280 lines
33 KiB
XML
280 lines
33 KiB
XML
<document id="3CB49AA8E718021A1E4D454E8FFB0779" ID-CLB-Dataset="31282" ID-DOI="10.11646/zootaxa.4358.2.12" ID-GBIF-Dataset="0f860d14-6b09-48af-8cdf-b2b92b65cbe3" ID-ISSN="1175-5326" ID-Zenodo-Dep="1068151" ID-ZooBank="F39DE060-F26E-431B-BB1E-10E03C7BA2F0" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1511955658708" checkinUser="plazi" docAuthor="Gharali, Babak & Evenhuis, Neal" docDate="2017" docId="03A287DD9465FFE6FF558E816C75FE3F" docLanguage="en" docName="zootaxa.4358.2.12.pdf.imf" docOrigin="Zootaxa 4358 (2)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Doliopteryx fratris Gharali & Evenhuis 2017, sp. nov." docType="treatment" docVersion="7" lastPageNumber="399" masterDocId="FF9BFFA59461FFE0FFC28E166F0CFFEF" masterDocTitle="First record of the genus Doliopteryx Hesse, 1956 (Diptera: Mythicomyiidae: Glabellulinae) from Iran, with description of two new species" masterLastPageNumber="400" masterPageNumber="393" pageNumber="397" updateTime="1698505101648" updateUser="ExternalLinkService">
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<mods:mods id="66C4F5450740909BA57ED42ED23FB62C" xmlns:mods="http://www.loc.gov/mods/v3">
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<mods:titleInfo id="29CA008F593FE603EE010D33BD5B0B4F">
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<mods:title id="DEBC3128207360C52BF0FC55DFDFB3C2">First record of the genus Doliopteryx Hesse, 1956 (Diptera: Mythicomyiidae: Glabellulinae) from Iran, with description of two new species</mods:title>
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<mods:namePart id="1FF1339097D2BAA35FF552AFC2B2BAC8">Gharali, Babak</mods:namePart>
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<mods:name id="12759ADBCD4AA425976B5EB865324049" type="personal">
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<mods:roleTerm id="9210EC42D1369259DA308D933AF0C0FD">Author</mods:roleTerm>
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<mods:namePart id="F61D314C118396F37933103454BF7186">Evenhuis, Neal</mods:namePart>
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<mods:typeOfResource id="722C5EFD90B5E8170A77186794BB1F24">text</mods:typeOfResource>
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<mods:title id="86B9D109BB730C54EFEED728C48D457C">Zootaxa</mods:title>
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<mods:part id="C48CDE10A809C8CE0A19FEA36939101E">
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<mods:date id="2BD4D929FD0839402DFBF519BF4C0E81">2017</mods:date>
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<mods:number id="922A8A0327B8E8513A9C2FD0D23473D5">2017-11-29</mods:number>
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<mods:number id="348582A4F2E1BDDCE31DF621B3E281CF">4358</mods:number>
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<mods:classification id="04C844C7759D092FB3464B50CA81658C">journal article</mods:classification>
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<mods:identifier id="82BB47F0731F7300F0B784EC61EE30D3" type="DOI">10.11646/zootaxa.4358.2.12</mods:identifier>
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<treatment id="03A287DD9465FFE6FF558E816C75FE3F" ID-DOI="http://doi.org/10.5281/zenodo.6023572" ID-GBIF-Taxon="137580841" ID-Zenodo-Dep="6023572" LSID="urn:lsid:plazi:treatment:03A287DD9465FFE6FF558E816C75FE3F" httpUri="http://treatment.plazi.org/id/03A287DD9465FFE6FF558E816C75FE3F" lastPageId="6" lastPageNumber="399" pageId="4" pageNumber="397">
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<subSubSection id="C31165409465FFE4FF558E816E06FF3C" pageId="4" pageNumber="397" type="nomenclature">
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<paragraph id="8BB436CB9465FFE4FF558E816EEFFF5D" blockId="4.[151,483,151,211]" box="[151,483,151,178]" pageId="4" pageNumber="397">
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<heading id="D0FC81A79465FFE4FF558E816EEFFF5D" bold="true" box="[151,483,151,178]" fontSize="11" level="1" pageId="4" pageNumber="397" reason="1">
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<emphasis id="B97FEAD99465FFE4FF558E816EEFFF5D" bold="true" box="[151,483,151,178]" pageId="4" pageNumber="397">
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<emphasis id="B97FEAD99465FFE4FF558E816E73FF5E" bold="true" box="[151,383,151,177]" italics="true" pageId="4" pageNumber="397">
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<taxonomicName id="4C0B4D489465FFE4FF558E816E75FF5E" authority="Gharali & Evenhuis, 2017" authorityName="Gharali & Evenhuis" authorityYear="2017" box="[151,377,151,177]" class="Insecta" family="Bombyliidae" genus="Doliopteryx" kingdom="Animalia" order="Diptera" pageId="4" pageNumber="397" phylum="Arthropoda" rank="species" species="fratris" status="sp. nov.">Doliopteryx fratris</taxonomicName>
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||
,
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||
</emphasis>
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<taxonomicNameLabel id="A24C57A29465FFE4FE478E8E6EEFFF5D" box="[389,483,152,178]" pageId="4" pageNumber="397" rank="species">sp. nov.</taxonomicNameLabel>
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</emphasis>
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||
</heading>
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||
</paragraph>
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<paragraph id="8BB436CB9465FFE4FF558EAC6E06FF3C" blockId="4.[151,483,151,211]" box="[151,266,186,211]" pageId="4" pageNumber="397">
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(
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||
<figureCitation id="13302A4E9465FFE4FF5D8EAC6FEBFF3C" box="[159,231,186,211]" captionStart="FIGURE3" captionStartId="4.[151,250,1912,1934]" captionTargetBox="[450,1137,983,1890]" captionTargetId="figure@4.[450,1137,983,1890]" captionTargetPageId="4" captionText="FIGURE3. Doliopteryx fratris, sp. nov., male holotype: a. dorsal view; b. lateral view; c. antenna(lateral view); d. wing" httpUri="https://zenodo.org/record/1068157/files/figure.png" pageId="4" pageNumber="397">Figs 3</figureCitation>
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,
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<figureCitation id="13302A4E9465FFE4FF318EAD6E0EFF3C" box="[243,258,187,211]" captionStart="FIGURE 4" captionStartId="5.[151,250,1835,1857]" captionTargetBox="[449,1137,916,1823]" captionTargetId="figure@5.[449,1137,916,1823]" captionTargetPageId="5" captionText="FIGURE 4. Doliopteryx fratris, sp. nov., male and female genitalia: a. epandrium; b. gonocoxites and gonostyli; c. genitalia without epandrium (dorsal view); d. female genitalia" httpUri="https://zenodo.org/record/1068159/files/figure.png" pageId="4" pageNumber="397">4</figureCitation>
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)
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||
</paragraph>
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||
</subSubSection>
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||
<subSubSection id="C31165409465FFE4FF558F146D67FE68" pageId="4" pageNumber="397" type="materials_examined">
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<paragraph id="8BB436CB9465FFE4FF558F146D67FE68" blockId="4.[151,1437,258,931]" pageId="4" pageNumber="397">
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<emphasis id="B97FEAD99465FFE4FF558F146E48FEF4" bold="true" box="[151,324,258,283]" pageId="4" pageNumber="397">Type material.</emphasis>
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Holotype ♂ (dried by HMDS), 20 ♂ and 20 ♀ paratypes (in 96% alcohol), Nodahak,
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<date id="FFB5100B9465FFE4FAC78F156A94FEF4" box="[1285,1432,258,283]" pageId="4" pageNumber="397" value="2017-07-13">13 July 2017</date>
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, N 35°58'32.86" E 49°35'45.57",
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<quantity id="4CF39B2E9465FFE4FDC98F316D6EFED1" box="[523,610,295,319]" metricMagnitude="3" metricUnit="m" metricValue="1.328" pageId="4" pageNumber="397" unit="m" value="1328.0">1328 m</quantity>
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a.s.l., Leg. B. Gharali (HMIM);
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<materialsCitation id="3B633C969465FFE4FC198F316EE3FE8C" ID-GBIF-Occurrence="1702642433" collectionCode="BPBM" pageId="4" pageNumber="397" specimenCount="100" specimenCount-female="56" specimenCount-male="44" typeStatus="paratype">
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||
<specimenCount id="9D0DFD429465FFE4FC198F316B18FED1" box="[987,1044,295,319]" pageId="4" pageNumber="397" type="male">44 ♂</specimenCount>
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||
and
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||
<specimenCount id="9D0DFD429465FFE4FB8C8F306B84FED1" box="[1102,1160,294,319]" pageId="4" pageNumber="397" type="female">56 ♀</specimenCount>
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||
<typeStatus id="54B088699465FFE4FB528F316BF2FED0" box="[1168,1278,295,319]" pageId="4" pageNumber="397" type="paratype">paratypes</typeStatus>
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||
, same data as holotype (in alcohol) (
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||
<collectionCode id="ED1AAE0E9465FFE4FE518F5D6EEBFE8C" box="[403,487,331,355]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34697" name="Bishop Museum" pageId="4" pageNumber="397">BPBM</collectionCode>
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||
)
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||
</materialsCitation>
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||
;
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||
<materialsCitation id="3B633C969465FFE4FE398F5D6A08FE8C" ID-GBIF-Occurrence="1702642431" box="[507,1284,330,355]" collectionCode="MTD" pageId="4" pageNumber="397" specimenCount="57" specimenCount-female="26" specimenCount-male="31" typeStatus="paratype">
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||
<specimenCount id="9D0DFD429465FFE4FE398F5D6D3FFE8D" box="[507,563,331,355]" pageId="4" pageNumber="397" type="male">31 ♂</specimenCount>
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||
and
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||
<specimenCount id="9D0DFD429465FFE4FDA98F5C6DAFFE8D" box="[619,675,330,355]" pageId="4" pageNumber="397" type="female">26 ♀</specimenCount>
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||
<typeStatus id="54B088699465FFE4FD698F5D6C15FE8C" box="[683,793,331,355]" pageId="4" pageNumber="397" type="paratype">paratypes</typeStatus>
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||
, same data as holotype (in alcohol) (
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||
<collectionCode id="ED1AAE0E9465FFE4FB7A8F5D6BF0FE8C" box="[1208,1276,331,355]" country="Germany" httpUri="http://grbio.org/cool/4j3t-w40g" name="Museum of Zoology Senckenberg Dresden" pageId="4" pageNumber="397">MTD</collectionCode>
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||
)
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||
</materialsCitation>
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||
;
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||
<materialsCitation id="3B633C969465FFE4FAD28F5D6D67FE68" ID-GBIF-Occurrence="1702642434" collectionCode="MTD" pageId="4" pageNumber="397" specimenCount="40" specimenCount-female="20" specimenCount-male="20" typeStatus="paratype">
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||
<specimenCount id="9D0DFD429465FFE4FAD28F5D6A44FE8D" box="[1296,1352,331,355]" pageId="4" pageNumber="397" type="male">20 ♂</specimenCount>
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||
and
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||
<specimenCount id="9D0DFD429465FFE4FA428F5D6FA0FE69" pageId="4" pageNumber="397" type="female">20 ♀</specimenCount>
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||
<typeStatus id="54B088699465FFE4FF718F796E2DFE68" box="[179,289,367,391]" pageId="4" pageNumber="397" type="paratype">paratypes</typeStatus>
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||
, same data as holotype (BG).
|
||
</materialsCitation>
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||
</paragraph>
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||
</subSubSection>
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||
<subSubSection id="C31165409465FFE4FF058F846DC7FD6C" pageId="4" pageNumber="397" type="diagnosis">
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||
<paragraph id="8BB436CB9465FFE4FF058F846DC7FD6C" blockId="4.[151,1437,258,931]" pageId="4" pageNumber="397">
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||
<emphasis id="B97FEAD99465FFE4FF058F846E4DFE44" bold="true" box="[199,321,402,427]" pageId="4" pageNumber="397">Diagnosis.</emphasis>
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||
In the key to the world species of
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||
<taxonomicName id="4C0B4D489465FFE4FD088F856B17FE44" authority="(Evenhuis, 2000)" authorityName="Evenhuis" authorityYear="2000" baseAuthorityName="Evenhuis" baseAuthorityYear="2000" box="[714,1051,402,427]" class="Insecta" family="Bombyliidae" genus="Doliopteryx" kingdom="Animalia" order="Diptera" pageId="4" pageNumber="397" phylum="Arthropoda" rank="genus">
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||
<emphasis id="B97FEAD99465FFE4FD088F856C5CFE44" box="[714,848,403,427]" italics="true" pageId="4" pageNumber="397">Doliopteryx</emphasis>
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||
(
|
||
<bibRefCitation id="EF9A4B3A9465FFE4FC9D8F846B1FFE44" author="Evenhuis" box="[863,1043,402,427]" pageId="4" pageNumber="397" refString="Evenhuis, N. L. (2000) A revision of the " microbombyliid " genus Doliopteryx Hesse (Diptera: Mythicomyiidae). Cimbebasia, 16, 117 - 135." type="journal article" year="2000">Evenhuis, 2000</bibRefCitation>
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||
)
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||
</taxonomicName>
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||
,
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||
<emphasis id="B97FEAD99465FFE4FBE58F856A0EFE44" box="[1063,1282,403,427]" italics="true" pageId="4" pageNumber="397">
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||
<taxonomicName id="4C0B4D489465FFE4FBE58F856BF2FE44" authority="Gharali & Evenhuis, 2017" authorityName="Gharali & Evenhuis" authorityYear="2017" box="[1063,1278,403,427]" class="Insecta" family="Bombyliidae" genus="Doliopteryx" kingdom="Animalia" order="Diptera" pageId="4" pageNumber="397" phylum="Arthropoda" rank="species" species="fratris" status="sp. nov.">Doliopteryx fratris</taxonomicName>
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||
,
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||
</emphasis>
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||
<emphasis id="B97FEAD99465FFE4FAC88F846A68FE44" bold="true" box="[1290,1380,402,427]" pageId="4" pageNumber="397">
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||
<taxonomicNameLabel id="A24C57A29465FFE4FAC88F846A68FE44" box="[1290,1380,402,427]" pageId="4" pageNumber="397" rank="species">sp. nov.</taxonomicNameLabel>
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||
</emphasis>
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runs to couplet 10 where our species is easily separated from the two described species,
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<taxonomicName id="4C0B4D489465FFE4FB898FA06A90FE20" authority=", Evenhuis" authorityName="Evenhuis" box="[1099,1436,438,463]" class="Insecta" family="Bombyliidae" genus="Doliopteryx" kingdom="Animalia" order="Diptera" pageId="4" pageNumber="397" phylum="Arthropoda" rank="species" species="tigrina">
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||
<emphasis id="B97FEAD99465FFE4FB898FA06A28FE21" box="[1099,1316,438,462]" italics="true" pageId="4" pageNumber="397">Doliopteryx tigrina</emphasis>
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||
, Evenhuis
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</taxonomicName>
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and
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<taxonomicName id="4C0B4D489465FFE4FF0B8FCD6EA5FE1C" authority=", Evenhuis" authorityName="Evenhuis" box="[201,425,474,499]" class="Insecta" family="Bombyliidae" genus="Doliopteryx" kingdom="Animalia" order="Diptera" pageId="4" pageNumber="397" phylum="Arthropoda" rank="species" species="arava">
|
||
<emphasis id="B97FEAD99465FFE4FF0B8FCD6FD1FE1D" box="[201,221,475,498]" italics="true" pageId="4" pageNumber="397">D</emphasis>
|
||
.
|
||
<emphasis id="B97FEAD99465FFE4FF2C8FCD6E23FE1D" box="[238,303,475,498]" italics="true" pageId="4" pageNumber="397">arava</emphasis>
|
||
, Evenhuis
|
||
</taxonomicName>
|
||
by the color patterns of mesonotum and scutellum. In
|
||
<emphasis id="B97FEAD99465FFE4FBEE8FCD6A05FE1C" box="[1068,1289,475,499]" italics="true" pageId="4" pageNumber="397">
|
||
<taxonomicName id="4C0B4D489465FFE4FBEE8FCD6A09FE1C" authority="Gharali & Evenhuis, 2017" authorityName="Gharali & Evenhuis" authorityYear="2017" box="[1068,1285,475,499]" class="Insecta" family="Bombyliidae" genus="Doliopteryx" kingdom="Animalia" order="Diptera" pageId="4" pageNumber="397" phylum="Arthropoda" rank="species" species="fratris" status="sp. nov.">Doliopteryx fratris</taxonomicName>
|
||
,
|
||
</emphasis>
|
||
<emphasis id="B97FEAD99465FFE4FAD18FCC6A7CFE1C" bold="true" box="[1299,1392,474,499]" pageId="4" pageNumber="397">
|
||
<taxonomicNameLabel id="A24C57A29465FFE4FAD18FCC6A7CFE1C" box="[1299,1392,474,499]" pageId="4" pageNumber="397" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
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||
the black lateral stripes of the mesonotum do not reach the scutellum; the lateral portion of the mesonotum posterior to the postpronotal lobe and the entire scutellum are yellow while in the two mentioned species, the mesonotal black lateral stripes reach the scutellum, and the lateral portion of the mesonotum posterior to the postpronotal lobe is black while the scutellum is yellow only medially.
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||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C31165409465FFE5FF058C986E4BFC88" lastPageId="5" lastPageNumber="398" pageId="4" pageNumber="397" type="description">
|
||
<paragraph id="8BB436CB9465FFE4FF058C986CE0FC4C" blockId="4.[151,1437,258,931]" pageId="4" pageNumber="397">
|
||
<emphasis id="B97FEAD99465FFE4FF058C986E55FD48" bold="true" box="[199,345,654,679]" pageId="4" pageNumber="397">Description.</emphasis>
|
||
Lengths: body:
|
||
<quantity id="4CF39B2E9465FFE4FDDA8C996D9CFD49" box="[536,656,655,679]" metricMagnitude="-3" metricUnit="m" metricValue="1.1" metricValueMax="1.2" metricValueMin="1.0" pageId="4" pageNumber="397" unit="mm" value="1.1" valueMax="1.2" valueMin="1.0">1–1.2 mm</quantity>
|
||
; wing:
|
||
<quantity id="4CF39B2E9465FFE4FD2A8C986C9DFD49" box="[744,913,654,679]" metricMagnitude="-4" metricUnit="m" metricValue="7.3" metricValueMax="7.6" metricValueMin="7.0" pageId="4" pageNumber="397" unit="mm" value="0.73" valueMax="0.76" valueMin="0.7">0.70–0.76 mm</quantity>
|
||
. Head (
|
||
<figureCitation id="13302A4E9465FFE4FC2E8C986B52FD48" box="[1004,1118,654,679]" captionStart="FIGURE3" captionStartId="4.[151,250,1912,1934]" captionTargetBox="[450,1137,983,1890]" captionTargetId="figure@4.[450,1137,983,1890]" captionTargetPageId="4" captionText="FIGURE3. Doliopteryx fratris, sp. nov., male holotype: a. dorsal view; b. lateral view; c. antenna(lateral view); d. wing" httpUri="https://zenodo.org/record/1068157/files/figure.png" pageId="4" pageNumber="397">Fig. 3a, b</figureCitation>
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||
) oval, occiput and mentum expanded posteriorly so that head is longer than high; mentum yellow, brownish basally; face yellow except oral margin narrowly black; frons yellow; occiput black dorsally, yellow laterally with oblong brownish mark on gena; vertex black except narrow yellow line just behind anterior ocellus; lateral ocelli situated at level equal to posterior eye margin, farther from each other than from anterior ocellus; eyes dichoptic; antenna (
|
||
<figureCitation id="13302A4E9465FFE4FBBE8D086BC0FCD8" box="[1148,1228,798,823]" captionStart="FIGURE3" captionStartId="4.[151,250,1912,1934]" captionTargetBox="[450,1137,983,1890]" captionTargetId="figure@4.[450,1137,983,1890]" captionTargetPageId="4" captionText="FIGURE3. Doliopteryx fratris, sp. nov., male holotype: a. dorsal view; b. lateral view; c. antenna(lateral view); d. wing" httpUri="https://zenodo.org/record/1068157/files/figure.png" pageId="4" pageNumber="397">Fig. 3c</figureCitation>
|
||
) with scape wider than long, yellow; pedicel brownish, wider than long; first flagellomere brown, oblong oval, length 1.6 times greatest width; second flagellomere brown, cylindrical, length 1.4 width, style hyaline, slightly smaller than second flagellomere; proboscis brownish, pointed apically, subequal to head length.
|
||
</paragraph>
|
||
<caption id="DF7466439465FFE4FF55896E6A63F861" box="[151,1391,1912,1934]" httpUri="https://zenodo.org/record/1068157/files/figure.png" pageId="4" pageNumber="397" targetBox="[450,1137,983,1890]" targetPageId="4">
|
||
<paragraph id="8BB436CB9465FFE4FF55896E6A63F861" blockId="4.[151,1391,1912,1934]" box="[151,1391,1912,1934]" pageId="4" pageNumber="397">
|
||
<emphasis id="B97FEAD99465FFE4FF55896E6E1EF861" bold="true" box="[151,274,1912,1934]" pageId="4" pageNumber="397">FIGURE 3.</emphasis>
|
||
<emphasis id="B97FEAD99465FFE4FEDA896E6ED2F861" box="[280,478,1912,1934]" italics="true" pageId="4" pageNumber="397">
|
||
<taxonomicName id="4C0B4D489465FFE4FEDA896E6ED6F861" authority="Gharali & Evenhuis, 2017" authorityName="Gharali & Evenhuis" authorityYear="2017" box="[280,474,1912,1934]" class="Insecta" family="Bombyliidae" genus="Doliopteryx" kingdom="Animalia" order="Diptera" pageId="4" pageNumber="397" phylum="Arthropoda" rank="species" species="fratris" status="sp. nov.">Doliopteryx fratris</taxonomicName>
|
||
,
|
||
</emphasis>
|
||
<emphasis id="B97FEAD99465FFE4FE26896E6D38F861" bold="true" box="[484,564,1912,1934]" pageId="4" pageNumber="397">
|
||
<taxonomicNameLabel id="A24C57A29465FFE4FE26896E6D38F861" box="[484,564,1912,1934]" pageId="4" pageNumber="397" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
, male holotype:
|
||
<emphasis id="B97FEAD99465FFE4FD1D896E6DE7F861" bold="true" box="[735,747,1912,1934]" pageId="4" pageNumber="397">a</emphasis>
|
||
. dorsal view;
|
||
<emphasis id="B97FEAD99465FFE4FCB9896E6C85F861" bold="true" box="[891,905,1912,1934]" pageId="4" pageNumber="397">b</emphasis>
|
||
. lateral view;
|
||
<emphasis id="B97FEAD99465FFE4FBD8896E6B29F861" bold="true" box="[1050,1061,1912,1934]" pageId="4" pageNumber="397">c</emphasis>
|
||
. antenna (lateral view);
|
||
<emphasis id="B97FEAD99465FFE4FAE0896E6A3CF861" bold="true" box="[1314,1328,1912,1934]" pageId="4" pageNumber="397">d</emphasis>
|
||
. wing
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="8BB436CB9465FFE5FF0589A36A3CFE12" blockId="4.[151,1436,1973,2034]" lastBlockId="5.[151,1437,151,872]" lastPageId="5" lastPageNumber="398" pageId="4" pageNumber="397">
|
||
Thorax. Mesonotum (
|
||
<figureCitation id="13302A4E9465FFE4FE0189A36D1AF821" box="[451,534,1973,1998]" captionStart="FIGURE3" captionStartId="4.[151,250,1912,1934]" captionTargetBox="[450,1137,983,1890]" captionTargetId="figure@4.[450,1137,983,1890]" captionTargetPageId="4" captionText="FIGURE3. Doliopteryx fratris, sp. nov., male holotype: a. dorsal view; b. lateral view; c. antenna(lateral view); d. wing" httpUri="https://zenodo.org/record/1068157/files/figure.png" pageId="4" pageNumber="397">Fig. 3a</figureCitation>
|
||
) yellow with three black stripes, lateral one starting just behind postpronotal lobes and extending to postalar calli, medial one starting from anterior margin of mesonotum and ending much before scutellum leaving yellow prescutellar area; postpronotal lobes yellow; notopleural stripe yellow from humeral callus to post alar calli; scutellum yellow; prosternum brown; lower half of katepisternum brown, upper half yellow; anepisternum yellow except brown mark at anteroventral corner; basal half of anepimeron brown, upper half yellow; katepisternum brown except upper margin narrowly yellow; meron anteriorly yellow, posteriorly brown; scutellum yellow; halter stem and knob yellow. Legs. Coxae brownish; femora yellow at base and apex, brownish medially, in lower margin darker; tibiae yellow; basal three segments of tarsi yellow, apical two segments brownish. Wing (
|
||
<figureCitation id="13302A4E9464FFE5FE238F796D31FE67" box="[481,573,367,392]" captionStart="FIGURE3" captionStartId="4.[151,250,1912,1934]" captionTargetBox="[450,1137,983,1890]" captionTargetId="figure@4.[450,1137,983,1890]" captionTargetPageId="4" captionText="FIGURE3. Doliopteryx fratris, sp. nov., male holotype: a. dorsal view; b. lateral view; c. antenna(lateral view); d. wing" httpUri="https://zenodo.org/record/1068157/files/figure.png" pageId="5" pageNumber="398">Fig. 3d</figureCitation>
|
||
). Hyaline; costa brown, ending at junction with R4+5; Sc very narrow, incomplete, ending much before costa; Rs shorter than r-m crossvein; R2+3 turning up perpendicularly, meeting R1 and forming a small triangular cell; remainder of veins evanescent; M1 straight, basally separated from M2 by a distance longer than r-m crossvein; CuA1 fairly straight to wing margin; CuA2 slightly curved at wing tip.
|
||
</paragraph>
|
||
<paragraph id="8BB436CB9464FFE5FF058C1D6BFEFD7F" blockId="5.[151,1437,151,872]" pageId="5" pageNumber="398">
|
||
Abdomen. Tergites brown except posterior and lateral margins yellow, abdominal sternites completely yellow; hypopygium yellow. Genitalia. Epandrium (
|
||
<figureCitation id="13302A4E9464FFE5FD528C396DEFFDA8" box="[656,739,559,584]" captionStart="FIGURE 4" captionStartId="5.[151,250,1835,1857]" captionTargetBox="[449,1137,916,1823]" captionTargetId="figure@5.[449,1137,916,1823]" captionTargetPageId="5" captionText="FIGURE 4. Doliopteryx fratris, sp. nov., male and female genitalia: a. epandrium; b. gonocoxites and gonostyli; c. genitalia without epandrium (dorsal view); d. female genitalia" httpUri="https://zenodo.org/record/1068159/files/figure.png" pageId="5" pageNumber="398">Fig. 4a</figureCitation>
|
||
) narrowing lateroventrally as two processes; cercus hyaline, triangular; gonocoxites (
|
||
<figureCitation id="13302A4E9464FFE5FE758C456D1CFD83" box="[439,528,595,620]" captionStart="FIGURE 4" captionStartId="5.[151,250,1835,1857]" captionTargetBox="[449,1137,916,1823]" captionTargetId="figure@5.[449,1137,916,1823]" captionTargetPageId="5" captionText="FIGURE 4. Doliopteryx fratris, sp. nov., male and female genitalia: a. epandrium; b. gonocoxites and gonostyli; c. genitalia without epandrium (dorsal view); d. female genitalia" httpUri="https://zenodo.org/record/1068159/files/figure.png" pageId="5" pageNumber="398">Fig. 4b</figureCitation>
|
||
) rectangular; gonostyli triangular, hollow basally; lateral aedeagal apodeme narrow, sinuous, basal aedeagal apodeme (
|
||
<figureCitation id="13302A4E9464FFE5FDB68C616DC8FD60" box="[628,708,631,656]" captionStart="FIGURE 4" captionStartId="5.[151,250,1835,1857]" captionTargetBox="[449,1137,916,1823]" captionTargetId="figure@5.[449,1137,916,1823]" captionTargetPageId="5" captionText="FIGURE 4. Doliopteryx fratris, sp. nov., male and female genitalia: a. epandrium; b. gonocoxites and gonostyli; c. genitalia without epandrium (dorsal view); d. female genitalia" httpUri="https://zenodo.org/record/1068159/files/figure.png" pageId="5" pageNumber="398">Fig. 4c</figureCitation>
|
||
) very large, with two small processes proximally.
|
||
</paragraph>
|
||
<paragraph id="8BB436CB9464FFE5FF058C8D6EEDFD5B" blockId="5.[151,1437,151,872]" box="[199,481,667,692]" pageId="5" pageNumber="398">
|
||
<emphasis id="B97FEAD99464FFE5FF058C8D6E29FD5B" bold="true" box="[199,293,667,692]" pageId="5" pageNumber="398">Female.</emphasis>
|
||
Similar to male.
|
||
</paragraph>
|
||
<paragraph id="8BB436CB9464FFE5FF058CA96E4BFC88" blockId="5.[151,1437,151,872]" pageId="5" pageNumber="398">
|
||
Female genitalia. Furca U-shaped (
|
||
<figureCitation id="13302A4E9464FFE5FD988CA96DBDFD37" box="[602,689,703,728]" captionStart="FIGURE 4" captionStartId="5.[151,250,1835,1857]" captionTargetBox="[449,1137,916,1823]" captionTargetId="figure@5.[449,1137,916,1823]" captionTargetPageId="5" captionText="FIGURE 4. Doliopteryx fratris, sp. nov., male and female genitalia: a. epandrium; b. gonocoxites and gonostyli; c. genitalia without epandrium (dorsal view); d. female genitalia" httpUri="https://zenodo.org/record/1068159/files/figure.png" pageId="5" pageNumber="398">Fig. 4d</figureCitation>
|
||
), indented in lateral arm, extending inwardly as two subquadrate plates; common spermathecal duct long, same as sperm pump in width and structure; two lateral spermathecal pumps shorter than median pump, median pump 1.5 times longer than lateral ones, apical valve of sperm pump present; apical spermathecal duct very short; spermathecal reservoir darkly sclerotized, bulbous, with long beakshaped process.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<caption id="DF7466439464FFE5FF55893D6DA2F8B0" httpUri="https://zenodo.org/record/1068159/files/figure.png" pageId="5" pageNumber="398" targetBox="[449,1137,916,1823]" targetPageId="5">
|
||
<paragraph id="8BB436CB9464FFE5FF55893D6DA2F8B0" blockId="5.[151,1436,1835,1887]" pageId="5" pageNumber="398">
|
||
<emphasis id="B97FEAD99464FFE5FF55893D6E1FF8AE" bold="true" box="[151,275,1835,1857]" pageId="5" pageNumber="398">FIGURE 4.</emphasis>
|
||
<emphasis id="B97FEAD99464FFE5FED9893D6EEDF8AE" box="[283,481,1835,1857]" italics="true" pageId="5" pageNumber="398">
|
||
<taxonomicName id="4C0B4D489464FFE5FED9893D6ED1F8AE" authority="Gharali & Evenhuis, 2017" authorityName="Gharali & Evenhuis" authorityYear="2017" box="[283,477,1835,1857]" class="Insecta" family="Bombyliidae" genus="Doliopteryx" kingdom="Animalia" order="Diptera" pageId="5" pageNumber="398" phylum="Arthropoda" rank="species" species="fratris" status="sp. nov.">Doliopteryx fratris</taxonomicName>
|
||
,
|
||
</emphasis>
|
||
<emphasis id="B97FEAD99464FFE5FE2A893D6D36F8AE" bold="true" box="[488,570,1835,1857]" pageId="5" pageNumber="398">
|
||
<taxonomicNameLabel id="A24C57A29464FFE5FE2A893D6D36F8AE" box="[488,570,1835,1857]" pageId="5" pageNumber="398" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
, male and female genitalia:
|
||
<emphasis id="B97FEAD99464FFE5FC9D893D6C60F8AE" bold="true" box="[863,876,1835,1857]" pageId="5" pageNumber="398">a</emphasis>
|
||
. epandrium;
|
||
<emphasis id="B97FEAD99464FFE5FC37893D6B0FF8AE" bold="true" box="[1013,1027,1835,1857]" pageId="5" pageNumber="398">b</emphasis>
|
||
. gonocoxites and gonostyli;
|
||
<emphasis id="B97FEAD99464FFE5FAEE893D6A31F8AE" bold="true" box="[1324,1341,1835,1857]" pageId="5" pageNumber="398">c.</emphasis>
|
||
genitalia without epandrium (dorsal view);
|
||
<emphasis id="B97FEAD99464FFE5FE33895F6D09F8B0" bold="true" box="[497,517,1865,1887]" pageId="5" pageNumber="398">d.</emphasis>
|
||
female genitalia
|
||
</paragraph>
|
||
</caption>
|
||
<subSubSection id="C31165409464FFE5FF05899A6A60F802" pageId="5" pageNumber="398" type="etymology">
|
||
<paragraph id="8BB436CB9464FFE5FF05899A6A60F802" blockId="5.[151,1436,1932,2029]" pageId="5" pageNumber="398">
|
||
<emphasis id="B97FEAD99464FFE5FF05899A6E42F84A" bold="true" box="[199,334,1932,1957]" pageId="5" pageNumber="398">Etymology.</emphasis>
|
||
The name of species,
|
||
<taxonomicName id="4C0B4D489464FFE5FD88899A6D9CF84B" authorityName="Gharali & Evenhuis" authorityYear="2017" box="[586,656,1932,1956]" class="Insecta" family="Bombyliidae" genus="Doliopteryx" kingdom="Animalia" order="Diptera" pageId="5" pageNumber="398" phylum="Arthropoda" rank="species" species="fratris">
|
||
<emphasis id="B97FEAD99464FFE5FD88899A6D9CF84B" box="[586,656,1932,1956]" italics="true" pageId="5" pageNumber="398">fratris</emphasis>
|
||
</taxonomicName>
|
||
, is Latin meaning “brother” and used in honor of Hamid Gharali, the brother of the senior author, whose curiosity led to finding the plant (
|
||
<taxonomicName id="4C0B4D489464FFE5FC7989A66BA1F827" box="[955,1197,1968,1992]" class="Magnoliopsida" family="Polygonaceae" genus="Polygonum" kingdom="Plantae" order="Polygonales" pageId="5" pageNumber="398" phylum="Tracheophyta" rank="species" species="aviculare">
|
||
<emphasis id="B97FEAD99464FFE5FC7989A66BA1F827" box="[955,1197,1968,1992]" italics="true" pageId="5" pageNumber="398">Polygonum aviculare</emphasis>
|
||
</taxonomicName>
|
||
L.) that seems to be adult food source of this new species. After placing pan traps close to this plant we collected many specimens.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C31165409467FFE6FF058E816BDCFF17" pageId="6" pageNumber="399" type="distribution">
|
||
<paragraph id="8BB436CB9467FFE6FF058E816BDCFF17" blockId="6.[151,1436,151,464]" pageId="6" pageNumber="399">
|
||
<emphasis id="B97FEAD99467FFE6FF058E816E6CFF5F" bold="true" box="[199,352,151,176]" pageId="6" pageNumber="399">Distribution.</emphasis>
|
||
This species is currently known only from Nodahak village (
|
||
<collectingRegion id="49CFF8299467FFE6FBCC8E816B6DFF5F" box="[1038,1121,151,176]" country="Iran" name="Qazvin" pageId="6" pageNumber="399">Qazvin</collectingRegion>
|
||
city) in northern
|
||
<collectingCountry id="F31C765B9467FFE6FAE68E8E6A59FF40" box="[1316,1365,152,175]" name="Iran" pageId="6" pageNumber="399">Iran</collectingCountry>
|
||
.
|
||
<emphasis id="B97FEAD99467FFE6FF058EAA6E41FF3A" bold="true" box="[199,333,188,213]" pageId="6" pageNumber="399">Variability.</emphasis>
|
||
Sometimes the black pattern on the mesonotum is coalesced into a large black mark instead of three separated stripes. The color of the prescutellar area varies from yellow to brownish yellow.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C31165409467FFE6FF058F126C75FE3F" pageId="6" pageNumber="399" type="discussion">
|
||
<paragraph id="8BB436CB9467FFE6FF058F126C75FE3F" blockId="6.[151,1436,151,464]" pageId="6" pageNumber="399">
|
||
<emphasis id="B97FEAD99467FFE6FF058F126E37FEF2" bold="true" box="[199,315,260,285]" pageId="6" pageNumber="399">Remarks.</emphasis>
|
||
Specimens were collected from an area (
|
||
<figureCitation id="13302A4E9467FFE6FCC58F126C44FEF2" box="[775,840,260,285]" captionStart="FIGURE 5" captionStartId="6.[151,249,1979,2001]" captionTargetBox="[213,1374,548,1967]" captionTargetId="figure@6.[213,1374,507,1967]" captionTargetPageId="6" captionText="FIGURE 5. Type locality:a. typelocality;b. Polygonum aviculare L.; c. a part of Polygonum aviculare L. with tiny flowers d. a white pan trap" httpUri="https://zenodo.org/record/1068161/files/figure.png" pageId="6" pageNumber="399">Fig. 5</figureCitation>
|
||
) densely populated by different species of composite plants such as
|
||
<taxonomicName id="4C0B4D489467FFE6FEFA8F3E6D93FEAF" authority="M. Bieb." authorityName="M.Bieb." box="[312,671,295,320]" class="Magnoliopsida" family="Asteraceae" genus="Carthamus" kingdom="Plantae" order="Asterales" pageId="6" pageNumber="399" phylum="Tracheophyta" rank="species" species="oxyacantha">
|
||
<emphasis id="B97FEAD99467FFE6FEFA8F3E6D30FEAF" box="[312,572,296,320]" italics="true" pageId="6" pageNumber="399">Carthamus oxyacantha</emphasis>
|
||
M.Bieb.
|
||
</taxonomicName>
|
||
but we think these tiny flies are likely pollen feeders on tiny flowers of
|
||
<taxonomicName id="4C0B4D489467FFE6FF748F5A6EAAFE8B" box="[182,422,332,356]" class="Magnoliopsida" family="Polygonaceae" genus="Polygonum" kingdom="Plantae" order="Polygonales" pageId="6" pageNumber="399" phylum="Tracheophyta" rank="species" species="aviculare">
|
||
<emphasis id="B97FEAD99467FFE6FF748F5A6EAAFE8B" box="[182,422,332,356]" italics="true" pageId="6" pageNumber="399">Polygonum aviculare</emphasis>
|
||
</taxonomicName>
|
||
L. because when we put our pans (about 100 pan traps) close to this plant, the number of adults increased noticeably and we were able to collect hundreds of specimens. It is worthy to note that many specimens of
|
||
<taxonomicName id="4C0B4D489467FFE6FEF08F826CF3FE42" authority="Winterton & Gharali, 2011" authorityName="Winterton & Gharali" authorityYear="2011" box="[306,1023,404,429]" class="Insecta" family="Scenopinidae" genus="Iranotrichia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Diptera" pageId="6" pageNumber="399" phylum="Arthropoda" rank="species" species="insolita">
|
||
<emphasis id="B97FEAD99467FFE6FEF08F826D1AFE43" box="[306,534,404,428]" italics="true" pageId="6" pageNumber="399">Iranotrichia insolita</emphasis>
|
||
<bibRefCitation id="EF9A4B3A9467FFE6FDDC8F826C42FE43" author="Winterton" box="[542,846,404,429]" pageId="6" pageNumber="399" refString="Winterton, S. L. & Gharali, B. (2011) Iranotrichia gen. n., a new genus of Scenopinidae (Diptera) from Iran, with a key to window fly genera of the world. ZooKeys, 138, 75 - 92." type="journal article" year="2011">Winterton & Gharali, 2011</bibRefCitation>
|
||
(Scenopinidae)
|
||
</taxonomicName>
|
||
and an undetermined species of gall midge (
|
||
<taxonomicName id="4C0B4D489467FFE6FF298FA16E94FE3F" box="[235,408,439,464]" class="Insecta" family="Cecidomyiidae" kingdom="Animalia" order="Diptera" pageId="6" pageNumber="399" phylum="Arthropoda" rank="family">Cecidomyiidae</taxonomicName>
|
||
) were collected along with these tiny flies.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |