139 lines
14 KiB
XML
139 lines
14 KiB
XML
<document id="23257D4A277C4016E87301760C0113CD" ID-DOI="10.5281/zenodo.162803" ID-GBIF-Dataset="f514e964-2196-461c-84ff-64243643bc81" ID-ISSN="11755326" ID-Zenodo-Dep="162803" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1459445184067" checkinUser="plazi" docAuthor="Erwin, Terry L." docDate="2002" docId="03A287CA7829FFB3FEECFE239F00F9F7" docLanguage="en" docName="zt00119.pdf" docOrigin="Zootaxa 119" docStyle="DocumentStyle:FA7E419B012A62B0FC3AC15A186C3DAF.3:Zootaxa.2001-2006.journal_article" docStyleId="FA7E419B012A62B0FC3AC15A186C3DAF" docStyleName="Zootaxa.2001-2006.journal_article" docStyleVersion="3" docTitle="Agra ichabod Erwin, new species" docType="treatment" docVersion="12" lastPageNumber="61" masterDocId="FF9BFFB27815FF8FFFE4FF829C34FFFF" masterDocTitle="The Beetle Family Carabidae of Costa Rica: Twentynine new species of Agra Fabricius 1801 (Coleoptera: Carabidae, Lebiini, Agrina)" masterLastPageNumber="68" masterPageNumber="1" pageNumber="61" updateTime="1698190273211" updateUser="plazi">
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<mods:mods id="4AE220EEAFCFBA857552E5AE112E76C4" xmlns:mods="http://www.loc.gov/mods/v3">
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<mods:titleInfo id="CD2405DD3C5687765AB0BC9C1980D99C">
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<mods:title id="3E6030D35C2E0984138D59AF6EBF4B69">The Beetle Family Carabidae of Costa Rica: Twentynine new species of Agra Fabricius 1801 (Coleoptera: Carabidae, Lebiini, Agrina)</mods:title>
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</mods:titleInfo>
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<mods:name id="690377116DB49A64D1C64410E854A861" type="personal">
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<mods:roleTerm id="646433DFA1D87925FC6C15D02B46E429">Author</mods:roleTerm>
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</mods:role>
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<mods:namePart id="92346611F542DE88CFD83903B5035CEA">Erwin, Terry L.</mods:namePart>
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<mods:typeOfResource id="FE4E1FD51ACDA2C80F235F736E269C73">text</mods:typeOfResource>
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<mods:title id="A69C683AC94C346498BDBC9FE9641867">Zootaxa</mods:title>
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</mods:titleInfo>
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<mods:part id="F4151EB38935EF8FFDB75B5BCC741583">
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<mods:date id="8C7DDEE4C9631630657A40F3425D1B85">2002</mods:date>
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<mods:detail id="333DD4AA350263EBD439C6FAD579211B" type="volume">
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<mods:number id="408B418E762DE0F2B6599F89940316D5">119</mods:number>
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</mods:detail>
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<mods:start id="35CF3088C1084763BD48F04EA8076B50">1</mods:start>
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<mods:end id="00D056491C58A71F1B736F475F2FDB7B">68</mods:end>
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<mods:classification id="5CA3A0FBB28F62BBCDB6240200695325">journal article</mods:classification>
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<mods:identifier id="C79A9416938D229FAD3A4C628050EF78" type="DOI">10.5281/zenodo.162803</mods:identifier>
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<mods:identifier id="6D3143C6ECBD959A1E2B307556843023" type="GBIF-Dataset">f514e964-2196-461c-84ff-64243643bc81</mods:identifier>
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<mods:identifier id="2D36FC06131C2812526B5CF95B86CEFD" type="ISSN">11755326</mods:identifier>
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<mods:identifier id="0F49A1263C863B58687FF9BC2CCA88C8" type="Zenodo-Dep">162803</mods:identifier>
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<treatment id="03A287CA7829FFB3FEECFE239F00F9F7" ID-DOI="http://doi.org/10.5281/zenodo.6278032" ID-GBIF-Taxon="119330631" ID-Zenodo-Dep="6278032" LSID="urn:lsid:plazi:treatment:03A287CA7829FFB3FEECFE239F00F9F7" httpUri="http://treatment.plazi.org/id/03A287CA7829FFB3FEECFE239F00F9F7" lastPageNumber="61" pageId="60" pageNumber="61">
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<subSubSection id="C31165577829FFB3FEECFE239DF5FE1C" pageId="60" pageNumber="61" type="nomenclature">
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<paragraph id="8BB436DC7829FFB3FEECFE239E94FE44" blockId="60.[264,672,417,483]" box="[264,672,417,443]" pageId="60" pageNumber="61">
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<heading id="D0FC81B07829FFB3FEECFE239E94FE44" bold="true" box="[264,672,417,443]" fontSize="11" level="1" pageId="60" pageNumber="61" reason="1">
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<emphasis id="B97FEACE7829FFB3FEECFE239E94FE44" bold="true" box="[264,672,417,443]" pageId="60" pageNumber="61">
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<taxonomicName id="4C0B4D5F7829FFB3FEECFE239E37FE44" ID-CoL="B2P2" authority="Erwin" authorityName="Erwin" box="[264,515,417,443]" class="Insecta" family="Carabidae" genus="Agra" kingdom="Animalia" order="Coleoptera" pageId="60" pageNumber="61" phylum="Arthropoda" rank="species" species="ichabod" status="sp. nov.">
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<emphasis id="B97FEACE7829FFB3FEECFE239D98FE44" bold="true" box="[264,428,417,443]" italics="true" pageId="60" pageNumber="61">Agra ichabod</emphasis>
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Erwin
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</taxonomicName>
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,
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<taxonomicNameLabel id="A24C57B57829FFB3FDF4FE239E94FE44" box="[528,672,417,443]" pageId="60" pageNumber="61" rank="species">new species</taxonomicNameLabel>
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</emphasis>
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</heading>
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</paragraph>
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<paragraph id="8BB436DC7829FFB3FEECFE4B9DF5FE1C" blockId="60.[264,672,417,483]" box="[264,449,457,483]" pageId="60" pageNumber="61">
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(
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<figureCitation id="13302A597829FFB3FEF5FE4B9D9EFE1C" box="[273,426,457,483]" captionStart="FIGURE 28. A" captionStartId="60.[264,367,1839,1863]" captionTargetBox="[558,1021,1610,1823]" captionTargetId="figure@60.[557,1031,1604,1825]" captionTargetPageId="60" captionText="FIGURE 28. A. ichabod, female, dorsal aspect; a) apex of elytron; b) Sternum VI, female, ventral aspect. Measures (mm): WE = 1.5; SW = 2.8." httpUri="https://zenodo.org/record/162842/files/figure.png" pageId="60" pageNumber="61">Figs. 28a, 28</figureCitation>
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b)
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</paragraph>
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</subSubSection>
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<subSubSection id="C31165577829FFB3FEECFD849E71FDB6" pageId="60" pageNumber="61" type="materials_examined">
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<paragraph id="8BB436DC7829FFB3FEECFD849E71FDB6" blockId="60.[264,1324,518,1544]" pageId="60" pageNumber="61">
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<emphasis id="B97FEACE7829FFB3FEECFD849DB5FDDF" bold="true" box="[264,385,518,544]" pageId="60" pageNumber="61">
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<typeStatus id="54B0887E7829FFB3FEECFD849D48FDDF" box="[264,380,518,544]" pageId="60" pageNumber="61" type="holotype">Holotype</typeStatus>
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.
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</emphasis>
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Female,
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<collectingCountry id="F31C764C7829FFB3FDEEFD859EFDFDDE" box="[522,713,519,545]" name="Costa Rica" pageId="60" pageNumber="61">COSTA RICA</collectingCountry>
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: Alajuela, Atenas,
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<geoCoordinate id="EE3F501B7829FFB3FC35FD859815FDDE" box="[977,1057,519,545]" direction="north" orientation="latitude" pageId="60" pageNumber="61" precision="925" value="9.966666">9º58'N</geoCoordinate>
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<geoCoordinate id="EE3F501B7829FFB3FBD2FD8598AFFDDE" box="[1078,1179,519,545]" direction="west" orientation="longitude" pageId="60" pageNumber="61" precision="925" value="-84.38333">84º23'W</geoCoordinate>
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, May, (C. Aviles)(
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<collectionCode id="ED1AAE197829FFB3FE80FDAD9D8DFDB6" box="[356,441,559,585]" pageId="60" pageNumber="61">UCOR</collectionCode>
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) ADP7607.
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</paragraph>
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</subSubSection>
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<subSubSection id="C31165577829FFB3FEDCFDD4984AFD66" pageId="60" pageNumber="61" type="diagnosis">
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<paragraph id="8BB436DC7829FFB3FEDCFDD4984AFD66" blockId="60.[264,1324,518,1544]" pageId="60" pageNumber="61">
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<emphasis id="B97FEACE7829FFB3FEDCFDD49D83FD8F" bold="true" box="[312,439,598,624]" pageId="60" pageNumber="61">Diagnosis.</emphasis>
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This is the only species of Costa Rican
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<taxonomicName id="4C0B4D5F7829FFB3FC7BFDD59FEEFD8F" box="[927,986,599,624]" class="Insecta" family="Carabidae" genus="Agra" kingdom="Animalia" order="Coleoptera" pageId="60" pageNumber="61" phylum="Arthropoda" rank="genus">
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<emphasis id="B97FEACE7829FFB3FC7BFDD59FEEFD8F" box="[927,986,599,624]" italics="true" pageId="60" pageNumber="61">Agra</emphasis>
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</taxonomicName>
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with aeneous elytral foveae and slightly oblique apex without a sutural tooth; foveae nearly contiguous.
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</paragraph>
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</subSubSection>
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<subSubSection id="C31165577829FFB3FEDCFD249DFFFBFE" pageId="60" pageNumber="61" type="description">
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<paragraph id="8BB436DC7829FFB3FEDCFD249DFFFBFE" blockId="60.[264,1324,518,1544]" pageId="60" pageNumber="61">
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<emphasis id="B97FEACE7829FFB3FEDCFD249DE4FD3F" bold="true" box="[312,464,678,704]" pageId="60" pageNumber="61">Description.</emphasis>
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Male unknown. Color and luster: Rufopiceous, aeneous. Legs rufopiceous with slightly infuscated knees; venter rufoaeneous. Foveae of interneurs with metallic blue reflections. Form: Head unknown. Pronotum with large punctures along each side of midline. Female slender, with elytra slightly flared behind middle. Elytron with apex (
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<figureCitation id="13302A597829FFB3FEF5FCC59D41FC9E" box="[273,373,839,865]" captionStart="FIGURE 28. A" captionStartId="60.[264,367,1839,1863]" captionTargetBox="[558,1021,1610,1823]" captionTargetId="figure@60.[557,1031,1604,1825]" captionTargetPageId="60" captionText="FIGURE 28. A. ichabod, female, dorsal aspect; a) apex of elytron; b) Sternum VI, female, ventral aspect. Measures (mm): WE = 1.5; SW = 2.8." httpUri="https://zenodo.org/record/162842/files/figure.png" pageId="60" pageNumber="61">Fig. 28a</figureCitation>
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) moderately oblique, lateral tooth small, sutural apex rounded; foveae of interneurs large and positioned irregularly, some separated by their own diameter, other coalesced. Sternum VI (
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<figureCitation id="13302A597829FFB3FDECFC159E56FC4E" box="[520,610,919,945]" captionStart="FIGURE 28. A" captionStartId="60.[264,367,1839,1863]" captionTargetBox="[558,1021,1610,1823]" captionTargetId="figure@60.[557,1031,1604,1825]" captionTargetPageId="60" captionText="FIGURE 28. A. ichabod, female, dorsal aspect; a) apex of elytron; b) Sternum VI, female, ventral aspect. Measures (mm): WE = 1.5; SW = 2.8." httpUri="https://zenodo.org/record/162842/files/figure.png" pageId="60" pageNumber="61">Fig. 28</figureCitation>
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b) of female nearly truncate. Size: small,
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<quantity id="4CF39B397829FFB3FB8CFC159887FC4E" box="[1128,1203,919,945]" metricMagnitude="-2" metricUnit="m" metricValue="1.2" pageId="60" pageNumber="61" unit="mm" value="12.0">12mm</quantity>
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in length (length of head estimated based on proportions of
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<taxonomicName id="4C0B4D5F7829FFB3FC8BFC3D98BAFC26" authority="Erwin" authorityName="Erwin" box="[879,1166,959,985]" class="Insecta" family="Carabidae" genus="Agra" kingdom="Animalia" order="Coleoptera" pageId="60" pageNumber="61" phylum="Arthropoda" rank="species" species="monteverde">
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<emphasis id="B97FEACE7829FFB3FC8BFC3D9808FC27" box="[879,1084,959,984]" italics="true" pageId="60" pageNumber="61">Agra monteverde</emphasis>
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Erwin
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</taxonomicName>
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(see below), 3.0mm in width.
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</paragraph>
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</subSubSection>
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<subSubSection id="C31165577829FFB3FEDCFB8C9F7EFB0E" pageId="60" pageNumber="61" type="materials_examined">
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<paragraph id="8BB436DC7829FFB3FEDCFB8C98E0FBD6" blockId="60.[264,1324,518,1544]" box="[312,1236,1038,1065]" pageId="60" pageNumber="61">
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<emphasis id="B97FEACE7829FFB3FEDCFB8C9EA4FBD7" bold="true" box="[312,656,1038,1064]" pageId="60" pageNumber="61">Other specimens examined.</emphasis>
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The female
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<typeStatus id="54B0887E7829FFB3FCC0FB8D9FA5FBD6" box="[804,913,1039,1065]" pageId="60" pageNumber="61" type="holotype">Holotype</typeStatus>
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is the only specimen I saw.
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</paragraph>
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<paragraph id="8BB436DC7829FFB3FEDCFBB49F7EFB0E" blockId="60.[264,1324,518,1544]" pageId="60" pageNumber="61">
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<emphasis id="B97FEACE7829FFB3FEDCFBB49E35FBAF" bold="true" box="[312,513,1078,1104]" pageId="60" pageNumber="61">Specific epithet.</emphasis>
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The specific epithet,
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<taxonomicName id="4C0B4D5F7829FFB3FCEDFBB59F53FBAF" box="[777,871,1079,1104]" class="Insecta" family="Carabidae" genus="Agra" kingdom="Animalia" order="Coleoptera" pageId="60" pageNumber="61" phylum="Arthropoda" rank="species" species="ichabod">
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<emphasis id="B97FEACE7829FFB3FCEDFBB59F53FBAF" box="[777,871,1079,1104]" italics="true" pageId="60" pageNumber="61">ichabod</emphasis>
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</taxonomicName>
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, is used as a noun in apposition, and refers to the fact that the
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<typeStatus id="54B0887E7829FFB3FDD4FBDD9EA9FB86" box="[560,669,1119,1145]" pageId="60" pageNumber="61" type="holotype">Holotype</typeStatus>
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is missing its head and the illusion is that of the frightened Schoolteacher
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<taxonomicName id="4C0B4D5F7829FFB3FE18FB059E6EFB5E" box="[508,602,1159,1185]" class="Insecta" family="Carabidae" genus="Agra" kingdom="Animalia" order="Coleoptera" pageId="60" pageNumber="61" phylum="Arthropoda" rank="species" species="ichabod">Ichabod</taxonomicName>
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Crane’s phantom nemesis, the Headless Horseman, in The Legend of Sleepy Hollow from The Sketchbook of Geoffrey Crayon, by the American author, Washington Irving, published in
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<date id="FFB5101C7829FFB3FD05FB559F73FB0E" box="[737,839,1239,1265]" pageId="60" pageNumber="61" value="1819" valueMax="1820">181920</date>
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.
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</paragraph>
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</subSubSection>
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<subSubSection id="C31165577829FFB3FEDCFB7C9F00F9F7" pageId="60" pageNumber="61" type="description">
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<paragraph id="8BB436DC7829FFB3FEDCFB7C9F00F9F7" blockId="60.[264,1324,518,1544]" pageId="60" pageNumber="61">
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<emphasis id="B97FEACE7829FFB3FEDCFB7C9DB1FAE7" bold="true" box="[312,389,1278,1304]" pageId="60" pageNumber="61">Notes.</emphasis>
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The
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<typeStatus id="54B0887E7829FFB3FE24FB7D9E19FAE6" box="[448,557,1279,1305]" pageId="60" pageNumber="61" type="holotype">Holotype</typeStatus>
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is from the Cordillera Volcánica Central Conservation Area. This very distinctive species is represented only by the female
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<typeStatus id="54B0887E7829FFB3FC34FAA59874FABE" box="[976,1088,1319,1345]" pageId="60" pageNumber="61" type="holotype">Holotype</typeStatus>
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, which is headless, however, its unique elytral attributes justify giving it a name and bringing it to the attention of collectors. The head may be described later, based on discovery of a second, complete, specimen. The closely related species,
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<taxonomicName id="4C0B4D5F7829FFB3FCC3FA1D9825FA46" authority="Bates" authorityName="Bates" box="[807,1041,1439,1465]" class="Insecta" family="Carabidae" genus="Agra" kingdom="Animalia" order="Coleoptera" pageId="60" pageNumber="61" phylum="Arthropoda" rank="species" species="soccata">
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<emphasis id="B97FEACE7829FFB3FCC3FA1D9FF2FA47" box="[807,966,1439,1464]" italics="true" pageId="60" pageNumber="61">Agra soccata</emphasis>
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Bates
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</taxonomicName>
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of
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<collectingCountry id="F31C764C7829FFB3FBDAFA1D98AFFA46" box="[1086,1179,1439,1465]" name="Panama" pageId="60" pageNumber="61">Panamá</collectingCountry>
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with which this female agrees in attributes provided the information on male attributes, hence my assignment of this species to the
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<taxonomicName id="4C0B4D5F7829FFB3FD6EFA6C9ED4F9F7" box="[650,736,1518,1544]" class="Insecta" family="Carabidae" genus="Agra" kingdom="Animalia" order="Coleoptera" pageId="60" pageNumber="61" phylum="Arthropoda" rank="species" species="soccata">soccata</taxonomicName>
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group.
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</paragraph>
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</subSubSection>
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</treatment>
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</document> |