174 lines
19 KiB
XML
174 lines
19 KiB
XML
<document id="5C6AB7375167E50D45E126A4F2A72B0E" ID-DOI="10.5281/zenodo.162803" ID-GBIF-Dataset="f514e964-2196-461c-84ff-64243643bc81" ID-ISSN="11755326" ID-Zenodo-Dep="162803" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1459445184067" checkinUser="plazi" docAuthor="Erwin, Terry L." docDate="2002" docId="03A287CA782FFFB4FEECF9D29927FBA9" docLanguage="en" docName="zt00119.pdf" docOrigin="Zootaxa 119" docStyle="DocumentStyle:FA7E419B012A62B0FC3AC15A186C3DAF.3:Zootaxa.2001-2006.journal_article" docStyleId="FA7E419B012A62B0FC3AC15A186C3DAF" docStyleName="Zootaxa.2001-2006.journal_article" docStyleVersion="3" docTitle="Agra julie Erwin, new species" docType="treatment" docVersion="11" lastPageNumber="60" masterDocId="FF9BFFB27815FF8FFFE4FF829C34FFFF" masterDocTitle="The Beetle Family Carabidae of Costa Rica: Twentynine new species of Agra Fabricius 1801 (Coleoptera: Carabidae, Lebiini, Agrina)" masterLastPageNumber="68" masterPageNumber="1" pageNumber="59" updateTime="1698190273211" updateUser="plazi">
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<mods:mods id="3DBB22C2304192A257218E2D974C917C" xmlns:mods="http://www.loc.gov/mods/v3">
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<mods:titleInfo id="B37E677AFA96D94DD6B3E35B0F61C8A3">
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<mods:title id="5C665404A30199EA77D5ABDDBEDDD03D">The Beetle Family Carabidae of Costa Rica: Twentynine new species of Agra Fabricius 1801 (Coleoptera: Carabidae, Lebiini, Agrina)</mods:title>
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</mods:titleInfo>
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<mods:name id="526355185EF389969EC5EADE0783D46E" type="personal">
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<mods:role id="312D421E2B32EDFD95348D548C174C25">
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<mods:roleTerm id="F9712EE85A1595333C7DB628078D9644">Author</mods:roleTerm>
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</mods:role>
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<mods:namePart id="E04B431995B0F8F86CEA21F25AE9E173">Erwin, Terry L.</mods:namePart>
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</mods:name>
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<mods:typeOfResource id="B3CF58121B4DD78F1E705E3843DBE8EA">text</mods:typeOfResource>
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<mods:title id="D3430E95F89D2DBD33C58A9DD772D88E">Zootaxa</mods:title>
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</mods:titleInfo>
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<mods:part id="7C9ED54E8C4C262B4F81C5D178EDB726">
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<mods:date id="76538FE0A00A340FFD75038E22F2AFEB">2002</mods:date>
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<mods:detail id="A83CB8E07D08B5639EAB2380F8A4A051" type="volume">
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<mods:number id="70156D8AB67D21D881B6930B0B1C4554">119</mods:number>
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<mods:end id="E093AC1403A2CD49804E8F9104E386DA">68</mods:end>
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<mods:classification id="F7740007BC34C5A82F69B3D0784ED732">journal article</mods:classification>
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<mods:identifier id="7A3938B1F6C09408F23C09C3A51071F3" type="DOI">10.5281/zenodo.162803</mods:identifier>
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<mods:identifier id="2289278F3FC26417D6C9516655DB9914" type="GBIF-Dataset">f514e964-2196-461c-84ff-64243643bc81</mods:identifier>
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<mods:identifier id="C6DB3BF169EB2EF9A4D645AE62D6B3B7" type="ISSN">11755326</mods:identifier>
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<mods:identifier id="D3D26EED313C517080623BE0623C4E7D" type="Zenodo-Dep">162803</mods:identifier>
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<treatment id="03A287CA782FFFB4FEECF9D29927FBA9" ID-DOI="http://doi.org/10.5281/zenodo.6278030" ID-GBIF-Taxon="121065203" ID-Zenodo-Dep="6278030" LSID="urn:lsid:plazi:treatment:03A287CA782FFFB4FEECF9D29927FBA9" httpUri="http://treatment.plazi.org/id/03A287CA782FFFB4FEECF9D29927FBA9" lastPageId="59" lastPageNumber="60" pageId="58" pageNumber="59">
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<subSubSection id="C3116557782FFFB5FEECF9D29DCFF972" pageId="58" pageNumber="59" type="nomenclature">
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<paragraph id="8BB436DC782FFFB5FEECF9D29E42F995" blockId="58.[264,630,1616,1677]" box="[264,630,1616,1642]" pageId="58" pageNumber="59">
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<heading id="D0FC81B0782FFFB5FEECF9D29E42F995" bold="true" box="[264,630,1616,1642]" fontSize="11" level="1" pageId="58" pageNumber="59" reason="1">
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<emphasis id="B97FEACE782FFFB5FEECF9D29E42F995" bold="true" box="[264,630,1616,1642]" pageId="58" pageNumber="59">
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<taxonomicName id="4C0B4D5F782FFFB5FEECF9D29DEDF995" ID-CoL="B2PR" authority="Erwin" authorityName="Erwin" box="[264,473,1616,1642]" class="Insecta" family="Carabidae" genus="Agra" kingdom="Animalia" order="Coleoptera" pageId="58" pageNumber="59" phylum="Arthropoda" rank="species" species="julie" status="sp. nov.">
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<emphasis id="B97FEACE782FFFB5FEECF9D29DB6F995" bold="true" box="[264,386,1616,1642]" italics="true" pageId="58" pageNumber="59">Agra julie</emphasis>
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Erwin
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</taxonomicName>
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,
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<taxonomicNameLabel id="A24C57B5782FFFB5FE02F9D29E42F995" box="[486,630,1616,1642]" pageId="58" pageNumber="59" rank="species">new species</taxonomicNameLabel>
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</emphasis>
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</heading>
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||
</paragraph>
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<paragraph id="8BB436DC782FFFB5FEECF9F19DCFF972" blockId="58.[264,630,1616,1677]" box="[264,507,1651,1677]" pageId="58" pageNumber="59">
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(
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<figureCitation id="13302A59782FFFB5FEF5F9F19D9FF972" box="[273,427,1651,1677]" captionStart="FIGURE 27. A" captionStartId="59.[264,368,1522,1546]" captionTargetBox="[408,1164,1195,1492]" captionTargetId="figure@59.[407,1179,1169,1505]" captionTargetPageId="59" captionText="FIGURE 27. A. julie, female, dorsal aspect; a) head; b) apex of elytron; c) Sternum VI, male (top), female (bottom), ventral aspect. Measures (mm): HL = 1.8; WE = 1.5; male SW = 2.7, female SW = 3.1." httpUri="https://zenodo.org/record/162841/files/figure.png" pageId="58" pageNumber="59">Figs. 27a, 27</figureCitation>
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b, 27c)
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</paragraph>
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||
</subSubSection>
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<subSubSection id="C3116557782FFFB5FEECF940991EF922" box="[264,1322,1730,1757]" pageId="58" pageNumber="59" type="materials_examined">
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<paragraph id="8BB436DC782FFFB5FEECF940991EF922" blockId="58.[264,1324,1730,1877]" box="[264,1322,1730,1757]" pageId="58" pageNumber="59">
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<emphasis id="B97FEACE782FFFB5FEECF9409D49F923" bold="true" box="[264,381,1730,1756]" pageId="58" pageNumber="59">
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<typeStatus id="54B0887E782FFFB5FEECF9409D4CF923" box="[264,376,1730,1756]" pageId="58" pageNumber="59" type="holotype">Holotype</typeStatus>
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.
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</emphasis>
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Female,
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<emphasis id="B97FEACE782FFFB5FE31F9409EB7F923" bold="true" box="[469,643,1730,1756]" pageId="58" pageNumber="59">
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<collectingCountry id="F31C764C782FFFB5FE31F9409E4AF923" box="[469,638,1730,1756]" name="Costa Rica" pageId="58" pageNumber="59">COSTARICA</collectingCountry>
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:
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</emphasis>
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Cartago,
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<taxonomicName id="4C0B4D5F782FFFB5FD06F9419F78F922" box="[738,844,1731,1757]" class="Insecta" family="Carabidae" genus="Agra" kingdom="Animalia" order="Coleoptera" pageId="58" pageNumber="25" phylum="Arthropoda" rank="species" species="turrialba">Turrialba</taxonomicName>
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,
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<quantity id="4CF39B39782FFFB5FCB4F9419FA5F922" box="[848,913,1731,1757]" metricMagnitude="2" metricUnit="m" metricValue="6.0" pageId="58" pageNumber="59" unit="m" value="600.0">600m</quantity>
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||
,
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||
<geoCoordinate id="EE3F501B782FFFB5FC72F9419FDEF922" box="[918,1002,1731,1757]" direction="north" orientation="latitude" pageId="58" pageNumber="59" precision="925" value="9.883333">09º53'N</geoCoordinate>
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<geoCoordinate id="EE3F501B782FFFB5FC0EF941987EF922" box="[1002,1098,1731,1757]" direction="west" orientation="longitude" pageId="58" pageNumber="59" precision="925" value="-83.63333">83º38'W</geoCoordinate>
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,(
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<collectionCode id="ED1AAE19782FFFB5FBB2F94198A4F922" LSID="urn:lsid:biocol.org:col:34700" box="[1110,1168,1731,1757]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:15690" name="California Academy of Sciences" pageId="58" pageNumber="59">CAS</collectionCode>
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:ADP10007).
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</paragraph>
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</subSubSection>
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<subSubSection id="C3116557782FFFB5FEDCF9689F21F8AA" pageId="58" pageNumber="59" type="diagnosis">
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<paragraph id="8BB436DC782FFFB5FEDCF9689F21F8AA" blockId="58.[264,1324,1730,1877]" pageId="58" pageNumber="59">
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<emphasis id="B97FEACE782FFFB5FEDCF9689D84F8FB" bold="true" box="[312,432,1770,1796]" pageId="58" pageNumber="59">Diagnosis</emphasis>
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. This is the only species of
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<taxonomicName id="4C0B4D5F782FFFB5FD1BF9699F0EF8FB" box="[767,826,1771,1796]" class="Insecta" family="Carabidae" genus="Agra" kingdom="Animalia" order="Coleoptera" pageId="58" pageNumber="59" phylum="Arthropoda" rank="genus">
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<emphasis id="B97FEACE782FFFB5FD1BF9699F0EF8FB" box="[767,826,1771,1796]" italics="true" pageId="58" pageNumber="59">Agra</emphasis>
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</taxonomicName>
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in
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<collectingCountry id="F31C764C782FFFB5FC85F9699FD7F8FA" box="[865,995,1771,1797]" name="Costa Rica" pageId="58" pageNumber="59">Costa Rica</collectingCountry>
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with brassy nearly truncate elytra and moderately long pubescence on the venter of the prothorax. Head quadrate behind with a marked dimple on the occiput.
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</paragraph>
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</subSubSection>
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<subSubSection id="C3116557782EFFB4FEDCFE99986DFDB1" pageId="59" pageNumber="60" type="description">
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<paragraph id="8BB436DC782EFFB4FEDCFE99986DFDB1" blockId="59.[264,1326,283,1110]" pageId="59" pageNumber="60">
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<emphasis id="B97FEACE782EFFB4FEDCFE999DFDFECA" bold="true" box="[312,457,283,309]" pageId="59" pageNumber="60">Description</emphasis>
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. Color and luster: Rufous. Elytra brassy green. Appendages rufotestaceous. Form: Head (
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<figureCitation id="13302A59782EFFB4FDE6FEC69E53FEA1" box="[514,615,324,350]" captionStart="FIGURE 27. A" captionStartId="59.[264,368,1522,1546]" captionTargetBox="[408,1164,1195,1492]" captionTargetId="figure@59.[407,1179,1169,1505]" captionTargetPageId="59" captionText="FIGURE 27. A. julie, female, dorsal aspect; a) head; b) apex of elytron; c) Sternum VI, male (top), female (bottom), ventral aspect. Measures (mm): HL = 1.8; WE = 1.5; male SW = 2.7, female SW = 3.1." httpUri="https://zenodo.org/record/162841/files/figure.png" pageId="59" pageNumber="60">Fig. 27a</figureCitation>
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) behind eyes short, hind angles quadrate; vertex dimpled. Pronotum and prosternum finely punctate, prothorax broad, widest at middle. Elytron with apex (
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<figureCitation id="13302A59782EFFB4FEB6FE169D9EFE51" box="[338,426,404,430]" captionStart="FIGURE 27. A" captionStartId="59.[264,368,1522,1546]" captionTargetBox="[408,1164,1195,1492]" captionTargetId="figure@59.[407,1179,1169,1505]" captionTargetPageId="59" captionText="FIGURE 27. A. julie, female, dorsal aspect; a) head; b) apex of elytron; c) Sternum VI, male (top), female (bottom), ventral aspect. Measures (mm): HL = 1.8; WE = 1.5; male SW = 2.7, female SW = 3.1." httpUri="https://zenodo.org/record/162841/files/figure.png" pageId="59" pageNumber="60">Fig. 27</figureCitation>
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||
b) slightly oblique, lateral tooth small, sutural apex rounded,
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<taxonomicName id="4C0B4D5F782EFFB4FB6AFE169887FE51" box="[1166,1203,404,430]" class="Insecta" family="Carabidae" genus="Agra" kingdom="Animalia" order="Coleoptera" pageId="59" pageNumber="60" phylum="Arthropoda" rank="species" species="not">not</taxonomicName>
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projected acutely, apical margin very slightly lobed. Metasternum of male densely pubescent; trochanter and abdominal sterna each with long scattered setae. Apex of sternum VI (
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<figureCitation id="13302A59782EFFB4FB1FFE669D13FDD9" captionStart="FIGURE 27. A" captionStartId="59.[264,368,1522,1546]" captionTargetBox="[408,1164,1195,1492]" captionTargetId="figure@59.[407,1179,1169,1505]" captionTargetPageId="59" captionText="FIGURE 27. A. julie, female, dorsal aspect; a) head; b) apex of elytron; c) Sternum VI, male (top), female (bottom), ventral aspect. Measures (mm): HL = 1.8; WE = 1.5; male SW = 2.7, female SW = 3.1." httpUri="https://zenodo.org/record/162841/files/figure.png" pageId="59" pageNumber="60">Fig. 27</figureCitation>
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c) angulate laterally in male, in female corner obtuse. Male legs unmodified, similar to females. Size: small, 9.0 to 11.0mm in length,
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||
<quantity id="4CF39B39782EFFB4FCCDFDB69FD7FDB1" box="[809,995,564,590]" metricMagnitude="-3" metricUnit="m" metricValue="3.1799999999999997" metricValueMax="3.38" metricValueMin="2.98" pageId="59" pageNumber="60" unit="mm" value="3.1799999999999997" valueMax="3.38" valueMin="2.98">2.98 to 3.38mm</quantity>
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in width..
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</paragraph>
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||
</subSubSection>
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||
<subSubSection id="C3116557782EFFB4FEA7FDD99F2EFBF9" pageId="59" pageNumber="60" type="materials_examined">
|
||
<paragraph id="8BB436DC782EFFB4FEA7FDD99E7EFC99" blockId="59.[264,1326,283,1110]" pageId="59" pageNumber="60">
|
||
<emphasis id="B97FEACE782EFFB4FEA7FDD99F0DFD8A" bold="true" box="[323,825,603,629]" pageId="59" pageNumber="60">
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||
Other specimens examined.
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||
<collectingCountry id="F31C764C782EFFB4FD49FDD99F0DFD8A" box="[685,825,603,629]" name="Costa Rica" pageId="59" pageNumber="60">Costa Rica</collectingCountry>
|
||
</emphasis>
|
||
: 4
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||
<typeStatus id="54B0887E782EFFB4FC81FDDE9FECFD89" box="[869,984,604,630]" pageId="59" pageNumber="60" type="paratype">paratypes</typeStatus>
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||
,
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<specimenCount id="9D0DFD55782EFFB4FC0CFDDE9854FD89" box="[1000,1120,604,630]" pageId="59" pageNumber="60" type="female">3 females</specimenCount>
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||
, 1 (very teneral) male, from the following Conservation Areas: Cordillera Volcánica Central, Osa, Pacifico Central.
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<collectingCountry id="F31C764C782EFFB4FE98FD2E9E39FD39" box="[380,525,684,710]" name="Costa Rica" pageId="59" pageNumber="60">Costa Rica</collectingCountry>
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||
, Cartago, 1f,
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||
<taxonomicName id="4C0B4D5F782EFFB4FD32FD2E9F71FD39" box="[726,837,684,710]" class="Insecta" family="Carabidae" genus="Agra" kingdom="Animalia" order="Coleoptera" pageId="59" pageNumber="25" phylum="Arthropoda" rank="species" species="turrialba">Turrialba</taxonomicName>
|
||
,
|
||
<geoCoordinate id="EE3F501B782EFFB4FC84FD2E9FF3FD39" box="[864,967,684,710]" direction="west" orientation="longitude" pageId="59" pageNumber="60" precision="925" value="-83.63333">83°38'W</geoCoordinate>
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<geoCoordinate id="EE3F501B782EFFB4FC39FD2E9806FD39" box="[989,1074,684,710]" direction="north" orientation="latitude" pageId="59" pageNumber="60" precision="925" value="9.883333">9°53'N</geoCoordinate>
|
||
, (O. Bryant)(
|
||
<collectionCode id="ED1AAE19782EFFB4FB0AFD2E991DFD39" LSID="urn:lsid:biocol.org:col:34700" box="[1262,1321,684,710]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:15690" name="California Academy of Sciences" pageId="59" pageNumber="60">CAS</collectionCode>
|
||
: ADP10006),
|
||
<quantity id="4CF39B39782EFFB4FE42FD569DFAFD11" box="[422,462,724,750]" metricMagnitude="0" metricUnit="m" metricValue="1.0" pageId="59" pageNumber="60" unit="m" value="1.0">1m</quantity>
|
||
, January, (V.M. Kirk)(
|
||
<collectionCode id="ED1AAE19782EFFB4FD38FD569F01FD11" LSID="urn:lsid:biocol.org:col:34404" box="[732,821,724,750]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34404" name="University of Alberta, E.H. Strickland Entomological Museum" pageId="59" pageNumber="60">UASM</collectionCode>
|
||
: ADP21784); Puntarenas, 1f, P.N. Manuel Antonio, Quepos,
|
||
<quantity id="4CF39B39782EFFB4FE3AFD7E9E21FCE9" box="[478,533,764,790]" metricMagnitude="1" metricUnit="m" metricValue="8.0" pageId="59" pageNumber="60" unit="m" value="80.0">80m</quantity>
|
||
,
|
||
<geoCoordinate id="EE3F501B782EFFB4FDC5FD7E9EBDFCE9" box="[545,649,764,790]" direction="west" orientation="longitude" pageId="59" pageNumber="60" precision="925" value="-84.15">84°09'W</geoCoordinate>
|
||
<geoCoordinate id="EE3F501B782EFFB4FD6BFD7E9ED0FCE9" box="[655,740,764,790]" direction="north" orientation="latitude" pageId="59" pageNumber="60" precision="925" value="9.4">9°24'N</geoCoordinate>
|
||
, LS 370900, 448800, August, (G. Valera)(
|
||
<taxonomicName id="4C0B4D5F782EFFB4FB32FD7E991EFCE9" box="[1238,1322,764,790]" class="Insecta" family="Carabidae" genus="Agra" kingdom="Animalia" order="Coleoptera" pageId="59" pageNumber="67" phylum="Arthropoda" rank="species" species="inbio">
|
||
<collectionCode id="ED1AAE19782EFFB4FB32FD7E991EFCE9" box="[1238,1322,764,790]" pageId="59" pageNumber="60">INBIO</collectionCode>
|
||
</taxonomicName>
|
||
:
|
||
<collectionCode id="ED1AAE19782EFFB4FEECFCA69D0FFCC1" LSID="urn:lsid:biocol.org:col:14323" box="[264,315,804,830]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:14323" name="Universidade do Extremo Sul Catarinense, Bairro Universitário" pageId="59" pageNumber="60">CRI</collectionCode>
|
||
001318173), 1f, Piedras Blancas, 24.0 km W,
|
||
<quantity id="4CF39B39782EFFB4FC47FCA69FDDFCC1" box="[931,1001,804,830]" metricMagnitude="2" metricUnit="m" metricValue="2.0" pageId="59" pageNumber="60" unit="m" value="200.0">200m</quantity>
|
||
, March May, (P. Hanson)(
|
||
<collectionCode id="ED1AAE19782EFFB4FEA6FCCE9DAEFC99" box="[322,410,844,870]" pageId="59" pageNumber="60">UCOR</collectionCode>
|
||
: ADP
|
||
<date id="FFB5101C782EFFB4FE00FCCE9E0AFC99" box="[484,574,844,870]" pageId="59" pageNumber="60" value="33-10-05">100533</date>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="8BB436DC782EFFB4FEDCFCF19F2EFBF9" blockId="59.[264,1326,283,1110]" pageId="59" pageNumber="60">
|
||
<emphasis id="B97FEACE782EFFB4FEDCFCF19DCBFC72" bold="true" box="[312,511,883,909]" pageId="59" pageNumber="60">Specific epithet.</emphasis>
|
||
The specific epithet,
|
||
<emphasis id="B97FEACE782EFFB4FD18FCF69F03FC72" box="[764,823,884,909]" italics="true" pageId="59" pageNumber="60">
|
||
<taxonomicName id="4C0B4D5F782EFFB4FD18FCF69F07FC72" box="[764,819,884,909]" class="Insecta" family="Carabidae" genus="Agra" kingdom="Animalia" order="Coleoptera" pageId="59" pageNumber="60" phylum="Arthropoda" rank="species" species="julie">julie</taxonomicName>
|
||
,
|
||
</emphasis>
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is a used as a noun in apposition honoring
|
||
<taxonomicName id="4C0B4D5F782EFFB4FEECFC1E9D0BFC49" box="[264,319,924,950]" class="Insecta" family="Carabidae" genus="Agra" kingdom="Animalia" order="Coleoptera" pageId="59" pageNumber="60" phylum="Arthropoda" rank="species" species="julie">Julie</taxonomicName>
|
||
Parenas, past Entomology Department secretary at
|
||
<collectionCode id="ED1AAE19782EFFB4FC57FC1E9FD8FC49" LSID="urn:lsid:biocol.org:col:34700" box="[947,1004,924,950]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:15690" name="California Academy of Sciences" pageId="59" pageNumber="60">CAS</collectionCode>
|
||
(Depository for the
|
||
<typeStatus id="54B0887E782EFFB4FB03FC1E9D0EFC21" pageId="59" pageNumber="60" type="holotype">Holotype</typeStatus>
|
||
of this species), who has contributed so much to the welfare of that department and its curators and staff beyond the job description.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C3116557782EFFB4FEDCFB919927FBA9" pageId="59" pageNumber="60" type="description">
|
||
<paragraph id="8BB436DC782EFFB4FEDCFB919927FBA9" blockId="59.[264,1326,283,1110]" pageId="59" pageNumber="60">
|
||
<emphasis id="B97FEACE782EFFB4FEDCFB919DB1FBD2" bold="true" box="[312,389,1043,1069]" pageId="59" pageNumber="60">Notes.</emphasis>
|
||
An illustration of rufiventrisgroup stylomere 2 is on the
|
||
<taxonomicName id="4C0B4D5F782EFFB4FBD0FB9698C5FBD2" box="[1076,1265,1044,1069]" class="Insecta" family="Carabidae" genus="Agra" kingdom="Animalia" order="Coleoptera" pageId="59" pageNumber="60" phylum="Arthropoda" rank="species" species="rufiventris">
|
||
<emphasis id="B97FEACE782EFFB4FBD0FB9698C5FBD2" box="[1076,1265,1044,1069]" italics="true" pageId="59" pageNumber="60">Agra rufiventris</emphasis>
|
||
</taxonomicName>
|
||
web page. The available male is so teneral that dissection of the aedeagus proved impossible.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |