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<mods:title id="31D77479494B8CFEB69B70044B1A86D0">Lance lacewings of the world (Neuroptera: Archeosmylidae, Osmylidae, Saucrosmylidae): review of living and fossil genera</mods:title>
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<mods:namePart id="01144277648B525E1A8D08663C36C845">Winterton, Shaun L.</mods:namePart>
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<subSubSection id="C37793EBFF908D277AD207D8FD29928F" box="[151,649,152,178]" pageId="19" pageNumber="20" type="nomenclature">
<paragraph id="8BD2C060FF908D277AD207D8FD29928F" blockId="19.[151,649,152,211]" box="[151,649,152,178]" pageId="19" pageNumber="20">
<heading id="D09A770CFF908D277AD207D8FD29928F" bold="true" box="[151,649,152,178]" fontSize="11" level="1" pageId="19" pageNumber="20" reason="1">
<taxonomicName id="4C6DBBE3FF908D277AD207D8FD29928F" ID-CoL="4D8L" authority="Panfilov in Dolin et al., 1980" authorityName="Panfilov in Dolin" authorityYear="1980" box="[151,649,152,178]" class="Insecta" family="Osmylidae" genus="Epiosmylus" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Neuroptera" pageId="19" pageNumber="20" phylum="Arthropoda" rank="genus">
<emphasis id="B9191C72FF908D277AD207D8FD29928F" bold="true" box="[151,649,152,178]" pageId="19" pageNumber="20">
<emphasis id="B9191C72FF908D277AD207D8FE83928F" bold="true" box="[151,291,152,178]" italics="true" pageId="19" pageNumber="20">Epiosmylus</emphasis>
Panfilov in Dolin
<emphasis id="B9191C72FF908D27784007D8FDE0928F" bold="true" box="[517,576,152,178]" italics="true" pageId="19" pageNumber="20">et al.</emphasis>
, 1980
</emphasis>
</taxonomicName>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C37793EBFF908D277AD207FAFF5E92EE" box="[151,254,186,211]" pageId="19" pageNumber="20" type="description">
<paragraph id="8BD2C060FF908D277AD207FAFF5E92EE" blockId="19.[151,649,152,211]" box="[151,254,186,211]" pageId="19" pageNumber="20">
(
<figureCitation id="1356DCE5FF908D277ADA07FAFF5592EE" box="[159,245,186,211]" captionStart="FIGURE 7" captionStartId="18.[151,250,1862,1884]" captionTargetBox="[184,1375,595,1775]" captionTargetId="figure@18.[153,1433,529,1871]" captionText="FIGURE 7. Gumillinae fossils. A. Allotriosmylus uniramosus Yang et al. (from Yang et al., 2010); B. Epiosmylus panfilovi Ren &amp; Yin (from Ren &amp; Yin, 2002); C. Enodinympha translucida Ren &amp; Engel (forewing); D. Nuddsia longiantennata Menon &amp; Makarkin (from Menon &amp; Makarkin, 2008); E. Nilionympha pulchella Ren &amp; Engel (from Ren &amp; Engel, 2007); F. Tenuosmylus brevineurus Wang et al. (from Wang et al., 2009b) (figures not to scale)." httpUri="https://zenodo.org/record/2644604/files/figure.png" pageId="19" pageNumber="20">Fig. 7B</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="C37793EBFF908D277AD20642FB219325" box="[151,1153,258,280]" pageId="19" pageNumber="20" type="materials_examined">
<paragraph id="8BD2C060FF908D277AD20642FB219325" blockId="19.[151,1153,258,280]" box="[151,1153,258,280]" pageId="19" pageNumber="20">
<emphasis id="B9191C72FF908D277AD20642FE829325" bold="true" box="[151,290,258,280]" pageId="19" pageNumber="20">
<typeStatus id="54D67EC2FF908D277AD20642FF6C9325" box="[151,204,258,280]" pageId="19" pageNumber="20">Type</typeStatus>
species.
</emphasis>
<taxonomicName id="4C6DBBE3FF908D277B6C0642FCD99325" authority="Panfilov in Dolin et al., 1980: 100" authorityName="Panfilov in Dolin" authorityPageNumber="100" authorityYear="1980" box="[297,889,258,280]" class="Insecta" family="Osmylidae" genus="Epiosmylus" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Neuroptera" pageId="19" pageNumber="20" phylum="Arthropoda" rank="species" species="longicornis">
<emphasis id="B9191C72FF908D277B6C0642FDB89325" box="[297,536,258,280]" italics="true" pageId="19" pageNumber="20">Epiosmylus longicornis</emphasis>
Panfilov in Dolin
<emphasis id="B9191C72FF908D2778910643FCA79325" box="[724,775,258,280]" italics="true" pageId="19" pageNumber="20">et al.</emphasis>
, 1980: 100
</taxonomicName>
(by original designation).
</paragraph>
</subSubSection>
<subSubSection id="C37793EBFF908D277AD20605FB5893D3" pageId="19" pageNumber="20" type="diagnosis">
<paragraph id="8BD2C060FF908D277AD20605FB5893D3" blockId="19.[151,1436,325,674]" pageId="19" pageNumber="20">
<emphasis id="B9191C72FF908D277AD20605FEB19363" bold="true" box="[151,273,325,350]" pageId="19" pageNumber="20">Diagnosis.</emphasis>
Wings narrow, subacute apically; RP with 56 branches; RP1 originating far from origin of RP (1/31/ 2 wing length); FW medial fork close to but distal to origin of RP; MA with few branches near wing margin in both wings; FW MP pectinately branched with 36 branches; HW MP elongate, pectinately branched; FW CuA pectinately branched (35 branches); FW CuP pectinately branched (37 branches); HW CuA elongate with widely spaced pectinately branches; HW CuP simple; FW A1 pectinate (3 widely spaced branches).
</paragraph>
</subSubSection>
<subSubSection id="C37793EBFF908D277A8206B9FEC190D9" pageId="19" pageNumber="20" type="discussion">
<paragraph id="8BD2C060FF908D277A8206B9FD1D909F" blockId="19.[151,1436,325,674]" pageId="19" pageNumber="20">
<emphasis id="B9191C72FF908D277A8206B9FEEF902F" bold="true" box="[199,335,505,530]" pageId="19" pageNumber="20">Comments.</emphasis>
The genus is the
<typeStatus id="54D67EC2FF908D27786F06BAFDFA902F" box="[554,602,506,530]" pageId="19" pageNumber="20">type</typeStatus>
of the family
<taxonomicName id="4C6DBBE3FF908D27794E06B9FAB1902C" authority="Panfilov in Dolin et al., 1980" authorityName="Panfilov in Dolin" authorityYear="1980" box="[779,1297,505,530]" class="Insecta" family="Epiosmylidae" higherTaxonomySource="GBIF" kingdom="Animalia" order="Neuroptera" pageId="19" pageNumber="20" phylum="Arthropoda" rank="family">
Epiosmylidae Panfilov in Dolin
<emphasis id="B9191C72FF908D277EC206BAFB64902F" box="[1159,1220,505,530]" italics="true" pageId="19" pageNumber="20">et al.</emphasis>
, 1980
</taxonomicName>
, which was synonymised with
<taxonomicName id="4C6DBBE3FF908D277B3D055DFE56900B" authorityName="Navas" authorityYear="1912" box="[376,502,541,566]" class="Insecta" family="Osmylidae" kingdom="Animalia" order="Neuroptera" pageId="19" pageNumber="20" phylum="Arthropoda" rank="subFamily" subFamily="Gumillinae">Gumillinae</taxonomicName>
by
<bibRefCitation id="EFFCBD91FF908D277868055EFCD6900B" author="Menon &amp; Makarkin" box="[557,886,541,566]" firstAuthor="Menon" pageId="19" pageNumber="20" pagination="149 - 162" refId="ref51782" refString="Menon, F. &amp; Makarkin, V. (2008) New fossil Lacewings and Antlion (Insecta, Neuroptera) from the lower Cretaceous Crato formation of Brazil. Paleontology, 51, 149 - 162. https: // doi. org / 10.1111 / j. 1475 - 4983.2007.00740. x" type="journal article" year="2008">Menon &amp; Makarkin (2008)</bibRefCitation>
.
<taxonomicName id="4C6DBBE3FF908D2779C2055DFBA9900B" box="[903,1033,541,566]" class="Insecta" family="Osmylidae" genus="Epiosmylus" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Neuroptera" pageId="19" pageNumber="20" phylum="Arthropoda" rank="genus">
<emphasis id="B9191C72FF908D2779C2055DFBA9900B" box="[903,1033,541,566]" italics="true" pageId="19" pageNumber="20">Epiosmylus</emphasis>
</taxonomicName>
is differentiated from most other gumilline genera by the first RP branch originating on RP at least 1/3 wing length and relatively few branches of RP (56). It appears closely related to
<taxonomicName id="4C6DBBE3FF908D2778130525FD529043" authorityName="Ren &amp; Engel" authorityYear="2007" box="[598,754,613,638]" class="Insecta" family="Osmylidae" genus="Enodinympha" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Neuroptera" pageId="19" pageNumber="20" phylum="Arthropoda" rank="genus">
<emphasis id="B9191C72FF908D2778130525FD529043" box="[598,754,613,638]" italics="true" pageId="19" pageNumber="20">Enodinympha</emphasis>
</taxonomicName>
and
<taxonomicName id="4C6DBBE3FF908D2779680525FC1D9043" box="[813,957,613,638]" class="Insecta" family="Osmylidae" genus="Tenuosmylus" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Neuroptera" pageId="19" pageNumber="20" phylum="Arthropoda" rank="genus">
<emphasis id="B9191C72FF908D2779680525FC1D9043" box="[813,957,613,638]" italics="true" pageId="19" pageNumber="20">Tenuosmylus</emphasis>
</taxonomicName>
. The wings of these genera are relatively narrow compared to other genera (e.g.,
<taxonomicName id="4C6DBBE3FF908D27781405C9FD0D909F" box="[593,685,649,674]" class="Insecta" family="Osmylidae" genus="Gumilla" kingdom="Animalia" order="Neuroptera" pageId="19" pageNumber="20" phylum="Arthropoda" rank="genus">
<emphasis id="B9191C72FF908D27781405C9FD0D909F" box="[593,685,649,674]" italics="true" pageId="19" pageNumber="20">Gumilla</emphasis>
</taxonomicName>
).
</paragraph>
<paragraph id="8BD2C060FF908D277AD2058CFEC190D9" blockId="19.[151,972,716,848]" box="[151,353,716,741]" pageId="19" pageNumber="20">
<emphasis id="B9191C72FF908D277AD2058CFEFA90D8" bold="true" box="[151,346,716,741]" pageId="19" pageNumber="20">Included species</emphasis>
.
</paragraph>
</subSubSection>
<subSubSection id="C37793EBFF908D277AD205B1FD449110" pageId="19" pageNumber="20" type="description">
<paragraph id="8BD2C060FF908D277AD205B1FC6C9135" blockId="19.[151,972,716,848]" box="[151,972,751,776]" pageId="19" pageNumber="20">
<taxonomicName id="4C6DBBE3FF908D277AD205B1FC809135" authority="Panfilov in Dolin et al., 1980" authorityName="Panfilov in Dolin" authorityYear="1980" box="[151,800,751,776]" class="Insecta" family="Osmylidae" genus="Epiosmylus" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Neuroptera" pageId="19" pageNumber="20" phylum="Arthropoda" rank="species" species="longicornis">
<emphasis id="B9191C72FF908D277AD205B1FF089135" box="[151,168,753,776]" italics="true" pageId="19" pageNumber="20">E</emphasis>
.
<emphasis id="B9191C72FF908D277AF305AFFE969135" box="[182,310,751,776]" italics="true" pageId="19" pageNumber="20">longicornis</emphasis>
Panfilov in Dolin
<emphasis id="B9191C72FF908D27784305B1FD9E9135" box="[518,574,751,776]" italics="true" pageId="19" pageNumber="20">et al.</emphasis>
, 1980 (
<collectingCountry id="F37A80F0FF908D2778D705AFFCB89135" box="[658,792,751,776]" name="Kazakhstan" pageId="19" pageNumber="20">Kazakhstan</collectingCountry>
)
</taxonomicName>
(Late Jurassic)
</paragraph>
<paragraph id="8BD2C060FF908D277AD20455FD449110" blockId="19.[151,972,716,848]" box="[151,740,788,813]" pageId="19" pageNumber="20">
<taxonomicName id="4C6DBBE3FF908D277AD20455FDBA9110" authority="Khramov, 2017" authorityName="Khramov" authorityYear="2017" box="[151,538,788,813]" class="Insecta" family="Osmylidae" genus="Enodinympha" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Neuroptera" pageId="19" pageNumber="20" phylum="Arthropoda" rank="species" species="longus">
<emphasis id="B9191C72FF908D277AD20455FEA19110" box="[151,257,788,813]" italics="true" pageId="19" pageNumber="20">E. longus</emphasis>
<bibRefCitation id="EFFCBD91FF908D277B4D0454FE159111" author="Khramov" box="[264,437,788,813]" firstAuthor="Khramov" pageId="19" pageNumber="20" pagination="59 - 68" refId="ref49424" refString="Khramov, A. V. (2017) Jurassic Lacewings (Insecta: Neuroptera) of Western Siberia. Paleontological Journal, 51, 59 - 68. https: // doi. org / 10.1134 / S 0031030117010063" type="journal article" year="2017">Khramov, 2017</bibRefCitation>
(
<collectingCountry id="F37A80F0FF908D277B810454FDB29110" box="[452,530,788,813]" name="Russia" pageId="19" pageNumber="20">Russia</collectingCountry>
)
</taxonomicName>
(Middle Jurassic)
</paragraph>
</subSubSection>
<subSubSection id="C37793EBFF908D277AD20479FCA7916D" box="[151,775,823,848]" pageId="19" pageNumber="20" type="reference_group">
<paragraph id="8BD2C060FF908D277AD20479FCA7916D" blockId="19.[151,972,716,848]" box="[151,775,823,848]" pageId="19" pageNumber="20">
<taxonomicName id="4C6DBBE3FF908D277AD20479FD9C916D" ID-CoL="6G7JD" authority="Ren &amp; Yin, 2002" authorityName="Ren &amp; Yin" authorityYear="2002" box="[151,572,823,848]" class="Insecta" family="Osmylidae" genus="Epiosmylus" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Neuroptera" pageId="19" pageNumber="20" phylum="Arthropoda" rank="species" species="panfilovi">
<emphasis id="B9191C72FF908D277AD20479FF08916D" box="[151,168,825,848]" italics="true" pageId="19" pageNumber="20">E</emphasis>
.
<emphasis id="B9191C72FF908D277AF30477FEBA916D" box="[182,282,823,848]" italics="true" pageId="19" pageNumber="20">panfilovi</emphasis>
<bibRefCitation id="EFFCBD91FF908D277B650478FE7F916D" author="Ren &amp; Yin" box="[288,479,823,848]" firstAuthor="Ren" pageId="19" pageNumber="20" pagination="274 - 277" refId="ref53792" refString="Ren, D. &amp; Yin, J. (2002) A new Middle Jurassic species of Epiosmylus from Inner Mongolia, China (Neuroptera: Osmylidae). Dong Wu Fen Lei Xue Bao, 27, 274 - 277." type="journal article" year="2002">Ren &amp; Yin, 2002</bibRefCitation>
(
<collectingCountry id="F37A80F0FF908D277BAB0477FD94916D" box="[494,564,823,848]" name="China" pageId="19" pageNumber="20">China</collectingCountry>
)
</taxonomicName>
(Middle Jurassic)
</paragraph>
</subSubSection>
</treatment>
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