603 lines
55 KiB
XML
603 lines
55 KiB
XML
<document ID-DOI="http://dx.doi.org/10.3897/zse.96.56885" ID-GBIF-Dataset="d6e6b764-5951-4cbe-96d0-50b49552970a" ID-GBIF-Taxon="170420543" ID-Pensoft-Pub="1860-0743-2-769" ID-Pensoft-UUID="BEE6C29A7DC253BBB1596C3FC268CAC5" ID-ZooBank="4D35899104044FF9A9BD0E9288B5E355" LSID="urn:lsid:zoobank.org:act:C6FF5DD9-BE35-4AEA-A049-3BB24048D9DB" ModsDocID="1860-0743-2-769" checkinTime="1605866846107" checkinUser="pensoft" docAuthor="Novak, Janos, Lorenz, Michelle & Harms, Danilo" docDate="2020" docId="544F3DEF88AF5097B055C5FB8E8F6274" docLanguage="en" docName="ZoosystEvol 96(2): 769-779" docOrigin="Zoosystematics and Evolution 96 (2)" docSource="http://dx.doi.org/10.3897/zse.96.56885" docTitle="Feaella (Tetrafeaella) obscura Novák & Lorenz & Harms 2020, sp. nov." docType="treatment" docUuid="C6FF5DD9-BE35-4AEA-A049-3BB24048D9DB" docUuidSource="ZooBank" docVersion="3" id="BEE6C29A7DC253BBB1596C3FC268CAC5" lastPageNumber="769" masterDocId="BEE6C29A7DC253BBB1596C3FC268CAC5" masterDocTitle="Feaella (Tetrafeaella) obscura sp. nov. - a new pseudoscorpion species from the Maldives (Arachnida, Pseudoscorpiones), and an updated identification key to the subgenus Feaella (Tetrafeaella)" masterLastPageNumber="779" masterPageNumber="769" pageId="0" pageNumber="769" updateTime="1643451882518" updateUser="ExternalLinkService">
|
||
<mods:mods xmlns:mods="http://www.loc.gov/mods/v3">
|
||
<mods:titleInfo>
|
||
<mods:title>Feaella (Tetrafeaella) obscura sp. nov. - a new pseudoscorpion species from the Maldives (Arachnida, Pseudoscorpiones), and an updated identification key to the subgenus Feaella (Tetrafeaella)</mods:title>
|
||
</mods:titleInfo>
|
||
<mods:name type="personal">
|
||
<mods:role>
|
||
<mods:roleTerm>Author</mods:roleTerm>
|
||
</mods:role>
|
||
<mods:namePart>Novak, Janos</mods:namePart>
|
||
<mods:affiliation>Hungarian Natural History Museum, Department of Zoology, Baross u. 13, H- 1088 Budapest, Hungary</mods:affiliation>
|
||
<mods:nameIdentifier type="email">novakjanos01@gmail.com</mods:nameIdentifier>
|
||
</mods:name>
|
||
<mods:name type="personal">
|
||
<mods:role>
|
||
<mods:roleTerm>Author</mods:roleTerm>
|
||
</mods:role>
|
||
<mods:namePart>Lorenz, Michelle</mods:namePart>
|
||
<mods:affiliation>Zoological Museum, Center of Natural History, Universitaet Hamburg, Martin-Luther-King-Platz 3, 20146 Hamburg, Germany</mods:affiliation>
|
||
</mods:name>
|
||
<mods:name type="personal">
|
||
<mods:role>
|
||
<mods:roleTerm>Author</mods:roleTerm>
|
||
</mods:role>
|
||
<mods:namePart>Harms, Danilo</mods:namePart>
|
||
<mods:affiliation>Zoological Museum, Center of Natural History, Universitaet Hamburg, Martin-Luther-King-Platz 3, 20146 Hamburg, Germany</mods:affiliation>
|
||
</mods:name>
|
||
<mods:typeOfResource>text</mods:typeOfResource>
|
||
<mods:relatedItem type="host">
|
||
<mods:titleInfo>
|
||
<mods:title>Zoosystematics and Evolution</mods:title>
|
||
</mods:titleInfo>
|
||
<mods:part>
|
||
<mods:date>2020</mods:date>
|
||
<mods:detail type="volume">
|
||
<mods:number>96</mods:number>
|
||
</mods:detail>
|
||
<mods:detail type="issue">
|
||
<mods:number>2</mods:number>
|
||
</mods:detail>
|
||
<mods:extent unit="page">
|
||
<mods:start>769</mods:start>
|
||
<mods:end>779</mods:end>
|
||
</mods:extent>
|
||
</mods:part>
|
||
</mods:relatedItem>
|
||
<mods:location>
|
||
<mods:url>http://dx.doi.org/10.3897/zse.96.56885</mods:url>
|
||
</mods:location>
|
||
<mods:classification>journal article</mods:classification>
|
||
<mods:identifier type="DOI">http://dx.doi.org/10.3897/zse.96.56885</mods:identifier>
|
||
<mods:identifier type="Pensoft-Pub">1860-0743-2-769</mods:identifier>
|
||
<mods:identifier type="ZooBank">4D35899104044FF9A9BD0E9288B5E355</mods:identifier>
|
||
<mods:identifier type="Pensoft-UUID">BEE6C29A7DC253BBB1596C3FC268CAC5</mods:identifier>
|
||
</mods:mods>
|
||
<treatment ID-GBIF-Taxon="170420543" LSID="urn:lsid:zoobank.org:act:C6FF5DD9-BE35-4AEA-A049-3BB24048D9DB" httpUri="http://treatment.plazi.org/id/544F3DEF88AF5097B055C5FB8E8F6274" lastPageNumber="769" pageId="0" pageNumber="769">
|
||
<subSubSection pageId="0" pageNumber="769" type="nomenclature">
|
||
<paragraph pageId="0" pageNumber="769">
|
||
<taxonomicName LSID="544F3DEF-88AF-5097-B055-C5FB8E8F6274" authority="Novák & Lorenz & Harms, 2020" authorityName="Novák & Lorenz & Harms" authorityYear="2020" class="Arachnida" family="Feaellidae" genus="Feaella" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Feaella (Tetrafeaella) obscura" order="Pseudoscorpiones" pageId="0" pageNumber="769" phylum="Arthropoda" rank="species" species="obscura" status="sp. nov." subGenus="Tetrafeaella">Feaella (Tetrafeaella) obscura</taxonomicName>
|
||
<taxonomicNameLabel pageId="0" pageNumber="769">sp. nov.</taxonomicNameLabel>
|
||
<figureCitation captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. Feaella (Tetrafeaella) obscura sp. nov, female holotype (HNHM Pseud- 2009). A. Body, dorsal view; B. Body, ventral view; C. Body, lateral view; D. Carapace, dorsal view; E. Coxal region; F. Left pedipalp, dorsal view." figureDoi="10.3897/zse.96.56885.figure2" httpUri="https://binary.pensoft.net/fig/477313" pageId="0" pageNumber="769">Figs 2A-F</figureCitation>
|
||
<figureCitation captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. Feaella (Tetrafeaella) obscura sp. nov, female holotype (HNHM Pseud- 2009): SEM images. A. Body, dorsal view; B. Coxal region; C. Coxae I and II with medioposterior depression and coxal spines (primary coxal spines of coxae I highlighted in purple); D. Anal region; E. Carapace, dorsal view; F. Female genital region with pedal coxae IV; G. Right chelal tip, dorsal view; H. Specialised, lanceolate setae (marked with arrow) midway between t and finger tip." figureDoi="10.3897/zse.96.56885.figure3" httpUri="https://binary.pensoft.net/fig/477314" pageId="0" pageNumber="769">, 3A-H</figureCitation>
|
||
<figureCitation captionStart="Figure 4" captionStartId="F4" captionText="Figure 4. Feaella (Tetrafeaella) obscura sp. nov, female holotype (HNHM Pseud- 2009): SEM images. A. Left chela, retrolateral view; B. Left chela, dorsomedial view; C. Left chela, ventral view. Male paratype (HNHM Pseud- 2010): SEM images. D. Left chela, retrolateral view; E. Left chela, ventral view; F. Left chela, medial view." figureDoi="10.3897/zse.96.56885.figure4" httpUri="https://binary.pensoft.net/fig/477315" pageId="0" pageNumber="769">, 4A-F</figureCitation>
|
||
<figureCitation captionStart="Figure 5" captionStartId="F5" captionText="Figure 5. Feaella (Tetrafeaella) obscura sp. nov. A. Right chelicera, dorsal view - female holotype (HNHM Pseud- 2009); B. Right pedipalp, dorsal view - female holotype; C. Left chela, retrolateral view - female holotype; D. Leg I, lateral view - female paratype (ZMH-A 0003101); E. Leg IV, lateral view - female paratype; F. Female genital region - female holotype; G. Male genital region - male paratype (HNHM Pseud- 2010)." figureDoi="10.3897/zse.96.56885.figure5" httpUri="https://binary.pensoft.net/fig/477316" pageId="0" pageNumber="769">, 5A-G</figureCitation>
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection pageId="0" pageNumber="769" type="type material">
|
||
<paragraph pageId="0" pageNumber="769">Type material.</paragraph>
|
||
<paragraph pageId="0" pageNumber="769">
|
||
Holotype female from the Maldives, Kabudu Island (might refer to Kaudu [
|
||
<geoCoordinate degrees="0" direction="north" minutes="17" orientation="latitude" precision="925" value="0.28333333">0°17'N</geoCoordinate>
|
||
,
|
||
<geoCoordinate degrees="73" direction="east" minutes="1" orientation="longitude" precision="925" value="73.01667">73°1'E</geoCoordinate>
|
||
] or Kandudu Islands [
|
||
<geoCoordinate degrees="2" direction="north" minutes="19" orientation="latitude" precision="925" value="2.3166666">2°19'N</geoCoordinate>
|
||
,
|
||
<geoCoordinate degrees="72" direction="east" minutes="55" orientation="longitude" precision="925" value="72.916664">72°55'E</geoCoordinate>
|
||
]), June 1984, leg:
|
||
<normalizedToken originalValue="Győző">Gyozo</normalizedToken>
|
||
<normalizedToken originalValue="Horváth">Horvath</normalizedToken>
|
||
. [On the label: Maldives, Kabudu sz., 1984.VII. leg.:
|
||
<normalizedToken originalValue="Győző">Gyozo</normalizedToken>
|
||
<normalizedToken originalValue="Horváth">Horvath</normalizedToken>
|
||
] (HNHM Pseud-2009). Paratypes: 1 male (HNHM Pseud-2010) and 1 female (ZMH-A0003101); all same data as holotype.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection pageId="0" pageNumber="769" type="etymology">
|
||
<paragraph pageId="0" pageNumber="769">Etymology.</paragraph>
|
||
<paragraph pageId="0" pageNumber="769">The name of the species refers to the obscure evolutionary and geographic origins of the species that is unlikely to have evolved on the young island that is the locus typicus.</paragraph>
|
||
</subSubSection>
|
||
<subSubSection pageId="0" pageNumber="769" type="distribution">
|
||
<paragraph pageId="0" pageNumber="769">Distribution.</paragraph>
|
||
<paragraph pageId="0" pageNumber="769">
|
||
Maldive Islands (Fig.
|
||
<figureCitation captionStart="Figure 1" captionStartId="F1" captionText="Figure 1. Known distribution of the family Feaellidae. Circle colours: blue = Recent records; green = fossil records; and red = F. (T.) obscura sp. nov." figureDoi="10.3897/zse.96.56885.figure1" httpUri="https://binary.pensoft.net/fig/477312" pageId="0" pageNumber="769">1</figureCitation>
|
||
).
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection pageId="0" pageNumber="769" type="diagnosis">
|
||
<paragraph pageId="0" pageNumber="769">Diagnosis.</paragraph>
|
||
<paragraph pageId="0" pageNumber="769">
|
||
A typical
|
||
<taxonomicName authority="" class="Arachnida" family="Feaellidae" genus="Feaella" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Feaella (Tetrafeaella)" order="Pseudoscorpiones" pageId="0" pageNumber="769" phylum="Arthropoda" rank="subGenus" subGenus="Tetrafeaella">Feaella (Tetrafeaella)</taxonomicName>
|
||
habitus (Figs
|
||
<figureCitation captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. Feaella (Tetrafeaella) obscura sp. nov, female holotype (HNHM Pseud- 2009). A. Body, dorsal view; B. Body, ventral view; C. Body, lateral view; D. Carapace, dorsal view; E. Coxal region; F. Left pedipalp, dorsal view." figureDoi="10.3897/zse.96.56885.figure2" httpUri="https://binary.pensoft.net/fig/477313" pageId="0" pageNumber="769">2A, B</figureCitation>
|
||
,
|
||
<figureCitation captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. Feaella (Tetrafeaella) obscura sp. nov, female holotype (HNHM Pseud- 2009): SEM images. A. Body, dorsal view; B. Coxal region; C. Coxae I and II with medioposterior depression and coxal spines (primary coxal spines of coxae I highlighted in purple); D. Anal region; E. Carapace, dorsal view; F. Female genital region with pedal coxae IV; G. Right chelal tip, dorsal view; H. Specialised, lanceolate setae (marked with arrow) midway between t and finger tip." figureDoi="10.3897/zse.96.56885.figure3" httpUri="https://binary.pensoft.net/fig/477314" pageId="0" pageNumber="769">3A</figureCitation>
|
||
) that differs from all the other species of the subgenus by the following combination of characters: with the anteromedian lobes of carapace being closer to each other than to the anterolateral ones; anterolateral pair approximately as broad as anteromedian pair, anteromedian lobes longer. Palpal trochanter with prolateral triangular protuberance, femur very robust (1.70-1.76
|
||
<normalizedToken originalValue="×">x</normalizedToken>
|
||
) with two pronounced triangular process on prolateral corner near base. Chelae are approximately as long as the palpal femora. Chelae with dorsal protuberance at basal third of finger, and a large medial tooth at the base of each fingers. Chelal trichobothriotaxy:
|
||
<emphasis italics="true" pageId="0" pageNumber="769">esb</emphasis>
|
||
and
|
||
<emphasis italics="true" pageId="0" pageNumber="769">est</emphasis>
|
||
situated on basal third of retrolateral face;
|
||
<emphasis italics="true" pageId="0" pageNumber="769">ib</emphasis>
|
||
,
|
||
<emphasis italics="true" pageId="0" pageNumber="769">isb</emphasis>
|
||
and
|
||
<emphasis italics="true" pageId="0" pageNumber="769">ist</emphasis>
|
||
situated basally;
|
||
<emphasis italics="true" pageId="0" pageNumber="769">eb</emphasis>
|
||
and
|
||
<emphasis italics="true" pageId="0" pageNumber="769">it</emphasis>
|
||
situated subdistally;
|
||
<emphasis italics="true" pageId="0" pageNumber="769">dt</emphasis>
|
||
situated distally;
|
||
<emphasis italics="true" pageId="0" pageNumber="769">esb</emphasis>
|
||
midway between
|
||
<emphasis italics="true" pageId="0" pageNumber="769">est</emphasis>
|
||
and
|
||
<emphasis italics="true" pageId="0" pageNumber="769">isb</emphasis>
|
||
;
|
||
<emphasis italics="true" pageId="0" pageNumber="769">ist</emphasis>
|
||
equidistant from
|
||
<emphasis italics="true" pageId="0" pageNumber="769">est</emphasis>
|
||
and
|
||
<emphasis italics="true" pageId="0" pageNumber="769">esb</emphasis>
|
||
;
|
||
<emphasis italics="true" pageId="0" pageNumber="769">isb</emphasis>
|
||
is closer to
|
||
<emphasis italics="true" pageId="0" pageNumber="769">ib</emphasis>
|
||
than to
|
||
<emphasis italics="true" pageId="0" pageNumber="769">ist</emphasis>
|
||
; on the movable chelal finger
|
||
<emphasis italics="true" pageId="0" pageNumber="769">t</emphasis>
|
||
and
|
||
<emphasis italics="true" pageId="0" pageNumber="769">sb</emphasis>
|
||
are situated midway between
|
||
<emphasis italics="true" pageId="0" pageNumber="769">b</emphasis>
|
||
and
|
||
<emphasis italics="true" pageId="0" pageNumber="769">st.</emphasis>
|
||
Pedal coxa I with medioposterior depression (coxal pit), a single primary coxal spine posteriorly; coxa II with 9 secondary spines in males and 8-11 in females. Pleural membrane with a dorsal row of 15 and a ventral row of 14 sclerotised pleural platelets. See detailed differentiation from the most similar species in the Differential diagnosis, and differentiation from all the other species of the subgenus in the key.
|
||
</paragraph>
|
||
<caption doi="10.3897/zse.96.56885.figure2" httpUri="https://binary.pensoft.net/fig/477313" pageId="0" pageNumber="769" start="Figure 2" startId="F2">
|
||
<paragraph pageId="0" pageNumber="769">
|
||
<emphasis bold="true" pageId="0" pageNumber="769">Figure 2.</emphasis>
|
||
<taxonomicName authorityName="Novák & Lorenz & Harms" authorityYear="2020" class="Arachnida" family="Feaellidae" genus="Feaella" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Feaella (Tetrafeaella) obscura" order="Pseudoscorpiones" pageId="0" pageNumber="769" phylum="Arthropoda" rank="species" species="obscura" subGenus="Tetrafeaella">Feaella (Tetrafeaella) obscura</taxonomicName>
|
||
sp. nov, female holotype (HNHM Pseud-2009).
|
||
<emphasis bold="true" pageId="0" pageNumber="769">A.</emphasis>
|
||
Body, dorsal view;
|
||
<emphasis bold="true" pageId="0" pageNumber="769">B.</emphasis>
|
||
Body, ventral view;
|
||
<emphasis bold="true" pageId="0" pageNumber="769">C.</emphasis>
|
||
Body, lateral view;
|
||
<emphasis bold="true" pageId="0" pageNumber="769">D.</emphasis>
|
||
Carapace, dorsal view;
|
||
<emphasis bold="true" pageId="0" pageNumber="769">E.</emphasis>
|
||
Coxal region;
|
||
<emphasis bold="true" pageId="0" pageNumber="769">F.</emphasis>
|
||
Left pedipalp, dorsal view.
|
||
</paragraph>
|
||
</caption>
|
||
</subSubSection>
|
||
<subSubSection pageId="0" pageNumber="769" type="description">
|
||
<paragraph pageId="0" pageNumber="769">Description.</paragraph>
|
||
<paragraph pageId="0" pageNumber="769">
|
||
(Holotype female, paratype male and female).
|
||
<emphasis italics="true" pageId="0" pageNumber="769">Colour and cuticular surface</emphasis>
|
||
(
|
||
<emphasis italics="true" pageId="0" pageNumber="769">in alcohol</emphasis>
|
||
): sclerotised parts reddish-brown, with reticulated cerotegument (Fig.
|
||
<figureCitation captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. Feaella (Tetrafeaella) obscura sp. nov, female holotype (HNHM Pseud- 2009). A. Body, dorsal view; B. Body, ventral view; C. Body, lateral view; D. Carapace, dorsal view; E. Coxal region; F. Left pedipalp, dorsal view." figureDoi="10.3897/zse.96.56885.figure2" httpUri="https://binary.pensoft.net/fig/477313" pageId="0" pageNumber="769">2A-F</figureCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph pageId="0" pageNumber="769">
|
||
<emphasis bold="true" italics="true" pageId="0" pageNumber="769">Setae</emphasis>
|
||
: most vestitural setae short, slightly curved, inconspicuous and acuminate.
|
||
</paragraph>
|
||
<paragraph pageId="0" pageNumber="769">
|
||
<emphasis bold="true" italics="true" pageId="0" pageNumber="769">Chelicera</emphasis>
|
||
(Fig.
|
||
<figureCitation captionStart="Figure 5" captionStartId="F5" captionText="Figure 5. Feaella (Tetrafeaella) obscura sp. nov. A. Right chelicera, dorsal view - female holotype (HNHM Pseud- 2009); B. Right pedipalp, dorsal view - female holotype; C. Left chela, retrolateral view - female holotype; D. Leg I, lateral view - female paratype (ZMH-A 0003101); E. Leg IV, lateral view - female paratype; F. Female genital region - female holotype; G. Male genital region - male paratype (HNHM Pseud- 2010)." figureDoi="10.3897/zse.96.56885.figure5" httpUri="https://binary.pensoft.net/fig/477316" pageId="0" pageNumber="769">5A</figureCitation>
|
||
): hand with five setae,
|
||
<emphasis italics="true" pageId="0" pageNumber="769">is</emphasis>
|
||
and
|
||
<emphasis italics="true" pageId="0" pageNumber="769">ls</emphasis>
|
||
adjacent to each other, movable finger with subdistal seta. Galea thick, without rami. Fingers without teeth. Serrula exterior with ca. 16 blades, lamina exterior absent. Rallum absent.
|
||
</paragraph>
|
||
<paragraph pageId="0" pageNumber="769">
|
||
<emphasis bold="true" italics="true" pageId="0" pageNumber="769">Pedipalps</emphasis>
|
||
(Figs
|
||
<figureCitation captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. Feaella (Tetrafeaella) obscura sp. nov, female holotype (HNHM Pseud- 2009). A. Body, dorsal view; B. Body, ventral view; C. Body, lateral view; D. Carapace, dorsal view; E. Coxal region; F. Left pedipalp, dorsal view." figureDoi="10.3897/zse.96.56885.figure2" httpUri="https://binary.pensoft.net/fig/477313" pageId="0" pageNumber="769">2F</figureCitation>
|
||
,
|
||
<figureCitation captionStart="Figure 5" captionStartId="F5" captionText="Figure 5. Feaella (Tetrafeaella) obscura sp. nov. A. Right chelicera, dorsal view - female holotype (HNHM Pseud- 2009); B. Right pedipalp, dorsal view - female holotype; C. Left chela, retrolateral view - female holotype; D. Leg I, lateral view - female paratype (ZMH-A 0003101); E. Leg IV, lateral view - female paratype; F. Female genital region - female holotype; G. Male genital region - male paratype (HNHM Pseud- 2010)." figureDoi="10.3897/zse.96.56885.figure5" httpUri="https://binary.pensoft.net/fig/477316" pageId="0" pageNumber="769">5B</figureCitation>
|
||
): trochanter with prolateral triangular protuberance, femur very robust with two pronounced triangular process on prolateral corner near base, 0.63-0.69/0.37-0.40 (1.70-1.73
|
||
<normalizedToken originalValue="×">x</normalizedToken>
|
||
) in females and 0.51/0.29 (1.76
|
||
<normalizedToken originalValue="×">x</normalizedToken>
|
||
) in male; patella conical, and with two lyrifissures at base, 0.51-0.54/0.18-0.19 (2.83-2.84
|
||
<normalizedToken originalValue="×">x</normalizedToken>
|
||
) in females and 0.41/0.14 (2.93
|
||
<normalizedToken originalValue="×">x</normalizedToken>
|
||
) in male; chela tubular (Figs
|
||
<figureCitation captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. Feaella (Tetrafeaella) obscura sp. nov, female holotype (HNHM Pseud- 2009): SEM images. A. Body, dorsal view; B. Coxal region; C. Coxae I and II with medioposterior depression and coxal spines (primary coxal spines of coxae I highlighted in purple); D. Anal region; E. Carapace, dorsal view; F. Female genital region with pedal coxae IV; G. Right chelal tip, dorsal view; H. Specialised, lanceolate setae (marked with arrow) midway between t and finger tip." figureDoi="10.3897/zse.96.56885.figure3" httpUri="https://binary.pensoft.net/fig/477314" pageId="0" pageNumber="769">3G</figureCitation>
|
||
,
|
||
<figureCitation captionStart="Figure 4" captionStartId="F4" captionText="Figure 4. Feaella (Tetrafeaella) obscura sp. nov, female holotype (HNHM Pseud- 2009): SEM images. A. Left chela, retrolateral view; B. Left chela, dorsomedial view; C. Left chela, ventral view. Male paratype (HNHM Pseud- 2010): SEM images. D. Left chela, retrolateral view; E. Left chela, ventral view; F. Left chela, medial view." figureDoi="10.3897/zse.96.56885.figure4" httpUri="https://binary.pensoft.net/fig/477315" pageId="0" pageNumber="769">4A-F</figureCitation>
|
||
,
|
||
<figureCitation captionStart="Figure 5" captionStartId="F5" captionText="Figure 5. Feaella (Tetrafeaella) obscura sp. nov. A. Right chelicera, dorsal view - female holotype (HNHM Pseud- 2009); B. Right pedipalp, dorsal view - female holotype; C. Left chela, retrolateral view - female holotype; D. Leg I, lateral view - female paratype (ZMH-A 0003101); E. Leg IV, lateral view - female paratype; F. Female genital region - female holotype; G. Male genital region - male paratype (HNHM Pseud- 2010)." figureDoi="10.3897/zse.96.56885.figure5" httpUri="https://binary.pensoft.net/fig/477316" pageId="0" pageNumber="769">5C</figureCitation>
|
||
), chela (with pedicel) 0.60-0.61 in females and 0.53 in male, chela (without pedicel) 0.57 in females and 0.50 in male; hand without pedicel and longer than broad. Fixed chelal finger and hand with eight major trichobothria, plus duplex trichobothrium (
|
||
<emphasis italics="true" pageId="0" pageNumber="769">dt</emphasis>
|
||
), movable chelal finger with four trichobothria:
|
||
<emphasis italics="true" pageId="0" pageNumber="769">esb</emphasis>
|
||
and
|
||
<emphasis italics="true" pageId="0" pageNumber="769">est</emphasis>
|
||
situated on basal third of retrolateral face;
|
||
<emphasis italics="true" pageId="0" pageNumber="769">ib</emphasis>
|
||
,
|
||
<emphasis italics="true" pageId="0" pageNumber="769">isb</emphasis>
|
||
and
|
||
<emphasis italics="true" pageId="0" pageNumber="769">ist</emphasis>
|
||
situated basally;
|
||
<emphasis italics="true" pageId="0" pageNumber="769">eb</emphasis>
|
||
and
|
||
<emphasis italics="true" pageId="0" pageNumber="769">it</emphasis>
|
||
situated subdistally;
|
||
<emphasis italics="true" pageId="0" pageNumber="769">dt</emphasis>
|
||
situated distally;
|
||
<emphasis italics="true" pageId="0" pageNumber="769">esb</emphasis>
|
||
midway between
|
||
<emphasis italics="true" pageId="0" pageNumber="769">est</emphasis>
|
||
and
|
||
<emphasis italics="true" pageId="0" pageNumber="769">isb</emphasis>
|
||
;
|
||
<emphasis italics="true" pageId="0" pageNumber="769">ist</emphasis>
|
||
equidistant from
|
||
<emphasis italics="true" pageId="0" pageNumber="769">est</emphasis>
|
||
and
|
||
<emphasis italics="true" pageId="0" pageNumber="769">esb</emphasis>
|
||
;
|
||
<emphasis italics="true" pageId="0" pageNumber="769">isb</emphasis>
|
||
is closer to
|
||
<emphasis italics="true" pageId="0" pageNumber="769">ib</emphasis>
|
||
than to
|
||
<emphasis italics="true" pageId="0" pageNumber="769">ist.</emphasis>
|
||
On the movable chelal finger
|
||
<emphasis italics="true" pageId="0" pageNumber="769">t</emphasis>
|
||
and
|
||
<emphasis italics="true" pageId="0" pageNumber="769">sb</emphasis>
|
||
are situated midway between
|
||
<emphasis italics="true" pageId="0" pageNumber="769">b</emphasis>
|
||
and
|
||
<emphasis italics="true" pageId="0" pageNumber="769">st</emphasis>
|
||
(Fig.
|
||
<figureCitation captionStart="Figure 5" captionStartId="F5" captionText="Figure 5. Feaella (Tetrafeaella) obscura sp. nov. A. Right chelicera, dorsal view - female holotype (HNHM Pseud- 2009); B. Right pedipalp, dorsal view - female holotype; C. Left chela, retrolateral view - female holotype; D. Leg I, lateral view - female paratype (ZMH-A 0003101); E. Leg IV, lateral view - female paratype; F. Female genital region - female holotype; G. Male genital region - male paratype (HNHM Pseud- 2010)." figureDoi="10.3897/zse.96.56885.figure5" httpUri="https://binary.pensoft.net/fig/477316" pageId="0" pageNumber="769">5C</figureCitation>
|
||
). Venom apparatus absent. Chelal hand very small; retrolateral condyle rounded; with dorsal protuberance at basal third of finger (Fig.
|
||
<figureCitation captionStart="Figure 5" captionStartId="F5" captionText="Figure 5. Feaella (Tetrafeaella) obscura sp. nov. A. Right chelicera, dorsal view - female holotype (HNHM Pseud- 2009); B. Right pedipalp, dorsal view - female holotype; C. Left chela, retrolateral view - female holotype; D. Leg I, lateral view - female paratype (ZMH-A 0003101); E. Leg IV, lateral view - female paratype; F. Female genital region - female holotype; G. Male genital region - male paratype (HNHM Pseud- 2010)." figureDoi="10.3897/zse.96.56885.figure5" httpUri="https://binary.pensoft.net/fig/477316" pageId="0" pageNumber="769">5C</figureCitation>
|
||
). Chelal teeth large, retrorse and diastemodentate: fixed finger with two parallel marginal dental line, one with 11 and one with 9 teeth, 9 prolateral and 7 distal teeth in male; 12-13 and 9-13 marginal, 9-10 prolateral and 3-4 distal teeth in females; movable finger with two parallel marginal dental line with 9 + 9, 9 prolateral and 7 distal teeth in male; 12 and 9 marginal, 9-10 prolateral and 6 distal teeth in females; plus a large medial tooth at the base of each finger. Movable chelal finger with several specialised, lanceolate setae midway between
|
||
<emphasis italics="true" pageId="0" pageNumber="769">t</emphasis>
|
||
and tip of finger; located in a depression (Fig.
|
||
<figureCitation captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. Feaella (Tetrafeaella) obscura sp. nov, female holotype (HNHM Pseud- 2009): SEM images. A. Body, dorsal view; B. Coxal region; C. Coxae I and II with medioposterior depression and coxal spines (primary coxal spines of coxae I highlighted in purple); D. Anal region; E. Carapace, dorsal view; F. Female genital region with pedal coxae IV; G. Right chelal tip, dorsal view; H. Specialised, lanceolate setae (marked with arrow) midway between t and finger tip." figureDoi="10.3897/zse.96.56885.figure3" httpUri="https://binary.pensoft.net/fig/477314" pageId="0" pageNumber="769">3H</figureCitation>
|
||
).
|
||
</paragraph>
|
||
<caption doi="10.3897/zse.96.56885.figure3" httpUri="https://binary.pensoft.net/fig/477314" pageId="0" pageNumber="769" start="Figure 3" startId="F3">
|
||
<paragraph pageId="0" pageNumber="769">
|
||
<emphasis bold="true" pageId="0" pageNumber="769">Figure 3.</emphasis>
|
||
<taxonomicName authorityName="Novák & Lorenz & Harms" authorityYear="2020" class="Arachnida" family="Feaellidae" genus="Feaella" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Feaella (Tetrafeaella) obscura" order="Pseudoscorpiones" pageId="0" pageNumber="769" phylum="Arthropoda" rank="species" species="obscura" subGenus="Tetrafeaella">Feaella (Tetrafeaella) obscura</taxonomicName>
|
||
sp. nov, female holotype (HNHM Pseud-2009): SEM images.
|
||
<emphasis bold="true" pageId="0" pageNumber="769">A.</emphasis>
|
||
Body, dorsal view;
|
||
<emphasis bold="true" pageId="0" pageNumber="769">B.</emphasis>
|
||
Coxal region;
|
||
<emphasis bold="true" pageId="0" pageNumber="769">C.</emphasis>
|
||
Coxae I and II with medioposterior depression and coxal spines (primary coxal spines of coxae I highlighted in purple);
|
||
<emphasis bold="true" pageId="0" pageNumber="769">D.</emphasis>
|
||
Anal region;
|
||
<emphasis bold="true" pageId="0" pageNumber="769">E.</emphasis>
|
||
Carapace, dorsal view;
|
||
<emphasis bold="true" pageId="0" pageNumber="769">F.</emphasis>
|
||
Female genital region with pedal coxae IV;
|
||
<emphasis bold="true" pageId="0" pageNumber="769">G.</emphasis>
|
||
Right chelal tip, dorsal view;
|
||
<emphasis bold="true" pageId="0" pageNumber="769">H.</emphasis>
|
||
Specialised, lanceolate setae (marked with arrow) midway between
|
||
<emphasis italics="true" pageId="0" pageNumber="769">t</emphasis>
|
||
and finger tip.
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph pageId="0" pageNumber="769">
|
||
<emphasis bold="true" italics="true" pageId="0" pageNumber="769">Carapace</emphasis>
|
||
: anterior margin with four lobes, the anteromedian lobes are closer to each other than to the anterolateral ones; anterolateral pair approximately as broad as anteromedian pair, anteromedian lobes longer (Figs
|
||
<figureCitation captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. Feaella (Tetrafeaella) obscura sp. nov, female holotype (HNHM Pseud- 2009). A. Body, dorsal view; B. Body, ventral view; C. Body, lateral view; D. Carapace, dorsal view; E. Coxal region; F. Left pedipalp, dorsal view." figureDoi="10.3897/zse.96.56885.figure2" httpUri="https://binary.pensoft.net/fig/477313" pageId="0" pageNumber="769">2D</figureCitation>
|
||
,
|
||
<figureCitation captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. Feaella (Tetrafeaella) obscura sp. nov, female holotype (HNHM Pseud- 2009): SEM images. A. Body, dorsal view; B. Coxal region; C. Coxae I and II with medioposterior depression and coxal spines (primary coxal spines of coxae I highlighted in purple); D. Anal region; E. Carapace, dorsal view; F. Female genital region with pedal coxae IV; G. Right chelal tip, dorsal view; H. Specialised, lanceolate setae (marked with arrow) midway between t and finger tip." figureDoi="10.3897/zse.96.56885.figure3" httpUri="https://binary.pensoft.net/fig/477314" pageId="0" pageNumber="769">3E</figureCitation>
|
||
). Four lobes between eyes. Lateral margins of carapace nearly parallel, slightly widened medially; carapace in females 1.27-1.28
|
||
<normalizedToken originalValue="×">x</normalizedToken>
|
||
, in male 1.24
|
||
<normalizedToken originalValue="×">x</normalizedToken>
|
||
longer than broad; with two pairs of eyes situated on tubercles away from anterior carapace margin; all eyes with tapetum; with numerous inconspicuous setae; with a pair of postero-lateral processes; shallow anterior and posterior furrows present. Three slit-like lyrifissures at the level of posterior one-fourth of carapace on each side, and two near posterior base.
|
||
</paragraph>
|
||
<paragraph pageId="0" pageNumber="769">
|
||
<emphasis bold="true" italics="true" pageId="0" pageNumber="769">Coxal region</emphasis>
|
||
(Figs
|
||
<figureCitation captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. Feaella (Tetrafeaella) obscura sp. nov, female holotype (HNHM Pseud- 2009). A. Body, dorsal view; B. Body, ventral view; C. Body, lateral view; D. Carapace, dorsal view; E. Coxal region; F. Left pedipalp, dorsal view." figureDoi="10.3897/zse.96.56885.figure2" httpUri="https://binary.pensoft.net/fig/477313" pageId="0" pageNumber="769">2E</figureCitation>
|
||
,
|
||
<figureCitation captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. Feaella (Tetrafeaella) obscura sp. nov, female holotype (HNHM Pseud- 2009): SEM images. A. Body, dorsal view; B. Coxal region; C. Coxae I and II with medioposterior depression and coxal spines (primary coxal spines of coxae I highlighted in purple); D. Anal region; E. Carapace, dorsal view; F. Female genital region with pedal coxae IV; G. Right chelal tip, dorsal view; H. Specialised, lanceolate setae (marked with arrow) midway between t and finger tip." figureDoi="10.3897/zse.96.56885.figure3" httpUri="https://binary.pensoft.net/fig/477314" pageId="0" pageNumber="769">3B</figureCitation>
|
||
): pedipalpal coxa with strong basal lateral processes; with numerous small setae and 3 acuminate apical setae. Coxa I with medioposterior depression (coxal pit), a single primary coxal spine posteriorly; coxa II with 9 secondary spines in male and 8-11 in females (Fig.
|
||
<figureCitation captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. Feaella (Tetrafeaella) obscura sp. nov, female holotype (HNHM Pseud- 2009): SEM images. A. Body, dorsal view; B. Coxal region; C. Coxae I and II with medioposterior depression and coxal spines (primary coxal spines of coxae I highlighted in purple); D. Anal region; E. Carapace, dorsal view; F. Female genital region with pedal coxae IV; G. Right chelal tip, dorsal view; H. Specialised, lanceolate setae (marked with arrow) midway between t and finger tip." figureDoi="10.3897/zse.96.56885.figure3" httpUri="https://binary.pensoft.net/fig/477314" pageId="0" pageNumber="769">3C</figureCitation>
|
||
). Pedipalpal coxa somewhat longer as combined length of leg coxae I-IV.
|
||
</paragraph>
|
||
<paragraph pageId="0" pageNumber="769">
|
||
<emphasis bold="true" italics="true" pageId="0" pageNumber="769">Legs</emphasis>
|
||
(Fig.
|
||
<figureCitation captionStart="Figure 5" captionStartId="F5" captionText="Figure 5. Feaella (Tetrafeaella) obscura sp. nov. A. Right chelicera, dorsal view - female holotype (HNHM Pseud- 2009); B. Right pedipalp, dorsal view - female holotype; C. Left chela, retrolateral view - female holotype; D. Leg I, lateral view - female paratype (ZMH-A 0003101); E. Leg IV, lateral view - female paratype; F. Female genital region - female holotype; G. Male genital region - male paratype (HNHM Pseud- 2010)." figureDoi="10.3897/zse.96.56885.figure5" httpUri="https://binary.pensoft.net/fig/477316" pageId="0" pageNumber="769">5D, E</figureCitation>
|
||
): claws simple, arolium shorter than claws. Metatarsi and tarsi fused. Each patella with a shallow dorsal depression. Tarsi without tactile setae, vestitural setae short and acuminate. Patellae I and II slightly shorter than femora I and II; patellae III and IV nearly twice as long as femora III and IV.
|
||
</paragraph>
|
||
<paragraph pageId="0" pageNumber="769">
|
||
<emphasis bold="true" italics="true" pageId="0" pageNumber="769">Abdomen</emphasis>
|
||
: longer than broad, somewhat ovoid; tergites II-IX and sternites IV-X with distinct median suture lines; tergite XI and sternite XI fused; tergite XII and sternite XII (anal sclerites) strongly sclerotised; most segments with numerous setae; tergite XII and sternite XII with two setae; anal region with raised circular rim (Fig.
|
||
<figureCitation captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. Feaella (Tetrafeaella) obscura sp. nov, female holotype (HNHM Pseud- 2009): SEM images. A. Body, dorsal view; B. Coxal region; C. Coxae I and II with medioposterior depression and coxal spines (primary coxal spines of coxae I highlighted in purple); D. Anal region; E. Carapace, dorsal view; F. Female genital region with pedal coxae IV; G. Right chelal tip, dorsal view; H. Specialised, lanceolate setae (marked with arrow) midway between t and finger tip." figureDoi="10.3897/zse.96.56885.figure3" httpUri="https://binary.pensoft.net/fig/477314" pageId="0" pageNumber="769">3D</figureCitation>
|
||
). Setae of sternites II and III longer than vestitural setae. Pleural membrane with a dorsal row of 15 and a ventral row of 14 sclerotised pleural platelets.
|
||
</paragraph>
|
||
<paragraph pageId="0" pageNumber="769">
|
||
<emphasis bold="true" italics="true" pageId="0" pageNumber="769">Genital region</emphasis>
|
||
: Female (Figs
|
||
<figureCitation captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. Feaella (Tetrafeaella) obscura sp. nov, female holotype (HNHM Pseud- 2009): SEM images. A. Body, dorsal view; B. Coxal region; C. Coxae I and II with medioposterior depression and coxal spines (primary coxal spines of coxae I highlighted in purple); D. Anal region; E. Carapace, dorsal view; F. Female genital region with pedal coxae IV; G. Right chelal tip, dorsal view; H. Specialised, lanceolate setae (marked with arrow) midway between t and finger tip." figureDoi="10.3897/zse.96.56885.figure3" httpUri="https://binary.pensoft.net/fig/477314" pageId="0" pageNumber="769">3F</figureCitation>
|
||
,
|
||
<figureCitation captionStart="Figure 5" captionStartId="F5" captionText="Figure 5. Feaella (Tetrafeaella) obscura sp. nov. A. Right chelicera, dorsal view - female holotype (HNHM Pseud- 2009); B. Right pedipalp, dorsal view - female holotype; C. Left chela, retrolateral view - female holotype; D. Leg I, lateral view - female paratype (ZMH-A 0003101); E. Leg IV, lateral view - female paratype; F. Female genital region - female holotype; G. Male genital region - male paratype (HNHM Pseud- 2010)." figureDoi="10.3897/zse.96.56885.figure5" httpUri="https://binary.pensoft.net/fig/477316" pageId="0" pageNumber="769">5F</figureCitation>
|
||
): 10 acuminate microsetae on each plate. Male (Fig.
|
||
<figureCitation captionStart="Figure 5" captionStartId="F5" captionText="Figure 5. Feaella (Tetrafeaella) obscura sp. nov. A. Right chelicera, dorsal view - female holotype (HNHM Pseud- 2009); B. Right pedipalp, dorsal view - female holotype; C. Left chela, retrolateral view - female holotype; D. Leg I, lateral view - female paratype (ZMH-A 0003101); E. Leg IV, lateral view - female paratype; F. Female genital region - female holotype; G. Male genital region - male paratype (HNHM Pseud- 2010)." figureDoi="10.3897/zse.96.56885.figure5" httpUri="https://binary.pensoft.net/fig/477316" pageId="0" pageNumber="769">5G</figureCitation>
|
||
): 40-45 acuminate microsetae on genital plate. Inner genital structures could not be clearly seen.
|
||
</paragraph>
|
||
<paragraph pageId="0" pageNumber="769">
|
||
Measurements (in mm, ratios in parentheses):
|
||
<emphasis bold="true" pageId="0" pageNumber="769">Male</emphasis>
|
||
paratype. Body 1.83. Carapace 0.51/0.41 (1.24
|
||
<normalizedToken originalValue="×">x</normalizedToken>
|
||
). Chelicera 0.19/0.10 (1.9
|
||
<normalizedToken originalValue="×">x</normalizedToken>
|
||
), movable finger 0.085. Palpal femur 0.51/0.29 (1.76
|
||
<normalizedToken originalValue="×">x</normalizedToken>
|
||
), patella 0.41/0.14 (2.93
|
||
<normalizedToken originalValue="×">x</normalizedToken>
|
||
), chela (with pedicel) 0.53/0.13 (4.08
|
||
<normalizedToken originalValue="×">x</normalizedToken>
|
||
), chela (without pedicel) 0.50, hand (with pedicel) 0.15, hand (without pedicel) 0.12, movable finger 0.37. Leg I. trochanter 0.12/0.19 (0.62
|
||
<normalizedToken originalValue="×">x</normalizedToken>
|
||
), femur 0.21/0.06 (3.50
|
||
<normalizedToken originalValue="×">x</normalizedToken>
|
||
), patella 0.18/0.07 (2.57
|
||
<normalizedToken originalValue="×">x</normalizedToken>
|
||
), tibia 0.17/0.06 (2.83
|
||
<normalizedToken originalValue="×">x</normalizedToken>
|
||
), tarsus 0.23/0.05 (4.60
|
||
<normalizedToken originalValue="×">x</normalizedToken>
|
||
). Leg IV. trochanter 0.22/0.12 (1.83
|
||
<normalizedToken originalValue="×">x</normalizedToken>
|
||
), femur 0.16/0.08 (2.00
|
||
<normalizedToken originalValue="×">x</normalizedToken>
|
||
), patella 0.28/0.10 (2.80
|
||
<normalizedToken originalValue="×">x</normalizedToken>
|
||
), tibia 0.32/0.06 (5.33
|
||
<normalizedToken originalValue="×">x</normalizedToken>
|
||
), tarsus 0.32/0.05 (6.4
|
||
<normalizedToken originalValue="×">x</normalizedToken>
|
||
).
|
||
</paragraph>
|
||
<caption doi="10.3897/zse.96.56885.figure4" httpUri="https://binary.pensoft.net/fig/477315" pageId="0" pageNumber="769" start="Figure 4" startId="F4">
|
||
<paragraph pageId="0" pageNumber="769">
|
||
<emphasis bold="true" pageId="0" pageNumber="769">Figure 4.</emphasis>
|
||
<taxonomicName authorityName="Novák & Lorenz & Harms" authorityYear="2020" class="Arachnida" family="Feaellidae" genus="Feaella" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Feaella (Tetrafeaella) obscura" order="Pseudoscorpiones" pageId="0" pageNumber="769" phylum="Arthropoda" rank="species" species="obscura" subGenus="Tetrafeaella">Feaella (Tetrafeaella) obscura</taxonomicName>
|
||
sp. nov, female holotype (HNHM Pseud-2009): SEM images.
|
||
<emphasis bold="true" pageId="0" pageNumber="769">A.</emphasis>
|
||
Left chela, retrolateral view;
|
||
<emphasis bold="true" pageId="0" pageNumber="769">B.</emphasis>
|
||
Left chela, dorsomedial view;
|
||
<emphasis bold="true" pageId="0" pageNumber="769">C.</emphasis>
|
||
Left chela, ventral view. Male paratype (HNHM Pseud-2010): SEM images.
|
||
<emphasis bold="true" pageId="0" pageNumber="769">D.</emphasis>
|
||
Left chela, retrolateral view;
|
||
<emphasis bold="true" pageId="0" pageNumber="769">E.</emphasis>
|
||
Left chela, ventral view;
|
||
<emphasis bold="true" pageId="0" pageNumber="769">F.</emphasis>
|
||
Left chela, medial view.
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph pageId="0" pageNumber="769">
|
||
<emphasis bold="true" pageId="0" pageNumber="769">Females</emphasis>
|
||
(holotype, followed by paratype in parentheses and then the ratios also in parentheses). Body 2.16 (2.46). Carapace 0.61/0.48 (0.64/0.50) (1.27-1.28
|
||
<normalizedToken originalValue="×">x</normalizedToken>
|
||
). Chelicera 0.20/0.12 (0.21/0.12) (1.67-1.75
|
||
<normalizedToken originalValue="×">x</normalizedToken>
|
||
), movable finger 0.095 (0.10). Palpal femur 0.63/0.37 (0.69/0.40) (1.70-1.73
|
||
<normalizedToken originalValue="×">x</normalizedToken>
|
||
), patella 0.51/0.18 (0.54/0.19) (2.83-2.84
|
||
<normalizedToken originalValue="×">x</normalizedToken>
|
||
), chela (with pedicel) 0.60/0.16 (0.61/0.16) (3.75-3.81
|
||
<normalizedToken originalValue="×">x</normalizedToken>
|
||
), chela (without pedicel) 0.57 (0.57), hand (with pedicel) 0.16 (0.17), hand (without pedicel) 0.12 (0.10), movable finger 0.40 (0.41). Leg I. trochanter 0.15/0.11 (0.16/0.12) (1.33-1.36
|
||
<normalizedToken originalValue="×">x</normalizedToken>
|
||
), femur 0.24/0.07 (0.27/0.08) (3.38-3.43
|
||
<normalizedToken originalValue="×">x</normalizedToken>
|
||
), patella 0.22/0.08 (0.21/0.09) (2.33-2.75
|
||
<normalizedToken originalValue="×">x</normalizedToken>
|
||
), tibia 0.20/0.07 (0.22/0.07) (2.85-3.14
|
||
<normalizedToken originalValue="×">x</normalizedToken>
|
||
), tarsus 0.22/0.05 (0.22/0.05) (4.4
|
||
<normalizedToken originalValue="×">x</normalizedToken>
|
||
). Leg IV. trochanter 0.26/0.14 (0.27/14) (1.86-1.93
|
||
<normalizedToken originalValue="×">x</normalizedToken>
|
||
), femur 0.18/0.10 (0.19/10) (1.8-1.9
|
||
<normalizedToken originalValue="×">x</normalizedToken>
|
||
), patella 0.36/0.11 (0.37/0.12) (3.08-3.27
|
||
<normalizedToken originalValue="×">x</normalizedToken>
|
||
), tibia 0.42/0.06 (0.44/0.07) (6.29-7.00
|
||
<normalizedToken originalValue="×">x</normalizedToken>
|
||
), tarsus 0.36/0.05 (0.38/0.06) (6.33-7.20
|
||
<normalizedToken originalValue="×">x</normalizedToken>
|
||
).
|
||
</paragraph>
|
||
<caption doi="10.3897/zse.96.56885.figure5" httpUri="https://binary.pensoft.net/fig/477316" pageId="0" pageNumber="769" start="Figure 5" startId="F5">
|
||
<paragraph pageId="0" pageNumber="769">
|
||
<emphasis bold="true" pageId="0" pageNumber="769">Figure 5.</emphasis>
|
||
<taxonomicName authorityName="Novák & Lorenz & Harms" authorityYear="2020" class="Arachnida" family="Feaellidae" genus="Feaella" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Feaella (Tetrafeaella) obscura" order="Pseudoscorpiones" pageId="0" pageNumber="769" phylum="Arthropoda" rank="species" species="obscura" subGenus="Tetrafeaella">Feaella (Tetrafeaella) obscura</taxonomicName>
|
||
sp. nov.
|
||
<emphasis bold="true" pageId="0" pageNumber="769">A.</emphasis>
|
||
Right chelicera, dorsal view - female holotype (HNHM Pseud-2009);
|
||
<emphasis bold="true" pageId="0" pageNumber="769">B.</emphasis>
|
||
Right pedipalp, dorsal view - female holotype;
|
||
<emphasis bold="true" pageId="0" pageNumber="769">C.</emphasis>
|
||
Left chela, retrolateral view - female holotype;
|
||
<emphasis bold="true" pageId="0" pageNumber="769">D.</emphasis>
|
||
Leg I, lateral view - female paratype (ZMH-A0003101);
|
||
<emphasis bold="true" pageId="0" pageNumber="769">E.</emphasis>
|
||
Leg IV, lateral view - female paratype;
|
||
<emphasis bold="true" pageId="0" pageNumber="769">F.</emphasis>
|
||
Female genital region - female holotype;
|
||
<emphasis bold="true" pageId="0" pageNumber="769">G.</emphasis>
|
||
Male genital region - male paratype (HNHM Pseud-2010).
|
||
</paragraph>
|
||
</caption>
|
||
</subSubSection>
|
||
<subSubSection pageId="0" pageNumber="769" type="differential diagnosis">
|
||
<paragraph pageId="0" pageNumber="769">Differential diagnosis.</paragraph>
|
||
<paragraph pageId="0" pageNumber="769">
|
||
Aside from
|
||
<taxonomicName authorityName="Novák & Lorenz & Harms" authorityYear="2020" class="Arachnida" family="Feaellidae" genus="Feaella" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Feaella (T.) obscura" order="Pseudoscorpiones" pageId="0" pageNumber="769" phylum="Arthropoda" rank="species" species="obscura" subGenus="T.">Feaella (T.) obscura</taxonomicName>
|
||
there are twelve species of
|
||
<taxonomicName authority="" class="Arachnida" family="Feaellidae" genus="Feaella" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Feaella (Tetrafeaella)" order="Pseudoscorpiones" pageId="0" pageNumber="769" phylum="Arthropoda" rank="subGenus" subGenus="Tetrafeaella">Feaella (Tetrafeaella)</taxonomicName>
|
||
:
|
||
<taxonomicName genus="F." lsidName="F. (T.) affinis" pageId="0" pageNumber="769" rank="species" species="affinis" subGenus="T.">F. (T.) affinis</taxonomicName>
|
||
Hirst, 1911 (Seychelles);
|
||
<taxonomicName genus="F." lsidName="F. (T.) capensis" pageId="0" pageNumber="769" rank="species" species="capensis" subGenus="T.">F. (T.) capensis</taxonomicName>
|
||
Beier, 1955,
|
||
<taxonomicName genus="F." lsidName="F. (T.) mucronata" pageId="0" pageNumber="769" rank="species" species="mucronata" subGenus="T.">F. (T.) mucronata</taxonomicName>
|
||
Tullgren, 1907 and
|
||
<taxonomicName genus="F." lsidName="F. (T.) parva" pageId="0" pageNumber="769" rank="species" species="parva" subGenus="T.">F. (T.) parva</taxonomicName>
|
||
Beier, 1947 (all South Africa),
|
||
<taxonomicName genus="F." lsidName="F. (T.) indica" pageId="0" pageNumber="769" rank="species" species="indica" subGenus="T.">F. (T.) indica</taxonomicName>
|
||
Chamberlin, 1931 (India and Sri Lanka),
|
||
<taxonomicName genus="F." lsidName="F. (T.) leleupi" pageId="0" pageNumber="769" rank="species" species="leleupi" subGenus="T.">F. (T.) leleupi</taxonomicName>
|
||
Beier, 1959 (Congo),
|
||
<taxonomicName genus="F." lsidName="F. (T.) nana" pageId="0" pageNumber="769" rank="species" species="nana" subGenus="T.">F. (T.) nana</taxonomicName>
|
||
Beier, 1966 comb. nov. (previously treated as a subspecies of
|
||
<taxonomicName genus="F." lsidName="capensis" pageId="0" pageNumber="769" rank="species" species="capensis">
|
||
<emphasis italics="true" pageId="0" pageNumber="769">capensis</emphasis>
|
||
</taxonomicName>
|
||
but see below),
|
||
<taxonomicName genus="F." lsidName="F. (T.) perreti" pageId="0" pageNumber="769" rank="species" species="perreti" subGenus="T.">F. (T.) perreti</taxonomicName>
|
||
Mahnert, 1982 (Kenya) and the western Australian species
|
||
<taxonomicName genus="F." lsidName="F. (T.) anderseni" pageId="0" pageNumber="769" rank="species" species="anderseni" subGenus="T.">F. (T.) anderseni</taxonomicName>
|
||
Harvey, 1989,
|
||
<taxonomicName genus="F." lsidName="F. (T.) callani" pageId="0" pageNumber="769" rank="species" species="callani" subGenus="T.">F. (T.) callani</taxonomicName>
|
||
Harvey et al., 2016,
|
||
<taxonomicName genus="F." lsidName="F. (T.) linetteae" pageId="0" pageNumber="769" rank="species" species="linetteae" subGenus="T.">F. (T.) linetteae</taxonomicName>
|
||
Harvey et al., 2016 and
|
||
<taxonomicName genus="F." lsidName="F. (T.) tealei" pageId="0" pageNumber="769" rank="species" species="tealei" subGenus="T.">F. (T.) tealei</taxonomicName>
|
||
Harvey et al., 2016.
|
||
</paragraph>
|
||
<paragraph pageId="0" pageNumber="769">
|
||
<taxonomicName authorityName="Novák & Lorenz & Harms" authorityYear="2020" class="Arachnida" family="Feaellidae" genus="Feaella" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Feaella (T.) obscura" order="Pseudoscorpiones" pageId="0" pageNumber="769" phylum="Arthropoda" rank="species" species="obscura" subGenus="T.">Feaella (T.) obscura</taxonomicName>
|
||
is most similar in general habitus of carapace and pedipalps and in most characters (see in the key below) to the following species from Africa, India and the Seychelles.
|
||
</paragraph>
|
||
<paragraph pageId="0" pageNumber="769">
|
||
It differs from
|
||
<taxonomicName genus="F." lsidName="F. (T.) capensis" pageId="0" pageNumber="769" rank="species" species="capensis" subGenus="T.">F. (T.) capensis</taxonomicName>
|
||
in having a pronounced and triangular dorsomedial process on the prolateral corner of palpal femur near its base (absent in
|
||
<taxonomicName genus="F." lsidName="F. (T.) capensis" pageId="0" pageNumber="769" rank="species" species="capensis" subGenus="T.">F. (T.) capensis</taxonomicName>
|
||
), conical shape of the frontal carapace lobes (long and triangular), pleural membrane with a dorsal row of 15 and ventral row of 14 platelets (15 and 15 in
|
||
<taxonomicName genus="F." lsidName="F. (T.) capensis" pageId="0" pageNumber="769" rank="species" species="capensis" subGenus="T.">F. (T.) capensis</taxonomicName>
|
||
) (
|
||
<bibRefCitation author="Beier, M" journalOrPublisher="Eos, Madrid" pageId="0" pageNumber="769" refId="B4" refString="Beier, M, 1955. Pseudoscorpionidea. In: Hanstrom B, Brinck P, Rudebeck G (Eds) South African Animal Life. Results of the Lund Expedition in 1950-1951, vol 1: 263-328. Almquist and Wiksell: Stockholm." title="Pseudoscorpionidea. In: Hanstrom B, Brinck P, Rudebeck G (Eds) South African Animal Life. Results of the Lund Expedition in 1950 - 1951, vol 1: 263 - 328. Almquist and Wiksell: Stockholm." year="1955">Beier 1955</bibRefCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph pageId="0" pageNumber="769">
|
||
<taxonomicName authorityName="Novák & Lorenz & Harms" authorityYear="2020" class="Arachnida" family="Feaellidae" genus="Feaella" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Feaella (T.) obscura" order="Pseudoscorpiones" pageId="0" pageNumber="769" phylum="Arthropoda" rank="species" species="obscura" subGenus="T.">Feaella (T.) obscura</taxonomicName>
|
||
differs from
|
||
<taxonomicName genus="F." lsidName="F. (T.) nana" pageId="0" pageNumber="769" rank="species" species="nana" subGenus="T.">F. (T.) nana</taxonomicName>
|
||
in having a pronounced and triangular dorsomedial process on the prolateral corner of the palpal femur near its base (absent in
|
||
<taxonomicName genus="F." lsidName="F. (T.) nana" pageId="0" pageNumber="769" rank="species" species="nana" subGenus="T.">F. (T.) nana</taxonomicName>
|
||
). Furthermore, in having pleural membrane with a dorsal row of 15 and a ventral row of 14 sclerotised pleural platelets (in
|
||
<taxonomicName genus="F." lsidName="F. (T.) nana" pageId="0" pageNumber="769" rank="species" species="nana" subGenus="T.">F. (T.) nana</taxonomicName>
|
||
the number of these are 14 and 14). In
|
||
<taxonomicName authorityName="Novák & Lorenz & Harms" authorityYear="2020" class="Arachnida" family="Feaellidae" genus="Feaella" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Feaella (T.) obscura" order="Pseudoscorpiones" pageId="0" pageNumber="769" phylum="Arthropoda" rank="species" species="obscura" subGenus="T.">Feaella (T.) obscura</taxonomicName>
|
||
on fixed chelal finger
|
||
<emphasis italics="true" pageId="0" pageNumber="769">isb</emphasis>
|
||
much closer to
|
||
<emphasis italics="true" pageId="0" pageNumber="769">ib</emphasis>
|
||
than to
|
||
<emphasis italics="true" pageId="0" pageNumber="769">ist</emphasis>
|
||
and these three trichobothria do not form a straight row, and trichobothrium
|
||
<emphasis italics="true" pageId="0" pageNumber="769">eb</emphasis>
|
||
situated clearly closer to the fingertip than to
|
||
<emphasis italics="true" pageId="0" pageNumber="769">ist</emphasis>
|
||
(in
|
||
<taxonomicName genus="F." lsidName="F. (T.) nana" pageId="0" pageNumber="769" rank="species" species="nana" subGenus="T.">F. (T.) nana</taxonomicName>
|
||
<emphasis italics="true" pageId="0" pageNumber="769">isb</emphasis>
|
||
,
|
||
<emphasis italics="true" pageId="0" pageNumber="769">ib</emphasis>
|
||
and
|
||
<emphasis italics="true" pageId="0" pageNumber="769">ist</emphasis>
|
||
are equally placed and form a straight row, and
|
||
<emphasis italics="true" pageId="0" pageNumber="769">eb</emphasis>
|
||
is situated midway between
|
||
<emphasis italics="true" pageId="0" pageNumber="769">ist</emphasis>
|
||
and the finger tip) (
|
||
<bibRefCitation author="Beier, M" journalOrPublisher="Annals of the Natal Museum" pageId="0" pageNumber="769" pagination="455 - 470" refId="B7" refString="Beier, M, 1966. Ergaenzungen zur Pseudoskorpioniden-Fauna des suedlichen Afrika. Annals of the Natal Museum 18: 455 - 470" title="Ergaenzungen zur Pseudoskorpioniden-Fauna des suedlichen Afrika." volume="18" year="1966">Beier 1966</bibRefCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph pageId="0" pageNumber="769">
|
||
The new species differs from
|
||
<taxonomicName genus="F." lsidName="F. (T.) indica" pageId="0" pageNumber="769" rank="species" species="indica" subGenus="T.">F. (T.) indica</taxonomicName>
|
||
in having a pronounced and triangular dorsomedial process on the prolateral corner of palpal femur near its base (in
|
||
<taxonomicName genus="F." lsidName="F. (T.) indica" pageId="0" pageNumber="769" rank="species" species="indica" subGenus="T.">F. (T.) indica</taxonomicName>
|
||
this process is absent). The chelae of
|
||
<taxonomicName genus="F." lsidName="F. (T.) obscura" pageId="0" pageNumber="769" rank="species" species="obscura" subGenus="T.">F. (T.) obscura</taxonomicName>
|
||
are approximately as long as the palpal femora (in
|
||
<taxonomicName genus="F." lsidName="F. (T.) indica" pageId="0" pageNumber="769" rank="species" species="indica" subGenus="T.">F. (T.) indica</taxonomicName>
|
||
these are distinctly shorter than the palpal femora). Furthermore, the palpal femur length/width ratio of
|
||
<taxonomicName genus="F." lsidName="F. (T.) obscura" pageId="0" pageNumber="769" rank="species" species="obscura" subGenus="T.">F. (T.) obscura</taxonomicName>
|
||
is 1.70-1.76
|
||
<normalizedToken originalValue="×">x</normalizedToken>
|
||
(this value is 1.90
|
||
<normalizedToken originalValue="×">x</normalizedToken>
|
||
in
|
||
<taxonomicName genus="F." lsidName="F. (T.) indica" pageId="0" pageNumber="769" rank="species" species="indica" subGenus="T.">F. (T.) indica</taxonomicName>
|
||
) (
|
||
<bibRefCitation author="Chamberlin, JC" journalOrPublisher="Stanford University Publications, Biological Sciences" pageId="0" pageNumber="769" pagination="1 - 284" refId="B9" refString="Chamberlin, JC, 1931. The arachnid order Chelonethida. Stanford University Publications, Biological Sciences 7: 1 - 284" title="The arachnid order Chelonethida." volume="7" year="1931">Chamberlin 1931</bibRefCitation>
|
||
;
|
||
<bibRefCitation DOI="https://doi.org/10.1515/9783111435107" author="Beier, M" journalOrPublisher="Das Tierreich" pageId="0" pageNumber="769" pagination="1 - 258" refId="B2" refString="Beier, M, 1932. Pseudoscorpionidea I. Subord. Chthoniinea et Neobisiinea. Das Tierreich 57: 1 - 258, DOI: https://doi.org/10.1515/9783111435107" title="Pseudoscorpionidea I. Subord. Chthoniinea et Neobisiinea." url="https://doi.org/10.1515/9783111435107" volume="57" year="1932">Beier 1932</bibRefCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph pageId="0" pageNumber="769">
|
||
In the case of
|
||
<taxonomicName genus="F." lsidName="F. (T.) affinis" pageId="0" pageNumber="769" rank="species" species="affinis" subGenus="T.">F. (T.) affinis</taxonomicName>
|
||
Hirst, 1911 trichobotria
|
||
<emphasis italics="true" pageId="0" pageNumber="769">sb</emphasis>
|
||
and
|
||
<emphasis italics="true" pageId="0" pageNumber="769">st</emphasis>
|
||
of movable chelal finger are closer to
|
||
<emphasis italics="true" pageId="0" pageNumber="769">b</emphasis>
|
||
than
|
||
<emphasis italics="true" pageId="0" pageNumber="769">t</emphasis>
|
||
(in the new species
|
||
<emphasis italics="true" pageId="0" pageNumber="769">sb</emphasis>
|
||
and
|
||
<emphasis italics="true" pageId="0" pageNumber="769">st</emphasis>
|
||
are situated midway between
|
||
<emphasis italics="true" pageId="0" pageNumber="769">b</emphasis>
|
||
and
|
||
<emphasis italics="true" pageId="0" pageNumber="769">t</emphasis>
|
||
). Furthermore, in
|
||
<taxonomicName genus="F." lsidName="F. (T.) affinis" pageId="0" pageNumber="769" rank="species" species="affinis" subGenus="T.">F. (T.) affinis</taxonomicName>
|
||
trichobothrium
|
||
<emphasis italics="true" pageId="0" pageNumber="769">ist</emphasis>
|
||
of fixed chelal finger is at the level of
|
||
<emphasis italics="true" pageId="0" pageNumber="769">esb</emphasis>
|
||
and
|
||
<emphasis italics="true" pageId="0" pageNumber="769">est</emphasis>
|
||
is placed distal to them (in
|
||
<taxonomicName genus="F." lsidName="F. (T.) obscura" pageId="0" pageNumber="769" rank="species" species="obscura" subGenus="T.">F. (T.) obscura</taxonomicName>
|
||
,
|
||
<emphasis italics="true" pageId="0" pageNumber="769">ist</emphasis>
|
||
is equidistant from
|
||
<emphasis italics="true" pageId="0" pageNumber="769">est</emphasis>
|
||
and
|
||
<emphasis italics="true" pageId="0" pageNumber="769">esb</emphasis>
|
||
) (
|
||
<bibRefCitation author="Mahnert, V" journalOrPublisher="Revue de Zoologie Africaine" pageId="0" pageNumber="769" pagination="867 - 888" refId="B27" refString="Mahnert, V, 1978. Contributions a l'etude de la faune terrestre des ȋles granitiques de l'archipel des Sechelles. Pseudoscorpiones. Revue de Zoologie Africaine 92: 867 - 888" title="Contributions a l'etude de la faune terrestre des ȋles granitiques de l'archipel des Sechelles. Pseudoscorpiones." volume="92" year="1978">Mahnert 1978</bibRefCitation>
|
||
).
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |