166 lines
18 KiB
XML
166 lines
18 KiB
XML
<document id="9872E0766859272B71EEA0E4C4A13265" ID-CLB-Dataset="28091" ID-DOI="10.11646/zootaxa.4504.1.3" ID-GBIF-Dataset="0b803003-244e-4511-aaf4-5e9c3f863861" ID-ISSN="1175-5326" ID-Zenodo-Dep="2606164" ID-ZooBank="52BA28B6-4280-4BD0-8B38-C794F04E4A66" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1553577945224" checkinUser="plazi" docAuthor="Abdalla, Ishtiag H., Deschodt, Christian M., Scholtz, Clarke H. & Sole, Catherine L." docDate="2018" docId="B276DC1EFFBDFFF9FF0DFC2CFE0B034F" docLanguage="en" docName="zootaxa.4504.1.3.pdf" docOrigin="Zootaxa 4504 (1)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Macroderes spectabilis Peringuey 1901" docType="treatment" docVersion="8" lastPageNumber="72" masterDocId="4E4FA466FFA2FFE6FF9AFFF0FFD20545" masterDocTitle="An update to the taxonomy of the genus Macroderes Westwood 1842 (Coleoptera: Scarabaeidae: Scarabaeinae) with descriptions of new species from South Africa" masterLastPageNumber="75" masterPageNumber="41" pageNumber="72" updateTime="1698712718888" updateUser="ExternalLinkService">
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<mods:titleInfo id="B8A813CCA61D3446B42D525FE06A310A">
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<mods:title id="6483632333EFD8B813A805E1A66FA55B">An update to the taxonomy of the genus Macroderes Westwood 1842 (Coleoptera: Scarabaeidae: Scarabaeinae) with descriptions of new species from South Africa</mods:title>
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<mods:namePart id="DE65F997DD00B9961FF8E4B152CBDF3D">Abdalla, Ishtiag H.</mods:namePart>
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<mods:namePart id="3F5DB8A9E503BE70A2C121631637EB0E">Deschodt, Christian M.</mods:namePart>
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<mods:namePart id="0DF92A8E9C0D3211144F2134E405626B">Scholtz, Clarke H.</mods:namePart>
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<mods:namePart id="434F6C52522880EDDE9C10070BCD4EB2">Sole, Catherine L.</mods:namePart>
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<mods:typeOfResource id="DCF23F2D010CD85CBECC97FFD721F8C0">text</mods:typeOfResource>
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<mods:title id="BDB0BC3C0307734C1097B2E5A6C2944F">Zootaxa</mods:title>
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<mods:date id="E2BD4D785655223EE0E2B2F96C1CE194">2018</mods:date>
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<mods:number id="504DE12B66C568847E6FA380BA0F6F0F">2018-10-23</mods:number>
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<mods:number id="F07C8ABBF03A832374E65888259286AF">4504</mods:number>
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<treatment id="B276DC1EFFBDFFF9FF0DFC2CFE0B034F" ID-DOI="http://doi.org/10.5281/zenodo.5996780" ID-GBIF-Taxon="154494808" ID-Zenodo-Dep="5996780" LSID="urn:lsid:plazi:treatment:B276DC1EFFBDFFF9FF0DFC2CFE0B034F" httpUri="http://treatment.plazi.org/id/B276DC1EFFBDFFF9FF0DFC2CFE0B034F" lastPageNumber="72" pageId="31" pageNumber="72">
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<subSubSection id="72C53E83FFBDFFF9FF0DFC2CFDAA06B3" box="[151,632,988,1014]" pageId="31" pageNumber="72" type="nomenclature">
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<paragraph id="3A606D08FFBDFFF9FF0DFC2CFDAA06B3" blockId="31.[151,632,988,1014]" box="[151,632,988,1014]" pageId="31" pageNumber="72">
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<heading id="6128DA64FFBDFFF9FF0DFC2CFDAA06B3" bold="true" box="[151,632,988,1014]" fontSize="11" level="1" pageId="31" pageNumber="72" reason="1">
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<taxonomicName id="FDDF168BFFBDFFF9FF0DFC2CFDAA06B3" ID-CoL="734DB" authority="Peringuey, 1901" authorityName="Peringuey" authorityYear="1901" box="[151,632,988,1014]" class="Insecta" family="Scarabaeidae" genus="Macroderes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="31" pageNumber="72" phylum="Arthropoda" rank="species" species="spectabilis">
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<emphasis id="08ABB11AFFBDFFF9FF0DFC2CFDAA06B3" bold="true" box="[151,632,988,1014]" pageId="31" pageNumber="72">
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<emphasis id="08ABB11AFFBDFFF9FF0DFC2CFE7806B3" bold="true" box="[151,426,988,1014]" italics="true" pageId="31" pageNumber="72">Macroderes spectabilis</emphasis>
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<bibRefCitation id="5E4E10F9FFBDFFF9FE28FC2CFDAA06B3" author="Peringuey, L." box="[434,632,988,1014]" pageId="31" pageNumber="72" pagination="1 - 563" refId="ref22154" refString="Peringuey, L. (1901) Descriptive catalogue of the Coleoptera of South Africa (Lucanidae and Scarabaeidae). Transactions of the South African Philosophical Society, 12 (1), 1 - 563. https: // doi. org / 10.5962 / bhl. title. 8963" type="journal article" year="1901">Péringuey, 1901</bibRefCitation>
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</emphasis>
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</taxonomicName>
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</heading>
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</paragraph>
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</subSubSection>
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<subSubSection id="72C53E83FFBDFFF9FF0DFBD2FACC017D" box="[151,1310,1058,1080]" pageId="31" pageNumber="72" type="reference_group">
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<paragraph id="3A606D08FFBDFFF9FF0DFBD2FACC017D" blockId="31.[151,1310,1058,1080]" box="[151,1310,1058,1080]" pageId="31" pageNumber="72">
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<treatmentCitationGroup id="1ACF4A26FFBDFFF9FF0DFBD2FACC017D" box="[151,1310,1058,1080]" pageId="31" pageNumber="72">
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<taxonomicName id="FDDF168BFFBDFFF9FF0DFBD2FDB1017D" ID-CoL="734DB" authority="Peringuey, 1901: 300" authorityName="Peringuey" authorityPageNumber="300" authorityYear="1901" box="[151,611,1058,1080]" class="Insecta" family="Scarabaeidae" genus="Macroderes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="31" pageNumber="72" phylum="Arthropoda" rank="species" species="spectabilis">
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<emphasis id="08ABB11AFFBDFFF9FF0DFBD2FE50017D" box="[151,386,1058,1080]" italics="true" pageId="31" pageNumber="72">Macroderes spectabilis</emphasis>
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<treatmentCitation id="BB7E4B19FFBDFFF9FE12FBD2FDB1017D" author="Peringuey, L." box="[392,611,1058,1080]" page="300" pageId="31" pageNumber="72" year="1901">
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<bibRefCitation id="5E4E10F9FFBDFFF9FE12FBD2FDE2017D" author="Peringuey, L." box="[392,560,1058,1080]" pageId="31" pageNumber="72" pagination="1 - 563" refId="ref22154" refString="Peringuey, L. (1901) Descriptive catalogue of the Coleoptera of South Africa (Lucanidae and Scarabaeidae). Transactions of the South African Philosophical Society, 12 (1), 1 - 563. https: // doi. org / 10.5962 / bhl. title. 8963" type="journal article" year="1901">Péringuey, 1901</bibRefCitation>
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: 300
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</treatmentCitation>
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</taxonomicName>
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;
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<treatmentCitation id="BB7E4B19FFBDFFF9FDF7FBD3FCF5017D" author="Janssens, A." box="[621,807,1058,1080]" page="27" pageId="31" pageNumber="72" year="1939">
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<bibRefCitation id="5E4E10F9FFBDFFF9FDF7FBD3FCD3017D" author="Janssens, A." box="[621,769,1058,1080]" pageId="31" pageNumber="72" pagination="1 - 104" refId="ref21780" refString="Janssens, A. (1939) Coprini. Exploration du Parc National Albert. Mission G. F. de Witte, 29, 1 - 104." type="journal article" year="1939">Janssens 1939</bibRefCitation>
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: 27
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</treatmentCitation>
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;
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<treatmentCitation id="BB7E4B19FFBDFFF9FCABFBD2FC23017D" author="Ferreira, M." box="[817,1009,1058,1080]" page="322" pageId="31" pageNumber="72" year="1969">
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<bibRefCitation id="5E4E10F9FFBDFFF9FCABFBD2FC6C017D" author="Ferreira, M." box="[817,958,1058,1080]" pageId="31" pageNumber="72" pagination="5 - 1088" refId="ref21605" refString="Ferreira, M. (1969) Os escarabideos de Africa (Sul do Saara), I. Revista de Entomologia de Mocambique, 11, 5 - 1088." type="journal article" year="1969">Ferreira 1969</bibRefCitation>
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: 322
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</treatmentCitation>
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;
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<treatmentCitation id="BB7E4B19FFBDFFF9FC61FBD2FAC8017D" author="Frolov, A. V. & Scholtz, C. H." box="[1019,1306,1058,1080]" page="392" pageId="31" pageNumber="72" year="2005">
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<bibRefCitation id="5E4E10F9FFBDFFF9FC61FBD2FB3A017D" author="Frolov, A. V. & Scholtz, C. H." box="[1019,1256,1058,1080]" pageId="31" pageNumber="72" pagination="373 - 393" refId="ref21694" refString="Frolov, A. V. & Scholtz, C. H. (2005) Revision of the southern African genus Macroderes Westwood (Coleoptera: Scarabaeidae: Scarabaeinae). Annales de la Societe Entomologique de France, 40 (3 - 4), 373 - 393. https: // doi. org / 10.1080 / 00379271.2004.10697429" type="journal article" year="2005">Frolov & Scholtz 2005</bibRefCitation>
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: 392
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</treatmentCitation>
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.
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</treatmentCitationGroup>
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</paragraph>
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</subSubSection>
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<subSubSection id="72C53E83FFBDFFF9FF0DFB95FA4301E4" pageId="31" pageNumber="72" type="materials_examined">
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<paragraph id="3A606D08FFBDFFF9FF0DFB95FD7B013B" blockId="31.[151,1437,1125,1546]" box="[151,681,1125,1150]" pageId="31" pageNumber="72">
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<materialsCitation id="8AB76755FFBDFFF9FF0DFB95FD77013B" ID-GBIF-Occurrence="2028137847" box="[151,677,1125,1150]" location="Colony" pageId="31" pageNumber="72" specimenCount="1" stateProvince="Western Cape" typeStatus="holotype">
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<emphasis id="08ABB11AFFBDFFF9FF0DFB95FEE7013B" bold="true" box="[151,309,1125,1150]" pageId="31" pageNumber="72">
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<typeStatus id="E564D3AAFFBDFFF9FF0DFB95FF00013B" box="[151,210,1125,1150]" pageId="31" pageNumber="72">Type</typeStatus>
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locality.
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</emphasis>
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<collectingRegion id="F81BA3EAFFBDFFF9FEA6FB96FEA4013B" box="[316,374,1126,1150]" country="South Africa" name="Western Cape" pageId="31" pageNumber="72">Cape</collectingRegion>
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<location id="3F003BD3FFBDFFF9FEE7FB95FE02013B" LSID="urn:lsid:plazi:treatment:B276DC1EFFBDFFF9FF0DFC2CFE0B034F:3F003BD3FFBDFFF9FEE7FB95FE02013B" box="[381,464,1125,1150]" name="Colony" pageId="31" pageNumber="72" stateProvince="Western Cape">Colony</location>
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(no exact locality)
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</materialsCitation>
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.
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</paragraph>
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<paragraph id="3A606D08FFBDFFF9FF5DFB79FA4301E4" blockId="31.[151,1437,1125,1546]" box="[199,1425,1161,1186]" pageId="31" pageNumber="72">
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<emphasis id="08ABB11AFFBDFFF9FF5DFB79FE3D01E7" bold="true" box="[199,495,1161,1186]" pageId="31" pageNumber="72">
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<typeStatus id="E564D3AAFFBDFFF9FF5DFB79FED001E7" box="[199,258,1161,1186]" pageId="31" pageNumber="72">Type</typeStatus>
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material examined.
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</emphasis>
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None.
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<materialsCitation id="8AB76755FFBDFFF9FDDAFB7AFA5F01E4" ID-GBIF-Occurrence="2028137821" box="[576,1421,1161,1186]" location="South African" pageId="31" pageNumber="72" specimenCount="1" typeStatus="holotype">
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<typeStatus id="E564D3AAFFBDFFF9FDDAFB7AFDA501E7" box="[576,631,1162,1186]" pageId="31" pageNumber="72">Type</typeStatus>
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specimen may be lost, it was not found at any
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<location id="3F003BD3FFBDFFF9FB1CFB79FAF101E7" LSID="urn:lsid:plazi:treatment:B276DC1EFFBDFFF9FF0DFC2CFE0B034F:3F003BD3FFBDFFF9FB1CFB79FAF101E7" box="[1158,1315,1161,1186]" name="South African" pageId="31" pageNumber="72">South African</location>
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museum
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</materialsCitation>
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.
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</paragraph>
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</subSubSection>
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<subSubSection id="72C53E83FFBDFFF9FF5DFB5DFE0B034F" pageId="31" pageNumber="72" type="discussion">
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<paragraph id="3A606D08FFBDFFF9FF5DFB5DFAE2003F" blockId="31.[151,1437,1125,1546]" pageId="31" pageNumber="72">
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<emphasis id="08ABB11AFFBDFFF9FF5DFB5DFEE90183" bold="true" box="[199,315,1197,1222]" pageId="31" pageNumber="72">Remarks.</emphasis>
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We have not seen the
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<typeStatus id="E564D3AAFFBDFFF9FDACFB5EFDB40183" box="[566,614,1198,1222]" pageId="31" pageNumber="72">type</typeStatus>
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specimen of this species at any of the South African museum throughout our study, thus we support
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<bibRefCitation id="5E4E10F9FFBDFFF9FE5CFB21FD0901AF" author="Frolov, A. V. & Scholtz, C. H." box="[454,731,1233,1258]" pageId="31" pageNumber="72" pagination="373 - 393" refId="ref21694" refString="Frolov, A. V. & Scholtz, C. H. (2005) Revision of the southern African genus Macroderes Westwood (Coleoptera: Scarabaeidae: Scarabaeinae). Annales de la Societe Entomologique de France, 40 (3 - 4), 373 - 393. https: // doi. org / 10.1080 / 00379271.2004.10697429" type="journal article" year="2005">Frolov & Scholtz (2005)</bibRefCitation>
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assumption that the
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<typeStatus id="E564D3AAFFBDFFF9FC5EFB22FC2601AF" box="[964,1012,1234,1258]" pageId="31" pageNumber="72">type</typeStatus>
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specimen of this species may be lost. However, Péringuey in his original description (1901: 300) mentioned that the length of the specimen is
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<quantity id="FD27C0EDFFBDFFF9FAB1FB05FA4F0048" box="[1323,1437,1269,1294]" metricMagnitude="-2" metricUnit="m" metricValue="1.275" pageId="31" pageNumber="72" unit="mm" value="12.75">12.75 mm</quantity>
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and the width is
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<quantity id="FD27C0EDFFBDFFF9FECCFAE9FE620074" box="[342,432,1305,1330]" metricMagnitude="-3" metricUnit="m" metricValue="8.5" pageId="31" pageNumber="72" unit="mm" value="8.5">8.5 mm</quantity>
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. The species is different from its congeners in that it has a remarkable retuse and lobate pronotum in front and the absence of elytral stria 9. In his description,
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<bibRefCitation id="5E4E10F9FFBDFFF9FC2AFACDFBA10013" author="Peringuey, L." box="[944,1139,1341,1366]" pageId="31" pageNumber="72" pagination="1 - 563" refId="ref22154" refString="Peringuey, L. (1901) Descriptive catalogue of the Coleoptera of South Africa (Lucanidae and Scarabaeidae). Transactions of the South African Philosophical Society, 12 (1), 1 - 563. https: // doi. org / 10.5962 / bhl. title. 8963" type="journal article" year="1901">Péringuey (1901)</bibRefCitation>
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added that the species has dense, equally distant and close pronotal punctures with the presence of deep impression antero-laterally.
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</paragraph>
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<paragraph id="3A606D08FFBDFFF9FF5DFA75FE0B034F" blockId="31.[151,1437,1125,1546]" pageId="31" pageNumber="72">
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The absence of elytral stria 9 is not known among
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<taxonomicName id="FDDF168BFFBDFFF9FCB6FA75FC6300D8" authorityName="Westwood" authorityYear="1842" box="[812,945,1413,1437]" class="Insecta" family="Scarabaeidae" genus="Macroderes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="31" pageNumber="72" phylum="Arthropoda" rank="genus">
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<emphasis id="08ABB11AFFBDFFF9FCB6FA75FC6300D8" box="[812,945,1413,1437]" italics="true" pageId="31" pageNumber="72">Macroderes</emphasis>
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</taxonomicName>
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species, however, other above mentioned characteristics put the species in close similarity to
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<taxonomicName id="FDDF168BFFBDFFF9FD73FA5AFC920084" baseAuthorityName="Olivier" baseAuthorityYear="1789" box="[745,832,1449,1473]" class="Insecta" family="Scarabaeidae" genus="Macroderes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="31" pageNumber="72" phylum="Arthropoda" rank="species" species="bias">
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<emphasis id="08ABB11AFFBDFFF9FD73FA5AFC920084" box="[745,832,1449,1473]" italics="true" pageId="31" pageNumber="72">M. bias</emphasis>
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</taxonomicName>
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.
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<bibRefCitation id="5E4E10F9FFBDFFF9FCCAFA59FBB90087" author="Frolov, A. V. & Scholtz, C. H." box="[848,1131,1449,1474]" pageId="31" pageNumber="72" pagination="373 - 393" refId="ref21694" refString="Frolov, A. V. & Scholtz, C. H. (2005) Revision of the southern African genus Macroderes Westwood (Coleoptera: Scarabaeidae: Scarabaeinae). Annales de la Societe Entomologique de France, 40 (3 - 4), 373 - 393. https: // doi. org / 10.1080 / 00379271.2004.10697429" type="journal article" year="2005">Frolov & Scholtz (2005)</bibRefCitation>
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suggested that the species might be “an aberrant specimen” of
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<taxonomicName id="FDDF168BFFBDFFF9FDAAFA3EFD5600A0" baseAuthorityName="Olivier" baseAuthorityYear="1789" box="[560,644,1485,1509]" class="Insecta" family="Scarabaeidae" genus="Macroderes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="31" pageNumber="72" phylum="Arthropoda" rank="species" species="bias">
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<emphasis id="08ABB11AFFBDFFF9FDAAFA3EFD5600A0" box="[560,644,1485,1509]" italics="true" pageId="31" pageNumber="72">M. bias</emphasis>
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</taxonomicName>
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and attributed the absence of elytral stria 9 to uncompleted separation of the striae from each other.
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</paragraph>
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</subSubSection>
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</treatment>
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</document> |