154 lines
17 KiB
XML
154 lines
17 KiB
XML
<document id="49D6D1C786EB489F8366E782DDCAF43D" ID-CLB-Dataset="27348" ID-DOI="10.11646/zootaxa.4581.1.1" ID-GBIF-Dataset="5b259853-01f6-4307-8bef-58c98ec73bd3" ID-ISSN="1175-5326" ID-Zenodo-Dep="2633842" ID-ZooBank="20A9776D-AE5F-41BC-A35B-0C5E42EDFE48" IM.bibliography_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1554804587469" checkinUser="plazi" docAuthor="Winterton, Shaun L., Martins, Caleb Califre, Makarkin, Vladimir, Ardila-Camacho, Adrian & Wang, Yongjie" docDate="2019" docId="03C47176FF968D217AD207D8FD8C916B" docLanguage="en" docName="zootaxa.4581.1.1.pdf.imd" docOrigin="Zootaxa 4581 (1)" docStyle="DocumentStyle:8B0D3ECF822058C8413568C103B59429.6:Zootaxa.2001-2006.monograph" docStyleId="8B0D3ECF822058C8413568C103B59429" docStyleName="Zootaxa.2001-2006.monograph" docStyleVersion="6" docTitle="Kolbasinella Khramov 2014" docType="treatment" docVersion="16" lastPageNumber="22" masterDocId="FFFD090EFF838D347A450740FFA0923D" masterDocTitle="Lance lacewings of the world (Neuroptera: Archeosmylidae, Osmylidae, Saucrosmylidae): review of living and fossil genera" masterLastPageNumber="99" masterPageNumber="1" pageNumber="22" updateTime="1698719428342" updateUser="ExternalLinkService">
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<mods:mods id="8920BD93DBC4593EA1B07F75519BF173" xmlns:mods="http://www.loc.gov/mods/v3">
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<mods:titleInfo id="C01A22868A3F43177747E712FC3125FE">
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<mods:title id="32A0EFF63B6C096A0313FB3A8730E491">Lance lacewings of the world (Neuroptera: Archeosmylidae, Osmylidae, Saucrosmylidae): review of living and fossil genera</mods:title>
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<mods:name id="BBAF2DE51C831A8E9F564887A528BDCA" type="personal">
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<mods:roleTerm id="FC158C6D4A68D6AED43C9A3591866A75">Author</mods:roleTerm>
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<mods:namePart id="6AC2C528D9FA537F28206EFB30D0A2CB">Winterton, Shaun L.</mods:namePart>
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</mods:name>
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<mods:name id="86269BDE41A75CE5C8FE6C464216550D" type="personal">
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<mods:roleTerm id="24CABF194552EB385DD54869971483E2">Author</mods:roleTerm>
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</mods:role>
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<mods:namePart id="0DFD84B66E28192B38BA717AB882E2D5">Martins, Caleb Califre</mods:namePart>
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</mods:name>
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<mods:name id="C628B37795B74128A2C03123808047CC" type="personal">
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<mods:roleTerm id="BA3E1C7858D8FF93DECBD9C6A43FCCB7">Author</mods:roleTerm>
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</mods:role>
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<mods:namePart id="C6458EAB523F8BC926A083F6A61D9DC4">Makarkin, Vladimir</mods:namePart>
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</mods:name>
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<mods:name id="17FBA6E0D077089B3104080B17172676" type="personal">
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<mods:roleTerm id="CDC064346AB5E896A026D050857B72D6">Author</mods:roleTerm>
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</mods:role>
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<mods:namePart id="318E05D05C9BA4E92D32A620A7A5B11A">Ardila-Camacho, Adrian</mods:namePart>
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</mods:name>
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<mods:name id="22633F997A4690F8ADFF9D7A692AE9F7" type="personal">
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<mods:roleTerm id="AB95AD1AA33CEE49C380357928A7F5E9">Author</mods:roleTerm>
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<mods:namePart id="56B2D34CF7ABF7B96B03B1DEEEE8155E">Wang, Yongjie</mods:namePart>
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<mods:titleInfo id="A1A2B7748D176A138A5070A1BE6936B2">
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<mods:title id="D75F234ABEE8237F085BC010641BC5AA">Zootaxa</mods:title>
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<mods:part id="FE8597C8A404FCC99BC4F49DEA32278F">
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<mods:date id="33DF229FBF88F95D21B41B75E3ED8D3A">2019</mods:date>
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<mods:number id="5EFCDA9520D067056AC7164E2D6CD83F">2019-04-09</mods:number>
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<mods:detail id="4CC420B5C7D02A3A762268157588A051" type="volume">
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<mods:number id="E4FC5B63DD0C9E79A58FAC664760536F">4581</mods:number>
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<mods:detail id="890ABE898C70BC90CE4F422B0D3DACD1" type="issue">
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<mods:identifier id="9BD9CFD403FB3265EF5960982A344DFC" type="CLB-Dataset">27348</mods:identifier>
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<mods:identifier id="1824BAEC642BF0CFFA4A7018C7603E42" type="DOI">10.11646/zootaxa.4581.1.1</mods:identifier>
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<mods:identifier id="28B7102F6A43413865479D160D90337B" type="ISSN">1175-5326</mods:identifier>
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</mods:mods>
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<treatment id="03C47176FF968D217AD207D8FD8C916B" ID-DOI="http://doi.org/10.5281/zenodo.5631435" ID-GBIF-Taxon="156146913" ID-Zenodo-Dep="5631435" LSID="urn:lsid:plazi:treatment:03C47176FF968D217AD207D8FD8C916B" httpUri="http://treatment.plazi.org/id/03C47176FF968D217AD207D8FD8C916B" lastPageNumber="22" pageId="21" pageNumber="22">
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<subSubSection id="C37793EBFF968D217AD207D8FDA4928F" box="[151,516,152,178]" pageId="21" pageNumber="22" type="nomenclature">
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<paragraph id="8BD2C060FF968D217AD207D8FDA4928F" blockId="21.[151,516,152,178]" box="[151,516,152,178]" pageId="21" pageNumber="22">
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<heading id="D09A770CFF968D217AD207D8FDA4928F" bold="true" box="[151,516,152,178]" fontSize="11" level="1" pageId="21" pageNumber="22" reason="1">
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<taxonomicName id="4C6DBBE3FF968D217AD207D8FDA4928F" ID-CoL="58R6" authority="Khramov, 2014 a" authorityName="Khramov" authorityYear="2014" box="[151,516,152,178]" class="Insecta" family="Osmylidae" genus="Kolbasinella" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Neuroptera" pageId="21" pageNumber="22" phylum="Arthropoda" rank="genus">
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<emphasis id="B9191C72FF968D217AD207D8FDA4928F" bold="true" box="[151,516,152,178]" pageId="21" pageNumber="22">
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<emphasis id="B9191C72FF968D217AD207D8FE8F928F" bold="true" box="[151,303,152,178]" italics="true" pageId="21" pageNumber="22">Kolbasinella</emphasis>
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<bibRefCitation id="EFFCBD91FF968D217B7007D8FDA4928F" author="Khramov" box="[309,516,152,178]" firstAuthor="Khramov" pageId="21" pageNumber="22" pagination="300 - 309" refId="ref49340" refString="Khramov, A. V. (2014 a) Lacewings of the family Osmylidae (Insecta: Neuroptera) from the Upper Jurassic of Asia. Paleontological Journal, 48 (3), 300 - 309. https: // doi. org / 10.1134 / S 0031030114030095" type="journal article" year="2014 a">Khramov, 2014a</bibRefCitation>
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</emphasis>
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</taxonomicName>
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</heading>
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</paragraph>
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</subSubSection>
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<subSubSection id="C37793EBFF968D217AD2079EFC5392C9" box="[151,1011,222,244]" pageId="21" pageNumber="22" type="materials_examined">
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<paragraph id="8BD2C060FF968D217AD2079EFC5392C9" blockId="21.[151,1011,222,244]" box="[151,1011,222,244]" pageId="21" pageNumber="22">
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<emphasis id="B9191C72FF968D217AD2079EFE8292C9" bold="true" box="[151,290,222,244]" pageId="21" pageNumber="22">
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<typeStatus id="54D67EC2FF968D217AD2079EFF6C92C9" box="[151,204,222,244]" pageId="21" pageNumber="22">Type</typeStatus>
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species.
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</emphasis>
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<taxonomicName id="4C6DBBE3FF968D217B6C079EFD4C92C9" authority="Khramov, 2014 a: 304" authorityName="Khramov" authorityPageNumber="304" authorityYear="2014" box="[297,748,222,244]" class="Insecta" family="Osmylidae" genus="Kolbasinella" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Neuroptera" pageId="21" pageNumber="22" phylum="Arthropoda" rank="species" species="elongata">
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<emphasis id="B9191C72FF968D217B6C079EFDAA92C9" box="[297,522,222,244]" italics="true" pageId="21" pageNumber="22">Kolbasinella elongata</emphasis>
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<bibRefCitation id="EFFCBD91FF968D217855079EFD1C92C9" author="Khramov" box="[528,700,222,244]" firstAuthor="Khramov" pageId="21" pageNumber="22" pagination="300 - 309" refId="ref49340" refString="Khramov, A. V. (2014 a) Lacewings of the family Osmylidae (Insecta: Neuroptera) from the Upper Jurassic of Asia. Paleontological Journal, 48 (3), 300 - 309. https: // doi. org / 10.1134 / S 0031030114030095" type="journal article" year="2014 a">Khramov, 2014a</bibRefCitation>
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: 304
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</taxonomicName>
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(by original designation).
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</paragraph>
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</subSubSection>
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<subSubSection id="C37793EBFF968D217AD20661FC2B939B" pageId="21" pageNumber="22" type="diagnosis">
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<paragraph id="8BD2C060FF968D217AD20661FC2B939B" blockId="21.[151,1437,289,854]" pageId="21" pageNumber="22">
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<emphasis id="B9191C72FF968D217AD20661FEB19307" bold="true" box="[151,273,289,314]" pageId="21" pageNumber="22">Diagnosis.</emphasis>
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Hind wing narrow; RP1 originating at approximately 1/5 wing length; RP with 12 branches MA pectinately branched near wing margin (7 closely spaced branches); numerous regularly arranged crossveins throughout wing, outer gradate series poorly formed near wing margin; MP with only a few pectinate branches; CuA strongly pectinate (
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<emphasis id="B9191C72FF968D217BEE06CEFE659398" box="[427,453,398,421]" italics="true" pageId="21" pageNumber="22">ca</emphasis>
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. 19 very short branches); FW unknown.
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</paragraph>
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</subSubSection>
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<subSubSection id="C37793EBFF968D217A8206F1FD8C916B" pageId="21" pageNumber="22" type="discussion">
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<paragraph id="8BD2C060FF968D217A8206F1FD8C916B" blockId="21.[151,1437,289,854]" pageId="21" pageNumber="22">
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<emphasis id="B9191C72FF968D217A8206F1FEE893F7" bold="true" box="[199,328,433,458]" pageId="21" pageNumber="22">Comments</emphasis>
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.
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<taxonomicName id="4C6DBBE3FF968D217B1306F1FE4593F7" authorityName="Khramov" authorityYear="2014" box="[342,485,433,458]" class="Insecta" family="Osmylidae" genus="Kolbasinella" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Neuroptera" pageId="21" pageNumber="22" phylum="Arthropoda" rank="genus">
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<emphasis id="B9191C72FF968D217B1306F1FE4593F7" box="[342,485,433,458]" italics="true" pageId="21" pageNumber="22">Kolbasinella</emphasis>
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</taxonomicName>
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was placed in
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<taxonomicName id="4C6DBBE3FF968D2178CE06F1FCA993F7" authorityName="Navas" authorityYear="1912" box="[651,777,433,458]" class="Insecta" family="Osmylidae" kingdom="Animalia" order="Neuroptera" pageId="21" pageNumber="22" phylum="Arthropoda" rank="subFamily" subFamily="Gumillinae">Gumillinae</taxonomicName>
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by
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<bibRefCitation id="EFFCBD91FF968D21797706F1FC5A93F4" author="Khramov" box="[818,1018,433,458]" firstAuthor="Khramov" pageId="21" pageNumber="22" pagination="300 - 309" refId="ref49340" refString="Khramov, A. V. (2014 a) Lacewings of the family Osmylidae (Insecta: Neuroptera) from the Upper Jurassic of Asia. Paleontological Journal, 48 (3), 300 - 309. https: // doi. org / 10.1134 / S 0031030114030095" type="journal article" year="2014 a">Khramov (2014a)</bibRefCitation>
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due to the crossvein density and lack of trichosors in the wing margin. While these are indeed traits exhibited (in part) by
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<taxonomicName id="4C6DBBE3FF968D217EC50695FAA193D3" authorityName="Navas" authorityYear="1912" box="[1152,1281,469,494]" class="Insecta" family="Osmylidae" kingdom="Animalia" order="Neuroptera" pageId="21" pageNumber="22" phylum="Arthropoda" rank="subFamily" subFamily="Gumillinae">Gumillinae</taxonomicName>
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, they are not diagnostic of the subfamily; some
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<taxonomicName id="4C6DBBE3FF968D21786E06B9FD09902F" authorityName="Navas" authorityYear="1912" box="[555,681,505,530]" class="Insecta" family="Osmylidae" kingdom="Animalia" order="Neuroptera" pageId="21" pageNumber="22" phylum="Arthropoda" rank="subFamily" subFamily="Gumillinae">Gumillinae</taxonomicName>
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do have trichosors in the distal part of the wing (e.g.,
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<taxonomicName id="4C6DBBE3FF968D217F7506B9FA2C902F" box="[1328,1420,505,530]" class="Insecta" family="Osmylidae" genus="Gumilla" kingdom="Animalia" order="Neuroptera" pageId="21" pageNumber="22" phylum="Arthropoda" rank="genus">
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<emphasis id="B9191C72FF968D217F7506B9FA2C902F" box="[1328,1420,505,530]" italics="true" pageId="21" pageNumber="22">Gumilla</emphasis>
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</taxonomicName>
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). Unfortunately, since
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<taxonomicName id="4C6DBBE3FF968D217BC5055DFDB0900B" authorityName="Khramov" authorityYear="2014" box="[384,528,541,566]" class="Insecta" family="Osmylidae" genus="Kolbasinella" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Neuroptera" pageId="21" pageNumber="22" phylum="Arthropoda" rank="genus">
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<emphasis id="B9191C72FF968D217BC5055DFDB0900B" box="[384,528,541,566]" italics="true" pageId="21" pageNumber="22">Kolbasinella</emphasis>
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</taxonomicName>
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is only known from a hind wing, the justification presented by
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<bibRefCitation id="EFFCBD91FF968D217E91055DFA3C900B" author="Khramov" box="[1236,1436,541,566]" firstAuthor="Khramov" pageId="21" pageNumber="22" pagination="300 - 309" refId="ref49340" refString="Khramov, A. V. (2014 a) Lacewings of the family Osmylidae (Insecta: Neuroptera) from the Upper Jurassic of Asia. Paleontological Journal, 48 (3), 300 - 309. https: // doi. org / 10.1134 / S 0031030114030095" type="journal article" year="2014 a">Khramov (2014a)</bibRefCitation>
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for placement in
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<taxonomicName id="4C6DBBE3FF968D217B1E0501FE799067" authorityName="Navas" authorityYear="1912" box="[347,473,577,602]" class="Insecta" family="Osmylidae" kingdom="Animalia" order="Neuroptera" pageId="21" pageNumber="22" phylum="Arthropoda" rank="subFamily" subFamily="Gumillinae">Gumillinae</taxonomicName>
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is equivocal as important diagnostic antennal and forewing features are absent. The hind wing of
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<taxonomicName id="4C6DBBE3FF968D217B690525FE1C9043" authorityName="Khramov" authorityYear="2014" box="[300,444,613,638]" class="Insecta" family="Osmylidae" genus="Kolbasinella" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Neuroptera" pageId="21" pageNumber="22" phylum="Arthropoda" rank="genus">
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<emphasis id="B9191C72FF968D217B690525FE1C9043" box="[300,444,613,638]" italics="true" pageId="21" pageNumber="22">Kolbasinella</emphasis>
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</taxonomicName>
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does not show the vein CuP but based on the shape of the other wing veins and the wing margin, it suggests that CuP is at least not elongate and pectinately branched, therefore placing it in one of three subfamilies,
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<taxonomicName id="4C6DBBE3FF968D217B6305EDFE6990FB" box="[294,457,685,710]" class="Insecta" family="Osmylidae" kingdom="Animalia" order="Neuroptera" pageId="21" pageNumber="22" phylum="Arthropoda" rank="subFamily" subFamily="Spilosmylinae">Spilosmylinae</taxonomicName>
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,
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<taxonomicName id="4C6DBBE3FF968D217B9105EDFDD690FB" authorityName="Kruger" authorityYear="1913" box="[468,630,685,710]" class="Insecta" family="Osmylidae" kingdom="Animalia" order="Neuroptera" pageId="21" pageNumber="22" phylum="Arthropoda" rank="subFamily" subFamily="Protosmylinae">Protosmylinae</taxonomicName>
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or
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<taxonomicName id="4C6DBBE3FF968D2178DF05EDFCBC90FB" authorityName="Navas" authorityYear="1912" box="[666,796,685,710]" class="Insecta" family="Osmylidae" kingdom="Animalia" order="Neuroptera" pageId="21" pageNumber="22" phylum="Arthropoda" rank="subFamily" subFamily="Gumillinae">Gumillinae</taxonomicName>
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. However, the genus is tentatively retained in
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<taxonomicName id="4C6DBBE3FF968D217F5B05EDFA3C90FB" authorityName="Navas" authorityYear="1912" box="[1310,1436,685,710]" class="Insecta" family="Osmylidae" kingdom="Animalia" order="Neuroptera" pageId="21" pageNumber="22" phylum="Arthropoda" rank="subFamily" subFamily="Gumillinae">Gumillinae</taxonomicName>
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here as its hind wing does not possess the basal sinuate crossvein
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<emphasis id="B9191C72FF968D2179370592FC0890D7" box="[882,936,722,746]" italics="true" pageId="21" pageNumber="22">1r-m</emphasis>
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, a state diagnostic for the subfamily (
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<emphasis id="B9191C72FF968D217F090592FA2290D7" box="[1356,1410,722,746]" italics="true" pageId="21" pageNumber="22">1r-m</emphasis>
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is present in
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<taxonomicName id="4C6DBBE3FF968D217B4805B5FE0D9133" box="[269,429,757,782]" class="Insecta" family="Osmylidae" kingdom="Animalia" order="Neuroptera" pageId="21" pageNumber="22" phylum="Arthropoda" rank="subFamily" subFamily="Spilosmylinae">Spilosmylinae</taxonomicName>
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and
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<taxonomicName id="4C6DBBE3FF968D217BA305B5FD2D9133" authorityName="Kruger" authorityYear="1913" box="[486,653,757,782]" class="Insecta" family="Osmylidae" kingdom="Animalia" order="Neuroptera" pageId="21" pageNumber="22" phylum="Arthropoda" rank="subFamily" subFamily="Protosmylinae">Protosmylinae</taxonomicName>
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). The lack of end-twigging in the posterior part of the wing is typical of
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<taxonomicName id="4C6DBBE3FF968D217AF30459FE94910F" authorityName="Navas" authorityYear="1912" box="[182,308,793,818]" class="Insecta" family="Osmylidae" kingdom="Animalia" order="Neuroptera" pageId="21" pageNumber="22" phylum="Arthropoda" rank="subFamily" subFamily="Gumillinae">Gumillinae</taxonomicName>
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but could also be due to the elongate shape of the wing and the proximity of the longitudinal veins (i.e., M and Cu) to the wing margin.
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</paragraph>
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</subSubSection>
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</treatment>
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</document> |