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<document id="8DC180E7B6729CBC9B2CDE56AED93871" ID-CLB-Dataset="28091" ID-DOI="10.11646/zootaxa.4504.1.3" ID-GBIF-Dataset="0b803003-244e-4511-aaf4-5e9c3f863861" ID-ISSN="1175-5326" ID-Zenodo-Dep="2606164" ID-ZooBank="52BA28B6-4280-4BD0-8B38-C794F04E4A66" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1553577945224" checkinUser="plazi" docAuthor="Abdalla, Ishtiag H., Deschodt, Christian M., Scholtz, Clarke H. &amp; Sole, Catherine L." docDate="2018" docId="B276DC1EFFACFFE9FF0DF894FF23003D" docLanguage="en" docName="zootaxa.4504.1.3.pdf" docOrigin="Zootaxa 4504 (1)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Macroderes gifboomi Abdalla &amp; Deschodt &amp; Scholtz &amp; Sole 2018, new species" docType="treatment" docUuid="41E8DF7E-27F6-4C30-8914-19B648371BB4" docUuidSource="ZooBank" docVersion="9" lastPageNumber="56" masterDocId="4E4FA466FFA2FFE6FF9AFFF0FFD20545" masterDocTitle="An update to the taxonomy of the genus Macroderes Westwood 1842 (Coleoptera: Scarabaeidae: Scarabaeinae) with descriptions of new species from South Africa" masterLastPageNumber="75" masterPageNumber="41" pageNumber="55" updateTime="1698712718888" updateUser="ExternalLinkService">
<mods:mods id="E254DE14F7657A523EC61CCD9F5270B1" xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo id="9B2E711A700C35A38A8DD1069BEF734A">
<mods:title id="3FEFB2940DE23B15E8D502668A0A23A5">An update to the taxonomy of the genus Macroderes Westwood 1842 (Coleoptera: Scarabaeidae: Scarabaeinae) with descriptions of new species from South Africa</mods:title>
</mods:titleInfo>
<mods:name id="73FA416345B231232385495699DF4D96" type="personal">
<mods:role id="035E53465E29A7028082B6E3C9E247C9">
<mods:roleTerm id="28DB333A3D038F37BB16840A6AC1BCA5">Author</mods:roleTerm>
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<mods:namePart id="86A556E9416BB34404B5061BE08C2803">Abdalla, Ishtiag H.</mods:namePart>
</mods:name>
<mods:name id="16E59ED9939D01E57698D4074E80503E" type="personal">
<mods:role id="4C0967359704958C4500FE93346A8177">
<mods:roleTerm id="B45B295DD93F9512942CBBB345A89A1B">Author</mods:roleTerm>
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<mods:namePart id="B785908EEDD59CC6064AEB93EB2A0801">Deschodt, Christian M.</mods:namePart>
</mods:name>
<mods:name id="6FEF0E54238C325AF8E66C05F311ADAB" type="personal">
<mods:role id="351F2DA0E5A2E251EFB44566E9BB477E">
<mods:roleTerm id="277D1DA5E85609EDD9A65A2F79605112">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="F13C2FE0F58FFCC1B4AF68B7F6959ACC">Scholtz, Clarke H.</mods:namePart>
</mods:name>
<mods:name id="FB334EE74B83296B93ACDCF2F151B878" type="personal">
<mods:role id="87E8C5ED04AE1D19314626183B4F3198">
<mods:roleTerm id="82310B10ED85C9CD09D9029455E1E490">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="500BA0D981A13F996D9D7AA5A2945DED">Sole, Catherine L.</mods:namePart>
</mods:name>
<mods:typeOfResource id="A59F8D9287661D451BAC107A0343FD6C">text</mods:typeOfResource>
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<mods:title id="E5AFB5B0D4ADA96571F1F86BAAA03ADE">Zootaxa</mods:title>
</mods:titleInfo>
<mods:part id="18252223F13A6008E4F87758B508694A">
<mods:date id="D7B3D1EE399F9D73CEF7CC83C40C161C">2018</mods:date>
<mods:detail id="C5AECCD62CBA43494505C616DA428252" type="pubDate">
<mods:number id="EE4F6A7C22E4E7593F37AED1037A7D9D">2018-10-23</mods:number>
</mods:detail>
<mods:detail id="DFDD65185E59ED7B03BB1B54DBBCBFA5" type="volume">
<mods:number id="7F79CAF83524B810F5D092F96760E065">4504</mods:number>
</mods:detail>
<mods:detail id="DFF52BE688057621A9C6E183ED4FB3B0" type="issue">
<mods:number id="C26121D5EF03424276348513CDC87C3C">1</mods:number>
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<mods:start id="6A5439D62650EFA9B6115E4B047FDDA1">41</mods:start>
<mods:end id="EF2097464064A3D828679D7DF72FBF6C">75</mods:end>
</mods:extent>
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<mods:classification id="0599DA786C49F1F2CE6F2AAB9DAE6F72">journal article</mods:classification>
<mods:identifier id="141F0DE372F440FD928F4A14C54E6C14" type="CLB-Dataset">28091</mods:identifier>
<mods:identifier id="053C504145F2182C7106D10645B59362" type="DOI">10.11646/zootaxa.4504.1.3</mods:identifier>
<mods:identifier id="23BA97F96139A1D10F16194B7A33D885" type="GBIF-Dataset">0b803003-244e-4511-aaf4-5e9c3f863861</mods:identifier>
<mods:identifier id="92D0DDE7657811B196BECE0A9768FC30" type="ISSN">1175-5326</mods:identifier>
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<treatment id="B276DC1EFFACFFE9FF0DF894FF23003D" ID-DOI="http://doi.org/10.5281/zenodo.5996736" ID-GBIF-Taxon="154494803" ID-Zenodo-Dep="5996736" LSID="urn:lsid:zoobank.org:act:41E8DF7E-27F6-4C30-8914-19B648371BB4" httpUri="http://treatment.plazi.org/id/B276DC1EFFACFFE9FF0DF894FF23003D" lastPageId="15" lastPageNumber="56" pageId="14" pageNumber="55">
<subSubSection id="72C53E83FFACFFE8FF0DF894FCF6023B" box="[151,804,1892,1918]" pageId="14" pageNumber="55" type="nomenclature">
<paragraph id="3A606D08FFACFFE8FF0DF894FCF6023B" blockId="14.[151,804,1892,1951]" box="[151,804,1892,1918]" pageId="14" pageNumber="55">
<heading id="6128DA64FFACFFE8FF0DF894FCF6023B" bold="true" box="[151,804,1892,1918]" fontSize="11" level="1" pageId="14" pageNumber="55" reason="1">
<emphasis id="08ABB11AFFACFFE8FF0DF894FCF6023B" bold="true" box="[151,804,1892,1918]" pageId="14" pageNumber="55">
<taxonomicName id="FDDF168BFFACFFE8FF0DF894FD55023B" ID-CoL="3X4TM" authority="Abdalla &amp; Scholtz" authorityName="Abdalla &amp; Deschodt &amp; Scholtz &amp; Sole" authorityYear="2018" box="[151,647,1892,1918]" class="Insecta" family="Scarabaeidae" genus="Macroderes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="14" pageNumber="55" phylum="Arthropoda" rank="species" species="gifboomi" status="sp. nov.">
<emphasis id="08ABB11AFFACFFE8FF0DF894FE4A023B" bold="true" box="[151,408,1892,1918]" italics="true" pageId="14" pageNumber="55">Macroderes gifboomi</emphasis>
Abdalla &amp; Scholtz
</taxonomicName>
,
<taxonomicNameLabel id="13980C61FFACFFE8FD0EF894FCF6023B" box="[660,804,1892,1918]" pageId="14" pageNumber="55" rank="species">new species</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="72C53E83FFACFFE8FF0DF876FCB802A1" pageId="14" pageNumber="55" type="description">
<paragraph id="3A606D08FFACFFE8FF0DF876FE7902DA" blockId="14.[151,804,1892,1951]" box="[151,427,1926,1951]" pageId="14" pageNumber="55">
(
<figureCitation id="A2E4718DFFACFFE8FF05F876FF2E02DA" box="[159,252,1926,1951]" captionStart="FIGURES 15–18" captionStartId="7.[151,264,1688,1711]" captionTargetBox="[314,1215,431,1621]" captionTargetId="figure@7.[309,1269,403,1653]" captionTargetPageId="7" captionText="FIGURES 15–18. Habitus of Macroderes species. 15, M. politulus Preudhomme de Borre, 1880; 16, M. undulatus Preudhomme de Borre, 1880; 17, M. porselinus Abdalla, new species; 18, M. gifboomi Abdalla &amp; Scholtz, new species." httpUri="https://zenodo.org/record/2606174/files/figure.png" pageId="14" pageNumber="55">Figs. 18</figureCitation>
,
<figureCitation id="A2E4718DFFACFFE8FE92F877FEF402DA" box="[264,294,1927,1951]" captionStart="FIGURES 24–39" captionStartId="10.[151,264,1343,1366]" captionTargetBox="[323,1250,205,1286]" captionTargetId="figure@10.[323,1283,195,1303]" captionTargetPageId="10" captionText="FIGURES 24–39. Internal sac of aedeagus (24), aedeagus (25) and internal sac sclerite (26-39) of Macroderes species. 26, M. endroedyi Frolov &amp; Scholtz, 2005; 27, M. amplior Frolov &amp; Scholtz, 2005; 28, M. bias (Olivier, 1789); 29, M. cederbergensis Abdalla &amp; Deschodt, new species; 30, M. cornutus Frolov &amp; Scholtz, 2005; 31, M. foveatus Frolov &amp; Scholtz, 2005; 32, M. gifboomi Abdalla &amp; Scholtz, new species; 33, M. leipoldti Abdalla &amp; Deschodt, new species; 34, M. mutilans Kolbe, 1908; 35, M. namakwanus Frolov &amp; Scholtz, 2005; 36, M. oreatus Abdalla &amp; Deschodt, new species; 37, M. porselinus Abdalla, new species; 38, M. soleiana Abdalla &amp; Deschodt, new species; 39, M. tourtosus Abdalla &amp; Scholtz, new species." httpUri="https://zenodo.org/record/2606178/files/figure.png" pageId="14" pageNumber="55">32</figureCitation>
,
<figureCitation id="A2E4718DFFACFFE8FEA8F877FE8302DB" box="[306,337,1927,1950]" captionStart="FIGURES 76–81" captionStartId="20.[151,264,1637,1660]" captionTargetBox="[171,1423,205,1613]" captionTargetId="figure@20.[171,1423,205,1613]" captionTargetPageId="20" captionText="FIGURES 76–81. Punctuation of the base of pronotum of Macroderes species. 76, M. cederbergensis Abdalla &amp; Deschodt, new species; 77, M. gifboomi Abdalla &amp; Scholtz, new species; 78, M. oreatus Abdalla &amp; Deschodt, new species; 79, M. porselinus Abdalla, new species; 80, M. soleiana Abdalla &amp; Deschodt, new species; 81, M. leipoldti Abdalla &amp; Deschodt, new species." httpUri="https://zenodo.org/record/2606198/files/figure.png" pageId="14" pageNumber="55">77</figureCitation>
,
<figureCitation id="A2E4718DFFACFFE8FEC7F877FEA902DA" box="[349,379,1927,1951]" captionStart="FIGURES 82–87" captionStartId="21.[151,264,1505,1528]" captionTargetBox="[316,1276,406,1475]" captionTargetId="figure@21.[316,1276,406,1475]" captionTargetPageId="21" captionText="FIGURES 82–87. Pronotum punctures shape of Macroderes species. 82, M. tourtosus Abdalla &amp; Scholtz, new species; 83, M. cederbergensis Abdalla &amp; Deschodt, new species; 84, M. gifboomi Abdalla &amp; Scholtz, new species; 85, M. leipoldti Abdalla &amp; Deschodt, new species; 86, M. oreatus Abdalla &amp; Deschodt, new species; 87, M. porselinus Abdalla, new species." httpUri="https://zenodo.org/record/2606200/files/figure.png" pageId="14" pageNumber="55">84</figureCitation>
,
<figureCitation id="A2E4718DFFACFFE8FE1CF877FE7102DA" box="[390,419,1927,1951]" captionStart="FIGURES 88–93" captionStartId="23.[151,264,1292,1315]" captionTargetBox="[316,1276,183,1264]" captionTargetId="figure@23.[316,1276,183,1264]" captionTargetPageId="23" captionText="FIGURES 88–93. Pronotum punctures shape (88, 89) and sculpture of second elytra intervals (90–93) of Macroderes species. 88, M. soleiana Abdalla &amp; Deschodt, new species; 89, M. tourtosus Abdalla and Scholtz, new species; 90, M. minutus Frolov and Scholtz, 2005; 91, M. greeni (Kirby, 1818); 92, M. politulus Preudhomme de Borre, 1880; 93, M. gifboomi Abdalla &amp; Scholtz, new species." httpUri="https://zenodo.org/record/2606202/files/figure.png" pageId="14" pageNumber="55">93</figureCitation>
)
</paragraph>
<paragraph id="3A606D08FFACFFE8FF0DF83EFCB802A1" blockId="14.[151,874,1998,2020]" box="[151,874,1998,2020]" pageId="14" pageNumber="55">
<uri id="4E4E610AFFACFFE8FF0DF83EFCB802A1" box="[151,874,1998,2020]" pageId="14" pageNumber="55">
urn:lsid:zoobank.org:act:
<uuid id="4E7957DDFFACFFE8FE0BF83EFCB802A1" box="[401,874,1998,2020]" pageId="14" pageNumber="55">41E8DF7E-27F6-4C30-8914-19B648371BB4</uuid>
</uri>
</paragraph>
</subSubSection>
<subSubSection id="72C53E83FFADFFE9FF0DFF67FAFC0778" pageId="15" pageNumber="56" type="materials_examined">
<paragraph id="3A606D08FFADFFE9FF0DFF67FA4205F5" blockId="15.[151,1437,151,1400]" box="[151,1424,151,176]" pageId="15" pageNumber="56">
<emphasis id="08ABB11AFFADFFE9FF0DFF67FEE705F5" bold="true" box="[151,309,151,176]" pageId="15" pageNumber="56">Type locality.</emphasis>
<materialsCitation id="8AB76755FFADFFE9FEA6FF67FAB805F5" ID-GBIF-Occurrence="2028137805" box="[316,1386,151,176]" country="South Africa" latitude="-31.801449" location="Vanrhynsdorp" municipality="Gifberg" pageId="15" pageNumber="56" specimenCount="1" stateProvince="Western Cape">
<collectingCountry id="42C82D98FFADFFE9FEA6FF67FE1D05F5" box="[316,463,151,176]" name="South Africa" pageId="15" pageNumber="56">South Africa</collectingCountry>
,
<collectingRegion id="F81BA3EAFFADFFE9FE40FF68FD3105F5" box="[474,739,151,176]" country="South Africa" name="Western Cape" pageId="15" pageNumber="56">Western Cape Province</collectingRegion>
,
<collectingMunicipality id="DA04F772FFADFFE9FD74FF67FC9A05F5" box="[750,840,151,176]" pageId="15" pageNumber="56">Gifberg</collectingMunicipality>
,
<quantity id="FD27C0EDFFADFFE9FCC9FF67FC4805F5" box="[851,922,151,176]" metricMagnitude="4" metricUnit="m" metricValue="2.6" pageId="15" pageNumber="56" unit="km" value="26.0">26 km</quantity>
SE of,
<location id="3F003BD3FFADFFE9FC77FF67FB5805F5" LSID="urn:lsid:plazi:treatment:B276DC1EFFACFFE9FF0DF894FF23003D:3F003BD3FFADFFE9FC77FF67FB5805F5" box="[1005,1162,151,176]" country="South Africa" latitude="-31.801449" municipality="Gifberg" name="Vanrhynsdorp" pageId="15" pageNumber="56" stateProvince="Western Cape">Vanrhynsdorp</location>
<geoCoordinate id="5FEB0BCFFFADFFE9FB0BFF67FAC805F5" box="[1169,1306,151,176]" direction="south" orientation="latitude" pageId="15" pageNumber="56" precision="1" value="-31.801449">31°48'5.22S</geoCoordinate>
18°55'
</materialsCitation>
7E.
</paragraph>
<paragraph id="3A606D08FFADFFE9FF5DFF4CFC6704CD" blockId="15.[151,1437,151,1400]" pageId="15" pageNumber="56">
<emphasis id="08ABB11AFFADFFE9FF5DFF4CFEAB0590" bold="true" box="[199,377,188,213]" pageId="15" pageNumber="56">Type material.</emphasis>
<materialsCitation id="8AB76755FFADFFE9FE19FF4CFC060458" ID-GBIF-Occurrence="2028137799" collectingDate="2016-06-16" collectionCode="TMSA" collectorName="C. Scholtz" country="South Africa" county="Gifberg" elevation="377" latitude="-31.801449" location="Alt" municipality="Vanrhynsdorp" pageId="15" pageNumber="56" specimenCount="1" specimenCount-male="1" stateProvince="Western Cape" typeStatus="holotype">
<typeStatus id="E564D3AAFFADFFE9FE19FF4CFE3C0590" box="[387,494,188,213]" pageId="15" pageNumber="56" type="holotype">Holotype</typeStatus>
,
<specimenCount id="2CD9A681FFADFFE9FE67FF4DFDC00591" box="[509,530,189,212]" pageId="15" pageNumber="56" type="male">♂</specimenCount>
(
<collectionCode id="5CCEF5CDFFADFFE9FDBCFF4CFDA90591" box="[550,635,188,212]" country="South Africa" httpUri="http://grbio.org/cool/s8gp-ktsi" name="Transvaal Museum" pageId="15" pageNumber="56">TMSA</collectionCode>
): “
<collectingCountry id="42C82D98FFADFFE9FD01FF4CFCE00590" box="[667,818,188,213]" name="South Africa" pageId="15" pageNumber="56">South Africa</collectingCountry>
/
<collectingRegion id="F81BA3EAFFADFFE9FCD5FF4DFB8C0590" box="[847,1118,188,213]" country="South Africa" name="Western Cape" pageId="15" pageNumber="56">Western Cape Province</collectingRegion>
/
<collectingCounty id="D3011584FFADFFE9FBE1FF4CFB000590" box="[1147,1234,188,213]" pageId="15" pageNumber="56">Gifberg</collectingCounty>
/
<quantity id="FD27C0EDFFADFFE9FB6AFF4CFAE90590" box="[1264,1339,188,213]" metricMagnitude="4" metricUnit="m" metricValue="2.6" pageId="15" pageNumber="56" unit="km" value="26.0">26 km</quantity>
SE of /
<collectingMunicipality id="DA04F772FFADFFE9FF0DFF2FFEE605BD" box="[151,308,223,248]" pageId="15" pageNumber="56">Vanrhynsdorp</collectingMunicipality>
/
<geoCoordinate id="5FEB0BCFFFADFFE9FED3FF10FE3505BD" box="[329,487,223,248]" direction="south" orientation="latitude" pageId="15" pageNumber="56" precision="1" value="-31.801449">31° 48' 5.22 S</geoCoordinate>
18° 55' 7 E / 16 Km E /
<location id="3F003BD3FFADFFE9FD63FF2FFCCF05BD" LSID="urn:lsid:plazi:treatment:B276DC1EFFACFFE9FF0DF894FF23003D:3F003BD3FFADFFE9FD63FF2FFCCF05BD" box="[761,797,223,248]" country="South Africa" county="Gifberg" latitude="-31.801449" municipality="Vanrhynsdorp" name="Alt" pageId="15" pageNumber="56" stateProvince="Western Cape">Alt</location>
<quantity id="FD27C0EDFFADFFE9FCBEFF10FCB905B2" box="[804,875,224,248]" metricMagnitude="2" metricUnit="m" metricValue="3.77" pageId="15" pageNumber="56" unit="m" value="377.0">
<elevation id="B1F28A3BFFADFFE9FCBEFF10FCB905B2" box="[804,875,224,248]" metricMagnitude="2" metricUnit="m" metricValue="3.77" pageId="15" pageNumber="56" unit="m" value="377.0">377 m</elevation>
</quantity>
/
<date id="4E614BC8FFADFFE9FC1AFF2FFC2A05BD" box="[896,1016,223,248]" pageId="15" pageNumber="56" value="2016-06-16">
<collectingDate id="5E25B220FFADFFE9FC1AFF2FFC2A05BD" box="[896,1016,223,248]" pageId="15" pageNumber="56" value="2016-06-16">16.vi.2016</collectingDate>
</date>
/
<collectorName id="972A08DEFFADFFE9FB94FF10FB5005BD" box="[1038,1154,223,248]" pageId="15" pageNumber="56">C. Scholtz</collectorName>
[printed] //
<typeStatus id="E564D3AAFFADFFE9FA92FF10FA4E05BD" box="[1288,1436,224,248]" pageId="15" pageNumber="56" type="holotype">HOLOTYPE</typeStatus>
/
<taxonomicName id="FDDF168BFFADFFE9FF3DFEF4FE4B0459" authorityName="Abdalla &amp; Deschodt &amp; Scholtz &amp; Sole" authorityYear="2018" box="[167,409,260,284]" class="Insecta" family="Scarabaeidae" genus="Macroderes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="15" pageNumber="56" phylum="Arthropoda" rank="species" species="gifboomi">
<emphasis id="08ABB11AFFADFFE9FF3DFEF4FE4B0459" box="[167,409,260,284]" italics="true" pageId="15" pageNumber="56">Macroderes gifboomi</emphasis>
</taxonomicName>
/ Abdalla &amp; Scholtz / 2018 / [red label, printed]”
</materialsCitation>
.
<materialsCitation id="8AB76755FFADFFE9FC45FEF5FE810405" ID-GBIF-Occurrence="2028137810" collectingDate="2016-06-16" collectionCode="TMSA" collectorName="C. Scholtz" country="South Africa" county="Gifberg" elevation="377" latitude="-31.801449" location="Alt" municipality="Vanrhynsdorp" pageId="15" pageNumber="56" specimenCount="3" specimenCount-female="2" specimenCount-male="1" stateProvince="Western Cape" typeStatus="paratype">
<typeStatus id="E564D3AAFFADFFE9FC45FEF5FB820458" box="[991,1104,261,285]" pageId="15" pageNumber="56" type="paratype">Paratypes</typeStatus>
:
<specimenCount id="2CD9A681FFADFFE9FBC6FEF4FBAD0459" box="[1116,1151,260,284]" pageId="15" pageNumber="56" type="male">1♂</specimenCount>
and
<specimenCount id="2CD9A681FFADFFE9FB23FEF4FB230459" box="[1209,1265,260,284]" pageId="15" pageNumber="56" type="female">2♀♀</specimenCount>
(
<collectionCode id="5CCEF5CDFFADFFE9FA9BFEF4FA800459" box="[1281,1362,260,284]" country="South Africa" httpUri="http://grbio.org/cool/s8gp-ktsi" name="Transvaal Museum" pageId="15" pageNumber="56">TMSA</collectionCode>
) same data as holotype
</materialsCitation>
;
<materialsCitation id="8AB76755FFADFFE9FEFAFED8FC6004CD" ID-GBIF-Occurrence="2028137792" collectingDate="2016-07-21" collectionCode="UPSA" collectorName="C. Deschodt &amp; W. P. Strumpher" country="South Africa" county="Gifberg" elevation="377" latitude="-31.801449" location="Alt" longLatPrecision="1" longitude="18.918655" municipality="Vanrhynsdorp" pageId="15" pageNumber="56" specimenCount="4" specimenCount-male="4" stateProvince="Western Cape" typeStatus="paratype">
<specimenCount id="2CD9A681FFADFFE9FEFAFED8FE59047A" box="[352,395,296,320]" pageId="15" pageNumber="56" type="male">4 ♂</specimenCount>
(
<collectionCode id="5CCEF5CDFFADFFE9FE06FED8FE380405" box="[412,490,296,320]" country="South Africa" httpUri="http://grbio.org/cool/73qr-ciny" name="University of Pretoria" pageId="15" pageNumber="56">UPSA</collectionCode>
): “
<collectingCountry id="42C82D98FFADFFE9FD92FED7FD490405" box="[520,667,295,320]" name="South Africa" pageId="15" pageNumber="56">South Africa</collectingCountry>
/
<collectingRegion id="F81BA3EAFFADFFE9FD2EFED8FC6C0405" box="[692,958,295,320]" country="South Africa" name="Western Cape" pageId="15" pageNumber="56">Western Cape Province</collectingRegion>
/
<collectingCounty id="D3011584FFADFFE9FC4CFED7FBFF0405" box="[982,1069,295,320]" pageId="15" pageNumber="56">Gifberg</collectingCounty>
/
<quantity id="FD27C0EDFFADFFE9FBDFFED7FB5F0405" box="[1093,1165,295,320]" metricMagnitude="4" metricUnit="m" metricValue="2.6" pageId="15" pageNumber="56" unit="km" value="26.0">26 km</quantity>
SE of /
<collectingMunicipality id="DA04F772FFADFFE9FB75FED7FA5E0405" box="[1263,1420,295,320]" pageId="15" pageNumber="56">Vanrhynsdorp</collectingMunicipality>
/
<geoCoordinate id="5FEB0BCFFFADFFE9FF0DFEBCFEF80421" box="[151,298,332,357]" direction="south" orientation="latitude" pageId="15" pageNumber="56" precision="1" value="-31.801449">31°48'5.22 S</geoCoordinate>
<geoCoordinate id="5FEB0BCFFFADFFE9FEA9FEBCFE150421" box="[307,455,332,357]" direction="east" orientation="longitude" pageId="15" pageNumber="56" precision="1" value="18.918655">18°55'7.16 E</geoCoordinate>
/
<location id="3F003BD3FFADFFE9FE79FEBCFDD50420" LSID="urn:lsid:plazi:treatment:B276DC1EFFACFFE9FF0DF894FF23003D:3F003BD3FFADFFE9FE79FEBCFDD50420" box="[483,519,332,357]" country="South Africa" county="Gifberg" latitude="-31.801449" longLatPrecision="1" longitude="18.918655" municipality="Vanrhynsdorp" name="Alt" pageId="15" pageNumber="56" stateProvince="Western Cape">Alt</location>
<quantity id="FD27C0EDFFADFFE9FD8BFEBCFD820421" box="[529,592,332,356]" metricMagnitude="2" metricUnit="m" metricValue="3.77" pageId="15" pageNumber="56" unit="m" value="377.0">
<elevation id="B1F28A3BFFADFFE9FD8BFEBCFD820421" box="[529,592,332,356]" metricMagnitude="2" metricUnit="m" metricValue="3.77" pageId="15" pageNumber="56" unit="m" value="377.0">377m</elevation>
</quantity>
/
<date id="4E614BC8FFADFFE9FDF6FEBCFD3E0420" box="[620,748,332,357]" pageId="15" pageNumber="56" value="2016-07-21">
<collectingDate id="5E25B220FFADFFE9FDF6FEBCFD3E0420" box="[620,748,332,357]" pageId="15" pageNumber="56" value="2016-07-21">21.vii.2016</collectingDate>
</date>
/
<collectorName id="972A08DEFFADFFE9FC9DFEBCFC470420" box="[775,917,332,357]" pageId="15" pageNumber="56">C. Deschodt</collectorName>
&amp;
<collectorName id="972A08DEFFADFFE9FC24FEBDFBA20420" box="[958,1136,332,357]" pageId="15" pageNumber="56">W.P. Strümpher</collectorName>
[printed] //
<typeStatus id="E564D3AAFFADFFE9FA9AFEBCFA590421" box="[1280,1419,332,356]" pageId="15" pageNumber="56" type="paratype">PARATYPE</typeStatus>
/
<taxonomicName id="FDDF168BFFADFFE9FF0DFE80FE5504CD" authorityName="Abdalla &amp; Deschodt &amp; Scholtz &amp; Sole" authorityYear="2018" box="[151,391,368,392]" class="Insecta" family="Scarabaeidae" genus="Macroderes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="15" pageNumber="56" phylum="Arthropoda" rank="species" species="gifboomi">
<emphasis id="08ABB11AFFADFFE9FF0DFE80FE5504CD" box="[151,391,368,392]" italics="true" pageId="15" pageNumber="56">Macroderes gifboomi</emphasis>
</taxonomicName>
/ Abdalla &amp; Scholtz / 2018 / [red label, printed]”
</materialsCitation>
.
</paragraph>
<paragraph id="3A606D08FFADFFE9FF5DFE64FB7A04E9" blockId="15.[151,1437,151,1400]" box="[199,1192,404,429]" pageId="15" pageNumber="56">
<emphasis id="08ABB11AFFADFFE9FF5DFE64FE1A04E8" bold="true" box="[199,456,404,429]" pageId="15" pageNumber="56">
<typeStatus id="E564D3AAFFADFFE9FF5DFE64FEE604E8" box="[199,308,404,429]" pageId="15" pageNumber="56" type="holotype">Holotype</typeStatus>
description.
</emphasis>
<typeStatus id="E564D3AAFFADFFE9FE54FE64FDEB04E8" box="[462,569,404,429]" pageId="15" pageNumber="56" type="holotype">Holotype</typeStatus>
,
<specimenCount id="2CD9A681FFADFFE9FDDFFE65FD8804E9" box="[581,602,405,428]" pageId="15" pageNumber="56" type="male">♂</specimenCount>
(
<figureCitation id="A2E4718DFFADFFE9FDF3FE64FD6F04E9" box="[617,701,404,429]" captionStart="FIGURES 15–18" captionStartId="7.[151,264,1688,1711]" captionTargetBox="[314,1215,431,1621]" captionTargetId="figure@7.[309,1269,403,1653]" captionTargetPageId="7" captionText="FIGURES 15–18. Habitus of Macroderes species. 15, M. politulus Preudhomme de Borre, 1880; 16, M. undulatus Preudhomme de Borre, 1880; 17, M. porselinus Abdalla, new species; 18, M. gifboomi Abdalla &amp; Scholtz, new species." httpUri="https://zenodo.org/record/2606174/files/figure.png" pageId="15" pageNumber="56">Fig. 18</figureCitation>
). Body length
<quantity id="FD27C0EDFFADFFE9FCFBFE64FC6B04E9" box="[865,953,404,428]" metricMagnitude="-3" metricUnit="m" metricValue="9.9" pageId="15" pageNumber="56" unit="mm" value="9.9">9.9 mm</quantity>
, body width
<quantity id="FD27C0EDFFADFFE9FBD1FE64FB7604E9" box="[1099,1188,404,428]" metricMagnitude="-3" metricUnit="m" metricValue="7.2" pageId="15" pageNumber="56" unit="mm" value="7.2">7.2 mm</quantity>
.
</paragraph>
<paragraph id="3A606D08FFADFFE9FF5DFE47FAFC0778" blockId="15.[151,1437,151,1400]" pageId="15" pageNumber="56">
<emphasis id="08ABB11AFFADFFE9FF5DFE47FEDF0495" bold="true" box="[199,269,439,464]" pageId="15" pageNumber="56">Head.</emphasis>
Clypeus broad, semi–circular, bordered by two teeth on anterior edge which are wider than longer. Genae right angled. Frontoclypeal suture clearly evident slightly curved without tubercle in the middle. Frons densely punctate with small round punctures. Clypeogenal suture present but poorly defined towards the clypeal edge. Clypeus densely punctate with fine punctures becoming rugose towards the clypeal frontal margin.
</paragraph>
</subSubSection>
<subSubSection id="72C53E83FFADFFE9FF5DFDB7FD040170" pageId="15" pageNumber="56" type="description">
<paragraph id="3A606D08FFADFFE9FF5DFDB7FCD607B5" blockId="15.[151,1437,151,1400]" pageId="15" pageNumber="56">
<emphasis id="08ABB11AFFADFFE9FF5DFDB7FE950725" bold="true" box="[199,327,583,608]" pageId="15" pageNumber="56">Pronotum.</emphasis>
Form trapezoidal, smooth, shiny, width greater than length, with two distinct depressions anterolaterally. Lateral edges rounded delimited with fine borders. Borders impunctate, non–crenulated, base not bordered. Anterior angles obtuse, posterior angles right to obtuse. Pronotal surface lacking a longitudinal depression and uniformly covered with regular round punctures with yellow setae located peripherally (
<figureCitation id="A2E4718DFFADFFE9FAA2FD44FA450789" box="[1336,1431,692,717]" captionStart="FIGURES 76–81" captionStartId="20.[151,264,1637,1660]" captionTargetBox="[171,1423,205,1613]" captionTargetId="figure@20.[171,1423,205,1613]" captionTargetPageId="20" captionText="FIGURES 76–81. Punctuation of the base of pronotum of Macroderes species. 76, M. cederbergensis Abdalla &amp; Deschodt, new species; 77, M. gifboomi Abdalla &amp; Scholtz, new species; 78, M. oreatus Abdalla &amp; Deschodt, new species; 79, M. porselinus Abdalla, new species; 80, M. soleiana Abdalla &amp; Deschodt, new species; 81, M. leipoldti Abdalla &amp; Deschodt, new species." httpUri="https://zenodo.org/record/2606198/files/figure.png" pageId="15" pageNumber="56">Figs. 77</figureCitation>
,
<figureCitation id="A2E4718DFFADFFE9FF0DFD28FF6507B5" box="[151,183,728,752]" captionStart="FIGURES 82–87" captionStartId="21.[151,264,1505,1528]" captionTargetBox="[316,1276,406,1475]" captionTargetId="figure@21.[316,1276,406,1475]" captionTargetPageId="21" captionText="FIGURES 82–87. Pronotum punctures shape of Macroderes species. 82, M. tourtosus Abdalla &amp; Scholtz, new species; 83, M. cederbergensis Abdalla &amp; Deschodt, new species; 84, M. gifboomi Abdalla &amp; Scholtz, new species; 85, M. leipoldti Abdalla &amp; Deschodt, new species; 86, M. oreatus Abdalla &amp; Deschodt, new species; 87, M. porselinus Abdalla, new species." httpUri="https://zenodo.org/record/2606200/files/figure.png" pageId="15" pageNumber="56">84</figureCitation>
). Punctures spaced by 2–4 times their own diameter.
</paragraph>
<paragraph id="3A606D08FFADFFE9FF5DFD0CFF2E06C5" blockId="15.[151,1437,151,1400]" pageId="15" pageNumber="56">
<emphasis id="08ABB11AFFADFFE9FF5DFD0CFEC80650" bold="true" box="[199,282,764,789]" pageId="15" pageNumber="56">Elytra.</emphasis>
Elytral intervals convex, shiny, densely punctate with small round punctures. Punctures separated by shiny elevated areas by approximately 1–4 their own puncture diameter (
<figureCitation id="A2E4718DFFADFFE9FC63FCEFFB83067D" box="[1017,1105,799,824]" captionStart="FIGURES 88–93" captionStartId="23.[151,264,1292,1315]" captionTargetBox="[316,1276,183,1264]" captionTargetId="figure@23.[316,1276,183,1264]" captionTargetPageId="23" captionText="FIGURES 88–93. Pronotum punctures shape (88, 89) and sculpture of second elytra intervals (90–93) of Macroderes species. 88, M. soleiana Abdalla &amp; Deschodt, new species; 89, M. tourtosus Abdalla and Scholtz, new species; 90, M. minutus Frolov and Scholtz, 2005; 91, M. greeni (Kirby, 1818); 92, M. politulus Preudhomme de Borre, 1880; 93, M. gifboomi Abdalla &amp; Scholtz, new species." httpUri="https://zenodo.org/record/2606202/files/figure.png" pageId="15" pageNumber="56">Fig. 93</figureCitation>
). Striae punctate with small punctures with short yellow setae/spaced by 2–4 punctures diameter, stria 9 closest to stria 10 about 2/3 the length of elytra.
</paragraph>
<paragraph id="3A606D08FFADFFE9FF5DFC7CFAB706E1" blockId="15.[151,1437,151,1400]" box="[199,1381,908,933]" pageId="15" pageNumber="56">
<emphasis id="08ABB11AFFADFFE9FF5DFC7CFE9306E0" bold="true" box="[199,321,908,933]" pageId="15" pageNumber="56">Aedeagus.</emphasis>
Sclerite of internal sac with long lateral process and rather short and strait basal part (
<figureCitation id="A2E4718DFFADFFE9FA9EFC7CFA8A06E1" box="[1284,1368,908,933]" captionStart="FIGURES 24–39" captionStartId="10.[151,264,1343,1366]" captionTargetBox="[323,1250,205,1286]" captionTargetId="figure@10.[323,1283,195,1303]" captionTargetPageId="10" captionText="FIGURES 24–39. Internal sac of aedeagus (24), aedeagus (25) and internal sac sclerite (26-39) of Macroderes species. 26, M. endroedyi Frolov &amp; Scholtz, 2005; 27, M. amplior Frolov &amp; Scholtz, 2005; 28, M. bias (Olivier, 1789); 29, M. cederbergensis Abdalla &amp; Deschodt, new species; 30, M. cornutus Frolov &amp; Scholtz, 2005; 31, M. foveatus Frolov &amp; Scholtz, 2005; 32, M. gifboomi Abdalla &amp; Scholtz, new species; 33, M. leipoldti Abdalla &amp; Deschodt, new species; 34, M. mutilans Kolbe, 1908; 35, M. namakwanus Frolov &amp; Scholtz, 2005; 36, M. oreatus Abdalla &amp; Deschodt, new species; 37, M. porselinus Abdalla, new species; 38, M. soleiana Abdalla &amp; Deschodt, new species; 39, M. tourtosus Abdalla &amp; Scholtz, new species." httpUri="https://zenodo.org/record/2606178/files/figure.png" pageId="15" pageNumber="56">Fig. 32</figureCitation>
).
</paragraph>
<paragraph id="3A606D08FFADFFE9FF5DFC5FFF1806A9" blockId="15.[151,1437,151,1400]" pageId="15" pageNumber="56">
<emphasis id="08ABB11AFFADFFE9FF5DFC5FFE9F068D" bold="true" box="[199,333,943,968]" pageId="15" pageNumber="56">Size range.</emphasis>
Males length:
<quantity id="FD27C0EDFFADFFE9FE64FC40FD4A0682" box="[510,664,944,968]" metricMagnitude="-3" metricUnit="m" metricValue="9.75" metricValueMax="10.1" metricValueMin="9.4" pageId="15" pageNumber="56" unit="mm" value="9.75" valueMax="10.1" valueMin="9.4">9.4–10.1 mm</quantity>
, width:
<quantity id="FD27C0EDFFADFFE9FD62FC5FFC540682" box="[760,902,943,968]" metricMagnitude="-3" metricUnit="m" metricValue="6.9" metricValueMax="7.1" metricValueMin="6.7" pageId="15" pageNumber="56" unit="mm" value="6.9" valueMax="7.1" valueMin="6.7">6.7–7.1 mm</quantity>
; females length:
<quantity id="FD27C0EDFFADFFE9FBD7FC5FFB3A0682" box="[1101,1256,943,968]" metricMagnitude="-3" metricUnit="m" metricValue="9.899999999999999" metricValueMax="10.6" metricValueMin="9.2" pageId="15" pageNumber="56" unit="mm" value="9.899999999999999" valueMax="10.6" valueMin="9.2">9.2–10.6 mm</quantity>
, width:
<quantity id="FD27C0EDFFADFFE9FAD2FC5FFF1406A9" metricMagnitude="-3" metricUnit="m" metricValue="6.949999999999999" metricValueMax="7.3" metricValueMin="6.6" pageId="15" pageNumber="56" unit="mm" value="6.949999999999999" valueMax="7.3" valueMin="6.6">6.6–7.3 mm</quantity>
.
</paragraph>
<paragraph id="3A606D08FFADFFE9FF5DFC07FD040170" blockId="15.[151,1437,151,1400]" pageId="15" pageNumber="56">
<emphasis id="08ABB11AFFADFFE9FF5DFC07FEF70155" bold="true" box="[199,293,1015,1040]" pageId="15" pageNumber="56">Female.</emphasis>
Female differs from male by having well developed tubercle in the middle of the frontoclypeal suture as well as less excavated pronotum anteriolaterally.
</paragraph>
</subSubSection>
<subSubSection id="72C53E83FFADFFE9FF5DFBCFFB930139" pageId="15" pageNumber="56" type="etymology">
<paragraph id="3A606D08FFADFFE9FF5DFBCFFB930139" blockId="15.[151,1437,151,1400]" pageId="15" pageNumber="56">
<emphasis id="08ABB11AFFADFFE9FF5DFBCFFE9C011D" bold="true" box="[199,334,1087,1112]" pageId="15" pageNumber="56">Etymology.</emphasis>
This species name reflects the Afrikaans name,
<emphasis id="08ABB11AFFADFFE9FC28FBB0FBDC011D" box="[946,1038,1088,1112]" italics="true" pageId="15" pageNumber="56">gifboom</emphasis>
, of the evergreen Fynbos shrub
<taxonomicName id="FDDF168BFFADFFE9FF0DFB94FD440138" authority="(Gaertn.) Lamb. &amp; Vahl" authorityName="Lamb. &amp; Vahl" baseAuthorityName="Gaertn." box="[151,662,1124,1149]" class="Magnoliopsida" family="Picrodendraceae" genus="Hyaenanche" higherTaxonomySource="GBIF" kingdom="Plantae" order="Malpighiales" pageId="15" pageNumber="56" phylum="Tracheophyta" rank="species" species="globosa">
<emphasis id="08ABB11AFFADFFE9FF0DFB94FE510139" box="[151,387,1124,1148]" italics="true" pageId="15" pageNumber="56">Hyaenanche globosa</emphasis>
(Gaertn.) Lamb. &amp; Vahl
</taxonomicName>
which is endemic to the Gifberg area.
</paragraph>
</subSubSection>
<subSubSection id="72C53E83FFADFFE9FF5DFB77FF1101AD" pageId="15" pageNumber="56" type="diagnosis">
<paragraph id="3A606D08FFADFFE9FF5DFB77FF1101AD" blockId="15.[151,1437,151,1400]" pageId="15" pageNumber="56">
<emphasis id="08ABB11AFFADFFE9FF5DFB77FE0301E5" bold="true" box="[199,465,1159,1184]" pageId="15" pageNumber="56">Differential diagnosis.</emphasis>
This species is most similar to
<taxonomicName id="FDDF168BFFADFFE9FCDEFB79FC6B01E5" authorityName="Harold /" authorityYear="1877" box="[836,953,1160,1184]" class="Insecta" family="Scarabaeidae" genus="Macroderes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="15" pageNumber="56" phylum="Arthropoda" rank="species" species="nitidus">
<emphasis id="08ABB11AFFADFFE9FCDEFB79FC6B01E5" box="[836,953,1160,1184]" italics="true" pageId="15" pageNumber="56">M. nitidus</emphasis>
</taxonomicName>
in having a pronotum with regular round punctures (
<figureCitation id="A2E4718DFFADFFE9FE8EFB5CFEA10181" box="[276,371,1196,1221]" captionStart="FIGURES 76–81" captionStartId="20.[151,264,1637,1660]" captionTargetBox="[171,1423,205,1613]" captionTargetId="figure@20.[171,1423,205,1613]" captionTargetPageId="20" captionText="FIGURES 76–81. Punctuation of the base of pronotum of Macroderes species. 76, M. cederbergensis Abdalla &amp; Deschodt, new species; 77, M. gifboomi Abdalla &amp; Scholtz, new species; 78, M. oreatus Abdalla &amp; Deschodt, new species; 79, M. porselinus Abdalla, new species; 80, M. soleiana Abdalla &amp; Deschodt, new species; 81, M. leipoldti Abdalla &amp; Deschodt, new species." httpUri="https://zenodo.org/record/2606198/files/figure.png" pageId="15" pageNumber="56">Figs. 77</figureCitation>
,
<figureCitation id="A2E4718DFFADFFE9FE1BFB5CFE720181" box="[385,416,1196,1220]" captionStart="FIGURES 82–87" captionStartId="21.[151,264,1505,1528]" captionTargetBox="[316,1276,406,1475]" captionTargetId="figure@21.[316,1276,406,1475]" captionTargetPageId="21" captionText="FIGURES 82–87. Pronotum punctures shape of Macroderes species. 82, M. tourtosus Abdalla &amp; Scholtz, new species; 83, M. cederbergensis Abdalla &amp; Deschodt, new species; 84, M. gifboomi Abdalla &amp; Scholtz, new species; 85, M. leipoldti Abdalla &amp; Deschodt, new species; 86, M. oreatus Abdalla &amp; Deschodt, new species; 87, M. porselinus Abdalla, new species." httpUri="https://zenodo.org/record/2606200/files/figure.png" pageId="15" pageNumber="56">84</figureCitation>
), however the pronotum is more convex and the whole body shinier than
<taxonomicName id="FDDF168BFFADFFE9FB71FB5DFA8F0181" authorityName="Harold /" authorityYear="1877" box="[1259,1373,1196,1220]" class="Insecta" family="Scarabaeidae" genus="Macroderes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="15" pageNumber="56" phylum="Arthropoda" rank="species" species="nitidus">
<emphasis id="08ABB11AFFADFFE9FB71FB5DFA8F0181" box="[1259,1373,1196,1220]" italics="true" pageId="15" pageNumber="56">M. nitidus</emphasis>
</taxonomicName>
(
<figureCitation id="A2E4718DFFADFFE9FAF4FB5CFF6401AD" captionStart="FIGURES 15–18" captionStartId="7.[151,264,1688,1711]" captionTargetBox="[314,1215,431,1621]" captionTargetId="figure@7.[309,1269,403,1653]" captionTargetPageId="7" captionText="FIGURES 15–18. Habitus of Macroderes species. 15, M. politulus Preudhomme de Borre, 1880; 16, M. undulatus Preudhomme de Borre, 1880; 17, M. porselinus Abdalla, new species; 18, M. gifboomi Abdalla &amp; Scholtz, new species." httpUri="https://zenodo.org/record/2606174/files/figure.png" pageId="15" pageNumber="56">Fig. 18</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="72C53E83FFADFFE9FF5DFB04FF23003D" pageId="15" pageNumber="56" type="description">
<paragraph id="3A606D08FFADFFE9FF5DFB04FF23003D" blockId="15.[151,1437,151,1400]" pageId="15" pageNumber="56">
<emphasis id="08ABB11AFFADFFE9FF5DFB04FE210048" bold="true" box="[199,499,1268,1293]" pageId="15" pageNumber="56">Habitat and distribution.</emphasis>
This species was collected from the base of the Gifberg (
<figureCitation id="A2E4718DFFADFFE9FBECFB04FB680049" box="[1142,1210,1268,1293]" captionStart="FIGURE 2" captionStartId="2.[151,250,1574,1596]" captionText="FIGURE 2. Distribution map of Macroderes species." pageId="15" pageNumber="56">Fig. 2</figureCitation>
) at the northern end of the Bokkeveld Escarpment Mountain Plateau just south of Vanrhynsdorp. These mountains consist of Bokkeveld Sandstone Fynbos (
<bibRefCitation id="5E4E10F9FFADFFE9FEE0FACCFD100010" author="Mucina, L. &amp; Rutherford, M. C." box="[378,706,1340,1365]" pageId="15" pageNumber="56" pagination="1 - 807" refId="ref21930" refString="Mucina, L. &amp; Rutherford, M. C. (Eds.) (2006) The Vegetation of South Africa, Lesotho and Swaziland. Strelitzia, 19, 1 - 807." type="journal article" year="2006">Mucina &amp; Rutherford 2006</bibRefCitation>
). The habitat is very rocky with shallow soil, poor in mineral content.
</paragraph>
</subSubSection>
</treatment>
</document>
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