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<document id="639F113E8EDECDC320E68FDB7EDDE969" ID-CLB-Dataset="28899" ID-DOI="10.5852/ejt.2018.462" ID-GBIF-Dataset="328c4344-2529-4aae-9c38-a3f9cb576c35" ID-Zenodo-Dep="1467972" ID-ZooBank="urn:lsid:zoobank.org:pub:60992A95-1F59-4407-9774-B8FC1545A8B4" IM.bibliography_approvedBy="felipe" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1540146986426" checkinUser="plazi" docAuthor="Ruttapon Srisonchai, Henrik Enghoff, Natdanai Likhitrakarn &amp; Somsak Panha" docDate="2018" docId="03A287AE0D78226FF7E3EC5FFCA45A4D" docLanguage="en" docName="ejt-462_srisonchai_enghoff_likhitrkarn_panha.pdf.imf" docOrigin="European Journal of Taxonomy 462" docStyle="DocumentStyle:EF2B578F1D15862ADE45B0C07C620911.14:EJT.2018-.journal_article.type1" docStyleId="EF2B578F1D15862ADE45B0C07C620911" docStyleName="EJT.2018-.journal_article.type1" docStyleVersion="14" docTitle="Nagaxytes Srisonchai, Enghoff &amp; Panha 2018, gen. nov." docType="treatment" docUuid="D3B82632-5FC4-4605-AC19-99EAFEB6A23B" docUuidSource="ZooBank" docVersion="10" lastPageNumber="11" masterDocId="FF9BFFD60D7B2265F629EF21FF8A5847" masterDocTitle="A revision of dragon millipedes II: the new genus Nagaxytes gen. nov., with the description of three new species (Diplopoda, Polydesmida, Paradoxosomatidae)" masterLastPageNumber="44" masterPageNumber="1" pageNumber="4" updateTime="1698580138259" updateUser="ExternalLinkService">
<mods:mods id="276A3D5BB36FDDFC44D92263E5427203" xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo id="454127E03C98462745CD0B408C303C90">
<mods:title id="B9D4CC70B0FF68DED41F84A54A061E27">A revision of dragon millipedes II: the new genus Nagaxytes gen. nov., with the description of three new species (Diplopoda, Polydesmida, Paradoxosomatidae)</mods:title>
</mods:titleInfo>
<mods:name id="490739180AB0FFA46E64CA14F9FDB531" type="personal">
<mods:role id="0832BEC17F14E34872DEEFE239C41E73">
<mods:roleTerm id="EA1F28E23813E0E6EC11D5B91AE9AE20">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="E758F78677E2F06CAF877F591F191F01">Ruttapon Srisonchai</mods:namePart>
</mods:name>
<mods:name id="804C190996340DDD7CF7851EDF9E1A0E" type="personal">
<mods:role id="A24D2B35A2C22AC68B7E3C08B1A33106">
<mods:roleTerm id="6B80DD983E6F298B598CAB4E723466DA">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="E539A941697A988FA3AD63B465F228F2">Henrik Enghoff</mods:namePart>
</mods:name>
<mods:name id="8F5EB4339A79EF5A1D6BD32EC1F32CB6" type="personal">
<mods:role id="82C961486F196438E6CDBD920454FCCE">
<mods:roleTerm id="03C0BF50ACE20CFBC0531955BC632209">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="9908C49D1568E1E1AE5FAD99C372BBE3">Natdanai Likhitrakarn</mods:namePart>
</mods:name>
<mods:name id="1F200658C6B765BB1463751030B50D13" type="personal">
<mods:role id="DAA7653C7DEB2ADB4F61F76F83BEF8A0">
<mods:roleTerm id="4003FBA91728286C7E7C81809A7AAC59">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="01C1731C137BD82D77AE620208BABDC7">Somsak Panha</mods:namePart>
</mods:name>
<mods:typeOfResource id="FA51C97EC24E60C5DDE717EA50B7CED5">text</mods:typeOfResource>
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<mods:titleInfo id="8D9EE6B96BFFED51D8224C1BD64514E5">
<mods:title id="4874F9227D6B02A3834287784E86FBEC">European Journal of Taxonomy</mods:title>
</mods:titleInfo>
<mods:part id="C973B49263BA02D6FDEA9BFE48881305">
<mods:date id="7255CC9AA893E52F5CFB7DDAEEABD45D">2018</mods:date>
<mods:detail id="C47E262BDEA2A64F7E74634D2979CF48" type="pubDate">
<mods:number id="6DEB635CCD945507CE9B079CC583BD51">2018-09-25</mods:number>
</mods:detail>
<mods:detail id="0BBCB307EC91C6575DD9E46B0206D297" type="volume">
<mods:number id="5C6548F4900E8AF775D82E4802E2D918">462</mods:number>
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<mods:start id="155A8D1FA2ABA1E75C6479A5BBAEE5F2">1</mods:start>
<mods:end id="CCF9957ACA7D442C371EAFF2C0FED9FA">44</mods:end>
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<mods:classification id="47B2F1D878BD26803A3D13CE3FECBE1D">journal article</mods:classification>
<mods:identifier id="8C4CE2A21C7BA1CC5C698BF8E9FA61F7" type="CLB-Dataset">28899</mods:identifier>
<mods:identifier id="EC06FFCEE87D0DCA2CB455E456AA2B1D" type="DOI">10.5852/ejt.2018.462</mods:identifier>
<mods:identifier id="987BBB565D331B290AB0FAB40EC175AC" type="GBIF-Dataset">328c4344-2529-4aae-9c38-a3f9cb576c35</mods:identifier>
<mods:identifier id="4D7662400ACFDBD377252E6B7CB36553" type="Zenodo-Dep">1467972</mods:identifier>
<mods:identifier id="95D7E5219FDB320D0D1E4E65A01278A5" type="ZooBank">urn:lsid:zoobank.org:pub:60992A95-1F59-4407-9774-B8FC1545A8B4</mods:identifier>
</mods:mods>
<treatment id="03A287AE0D78226FF7E3EC5FFCA45A4D" ID-DOI="http://doi.org/10.5281/zenodo.5946400" ID-GBIF-Taxon="148885083" ID-Zenodo-Dep="5946400" LSID="urn:lsid:zoobank.org:act:D3B82632-5FC4-4605-AC19-99EAFEB6A23B" httpUri="http://treatment.plazi.org/id/03A287AE0D78226FF7E3EC5FFCA45A4D" lastPageId="10" lastPageNumber="11" pageId="3" pageNumber="4">
<subSubSection id="C31165330D782266F7E3EC5FFB435BFB" pageId="3" pageNumber="4" type="nomenclature">
<paragraph id="8BB436B80D782266F7E3EC5FFBE05BDF" blockId="3.[363,1225,894,956]" box="[458,1130,894,920]" pageId="3" pageNumber="4">
Genus
<taxonomicName id="4C0B4D3B0D782266F435EC5FFC725BDF" ID-CoL="7PPPD" authority="Srisonchai, Enghoff &amp; Panha" authorityName="Srisonchai, Enghoff &amp; Panha" authorityYear="2018" box="[540,1016,894,920]" class="Diplopoda" family="Paradoxosomatidae" genus="Nagaxytes" higherTaxonomySource="CoL" kingdom="Animalia" order="Polydesmida" pageId="3" pageNumber="4" phylum="Arthropoda" rank="genus" status="gen. nov.">
<emphasis id="B97FEAAA0D782266F435EC5FFD135BDF" bold="true" box="[540,665,894,920]" italics="true" pageId="3" pageNumber="4">Nagaxytes</emphasis>
Srisonchai, Enghoff &amp; Panha
</taxonomicName>
<taxonomicNameLabel id="A24C57D10D782266F5D6EC5FFBE05BDF" box="[1023,1130,894,920]" pageId="3" pageNumber="4" rank="genus">gen. nov.</taxonomicNameLabel>
</paragraph>
<paragraph id="8BB436B80D782266F742EC80FB435BFB" blockId="3.[363,1225,894,956]" box="[363,1225,929,956]" pageId="3" pageNumber="4">urn:lsid:zoobank.org:act:D3B82632-5FC4-4605-AC19-99EAFEB6A23B</paragraph>
</subSubSection>
<subSubSection id="C31165330D782266F694ECC2FCDB5C6C" pageId="3" pageNumber="4" type="type_taxon">
<paragraph id="8BB436B80D782266F694ECC2FED25BBB" blockId="3.[189,344,994,1020]" box="[189,344,994,1021]" pageId="3" pageNumber="4">
<emphasis id="B97FEAAA0D782266F694ECC2FED25BBB" bold="true" box="[189,344,994,1021]" pageId="3" pageNumber="4">
<typeStatus id="54B0881A0D782266F694ECC2FF705BBA" box="[189,250,995,1021]" pageId="3" pageNumber="4">Type</typeStatus>
species
</emphasis>
</paragraph>
<paragraph id="8BB436B80D782266F694EB33FCDB5C6C" blockId="3.[189,849,1041,1067]" box="[189,849,1041,1067]" pageId="3" pageNumber="4">
<taxonomicName id="4C0B4D3B0D782266F694EB33FCDB5C6C" ID-CoL="6CNL8" authority="Golovatch &amp; Enghoff, 1994" authorityName="Golovatch &amp; Enghoff" authorityYear="1994" box="[189,849,1041,1067]" class="Diplopoda" family="Paradoxosomatidae" genus="Desmoxytes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Polydesmida" pageId="3" pageNumber="4" phylum="Arthropoda" rank="species" species="acantherpestes">
<emphasis id="B97FEAAA0D782266F694EB33FD8A5C6C" box="[189,512,1041,1067]" italics="true" pageId="3" pageNumber="4">Desmoxytes acantherpestes</emphasis>
<bibRefCitation id="EF9A4B490D782266F421EB30FCDB5C6C" box="[520,849,1041,1067]" pageId="3" pageNumber="4" refId="ref11554" refString="Golovatch S. I. &amp; Enghoff H. 1994. Review of the dragon millipedes, genus Desmoxytes Chamberlin, 1923 (Diplopoda, Polydesmida, Paradoxosomatidae). Steenstrupia 20 (2): 45 - 71." type="journal article">Golovatch &amp; Enghoff, 1994</bibRefCitation>
</taxonomicName>
</paragraph>
</subSubSection>
<subSubSection id="C31165330D782266F694EB73FC135DF7" pageId="3" pageNumber="4" type="diagnosis">
<paragraph id="8BB436B80D782266F694EB73FEBC5C2B" blockId="3.[189,310,1106,1132]" box="[189,310,1106,1132]" pageId="3" pageNumber="4">
<heading id="D0FC81D40D782266F694EB73FEBC5C2B" bold="true" box="[189,310,1106,1132]" fontSize="11" level="3" pageId="3" pageNumber="4" reason="3">
<emphasis id="B97FEAAA0D782266F694EB73FEBC5C2B" bold="true" box="[189,310,1106,1132]" pageId="3" pageNumber="4">Diagnosis</emphasis>
</heading>
</paragraph>
<paragraph id="8BB436B80D782266F694EBA0FC9F5CDC" blockId="3.[189,789,1153,1180]" box="[189,789,1153,1180]" pageId="3" pageNumber="4">
The genus
<taxonomicName id="4C0B4D3B0D782266F717EBA3FE325CDC" authorityName="Srisonchai, Enghoff &amp; Panha" authorityYear="2018" box="[318,440,1154,1179]" class="Diplopoda" family="Paradoxosomatidae" genus="Nagaxytes" higherTaxonomySource="CoL" kingdom="Animalia" order="Polydesmida" pageId="3" pageNumber="4" phylum="Arthropoda" rank="genus" status="gen. nov.">
<emphasis id="B97FEAAA0D782266F717EBA3FE325CDC" box="[318,440,1154,1179]" italics="true" pageId="3" pageNumber="4">Nagaxytes</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A24C57D10D782266F796EBA3FDA05CDB" box="[447,554,1154,1180]" pageId="3" pageNumber="4" rank="genus">gen. nov.</taxonomicNameLabel>
is characterized by:
</paragraph>
<paragraph id="8BB436B80D782266F694EBE2FE715C9B" blockId="3.[189,1398,1218,1456]" box="[189,507,1218,1245]" pageId="3" pageNumber="4">1. Paraterga subspiniform.</paragraph>
<paragraph id="8BB436B80D782266F694EBC7FA9A5D46" blockId="3.[189,1398,1218,1456]" box="[189,1296,1254,1281]" pageId="3" pageNumber="4">2. Metaterga with 2 rows of tubercles/cones/spines (lateral spines of posterior row very long).</paragraph>
<paragraph id="8BB436B80D782266F694EA28FD565D64" blockId="3.[189,1398,1218,1456]" box="[189,732,1289,1316]" pageId="3" pageNumber="4">3. Male femora 5 and 6 without modification.</paragraph>
<paragraph id="8BB436B80D782266F694EA0DFDAD5D2E" blockId="3.[189,1398,1218,1456]" pageId="3" pageNumber="4">4. Postfemoral part of gonopod conspicuous, demarcated from femur by a deep mesal sulcus and a shallow/deep lateral sulcus.</paragraph>
<paragraph id="8BB436B80D782266F694EA53FC165DCB" blockId="3.[189,1398,1218,1456]" box="[189,924,1394,1421]" pageId="3" pageNumber="4">5. Lamina lateralis obviously separated from lamina medialis.</paragraph>
<paragraph id="8BB436B80D782266F694EAB7FC135DF7" blockId="3.[189,1398,1218,1456]" box="[189,921,1430,1456]" pageId="3" pageNumber="4">6. Lamina medialis long and curved, apically fringed/hooked.</paragraph>
</subSubSection>
<subSubSection id="C31165330D782266F694EAF6FC795E21" pageId="3" pageNumber="4" type="etymology">
<paragraph id="8BB436B80D782266F694EAF6FECE5DB6" blockId="3.[189,324,1495,1521]" box="[189,324,1495,1521]" pageId="3" pageNumber="4">
<heading id="D0FC81D40D782266F694EAF6FECE5DB6" bold="true" box="[189,324,1495,1521]" fontSize="11" level="3" pageId="3" pageNumber="4" reason="3">
<emphasis id="B97FEAAA0D782266F694EAF6FECE5DB6" bold="true" box="[189,324,1495,1521]" pageId="3" pageNumber="4">Etymology</emphasis>
</heading>
</paragraph>
<paragraph id="8BB436B80D782266F694E927FC795E21" blockId="3.[189,1398,1542,1639]" pageId="3" pageNumber="4">
‘
<emphasis id="B97FEAAA0D782266F6EEE926FE8D5E67" box="[199,263,1543,1568]" italics="true" pageId="3" pageNumber="4">Naga</emphasis>
’ is a Sanskrit and Pali word, relating to a category of snake-like spirits in Buddhist and Hindu mythology, and refers to the snake-like shape of the lamina medialis of the gonopod; ‘-
<emphasis id="B97FEAAA0D782266F2FAE90BFA845E04" box="[1235,1294,1578,1603]" italics="true" pageId="3" pageNumber="4">xytes</emphasis>
’ ensures harmony with
<taxonomicName id="4C0B4D3B0D782266F741E96CFE7E5E21" authorityName="Chamberlin" authorityYear="1923" box="[360,500,1613,1638]" class="Diplopoda" family="Paradoxosomatidae" genus="Desmoxytes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Polydesmida" pageId="3" pageNumber="4" phylum="Arthropoda" rank="genus">
<emphasis id="B97FEAAA0D782266F741E96CFE7E5E21" box="[360,500,1613,1638]" italics="true" pageId="3" pageNumber="4">Desmoxytes</emphasis>
</taxonomicName>
(and its synonym
<taxonomicName id="4C0B4D3B0D782266F4E4E96CFC6C5E21" authority="Jeekel, 1980" authorityName="Jeekel" authorityYear="1980" box="[717,998,1612,1638]" class="Diplopoda" family="Paradoxosomatidae" genus="Pteroxytes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Polydesmida" pageId="3" pageNumber="4" phylum="Arthropoda" rank="genus">
<emphasis id="B97FEAAA0D782266F4E4E96CFCC25E21" box="[717,840,1613,1638]" italics="true" pageId="3" pageNumber="4">Pteroxytes</emphasis>
<bibRefCitation id="EF9A4B490D782266F566E96DFC6C5E21" box="[847,998,1612,1638]" pageId="3" pageNumber="4" refId="ref11662" refString="Jeekel C. A. W. 1980. The generic allocation of some little-known Paradoxosomatidae from South- East Asia (Diplopoda, Polydesmida). Revue suisse de Zoologie 87 (3): 651 - 670. Available from https: // biodiversitylibrary. org / page / 41420961 [accessed 22 Aug. 2018]." type="journal article">Jeekel, 1980</bibRefCitation>
</taxonomicName>
).
</paragraph>
</subSubSection>
<subSubSection id="C31165330D782266F694E9AFFCD75F14" pageId="3" pageNumber="4" type="synonymic_list">
<paragraph id="8BB436B80D782266F694E9AFFE395EE0" blockId="3.[189,435,1677,1704]" box="[189,435,1677,1704]" pageId="3" pageNumber="4">
<heading id="D0FC81D40D782266F694E9AFFE395EE0" bold="true" box="[189,435,1677,1704]" fontSize="11" level="3" pageId="3" pageNumber="4" reason="3">
<emphasis id="B97FEAAA0D782266F694E9AFFE395EE0" bold="true" box="[189,435,1677,1704]" pageId="3" pageNumber="4">Included species (4)</emphasis>
</heading>
</paragraph>
<paragraph id="8BB436B80D782266F694E9F1FC685EAD" blockId="3.[189,994,1743,1875]" box="[189,994,1743,1770]" pageId="3" pageNumber="4">
-
<taxonomicName id="4C0B4D3B0D782266F6E7E9F1FC895EAE" authority="(Golovatch &amp; Enghoff, 1994)" baseAuthorityName="Golovatch &amp; Enghoff" baseAuthorityYear="1994" box="[206,771,1743,1769]" class="Diplopoda" family="Paradoxosomatidae" genus="Desmoxytes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Polydesmida" pageId="3" pageNumber="4" phylum="Arthropoda" rank="species" species="acantherpestes" status="gen. et comb. nov.">
<emphasis id="B97FEAAA0D782266F6E7E9F1FE2A5EAE" box="[206,416,1743,1769]" italics="true" pageId="3" pageNumber="4">N. acantherpestes</emphasis>
(
<bibRefCitation id="EF9A4B490D782266F786E9EEFD715EAE" box="[431,763,1743,1769]" pageId="3" pageNumber="4" refId="ref11554" refString="Golovatch S. I. &amp; Enghoff H. 1994. Review of the dragon millipedes, genus Desmoxytes Chamberlin, 1923 (Diplopoda, Polydesmida, Paradoxosomatidae). Steenstrupia 20 (2): 45 - 71." type="journal article">Golovatch &amp; Enghoff, 1994</bibRefCitation>
)
</taxonomicName>
<taxonomicNameLabel id="A24C57D10D782266F522E9F1FC685EAD" box="[779,994,1743,1770]" pageId="3" pageNumber="4">gen. et comb. nov.</taxonomicNameLabel>
</paragraph>
<paragraph id="8BB436B80D782266F694E9D2FCC55F4A" blockId="3.[189,994,1743,1875]" box="[189,847,1778,1805]" pageId="3" pageNumber="4">
-
<taxonomicName id="4C0B4D3B0D782266F6E7E9D2FD1D5F4B" authority="Srisonchai, Enghoff &amp; Panha" authorityName="Srisonchai, Enghoff &amp; Panha" authorityYear="2018" box="[206,663,1778,1804]" class="Diplopoda" family="Paradoxosomatidae" genus="Nagaxytes" higherTaxonomySource="CoL" kingdom="Animalia" order="Polydesmida" pageId="3" pageNumber="4" phylum="Arthropoda" rank="species" species="erecta" status="gen. et sp. nov.">
<emphasis id="B97FEAAA0D782266F6E7E9D2FEB25F4B" box="[206,312,1779,1804]" italics="true" pageId="3" pageNumber="4">N. erecta</emphasis>
Srisonchai, Enghoff &amp; Panha
</taxonomicName>
<taxonomicNameLabel id="A24C57D10D782266F4B7E9D2FCC55F4A" box="[670,847,1779,1805]" pageId="3" pageNumber="4" rank="species">gen. et sp. nov.</taxonomicNameLabel>
</paragraph>
<paragraph id="8BB436B80D782266F694E837FCEA5F77" blockId="3.[189,994,1743,1875]" box="[189,864,1813,1840]" pageId="3" pageNumber="4">
-
<taxonomicName id="4C0B4D3B0D782266F6E7E837FD235F68" authority="Srisonchai, Enghoff &amp; Panha" authorityName="Srisonchai, Enghoff &amp; Panha" authorityYear="2018" box="[206,681,1813,1840]" class="Diplopoda" family="Paradoxosomatidae" genus="Nagaxytes" higherTaxonomySource="CoL" kingdom="Animalia" order="Polydesmida" pageId="3" pageNumber="4" phylum="Arthropoda" rank="species" species="gracilis" status="gen. et sp. nov.">
<emphasis id="B97FEAAA0D782266F6E7E837FEC05F68" box="[206,330,1813,1839]" italics="true" pageId="3" pageNumber="4">N. gracilis</emphasis>
Srisonchai, Enghoff &amp; Panha
</taxonomicName>
<taxonomicNameLabel id="A24C57D10D782266F499E837FCEA5F77" box="[688,864,1814,1840]" pageId="3" pageNumber="4" rank="species">gen. et sp. nov.</taxonomicNameLabel>
</paragraph>
<paragraph id="8BB436B80D782266F694E818FCD75F14" blockId="3.[189,994,1743,1875]" box="[189,861,1848,1875]" pageId="3" pageNumber="4">
-
<taxonomicName id="4C0B4D3B0D782266F6E7E818FD2C5F15" authority="Srisonchai, Enghoff &amp; Panha" authorityName="Srisonchai, Enghoff &amp; Panha" authorityYear="2018" box="[206,678,1848,1875]" class="Diplopoda" family="Paradoxosomatidae" genus="Nagaxytes" higherTaxonomySource="CoL" kingdom="Animalia" order="Polydesmida" pageId="3" pageNumber="4" phylum="Arthropoda" rank="species" species="spatula" status="gen. et sp. nov.">
<emphasis id="B97FEAAA0D782266F6E7E818FECC5F14" box="[206,326,1849,1875]" italics="true" pageId="3" pageNumber="4">N. spatula</emphasis>
Srisonchai, Enghoff &amp; Panha
</taxonomicName>
<taxonomicNameLabel id="A24C57D10D782266F484E818FCD75F14" box="[685,861,1849,1875]" pageId="3" pageNumber="4" rank="species">gen. et sp. nov.</taxonomicNameLabel>
</paragraph>
</subSubSection>
<subSubSection id="C31165330D782261F694E85BFCA2592B" lastPageId="4" lastPageNumber="5" pageId="3" pageNumber="4" type="discussion">
<paragraph id="8BB436B80D782266F694E85BFEA55FD3" blockId="3.[189,303,1914,1940]" box="[189,303,1914,1940]" pageId="3" pageNumber="4">
<heading id="D0FC81D40D782266F694E85BFEA55FD3" bold="true" box="[189,303,1914,1940]" fontSize="11" level="3" pageId="3" pageNumber="4" reason="3">
<emphasis id="B97FEAAA0D782266F694E85BFEA55FD3" bold="true" box="[189,303,1914,1940]" pageId="3" pageNumber="4">Remarks</emphasis>
</heading>
</paragraph>
<paragraph id="8BB436B80D782261F694E889FCA2592B" blockId="3.[189,1398,1960,2022]" lastBlockId="4.[189,1399,267,364]" lastPageId="4" lastPageNumber="5" pageId="3" pageNumber="4">
<bibRefCitation id="EF9A4B490D782266F694E889FE455F84" box="[189,463,1960,1987]" pageId="3" pageNumber="4" refId="ref12078" refString="Srisonchai R., Enghoff H., Likhitrakarn N. &amp; Panha S. 2018. A revision of dragon millipedes I: genus Desmoxytes Chamberlin, 1923, with the description of eight new species (Diplopoda, Polydesmida, Paradoxosomatidae). ZooKeys 761: 1 - 177. https: // doi. org / 10.3897 / zookeys. 761.24214" type="journal article">
Srisonchai
<emphasis id="B97FEAAA0D782266F769E888FEF05F84" box="[320,378,1961,1987]" italics="true" pageId="3" pageNumber="4">et al.</emphasis>
(2018)
</bibRefCitation>
proposed to subdivide
<taxonomicName id="4C0B4D3B0D782266F4C9E888FA845F84" authority="sensu Golovatch &amp; Enghoff (1994)" authorityName="sensu Golovatch &amp; Enghoff" authorityYear="1994" box="[736,1294,1960,1987]" class="Diplopoda" family="Paradoxosomatidae" genus="Desmoxytes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Polydesmida" pageId="3" pageNumber="4" phylum="Arthropoda" rank="genus">
<emphasis id="B97FEAAA0D782266F4C9E888FC3E5F85" box="[736,948,1961,1986]" italics="true" pageId="3" pageNumber="4">Desmoxytes sensu</emphasis>
<bibRefCitation id="EF9A4B490D782266F592E889FA845F84" box="[955,1294,1960,1987]" pageId="3" pageNumber="4" refId="ref11554" refString="Golovatch S. I. &amp; Enghoff H. 1994. Review of the dragon millipedes, genus Desmoxytes Chamberlin, 1923 (Diplopoda, Polydesmida, Paradoxosomatidae). Steenstrupia 20 (2): 45 - 71." type="journal article">Golovatch &amp; Enghoff (1994)</bibRefCitation>
</taxonomicName>
into five genera. The ‘
<taxonomicName id="4C0B4D3B0D782266F749E8EDFD9A5FA1" baseAuthorityName="Golovatch &amp; Enghoff" baseAuthorityYear="1994" box="[352,528,1996,2022]" class="Diplopoda" family="Paradoxosomatidae" genus="Desmoxytes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Polydesmida" pageId="3" pageNumber="4" phylum="Arthropoda" rank="species" species="acantherpestes">
<emphasis id="B97FEAAA0D782266F749E8EDFD9A5FA1" box="[352,528,1996,2022]" italics="true" pageId="3" pageNumber="4">acantherpestes</emphasis>
</taxonomicName>
’ group (=
<taxonomicName id="4C0B4D3B0D782266F4BEE8ECFC9B5FA1" authorityName="Srisonchai, Enghoff &amp; Panha" authorityYear="2018" box="[663,785,1997,2022]" class="Diplopoda" family="Paradoxosomatidae" genus="Nagaxytes" higherTaxonomySource="CoL" kingdom="Animalia" order="Polydesmida" pageId="3" pageNumber="43" phylum="Arthropoda" rank="genus" status="gen. nov.">
<emphasis id="B97FEAAA0D782266F4BEE8ECFC9B5FA1" box="[663,785,1997,2022]" italics="true" pageId="3" pageNumber="4">Nagaxytes</emphasis>
</taxonomicName>
) is well-defined by several distinct morphological characters (see diagnosis), especially the distinctive subspiniform paraterga and the very long lamina medialis of the gonopods. Even though the subspiniform paraterga are relatively similar to those of the ‘
<emphasis id="B97FEAAA0D7F2261F6EEEE72FE8D592B" box="[199,263,339,364]" italics="true" pageId="4" pageNumber="5">gigas</emphasis>
’ group, the gonopod details are very different.
</paragraph>
</subSubSection>
<subSubSection id="C31165330D7F2262F694EEB2FBA95FD0" lastPageId="7" lastPageNumber="8" pageId="4" pageNumber="5" type="description">
<paragraph id="8BB436B80D7F2261F694EEB2FD4559EA" blockId="4.[189,1399,403,547]" box="[189,719,403,430]" pageId="4" pageNumber="5">
<heading id="D0FC81D40D7F2261F694EEB2FD4559EA" bold="true" box="[189,719,403,430]" fontSize="11" level="3" pageId="4" pageNumber="5" reason="3">
<emphasis id="B97FEAAA0D7F2261F694EEB2FD4559EA" bold="true" box="[189,719,403,430]" pageId="4" pageNumber="5">
General description of
<taxonomicName id="4C0B4D3B0D7F2261F7F4EEB5FDD059E9" authorityName="Srisonchai, Enghoff &amp; Panha" authorityYear="2018" box="[477,602,404,430]" class="Diplopoda" family="Paradoxosomatidae" genus="Nagaxytes" higherTaxonomySource="CoL" kingdom="Animalia" order="Polydesmida" pageId="4" pageNumber="5" phylum="Arthropoda" rank="genus" status="gen. nov.">
<emphasis id="B97FEAAA0D7F2261F7F4EEB5FDD059E9" bold="true" box="[477,602,404,430]" italics="true" pageId="4" pageNumber="5">Nagaxytes</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A24C57D10D7F2261F448EEB2FD4559EA" box="[609,719,403,429]" pageId="4" pageNumber="5" rank="genus">gen. nov.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
<paragraph id="8BB436B80D7F2261F694EEE3FEB45A64" blockId="4.[189,1399,403,547]" pageId="4" pageNumber="5">
The description applies to adult males and females, except for the gonopods or when ‘male’ is specified (
<figureCitation id="13302A3D0D7F2261F6ECEEC4FEA559B8" box="[197,303,485,511]" captionStart-0="Fig. 1" captionStart-1="Fig. 2" captionStart-2="Fig. 3" captionStartId-0="4.[189,232,1958,1984]" captionStartId-1="5.[189,232,1872,1898]" captionStartId-2="6.[189,231,1809,1835]" captionTargetBox-0="[189,1399,1030,1905]" captionTargetBox-1="[192,1396,282,1802]" captionTargetBox-2="[189,1398,264,1781]" captionTargetId-0="figure@4.[189,1399,1030,1915]" captionTargetId-1="figure@5.[189,1398,269,1817]" captionTargetId-2="figure@6.[189,1398,264,1781]" captionTargetPageId-0="4" captionTargetPageId-1="5" captionTargetPageId-2="6" captionText-0="Fig. 1. Nagaxytes gracilis Srisonchai, Enghoff &amp; Panha gen. et sp. nov., ♂, paratype (CUMZpxDGT00095). Watercolour by R. Srisonchai and P. Kriatpraprai." captionText-1="Fig. 2. General body characters of Nagaxytes Srisonchai, Enghoff &amp; Panha gen. nov. – N. acantherpestes (Golovatch &amp; Enghoff, 1994) gen. et comb. nov., ♂, holotype, Prachuap Khiri Khan Province, Hua Hin District (ZMUC 000101457). A. Entire body. B. Anterior body part. C. Body rings 5–8, showing sternal lobe between coxae 4 and gonopods on ring 7. D. Head and antennae." captionText-2="Fig. 3. General body characters of Nagaxytes Srisonchai, Enghoff &amp; Panha gen. nov. (N. erecta Srisonchai, Enghoff &amp; Panha gen. et sp. nov., specimen from Prang Ka Sri Temple, Thailand), SEM images. A–B. Anterior body part (arrowheads point to pleurosternal carinae). C. Collum (arrowheads point to the setae/tubercles of the anterior and intermediate rows). D. Body rings 9–10. E. Body rings 8–10. F. Telson. G. Posteriormost rings and telson. H. Body ring 10. I. Mouth parts, ventral view. J. Gonopods in situ. K. Sternal lobe between male coxae 4. L, N. Tip of epiproct. M. Tarsus and claw of leg 13." httpUri-0="https://zenodo.org/record/1467974/files/figure.png" httpUri-1="https://zenodo.org/record/1467976/files/figure.png" httpUri-2="https://zenodo.org/record/1467978/files/figure.png" pageId="4" pageNumber="5">Figs 1–3</figureCitation>
). The general description of gonopods is based mainly on
<taxonomicName id="4C0B4D3B0D7F2261F5CBEEC7FB3459B8" authorityName="Srisonchai, Enghoff &amp; Panha" authorityYear="2018" box="[994,1214,485,511]" class="Diplopoda" family="Paradoxosomatidae" genus="Nagaxytes" higherTaxonomySource="CoL" kingdom="Animalia" order="Polydesmida" pageId="4" pageNumber="5" phylum="Arthropoda" rank="species" species="gracilis" status="gen. et sp. nov.">
<emphasis id="B97FEAAA0D7F2261F5CBEEC7FB3459B8" box="[994,1214,485,511]" italics="true" pageId="4" pageNumber="5">Nagaxytes gracilis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A24C57D10D7F2261F2ECEEC4FAFD59B8" box="[1221,1399,485,511]" pageId="4" pageNumber="5" rank="species">gen. et sp. nov.</taxonomicNameLabel>
(
<figureCitation id="13302A3D0D7F2261F6ECED29FEBB5A64" box="[197,305,520,547]" captionStart-0="Fig. 4" captionStart-1="Fig. 5" captionStartId-0="8.[189,232,1919,1945]" captionStartId-1="9.[190,233,1883,1909]" captionTargetBox-0="[196,1381,311,1854]" captionTargetBox-1="[189,1398,279,1828]" captionTargetId-0="figure@8.[189,1398,305,1854]" captionTargetId-1="figure@9.[189,1398,279,1828]" captionTargetPageId-0="8" captionTargetPageId-1="9" captionText-0="Fig. 4. Diagrammatic drawing of right gonopod of Nagaxytes Srisonchai, Enghoff &amp; Panha gen. nov. – N. gracilis Srisonchai, Enghoff &amp; Panha gen. et sp. nov., specimen from Wat Vimooddhi Wanaram, Thailand. A. Mesal view. B. Dorsal view. C. Ventral view." captionText-1="Fig. 5. SEM images of right gonopod of Nagaxytes Srisonchai, Enghoff &amp; Panha gen. nov. – N. gracilis Srisonchai, Enghoff &amp; Panha gen. et sp. nov., specimen from Wat Vimooddhi Wanaram, Thailand. A. Lateral view. B. Mesal view. C. Dorsal view. D. Ventral view. Colours: red = lamina medialis, green = lamina lateralis, yellow = solenomere, purple = seminal groove." httpUri-0="https://zenodo.org/record/1467980/files/figure.png" httpUri-1="https://zenodo.org/record/1467982/files/figure.png" pageId="4" pageNumber="5">Figs 4–5</figureCitation>
).
</paragraph>
<paragraph id="8BB436B80D7F2261F694ED6BFBCF5AC0" blockId="4.[189,1399,585,648]" pageId="4" pageNumber="5">
SIZE. Body length
<quantity id="4CF39B5D0D7F2261F7B5ED6BFD975A23" box="[412,541,586,612]" metricMagnitude="-2" metricUnit="m" metricValue="2.8" metricValueMax="3.4" metricValueMin="2.2" pageId="4" pageNumber="5" unit="mm" value="28.0" valueMax="34.0" valueMin="22.0">22–34 mm</quantity>
(male)
<quantity id="4CF39B5D0D7F2261F457ED6BFD755A23" box="[638,767,586,612]" metricMagnitude="-2" metricUnit="m" metricValue="3.1" metricValueMax="3.8" metricValueMin="2.4" pageId="4" pageNumber="5" unit="mm" value="31.0" valueMax="38.0" valueMin="24.0">24–38 mm</quantity>
(female), width ca. 2.0–
<quantity id="4CF39B5D0D7F2261F203ED6BFB0C5A23" box="[1066,1158,586,612]" metricMagnitude="-3" metricUnit="m" metricValue="2.4" pageId="4" pageNumber="5" unit="mm" value="2.4">2.4 mm</quantity>
(male)
<quantity id="4CF39B5D0D7F2261F2CEED6BFAFD5A23" box="[1255,1399,586,612]" metricMagnitude="-3" metricUnit="m" metricValue="3.0999999999999996" metricValueMax="3.4" metricValueMin="2.8" pageId="4" pageNumber="5" unit="mm" value="3.0999999999999996" valueMax="3.4" valueMin="2.8">2.8–3.4 mm</quantity>
(female), varies between species, usually female wider and longer than male.
</paragraph>
<paragraph id="8BB436B80D7F2261F694ED8FFB345B49" blockId="4.[189,1398,686,783]" pageId="4" pageNumber="5">
COLOUR (
<figureCitation id="13302A3D0D7F2261F70EED8FFEE55A8F" box="[295,367,686,712]" captionStart="Fig. 1" captionStartId="4.[189,232,1958,1984]" captionTargetBox="[189,1399,1030,1905]" captionTargetId="figure@4.[189,1399,1030,1915]" captionTargetPageId="4" captionText="Fig. 1. Nagaxytes gracilis Srisonchai, Enghoff &amp; Panha gen. et sp. nov., ♂, paratype (CUMZpxDGT00095). Watercolour by R. Srisonchai and P. Kriatpraprai." httpUri="https://zenodo.org/record/1467974/files/figure.png" pageId="4" pageNumber="5">Figs 1</figureCitation>
,
<figureCitation id="13302A3D0D7F2261F754ED8FFE4D5A8F" box="[381,455,686,712]" captionStart-0="Fig. 11" captionStart-1="Fig. 12" captionStartId-0="19.[189,232,1883,1909]" captionStartId-1="20.[189,232,1926,1952]" captionTargetBox-0="[192,1395,283,1829]" captionTargetBox-1="[192,1394,306,1855]" captionTargetId-0="figure@19.[189,1398,283,1832]" captionTargetId-1="figure@20.[189,1398,306,1855]" captionTargetPageId-0="19" captionTargetPageId-1="20" captionText-0="Fig. 11. Live photographs of Nagaxytes erecta Srisonchai, Enghoff &amp; Panha gen. et sp. nov. A–B. Specimens from Phuphrai Thannam Resort, Thailand (♂♂, CUMZ-pxDGT00002-3). C. Specimen from Ban Nong Bang, Thailand (♂). D. Specimen from Phuphrai Thannam Resort, Thailand (♀, CUMZpxDGT00054)." captionText-1="Fig. 12. Live photographs and habitat of Nagaxytes erecta Srisonchai, Enghoff &amp; Panha gen. et sp. nov. A. Specimen from Ban Nong Bang, Thailand (♀). B. Mating couple. C. ♂ with parasitic mites. D. Habitat." httpUri-0="https://zenodo.org/record/1467994/files/figure.png" httpUri-1="https://zenodo.org/record/1467996/files/figure.png" pageId="4" pageNumber="5">11–12</figureCitation>
,
<figureCitation id="13302A3D0D7F2261F7FAED8FFE785A8F" box="[467,498,686,712]" captionStart="Fig. 17" captionStartId="28.[189,232,1926,1952]" captionTargetBox="[193,1397,304,1847]" captionTargetId="figure@28.[189,1399,300,1849]" captionTargetPageId="28" captionText="Fig. 17. Live photographs and habitat of Nagaxytes gracilis Srisonchai, Enghoff &amp; Panha gen. et sp. nov. (specimens from Daowadueng Cave, Thailand). A. ♂, paratype (CUMZ-pxDGT00091). B. ♀, paratype (CUMZ-pxDGT00098). C. Habitat." httpUri="https://zenodo.org/record/1468006/files/figure.png" pageId="4" pageNumber="5">17</figureCitation>
,
<figureCitation id="13302A3D0D7F2261F7D7ED8FFD955A8F" box="[510,543,686,712]" captionStart="Fig. 22" captionStartId="35.[189,232,1890,1916]" captionTargetBox="[191,1395,316,1855]" captionTargetId="figure@35.[187,1397,306,1855]" captionTargetPageId="35" captionText="Fig. 22. Live photographs and habitat of Nagaxytes spatula Srisonchai, Enghoff &amp; Panha gen. et sp. nov. (specimens from Chao Por Phawo Shrine, Thailand). A. ♂, paratype (CUMZ-pxDGT00120). B. ♀, paratype (CUMZ-pxDGT00147). C. ♂, paratype (CUMZ-pxDGT00146), newly moulted adult. D. Habitat." httpUri="https://zenodo.org/record/1468016/files/figure.png" pageId="4" pageNumber="5">22</figureCitation>
). Specimens in life with brown or reddish brown colour or pinkish brown (possibly aposematic colouration). Colour in alcohol: all specimens partly faded to pale whitish brown after one year’s preservation in alcohol; specimens kept in darkness faded more slowly.
</paragraph>
<paragraph id="8BB436B80D7F2261F694EC17FAFC5B34" blockId="4.[189,1399,822,884]" pageId="4" pageNumber="5">
ANTENNAE (
<figureCitation id="13302A3D0D7F2261F76BEC17FE685B16" box="[322,482,822,849]" captionStart="Fig. 2" captionStartId="5.[189,232,1872,1898]" captionTargetBox="[192,1396,282,1802]" captionTargetId="figure@5.[189,1398,269,1817]" captionTargetPageId="5" captionText="Fig. 2. General body characters of Nagaxytes Srisonchai, Enghoff &amp; Panha gen. nov. – N. acantherpestes (Golovatch &amp; Enghoff, 1994) gen. et comb. nov., ♂, holotype, Prachuap Khiri Khan Province, Hua Hin District (ZMUC 000101457). A. Entire body. B. Anterior body part. C. Body rings 5–8, showing sternal lobe between coxae 4 and gonopods on ring 7. D. Head and antennae." httpUri="https://zenodo.org/record/1467976/files/figure.png" pageId="4" pageNumber="5">Fig. 2A–B, D</figureCitation>
). Long and slender, covered by delicate setation, usually reaching backwards to body ring 5–7 (male) and 4–6 (female) when stretched dorsally. Antennomere 3 = 4&gt; 5 ≥ 2&gt; 6&gt; 1&gt; 7&gt; 8.
</paragraph>
<paragraph id="8BB436B80D7F2261F694ECBAFED25B9F" blockId="4.[189,1398,923,984]" pageId="4" pageNumber="5">HEAD. Delicately setose; vertex, labrum and genae delicately setose; epicranial suture conspicuous as brown stripe.</paragraph>
<caption id="DF7466300D7F2261F694E887FC345FA4" httpUri="https://zenodo.org/record/1467974/files/figure.png" pageId="4" pageNumber="5" startId="4.[189,232,1958,1984]" targetBox="[189,1399,1030,1905]" targetPageId="4">
<paragraph id="8BB436B80D7F2261F694E887FC345FA4" blockId="4.[189,1398,1958,2020]" pageId="4" pageNumber="5">
<emphasis id="B97FEAAA0D7F2261F694E887FE985F87" bold="true" box="[189,274,1958,1984]" pageId="4" pageNumber="5">Fig. 1.</emphasis>
<taxonomicName id="4C0B4D3B0D7F2261F708E886FC085F87" authority="Srisonchai, Enghoff &amp; Panha" authorityName="Srisonchai, Enghoff &amp; Panha" authorityYear="2018" box="[289,898,1958,1984]" class="Diplopoda" family="Paradoxosomatidae" genus="Nagaxytes" higherTaxonomySource="CoL" kingdom="Animalia" order="Polydesmida" pageId="4" pageNumber="5" phylum="Arthropoda" rank="species" species="gracilis" status="gen. et sp. nov.">
<emphasis id="B97FEAAA0D7F2261F708E886FD8E5F87" box="[289,516,1958,1984]" italics="true" pageId="4" pageNumber="5">Nagaxytes gracilis</emphasis>
Srisonchai, Enghoff &amp; Panha
</taxonomicName>
<taxonomicNameLabel id="A24C57D10D7F2261F5B8E886FBD05F86" box="[913,1114,1959,1985]" pageId="4" pageNumber="5" rank="species">gen. et sp. nov.</taxonomicNameLabel>
, ♂, paratype (CUMZpxDGT00095). Watercolour by R. Srisonchai and P. Kriatpraprai.
</paragraph>
</caption>
<caption id="DF7466300D7E2260F694E871FC7A5F93" httpUri="https://zenodo.org/record/1467976/files/figure.png" pageId="5" pageNumber="6" startId="5.[189,232,1872,1898]" targetBox="[192,1396,282,1802]" targetPageId="5">
<paragraph id="8BB436B80D7E2260F694E871FC7A5F93" blockId="5.[189,1398,1872,2004]" pageId="5" pageNumber="6">
<emphasis id="B97FEAAA0D7E2260F694E871FE835F2D" bold="true" box="[189,265,1872,1898]" pageId="5" pageNumber="6">Fig. 2.</emphasis>
General body characters of
<taxonomicName id="4C0B4D3B0D7E2260F465E870FB945F2D" authority="Srisonchai, Enghoff &amp; Panha" authorityName="Srisonchai, Enghoff &amp; Panha" authorityYear="2018" box="[588,1054,1872,1898]" class="Diplopoda" family="Paradoxosomatidae" genus="Nagaxytes" higherTaxonomySource="CoL" kingdom="Animalia" order="Polydesmida" pageId="5" pageNumber="6" phylum="Arthropoda" rank="genus" status="gen. nov.">
<emphasis id="B97FEAAA0D7E2260F465E870FD4C5F2D" box="[588,710,1873,1898]" italics="true" pageId="5" pageNumber="6">Nagaxytes</emphasis>
Srisonchai, Enghoff &amp; Panha
</taxonomicName>
<taxonomicNameLabel id="A24C57D10D7E2260F20AE871FB065F2D" box="[1059,1164,1872,1898]" pageId="5" pageNumber="6" rank="genus">gen. nov.</taxonomicNameLabel>
–
<taxonomicName id="4C0B4D3B0D7E2260F28FE870FD925FCA" authority="(Golovatch &amp; Enghoff, 1994)" baseAuthorityName="Golovatch &amp; Enghoff" baseAuthorityYear="1994" class="Diplopoda" family="Paradoxosomatidae" genus="Desmoxytes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Polydesmida" pageId="5" pageNumber="6" phylum="Arthropoda" rank="species" species="acantherpestes" status="gen. et comb. nov.">
<emphasis id="B97FEAAA0D7E2260F28FE870FAFC5F2D" box="[1190,1398,1872,1898]" italics="true" pageId="5" pageNumber="6">N. acantherpestes</emphasis>
(Golovatch &amp; Enghoff, 1994)
</taxonomicName>
<taxonomicNameLabel id="A24C57D10D7E2260F436E852FD7D5FCA" box="[543,759,1907,1933]" pageId="5" pageNumber="6">gen. et comb. nov.</taxonomicNameLabel>
, ♂, holotype, Prachuap Khiri Khan Province, Hua Hin District (ZMUC 000101457).
<emphasis id="B97FEAAA0D7E2260F435E8B7FDBB5FF7" bold="true" box="[540,561,1942,1968]" pageId="5" pageNumber="6">A</emphasis>
. Entire body.
<emphasis id="B97FEAAA0D7E2260F4FDE8B7FD625FF7" bold="true" box="[724,744,1942,1968]" pageId="5" pageNumber="6">B</emphasis>
. Anterior body part.
<emphasis id="B97FEAAA0D7E2260F5F3E8B7FC655FF7" bold="true" box="[986,1007,1942,1968]" pageId="5" pageNumber="6">C</emphasis>
. Body rings 5–8, showing sternal lobe between coxae 4 and gonopods on ring 7.
<emphasis id="B97FEAAA0D7E2260F4CCE89BFD705F93" bold="true" box="[741,762,1978,2004]" pageId="5" pageNumber="6">D</emphasis>
. Head and antennae.
</paragraph>
</caption>
<caption id="DF7466300D7D2263F694E830FAB95F9B" httpUri="https://zenodo.org/record/1467978/files/figure.png" pageId="6" pageNumber="7" startId="6.[189,231,1809,1835]" targetBox="[189,1398,264,1781]" targetPageId="6">
<paragraph id="8BB436B80D7D2263F694E830FAB95F9B" blockId="6.[189,1399,1809,2012]" pageId="6" pageNumber="7">
<emphasis id="B97FEAAA0D7D2263F694E830FE8D5F6C" bold="true" box="[189,263,1809,1835]" pageId="6" pageNumber="7">Fig. 3.</emphasis>
General body characters of
<taxonomicName id="4C0B4D3B0D7D2263F46DE833FB845F6C" authority="Srisonchai, Enghoff &amp; Panha" authorityName="Srisonchai, Enghoff &amp; Panha" authorityYear="2018" box="[580,1038,1809,1836]" class="Diplopoda" family="Paradoxosomatidae" genus="Nagaxytes" higherTaxonomySource="CoL" kingdom="Animalia" order="Polydesmida" pageId="6" pageNumber="7" phylum="Arthropoda" rank="genus" status="gen. nov.">
<emphasis id="B97FEAAA0D7D2263F46DE833FD375F6C" box="[580,701,1810,1835]" italics="true" pageId="6" pageNumber="7">Nagaxytes</emphasis>
Srisonchai, Enghoff &amp; Panha
</taxonomicName>
<taxonomicNameLabel id="A24C57D10D7D2263F23AE833FBF15F6B" box="[1043,1147,1810,1836]" pageId="6" pageNumber="7" rank="genus">gen. nov.</taxonomicNameLabel>
(
<taxonomicName id="4C0B4D3B0D7D2263F2A0E833FE1E5F08" authority="Srisonchai, Enghoff &amp; Panha" authorityName="Srisonchai, Enghoff &amp; Panha" authorityYear="2018" class="Diplopoda" family="Paradoxosomatidae" genus="Nagaxytes" higherTaxonomySource="CoL" kingdom="Animalia" order="Polydesmida" pageId="6" pageNumber="7" phylum="Arthropoda" rank="species" species="erecta" status="gen. et sp. nov.">
<emphasis id="B97FEAAA0D7D2263F2A0E833FB7A5F6C" box="[1161,1264,1810,1835]" italics="true" pageId="6" pageNumber="7">N. erecta</emphasis>
Srisonchai, Enghoff &amp; Panha
</taxonomicName>
<taxonomicNameLabel id="A24C57D10D7D2263F788E814FDE95F08" box="[417,611,1845,1871]" pageId="6" pageNumber="7" rank="species">gen. et sp. nov.</taxonomicNameLabel>
, specimen from Prang Ka Sri Temple, Thailand), SEM images.
<emphasis id="B97FEAAA0D7D2263F694E879FF7E5F35" bold="true" box="[189,244,1880,1906]" pageId="6" pageNumber="7">A–B</emphasis>
. Anterior body part (arrowheads point to pleurosternal
<taxonomicName id="4C0B4D3B0D7D2263F5A2E879FC6E5F34" authorityName="" authorityYear="1992" box="[907,996,1880,1907]" class="Insecta" family="Eccoptarthridae" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="7" phylum="Arthropoda" rank="subFamily" subFamily="Carinae">carinae</taxonomicName>
).
<emphasis id="B97FEAAA0D7D2263F5D3E879FB855F35" bold="true" box="[1018,1039,1880,1906]" pageId="6" pageNumber="7">C</emphasis>
. Collum (arrowheads point to the setae/tubercles of the anterior and intermediate rows).
<emphasis id="B97FEAAA0D7D2263F554E85AFC185FD2" bold="true" box="[893,914,1915,1941]" pageId="6" pageNumber="7">D</emphasis>
. Body rings 9–10.
<emphasis id="B97FEAAA0D7D2263F2A9E85AFB195FD2" bold="true" box="[1152,1171,1915,1941]" pageId="6" pageNumber="7">E</emphasis>
. Body rings 8–10.
<emphasis id="B97FEAAA0D7D2263F694E8BFFF455FFF" bold="true" box="[189,207,1950,1976]" pageId="6" pageNumber="7">F</emphasis>
. Telson.
<emphasis id="B97FEAAA0D7D2263F71EE8BFFEC45FFF" bold="true" box="[311,334,1950,1976]" pageId="6" pageNumber="7">G</emphasis>
. Posteriormost rings and telson.
<emphasis id="B97FEAAA0D7D2263F4E1E8BFFD555FFF" bold="true" box="[712,735,1950,1976]" pageId="6" pageNumber="7">H</emphasis>
. Body ring 10.
<emphasis id="B97FEAAA0D7D2263F5BAE8BFFC145FFF" bold="true" box="[915,926,1950,1976]" pageId="6" pageNumber="7">I</emphasis>
. Mouth parts, ventral view.
<emphasis id="B97FEAAA0D7D2263F2CBE8BFFB7A5FFF" bold="true" box="[1250,1264,1950,1976]" pageId="6" pageNumber="7">J</emphasis>
. Gonopods
<emphasis id="B97FEAAA0D7D2263F694E8E3FE865F9C" box="[189,268,1986,2011]" italics="true" pageId="6" pageNumber="7">in situ.</emphasis>
<emphasis id="B97FEAAA0D7D2263F73AE8E3FEA05F9B" bold="true" box="[275,298,1986,2012]" pageId="6" pageNumber="7">K</emphasis>
. Sternal lobe between male coxae 4.
<emphasis id="B97FEAAA0D7D2263F4F3E8E3FD675F9B" bold="true" box="[730,749,1986,2012]" pageId="6" pageNumber="7">L</emphasis>
,
<emphasis id="B97FEAAA0D7D2263F4D2E8E3FC9A5F9B" bold="true" box="[763,784,1986,2012]" pageId="6" pageNumber="7">N</emphasis>
. Tip of epiproct.
<emphasis id="B97FEAAA0D7D2263F5F3E8E3FC7C5F9B" bold="true" box="[986,1014,1986,2012]" pageId="6" pageNumber="7">M</emphasis>
. Tarsus and claw of leg 13.
</paragraph>
</caption>
<paragraph id="8BB436B80D7C2262F694EE2AFDF459C8" blockId="7.[189,1399,267,399]" pageId="7" pageNumber="8">
COLLUM (
<figureCitation id="13302A3D0D7C2262F706EE2AFE3D5962" box="[303,439,267,294]" captionStart="Fig. 3" captionStartId="6.[189,231,1809,1835]" captionTargetBox="[189,1398,264,1781]" captionTargetId="figure@6.[189,1398,264,1781]" captionTargetPageId="6" captionText="Fig. 3. General body characters of Nagaxytes Srisonchai, Enghoff &amp; Panha gen. nov. (N. erecta Srisonchai, Enghoff &amp; Panha gen. et sp. nov., specimen from Prang Ka Sri Temple, Thailand), SEM images. A–B. Anterior body part (arrowheads point to pleurosternal carinae). C. Collum (arrowheads point to the setae/tubercles of the anterior and intermediate rows). D. Body rings 9–10. E. Body rings 8–10. F. Telson. G. Posteriormost rings and telson. H. Body ring 10. I. Mouth parts, ventral view. J. Gonopods in situ. K. Sternal lobe between male coxae 4. L, N. Tip of epiproct. M. Tarsus and claw of leg 13." httpUri="https://zenodo.org/record/1467978/files/figure.png" pageId="7" pageNumber="8">Fig. 3A, C</figureCitation>
). With three transverse rows of setae/tubercles and spines; 3+3 setae/tubercles (anterior row), 1+1 setae/tubercles (intermediate row) and 2+2 spines (posterior row); lateral spines of posterior row very long. Paraterga of collum wing-like, usually elevated at ca. 25°–40°, with two conspicuous notches at lateral margin.
</paragraph>
<paragraph id="8BB436B80D7C2262F694EE96FDA15A7D" blockId="7.[189,1399,438,570]" pageId="7" pageNumber="8">
TEGUMENT (
<figureCitation id="13302A3D0D7C2262F760EE97FE5A5997" box="[329,464,438,465]" captionStart="Fig. 3" captionStartId="6.[189,231,1809,1835]" captionTargetBox="[189,1398,264,1781]" captionTargetId="figure@6.[189,1398,264,1781]" captionTargetPageId="6" captionText="Fig. 3. General body characters of Nagaxytes Srisonchai, Enghoff &amp; Panha gen. nov. (N. erecta Srisonchai, Enghoff &amp; Panha gen. et sp. nov., specimen from Prang Ka Sri Temple, Thailand), SEM images. A–B. Anterior body part (arrowheads point to pleurosternal carinae). C. Collum (arrowheads point to the setae/tubercles of the anterior and intermediate rows). D. Body rings 9–10. E. Body rings 8–10. F. Telson. G. Posteriormost rings and telson. H. Body ring 10. I. Mouth parts, ventral view. J. Gonopods in situ. K. Sternal lobe between male coxae 4. L, N. Tip of epiproct. M. Tarsus and claw of leg 13." httpUri="https://zenodo.org/record/1467978/files/figure.png" pageId="7" pageNumber="8">Fig. 3A–G</figureCitation>
). Often dull, sometimes quite shiny; collum, metaterga (except anterior part of metaterga smooth) and surface below paraterga finely microgranulate; prozona finely shagreened; paraterga, sterna and epiproct smooth. Stricture between prozona and metazona wide, usually quite shallow, sometimes quite deep.
</paragraph>
<paragraph id="8BB436B80D7C2262F694ED43FCDA5B4F" blockId="7.[189,1399,609,777]" pageId="7" pageNumber="8">
METATERGA (
<figureCitation id="13302A3D0D7C2262F761ED40FE225A3B" box="[328,424,609,636]" captionStart="Fig. 2" captionStartId="5.[189,232,1872,1898]" captionTargetBox="[192,1396,282,1802]" captionTargetId="figure@5.[189,1398,269,1817]" captionTargetPageId="5" captionText="Fig. 2. General body characters of Nagaxytes Srisonchai, Enghoff &amp; Panha gen. nov. – N. acantherpestes (Golovatch &amp; Enghoff, 1994) gen. et comb. nov., ♂, holotype, Prachuap Khiri Khan Province, Hua Hin District (ZMUC 000101457). A. Entire body. B. Anterior body part. C. Body rings 5–8, showing sternal lobe between coxae 4 and gonopods on ring 7. D. Head and antennae." httpUri="https://zenodo.org/record/1467976/files/figure.png" pageId="7" pageNumber="8">Figs 2A</figureCitation>
,
<figureCitation id="13302A3D0D7C2262F79CED43FDAB5A3B" box="[437,545,610,636]" captionStart="Fig. 3" captionStartId="6.[189,231,1809,1835]" captionTargetBox="[189,1398,264,1781]" captionTargetId="figure@6.[189,1398,264,1781]" captionTargetPageId="6" captionText="Fig. 3. General body characters of Nagaxytes Srisonchai, Enghoff &amp; Panha gen. nov. (N. erecta Srisonchai, Enghoff &amp; Panha gen. et sp. nov., specimen from Prang Ka Sri Temple, Thailand), SEM images. A–B. Anterior body part (arrowheads point to pleurosternal carinae). C. Collum (arrowheads point to the setae/tubercles of the anterior and intermediate rows). D. Body rings 9–10. E. Body rings 8–10. F. Telson. G. Posteriormost rings and telson. H. Body ring 10. I. Mouth parts, ventral view. J. Gonopods in situ. K. Sternal lobe between male coxae 4. L, N. Tip of epiproct. M. Tarsus and claw of leg 13." httpUri="https://zenodo.org/record/1467978/files/figure.png" pageId="7" pageNumber="8">3A, D, G</figureCitation>
). With one or two transverse rows of setae/tubercles and spines; usually with 2+2 setae/tubercles/cones/spines in anterior row and 2+2 spines in posterior row (sometimes setae/ tubercles/cones/spines in anterior row poorly developed or absent); lateral spines of posterior row longer and larger than mesal ones. Suture (transverse sulcus) on metaterga quite deep, conspicuous on body ring
<quantity id="4CF39B5D0D7C2262F6DDEDCFFEC75B4F" box="[244,333,750,777]" metricMagnitude="-1" metricUnit="m" metricValue="2.794" metricValueMax="4.318" metricValueMin="1.27" pageId="7" pageNumber="8" unit="in" value="11.0" valueMax="17.0" valueMin="5.0">5–17 in</quantity>
all species. Mid-dorsal (axial) line missing.
</paragraph>
<paragraph id="8BB436B80D7C2262F694EC11FDCD5B2A" blockId="7.[189,1398,816,877]" pageId="7" pageNumber="8">
PLEUROSTERNAL CARINAE (
<figureCitation id="13302A3D0D7C2262F7C8EC11FDB65B0D" box="[481,572,816,842]" captionStart="Fig. 3" captionStartId="6.[189,231,1809,1835]" captionTargetBox="[189,1398,264,1781]" captionTargetId="figure@6.[189,1398,264,1781]" captionTargetPageId="6" captionText="Fig. 3. General body characters of Nagaxytes Srisonchai, Enghoff &amp; Panha gen. nov. (N. erecta Srisonchai, Enghoff &amp; Panha gen. et sp. nov., specimen from Prang Ka Sri Temple, Thailand), SEM images. A–B. Anterior body part (arrowheads point to pleurosternal carinae). C. Collum (arrowheads point to the setae/tubercles of the anterior and intermediate rows). D. Body rings 9–10. E. Body rings 8–10. F. Telson. G. Posteriormost rings and telson. H. Body ring 10. I. Mouth parts, ventral view. J. Gonopods in situ. K. Sternal lobe between male coxae 4. L, N. Tip of epiproct. M. Tarsus and claw of leg 13." httpUri="https://zenodo.org/record/1467978/files/figure.png" pageId="7" pageNumber="8">Fig. 3B</figureCitation>
). Forming a complete crest on ring 2, smalls ridges on ring 3 and/or 4, missing on remaining body rings.
</paragraph>
<paragraph id="8BB436B80D7C2262F694ECB4FAFA5C19" blockId="7.[189,1399,916,1118]" pageId="7" pageNumber="8">
PARATERGA (
<figureCitation id="13302A3D0D7C2262F766ECB5FDED5BE9" box="[335,615,916,943]" captionStart="Fig. 3" captionStartId="6.[189,231,1809,1835]" captionTargetBox="[189,1398,264,1781]" captionTargetId="figure@6.[189,1398,264,1781]" captionTargetPageId="6" captionText="Fig. 3. General body characters of Nagaxytes Srisonchai, Enghoff &amp; Panha gen. nov. (N. erecta Srisonchai, Enghoff &amp; Panha gen. et sp. nov., specimen from Prang Ka Sri Temple, Thailand), SEM images. A–B. Anterior body part (arrowheads point to pleurosternal carinae). C. Collum (arrowheads point to the setae/tubercles of the anterior and intermediate rows). D. Body rings 9–10. E. Body rings 8–10. F. Telson. G. Posteriormost rings and telson. H. Body ring 10. I. Mouth parts, ventral view. J. Gonopods in situ. K. Sternal lobe between male coxae 4. L, N. Tip of epiproct. M. Tarsus and claw of leg 13." httpUri="https://zenodo.org/record/1467978/files/figure.png" pageId="7" pageNumber="8">Fig. 3A–B, D–E, G–H</figureCitation>
). Subspiniform, long, elevated at ca. 50°–70° (male) or 45°–70° (female), directed caudolaterad on rings 2–17 or 2–18, directed increasingly caudad on ring 19 or 18–19. Callus and shoulder poorly developed. Anterior margin with two distinct notches; on body rings 9, 10, 12, 13, 15–18 a denticle usually present at lateral margin near tip (denticle absent in some rings). Degree of elevation of paraterga in male usually higher than in female. Posterior edge concave; posterior angle pointed and sharp. Ozopore visible in lateral and dorsal views, round, small, somewhat inconspicuous.
</paragraph>
<paragraph id="8BB436B80D7C2262F694EBA7FEA55D34" blockId="7.[189,1399,1158,1395]" pageId="7" pageNumber="8">
TELSON (
<figureCitation id="13302A3D0D7C2262F70AEBA7FE6D5CE7" box="[291,487,1158,1184]" captionStart="Fig. 3" captionStartId="6.[189,231,1809,1835]" captionTargetBox="[189,1398,264,1781]" captionTargetId="figure@6.[189,1398,264,1781]" captionTargetPageId="6" captionText="Fig. 3. General body characters of Nagaxytes Srisonchai, Enghoff &amp; Panha gen. nov. (N. erecta Srisonchai, Enghoff &amp; Panha gen. et sp. nov., specimen from Prang Ka Sri Temple, Thailand), SEM images. A–B. Anterior body part (arrowheads point to pleurosternal carinae). C. Collum (arrowheads point to the setae/tubercles of the anterior and intermediate rows). D. Body rings 9–10. E. Body rings 8–10. F. Telson. G. Posteriormost rings and telson. H. Body ring 10. I. Mouth parts, ventral view. J. Gonopods in situ. K. Sternal lobe between male coxae 4. L, N. Tip of epiproct. M. Tarsus and claw of leg 13." httpUri="https://zenodo.org/record/1467978/files/figure.png" pageId="7" pageNumber="8">Fig. 3F–G, L, N</figureCitation>
). Epiproct quite short, often flattened dorsoventrally, tip usually subtruncate, sometimes emarginate; lateral setiferous tubercles conspicuous, digitiform; apical tubercles mostly conspicuous, sometimes inconspicuous; setiferous tubercles beyond lateral setiferous tubercles long, digitiform; epiproct apically with four spinnerets at the corners of a square, not in a depression, anterior pair close to apical tubercles (
<figureCitation id="13302A3D0D7C2262F434EA33FDF35D6A" box="[541,633,1298,1325]" captionStart="Fig. 3" captionStartId="6.[189,231,1809,1835]" captionTargetBox="[189,1398,264,1781]" captionTargetId="figure@6.[189,1398,264,1781]" captionTargetPageId="6" captionText="Fig. 3. General body characters of Nagaxytes Srisonchai, Enghoff &amp; Panha gen. nov. (N. erecta Srisonchai, Enghoff &amp; Panha gen. et sp. nov., specimen from Prang Ka Sri Temple, Thailand), SEM images. A–B. Anterior body part (arrowheads point to pleurosternal carinae). C. Collum (arrowheads point to the setae/tubercles of the anterior and intermediate rows). D. Body rings 9–10. E. Body rings 8–10. F. Telson. G. Posteriormost rings and telson. H. Body ring 10. I. Mouth parts, ventral view. J. Gonopods in situ. K. Sternal lobe between male coxae 4. L, N. Tip of epiproct. M. Tarsus and claw of leg 13." httpUri="https://zenodo.org/record/1467978/files/figure.png" pageId="7" pageNumber="8">Fig. 3L</figureCitation>
). Paraprocts convex. Hypoproct subtriangular or subtrapeziform; caudal margin often subtriangular, sometimes round, with two conspicuous or inconspicuous setiferous tubercles.
</paragraph>
<paragraph id="8BB436B80D7C2262F694EABBFD685E59" blockId="7.[189,1399,1434,1566]" pageId="7" pageNumber="8">
STERNA (
<figureCitation id="13302A3D0D7C2262F70FEABBFE025DF3" box="[294,392,1434,1460]" captionStart="Fig. 2" captionStartId="5.[189,232,1872,1898]" captionTargetBox="[192,1396,282,1802]" captionTargetId="figure@5.[189,1398,269,1817]" captionTargetPageId="5" captionText="Fig. 2. General body characters of Nagaxytes Srisonchai, Enghoff &amp; Panha gen. nov. – N. acantherpestes (Golovatch &amp; Enghoff, 1994) gen. et comb. nov., ♂, holotype, Prachuap Khiri Khan Province, Hua Hin District (ZMUC 000101457). A. Entire body. B. Anterior body part. C. Body rings 5–8, showing sternal lobe between coxae 4 and gonopods on ring 7. D. Head and antennae." httpUri="https://zenodo.org/record/1467976/files/figure.png" pageId="7" pageNumber="8">Figs 2C</figureCitation>
,
<figureCitation id="13302A3D0D7C2262F7BEEABBFE355DF3" box="[407,447,1434,1460]" captionStart="Fig. 3" captionStartId="6.[189,231,1809,1835]" captionTargetBox="[189,1398,264,1781]" captionTargetId="figure@6.[189,1398,264,1781]" captionTargetPageId="6" captionText="Fig. 3. General body characters of Nagaxytes Srisonchai, Enghoff &amp; Panha gen. nov. (N. erecta Srisonchai, Enghoff &amp; Panha gen. et sp. nov., specimen from Prang Ka Sri Temple, Thailand), SEM images. A–B. Anterior body part (arrowheads point to pleurosternal carinae). C. Collum (arrowheads point to the setae/tubercles of the anterior and intermediate rows). D. Body rings 9–10. E. Body rings 8–10. F. Telson. G. Posteriormost rings and telson. H. Body ring 10. I. Mouth parts, ventral view. J. Gonopods in situ. K. Sternal lobe between male coxae 4. L, N. Tip of epiproct. M. Tarsus and claw of leg 13." httpUri="https://zenodo.org/record/1467978/files/figure.png" pageId="7" pageNumber="8">3K</figureCitation>
). Sparsely setose; cross-impressions somewhat deep, sometimes quite shallow. Sternal lobe between male coxae 4 modified; usually subtrapeziform, sometimes incompletely bilobed, varying between species; tip slightly emarginate, sometimes deeply emarginate or even incompletely bilobed; with two pores seen in posterior view.
</paragraph>
<paragraph id="8BB436B80D7C2262F694E967FA9A5EC5" blockId="7.[189,1399,1605,1667]" pageId="7" pageNumber="8">
LEGS (
<figureCitation id="13302A3D0D7C2262F722E964FE1A5E18" box="[267,400,1605,1632]" captionStart="Fig. 2" captionStartId="5.[189,232,1872,1898]" captionTargetBox="[192,1396,282,1802]" captionTargetId="figure@5.[189,1398,269,1817]" captionTargetPageId="5" captionText="Fig. 2. General body characters of Nagaxytes Srisonchai, Enghoff &amp; Panha gen. nov. – N. acantherpestes (Golovatch &amp; Enghoff, 1994) gen. et comb. nov., ♂, holotype, Prachuap Khiri Khan Province, Hua Hin District (ZMUC 000101457). A. Entire body. B. Anterior body part. C. Body rings 5–8, showing sternal lobe between coxae 4 and gonopods on ring 7. D. Head and antennae." httpUri="https://zenodo.org/record/1467976/files/figure.png" pageId="7" pageNumber="8">Fig. 2A–C</figureCitation>
). Very long and slender. Relative lengths of podomeres: femur&gt; tarsus&gt; tibia&gt; prefemur = postfemur&gt; coxa&gt; claw. Male femora 5 and 6 without modification in all species.
</paragraph>
<paragraph id="8BB436B80D7C2262F694E98BFBA95FD0" blockId="7.[189,1399,1706,1943]" pageId="7" pageNumber="8">
GONOPODS (
<figureCitation id="13302A3D0D7C2262F76FE98BFE385E82" box="[326,434,1706,1733]" captionStart-0="Fig. 4" captionStart-1="Fig. 5" captionStartId-0="8.[189,232,1919,1945]" captionStartId-1="9.[190,233,1883,1909]" captionTargetBox-0="[196,1381,311,1854]" captionTargetBox-1="[189,1398,279,1828]" captionTargetId-0="figure@8.[189,1398,305,1854]" captionTargetId-1="figure@9.[189,1398,279,1828]" captionTargetPageId-0="8" captionTargetPageId-1="9" captionText-0="Fig. 4. Diagrammatic drawing of right gonopod of Nagaxytes Srisonchai, Enghoff &amp; Panha gen. nov. – N. gracilis Srisonchai, Enghoff &amp; Panha gen. et sp. nov., specimen from Wat Vimooddhi Wanaram, Thailand. A. Mesal view. B. Dorsal view. C. Ventral view." captionText-1="Fig. 5. SEM images of right gonopod of Nagaxytes Srisonchai, Enghoff &amp; Panha gen. nov. – N. gracilis Srisonchai, Enghoff &amp; Panha gen. et sp. nov., specimen from Wat Vimooddhi Wanaram, Thailand. A. Lateral view. B. Mesal view. C. Dorsal view. D. Ventral view. Colours: red = lamina medialis, green = lamina lateralis, yellow = solenomere, purple = seminal groove." httpUri-0="https://zenodo.org/record/1467980/files/figure.png" httpUri-1="https://zenodo.org/record/1467982/files/figure.png" pageId="7" pageNumber="8">Figs 4–5</figureCitation>
). Coxa longer than prefemoral part, with a distoanterior group of setae. Cannula long or short. Telopodite straight. Prefemoral part usually shorter than femur, sometimes subequal in length to femur. Femur usually long, sometimes quite short. Seminal groove running entirely on mesal surface of femur. Mesal sulcus and lateral sulcus conspicuous. Postfemoral part conspicuous, shorter than femur. Solenophore well-developed: lamina lateralis variously modified; swollen, sometimes long digitiform/lamella-like and projecting, sometimes with spatula-like lobe: lamina medialis longer than lamina lateralis, snake-like, tip curving down. Solenomere relatively long.
</paragraph>
</subSubSection>
<caption id="DF7466300D73226DF694E85EFCE25F98" httpUri="https://zenodo.org/record/1467980/files/figure.png" pageId="8" pageNumber="9" startId="8.[189,232,1919,1945]" targetBox="[196,1381,311,1854]" targetPageId="8">
<paragraph id="8BB436B80D73226DF694E85EFCE25F98" blockId="8.[189,1399,1919,2016]" pageId="8" pageNumber="9">
<emphasis id="B97FEAAA0D73226DF694E85EFE865FDE" bold="true" box="[189,268,1919,1945]" pageId="8" pageNumber="9">Fig. 4.</emphasis>
Diagrammatic drawing of right gonopod of
<taxonomicName id="4C0B4D3B0D73226DF50BE8A1FA885FDE" authority="Srisonchai, Enghoff &amp; Panha" authorityName="Srisonchai, Enghoff &amp; Panha" authorityYear="2018" box="[802,1282,1919,1945]" class="Diplopoda" family="Paradoxosomatidae" genus="Nagaxytes" higherTaxonomySource="CoL" kingdom="Animalia" order="Polydesmida" pageId="8" pageNumber="9" phylum="Arthropoda" rank="genus" status="gen. nov.">
<emphasis id="B97FEAAA0D73226DF50BE8A1FC165FDE" box="[802,924,1920,1945]" italics="true" pageId="8" pageNumber="9">Nagaxytes</emphasis>
Srisonchai, Enghoff &amp; Panha
</taxonomicName>
<taxonomicNameLabel id="A24C57D10D73226DF323E8A1FAFD5FDD" box="[1290,1399,1920,1946]" pageId="8" pageNumber="9" rank="genus">gen. nov.</taxonomicNameLabel>
–
<taxonomicName id="4C0B4D3B0D73226DF6FDE882FD3D5FFB" authority="Srisonchai, Enghoff &amp; Panha" authorityName="Srisonchai, Enghoff &amp; Panha" authorityYear="2018" box="[212,695,1954,1980]" class="Diplopoda" family="Paradoxosomatidae" genus="Nagaxytes" higherTaxonomySource="CoL" kingdom="Animalia" order="Polydesmida" pageId="8" pageNumber="9" phylum="Arthropoda" rank="species" species="gracilis" status="gen. et sp. nov.">
<emphasis id="B97FEAAA0D73226DF6FDE882FED85FFB" box="[212,338,1954,1980]" italics="true" pageId="8" pageNumber="9">N. gracilis</emphasis>
Srisonchai, Enghoff &amp; Panha
</taxonomicName>
<taxonomicNameLabel id="A24C57D10D73226DF496E882FCFC5FFA" box="[703,886,1955,1981]" pageId="8" pageNumber="9" rank="species">gen. et sp. nov.</taxonomicNameLabel>
, specimen from Wat Vimooddhi Wanaram, Thailand.
<emphasis id="B97FEAAA0D73226DF71DE8E4FEC35F98" bold="true" box="[308,329,1989,2015]" pageId="8" pageNumber="9">A</emphasis>
. Mesal view.
<emphasis id="B97FEAAA0D73226DF7C4E8E7FD8B5FA7" bold="true" box="[493,513,1990,2016]" pageId="8" pageNumber="9">B</emphasis>
. Dorsal view.
<emphasis id="B97FEAAA0D73226DF483E8E4FD355F98" bold="true" box="[682,703,1989,2015]" pageId="8" pageNumber="9">C</emphasis>
. Ventral view.
</paragraph>
</caption>
<caption id="DF7466300D72226CF697E87AFB9C5F98" httpUri="https://zenodo.org/record/1467982/files/figure.png" pageId="9" pageNumber="10" startId="9.[190,233,1883,1909]" targetBox="[189,1398,279,1828]" targetPageId="9">
<paragraph id="8BB436B80D72226CF697E87AFB9C5F98" blockId="9.[190,1400,1883,2015]" pageId="9" pageNumber="10">
<emphasis id="B97FEAAA0D72226CF697E87AFE805F32" bold="true" box="[190,266,1883,1909]" pageId="9" pageNumber="10">Fig. 5.</emphasis>
SEM images of right gonopod of
<taxonomicName id="4C0B4D3B0D72226CF4BEE87DFBE55F32" authority="Srisonchai, Enghoff &amp; Panha" authorityName="Srisonchai, Enghoff &amp; Panha" authorityYear="2018" box="[663,1135,1883,1909]" class="Diplopoda" family="Paradoxosomatidae" genus="Nagaxytes" higherTaxonomySource="CoL" kingdom="Animalia" order="Polydesmida" pageId="9" pageNumber="10" phylum="Arthropoda" rank="genus" status="gen. nov.">
<emphasis id="B97FEAAA0D72226CF4BEE87DFC995F32" box="[663,787,1884,1909]" italics="true" pageId="9" pageNumber="10">Nagaxytes</emphasis>
Srisonchai, Enghoff &amp; Panha
</taxonomicName>
<taxonomicNameLabel id="A24C57D10D72226CF25FE87DFB6B5F31" box="[1142,1249,1884,1910]" pageId="9" pageNumber="10" rank="genus">gen. nov.</taxonomicNameLabel>
–
<taxonomicName id="4C0B4D3B0D72226CF2D2E87DFDA85FDF" authority="Srisonchai, Enghoff &amp; Panha" authorityName="Srisonchai, Enghoff &amp; Panha" authorityYear="2018" class="Diplopoda" family="Paradoxosomatidae" genus="Nagaxytes" higherTaxonomySource="CoL" kingdom="Animalia" order="Polydesmida" pageId="9" pageNumber="10" phylum="Arthropoda" rank="species" species="gracilis" status="gen. et sp. nov.">
<emphasis id="B97FEAAA0D72226CF2D2E87DFAFD5F32" box="[1275,1399,1883,1909]" italics="true" pageId="9" pageNumber="10">N. gracilis</emphasis>
Srisonchai, Enghoff &amp; Panha
</taxonomicName>
<taxonomicNameLabel id="A24C57D10D72226CF407E85EFD675FDE" box="[558,749,1919,1945]" pageId="9" pageNumber="10" rank="species">gen. et sp. nov.</taxonomicNameLabel>
, specimen from Wat Vimooddhi Wanaram, Thailand.
<emphasis id="B97FEAAA0D72226CF697E880FF595FFC" bold="true" box="[190,211,1953,1979]" pageId="9" pageNumber="10">A</emphasis>
. Lateral view.
<emphasis id="B97FEAAA0D72226CF7A6E883FE295FFB" bold="true" box="[399,419,1954,1980]" pageId="9" pageNumber="10">B</emphasis>
. Mesal view.
<emphasis id="B97FEAAA0D72226CF47AE880FDE25FFC" bold="true" box="[595,616,1953,1979]" pageId="9" pageNumber="10">C</emphasis>
. Dorsal view.
<emphasis id="B97FEAAA0D72226CF536E883FCBE5FFB" bold="true" box="[799,820,1954,1980]" pageId="9" pageNumber="10">D</emphasis>
. Ventral view. Colours: red = lamina medialis, green = lamina lateralis, yellow = solenomere, purple = seminal groove.
</paragraph>
</caption>
<subSubSection id="C31165330D71226FF694EE2AFCA45A4D" pageId="10" pageNumber="11" type="distribution">
<paragraph id="8BB436B80D71226FF694EE2AFE645962" blockId="10.[189,494,267,293]" box="[189,494,267,293]" pageId="10" pageNumber="11">
<heading id="D0FC81D40D71226FF694EE2AFE645962" bold="true" box="[189,494,267,293]" fontSize="11" level="3" pageId="10" pageNumber="11" reason="3">
<emphasis id="B97FEAAA0D71226FF694EE2AFE645962" bold="true" box="[189,494,267,293]" pageId="10" pageNumber="11">Distribution and habitat</emphasis>
</heading>
</paragraph>
<paragraph id="8BB436B80D71226FF694EE1BFCA45A4D" blockId="10.[189,1399,314,522]" pageId="10" pageNumber="11">
This genus is known only from the western part of
<collectingCountry id="F31C76280D71226FF4D7EE1BFCE15913" box="[766,875,314,340]" name="Thailand" pageId="10" pageNumber="11">Thailand</collectingCountry>
:
<collectingRegion id="49CFF85A0D71226FF55DEE1BFB5C5913" box="[884,1238,314,340]" country="Thailand" name="Prachuap Khiri Khan" pageId="10" pageNumber="11">Prachuap Khiri Khan Province</collectingRegion>
(Hua Hin and Kui Buri Districts),
<collectingRegion id="49CFF85A0D71226FF781EE7EFD36593E" box="[424,700,351,377]" country="Thailand" name="Kanchanaburi" pageId="10" pageNumber="11">Kanchanaburi Province</collectingRegion>
(Sai Yok and Thong Pha Phum Districts) and
<collectingRegion id="49CFF85A0D71226FF2F2EE7EFAFC593E" box="[1243,1398,351,377]" country="Thailand" name="Tak" pageId="10" pageNumber="11">Tak Province</collectingRegion>
(Mae Sot and Umphang Districts) (
<figureCitation id="13302A3D0D71226FF47CEEA2FD2659DA" box="[597,684,387,413]" captionStart="Fig. 27" captionStartId="41.[189,232,1941,1967]" captionTargetBox="[189,1399,302,1851]" captionTargetId="figure@41.[189,1399,302,1851]" captionTargetPageId="41" captionText="Fig. 27. Known distribution of all species of Nagaxytes Srisonchai, Enghoff &amp; Panha gen. nov. (black symbol = type locality, white symbol = other localities)." httpUri="https://zenodo.org/record/1468026/files/figure.png" pageId="10" pageNumber="11">Fig. 27</figureCitation>
). All four species seem to be local endemics, highly restricted to limestone habitats, and all have narrow distribution ranges. The specimens were usually found on humid rocks, rock walls, leaf litter or surface roots of trees. Some species can be found syntopically in the same habitat with other dragon millipede species.
</paragraph>
</subSubSection>
</treatment>
</document>
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