||
had been rarely treated since important taxonomic and faunistic revisions by
|
||
<bibRefCitation id="EFFCBD91FFB98D0E7EF501BDFACB9528" author="Kimmins" box="[1200,1387,1789,1814]" firstAuthor="Kimmins" pageId="58" pageNumber="59" pagination="848 - 855" refId="ref49549" refString="Kimmins, D. E. (1942) The genus Thyridosmylus Kruger, with notes on the subfamily Spilosmylinae (Neuroptera, Osmylidae). Annals and Magazine of Natural History, 9, 848 - 855. https: // doi. org / 10.1080 / 03745481.1942.9755527" type="journal article" year="1942">Kimmins (1942)</bibRefCitation>
|
||
and
|
||
<bibRefCitation id="EFFCBD91FFB98D0E7AD20062FEB79507" author="New" box="[151,279,1825,1850]" firstAuthor="New" pageId="58" pageNumber="59" pagination="447 - 452" refId="ref52722" refString="New, T. R. (1986 a) A new Australian genus of Stenosmylinae (Neuroptera: Osmylidae). Systematic Entomology, 11, 447 - 452. https: // doi. org / 10.1111 / j. 1365 - 3113.1986. tb 00537. x" type="journal article" year="1986">New (1986</bibRefCitation>
|
||
, 1988,
|
||
<bibRefCitation id="EFFCBD91FFB98D0E7B2F0062FE059507" author="New" box="[362,421,1826,1850]" firstAuthor="New" pageId="58" pageNumber="59" pagination="1 - 31" refId="ref53041" refString="New, T. R. (1991) Osmylidae (Insecta: Neuroptera) from the Oriental Region. Invertebrate Taxonomy, 5, 1 - 31. https: // doi. org / 10.1071 / IT 9910001" type="journal article" year="1991">1991</bibRefCitation>
|
||
), although several species of spilosmylines have been described recently (
|
||
<bibRefCitation id="EFFCBD91FFB98D0E7EAD0062FA349507" author="Xu" box="[1256,1428,1825,1850]" firstAuthor="Xu" pageId="58" pageNumber="59" pagination="43 - 52" refId="ref55544" refString="Xu, H., Winterton, S. L., Wang, Y. & Liu, Z. (2017) Two new species of Thyridosmylus Kruger, 1913 from Madagascar (Neuroptera, Osmylidae). Zookeys, 724, 43 - 52. https: // doi. org / 10.3897 / zookeys. 724.21057" type="journal article" year="2017">
|
||
Xu
|
||
<emphasis id="B9191C72FFB98D0E7F510062FAE49507" box="[1300,1348,1825,1850]" italics="true" pageId="58" pageNumber="59">et al</emphasis>
|
||
., 2017
|
||
</bibRefCitation>
|
||
) (
|
||
<taxonomicName id="4C6DBBE3FFB98D0E7AE50005FEE59563" box="[160,325,1861,1886]" class="Insecta" family="Osmylidae" genus="Thyridosmylus" kingdom="Animalia" order="Neuroptera" pageId="58" pageNumber="59" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B9191C72FFB98D0E7AE50005FEE59563" box="[160,325,1861,1886]" italics="true" pageId="58" pageNumber="59">Thyridosmylus</emphasis>
|
||
</taxonomicName>
|
||
) and
|
||
<bibRefCitation id="EFFCBD91FFB98D0E7BD90005FD5C9560" author="Badano & Winterton" box="[412,764,1861,1886]" firstAuthor="Badano" pageId="58" pageNumber="59" pagination="29 - 42" refId="ref47358" refString="Badano, D. & Winterton, S. L. (2017) New Philippine species of Spilosmylus Kolbe (Neuroptera, Osmylidae). Zookeys, 712, 29 - 42. https: // doi. org / 10.3897 / zookeys. 712.19883" type="journal article" year="2017">Badano & Winterton (2017)</bibRefCitation>
|
||
(
|
||
<taxonomicName id="4C6DBBE3FFB98D0E795D0005FC3E9563" box="[792,926,1861,1886]" class="Insecta" family="Osmylidae" genus="Spilosmylus" kingdom="Animalia" order="Neuroptera" pageId="58" pageNumber="59" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B9191C72FFB98D0E795D0005FC3E9563" box="[792,926,1861,1886]" italics="true" pageId="58" pageNumber="59">Spilosmylus</emphasis>
|
||
</taxonomicName>
|
||
) to complement phylogenetic studies of
|
||
<taxonomicName id="4C6DBBE3FFB98D0E7AD20029FE9C95BF" box="[151,316,1897,1922]" class="Insecta" family="Osmylidae" genus="Thyridosmylus" kingdom="Animalia" order="Neuroptera" pageId="58" pageNumber="59" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B9191C72FFB98D0E7AD20029FE9C95BF" box="[151,316,1897,1922]" italics="true" pageId="58" pageNumber="59">Thyridosmylus</emphasis>
|
||
</taxonomicName>
|
||
by
|
||
<bibRefCitation id="EFFCBD91FFB98D0E7B23002AFD9695BF" author="Wang" box="[358,566,1897,1922]" firstAuthor="Wang" pageId="58" pageNumber="59" pagination="865 - 869" refId="ref54921" refString="Wang, Y., Liu, Z., Ren, D. & Shih, C (2011 a) New Middle Jurassic kempynin osmylid lacewings from China. Acta Palaeontologica Polonica, 56, 865 - 869. https: // doi. org / 10.4202 / app. 2010.0050" type="journal article" year="2011">
|
||
Wang
|
||
<emphasis id="B9191C72FFB98D0E7BE8002AFE4595BF" box="[429,485,1897,1922]" italics="true" pageId="58" pageNumber="59">et al.</emphasis>
|
||
(2011)
|
||
</bibRefCitation>
|
||
and
|
||
<bibRefCitation id="EFFCBD91FFB98D0E78290029FC9A95BF" author="Zhao" box="[620,826,1897,1922]" firstAuthor="Zhao" pageId="58" pageNumber="59" pagination="62943" refId="ref56300" refString="Zhao, J., Winterton, S. L. & Liu, Z. (2013) Ancestral gene organization in the mitochondrial genome of Thyridosmylus langii (McLachlan, 1870) (Neuroptera: Osmylidae) and implications for lacewing evolution. PLoS ONE, 8, e 62943. https: // doi. org / 10.1371 / journal. pone. 0062943" type="journal article" year="2013">
|
||
Zhao
|
||
<emphasis id="B9191C72FFB98D0E78E8002AFD4595BF" box="[685,741,1897,1922]" italics="true" pageId="58" pageNumber="59">et al.</emphasis>
|
||
(2013)
|
||
</bibRefCitation>
|
||
. Larvae inhabit riparian zones of lotic watercourses.
|
||
</paragraph>
|
||
<paragraph id="8BD2C060FFB98D0E7A8200CDFE5495F7" blockId="58.[151,1437,925,1994]" pageId="58" pageNumber="59">
|
||
<emphasis id="B9191C72FFB98D0E7A8200CDFE39959B" bold="true" box="[199,409,1933,1958]" pageId="58" pageNumber="59">Included genera.</emphasis>
|
||
<taxonomicName id="4C6DBBE3FFB98D0E7BEC00CDFD10959B" authority="Khramov" authorityName="Khramov" authorityYear="2014" box="[425,688,1933,1958]" class="Insecta" family="Osmylidae" genus="Ensiosmylus" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Neuroptera" pageId="58" pageNumber="59" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B9191C72FFB98D0E7BEC00CDFD95959B" box="[425,565,1933,1958]" italics="true" pageId="58" pageNumber="59">Ensiosmylus</emphasis>
|
||
Khramov
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="4C6DBBE3FFB98D0E788100CDFC78959B" authority="Makarkin" authorityName="Makarkin" authorityYear="1994" box="[708,984,1933,1958]" class="Insecta" family="Osmylidae" genus="Imanosmylus" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Neuroptera" pageId="58" pageNumber="59" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B9191C72FFB98D0E788100CDFCF8959B" box="[708,856,1933,1958]" italics="true" pageId="58" pageNumber="59">Imanosmylus</emphasis>
|
||
Makarkin
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="4C6DBBE3FFB98D0E79A900CDFB6A959B" authority="Kolbe" authorityName="Kolbe" box="[1004,1226,1933,1958]" class="Insecta" family="Osmylidae" genus="Spilosmylus" kingdom="Animalia" order="Neuroptera" pageId="58" pageNumber="59" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B9191C72FFB98D0E79A900CDFBD2959B" box="[1004,1138,1933,1958]" italics="true" pageId="58" pageNumber="59">Spilosmylus</emphasis>
|
||
Kolbe
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="4C6DBBE3FFB98D0E7E9B00CDFF4895F7" authority="Kruger" authorityName="Kruger" class="Insecta" family="Osmylidae" genus="Thaumatosmylus" kingdom="Animalia" order="Neuroptera" pageId="58" pageNumber="59" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B9191C72FFB98D0E7E9B00CDFA3C959B" box="[1246,1436,1933,1958]" italics="true" pageId="58" pageNumber="59">Thaumatosmylus</emphasis>
|
||
Krüger
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="4C6DBBE3FFB98D0E7AB600F1FE5495F7" authority="Kruger." authorityName="Kruger." box="[243,500,1969,1994]" class="Insecta" family="Osmylidae" genus="Thyridosmylus" kingdom="Animalia" order="Neuroptera" pageId="58" pageNumber="59" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B9191C72FFB98D0E7AB600F1FE3895F7" box="[243,408,1969,1994]" italics="true" pageId="58" pageNumber="59">Thyridosmylus</emphasis>
|
||
Krüger.
|
||
</taxonomicName>
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |