545 lines
61 KiB
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545 lines
61 KiB
XML
<document ID-DOI="http://dx.doi.org/10.3897/zookeys.1142.90097" ID-Pensoft-Pub="1313-2970-1142-1" ID-Pensoft-UUID="1D4BDF04F72B59B9984E00D0B4FB338D" ID-ZooBank="A1129EE50F9941CFB73AE771B66E2486" ModsDocID="1313-2970-1142-1" checkinTime="1674655519517" checkinUser="pensoft" docAuthor="Tongkerd, Piyoros, Tumpeesuwan, Sakboworn, Inkhavilay, Khamla, Prasankok, Pongpun, Jeratthitikul, Ekgachai, Panha, Somsak & Sutcharit, Chirasak" docDate="2023" docId="03122B5FADC55EFE8BC5CF4838D12522" docLanguage="en" docName="ZooKeys 1142: 1-144" docOrigin="ZooKeys 1142" docPubDate="2023-01-24" docSource="http://dx.doi.org/10.3897/zookeys.1142.90097" docTitle="Rhiostoma housei" docType="treatment" docVersion="1" id="1D4BDF04F72B59B9984E00D0B4FB338D" lastPageNumber="1" masterDocId="1D4BDF04F72B59B9984E00D0B4FB338D" masterDocTitle="Systematic revision of the snorkel snail genus Rhiostoma Benson, 1860 (Gastropoda, Caenogastropoda, Cyclophoridae) with descriptions of new species" masterLastPageNumber="144" masterPageNumber="1" pageNumber="1" updateTime="1674655519517" updateUser="pensoft">
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<mods:titleInfo>
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<mods:title>Systematic revision of the snorkel snail genus Rhiostoma Benson, 1860 (Gastropoda, Caenogastropoda, Cyclophoridae) with descriptions of new species</mods:title>
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</mods:titleInfo>
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<mods:name type="personal">
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<mods:roleTerm>Author</mods:roleTerm>
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<mods:namePart>Tongkerd, Piyoros</mods:namePart>
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<mods:affiliation>Animal Systematics Research Unit, Department of Biology, Faculty of Science, Chulalongkorn University, Bangkok 10330, Thailand</mods:affiliation>
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</mods:name>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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<mods:namePart>Tumpeesuwan, Sakboworn</mods:namePart>
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<mods:affiliation>Animal Systematics Research Unit, Department of Biology, Faculty of Science, Chulalongkorn University, Bangkok 10330, Thailand & Present address: Department of Biology, Faculty of Science, Mahasarakham University, Kantharawichai, Maha Sarakham 44150 Thailand</mods:affiliation>
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</mods:name>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Inkhavilay, Khamla</mods:namePart>
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<mods:affiliation>Department of Biology, Faculty of Natural Science, National University of Laos, P. O. Box 7322, Dongdok, Vientiane, Laos</mods:affiliation>
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</mods:name>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Prasankok, Pongpun</mods:namePart>
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<mods:affiliation>School of Biology, Institute of Science, Suranaree University of Technology, Nakhon Ratchasima 30000, Thailand</mods:affiliation>
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</mods:name>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Jeratthitikul, Ekgachai</mods:namePart>
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<mods:affiliation>Animal Systematics and Molecular Ecology Laboratory, Department of Biology, Faculty of Science, Mahidol University, Bangkok 10400, Thailand</mods:affiliation>
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</mods:name>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Panha, Somsak</mods:namePart>
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<mods:affiliation>Animal Systematics Research Unit, Department of Biology, Faculty of Science, Chulalongkorn University, Bangkok 10330, Thailand</mods:affiliation>
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<mods:nameIdentifier type="email">somsak.pan@chula.ac.th</mods:nameIdentifier>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Sutcharit, Chirasak</mods:namePart>
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<mods:affiliation>Animal Systematics Research Unit, Department of Biology, Faculty of Science, Chulalongkorn University, Bangkok 10330, Thailand</mods:affiliation>
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<mods:nameIdentifier type="email">jirasak4@yahoo.com</mods:nameIdentifier>
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<mods:title>ZooKeys</mods:title>
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<mods:date>2023</mods:date>
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<mods:number>2023-01-24</mods:number>
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<mods:detail type="volume">
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<mods:number>1142</mods:number>
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<mods:extent unit="page">
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<mods:start>1</mods:start>
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<mods:end>144</mods:end>
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<mods:url>http://dx.doi.org/10.3897/zookeys.1142.90097</mods:url>
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<mods:classification>journal article</mods:classification>
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<mods:identifier type="DOI">http://dx.doi.org/10.3897/zookeys.1142.90097</mods:identifier>
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<treatment LSID="urn:lsid:plazi:treatment:03122B5FADC55EFE8BC5CF4838D12522" httpUri="http://treatment.plazi.org/id/03122B5FADC55EFE8BC5CF4838D12522" lastPageNumber="1" pageId="0" pageNumber="1">
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<subSubSection pageId="0" pageNumber="1" type="nomenclature">
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<paragraph pageId="0" pageNumber="1">
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4.
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<taxonomicName ID-CoL="7SZX2" LSID="03122B5F-ADC5-5EFE-8BC5-CF4838D12522" authority="(Haines, 1855)" baseAuthorityName="Haines" baseAuthorityYear="1855" class="Gastropoda" family="Cyclophoridae" genus="Rhiostoma" higherTaxonomySource="treatment-meta" kingdom="Animalia" lsidName="Rhiostoma housei" order="Architaenioglossa" pageId="0" pageNumber="1" phylum="Mollusca" rank="species" species="housei">Rhiostoma housei (Haines, 1855)</taxonomicName>
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</paragraph>
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</subSubSection>
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<subSubSection pageId="0" pageNumber="1" type="description">
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<paragraph pageId="0" pageNumber="1">
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<figureCitation captionStart="Figure 17" captionStartId="F17" captionText="Figure 17. Live snails of the species group II A Rhiostoma housei specimen CUMZ 3987 from Ao Manow, Prachuap Khiri Khan B, C Rhiostoma hainesi from Khao Soi Dao, Chanthaburi B without breathing device and C with long breathing device D Rhiostoma jalorensis specimen CUMZ 3994 from Wat Tham Sue, Krabi. All figures are not to scale. Photographs: P. Tongkerd (B, C)." figureDoi="10.3897/zookeys.1142.90097.figure17" httpUri="https://binary.pensoft.net/fig/799875" pageId="0" pageNumber="1">Figs 17A</figureCitation>
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<figureCitation captionStart="Figure 18" captionStartId="F18" captionText="Figure 18. Distribution map of Rhiostoma housei - species group." figureDoi="10.3897/zookeys.1142.90097.figure18" httpUri="https://binary.pensoft.net/fig/799876" pageId="0" pageNumber="1">, 18</figureCitation>
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<figureCitation captionStart="Figure 19" captionStartId="F19" captionText="Figure 19. Shell of Rhiostoma housei A, B syntype AMNH 42923 from Siam A figured specimen in Haines (1855: figs 12, 13) anb B other specimen from the same syntype lot C specimens CUMZ 4383 from Tham Phitsadan, Chumphon D specimen CUMZ 3982 from Tham Sriwilai, Saraburi E specimen CUMZ 4384 from Pak Chong, Nakhon Ratchasrima F specimen CUMZ 4879 from Hua Hin, Prachuap Khiri Khan." figureDoi="10.3897/zookeys.1142.90097.figure19" httpUri="https://binary.pensoft.net/fig/799877" pageId="0" pageNumber="1">, 19</figureCitation>
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<figureCitation captionStart="Figure 20" captionStartId="F20" captionText="Figure 20. Drawing of general anatomy of Rhiostoma housei specimen CUMZ 3982 from Saraburi Province, Thailand. A female B male. Abbreviations: bc, bursa copulatrix;; cm, columellar muscle; ct, cephalic tentacle; dg, digestive gland; es, eyespot; f, foot; fc, faeces; int, intestine; lc, lung cavity; mc, mantle collar; od, oviduct; op, operculum; ov, ovary; p, external penis; pg, prostate gland; sg, sperm groove; sn, snout; te, testis; ut, uterus; vd, vas deferens; vg, vaginal groove." figureDoi="10.3897/zookeys.1142.90097.figure20" httpUri="https://binary.pensoft.net/fig/799878" pageId="0" pageNumber="1">, 20</figureCitation>
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<figureCitation captionStart="Figure 32" captionStartId="F32" captionText="Figure 32. Radula morphology A Rhiostoma housei specimen CUMZ 3982 from Wat Tham Srivilai, Saraburi B Rhiostoma hainesi specimen from Chanthaburi C Rhiostoma jalorensis specimen CUMZ 3994 from Wat Tham Sua, Krabi D Rhiostoma ebenozostera sp. nov. paratype CUMZ 4704 from Tham Pha Poo, Loei E Rhiostoma lannaense sp. nov. paratype CUMZ 4701 from Chiang Dao, Chiang Mai F Rhiostoma tigrina sp. nov. paratype CUMZ 4497 / 2 from Mae Sai, Chiang Rai." figureDoi="10.3897/zookeys.1142.90097.figure32" httpUri="https://binary.pensoft.net/fig/799890" pageId="0" pageNumber="1">, 32A</figureCitation>
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</paragraph>
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</subSubSection>
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<subSubSection pageId="0" pageNumber="1" type="reference_group">
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<paragraph pageId="0" pageNumber="1">
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<taxonomicName authorityName="Haines" authorityYear="1855" class="Gastropoda" family="Cyclophoridae" genus="Cyclostoma" higherTaxonomySource="treatment-meta" kingdom="Animalia" lsidName="Cyclostoma housei" order="Architaenioglossa" pageId="0" pageNumber="1" phylum="Mollusca" rank="species" species="housei">Cyclostoma housei</taxonomicName>
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Haines, 1855: 157, pl. 5, figs 12-15. Type locality: Siam [= Thailand].
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<bibRefCitation DOI="https://doi.org/10.1206/0003-0090(2001)262<0001:CORTSI>2.0.CO;2" author="Boyko, CB" journalOrPublisher="Bulletin of the American Museum of Natural History" pageId="0" pageNumber="1" pagination="1 - 158" refId="B24" refString="Boyko, CB, Cordeiro, JR, 2001. Catalog of recent type specimens in the Division of Invertebrate Zoology, American Museum of Natural History. V. Mollusca, Part 2 (Class Gastropoda [exclusive of Opisthobranchia and Pulmonata], with supplements to Gastropoda [Opisthobranchia], and Bivalvia). Bulletin of the American Museum of Natural History 262: 1 - 158, DOI: https://doi.org/10.1206/0003-0090(2001)262<0001:CORTSI>2.0.CO;2" title="Catalog of recent type specimens in the Division of Invertebrate Zoology, American Museum of Natural History. V. Mollusca, Part 2 (Class Gastropoda [exclusive of Opisthobranchia and Pulmonata], with supplements to Gastropoda [Opisthobranchia], and Bivalvia)." url="https://doi.org/10.1206/0003-0090(2001)262<0001:CORTSI>2.0.CO;2" volume="262" year="2001">Boyko and Cordeiro 2001</bibRefCitation>
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: 52.
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</paragraph>
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<paragraph pageId="0" pageNumber="1">
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<taxonomicName class="Gastropoda" family="Cyclophoridae" genus="Pterocyclos" higherTaxonomySource="treatment-meta" kingdom="Animalia" lsidName="Pterocyclos housei" order="Architaenioglossa" pageId="0" pageNumber="1" phylum="Mollusca" rank="species" species="housei">Pterocyclos housei</taxonomicName>
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-
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<bibRefCitation DOI="https://doi.org/10.5962/bhl.title.11217" author="Pfeiffer, L" journalOrPublisher="Supplementum primum, Volume 2. Cassellis: Theodori Fischeri" pageId="0" pageNumber="1" refId="B133" refString="Pfeiffer, L, 1858. Monographia Pneumonopomorum viventium. Supplementum primum, Volume 2. Cassellis: Theodori Fischeri, DOI: https://doi.org/10.5962/bhl.title.11217" title="Monographia Pneumonopomorum viventium." url="https://doi.org/10.5962/bhl.title.11217" year="1858">Pfeiffer 1858</bibRefCitation>
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: 29.
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<bibRefCitation author="Reeve, LA" journalOrPublisher="Proceedings of the Malacological Society of London" pageId="0" pageNumber="1" refId="B145" refString="Reeve, LA, 1863. Conchologia Iconica: Illustrations of the shells of molluscous animals. Volume 14, Pterocyclos, plates 1-5. Lovell Reeve and Co., London." title="Conchologia Iconica: Illustrations of the shells of molluscous animals. Volume 14, Pterocyclos, plates 1 - 5. Lovell Reeve and Co., London." year="1863">Reeve 1863</bibRefCitation>
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:
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<taxonomicName authorityName="Benson" authorityYear="1832" class="Gastropoda" family="Cyclophoridae" genus="Pterocyclos" higherTaxonomySource="treatment-meta" kingdom="Animalia" lsidName="Pterocyclos" order="Architaenioglossa" pageId="0" pageNumber="1" phylum="Mollusca" rank="genus">Pterocyclos</taxonomicName>
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pl. 4, species 21.
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</paragraph>
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<paragraph pageId="0" pageNumber="1">
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<taxonomicName baseAuthorityName="Haines" baseAuthorityYear="1855" class="Gastropoda" family="Cyclophoridae" genus="Rhiostoma" higherTaxonomySource="treatment-meta" kingdom="Animalia" lsidName="Rhiostoma housei" order="Architaenioglossa" pageId="0" pageNumber="1" phylum="Mollusca" rank="species" species="housei">Rhiostoma housei</taxonomicName>
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-
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<bibRefCitation DOI="https://doi.org/10.1080/00222936008697183" author="Benson, WH" journalOrPublisher="Journal of the Asiatic Society of Bengal" pageId="0" pageNumber="1" refId="B14" refString="Benson, WH, 1860. On Clostophis and Rhiostoma, new Burmese genera of land shells. Annals and Magazine of Natural History, Series 3 5(26): 95-97. https://doi.org/10.1080/00222936008697183" title="On Clostophis and Rhiostoma, new Burmese genera of land shells. Annals and Magazine of Natural History, Series 3 5 (26): 95 - 97." url="https://doi.org/10.1080/00222936008697183" year="1860">Benson 1860</bibRefCitation>
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: 97.
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<bibRefCitation author="Pfeiffer, L" journalOrPublisher="Proceedings of the Zoological Society of London" pageId="0" pageNumber="1" pagination="115 - 117" refId="B134" refString="Pfeiffer, L, 1862a. Description of eight new species of Cyclostomacea from the collection of H. Cumming. Esq. Proceedings of the Zoological Society of London 30: 115 - 117" title="Description of eight new species of Cyclostomacea from the collection of H. Cumming. Esq." volume="30" year="1862 a">Pfeiffer 1862a</bibRefCitation>
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||
: 117, pl. 12, fig. 9.
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<bibRefCitation DOI="https://doi.org/10.5962/bhl.title.10621" author="Pfeiffer, L" journalOrPublisher="Supplementum Secundum, Volume 3. Cassellis: Theodori Fischeri" pageId="0" pageNumber="1" refId="B136" refString="Pfeiffer, L, 1865. Monographia Pneumonopomorum viventium. Supplementum Secundum, Volume 3. Cassellis: Theodori Fischeri, DOI: https://doi.org/10.5962/bhl.title.10621" title="Monographia Pneumonopomorum viventium." url="https://doi.org/10.5962/bhl.title.10621" year="1865">Pfeiffer 1865</bibRefCitation>
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: 38.
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<bibRefCitation author="Martens, E von" journalOrPublisher="Proceedings of the Zoological Society of London" pageId="0" pageNumber="1" refId="B91" refString="Martens, E von, 1867. Die Preussische Expedition nach Ost-Asien. Verlag der Koeniglichen Geheimen Ober-Hofbuchdruckerei, 477 pp." title="Die Preussische Expedition nach Ost-Asien. Verlag der Koeniglichen Geheimen Ober-Hofbuchdruckerei, 477 pp." year="1867">Martens 1867</bibRefCitation>
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: 63.
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<bibRefCitation DOI="https://doi.org/10.5962/bhl.title.14809" author="Fischer, P" journalOrPublisher="Extrait du Bulletin de la Societe ďHistoire Naturelle ďAutun" pageId="0" pageNumber="1" pagination="1 - 193" refId="B40" refString="Fischer, P, 1891. Catalogue et distribution geographique des mollusques terrestres, fluviatiles & marins ďune partie de ľIndo-Chine (Siam, Laos, Cambodge, Cochinchine, Annam, Tonkin). Extrait du Bulletin de la Societe ďHistoire Naturelle ďAutun 4: 1 - 193, DOI: https://doi.org/10.5962/bhl.title.14809" title="Catalogue et distribution geographique des mollusques terrestres, fluviatiles & marins ďune partie de ľIndo-Chine (Siam, Laos, Cambodge, Cochinchine, Annam, Tonkin)." url="https://doi.org/10.5962/bhl.title.14809" volume="4" year="1891">Fischer 1891</bibRefCitation>
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: 101.
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<bibRefCitation author="Moellendorff, OF von" journalOrPublisher="Proceedings of the Zoological Society of London" pageId="0" pageNumber="1" pagination="146 - 156" refId="B98" refString="Moellendorff, OF von, 1894. On a collection of land-shells from the Samui Islands, Gulf of Siam. Proceedings of the Zoological Society of London 62: 146 - 156" title="On a collection of land-shells from the Samui Islands, Gulf of Siam." volume="62" year="1894">
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<normalizedToken originalValue="Möllendorff">Moellendorff</normalizedToken>
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1894
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</bibRefCitation>
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||
: 152. Kobelt and
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<normalizedToken originalValue="Möllendorff">Moellendorff</normalizedToken>
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||
1897: 115.
|
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<bibRefCitation author="Kobelt, W" journalOrPublisher="Das Tierreich" pageId="0" pageNumber="1" pagination="1 - 663" refId="B71" refString="Kobelt, W, 1902. Cyclophoridae. Das Tierreich 16: 1 - 663" title="Cyclophoridae." volume="16" year="1902">Kobelt 1902</bibRefCitation>
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: 178.
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<bibRefCitation author="Fischer, H" editor="Pavie, A" journalOrPublisher="Ernest Leroux, Editeur, Paris" pageId="0" pageNumber="1" pagination="390 - 450" refId="B42" refString="Fischer, H, Dautzenberg, P, 1904. Catalogue des mollusques terrestres et fluviatites de ľIndo-Chine orientale cites jusqu a ce jour. In: Pavie, A, Ed., Mission Pavie, Indo-Chine 1879-1895, Etudes Diverse Recherches sur Ľhistoire Naturelle de Ľhistoire Indo-Chine Orientale, Tome III. Ernest Leroux, Editeur, Paris: 390 - 450" title="Catalogue des mollusques terrestres et fluviatites de ľIndo-Chine orientale cites jusqu a ce jour." volumeTitle="Mission Pavie, Indo-Chine 1879 - 1895, Etudes Diverse Recherches sur Ľhistoire Naturelle de Ľhistoire Indo-Chine Orientale, Tome III." year="1904">Fischer and Dautzenberg 1904</bibRefCitation>
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||
: 427. Kobelt 1911: 757, 758, pl. 110, figs 8-10, pl. 113, fig. 2.
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<bibRefCitation author="Zilch, A" journalOrPublisher="Archiv fuer Molluskenkunde" pageId="0" pageNumber="1" pagination="171 - 196" refId="B188" refString="Zilch, A, 1956. Die typen und typoide des Natur-Museums Senckenberg, 18, Mollusca Cyclophoridae, Cyclophorinae (4). Archiv fuer Molluskenkunde 85: 171 - 196" title="Die typen und typoide des Natur-Museums Senckenberg, 18, Mollusca Cyclophoridae, Cyclophorinae (4)." volume="85" year="1956">Zilch 1956</bibRefCitation>
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: 174.
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||
<bibRefCitation author="Rees, WJ" journalOrPublisher="Proceedings of the Malacological Society of London" pageId="0" pageNumber="1" pagination="55 - 67" refId="B144" refString="Rees, WJ, 1964. A review of breathing devices in land operculated snail. Proceedings of the Malacological Society of London 36: 55 - 67" title="A review of breathing devices in land operculated snail." volume="36" year="1964">Rees 1964</bibRefCitation>
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||
: 67, pl. 4, fig. 15.
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<bibRefCitation author="Solem, A" journalOrPublisher="Sapolia Zoologica Musei Hauniensis" pageId="0" pageNumber="1" pagination="1 - 110" refId="B155" refString="Solem, A, 1966. Some non-marine mollusks from Thailand, with notes on classification of the Helicarionidae. Sapolia Zoologica Musei Hauniensis 24: 1 - 110" title="Some non-marine mollusks from Thailand, with notes on classification of the Helicarionidae." volume="24" year="1966">Solem 1966</bibRefCitation>
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: 11.
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<bibRefCitation author="Fischer, P-H" journalOrPublisher="Journal de Conchyliologie" pageId="0" pageNumber="1" pagination="40 - 64" refId="B41" refString="Fischer, P-H, 1973. Les mollusques testaces du Cambodge, prmiere partie: Introduction et Gastropodes Prosobranches. Journal de Conchyliologie 90: 40 - 64" title="Les mollusques testaces du Cambodge, prmiere partie: Introduction et Gastropodes Prosobranches." volume="90" year="1973">Fischer 1973</bibRefCitation>
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||
: 47.
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<bibRefCitation author="Abbott, RT" journalOrPublisher=", Melbourne, Florida" pageId="0" pageNumber="1" refId="B1" refString="Abbott, RT, 1989. Compendium of Landshells. American Malacologists Inc., Melbourne, Florida" title="Compendium of Landshells. American Malacologists Inc." year="1989">Abbott 1989</bibRefCitation>
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||
: 29, 42, with 3 figures.
|
||
<bibRefCitation author="Tumpeesuwan, S" journalOrPublisher="Master Thesis, Department of Biology, Faculty of Science, Chulalongkorn University" pageId="0" pageNumber="1" refId="B175" refString="Tumpeesuwan, S, 2001. Taxonomy of snorkel snails genus Rhiostoma Benson, 1860 in Thailand. Master Thesis, Department of Biology, Faculty of Science, Chulalongkorn University" title="Taxonomy of snorkel snails genus Rhiostoma Benson, 1860 in Thailand." year="2001">Tumpeesuwan 2001</bibRefCitation>
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||
: 41-46, figs 4.10-4.12.
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||
<bibRefCitation author="Kongim, B" journalOrPublisher="The Raffles Bulletin of Zoology" pageId="0" pageNumber="1" pagination="13 - 20" refId="B78" refString="Kongim, B, Sutcharit, C, Tongkerd, P, Panha, S, 2013b. Karyotype of the snorkel snails genera Pterocyclos and Rhiostoma (Prosobranchia: Cyclophoridae). The Raffles Bulletin of Zoology 61: 13 - 20" title="Karyotype of the snorkel snails genera Pterocyclos and Rhiostoma (Prosobranchia: Cyclophoridae)." volume="61" year="2013 b">Kongim et al. 2013b</bibRefCitation>
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||
: 16, fig. 2f, g.
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||
</paragraph>
|
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<paragraph pageId="0" pageNumber="1">
|
||
<taxonomicName class="Squamata" family="Cyclophoridae" genus="Rhinostoma" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Rhinostoma" order="Architaenioglossa" pageId="0" pageNumber="1" phylum="Chordata" rank="genus">Rhinostoma</taxonomicName>
|
||
[sic]
|
||
<taxonomicName baseAuthorityName="Haines" baseAuthorityYear="1855" class="Squamata" family="Cyclophoridae" genus="Rhinostoma" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Rhinostoma housei" order="Architaenioglossa" pageId="0" pageNumber="1" phylum="Chordata" rank="species" species="housei">Rhinostoma housei</taxonomicName>
|
||
-
|
||
<bibRefCitation author="Habe, T" journalOrPublisher="Nature and Life in Southeast Asia, Kyoto" pageId="0" pageNumber="1" pagination="111 - 128" refId="B53" refString="Habe, T, 1965. Operculated land molluscs from Southeast Asia. Nature and Life in Southeast Asia, Kyoto 4: 111 - 128" title="Operculated land molluscs from Southeast Asia." volume="4" year="1965">Habe 1965</bibRefCitation>
|
||
: 128 (in part), pl. 2, fig. 10.
|
||
</paragraph>
|
||
<paragraph pageId="0" pageNumber="1">
|
||
<taxonomicName class="Gastropoda" family="Cyclophoridae" genus="Pterocyclos" higherTaxonomySource="treatment-meta" kingdom="Animalia" lsidName="Pterocyclos (Spiraculum) housei" order="Architaenioglossa" pageId="0" pageNumber="1" phylum="Mollusca" rank="species" species="housei" subGenus="Spiraculum">Pterocyclos (Spiraculum) housei</taxonomicName>
|
||
-
|
||
<bibRefCitation author="Martens, E von" journalOrPublisher="Proceedings of the Zoological Society of London" pageId="0" pageNumber="1" pagination="6 - 18" refId="B90" refString="Martens, E von, 1860. On the Mollusca of Siam. Proceedings of the Zoological Society of London 28: 6 - 18" title="On the Mollusca of Siam." volume="28" year="1860">Martens 1860</bibRefCitation>
|
||
: 10.
|
||
</paragraph>
|
||
<paragraph pageId="0" pageNumber="1">
|
||
<taxonomicName class="Gastropoda" family="Cyclophoridae" genus="Pterocyclus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Pterocyclus" order="Architaenioglossa" pageId="0" pageNumber="1" phylum="Mollusca" rank="genus">Pterocyclus</taxonomicName>
|
||
[sic] (
|
||
<taxonomicName authorityName="Tongkerd & Tumpeesuwan & Inkhavilay & Prasankok & Jeratthitikul & Panha & Sutcharit" authorityYear="2023" class="Gastropoda" family="Cyclophoridae" genus="Rhiostoma" higherTaxonomySource="treatment-meta" kingdom="Animalia" lsidName="Rhiostoma" order="Architaenioglossa" pageId="0" pageNumber="1" phylum="Mollusca" rank="genus">Rhiostoma</taxonomicName>
|
||
)
|
||
<taxonomicName baseAuthorityName="Haines" baseAuthorityYear="1855" class="Gastropoda" family="Cyclophoridae" genus="Rhiostoma" higherTaxonomySource="treatment-meta" kingdom="Animalia" lsidName="Rhiostoma housei" order="Architaenioglossa" pageId="0" pageNumber="1" phylum="Mollusca" rank="species" species="housei">Rhiostoma housei</taxonomicName>
|
||
-
|
||
<bibRefCitation DOI="https://doi.org/10.5962/bhl.title.11957" author="Nevill, G" journalOrPublisher="Calcutta, Trustees" pageId="0" pageNumber="1" refId="B114" refString="Nevill, G, 1878. Hand list of Mollusca in the Indian Museum, Calcutta. Calcutta, Trustees, DOI: https://doi.org/10.5962/bhl.title.11957" title="Hand list of Mollusca in the Indian Museum, Calcutta." url="https://doi.org/10.5962/bhl.title.11957" year="1878">Nevill 1878</bibRefCitation>
|
||
: 263.
|
||
</paragraph>
|
||
<paragraph pageId="0" pageNumber="1">
|
||
<taxonomicName authorityName="Haines" authorityYear="1855" class="Gastropoda" family="Cyclophoridae" genus="Cyclostoma" higherTaxonomySource="treatment-meta" kingdom="Animalia" lsidName="Cyclostoma (Pterocyclos) housei" order="Architaenioglossa" pageId="0" pageNumber="1" phylum="Mollusca" rank="species" species="housei" subGenus="Pterocyclos">Cyclostoma (Pterocyclos) housei</taxonomicName>
|
||
-
|
||
<bibRefCitation DOI="https://doi.org/10.1206/0003-0090(2001)262<0001:CORTSI>2.0.CO;2" author="Boyko, CB" journalOrPublisher="Bulletin of the American Museum of Natural History" pageId="0" pageNumber="1" pagination="1 - 158" refId="B24" refString="Boyko, CB, Cordeiro, JR, 2001. Catalog of recent type specimens in the Division of Invertebrate Zoology, American Museum of Natural History. V. Mollusca, Part 2 (Class Gastropoda [exclusive of Opisthobranchia and Pulmonata], with supplements to Gastropoda [Opisthobranchia], and Bivalvia). Bulletin of the American Museum of Natural History 262: 1 - 158, DOI: https://doi.org/10.1206/0003-0090(2001)262<0001:CORTSI>2.0.CO;2" title="Catalog of recent type specimens in the Division of Invertebrate Zoology, American Museum of Natural History. V. Mollusca, Part 2 (Class Gastropoda [exclusive of Opisthobranchia and Pulmonata], with supplements to Gastropoda [Opisthobranchia], and Bivalvia)." url="https://doi.org/10.1206/0003-0090(2001)262<0001:CORTSI>2.0.CO;2" volume="262" year="2001">Boyko and Cordeiro 2001</bibRefCitation>
|
||
: 52.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection pageId="0" pageNumber="1" type="type material">
|
||
<paragraph pageId="0" pageNumber="1">Type material.</paragraph>
|
||
<paragraph pageId="0" pageNumber="1">
|
||
This species was described from specimens from the Haines collection (AMNH). The unique name-bearing type was not designated; however, at least two specimens were illustrated in the original description. The specimen identical to the illustration (shape of breathing device) and measurements in
|
||
<bibRefCitation DOI="https://doi.org/10.1111/j.1749-6632.1858.tb00358.x" author="Haines, WA" journalOrPublisher="Nature and Life in Southeast Asia, Kyoto" pageId="0" pageNumber="1" refId="B54" refString="Haines, WA, 1855. Descriptions of four new species of terrestrial shells from Siam. Annals of the Lyceum of Natural History of New York 6: 157, 158. https://doi.org/10.1111/j.1749-6632.1858.tb00358.x" title="Descriptions of four new species of terrestrial shells from Siam. Annals of the Lyceum of Natural History of New York 6: 157, 158." url="https://doi.org/10.1111/j.1749-6632.1858.tb00358.x" year="1855">Haines (1855</bibRefCitation>
|
||
: figs 12, 13) is figured herein,
|
||
<emphasis bold="true" italics="true" pageId="0" pageNumber="1">syntype</emphasis>
|
||
AMNH 42923 (5 shells; Fig.
|
||
<figureCitation captionStart="Figure 19" captionStartId="F19" captionText="Figure 19. Shell of Rhiostoma housei A, B syntype AMNH 42923 from Siam A figured specimen in Haines (1855: figs 12, 13) anb B other specimen from the same syntype lot C specimens CUMZ 4383 from Tham Phitsadan, Chumphon D specimen CUMZ 3982 from Tham Sriwilai, Saraburi E specimen CUMZ 4384 from Pak Chong, Nakhon Ratchasrima F specimen CUMZ 4879 from Hua Hin, Prachuap Khiri Khan." figureDoi="10.3897/zookeys.1142.90097.figure19" httpUri="https://binary.pensoft.net/fig/799877" pageId="0" pageNumber="1">19A, B</figureCitation>
|
||
) from Siam [Thailand].
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection pageId="0" pageNumber="1" type="materials_examined">
|
||
<paragraph pageId="0" pageNumber="1">Other material examined.</paragraph>
|
||
<paragraph pageId="0" pageNumber="1">
|
||
<materialsCitation collectingDate="2023-01-01" collectingDateMax="2023-12-31" collectingDateMin="2023-01-01" country="Thailand" county="East Siam" location="East Siam" municipality="East Siam" specimenCount="1" stateProvince="Saraburi">
|
||
<emphasis bold="true" pageId="0" pageNumber="1">
|
||
<collectingCountry name="Thailand">Thailand</collectingCountry>
|
||
</emphasis>
|
||
: Boong Tuey,
|
||
<collectingCounty>East Siam</collectingCounty>
|
||
: RBINS 659990, 659991; Pak Chong,
|
||
<collectingMunicipality>East Siam</collectingMunicipality>
|
||
: RBINS 659978, 659979, 659980; Lam Ton Lang,
|
||
<location LSID="urn:lsid:plazi:treatment:03122B5FADC55EFE8BC5CF4838D12522:09A020DC20452C1AB7A3BDD344678012" country="Thailand" county="East Siam" municipality="East Siam" name="East Siam" stateProvince="Saraburi">East Siam</location>
|
||
: RBINS 659982, 659983, 659984. Siam: NHMW 75000/E/23126 (1 shell). Samui, Siam: NHMW Rusnov (1 shell). Tham Phet Tham Thong, Takhli, Nakhonsawan: CUMZ 4385. Pukae Botanical Garden, Chaloem Phra Kiat,
|
||
<collectingRegion country="Thailand" name="Sara Buri">Saraburi</collectingRegion>
|
||
: CUMZ 4742. Wat Tham Srivilai, Chaloem Phra Kiat,
|
||
<collectingRegion country="Thailand" name="Sara Buri">Saraburi</collectingRegion>
|
||
: CUMZ 3982 (Figs
|
||
<figureCitation captionStart="Figure 7" captionStartId="F7" captionText="Figure 7. A-C Internal anatomy of Rhiostoma housei specimen CUMZ 3982 from Tham Sriwilai, Saraburi, A inside view of lung cavity shows an arrangement of the kidney, circulatory system and the terminal part of the digestive tract B male genitalia and C female genitalia D spermatophore of Rhiostoma samuiense specimen CUMZ 4713 from Kanchanadit, Surat Thani with its cross sections showing spermatophore shape in each position. Abbreviations: an, anus; bc, bursa copulatrix; fc, faeces; go, genital opening; h, heart; kd, kidney; mc, mantle collar; od, oviduct; pg, prostate gland; re, rectum; sr, seminal receptacle; ut, uterus; v, vein; vd, vas deferens." figureDoi="10.3897/zookeys.1142.90097.figure7" httpUri="https://binary.pensoft.net/fig/799865" pageId="0" pageNumber="1">7A-C</figureCitation>
|
||
,
|
||
<figureCitation captionStart="Figure 19" captionStartId="F19" captionText="Figure 19. Shell of Rhiostoma housei A, B syntype AMNH 42923 from Siam A figured specimen in Haines (1855: figs 12, 13) anb B other specimen from the same syntype lot C specimens CUMZ 4383 from Tham Phitsadan, Chumphon D specimen CUMZ 3982 from Tham Sriwilai, Saraburi E specimen CUMZ 4384 from Pak Chong, Nakhon Ratchasrima F specimen CUMZ 4879 from Hua Hin, Prachuap Khiri Khan." figureDoi="10.3897/zookeys.1142.90097.figure19" httpUri="https://binary.pensoft.net/fig/799877" pageId="0" pageNumber="1">19D</figureCitation>
|
||
,
|
||
<figureCitation captionStart="Figure 20" captionStartId="F20" captionText="Figure 20. Drawing of general anatomy of Rhiostoma housei specimen CUMZ 3982 from Saraburi Province, Thailand. A female B male. Abbreviations: bc, bursa copulatrix;; cm, columellar muscle; ct, cephalic tentacle; dg, digestive gland; es, eyespot; f, foot; fc, faeces; int, intestine; lc, lung cavity; mc, mantle collar; od, oviduct; op, operculum; ov, ovary; p, external penis; pg, prostate gland; sg, sperm groove; sn, snout; te, testis; ut, uterus; vd, vas deferens; vg, vaginal groove." figureDoi="10.3897/zookeys.1142.90097.figure20" httpUri="https://binary.pensoft.net/fig/799878" pageId="0" pageNumber="1">20</figureCitation>
|
||
,
|
||
<figureCitation captionStart="Figure 32" captionStartId="F32" captionText="Figure 32. Radula morphology A Rhiostoma housei specimen CUMZ 3982 from Wat Tham Srivilai, Saraburi B Rhiostoma hainesi specimen from Chanthaburi C Rhiostoma jalorensis specimen CUMZ 3994 from Wat Tham Sua, Krabi D Rhiostoma ebenozostera sp. nov. paratype CUMZ 4704 from Tham Pha Poo, Loei E Rhiostoma lannaense sp. nov. paratype CUMZ 4701 from Chiang Dao, Chiang Mai F Rhiostoma tigrina sp. nov. paratype CUMZ 4497 / 2 from Mae Sai, Chiang Rai." figureDoi="10.3897/zookeys.1142.90097.figure32" httpUri="https://binary.pensoft.net/fig/799890" pageId="0" pageNumber="1">32A</figureCitation>
|
||
), 4449. Kaeng Khoi,
|
||
<collectingRegion country="Thailand" name="Sara Buri">Saraburi</collectingRegion>
|
||
: CUMZ 4315.
|
||
<location LSID="urn:lsid:plazi:treatment:03122B5FADC55EFE8BC5CF4838D12522:481EF7EA020CEF01EC7556F3C157DBA8" country="Thailand" county="East Siam" municipality="East Siam" name="Ban Tha Sao Village" stateProvince="Saraburi">Ban Tha Sao Village</location>
|
||
, Kaeng Khoi,
|
||
<collectingRegion country="Thailand" name="Sara Buri">Saraburi</collectingRegion>
|
||
: CUMZ 10135. Wat Tham Phra Phothisat, Kaeng Khoi,
|
||
<collectingRegion country="Thailand" name="Sara Buri">Saraburi</collectingRegion>
|
||
: CUMZ 10026, 10158. Chet Sao Noi Waterfall, Muaklek,
|
||
<collectingRegion country="Thailand" name="Sara Buri">Saraburi</collectingRegion>
|
||
: CUMZ 4732. Muaklek,
|
||
<collectingRegion country="Thailand" name="Sara Buri">Saraburi</collectingRegion>
|
||
: CUMZ 4390, 4460. Wat Tham Dao Khao Kaew, Muaklek,
|
||
<collectingRegion country="Thailand" name="Sara Buri">Saraburi</collectingRegion>
|
||
: CUMZ 3802, 3934, 4389, 4760, 4842, 4897, 10131, 10133. Khao Sompod, Chai Ba Dan,
|
||
<collectingRegion country="Thailand" name="Lop Buri">Lopburi</collectingRegion>
|
||
: CUMZ 3932, 3936, 3937. Wat Tham Promlok, Chai Ba Dan,
|
||
<collectingRegion country="Thailand" name="Lop Buri">Lopburi</collectingRegion>
|
||
: CUMZ 10128. Sub Langka, Lamsonthi,
|
||
<collectingRegion country="Thailand" name="Lop Buri">Lopburi</collectingRegion>
|
||
: CUMZ 4736. Khao Look Chang, Pak Chong,
|
||
<collectingRegion country="Thailand" name="Nakhon Ratchasima">Nakhon Ratchasima</collectingRegion>
|
||
: CUMZ 3805, 3981, 4316, 4318, 4320, 4379, 4384 (Fig.
|
||
<figureCitation captionStart="Figure 19" captionStartId="F19" captionText="Figure 19. Shell of Rhiostoma housei A, B syntype AMNH 42923 from Siam A figured specimen in Haines (1855: figs 12, 13) anb B other specimen from the same syntype lot C specimens CUMZ 4383 from Tham Phitsadan, Chumphon D specimen CUMZ 3982 from Tham Sriwilai, Saraburi E specimen CUMZ 4384 from Pak Chong, Nakhon Ratchasrima F specimen CUMZ 4879 from Hua Hin, Prachuap Khiri Khan." figureDoi="10.3897/zookeys.1142.90097.figure19" httpUri="https://binary.pensoft.net/fig/799877" pageId="0" pageNumber="1">19E</figureCitation>
|
||
), 4454. The way to Khao Yai, Pak Chong,
|
||
<collectingRegion country="Thailand" name="Nakhon Ratchasima">Nakhon Ratchasima</collectingRegion>
|
||
: CUMZ 4454. Wat Tham Prommachan Thammaram, Pak Chong,
|
||
<collectingRegion country="Thailand" name="Nakhon Ratchasima">Nakhon Ratchasima</collectingRegion>
|
||
: CUMZ 10160. Wat Tham Sub Muet, Pak Chong,
|
||
<collectingRegion country="Thailand" name="Nakhon Ratchasima">Nakhon Ratchasima</collectingRegion>
|
||
: CUMZ 10132. Wat Tham Thep Nimit, Pak Chong,
|
||
<collectingRegion country="Thailand" name="Nakhon Ratchasima">Nakhon Ratchasima</collectingRegion>
|
||
: CUMZ 10126. Wat Thepphitak Punnaram, Pak Chong,
|
||
<collectingRegion country="Thailand" name="Nakhon Ratchasima">Nakhon Ratchasima</collectingRegion>
|
||
: CUMZ 4321, 4416. Sakaerat Biosphere, Pak Thong Chai,
|
||
<collectingRegion country="Thailand" name="Nakhon Ratchasima">Nakhon Ratchasima</collectingRegion>
|
||
: CUMZ 4391.
|
||
<location LSID="urn:lsid:plazi:treatment:03122B5FADC55EFE8BC5CF4838D12522:181AF75D22508CB0BB8E8430244AE702" country="Thailand" county="East Siam" municipality="East Siam" name="Sakaerat Environmental Research Station" stateProvince="Saraburi">Sakaerat Environmental Research Station</location>
|
||
, Wang Nam Khiao,
|
||
<collectingRegion country="Thailand" name="Nakhon Ratchasima">Nakhon Ratchasima</collectingRegion>
|
||
: CUMZ 10134. Ban Bueng, Ban Bueng,
|
||
<collectingRegion country="Thailand" name="Chon Buri">Chonburi</collectingRegion>
|
||
: CUMZ 4312. Khao Chakan, Srakeo: CUMZ 10156. Khao Nang Panturat, Cha Am,
|
||
<collectingRegion country="Thailand" name="Phetchaburi">Phetchaburi</collectingRegion>
|
||
: CUMZ 4879 (Fig.
|
||
<figureCitation captionStart="Figure 19" captionStartId="F19" captionText="Figure 19. Shell of Rhiostoma housei A, B syntype AMNH 42923 from Siam A figured specimen in Haines (1855: figs 12, 13) anb B other specimen from the same syntype lot C specimens CUMZ 4383 from Tham Phitsadan, Chumphon D specimen CUMZ 3982 from Tham Sriwilai, Saraburi E specimen CUMZ 4384 from Pak Chong, Nakhon Ratchasrima F specimen CUMZ 4879 from Hua Hin, Prachuap Khiri Khan." figureDoi="10.3897/zookeys.1142.90097.figure19" httpUri="https://binary.pensoft.net/fig/799877" pageId="0" pageNumber="1">19F</figureCitation>
|
||
), 10129, 10130, 10164, 10165.
|
||
<location LSID="urn:lsid:plazi:treatment:03122B5FADC55EFE8BC5CF4838D12522:0CAE6503CB067FBB1C0B54954B8704AE" country="Thailand" county="East Siam" municipality="East Siam" name="Khao Nang Panturat Nature Reserve" stateProvince="Saraburi">Khao Nang Panturat Nature Reserve</location>
|
||
, Cha Am,
|
||
<collectingRegion country="Thailand" name="Phetchaburi">Phetchaburi</collectingRegion>
|
||
: CUMZ 4480.
|
||
<location LSID="urn:lsid:plazi:treatment:03122B5FADC55EFE8BC5CF4838D12522:4A21BB501F97840176D365B9116B9D6F" country="Thailand" county="East Siam" municipality="East Siam" name="Tham Na Kwang" stateProvince="Saraburi">Tham Na Kwang</location>
|
||
, Cha Am,
|
||
<collectingRegion country="Thailand" name="Phetchaburi">Phetchaburi</collectingRegion>
|
||
: CUMZ 3988, 4452, 4719, 10162. 30 km. before
|
||
<location LSID="urn:lsid:plazi:treatment:03122B5FADC55EFE8BC5CF4838D12522:865DDE916C2BFBDC42304B33904FE650" country="Thailand" county="East Siam" municipality="East Siam" name="Pa La Au Waterfall" stateProvince="Saraburi">Pa La Au Waterfall</location>
|
||
, Hua Hin,
|
||
<collectingRegion country="Thailand" name="Phetchaburi">Phetchaburi</collectingRegion>
|
||
: CUMZ 3827, 4740. Kaeng Kachan,
|
||
<collectingRegion country="Thailand" name="Phetchaburi">Phetchaburi</collectingRegion>
|
||
: CUMZ 4409, 4412. Khao Lom Muak, Muang,
|
||
<collectingRegion country="Thailand" name="Prachuap Khiri Khan">Prachuap Khiri Khan</collectingRegion>
|
||
: CUMZ 3870, 3987 (Fig.
|
||
<figureCitation captionStart="Figure 17" captionStartId="F17" captionText="Figure 17. Live snails of the species group II A Rhiostoma housei specimen CUMZ 3987 from Ao Manow, Prachuap Khiri Khan B, C Rhiostoma hainesi from Khao Soi Dao, Chanthaburi B without breathing device and C with long breathing device D Rhiostoma jalorensis specimen CUMZ 3994 from Wat Tham Sue, Krabi. All figures are not to scale. Photographs: P. Tongkerd (B, C)." figureDoi="10.3897/zookeys.1142.90097.figure17" httpUri="https://binary.pensoft.net/fig/799875" pageId="0" pageNumber="1">17A</figureCitation>
|
||
), 4322, 10163. Bang Poo, Sam Roi Yod,
|
||
<collectingRegion country="Thailand" name="Prachuap Khiri Khan">Prachuap Khiri Khan</collectingRegion>
|
||
: CUMZ 4414, 4417, 4429, 4430.
|
||
<location LSID="urn:lsid:plazi:treatment:03122B5FADC55EFE8BC5CF4838D12522:F42B3B25FEEB327418C3E6483928E409" country="Thailand" county="East Siam" municipality="East Siam" name="Saun Wiweg Bureau of Monks" stateProvince="Saraburi">Saun Wiweg Bureau of Monks</location>
|
||
, Sam Roi Yod,
|
||
<collectingRegion country="Thailand" name="Prachuap Khiri Khan">Prachuap Khiri Khan</collectingRegion>
|
||
: CUMZ 10127. Ao Noi,
|
||
<collectingRegion country="Thailand" name="Prachuap Khiri Khan">Prachuap Khiri Khan</collectingRegion>
|
||
: CUMZ 3868. Tham Pissadarn, Tha Sae,
|
||
<collectingRegion country="Thailand" name="Chumphon">Chumphon</collectingRegion>
|
||
: CUMZ 4383 (Fig.
|
||
<figureCitation captionStart="Figure 19" captionStartId="F19" captionText="Figure 19. Shell of Rhiostoma housei A, B syntype AMNH 42923 from Siam A figured specimen in Haines (1855: figs 12, 13) anb B other specimen from the same syntype lot C specimens CUMZ 4383 from Tham Phitsadan, Chumphon D specimen CUMZ 3982 from Tham Sriwilai, Saraburi E specimen CUMZ 4384 from Pak Chong, Nakhon Ratchasrima F specimen CUMZ 4879 from Hua Hin, Prachuap Khiri Khan." figureDoi="10.3897/zookeys.1142.90097.figure19" httpUri="https://binary.pensoft.net/fig/799877" pageId="0" pageNumber="1">19C</figureCitation>
|
||
), 4453, 4851
|
||
</materialsCitation>
|
||
.
|
||
</paragraph>
|
||
<caption doi="10.3897/zookeys.1142.90097.figure17" httpUri="https://binary.pensoft.net/fig/799875" pageId="0" pageNumber="1" start="Figure 17" startId="F17">
|
||
<paragraph pageId="0" pageNumber="1">
|
||
<emphasis bold="true" pageId="0" pageNumber="1">Figure 17.</emphasis>
|
||
Live snails of the species group II
|
||
<emphasis bold="true" pageId="0" pageNumber="1">A</emphasis>
|
||
<taxonomicName baseAuthorityName="Haines" baseAuthorityYear="1855" class="Gastropoda" family="Cyclophoridae" genus="Rhiostoma" kingdom="Animalia" lsidName="Rhiostoma housei" order="Architaenioglossa" pageId="0" pageNumber="1" phylum="Mollusca" rank="species" species="housei">
|
||
<emphasis italics="true" pageId="0" pageNumber="1">Rhiostoma housei</emphasis>
|
||
</taxonomicName>
|
||
specimen CUMZ 3987 from Ao Manow,
|
||
<collectingRegion country="Thailand" name="Prachuap Khiri Khan">Prachuap Khiri Khan</collectingRegion>
|
||
<emphasis bold="true" pageId="0" pageNumber="1">B, C</emphasis>
|
||
<taxonomicName authorityName="Pfeiffer" authorityYear="1862" class="Gastropoda" family="Cyclophoridae" genus="Rhiostoma" kingdom="Animalia" lsidName="Rhiostoma hainesi" order="Architaenioglossa" pageId="0" pageNumber="1" phylum="Mollusca" rank="species" species="hainesi">
|
||
<emphasis italics="true" pageId="0" pageNumber="1">Rhiostoma hainesi</emphasis>
|
||
</taxonomicName>
|
||
from Khao Soi Dao,
|
||
<collectingRegion country="Thailand" name="Chanthaburi">Chanthaburi</collectingRegion>
|
||
<emphasis bold="true" pageId="0" pageNumber="1">B</emphasis>
|
||
without breathing device and
|
||
<emphasis bold="true" pageId="0" pageNumber="1">C</emphasis>
|
||
with long breathing device
|
||
<emphasis bold="true" pageId="0" pageNumber="1">D</emphasis>
|
||
<taxonomicName authorityName="Sykes" authorityYear="1903" class="Gastropoda" family="Cyclophoridae" genus="Rhiostoma" kingdom="Animalia" lsidName="Rhiostoma jalorensis" order="Architaenioglossa" pageId="0" pageNumber="1" phylum="Mollusca" rank="species" species="jalorensis">
|
||
<emphasis italics="true" pageId="0" pageNumber="1">Rhiostoma jalorensis</emphasis>
|
||
</taxonomicName>
|
||
specimen CUMZ 3994 from Wat Tham Sue,
|
||
<collectingRegion country="Thailand" name="Krabi">Krabi</collectingRegion>
|
||
. All figures are not to scale. Photographs: P. Tongkerd (
|
||
<emphasis bold="true" pageId="0" pageNumber="1">B, C</emphasis>
|
||
).
|
||
</paragraph>
|
||
</caption>
|
||
</subSubSection>
|
||
<subSubSection pageId="0" pageNumber="1" type="diagnosis">
|
||
<paragraph pageId="0" pageNumber="1">Diagnosis.</paragraph>
|
||
<paragraph pageId="0" pageNumber="1">Shell medium to large, thick, and depressed. Detached whorl longer than aperture width, curved, and descending. Breathing device straight with complete tube. Shell usually with dark brown zigzag patterns.</paragraph>
|
||
</subSubSection>
|
||
<subSubSection pageId="0" pageNumber="1" type="differential diagnosis">
|
||
<paragraph pageId="0" pageNumber="1">Differential diagnosis.</paragraph>
|
||
<paragraph pageId="0" pageNumber="1">
|
||
<taxonomicName baseAuthorityName="Haines" baseAuthorityYear="1855" class="Gastropoda" family="Cyclophoridae" genus="Rhiostoma" kingdom="Animalia" lsidName="Rhiostoma housei" order="Architaenioglossa" pageId="0" pageNumber="1" phylum="Mollusca" rank="species" species="housei">
|
||
<emphasis italics="true" pageId="0" pageNumber="1">Rhiostoma housei</emphasis>
|
||
</taxonomicName>
|
||
can be distinguished from
|
||
<taxonomicName lsidName="R. tigrina" pageId="0" pageNumber="1" rank="species" species="tigrina">
|
||
<emphasis italics="true" pageId="0" pageNumber="1">R. tigrina</emphasis>
|
||
</taxonomicName>
|
||
sp. nov. in usually having a stout detached whorl, generally a curved tubular breathing device, and a faded colour pattern. In contrast,
|
||
<taxonomicName lsidName="R. tigrina" pageId="0" pageNumber="1" rank="species" species="tigrina">
|
||
<emphasis italics="true" pageId="0" pageNumber="1">R. tigrina</emphasis>
|
||
</taxonomicName>
|
||
sp. nov. has a slender detached whorl, generally a straight tubular breathing device, and a brownish zigzag colour pattern.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection pageId="0" pageNumber="1" type="description">
|
||
<paragraph pageId="0" pageNumber="1">Description.</paragraph>
|
||
<paragraph pageId="0" pageNumber="1">
|
||
<emphasis bold="true" pageId="0" pageNumber="1">
|
||
<emphasis italics="true" pageId="0" pageNumber="1">Shell</emphasis>
|
||
.
|
||
</emphasis>
|
||
Shell medium to large, cW 22.1-26.4 mm, cH 12.1-13.8 mm, thick, and sub-discoidal to discoidal shape; detached-whorl length 8.0-14.5 mm. Apex acute with dark colouration; spire slightly elevated. Whorls 5 to 6, convex and increasing regularly; suture wide and deep; last whorl rounded and stout. Shell surface with fine growth lines. Periostracum thin corneous, and transparent to thick with brownish colour. Shell colour whitish with dark brown zigzag pattern; dorsal shell surface with darker colour pattern than ventral surface; with narrow brownish spiral band on periphery (sometimes absent). Detached whorl of medium length, approximately the same as apertural width. Peristome circular and double; lip thickened, slightly expanded and multi-layered. Aperture opened sub-laterally. Breathing device tubular and its tip usually attached to preceding whorl; outer lip forms a long, curved or straight, closed tube; inner lip with small hole inside aperture. Umbilicus widely opened and deep. Operculum calcareous, tall cup-shaped, and multispiral (Fig.
|
||
<figureCitation captionStart="Figure 19" captionStartId="F19" captionText="Figure 19. Shell of Rhiostoma housei A, B syntype AMNH 42923 from Siam A figured specimen in Haines (1855: figs 12, 13) anb B other specimen from the same syntype lot C specimens CUMZ 4383 from Tham Phitsadan, Chumphon D specimen CUMZ 3982 from Tham Sriwilai, Saraburi E specimen CUMZ 4384 from Pak Chong, Nakhon Ratchasrima F specimen CUMZ 4879 from Hua Hin, Prachuap Khiri Khan." figureDoi="10.3897/zookeys.1142.90097.figure19" httpUri="https://binary.pensoft.net/fig/799877" pageId="0" pageNumber="1">19</figureCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph pageId="0" pageNumber="1">
|
||
<emphasis bold="true" pageId="0" pageNumber="1">
|
||
<emphasis italics="true" pageId="0" pageNumber="1">Radula</emphasis>
|
||
.
|
||
</emphasis>
|
||
Taenioglossate radula arranged in inverted V-shaped rows. Central tooth with well-developed central cusp and two lateral cusps on each side tapering in size; central cusp large with blunt tip; lateral cusps with triangular shape and pointed tip. Lateral teeth consisting of four cusps; central cusp large, elongate, and flanked by pointed tips of two small inner cusps and one outer cusp. Inner and outer marginal teeth each composed of three cusps; central cusp with large and dull tip, flanked by small pointed tips of one inner cusp and one outer cusp (Fig.
|
||
<figureCitation captionStart="Figure 32" captionStartId="F32" captionText="Figure 32. Radula morphology A Rhiostoma housei specimen CUMZ 3982 from Wat Tham Srivilai, Saraburi B Rhiostoma hainesi specimen from Chanthaburi C Rhiostoma jalorensis specimen CUMZ 3994 from Wat Tham Sua, Krabi D Rhiostoma ebenozostera sp. nov. paratype CUMZ 4704 from Tham Pha Poo, Loei E Rhiostoma lannaense sp. nov. paratype CUMZ 4701 from Chiang Dao, Chiang Mai F Rhiostoma tigrina sp. nov. paratype CUMZ 4497 / 2 from Mae Sai, Chiang Rai." figureDoi="10.3897/zookeys.1142.90097.figure32" httpUri="https://binary.pensoft.net/fig/799890" pageId="0" pageNumber="1">32A</figureCitation>
|
||
).
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection pageId="0" pageNumber="1" type="distribution">
|
||
<paragraph pageId="0" pageNumber="1">Distribution.</paragraph>
|
||
<paragraph pageId="0" pageNumber="1">
|
||
The type locality was given as a broad location of
|
||
<normalizedToken originalValue="“Siam”">"Siam"</normalizedToken>
|
||
; however, the specimens from central Thailand are the most similar to the syntype illustrated herein (Fig.
|
||
<figureCitation captionStart="Figure 19" captionStartId="F19" captionText="Figure 19. Shell of Rhiostoma housei A, B syntype AMNH 42923 from Siam A figured specimen in Haines (1855: figs 12, 13) anb B other specimen from the same syntype lot C specimens CUMZ 4383 from Tham Phitsadan, Chumphon D specimen CUMZ 3982 from Tham Sriwilai, Saraburi E specimen CUMZ 4384 from Pak Chong, Nakhon Ratchasrima F specimen CUMZ 4879 from Hua Hin, Prachuap Khiri Khan." figureDoi="10.3897/zookeys.1142.90097.figure19" httpUri="https://binary.pensoft.net/fig/799877" pageId="0" pageNumber="1">19A, B</figureCitation>
|
||
). In this study,
|
||
<taxonomicName lsidName="R. housei" pageId="0" pageNumber="1" rank="species" species="housei">
|
||
<emphasis italics="true" pageId="0" pageNumber="1">R. housei</emphasis>
|
||
</taxonomicName>
|
||
has a wide distribution ranging from central to peninsular Thailand. The southernmost limits are around the Isthmus of Kra (Pratiew, Chumphon Province), and the northern limit is approximately in Nakhon Sawan Province (Fig.
|
||
<figureCitation captionStart="Figure 18" captionStartId="F18" captionText="Figure 18. Distribution map of Rhiostoma housei - species group." figureDoi="10.3897/zookeys.1142.90097.figure18" httpUri="https://binary.pensoft.net/fig/799876" pageId="0" pageNumber="1">18</figureCitation>
|
||
).
|
||
</paragraph>
|
||
<caption doi="10.3897/zookeys.1142.90097.figure18" httpUri="https://binary.pensoft.net/fig/799876" pageId="0" pageNumber="1" start="Figure 18" startId="F18">
|
||
<paragraph pageId="0" pageNumber="1">
|
||
<emphasis bold="true" pageId="0" pageNumber="1">Figure 18.</emphasis>
|
||
Distribution map of
|
||
<taxonomicName baseAuthorityName="Haines" baseAuthorityYear="1855" class="Gastropoda" family="Cyclophoridae" genus="Rhiostoma" kingdom="Animalia" lsidName="Rhiostoma housei" order="Architaenioglossa" pageId="0" pageNumber="1" phylum="Mollusca" rank="species" species="housei">
|
||
<emphasis italics="true" pageId="0" pageNumber="1">Rhiostoma housei</emphasis>
|
||
</taxonomicName>
|
||
-species group.
|
||
</paragraph>
|
||
</caption>
|
||
<caption doi="10.3897/zookeys.1142.90097.figure19" httpUri="https://binary.pensoft.net/fig/799877" pageId="0" pageNumber="1" start="Figure 19" startId="F19">
|
||
<paragraph pageId="0" pageNumber="1">
|
||
<emphasis bold="true" pageId="0" pageNumber="1">Figure 19.</emphasis>
|
||
Shell of
|
||
<taxonomicName baseAuthorityName="Haines" baseAuthorityYear="1855" class="Gastropoda" family="Cyclophoridae" genus="Rhiostoma" kingdom="Animalia" lsidName="Rhiostoma housei" order="Architaenioglossa" pageId="0" pageNumber="1" phylum="Mollusca" rank="species" species="housei">
|
||
<emphasis italics="true" pageId="0" pageNumber="1">Rhiostoma housei</emphasis>
|
||
</taxonomicName>
|
||
<emphasis bold="true" pageId="0" pageNumber="1">A, B</emphasis>
|
||
syntype AMNH 42923 from Siam
|
||
<emphasis bold="true" pageId="0" pageNumber="1">A</emphasis>
|
||
figured specimen in
|
||
<bibRefCitation DOI="https://doi.org/10.1111/j.1749-6632.1858.tb00358.x" author="Haines, WA" journalOrPublisher="Nature and Life in Southeast Asia, Kyoto" pageId="0" pageNumber="1" refId="B54" refString="Haines, WA, 1855. Descriptions of four new species of terrestrial shells from Siam. Annals of the Lyceum of Natural History of New York 6: 157, 158. https://doi.org/10.1111/j.1749-6632.1858.tb00358.x" title="Descriptions of four new species of terrestrial shells from Siam. Annals of the Lyceum of Natural History of New York 6: 157, 158." url="https://doi.org/10.1111/j.1749-6632.1858.tb00358.x" year="1855">Haines (1855</bibRefCitation>
|
||
: figs 12, 13) anb
|
||
<emphasis bold="true" pageId="0" pageNumber="1">B</emphasis>
|
||
other specimen from the same syntype lot
|
||
<emphasis bold="true" pageId="0" pageNumber="1">C</emphasis>
|
||
specimens CUMZ 4383 from Tham Phitsadan, Chumphon
|
||
<emphasis bold="true" pageId="0" pageNumber="1">D</emphasis>
|
||
specimen CUMZ 3982 from Tham Sriwilai, Saraburi
|
||
<emphasis bold="true" pageId="0" pageNumber="1">E</emphasis>
|
||
specimen CUMZ 4384 from Pak Chong, Nakhon Ratchasrima
|
||
<emphasis bold="true" pageId="0" pageNumber="1">F</emphasis>
|
||
specimen CUMZ 4879 from Hua Hin, Prachuap Khiri Khan.
|
||
</paragraph>
|
||
</caption>
|
||
<caption doi="10.3897/zookeys.1142.90097.figure20" httpUri="https://binary.pensoft.net/fig/799878" pageId="0" pageNumber="1" start="Figure 20" startId="F20">
|
||
<paragraph pageId="0" pageNumber="1">
|
||
<emphasis bold="true" pageId="0" pageNumber="1">Figure 20.</emphasis>
|
||
Drawing of general anatomy of
|
||
<taxonomicName baseAuthorityName="Haines" baseAuthorityYear="1855" class="Gastropoda" family="Cyclophoridae" genus="Rhiostoma" kingdom="Animalia" lsidName="Rhiostoma housei" order="Architaenioglossa" pageId="0" pageNumber="1" phylum="Mollusca" rank="species" species="housei">
|
||
<emphasis italics="true" pageId="0" pageNumber="1">Rhiostoma housei</emphasis>
|
||
</taxonomicName>
|
||
specimen CUMZ 3982 from Saraburi Province, Thailand.
|
||
<emphasis bold="true" pageId="0" pageNumber="1">A</emphasis>
|
||
female
|
||
<emphasis bold="true" pageId="0" pageNumber="1">B</emphasis>
|
||
male. Abbreviations:
|
||
<emphasis bold="true" pageId="0" pageNumber="1">bc</emphasis>
|
||
, bursa copulatrix;;
|
||
<emphasis bold="true" pageId="0" pageNumber="1">cm</emphasis>
|
||
, columellar muscle;
|
||
<emphasis bold="true" pageId="0" pageNumber="1">ct</emphasis>
|
||
, cephalic tentacle;
|
||
<emphasis bold="true" pageId="0" pageNumber="1">dg</emphasis>
|
||
, digestive gland;
|
||
<emphasis bold="true" pageId="0" pageNumber="1">es</emphasis>
|
||
, eyespot;
|
||
<emphasis bold="true" pageId="0" pageNumber="1">f</emphasis>
|
||
, foot;
|
||
<emphasis bold="true" pageId="0" pageNumber="1">fc</emphasis>
|
||
, faeces;
|
||
<emphasis bold="true" pageId="0" pageNumber="1">int</emphasis>
|
||
, intestine;
|
||
<emphasis bold="true" pageId="0" pageNumber="1">lc</emphasis>
|
||
, lung cavity;
|
||
<emphasis bold="true" pageId="0" pageNumber="1">mc</emphasis>
|
||
, mantle collar;
|
||
<emphasis bold="true" pageId="0" pageNumber="1">od</emphasis>
|
||
, oviduct;
|
||
<emphasis bold="true" pageId="0" pageNumber="1">op</emphasis>
|
||
, operculum;
|
||
<emphasis bold="true" pageId="0" pageNumber="1">ov</emphasis>
|
||
, ovary;
|
||
<emphasis bold="true" pageId="0" pageNumber="1">p</emphasis>
|
||
, external penis;
|
||
<emphasis bold="true" pageId="0" pageNumber="1">pg</emphasis>
|
||
, prostate gland;
|
||
<emphasis bold="true" pageId="0" pageNumber="1">sg</emphasis>
|
||
, sperm groove;
|
||
<emphasis bold="true" pageId="0" pageNumber="1">sn</emphasis>
|
||
, snout;
|
||
<emphasis bold="true" pageId="0" pageNumber="1">te</emphasis>
|
||
, testis;
|
||
<emphasis bold="true" pageId="0" pageNumber="1">ut</emphasis>
|
||
, uterus;
|
||
<emphasis bold="true" pageId="0" pageNumber="1">vd</emphasis>
|
||
, vas deferens;
|
||
<emphasis bold="true" pageId="0" pageNumber="1">vg</emphasis>
|
||
, vaginal groove.
|
||
</paragraph>
|
||
</caption>
|
||
</subSubSection>
|
||
<subSubSection pageId="0" pageNumber="1" type="remarks">
|
||
<paragraph pageId="0" pageNumber="1">Remarks.</paragraph>
|
||
<paragraph pageId="0" pageNumber="1">
|
||
Previous literature recorded
|
||
<taxonomicName lsidName="R. housei" pageId="0" pageNumber="1" rank="species" species="housei">
|
||
<emphasis italics="true" pageId="0" pageNumber="1">R. housei</emphasis>
|
||
</taxonomicName>
|
||
from several places in Laos, Cambodia, and Vietnam [Tonkin] (
|
||
<bibRefCitation author="Moellendorff, von OF" journalOrPublisher="Jahrbuecher der Deutschen Malakozoologischen Gesellschaft" pageId="0" pageNumber="1" pagination="349 - 398" refId="B95" refString="Moellendorff, von OF, 1885. Materialien zur fauna von China (Die Auriculaceen, Nachtraege und Berichtigungen). Jahrbuecher der Deutschen Malakozoologischen Gesellschaft 12: 349 - 398" title="Materialien zur fauna von China (Die Auriculaceen, Nachtraege und Berichtigungen)." volume="12" year="1885">
|
||
<normalizedToken originalValue="Möllendorff">Moellendorff</normalizedToken>
|
||
1885
|
||
</bibRefCitation>
|
||
;
|
||
<bibRefCitation author="Ancey, C-F" journalOrPublisher="Bulletin du Musee ďHistoire Naturelle de Marseille" pageId="0" pageNumber="1" pagination="125 - 150" refId="B3" refString="Ancey, C-F, 1898. Observations sur les mollusques terrestres et fluviatiles recueillies dans I'Indo-Chine et particulaierement au Laos, par M. Henri Counillon, accompagnees de remarques sur ďautres mollusques de cette region. Bulletin du Musee ďHistoire Naturelle de Marseille 1: 125 - 150" title="Observations sur les mollusques terrestres et fluviatiles recueillies dans I'Indo-Chine et particulaierement au Laos, par M. Henri Counillon, accompagnees de remarques sur ďautres mollusques de cette region." volume="1" year="1898">Ancey 1898</bibRefCitation>
|
||
;
|
||
<bibRefCitation author="Fischer, P-H" journalOrPublisher="Journal de Conchyliologie" pageId="0" pageNumber="1" pagination="40 - 64" refId="B41" refString="Fischer, P-H, 1973. Les mollusques testaces du Cambodge, prmiere partie: Introduction et Gastropodes Prosobranches. Journal de Conchyliologie 90: 40 - 64" title="Les mollusques testaces du Cambodge, prmiere partie: Introduction et Gastropodes Prosobranches." volume="90" year="1973">Fischer 1973</bibRefCitation>
|
||
). However, these records seem ambiguous because the type specimens have never been examined, were very limited in the number of specimens, and many species have a similar shell morphology to
|
||
<taxonomicName lsidName="R. housei" pageId="0" pageNumber="1" rank="species" species="housei">
|
||
<emphasis italics="true" pageId="0" pageNumber="1">R. housei</emphasis>
|
||
</taxonomicName>
|
||
. Moreover, the species delimitation by previous authors is probably different from the concept of the type specimens. For example, this species has been reported in Chiangdao, Chiangmai, northern Thailand (sensu
|
||
<bibRefCitation author="Solem, A" journalOrPublisher="Sapolia Zoologica Musei Hauniensis" pageId="0" pageNumber="1" pagination="1 - 110" refId="B155" refString="Solem, A, 1966. Some non-marine mollusks from Thailand, with notes on classification of the Helicarionidae. Sapolia Zoologica Musei Hauniensis 24: 1 - 110" title="Some non-marine mollusks from Thailand, with notes on classification of the Helicarionidae." volume="24" year="1966">Solem 1966</bibRefCitation>
|
||
). We collected many specimens from several populations in the same area and examined
|
||
<normalizedToken originalValue="Solem’s">Solem's</normalizedToken>
|
||
specimens in ZMUC. Due to the clear differences in shell characters, we recognise these populations as distinct species (
|
||
<taxonomicName lsidName="R. tigrina" pageId="0" pageNumber="1" rank="species" species="tigrina">
|
||
<emphasis italics="true" pageId="0" pageNumber="1">R. tigrina</emphasis>
|
||
</taxonomicName>
|
||
sp. nov.). For that reason, the distribution range of
|
||
<taxonomicName lsidName="R. housei" pageId="0" pageNumber="1" rank="species" species="housei">
|
||
<emphasis italics="true" pageId="0" pageNumber="1">R. housei</emphasis>
|
||
</taxonomicName>
|
||
, especially from Laos, Cambodia, and Vietnam, needs to be re-examined with new specimen collections with more precise locality.
|
||
</paragraph>
|
||
<paragraph pageId="0" pageNumber="1">
|
||
<bibRefCitation DOI="https://doi.org/10.35885/ruthenica.2021.30(3).4" author="Do, DS" journalOrPublisher="Ruthenica" pageId="0" pageNumber="1" pagination="165 - 172" refId="B36" refString="Do, DS, Nguyen, TS, Do, HL, 2020b. The land snail genus Rhiostoma Benson, 1860 from Vietnam (Gastropoda: Caenogastropoda: Cyclophoridae), with description of a new species. Ruthenica 30 (3): 165 - 172, DOI: https://doi.org/10.35885/ruthenica.2021.30(3).4" title="The land snail genus Rhiostoma Benson, 1860 from Vietnam (Gastropoda: Caenogastropoda: Cyclophoridae), with description of a new species." url="https://doi.org/10.35885/ruthenica.2021.30(3).4" volume="30" year="2020 b">Do et al. (2020b)</bibRefCitation>
|
||
reported several "
|
||
<taxonomicName lsidName="R. housei" pageId="0" pageNumber="1" rank="species" species="housei">
|
||
<emphasis italics="true" pageId="0" pageNumber="1">R. housei</emphasis>
|
||
</taxonomicName>
|
||
" populations from Northern Vietnam in Dien Bien, Hao Binh, and Son La provinces. The figured specimen from Dien Bien Province shows the distance ~800 km away from the central population of this species from central Thailand. This specimen (
|
||
<bibRefCitation DOI="https://doi.org/10.35885/ruthenica.2021.30(3).4" author="Do, DS" journalOrPublisher="Ruthenica" pageId="0" pageNumber="1" pagination="165 - 172" refId="B36" refString="Do, DS, Nguyen, TS, Do, HL, 2020b. The land snail genus Rhiostoma Benson, 1860 from Vietnam (Gastropoda: Caenogastropoda: Cyclophoridae), with description of a new species. Ruthenica 30 (3): 165 - 172, DOI: https://doi.org/10.35885/ruthenica.2021.30(3).4" title="The land snail genus Rhiostoma Benson, 1860 from Vietnam (Gastropoda: Caenogastropoda: Cyclophoridae), with description of a new species." url="https://doi.org/10.35885/ruthenica.2021.30(3).4" volume="30" year="2020 b">Do et al. 2020b</bibRefCitation>
|
||
: fig. 6b, c) had a depressed spire, and a curved, tubular breathing device, and is probably
|
||
<taxonomicName lsidName="R. simplicilabre" pageId="0" pageNumber="1" rank="species" species="simplicilabre">
|
||
<emphasis italics="true" pageId="0" pageNumber="1">R. simplicilabre</emphasis>
|
||
</taxonomicName>
|
||
rather than
|
||
<taxonomicName lsidName="R. housei" pageId="0" pageNumber="1" rank="species" species="housei">
|
||
<emphasis italics="true" pageId="0" pageNumber="1">R. housei</emphasis>
|
||
</taxonomicName>
|
||
.
|
||
</paragraph>
|
||
<paragraph pageId="0" pageNumber="1">
|
||
The specimens collected from southern Thailand (Chumporn, Phetchaburi, and Prachuapkhirikhan provinces) tended to have longer detached whorls, longer breathing devices, and brighter shell colour than the populations from central Thailand (typical). However, the shell colour pattern, detached whorl, and breathing device structures are identical to the type specimens (Fig.
|
||
<figureCitation captionStart="Figure 19" captionStartId="F19" captionText="Figure 19. Shell of Rhiostoma housei A, B syntype AMNH 42923 from Siam A figured specimen in Haines (1855: figs 12, 13) anb B other specimen from the same syntype lot C specimens CUMZ 4383 from Tham Phitsadan, Chumphon D specimen CUMZ 3982 from Tham Sriwilai, Saraburi E specimen CUMZ 4384 from Pak Chong, Nakhon Ratchasrima F specimen CUMZ 4879 from Hua Hin, Prachuap Khiri Khan." figureDoi="10.3897/zookeys.1142.90097.figure19" httpUri="https://binary.pensoft.net/fig/799877" pageId="0" pageNumber="1">19A, B</figureCitation>
|
||
). Therefore, these are attributed to morphological variation across the wide distribution range of this species.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |