treatments-xml/data/20/2D/EE/202DEE3AFFDA670AE7C8FA88FC9AF862.xml

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<mods:title id="C121A51240624F4D6DE9E1FC45541C25">A new species of the greeneye genus Chlorophthalmus (Teleostei: Chlorophthalmidae) from the central North Pacific</mods:title>
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<mods:namePart id="88E9CADAB455BA1A87642B1C35CB33F8">Wada, Hidetoshi</mods:namePart>
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<emphasis id="9AF0833EFFDA6703E7C8FA88FDE2FAFA" bold="true" box="[151,592,1362,1389]" pageId="1" pageNumber="397">
<taxonomicName id="6F8424AFFFDA6703E7C8FA88FE59FAFB" authority="Fujiwara &amp; Wada &amp; Motomura, 2019" authorityName="Fujiwara &amp; Wada &amp; Motomura" authorityYear="2019" box="[151,491,1362,1388]" class="Actinopterygii" family="Chlorophthalmidae" genus="Chlorophthalmus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Aulopiformes" pageId="1" pageNumber="397" phylum="Chordata" rank="species" species="imperator" status="sp. nov.">
<emphasis id="9AF0833EFFDA6703E7C8FA88FE59FAFB" bold="true" box="[151,491,1362,1388]" italics="true" pageId="1" pageNumber="397">Chlorophthalmus imperator</emphasis>
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<paragraph id="A83B5F2CFFDA6703E7C8FAACFDFDFA19" blockId="1.[151,592,1362,1458]" box="[151,591,1397,1422]" pageId="1" pageNumber="397">New English name: Emperor Greeneye</paragraph>
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<subSubSection id="E09E0CA7FFDA6703E7C8FA43FE2BFA25" box="[151,409,1433,1458]" pageId="1" pageNumber="397" type="description">
<paragraph id="A83B5F2CFFDA6703E7C8FA43FE2BFA25" blockId="1.[151,592,1362,1458]" box="[151,409,1433,1458]" pageId="1" pageNumber="397">
<figureCitation id="30BF43A9FFDA6703E7C8FA43FEACFA25" box="[151,286,1433,1458]" captionStart-0="FIGURE 1" captionStart-1="FIGURE 2" captionStart-2="FIGURE 3" captionStart-3="FIGURE 4" captionStartId-0="2.[151,250,1820,1842]" captionStartId-1="3.[151,250,964,986]" captionStartId-2="4.[151,250,1793,1815]" captionStartId-3="5.[151,250,1070,1092]" captionTargetBox-0="[340,1247,483,1795]" captionTargetBox-1="[331,1238,213,937]" captionTargetBox-2="[340,1247,481,1732]" captionTargetBox-3="[376,1190,301,1043]" captionTargetId-0="figure@2.[340,1247,483,1795]" captionTargetId-1="figure@3.[331,1238,213,937]" captionTargetId-2="figure@4.[340,1247,481,1732]" captionTargetId-3="figure@5.[340,1247,265,1048]" captionTargetPageId-0="2" captionTargetPageId-1="3" captionTargetPageId-2="4" captionTargetPageId-3="5" captionText-0="FIGURE 1. Preserved specimens of Chlorophthalmus imperator sp. nov. collected from the Emperor Seamount Chain in the central North Pacific. (A–C), KAUM–I. 120091, 130.0 mm SL, holotype; (D), KAUM–I. 120099, 174.8 mm SL, paratype; (E), KAUM–I. 120094, 146.6 mm SL, paratype; (F), KAUM–I. 120097, 98.1 mm SL, paratype. (A, D–F), lateral views; (B), dorsal view; (C), ventral view." captionText-1="FIGURE 2. Lower-jaw tips of Chlorophthalmus imperator sp. nov. in dorsal view. (A), KAUM–I. 120097, paratype; (B), KAUM–I. 120098, paratype; (C), KAUM–I. 120091, holotype; (D), KAUM–I. 120093, paratype; (E), KAUM–I. 120099, paratype; (F), KAUM–I. 120096, paratype (damaged)." captionText-2="FIGURE 3. Fresh specimens of Chlorophthalmus imperator sp. nov. (A), KAUM–I. 120091, 130.0 mm SL, holotype; (B), KAUM–I. 120095, 122.1 mm SL, paratype; (C), KAUM–I. 120098, 123.6 mm SL, paratype; (D), KAUM–I. 120094, 146.6 mm SL, paratype. Photos by SNFR." captionText-3="FIGURE 4. Distribution of Chlorophthalmus imperator sp. nov. (red squares), arrowhead indicates type locality." httpUri-0="https://zenodo.org/record/2624532/files/figure.png" httpUri-1="https://zenodo.org/record/2624534/files/figure.png" httpUri-2="https://zenodo.org/record/2624536/files/figure.png" httpUri-3="https://zenodo.org/record/2624538/files/figure.png" pageId="1" pageNumber="397">Figures 1–4</figureCitation>
, 
<figureCitation id="30BF43A9FFDA6703E675FA43FE88FA25" box="[298,314,1433,1458]" captionStart="FIGURE 6" captionStartId="7.[151,250,1097,1119]" captionTargetBox="[316,1206,202,995]" captionTargetId="figure@7.[314,1221,195,997]" captionTargetPageId="7" captionText="FIGURE 6. Relationships of (A) head length, (B) snout length, (C) upper-jaw length, (D) maxillary depth, (E) horizontal orbit diamter, and (F) pre-pectoral-fin length (all as % of standard length) to standard length (mm) in Chlorophthalmus imperator sp. nov. (red squares) and C. proridens (blue circles). Arrowheads indicate holotype." httpUri="https://zenodo.org/record/2624542/files/figure.png" pageId="1" pageNumber="397">6</figureCitation>
; 
<tableCitation id="E5066A97FFDA6703E619FA43FE2BFA25" box="[326,409,1433,1458]" captionStart="TABLE 1" captionStartId="7.[151,239,1220,1243]" captionText="TABLE 1. Counts and measurements, expressed as percentages of standard length, of Chlorophthalmus imperator sp. nov." pageId="1" pageNumber="397">Table 1</tableCitation>
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<materialsCitation id="18EC5571FFDA6703E7C8FA3BFE4BF9D5" ID-GBIF-Occurrence="2234227369" collectingDate="2015-08-06" collectionCode="KAUM-I, RV" county="Kaiyou-maru" elevation="601" latitude="32.260834" location="Daikakuji Seamount" longLatPrecision="20" longitude="172.88805" municipality="central North Pacific" pageId="1" pageNumber="397" specimenCode="SNFR 21135" specimenCount="1" typeStatus="holotype">
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<typeStatus id="773FE18EFFDA6703E7C8FA3BFEB4FA6D" box="[151,262,1505,1530]" pageId="1" pageNumber="397" type="holotype">Holotype</typeStatus>
.
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. 120091 (formerly 
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), 130.0 mm SL, 
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, 
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, 
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, 
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, 
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, midwater trawl, 
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, 
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, 
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<date id="DC3A79ECFFDA6703E639F9F3FE47F9D5" box="[358,501,1577,1602]" pageId="1" pageNumber="397" value="2015-08-06">6 Aug. 2015</date>
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.
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, 
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SL, all from the 
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in the 
<collectingMunicipality id="485FC556FFDA6703E25CF997FF57F91D" pageId="1" pageNumber="397">central North Pacific</collectingMunicipality>
(formerly deposited at 
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). 
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. 
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SL, 
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SL, 
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, 
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, 
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, 
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, bottom trawl, 
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, 
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FV 
<emphasis id="9AF0833EFFDA6703E24FF94CFA27F93A" box="[1296,1429,1686,1709]" italics="true" pageId="1" pageNumber="397">Hiroe-maru</emphasis>
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</materialsCitation>
; 
<materialsCitation id="18EC5571FFDA6703E614F960FC37F961" ID-GBIF-Occurrence="2234227269" collectingDate="2008-12-08" collectionCode="KAUM-I" collectorName="FV Tomi-maru" elevation="366" latitude="32.283333" location="Daikakuji Seamount" longLatPrecision="20" longitude="172.86667" pageId="1" pageNumber="397" specimenCount="1" typeStatus="paratype">
<collectionCode id="CE95C7E9FFDA6703E614F960FE0EF945" box="[331,444,1722,1746]" pageId="1" pageNumber="397">KAUM–I</collectionCode>
. 
<quantity id="6F7CF2C9FFDA6703E696F960FD16F946" box="[457,676,1721,1746]" metricMagnitude="5" metricUnit="m" metricValue="1.200941466" pageId="1" pageNumber="397" unit="mm" value="1.200941466E8">120094, 146.6 mm</quantity>
SL, 
<location id="AD5B09F7FFDA6703E5BFF963FC7EF945" LSID="urn:lsid:plazi:treatment:202DEE3AFFDA670AE7C8FA88FC9AF862:AD5B09F7FFDA6703E5BFF963FC7EF945" box="[736,972,1721,1746]" latitude="32.283333" longLatPrecision="20" longitude="172.86667" name="Daikakuji Seamount" pageId="1" pageNumber="397">Daikakuji Seamount</location>
, 
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, 
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, 
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, bottom trawl, 
<quantity id="6F7CF2C9FFDA6703E682F907FD9BF962" box="[477,553,1757,1782]" metricMagnitude="2" metricUnit="m" metricValue="3.66" pageId="1" pageNumber="397" unit="m" value="366.0">
<elevation id="23A9B81FFFDA6703E682F907FD9BF962" box="[477,553,1757,1782]" metricMagnitude="2" metricUnit="m" metricValue="3.66" pageId="1" pageNumber="397" unit="m" value="366.0">366 m</elevation>
</quantity>
, 
<collectorName id="05713AFAFFDA6703E567F904FD53F962" box="[568,737,1758,1781]" pageId="1" pageNumber="397">
FV 
<emphasis id="9AF0833EFFDA6703E539F904FD53F962" box="[614,737,1758,1781]" italics="true" pageId="1" pageNumber="397">Tomi-maru</emphasis>
</collectorName>
, 
<date id="DC3A79ECFFDA6703E5ACF904FC37F961" box="[755,901,1758,1782]" pageId="1" pageNumber="397" value="2008-12-08">
<collectingDate id="CC7E8004FFDA6703E5ACF904FC37F961" box="[755,901,1758,1782]" pageId="1" pageNumber="397" value="2008-12-08">8 Dec. 2008</collectingDate>
</date>
</materialsCitation>
; 
<materialsCitation id="18EC5571FFDA6703E4CCF904FA25F8A9" ID-GBIF-Occurrence="2234227293" collectingDate="2015-08-04" collectionCode="KAUM-I, RV" collectorName="Kaiyou-maru" elevation="423" latitude="32.000557" location="Kammu Seamount" longLatPrecision="20" longitude="173.13777" pageId="1" pageNumber="397" specimenCount="1" typeStatus="paratype">
<collectionCode id="CE95C7E9FFDA6703E4CCF904FBB6F961" box="[915,1028,1758,1782]" pageId="1" pageNumber="397">KAUM–I</collectionCode>
. 
<quantity id="6F7CF2C9FFDA6703E34EF907FB5EF962" box="[1041,1260,1757,1782]" metricMagnitude="5" metricUnit="m" metricValue="1.200951221" pageId="1" pageNumber="397" unit="mm" value="1.200951221E8">120095, 122.1 mm</quantity>
SL, 
<collectionCode id="CE95C7E9FFDA6703E277F904FA2BF961" box="[1320,1433,1758,1782]" pageId="1" pageNumber="397">KAUM–I</collectionCode>
. 120096, 103.0 mm SL, 
<collectionCode id="CE95C7E9FFDA6703E6F4F8D8FDAEF88D" box="[427,540,1794,1818]" pageId="1" pageNumber="397">KAUM–I</collectionCode>
. 120097, 
<quantity id="6F7CF2C9FFDA6703E5D1F8D8FD46F88E" box="[654,756,1794,1818]" metricMagnitude="-2" metricUnit="m" metricValue="9.809999999999999" pageId="1" pageNumber="397" unit="mm" value="98.1">98.1 mm</quantity>
SL, 
<collectionCode id="CE95C7E9FFDA6703E470F8D8FC12F88D" box="[815,928,1794,1818]" pageId="1" pageNumber="397">KAUM–I</collectionCode>
. 
<quantity id="6F7CF2C9FFDA6703E4F2F8D8FB34F88E" box="[941,1158,1793,1818]" metricMagnitude="5" metricUnit="m" metricValue="1.200981236" pageId="1" pageNumber="397" unit="mm" value="1.200981236E8">120098, 123.6 mm</quantity>
SL, 
<location id="AD5B09F7FFDA6703E39EF8D8FA2AF88D" LSID="urn:lsid:plazi:treatment:202DEE3AFFDA670AE7C8FA88FC9AF862:AD5B09F7FFDA6703E39EF8D8FA2AF88D" box="[1217,1432,1794,1818]" latitude="32.000557" longLatPrecision="20" longitude="173.13777" name="Kammu Seamount" pageId="1" pageNumber="397">Kammu Seamount</location>
, 
<geoCoordinate id="CDB039EBFFDA6703E7C8F8FFFEADF8A9" box="[151,287,1829,1854]" direction="north" orientation="latitude" pageId="1" pageNumber="397" precision="15" value="32.024445">32°01′28″N</geoCoordinate>
, 
<geoCoordinate id="CDB039EBFFDA6703E675F8FFFE74F8A9" box="[298,454,1829,1854]" direction="east" orientation="longitude" pageId="1" pageNumber="397" precision="15" value="173.18889">173°11′20″E–</geoCoordinate>
<geoCoordinate id="CDB039EBFFDA6703E699F8FFFDFDF8A9" box="[454,591,1829,1854]" direction="north" orientation="latitude" pageId="1" pageNumber="397" precision="15" value="32.000557">32°00′02″N</geoCoordinate>
, 
<geoCoordinate id="CDB039EBFFDA6703E505F8FFFD5FF8A9" box="[602,749,1829,1854]" direction="east" orientation="longitude" pageId="1" pageNumber="397" precision="15" value="173.13777">173°08′16″E</geoCoordinate>
, midwater trawl, 
<quantity id="6F7CF2C9FFDA6703E4F0F8FCFB82F8AA" box="[943,1072,1830,1854]" metricMagnitude="2" metricUnit="m" metricValue="4.2299999999999995" metricValueMax="4.970000000000001" metricValueMin="3.4899999999999998" pageId="1" pageNumber="397" unit="m" value="423.0" valueMax="497.0" valueMin="349.0">
<elevation id="23A9B81FFFDA6703E4F0F8FCFB82F8AA" box="[943,1072,1830,1854]" metricMagnitude="2" metricUnit="m" metricValue="4.2299999999999995" metricValueMax="4.970000000000001" metricValueMin="3.4899999999999998" pageId="1" pageNumber="397" unit="m" value="423.0" valueMax="497.0" valueMin="349.0">349–497 m</elevation>
</quantity>
, 
<collectionCode id="CE95C7E9FFDA6703E364F8FCFBD2F8AA" box="[1083,1120,1830,1853]" country="Italy" name="Collection of Leptospira Strains" pageId="1" pageNumber="397">RV</collectionCode>
<emphasis id="9AF0833EFFDA6703E339F8FCFB49F8AA" box="[1126,1275,1830,1853]" italics="true" pageId="1" pageNumber="397">
<collectorName id="05713AFAFFDA6703E339F8FCFB49F8AA" box="[1126,1275,1830,1853]" pageId="1" pageNumber="397">Kaiyou-maru</collectorName>
</emphasis>
, 
<date id="DC3A79ECFFDA6703E255F8FCFA25F8A9" box="[1290,1431,1829,1854]" pageId="1" pageNumber="397" value="2015-08-04">
<collectingDate id="CC7E8004FFDA6703E255F8FCFA25F8A9" box="[1290,1431,1829,1854]" pageId="1" pageNumber="397" value="2015-08-04">4 Aug. 2015</collectingDate>
</date>
</materialsCitation>
; 
<materialsCitation id="18EC5571FFDA6703E7C8F890FC3DF8F5" ID-GBIF-Occurrence="2234227394" box="[151,911,1865,1890]" collectingDate="2015-07-31" collectionCode="KAUM-I" collectorName="Kammu Seamount" pageId="1" pageNumber="397" specimenCount="1" typeStatus="paratype">
<collectionCode id="CE95C7E9FFDA6703E7C8F890FEBAF8F5" box="[151,264,1866,1890]" pageId="1" pageNumber="397">KAUM–I</collectionCode>
. 
<quantity id="6F7CF2C9FFDA6703E64DF890FE54F8F6" box="[274,486,1866,1890]" metricMagnitude="5" metricUnit="m" metricValue="1.2009917479999999" pageId="1" pageNumber="397" unit="mm" value="1.200991748E8">120099, 174.8 mm</quantity>
SL, 
<collectorName id="05713AFAFFDA6703E544F890FD5DF8F5" box="[539,751,1866,1890]" pageId="1" pageNumber="397">Kammu Seamount</collectorName>
, 
<date id="DC3A79ECFFDA6703E5A5F890FC3DF8F5" box="[762,911,1865,1890]" pageId="1" pageNumber="397" value="2015-07-31">
<collectingDate id="CC7E8004FFDA6703E5A5F890FC3DF8F5" box="[762,911,1865,1890]" pageId="1" pageNumber="397" value="2015-07-31">31 July 2015</collectingDate>
</date>
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="E09E0CA7FFDA6700E798F8B7FBAEFED7" lastPageId="2" lastPageNumber="398" pageId="1" pageNumber="397" type="diagnosis">
<paragraph id="A83B5F2CFFDA6700E798F8B7FBAEFED7" blockId="1.[151,1437,1505,2034]" lastBlockId="2.[151,1437,151,429]" lastPageId="2" lastPageNumber="398" pageId="1" pageNumber="397">
<emphasis id="9AF0833EFFDA6703E798F8B7FEF3F811" bold="true" box="[199,321,1901,1926]" pageId="1" pageNumber="397">Diagnosis.</emphasis>
A relatively large species (standard length exceeding 
<quantity id="6F7CF2C9FFDA6703E496F8B4FB9DF812" box="[969,1071,1902,1926]" metricMagnitude="-1" metricUnit="m" metricValue="1.7" pageId="1" pageNumber="397" unit="mm" value="170.0">170 mm</quantity>
: 
<figureCitation id="30BF43A9FFDA6703E361F8B7FB2BF811" box="[1086,1177,1901,1926]" captionStart="FIGURE 1" captionStartId="2.[151,250,1820,1842]" captionTargetBox="[340,1247,483,1795]" captionTargetId="figure@2.[340,1247,483,1795]" captionTargetPageId="2" captionText="FIGURE 1. Preserved specimens of Chlorophthalmus imperator sp. nov. collected from the Emperor Seamount Chain in the central North Pacific. (A–C), KAUM–I. 120091, 130.0 mm SL, holotype; (D), KAUM–I. 120099, 174.8 mm SL, paratype; (E), KAUM–I. 120094, 146.6 mm SL, paratype; (F), KAUM–I. 120097, 98.1 mm SL, paratype. (A, D–F), lateral views; (B), dorsal view; (C), ventral view." httpUri="https://zenodo.org/record/2624532/files/figure.png" pageId="1" pageNumber="397">Fig. 1D</figureCitation>
) of 
<taxonomicName id="6F8424AFFFDA6703E38DF8B7FA27F811" authorityName="Bonaparte" authorityYear="1840" box="[1234,1429,1901,1926]" class="Actinopterygii" family="Chlorophthalmidae" genus="Chlorophthalmus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Aulopiformes" pageId="1" pageNumber="397" phylum="Chordata" rank="genus">
<emphasis id="9AF0833EFFDA6703E38DF8B7FA27F811" box="[1234,1429,1901,1926]" italics="true" pageId="1" pageNumber="397">Chlorophthalmus</emphasis>
</taxonomicName>
, characterized by the following combination of characters: 49–51 lateral-line scales; 6 scale rows above lateral line; 3 + 19–22 = 22–25 (modally 22) gill rakers; outermost tooth patches on lower jaw with 6–14 large thorn-shaped teeth, its tips projecting in advance of lower-jaw profile (
<figureCitation id="30BF43A9FFDA6703E400F803FC19F865" box="[863,939,2009,2034]" captionStart="FIGURE 2" captionStartId="3.[151,250,964,986]" captionTargetBox="[331,1238,213,937]" captionTargetId="figure@3.[331,1238,213,937]" captionTargetPageId="3" captionText="FIGURE 2. Lower-jaw tips of Chlorophthalmus imperator sp. nov. in dorsal view. (A), KAUM–I. 120097, paratype; (B), KAUM–I. 120098, paratype; (C), KAUM–I. 120091, holotype; (D), KAUM–I. 120093, paratype; (E), KAUM–I. 120099, paratype; (F), KAUM–I. 120096, paratype (damaged)." httpUri="https://zenodo.org/record/2624534/files/figure.png" pageId="1" pageNumber="397">Fig. 2</figureCitation>
); lower-jaw symphysis with two distinct moderately-sized projections (
<figureCitation id="30BF43A9FFD96700E6B1FF4DFD87FF27" box="[494,565,151,176]" captionStart="FIGURE 2" captionStartId="3.[151,250,964,986]" captionTargetBox="[331,1238,213,937]" captionTargetId="figure@3.[331,1238,213,937]" captionTargetPageId="3" captionText="FIGURE 2. Lower-jaw tips of Chlorophthalmus imperator sp. nov. in dorsal view. (A), KAUM–I. 120097, paratype; (B), KAUM–I. 120098, paratype; (C), KAUM–I. 120091, holotype; (D), KAUM–I. 120093, paratype; (E), KAUM–I. 120099, paratype; (F), KAUM–I. 120096, paratype (damaged)." httpUri="https://zenodo.org/record/2624534/files/figure.png" pageId="2" pageNumber="398">Fig. 2</figureCitation>
); tongue without teeth; pelvic-fin origin vertically below 4th or 5th (usually 5th) dorsal-fin ray base; head length 26.7–28.9 (mean 27.7) % SL; snout length 6.9–7.7 (7.3) % SL; horizontal orbit diameter 11.9–12.9 (12.3) % SL; upper-jaw length 11.7–13.2 (12.6) % SL; maxillary depth 3.1–3.8 (3.4) % SL; pre-dorsal-fin length 34.2–36.6 (35.3) % SL; pre-pectoral-fin length 27.3–29.8 (27.9) % SL; anus to anal-fin origin length 27.3–29.9 (28.5) % SL; pectoral-fin length 21.0–24.9 (22.7) % SL.
</paragraph>
</subSubSection>
<subSubSection id="E09E0CA7FFD96706E798FE96FD39F875" lastPageId="4" lastPageNumber="400" pageId="2" pageNumber="398" type="description">
<paragraph id="A83B5F2CFFD96700E798FE96FE1FFE3B" blockId="2.[151,1437,151,429]" pageId="2" pageNumber="398">
<emphasis id="9AF0833EFFD96700E798FE96FEEBFEF2" bold="true" box="[199,345,332,357]" pageId="2" pageNumber="398">Description.</emphasis>
Data for the 
<typeStatus id="773FE18EFFD96700E55DFE96FDD6FEF2" box="[514,612,332,357]" pageId="2" pageNumber="398" type="holotype">holotype</typeStatus>
are presented first, followed by data for 
<typeStatus id="773FE18EFFD96700E307FE97FB71FEF2" box="[1112,1219,333,357]" pageId="2" pageNumber="398" type="paratype">paratypes</typeStatus>
in parentheses (if different). Characters included in the diagnosis are not repeated here. Counts and measurements (percentages of SL) are given in 
<tableCitation id="E5066A97FFD96700E60CFE4EFE15FE3B" box="[339,423,404,429]" captionStart="TABLE 1" captionStartId="7.[151,239,1220,1243]" captionText="TABLE 1. Counts and measurements, expressed as percentages of standard length, of Chlorophthalmus imperator sp. nov." pageId="2" pageNumber="398">Table 1</tableCitation>
.
</paragraph>
<caption id="FCFB0FA4FFD96700E7C8F8C6FE35F819" httpUri="https://zenodo.org/record/2624532/files/figure.png" pageId="2" pageNumber="398" startId="2.[151,250,1820,1842]" targetBox="[340,1247,483,1795]" targetPageId="2">
<paragraph id="A83B5F2CFFD96700E7C8F8C6FE35F819" blockId="2.[151,1436,1820,1934]" pageId="2" pageNumber="398">
<emphasis id="9AF0833EFFD96700E7C8F8C6FEA6F8A5" bold="true" box="[151,276,1820,1842]" pageId="2" pageNumber="398">FIGURE 1.</emphasis>
Preserved specimens of 
<taxonomicName id="6F8424AFFFD96700E54CF8C6FC9DF8A5" authority="Fujiwara &amp; Wada &amp; Motomura, 2019" authorityName="Fujiwara &amp; Wada &amp; Motomura" authorityYear="2019" box="[531,815,1820,1842]" class="Actinopterygii" family="Chlorophthalmidae" genus="Chlorophthalmus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Aulopiformes" pageId="2" pageNumber="398" phylum="Chordata" rank="species" species="imperator" status="sp. nov.">
<emphasis id="9AF0833EFFD96700E54CF8C6FC9DF8A5" box="[531,815,1820,1842]" italics="true" pageId="2" pageNumber="398">Chlorophthalmus imperator</emphasis>
</taxonomicName>
<emphasis id="9AF0833EFFD96700E468F8C6FC3BF8A5" bold="true" box="[823,905,1820,1842]" pageId="2" pageNumber="398">
<taxonomicNameLabel id="81C33E45FFD96700E468F8C6FC3BF8A5" box="[823,905,1820,1842]" pageId="2" pageNumber="398" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
collected from the Emperor Seamount Chain in the central North Pacific. (A–C), KAUM–I. 120091, 130.0 mm SL, holotype; (D), KAUM–I. 120099, 174.8 mm SL, paratype; (E), KAUM–I. 120094, 146.6 mm SL, paratype; (F), KAUM–I. 120097, 98.1 mm SL, paratype. (A, D–F), lateral views; (B), dorsal view; (C), ventral view.
</paragraph>
</caption>
<caption id="FCFB0FA4FFD86701E7C8FC1EFD0DFB8F" httpUri="https://zenodo.org/record/2624534/files/figure.png" pageId="3" pageNumber="399" startId="3.[151,250,964,986]" targetBox="[331,1238,213,937]" targetPageId="3">
<paragraph id="A83B5F2CFFD86701E7C8FC1EFD0DFB8F" blockId="3.[151,1436,964,1048]" pageId="3" pageNumber="399">
<emphasis id="9AF0833EFFD86701E7C8FC1EFEA4FC4D" bold="true" box="[151,278,964,986]" pageId="3" pageNumber="399">FIGURE 2.</emphasis>
Lower-jaw tips of 
<taxonomicName id="6F8424AFFFD86701E6BCFC1EFCB3FC4D" authority="Fujiwara &amp; Wada &amp; Motomura, 2019" authorityName="Fujiwara &amp; Wada &amp; Motomura" authorityYear="2019" box="[483,769,964,986]" class="Actinopterygii" family="Chlorophthalmidae" genus="Chlorophthalmus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Aulopiformes" pageId="3" pageNumber="399" phylum="Chordata" rank="species" species="imperator" status="sp. nov.">
<emphasis id="9AF0833EFFD86701E6BCFC1EFCB3FC4D" box="[483,769,964,986]" italics="true" pageId="3" pageNumber="399">Chlorophthalmus imperator</emphasis>
</taxonomicName>
<emphasis id="9AF0833EFFD86701E454FC1EFCEDFC4D" bold="true" box="[779,863,964,986]" pageId="3" pageNumber="399">
<taxonomicNameLabel id="81C33E45FFD86701E454FC1EFCEDFC4D" box="[779,863,964,986]" pageId="3" pageNumber="399" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
in dorsal view. (A), KAUM–I. 120097, paratype; (B), KAUM–I. 120098, paratype; (C), KAUM–I. 120091, holotype; (D), KAUM–I. 120093, paratype; (E), KAUM–I. 120099, paratype; (F), KAUM–I. 120096, paratype (damaged).
</paragraph>
</caption>
<paragraph id="A83B5F2CFFD86701E798FB9EFC08FACF" blockId="3.[151,1437,1092,2017]" pageId="3" pageNumber="399">Body slender, cylindrical, compressed at caudal peduncle; maximum depth at dorsal-fin origin. Snout pointed in lateral view, semicircular in dorsal view, its profile straight (weakly concave). Dorsal surface of snout tip with two small bluntly pointed spines. Anterior and posterior nostrils slit-like and more-or-less triangular (circular), respectively, latter much larger; closely positioned just forward eye, slightly below horizontal level of snout tip. Eye very large with diameter greater than snout length, upper profile included in head profile in lateral view. Interorbital region slightly concave in midline; least interorbital very narrow, width less than pelvic-fin base length. Preopercle rounded. Upper portion of opercle bluntly pointed, lower portion rounded. Anus very close to pelvic-fin base, hidden by pelvic fin when depressed. Perianal light organ present.</paragraph>
<paragraph id="A83B5F2CFFD86701E798FABEFC04F956" blockId="3.[151,1437,1092,2017]" pageId="3" pageNumber="399">
Mouth terminal. Upper jaw long, posterior end of maxillary somewhat rounded, reaching to between vertical through anterior margin and center of eye; with ca. 1 to 3 rows of villiform teeth, tooth band and size, respectively, becoming wider and smaller posteriorly. Anterior tip of lower jaw extending beyond that of upper jaw, with two tooth patches including ca. 4 (5) irregular rows of thoron shaped teeth, its size becoming large anteriorly, its tips strongly curved posteriorly (4 of 
<specimenCount id="BE8294A5FFD86701E574FA2FFD09F99A" box="[555,699,1524,1549]" pageId="3" pageNumber="399" type="generic">9 specimens</specimenCount>
variously damaged, but teeth fragments confirmed: 
<figureCitation id="30BF43A9FFD86701E26CFA2EFA3CF99A" box="[1331,1422,1524,1549]" captionStart="FIGURE 2" captionStartId="3.[151,250,964,986]" captionTargetBox="[331,1238,213,937]" captionTargetId="figure@3.[331,1238,213,937]" captionTargetPageId="3" captionText="FIGURE 2. Lower-jaw tips of Chlorophthalmus imperator sp. nov. in dorsal view. (A), KAUM–I. 120097, paratype; (B), KAUM–I. 120098, paratype; (C), KAUM–I. 120091, holotype; (D), KAUM–I. 120093, paratype; (E), KAUM–I. 120099, paratype; (F), KAUM–I. 120096, paratype (damaged)." httpUri="https://zenodo.org/record/2624534/files/figure.png" pageId="3" pageNumber="399">Fig. 2F</figureCitation>
); posteriorly 1 or 2 rows more-or-less of relatively large (larger than upper-jaw teeth), slightly recurved pointed villiform teeth; jaw symphyses without villiform teeth. Vomer with 1 (1–4) teeth on anterior margin, thereafter margin (sides of vomer) with 6–17 (usually 6 or 8) sharp, moderately-sized teeth (some lost in 
<typeStatus id="773FE18EFFD86701E39CF9BAFA98F9EE" box="[1219,1322,1632,1657]" pageId="3" pageNumber="399" type="holotype">holotype</typeStatus>
). Anterior half of palatine with minute pointed teeth in ca. 2–4 rows, tapering to single row posteriorly. Two pores just behind projections of lower-jaw symphysis. Gill rakers long, with pointed tips.
</paragraph>
<paragraph id="A83B5F2CFFD86706E798F916FF7AFF42" blockId="3.[151,1437,1092,2017]" lastBlockId="4.[151,1437,151,429]" lastPageId="4" lastPageNumber="400" pageId="3" pageNumber="399">Dorsal fin triangular, positioned anteriorly, its origin anterior to vertical through pelvic-fin origin; 3rd (rarely 2nd) dorsal-fin ray longest; 1st and 2nd dorsal-fin rays unbranched, other rays with 2 or 3 branches. Pectoral fin base low, just behind pointed portion of opercle; uppermost pectoral-fin ray unbranched; tip of fin pointed, extending to between vertical through last dorsal-fin ray base and tip of depressed pelvic fin. Pelvic fin short, its tip pointed; 2nd and 3rd rays longest. Anal fin short, its base shorter than longest anal-fin ray length; 4th (3th) ray longest; 1st to 3rd (4th) rays unbranched, other rays with 2 or 3 branches. Adipose fin small, slightly behind vertical through anal-fin origin. Caudal fin forked, tips pointed. Body covered with deciduous cycloid scales, scale pockets distinct; scale margins weakly serrated. Anterior extent of pre-dorsal scales slightly forward of posterior margin of eyes. Head without scales, except for cheek; scales on isthmus and thorax smaller. Scales absent on all fins.</paragraph>
<paragraph id="A83B5F2CFFDF6706E798FF05FB01FE3A" blockId="4.[151,1437,151,429]" pageId="4" pageNumber="400">
<emphasis id="9AF0833EFFDF6706E798FF05FE36FF6F" box="[199,388,223,248]" italics="true" pageId="4" pageNumber="400">Fresh coloration</emphasis>
(
<figureCitation id="30BF43A9FFDF6706E6CCFF05FE6AFF6F" box="[403,472,223,248]" captionStart="FIGURE 3" captionStartId="4.[151,250,1793,1815]" captionTargetBox="[340,1247,481,1732]" captionTargetId="figure@4.[340,1247,481,1732]" captionTargetPageId="4" captionText="FIGURE 3. Fresh specimens of Chlorophthalmus imperator sp. nov. (A), KAUM–I. 120091, 130.0 mm SL, holotype; (B), KAUM–I. 120095, 122.1 mm SL, paratype; (C), KAUM–I. 120098, 123.6 mm SL, paratype; (D), KAUM–I. 120094, 146.6 mm SL, paratype. Photos by SNFR." httpUri="https://zenodo.org/record/2624536/files/figure.png" pageId="4" pageNumber="400">Fig. 3</figureCitation>
): Body generally faint gray. Anterodorsal surface (between posterior margins of orbit and dorsal-fin base) black. Dorsal surface (upper lateral line) with black diagonal lines along scale margins. Tip of snout black. Branchial cavity and opercular margin black. Cheek and thorax with many black melanophores. Dorsal fin blackish-gray, anteriormost 3–4 rays somewhat darker. Anal and adipose fins translucent to light gray, bases black. Pectoral fin translucent to light gray, upper half part slightly darker pigmentation. Pelvic fin generally translucent to light gray, inner 4–5 rays black, fin tip blackish. Caudal fin gray, fin base black.
</paragraph>
<caption id="FCFB0FA4FFDF6706E7C8F8DBFDB6F8C3" httpUri="https://zenodo.org/record/2624536/files/figure.png" pageId="4" pageNumber="400" startId="4.[151,250,1793,1815]" targetBox="[340,1247,481,1732]" targetPageId="4">
<paragraph id="A83B5F2CFFDF6706E7C8F8DBFDB6F8C3" blockId="4.[151,1436,1793,1876]" pageId="4" pageNumber="400">
<emphasis id="9AF0833EFFDF6706E7C8F8DBFEA7F880" bold="true" box="[151,277,1793,1815]" pageId="4" pageNumber="400">FIGURE 3.</emphasis>
Fresh specimens of 
<taxonomicName id="6F8424AFFFDF6706E6B2F8DBFCB9F880" authority="Fujiwara &amp; Wada &amp; Motomura, 2019" authorityName="Fujiwara &amp; Wada &amp; Motomura" authorityYear="2019" box="[493,779,1793,1815]" class="Actinopterygii" family="Chlorophthalmidae" genus="Chlorophthalmus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Aulopiformes" pageId="4" pageNumber="400" phylum="Chordata" rank="species" species="imperator" status="sp. nov.">
<emphasis id="9AF0833EFFDF6706E6B2F8DBFCB9F880" box="[493,779,1793,1815]" italics="true" pageId="4" pageNumber="400">Chlorophthalmus imperator</emphasis>
</taxonomicName>
<emphasis id="9AF0833EFFDF6706E44BF8DBFCD5F880" bold="true" box="[788,871,1793,1815]" pageId="4" pageNumber="400">
<taxonomicNameLabel id="81C33E45FFDF6706E44BF8DBFCD5F880" box="[788,871,1793,1815]" pageId="4" pageNumber="400" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
(A), KAUM–I. 120091, 130.0 mm SL, holotype; (B), KAUM–I. 120095, 122.1 mm SL, paratype; (C), KAUM–I. 120098, 123.6 mm SL, paratype; (D), KAUM–I. 120094, 146.6 mm SL, paratype. Photos by SNFR.
</paragraph>
</caption>
<paragraph id="A83B5F2CFFDF6706E798F85BFD39F875" blockId="4.[151,1436,1921,2018]" pageId="4" pageNumber="400">
<emphasis id="9AF0833EFFDF6706E798F85BFE07F80D" box="[199,437,1921,1946]" italics="true" pageId="4" pageNumber="400">Preserved coloration</emphasis>
: Similar to fresh coloration but body generally light yellow to brown. Oral cavity and tongue yellowish [upper and posterior surface of oral cavity with black pigmentation in largest specimen (
<quantity id="6F7CF2C9FFDF6706E203F87CFF71F876" metricMagnitude="-1" metricUnit="m" metricValue="1.748" pageId="4" pageNumber="400" unit="mm" value="174.8">174.8 mm</quantity>
SL)]. Anal region and light organ black.
</paragraph>
</subSubSection>
<subSubSection id="E09E0CA7FFDE6707E798FF4DFCB2FF43" pageId="5" pageNumber="401" type="distribution">
<paragraph id="A83B5F2CFFDE6707E798FF4DFA2EFF27" blockId="5.[151,1436,151,213]" box="[199,1436,151,176]" pageId="5" pageNumber="401">
<emphasis id="9AF0833EFFDE6707E798FF4DFED2FF27" bold="true" box="[199,352,151,176]" pageId="5" pageNumber="401">Distribution.</emphasis>
Currently known only from the Emperor Seamount Chain in the central North Pacific (
<figureCitation id="30BF43A9FFDE6707E217FF4DFA3DFF27" box="[1352,1423,151,176]" captionStart="FIGURE 4" captionStartId="5.[151,250,1070,1092]" captionTargetBox="[376,1190,301,1043]" captionTargetId="figure@5.[340,1247,265,1048]" captionTargetPageId="5" captionText="FIGURE 4. Distribution of Chlorophthalmus imperator sp. nov. (red squares), arrowhead indicates type locality." httpUri="https://zenodo.org/record/2624538/files/figure.png" pageId="5" pageNumber="401">Fig. 4</figureCitation>
).
</paragraph>
<paragraph id="A83B5F2CFFDE6707E7C8FF66FCB2FF43" blockId="5.[151,1436,151,213]" box="[151,768,188,213]" pageId="5" pageNumber="401">
The specimens were collected in depths of 
<quantity id="6F7CF2C9FFDE6707E525FF66FD4EFF43" box="[634,764,188,213]" metricMagnitude="2" metricUnit="m" metricValue="4.77" metricValueMax="6.05" metricValueMin="3.4899999999999998" pageId="5" pageNumber="401" unit="m" value="477.0" valueMax="605.0" valueMin="349.0">
349– 
<specimenCount id="BE8294A5FFDE6707E5EDFF66FD4EFF43" box="[690,764,188,213]" pageId="5" pageNumber="401" type="generic">605 m</specimenCount>
</quantity>
.
</paragraph>
</subSubSection>
<caption id="FCFB0FA4FFDE6707E7C8FBF4FAA1FBD3" box="[151,1299,1070,1092]" httpUri="https://zenodo.org/record/2624538/files/figure.png" pageId="5" pageNumber="401" startId="5.[151,250,1070,1092]" targetBox="[376,1190,301,1043]" targetPageId="5">
<paragraph id="A83B5F2CFFDE6707E7C8FBF4FAA1FBD3" blockId="5.[151,1299,1070,1092]" box="[151,1299,1070,1092]" pageId="5" pageNumber="401">
<emphasis id="9AF0833EFFDE6707E7C8FBF4FEA0FBD3" bold="true" box="[151,274,1070,1092]" pageId="5" pageNumber="401">FIGURE 4.</emphasis>
Distribution of 
<taxonomicName id="6F8424AFFFDE6707E6EAFBF4FD63FBD3" authority="Fujiwara &amp; Wada &amp; Motomura, 2019" authorityName="Fujiwara &amp; Wada &amp; Motomura" authorityYear="2019" box="[437,721,1070,1092]" class="Actinopterygii" family="Chlorophthalmidae" genus="Chlorophthalmus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Aulopiformes" pageId="5" pageNumber="401" phylum="Chordata" rank="species" species="imperator" status="sp. nov.">
<emphasis id="9AF0833EFFDE6707E6EAFBF4FD63FBD3" box="[437,721,1070,1092]" italics="true" pageId="5" pageNumber="401">Chlorophthalmus imperator</emphasis>
</taxonomicName>
<emphasis id="9AF0833EFFDE6707E588FBF4FC9AFBD3" bold="true" box="[727,808,1070,1092]" pageId="5" pageNumber="401">
<taxonomicNameLabel id="81C33E45FFDE6707E588FBF4FC9AFBD3" box="[727,808,1070,1092]" pageId="5" pageNumber="401" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
(red squares), arrowhead indicates type locality.
</paragraph>
</caption>
<subSubSection id="E09E0CA7FFDE6707E798FBAAFDC1FB3A" pageId="5" pageNumber="401" type="etymology">
<paragraph id="A83B5F2CFFDE6707E798FBAAFDC1FB3A" blockId="5.[151,1437,1136,2029]" pageId="5" pageNumber="401">
<emphasis id="9AF0833EFFDE6707E798FBAAFEFCFB1E" bold="true" box="[199,334,1136,1161]" pageId="5" pageNumber="401">Etymology.</emphasis>
The species name 
<taxonomicName id="6F8424AFFFDE6707E57AFBA8FD26FB1E" authorityName="Fujiwara &amp; Wada &amp; Motomura" authorityYear="2019" box="[549,660,1138,1161]" class="Actinopterygii" family="Chlorophthalmidae" genus="Chlorophthalmus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Aulopiformes" pageId="5" pageNumber="401" phylum="Chordata" rank="species" species="imperator">
<emphasis id="9AF0833EFFDE6707E57AFBA8FD26FB1E" box="[549,660,1138,1161]" italics="true" pageId="5" pageNumber="401">imperator</emphasis>
</taxonomicName>
, derived from Latin, means emperor in reference to the 
<typeStatus id="773FE18EFFDE6707E24FFBABFAF2FB1E" box="[1296,1344,1137,1161]" pageId="5" pageNumber="401">type</typeStatus>
locality of the species (Emperor Seamount Chain).
</paragraph>
</subSubSection>
<subSubSection id="E09E0CA7FFDE670AE798FB66FC9AF862" lastPageId="8" lastPageNumber="404" pageId="5" pageNumber="401" type="discussion">
<paragraph id="A83B5F2CFFDE6707E798FB66FEE9FAD6" blockId="5.[151,1437,1136,2029]" pageId="5" pageNumber="401">
<emphasis id="9AF0833EFFDE6707E798FB66FE89FB42" bold="true" box="[199,315,1212,1237]" pageId="5" pageNumber="401">Remarks.</emphasis>
The number and size of thorn-shaped teeth in the outermost tooth patches on the lower jaw probably tend to increase in number and become relatively smaller with growth (
<figureCitation id="30BF43A9FFDE6707E48CFB3AFBA9FB6E" box="[979,1051,1248,1273]" captionStart="FIGURE 2" captionStartId="3.[151,250,964,986]" captionTargetBox="[331,1238,213,937]" captionTargetId="figure@3.[331,1238,213,937]" captionTargetPageId="3" captionText="FIGURE 2. Lower-jaw tips of Chlorophthalmus imperator sp. nov. in dorsal view. (A), KAUM–I. 120097, paratype; (B), KAUM–I. 120098, paratype; (C), KAUM–I. 120091, holotype; (D), KAUM–I. 120093, paratype; (E), KAUM–I. 120099, paratype; (F), KAUM–I. 120096, paratype (damaged)." httpUri="https://zenodo.org/record/2624534/files/figure.png" pageId="5" pageNumber="401">Fig. 2</figureCitation>
). Similar changes have also been noted in 
<taxonomicName id="6F8424AFFFDE6707E7A0FADFFE3DFA8A" authorityName="Gilbert &amp; Cramer" authorityYear="1897" box="[255,399,1284,1309]" class="Actinopterygii" family="Chlorophthalmidae" genus="Chlorophthalmus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Aulopiformes" pageId="5" pageNumber="401" phylum="Chordata" rank="species" species="proridens">
<emphasis id="9AF0833EFFDE6707E7A0FADFFE3DFA8A" box="[255,399,1284,1309]" italics="true" pageId="5" pageNumber="401">C. proridens</emphasis>
</taxonomicName>
(
<figureCitation id="30BF43A9FFDE6707E6FEFADEFE57FA8A" box="[417,485,1284,1309]" captionStart="FIGURE 5" captionStartId="6.[151,250,1882,1904]" captionTargetBox="[340,1247,772,1860]" captionTargetId="figure@6.[340,1247,772,1861]" captionTargetPageId="6" captionText="FIGURE 5. Dorsal view of lower-jaw tips of (A–E) Chlorophthalmus proridens, (F) C. acutifrons, (G) C. nigromarginatus and (H, I) C. albatrossis. (A, C), USNM 47715 (paratypes); (B, D), CAS-SU 4916 (paratypes); (E), USNM 47715 (holotype); (F), KAUM–I. 151740; (G), KAUM–I. 111668; (H), KAUM–I. 151705; (I), KAUM–I. 151739." httpUri="https://zenodo.org/record/2624540/files/figure.png" pageId="5" pageNumber="401">Fig. 5</figureCitation>
A–E). However, additional specimens are required to understand such ontogenetic changes in detail.
</paragraph>
<paragraph id="A83B5F2CFFDE6707E798FA96FDB6FA46" blockId="5.[151,1437,1136,2029]" pageId="5" pageNumber="401">
<bibRefCitation id="CC1522DDFFDE6707E798FA96FE6FFAF2" author="Humphreys, R. L. &amp; Tagami, D. T. &amp; Seki, M. P." box="[199,477,1356,1381]" pageId="5" pageNumber="401" pagination="283 - 327" refId="ref4494" refString="Humphreys, R. L., Tagami, D. T. &amp; Seki, M. P. (1984) Seamount fishery resources within the southern Emperor-northern Hawaiian Ridge area. In: Grigg, R. W. &amp; Tanoue, K. Y. (Eds), Proceedings of the Second Symposium on Resource Investigations in the Northwestern Hawaiian Islands. Lolume 1. University of Hawaii Sea Grant College Program, Honolulu, pp. 283 - 327." type="book chapter" year="1984">
Humphreys 
<emphasis id="9AF0833EFFDE6707E60EFA97FE31FAF2" box="[337,387,1356,1381]" italics="true" pageId="5" pageNumber="401">et al</emphasis>
. (1984)
</bibRefCitation>
recorded 
<taxonomicName id="6F8424AFFFDE6707E510FA97FD58FAF2" authorityName="Jordan &amp; Starks" authorityYear="1904" box="[591,746,1356,1381]" class="Actinopterygii" family="Chlorophthalmidae" genus="Chlorophthalmus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Aulopiformes" pageId="5" pageNumber="401" phylum="Chordata" rank="species" species="albatrossis">
<emphasis id="9AF0833EFFDE6707E510FA97FD58FAF2" box="[591,746,1356,1381]" italics="true" pageId="5" pageNumber="401">C. albatrossis</emphasis>
</taxonomicName>
<bibRefCitation id="CC1522DDFFDE6707E5ADFA96FC5AFAF2" author="Jordan, D. S. &amp; Starks, E. C." box="[754,1000,1356,1381]" pageId="5" pageNumber="401" pagination="577 - 630" refId="ref4580" refString="Jordan, D. S. &amp; Starks, E. C. (1904) List of fishes dredged by the steamer Albatross off the coast of Japan in the summer of 1900, with descriptions of new species and a review of the Japanese Macrouridae. Bulletin of the United States Fish Commission, 22 (1902), 577 - 630, pls. 1 - 8." type="journal article" year="1904">
<collectingCountry id="D0931FBCFFDE6707E5ADFA96FC8FFAF2" box="[754,829,1356,1381]" name="Jordan" pageId="5" pageNumber="401">Jordan</collectingCountry>
&amp; Starks 1904
</bibRefCitation>
from Kammu Seamount, the Emperor Seamount Chain. That record may have in fact been based on the new species, since the main distributional range of 
<taxonomicName id="6F8424AFFFDE6707E7E8FA4FFEE1FA3A" authorityName="Jordan &amp; Starks" authorityYear="1904" box="[183,339,1428,1453]" class="Actinopterygii" family="Chlorophthalmidae" genus="Chlorophthalmus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Aulopiformes" pageId="5" pageNumber="401" phylum="Chordata" rank="species" species="albatrossis">
<emphasis id="9AF0833EFFDE6707E7E8FA4FFEE1FA3A" box="[183,339,1428,1453]" italics="true" pageId="5" pageNumber="401">C. albatrossis</emphasis>
</taxonomicName>
is in the western Pacific Ocean (
<bibRefCitation id="CC1522DDFFDE6707E592FA4FFCD3FA3A" author="Paxton, J. R." box="[717,865,1429,1453]" pageId="5" pageNumber="401" pagination="591" refId="ref5090" refString="Paxton, J. R. (2000) Chlorophthalmidae. In: Randall, J. E. &amp; Lim, K. K. P. (Eds), A checklist of the fishes of the South China Sea. The Ruffles Bulletin of Zoology Supplement, 8, pp. 591." type="journal article" year="2000">Paxton 2000</bibRefCitation>
; 
<bibRefCitation id="CC1522DDFFDE6707E432FA4EFBEAFA3A" author="Nakabo, T. &amp; Kai, Y." box="[877,1112,1428,1453]" pageId="5" pageNumber="401" pagination="429 - 430" refId="ref4930" refString="Nakabo, T. &amp; Kai, Y. (2013) Chlorophthalmidae. In: Nakabo, T. (Ed), Fishes of Japan with pictorial keys to the species. 3 rd edition. Tokai University Press, Hadano, pp. 429 - 430, 1853." type="journal article" year="2013">Nakabo &amp; Kai 2013</bibRefCitation>
), but must remain equivocal due to the lack of further details.
</paragraph>
<paragraph id="A83B5F2CFFDE6704E798FA06FAC0FF6F" blockId="5.[151,1437,1136,2029]" lastBlockId="6.[151,1436,151,717]" lastPageId="6" lastPageNumber="402" pageId="5" pageNumber="401">
<emphasis id="9AF0833EFFDE6707E798FA06FEDEFA62" bold="true" box="[199,364,1500,1525]" pageId="5" pageNumber="401">Comparisons.</emphasis>
<taxonomicName id="6F8424AFFFDE6707E62BFA06FD1DFA63" authority="Fujiwara &amp; Wada &amp; Motomura, 2019" authorityName="Fujiwara &amp; Wada &amp; Motomura" authorityYear="2019" box="[372,687,1500,1525]" class="Actinopterygii" family="Chlorophthalmidae" genus="Chlorophthalmus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Aulopiformes" pageId="5" pageNumber="401" phylum="Chordata" rank="species" species="imperator" status="sp. nov.">
<emphasis id="9AF0833EFFDE6707E62BFA06FD1DFA63" box="[372,687,1500,1525]" italics="true" pageId="5" pageNumber="401">Chlorophthalmus imperator</emphasis>
</taxonomicName>
<emphasis id="9AF0833EFFDE6707E5E8FA06FCA0FA62" bold="true" box="[695,786,1500,1525]" pageId="5" pageNumber="401">
<taxonomicNameLabel id="81C33E45FFDE6707E5E8FA06FCA0FA62" box="[695,786,1500,1525]" pageId="5" pageNumber="401" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
is very similar to 
<taxonomicName id="6F8424AFFFDE6707E4BDFA07FBC2FA62" authorityName="Gilbert &amp; Cramer" authorityYear="1897" box="[994,1136,1500,1525]" class="Actinopterygii" family="Chlorophthalmidae" genus="Chlorophthalmus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Aulopiformes" pageId="5" pageNumber="401" phylum="Chordata" rank="species" species="proridens">
<emphasis id="9AF0833EFFDE6707E4BDFA07FBC2FA62" box="[994,1136,1500,1525]" italics="true" pageId="5" pageNumber="401">C. proridens</emphasis>
</taxonomicName>
(in the Hawaiian Islands), 
<taxonomicName id="6F8424AFFFDE6707E7C8F9DBFE8EF98E" authorityName="Kotlyar &amp; Parin" authorityYear="1986" box="[151,316,1536,1561]" class="Actinopterygii" family="Chlorophthalmidae" genus="Chlorophthalmus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Aulopiformes" pageId="5" pageNumber="401" phylum="Chordata" rank="species" species="ichthyandri">
<emphasis id="9AF0833EFFDE6707E7C8F9DBFE8EF98E" box="[151,316,1536,1561]" italics="true" pageId="5" pageNumber="401">C. ichthyandri</emphasis>
</taxonomicName>
(in the southeastern Pacific Ocean) and 
<taxonomicName id="6F8424AFFFDE6707E449F9DBFC70F98F" authorityName="Kobylianskii" authorityYear="2013" box="[790,962,1537,1560]" class="Actinopterygii" family="Chlorophthalmidae" genus="Chlorophthalmus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Aulopiformes" pageId="5" pageNumber="401" phylum="Chordata" rank="species" species="mascarensis">
<emphasis id="9AF0833EFFDE6707E449F9DBFC70F98F" box="[790,962,1537,1560]" italics="true" pageId="5" pageNumber="401">C. mascarensis</emphasis>
</taxonomicName>
(in the central western Indian Ocean) in having large thorn-shaped teeth in the outermost tooth patches on the lower jaw (
<figureCitation id="30BF43A9FFDE6707E311F9FEFB12F9AA" box="[1102,1184,1572,1597]" captionStart="FIGURE 2" captionStartId="3.[151,250,964,986]" captionTargetBox="[331,1238,213,937]" captionTargetId="figure@3.[331,1238,213,937]" captionTargetPageId="3" captionText="FIGURE 2. Lower-jaw tips of Chlorophthalmus imperator sp. nov. in dorsal view. (A), KAUM–I. 120097, paratype; (B), KAUM–I. 120098, paratype; (C), KAUM–I. 120091, holotype; (D), KAUM–I. 120093, paratype; (E), KAUM–I. 120099, paratype; (F), KAUM–I. 120096, paratype (damaged)." httpUri="https://zenodo.org/record/2624534/files/figure.png" pageId="5" pageNumber="401">Figs. 2</figureCitation>
, 
<figureCitation id="30BF43A9FFDE6707E3EFF9FEFB0FF9AA" box="[1200,1213,1572,1597]" captionStart="FIGURE 5" captionStartId="6.[151,250,1882,1904]" captionTargetBox="[340,1247,772,1860]" captionTargetId="figure@6.[340,1247,772,1861]" captionTargetPageId="6" captionText="FIGURE 5. Dorsal view of lower-jaw tips of (A–E) Chlorophthalmus proridens, (F) C. acutifrons, (G) C. nigromarginatus and (H, I) C. albatrossis. (A, C), USNM 47715 (paratypes); (B, D), CAS-SU 4916 (paratypes); (E), USNM 47715 (holotype); (F), KAUM–I. 151740; (G), KAUM–I. 111668; (H), KAUM–I. 151705; (I), KAUM–I. 151739." httpUri="https://zenodo.org/record/2624540/files/figure.png" pageId="5" pageNumber="401">5</figureCitation>
A–E). However, 
<taxonomicName id="6F8424AFFFDE6707E2DDF9FFFEB5F9F7" authorityName="Fujiwara &amp; Wada &amp; Motomura" authorityYear="2019" class="Actinopterygii" family="Chlorophthalmidae" genus="Chlorophthalmus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Aulopiformes" pageId="5" pageNumber="401" phylum="Chordata" rank="species" species="imperator">
<emphasis id="9AF0833EFFDE6707E2DDF9FFFEB5F9F7" italics="true" pageId="5" pageNumber="401">C. imperator</emphasis>
</taxonomicName>
differs from the latter three species as follows: head length 26.7–28.9 (mean 27.7) % SL [vs. 30.8–32.2 (31.5) % SL in 
<taxonomicName id="6F8424AFFFDE6707E61AF9B7FE60F912" authorityName="Gilbert &amp; Cramer" authorityYear="1897" box="[325,466,1644,1669]" class="Actinopterygii" family="Chlorophthalmidae" genus="Chlorophthalmus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Aulopiformes" pageId="5" pageNumber="401" phylum="Chordata" rank="species" species="proridens">
<emphasis id="9AF0833EFFDE6707E61AF9B7FE60F912" box="[325,466,1644,1669]" italics="true" pageId="5" pageNumber="401">C. proridens</emphasis>
</taxonomicName>
, 29.8–32.7% SL in 
<taxonomicName id="6F8424AFFFDE6707E5ECF9B7FCE7F912" authorityName="Kotlyar &amp; Parin" authorityYear="1986" box="[691,853,1644,1669]" class="Actinopterygii" family="Chlorophthalmidae" genus="Chlorophthalmus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Aulopiformes" pageId="5" pageNumber="401" phylum="Chordata" rank="species" species="ichthyandri">
<emphasis id="9AF0833EFFDE6707E5ECF9B7FCE7F912" box="[691,853,1644,1669]" italics="true" pageId="5" pageNumber="401">C. ichthyandri</emphasis>
</taxonomicName>
and 30.3–33.9% SL in 
<taxonomicName id="6F8424AFFFDE6707E301F9B7FABAF913" authorityName="Kobylianskii" authorityYear="2013" box="[1118,1288,1645,1668]" class="Actinopterygii" family="Chlorophthalmidae" genus="Chlorophthalmus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Aulopiformes" pageId="5" pageNumber="401" phylum="Chordata" rank="species" species="mascarensis">
<emphasis id="9AF0833EFFDE6707E301F9B7FABAF913" box="[1118,1288,1645,1668]" italics="true" pageId="5" pageNumber="401">C. mascarensis</emphasis>
</taxonomicName>
]; pre-dorsalfin length 34.2–36.6 (35.3) % SL [vs. 36.3–37.9 (37.3) % SL, 35.9–39.1% SL and 37.6–41.7% SL]; and prepectoral-fin length 27.3–29.8 (27.9) % SL [vs. 30.0–32.0 (31.4) % SL, 28.7–31.1% SL and 31.5–33.1% SL] (
<figureCitation id="30BF43A9FFDE6707E230F96EFF6EF966" captionStart="FIGURE 6" captionStartId="7.[151,250,1097,1119]" captionTargetBox="[316,1206,202,995]" captionTargetId="figure@7.[314,1221,195,997]" captionTargetPageId="7" captionText="FIGURE 6. Relationships of (A) head length, (B) snout length, (C) upper-jaw length, (D) maxillary depth, (E) horizontal orbit diamter, and (F) pre-pectoral-fin length (all as % of standard length) to standard length (mm) in Chlorophthalmus imperator sp. nov. (red squares) and C. proridens (blue circles). Arrowheads indicate holotype." httpUri="https://zenodo.org/record/2624542/files/figure.png" pageId="5" pageNumber="401">Fig. 6A, F</figureCitation>
). Although the new species most closely resembles 
<taxonomicName id="6F8424AFFFDE6707E46BF903FC71F966" authorityName="Gilbert &amp; Cramer" authorityYear="1897" box="[820,963,1752,1777]" class="Actinopterygii" family="Chlorophthalmidae" genus="Chlorophthalmus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Aulopiformes" pageId="5" pageNumber="401" phylum="Chordata" rank="species" species="proridens">
<emphasis id="9AF0833EFFDE6707E46BF903FC71F966" box="[820,963,1752,1777]" italics="true" pageId="5" pageNumber="401">C. proridens</emphasis>
</taxonomicName>
(
<figureCitation id="30BF43A9FFDE6707E48BF902FBA9F966" box="[980,1051,1752,1777]" captionStart="FIGURE 7" captionStartId="9.[151,250,1235,1257]" captionTargetBox="[340,1247,195,1213]" captionTargetId="figure@9.[340,1247,195,1213]" captionTargetPageId="9" captionText="FIGURE 7. Preserved holotype of Chlorophthalmus proridens, USNM 47715, 149.9 mm SL. (A), lateral view; (B), dorsal view; (C), ventral view." httpUri="https://zenodo.org/record/2624544/files/figure.png" pageId="5" pageNumber="401">Fig. 7</figureCitation>
), it also differs in having slightly smaller eyes with a horizontal diameter of 11.9–12.9 (12.3) % SL vs. 13.1–14.7 (14.1), a shorter snout, its length 6.9–7.7 (7.3) % SL vs. 8.0–8.8 (8.3), a shorter upper jaw with a length of 11.7–13.2 (12.6) % SL vs. 14.1–15.0 (14.5) and a narrower maxilla having a depth of 3.1–3.8 (3.4) % SL vs. 3.8–4.9 (4.2) (
<figureCitation id="30BF43A9FFDE6707E307F89EFB28F8CA" box="[1112,1178,1860,1885]" captionStart="FIGURE 6" captionStartId="7.[151,250,1097,1119]" captionTargetBox="[316,1206,202,995]" captionTargetId="figure@7.[314,1221,195,997]" captionTargetPageId="7" captionText="FIGURE 6. Relationships of (A) head length, (B) snout length, (C) upper-jaw length, (D) maxillary depth, (E) horizontal orbit diamter, and (F) pre-pectoral-fin length (all as % of standard length) to standard length (mm) in Chlorophthalmus imperator sp. nov. (red squares) and C. proridens (blue circles). Arrowheads indicate holotype." httpUri="https://zenodo.org/record/2624542/files/figure.png" pageId="5" pageNumber="401">Fig. 6</figureCitation>
B–E). The new species is also of distinct from 
<taxonomicName id="6F8424AFFFDE6707E6C7F8B3FD8BF816" authorityName="Kotlyar &amp; Parin" authorityYear="1986" box="[408,569,1896,1921]" class="Actinopterygii" family="Chlorophthalmidae" genus="Chlorophthalmus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Aulopiformes" pageId="5" pageNumber="401" phylum="Chordata" rank="species" species="ichthyandri">
<emphasis id="9AF0833EFFDE6707E6C7F8B3FD8BF816" box="[408,569,1896,1921]" italics="true" pageId="5" pageNumber="401">C. ichthyandri</emphasis>
</taxonomicName>
and 
<taxonomicName id="6F8424AFFFDE6707E531F8B3FCAAF817" authorityName="Kobylianskii" authorityYear="2013" box="[622,792,1897,1920]" class="Actinopterygii" family="Chlorophthalmidae" genus="Chlorophthalmus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Aulopiformes" pageId="5" pageNumber="401" phylum="Chordata" rank="species" species="mascarensis">
<emphasis id="9AF0833EFFDE6707E531F8B3FCAAF817" box="[622,792,1897,1920]" italics="true" pageId="5" pageNumber="401">C. mascarensis</emphasis>
</taxonomicName>
in having a greater anus to anal-fin origin length of 27.3– 29.9 (28.5) % SL vs. 23.2–28.2% SL in 
<taxonomicName id="6F8424AFFFDE6707E504F857FD4FF832" authorityName="Kotlyar &amp; Parin" authorityYear="1986" box="[603,765,1932,1957]" class="Actinopterygii" family="Chlorophthalmidae" genus="Chlorophthalmus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Aulopiformes" pageId="5" pageNumber="401" phylum="Chordata" rank="species" species="ichthyandri">
<emphasis id="9AF0833EFFDE6707E504F857FD4FF832" box="[603,765,1932,1957]" italics="true" pageId="5" pageNumber="401">C. ichthyandri</emphasis>
</taxonomicName>
and 21.7–24.9% SL in 
<taxonomicName id="6F8424AFFFDE6707E358F857FB03F833" authorityName="Kobylianskii" authorityYear="2013" box="[1031,1201,1933,1956]" class="Actinopterygii" family="Chlorophthalmidae" genus="Chlorophthalmus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Aulopiformes" pageId="5" pageNumber="401" phylum="Chordata" rank="species" species="mascarensis">
<emphasis id="9AF0833EFFDE6707E358F857FB03F833" box="[1031,1201,1933,1956]" italics="true" pageId="5" pageNumber="401">C. mascarensis</emphasis>
</taxonomicName>
and lacking teeth on the tongue. Moreover, compared with 
<taxonomicName id="6F8424AFFFDE6707E513F86BFD45F85F" authorityName="Kobylianskii" authorityYear="2013" box="[588,759,1969,1992]" class="Actinopterygii" family="Chlorophthalmidae" genus="Chlorophthalmus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Aulopiformes" pageId="5" pageNumber="401" phylum="Chordata" rank="species" species="mascarensis">
<emphasis id="9AF0833EFFDE6707E513F86BFD45F85F" box="[588,759,1969,1992]" italics="true" pageId="5" pageNumber="401">C. mascarensis</emphasis>
</taxonomicName>
, the new species has a shorter pectoral fin with a length of 21.0–24.9 (22.7) % SL vs. 25.7–29.0% SL, a shorter snout with a length of 1.6–1.8 times in its horizontal orbit diameter vs. 1.1–1.2 times, a more anteriorly positioned pelvic-fin origin located vertically below the 4th or 5th dorsal-fin ray base vs. below the 7th or 8th ray and fewer lateral-line scales, 49–51 vs. 53–55. The maximum recorded length of the new species (
<quantity id="6F7CF2C9FFDD6704E575FF3AFD2FFF60" box="[554,669,224,248]" metricMagnitude="-1" metricUnit="m" metricValue="1.748" pageId="6" pageNumber="402" unit="mm" value="174.8">174.8 mm</quantity>
SL) also exceeds that of 
<taxonomicName id="6F8424AFFFDD6704E4E7FF3AFBEBFF6F" authorityName="Kotlyar &amp; Parin" authorityYear="1986" box="[952,1113,223,248]" class="Actinopterygii" family="Chlorophthalmidae" genus="Chlorophthalmus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Aulopiformes" pageId="6" pageNumber="402" phylum="Chordata" rank="species" species="ichthyandri">
<emphasis id="9AF0833EFFDD6704E4E7FF3AFBEBFF6F" box="[952,1113,223,248]" italics="true" pageId="6" pageNumber="402">C. ichthyandri</emphasis>
</taxonomicName>
recorded to 
<quantity id="6F7CF2C9FFDD6704E3B9FF3AFAF1FF60" box="[1254,1347,224,248]" metricMagnitude="-1" metricUnit="m" metricValue="1.22" pageId="6" pageNumber="402" unit="mm" value="122.0">122 mm</quantity>
SL.
</paragraph>
<paragraph id="A83B5F2CFFDD6704E798FEDEFCE7FDF7" blockId="6.[151,1436,151,717]" pageId="6" pageNumber="402">
The outermost lower-jaw tooth patches in 
<taxonomicName id="6F8424AFFFDD6704E5C0FEDFFC82FE8B" authorityName="Fujiwara &amp; Wada &amp; Motomura" authorityYear="2019" box="[671,816,261,284]" class="Actinopterygii" family="Chlorophthalmidae" genus="Chlorophthalmus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Aulopiformes" pageId="6" pageNumber="402" phylum="Chordata" rank="species" species="imperator">
<emphasis id="9AF0833EFFDD6704E5C0FEDFFC82FE8B" box="[671,816,261,284]" italics="true" pageId="6" pageNumber="402">C. imperator</emphasis>
</taxonomicName>
extend beyond the lower-jaw profile, as they also do in 
<taxonomicName id="6F8424AFFFDD6704E7C8FEF2FE83FED7" authorityName="Jordan &amp; Starks" authorityYear="1904" box="[151,305,295,320]" class="Actinopterygii" family="Chlorophthalmidae" genus="Chlorophthalmus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Aulopiformes" pageId="6" pageNumber="402" phylum="Chordata" rank="species" species="albatrossis">
<emphasis id="9AF0833EFFDD6704E7C8FEF2FE83FED7" box="[151,305,295,320]" italics="true" pageId="6" pageNumber="402">C. albatrossis</emphasis>
</taxonomicName>
, 
<taxonomicName id="6F8424AFFFDD6704E660FEF2FE08FED7" authorityName="Kuronuma &amp; Yamaguchi" authorityYear="1941" box="[319,442,295,320]" class="Actinopterygii" family="Chlorophthalmidae" genus="Chlorophthalmus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Aulopiformes" pageId="6" pageNumber="402" phylum="Chordata" rank="species" species="borealis">
<emphasis id="9AF0833EFFDD6704E660FEF2FE08FED7" box="[319,442,295,320]" italics="true" pageId="6" pageNumber="402">C. borealis</emphasis>
</taxonomicName>
, 
<taxonomicName id="6F8424AFFFDD6704E698FEF2FDE7FED7" authorityName="Gilchrist" authorityYear="1904" box="[455,597,296,320]" class="Actinopterygii" family="Chlorophthalmidae" genus="Chlorophthalmus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Aulopiformes" pageId="6" pageNumber="402" phylum="Chordata" rank="species" species="punctatus">
<emphasis id="9AF0833EFFDD6704E698FEF2FDE7FED7" box="[455,597,296,320]" italics="true" pageId="6" pageNumber="402">C. punctatus</emphasis>
</taxonomicName>
and 
<emphasis id="9AF0833EFFDD6704E5D4FEF2FC91FED7" box="[651,803,295,320]" italics="true" pageId="6" pageNumber="402">
<taxonomicName id="6F8424AFFFDD6704E5D4FEF2FCADFED7" authorityName="Okamura &amp; Doi" authorityYear="1984" box="[651,799,295,320]" class="Actinopterygii" family="Chlorophthalmidae" genus="Chlorophthalmus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Aulopiformes" pageId="6" pageNumber="402" phylum="Chordata" rank="species" species="pectoralis">C. pectoralis</taxonomicName>
.
</emphasis>
Counts of lateral-line scales and/or scale rows above the lateral line differ between the new species and the above four species [
<emphasis id="9AF0833EFFDD6704E498FE97FC54FEF3" box="[967,998,333,356]" italics="true" pageId="6" pageNumber="402">viz</emphasis>
., 49–51 lateral-line scales and 6 scale rows above lateral line in 
<taxonomicName id="6F8424AFFFDD6704E6E8FEAAFDFAFE1F" authorityName="Fujiwara &amp; Wada &amp; Motomura" authorityYear="2019" box="[439,584,368,392]" class="Actinopterygii" family="Chlorophthalmidae" genus="Chlorophthalmus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Aulopiformes" pageId="6" pageNumber="402" phylum="Chordata" rank="species" species="imperator">
<emphasis id="9AF0833EFFDD6704E6E8FEAAFDFAFE1F" box="[439,584,368,392]" italics="true" pageId="6" pageNumber="402">C. imperator</emphasis>
</taxonomicName>
vs. 52–56 (given in error as 53– 
<quantity id="6F7CF2C9FFDD6704E4EBFEB5FC59FE1F" box="[948,1003,367,392]" metricMagnitude="-1" metricUnit="m" metricValue="4.064" pageId="6" pageNumber="402" unit="in" value="16.0">16 in</quantity>
original description) and 
<quantity id="6F7CF2C9FFDD6704E254FEB5FACEFE1F" box="[1291,1404,367,392]" metricMagnitude="-1" metricUnit="m" metricValue="2.032" metricValueMax="2.159" metricValueMin="1.905" pageId="6" pageNumber="402" unit="in" value="8.0" valueMax="8.5" valueMin="7.5">7.5–8.5 in</quantity>
<taxonomicName id="6F8424AFFFDD6704E2DDFEAAFEA3FE3A" authorityName="Jordan &amp; Starks" authorityYear="1904" class="Actinopterygii" family="Chlorophthalmidae" genus="Chlorophthalmus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Aulopiformes" pageId="6" pageNumber="402" phylum="Chordata" rank="species" species="albatrossis">
<emphasis id="9AF0833EFFDD6704E2DDFEAAFEA3FE3A" italics="true" pageId="6" pageNumber="402">C. albatrossis</emphasis>
</taxonomicName>
, 56 and unknown in 
<taxonomicName id="6F8424AFFFDD6704E6A3FE4FFD3BFE3B" authorityName="Gilchrist" authorityYear="1904" box="[508,649,405,428]" class="Actinopterygii" family="Chlorophthalmidae" genus="Chlorophthalmus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Aulopiformes" pageId="6" pageNumber="402" phylum="Chordata" rank="species" species="punctatus">
<emphasis id="9AF0833EFFDD6704E6A3FE4FFD3BFE3B" box="[508,649,405,428]" italics="true" pageId="6" pageNumber="402">C. punctatus</emphasis>
</taxonomicName>
, 51–52 and 
<quantity id="6F7CF2C9FFDD6704E44CFE4EFCE1FE3A" box="[787,851,404,429]" metricMagnitude="-1" metricUnit="m" metricValue="1.905" pageId="6" pageNumber="402" unit="in" value="7.5">7.5 in</quantity>
<taxonomicName id="6F8424AFFFDD6704E406FE4FFC58FE3A" authorityName="Okamura &amp; Doi" authorityYear="1984" box="[857,1002,404,429]" class="Actinopterygii" family="Chlorophthalmidae" genus="Chlorophthalmus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Aulopiformes" pageId="6" pageNumber="402" phylum="Chordata" rank="species" species="pectoralis">
<emphasis id="9AF0833EFFDD6704E406FE4FFC58FE3A" box="[857,1002,404,429]" italics="true" pageId="6" pageNumber="402">C. pectoralis</emphasis>
</taxonomicName>
and 50–56 (modally 54) and unknown in 
<taxonomicName id="6F8424AFFFDD6704E7E8FE62FE84FE47" authorityName="Kuronuma &amp; Yamaguchi" authorityYear="1941" box="[183,310,439,464]" class="Actinopterygii" family="Chlorophthalmidae" genus="Chlorophthalmus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Aulopiformes" pageId="6" pageNumber="402" phylum="Chordata" rank="species" species="borealis">
<emphasis id="9AF0833EFFDD6704E7E8FE62FE84FE47" box="[183,310,439,464]" italics="true" pageId="6" pageNumber="402">C. borealis</emphasis>
</taxonomicName>
]. Compared with 
<taxonomicName id="6F8424AFFFDD6704E553FE62FD1BFE47" authorityName="Jordan &amp; Starks" authorityYear="1904" box="[524,681,439,464]" class="Actinopterygii" family="Chlorophthalmidae" genus="Chlorophthalmus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Aulopiformes" pageId="6" pageNumber="402" phylum="Chordata" rank="species" species="albatrossis">
<emphasis id="9AF0833EFFDD6704E553FE62FD1BFE47" box="[524,681,439,464]" italics="true" pageId="6" pageNumber="402">C. albatrossis</emphasis>
</taxonomicName>
, 
<taxonomicName id="6F8424AFFFDD6704E5E4FE62FCFDFE47" authorityName="Okamura &amp; Doi" authorityYear="1984" box="[699,847,439,464]" class="Actinopterygii" family="Chlorophthalmidae" genus="Chlorophthalmus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Aulopiformes" pageId="6" pageNumber="402" phylum="Chordata" rank="species" species="pectoralis">
<emphasis id="9AF0833EFFDD6704E5E4FE62FCFDFE47" box="[699,847,439,464]" italics="true" pageId="6" pageNumber="402">C. pectoralis</emphasis>
</taxonomicName>
and 
<taxonomicName id="6F8424AFFFDD6704E4D3FE62FBB8FE47" authorityName="Kuronuma &amp; Yamaguchi" authorityYear="1941" box="[908,1034,439,464]" class="Actinopterygii" family="Chlorophthalmidae" genus="Chlorophthalmus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Aulopiformes" pageId="6" pageNumber="402" phylum="Chordata" rank="species" species="borealis">
<emphasis id="9AF0833EFFDD6704E4D3FE62FBB8FE47" box="[908,1034,439,464]" italics="true" pageId="6" pageNumber="402">C. borealis</emphasis>
</taxonomicName>
, the new species has larger thornshaped teeth in the outermost lower-jaw tooth patches vs. relatively small teeth (
<figureCitation id="30BF43A9FFDD6704E34AFE06FBD6FE63" box="[1045,1124,476,501]" captionStart="FIGURE 2" captionStartId="3.[151,250,964,986]" captionTargetBox="[331,1238,213,937]" captionTargetId="figure@3.[331,1238,213,937]" captionTargetPageId="3" captionText="FIGURE 2. Lower-jaw tips of Chlorophthalmus imperator sp. nov. in dorsal view. (A), KAUM–I. 120097, paratype; (B), KAUM–I. 120098, paratype; (C), KAUM–I. 120091, holotype; (D), KAUM–I. 120093, paratype; (E), KAUM–I. 120099, paratype; (F), KAUM–I. 120096, paratype (damaged)." httpUri="https://zenodo.org/record/2624534/files/figure.png" pageId="6" pageNumber="402">Figs. 2</figureCitation>
, 
<figureCitation id="30BF43A9FFDD6704E330FE06FB1BFE63" box="[1135,1193,476,500]" captionStart="FIGURE 4" captionStartId="5.[151,250,1070,1092]" captionTargetBox="[376,1190,301,1043]" captionTargetId="figure@5.[340,1247,265,1048]" captionTargetPageId="5" captionText="FIGURE 4. Distribution of Chlorophthalmus imperator sp. nov. (red squares), arrowhead indicates type locality." httpUri="https://zenodo.org/record/2624538/files/figure.png" pageId="6" pageNumber="402">4H, I</figureCitation>
) and a slightly higher gill raker count: 3 + 19–22 = 22–25 (22) vs. 3 + 18–19 = 
<quantity id="6F7CF2C9FFDD6704E460FDDAFC17FD8F" box="[831,933,511,536]" metricMagnitude="-1" metricUnit="m" metricValue="5.461" metricValueMax="5.588" metricValueMin="5.334" pageId="6" pageNumber="402" unit="in" value="21.5" valueMax="22.0" valueMin="21.0">21–22 in</quantity>
<taxonomicName id="6F8424AFFFDD6704E4F1FDDAFBFEFD8F" authorityName="Jordan &amp; Starks" authorityYear="1904" box="[942,1100,511,536]" class="Actinopterygii" family="Chlorophthalmidae" genus="Chlorophthalmus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Aulopiformes" pageId="6" pageNumber="402" phylum="Chordata" rank="species" species="albatrossis">
<emphasis id="9AF0833EFFDD6704E4F1FDDAFBFEFD8F" box="[942,1100,511,536]" italics="true" pageId="6" pageNumber="402">C. albatrossis</emphasis>
</taxonomicName>
, 2–3 + 18–19 = 
<quantity id="6F7CF2C9FFDD6704E24CFDDAFACBFD8F" box="[1299,1401,511,536]" metricMagnitude="-1" metricUnit="m" metricValue="5.334" metricValueMax="5.588" metricValueMin="5.08" pageId="6" pageNumber="402" unit="in" value="21.0" valueMax="22.0" valueMin="20.0">20–22 in</quantity>
<taxonomicName id="6F8424AFFFDD6704E2DDFDDAFEB5FDAA" authorityName="Okamura &amp; Doi" authorityYear="1984" class="Actinopterygii" family="Chlorophthalmidae" genus="Chlorophthalmus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Aulopiformes" pageId="6" pageNumber="402" phylum="Chordata" rank="species" species="pectoralis">
<emphasis id="9AF0833EFFDD6704E2DDFDDAFEB5FDAA" italics="true" pageId="6" pageNumber="402">C. pectoralis</emphasis>
</taxonomicName>
and total 17–22 (19) in 
<taxonomicName id="6F8424AFFFDD6704E54AFDFFFD23FDAA" authorityName="Kuronuma &amp; Yamaguchi" authorityYear="1941" box="[533,657,548,573]" class="Actinopterygii" family="Chlorophthalmidae" genus="Chlorophthalmus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Aulopiformes" pageId="6" pageNumber="402" phylum="Chordata" rank="species" species="borealis">
<emphasis id="9AF0833EFFDD6704E54AFDFFFD23FDAA" box="[533,657,548,573]" italics="true" pageId="6" pageNumber="402">C. borealis</emphasis>
</taxonomicName>
. The new species also has larger eyes than 
<taxonomicName id="6F8424AFFFDD6704E327FDFFFB41FDAA" authorityName="Kuronuma &amp; Yamaguchi" authorityYear="1941" box="[1144,1267,548,573]" class="Actinopterygii" family="Chlorophthalmidae" genus="Chlorophthalmus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Aulopiformes" pageId="6" pageNumber="402" phylum="Chordata" rank="species" species="borealis">
<emphasis id="9AF0833EFFDD6704E327FDFFFB41FDAA" box="[1144,1267,548,573]" italics="true" pageId="6" pageNumber="402">C. borealis</emphasis>
</taxonomicName>
, the horizontal orbit diameter 11.9–12.9 (12.3) % SL vs. 8.7–10.4 (9.6) % SL.
</paragraph>
<paragraph id="A83B5F2CFFDD6704E798FDB6FBA0FD5A" blockId="6.[151,1436,151,717]" pageId="6" pageNumber="402">
Projections of the lower-jaw symphysis also distinguish 
<taxonomicName id="6F8424AFFFDD6704E462FDB7FC7EFD13" authorityName="Fujiwara &amp; Wada &amp; Motomura" authorityYear="2019" box="[829,972,621,644]" class="Actinopterygii" family="Chlorophthalmidae" genus="Chlorophthalmus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Aulopiformes" pageId="6" pageNumber="402" phylum="Chordata" rank="species" species="imperator">
<emphasis id="9AF0833EFFDD6704E462FDB7FC7EFD13" box="[829,972,621,644]" italics="true" pageId="6" pageNumber="402">C. imperator</emphasis>
</taxonomicName>
, which has two distinct moderately-sized projections from 
<taxonomicName id="6F8424AFFFDD6704E602FD4AFE52FD3F" authorityName="Alcock" authorityYear="1894" box="[349,480,656,680]" class="Actinopterygii" family="Chlorophthalmidae" genus="Chlorophthalmus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Aulopiformes" pageId="6" pageNumber="402" phylum="Chordata" rank="species" species="corniger">
<emphasis id="9AF0833EFFDD6704E602FD4AFE52FD3F" box="[349,480,656,680]" italics="true" pageId="6" pageNumber="402">C. corniger</emphasis>
</taxonomicName>
with two very large spine-like projections, 
<taxonomicName id="6F8424AFFFDD6704E485FD4AFBC5FD3F" authorityName="Jordan &amp; Starks" authorityYear="1904" box="[986,1143,655,680]" class="Actinopterygii" family="Chlorophthalmidae" genus="Chlorophthalmus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Aulopiformes" pageId="6" pageNumber="402" phylum="Chordata" rank="species" species="albatrossis">
<emphasis id="9AF0833EFFDD6704E485FD4AFBC5FD3F" box="[986,1143,655,680]" italics="true" pageId="6" pageNumber="402">C. albatrossis</emphasis>
</taxonomicName>
without projections (
<figureCitation id="30BF43A9FFDD6704E231FD55FF63FD5B" captionStart="FIGURE 5" captionStartId="6.[151,250,1882,1904]" captionTargetBox="[340,1247,772,1860]" captionTargetId="figure@6.[340,1247,772,1861]" captionTargetPageId="6" captionText="FIGURE 5. Dorsal view of lower-jaw tips of (A–E) Chlorophthalmus proridens, (F) C. acutifrons, (G) C. nigromarginatus and (H, I) C. albatrossis. (A, C), USNM 47715 (paratypes); (B, D), CAS-SU 4916 (paratypes); (E), USNM 47715 (holotype); (F), KAUM–I. 151740; (G), KAUM–I. 111668; (H), KAUM–I. 151705; (I), KAUM–I. 151739." httpUri="https://zenodo.org/record/2624540/files/figure.png" pageId="6" pageNumber="402">Fig. 5H, I</figureCitation>
) and 
<taxonomicName id="6F8424AFFFDD6704E64FFD6FFE13FD5A" authorityName="Okamura &amp; Doi" authorityYear="1984" box="[272,417,692,717]" class="Actinopterygii" family="Chlorophthalmidae" genus="Chlorophthalmus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Aulopiformes" pageId="6" pageNumber="402" phylum="Chordata" rank="species" species="pectoralis">
<emphasis id="9AF0833EFFDD6704E64FFD6FFE13FD5A" box="[272,417,692,717]" italics="true" pageId="6" pageNumber="402">C. pectoralis</emphasis>
</taxonomicName>
, which has three distinct, moderately-sized projections.
</paragraph>
<caption id="FCFB0FA4FFDD6704E7C8F880FB87F83A" httpUri="https://zenodo.org/record/2624540/files/figure.png" pageId="6" pageNumber="402" startId="6.[151,250,1882,1904]" targetBox="[340,1247,772,1860]" targetPageId="6">
<paragraph id="A83B5F2CFFDD6704E7C8F880FB87F83A" blockId="6.[151,1435,1882,1965]" pageId="6" pageNumber="402">
<emphasis id="9AF0833EFFDD6704E7C8F880FEA0F8E7" bold="true" box="[151,274,1882,1904]" pageId="6" pageNumber="402">FIGURE 5.</emphasis>
Dorsal view of lower-jaw tips of (A–E) 
<taxonomicName id="6F8424AFFFDD6704E5F4F880FC70F8E7" authorityName="Gilbert &amp; Cramer" authorityYear="1897" box="[683,962,1882,1904]" class="Actinopterygii" family="Chlorophthalmidae" genus="Chlorophthalmus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Aulopiformes" pageId="6" pageNumber="402" phylum="Chordata" rank="species" species="proridens">
<emphasis id="9AF0833EFFDD6704E5F4F880FC70F8E7" box="[683,962,1882,1904]" italics="true" pageId="6" pageNumber="402">Chlorophthalmus proridens</emphasis>
</taxonomicName>
, (F) 
<taxonomicName id="6F8424AFFFDD6704E4ABF880FBC5F8E7" authorityName="Hiyama" authorityYear="1940" box="[1012,1143,1882,1904]" class="Actinopterygii" family="Chlorophthalmidae" genus="Chlorophthalmus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Aulopiformes" pageId="6" pageNumber="402" phylum="Chordata" rank="species" species="acutifrons">
<emphasis id="9AF0833EFFDD6704E4ABF880FBC5F8E7" box="[1012,1143,1882,1904]" italics="true" pageId="6" pageNumber="402">C. acutifrons</emphasis>
</taxonomicName>
, (G) 
<taxonomicName id="6F8424AFFFDD6704E3F3F880FAC3F8E7" authorityName="Kamohara" authorityYear="1953" box="[1196,1393,1882,1904]" class="Actinopterygii" family="Chlorophthalmidae" genus="Chlorophthalmus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Aulopiformes" pageId="6" pageNumber="402" phylum="Chordata" rank="species" species="nigromarginatus">
<emphasis id="9AF0833EFFDD6704E3F3F880FAC3F8E7" box="[1196,1393,1882,1904]" italics="true" pageId="6" pageNumber="402">C. nigromarginatus</emphasis>
</taxonomicName>
and (H, I) 
<taxonomicName id="6F8424AFFFDD6704E789F8A3FED0F819" authorityName="Jordan &amp; Starks" authorityYear="1904" box="[214,354,1912,1934]" class="Actinopterygii" family="Chlorophthalmidae" genus="Chlorophthalmus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Aulopiformes" pageId="6" pageNumber="402" phylum="Chordata" rank="species" species="albatrossis">
<emphasis id="9AF0833EFFDD6704E789F8A3FED0F819" box="[214,354,1912,1934]" italics="true" pageId="6" pageNumber="402">C. albatrossis</emphasis>
</taxonomicName>
. (A, C), USNM 47715 (paratypes); (B, D), CAS-SU 4916 (paratypes); (E), USNM 47715 (holotype); (F), KAUM–I. 151740; (G), KAUM–I. 111668; (H), KAUM–I. 151705; (I), KAUM–I. 151739.
</paragraph>
</caption>
<caption id="FCFB0FA4FFDC6705E7C8FB93FC7FFB0B" httpUri="https://zenodo.org/record/2624542/files/figure.png" pageId="7" pageNumber="403" startId="7.[151,250,1097,1119]" targetBox="[316,1206,202,995]" targetPageId="7">
<paragraph id="A83B5F2CFFDC6705E7C8FB93FC7FFB0B" blockId="7.[151,1436,1097,1180]" pageId="7" pageNumber="403">
<emphasis id="9AF0833EFFDC6705E7C8FB93FEA0FBC8" bold="true" box="[151,274,1097,1119]" pageId="7" pageNumber="403">FIGURE 6.</emphasis>
Relationships of (A) head length, (B) snout length, (C) upper-jaw length, (D) maxillary depth, (E) horizontal orbit diamter, and (F) pre-pectoral-fin length (all as % of standard length) to standard length (mm) in 
<taxonomicName id="6F8424AFFFDC6705E303FBB2FAC5FBE9" authority="Fujiwara &amp; Wada &amp; Motomura, 2019" authorityName="Fujiwara &amp; Wada &amp; Motomura" authorityYear="2019" box="[1116,1399,1128,1150]" class="Actinopterygii" family="Chlorophthalmidae" genus="Chlorophthalmus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Aulopiformes" pageId="7" pageNumber="403" phylum="Chordata" rank="species" species="imperator" status="sp. nov.">
<emphasis id="9AF0833EFFDC6705E303FBB2FAC5FBE9" box="[1116,1399,1128,1150]" italics="true" pageId="7" pageNumber="403">Chlorophthalmus imperator</emphasis>
</taxonomicName>
<emphasis id="9AF0833EFFDC6705E222FBB2FF71FB0B" bold="true" pageId="7" pageNumber="403">
<taxonomicNameLabel id="81C33E45FFDC6705E222FBB2FF71FB0B" pageId="7" pageNumber="403" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
(red squares) and 
<taxonomicName id="6F8424AFFFDC6705E621FB5DFE4CFB0B" authorityName="Gilbert &amp; Cramer" authorityYear="1897" box="[382,510,1158,1180]" class="Actinopterygii" family="Chlorophthalmidae" genus="Chlorophthalmus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Aulopiformes" pageId="7" pageNumber="403" phylum="Chordata" rank="species" species="proridens">
<emphasis id="9AF0833EFFDC6705E621FB5DFE4CFB0B" box="[382,510,1158,1180]" italics="true" pageId="7" pageNumber="403">C. proridens</emphasis>
</taxonomicName>
(blue circles). Arrowheads indicate holotype.
</paragraph>
</caption>
<caption id="FCFB0FA4FFDC6705E7C8FB1EFF74FB6C" pageId="7" pageNumber="403">
<paragraph id="A83B5F2CFFDC6705E7C8FB1EFF74FB6C" blockId="7.[151,1436,1219,1275]" pageId="7" pageNumber="403">
<emphasis id="9AF0833EFFDC6705E7C8FB1EFEB8FB4C" bold="true" box="[151,266,1220,1243]" pageId="7" pageNumber="403">TABLE 1.</emphasis>
Counts and measurements, expressed as percentages of standard length, of 
<taxonomicName id="6F8424AFFFDC6705E318FB19FAC6FB4C" authority="Fujiwara &amp; Wada &amp; Motomura, 2019" authorityName="Fujiwara &amp; Wada &amp; Motomura" authorityYear="2019" box="[1095,1396,1219,1243]" class="Actinopterygii" family="Chlorophthalmidae" genus="Chlorophthalmus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Aulopiformes" pageId="7" pageNumber="403" phylum="Chordata" rank="species" species="imperator" status="sp. nov.">
<emphasis id="9AF0833EFFDC6705E318FB19FAC6FB4C" box="[1095,1396,1219,1243]" italics="true" pageId="7" pageNumber="403">Chlorophthalmus imperator</emphasis>
</taxonomicName>
<taxonomicNameLabel id="81C33E45FFDC6705E223FB1EFF74FB6C" pageId="7" pageNumber="403" rank="species">sp. nov.</taxonomicNameLabel>
</paragraph>
</caption>
<paragraph id="A83B5F2CFFDC6705E5C7FACCFB83FA15" pageId="7" pageNumber="403">
<table id="DA84AD8CFFDC98FDE7FAFACCFB83FA15" box="[165,1073,1302,1410]" gridcols="3" gridrows="3" pageId="7" pageNumber="403">
<tr id="16B45D6EFFDC98FDE7FAFACCFB83FABB" box="[165,1073,1302,1324]" gridrow="0" pageId="7" pageNumber="403" rowspan-0="1">
<th id="55653412FFDC98FDE5C7FACCFCE1FABB" box="[664,851,1302,1324]" gridcol="1" gridrow="0" pageId="7" pageNumber="403">Holotype</th>
<th id="55653412FFDC98FDE4E0FACCFB83FABB" box="[959,1073,1302,1324]" gridcol="2" gridrow="0" pageId="7" pageNumber="403">Paratypes</th>
</tr>
<tr id="16B45D6EFFDC98FDE7FAFA99FB83FACE" box="[165,1073,1347,1369]" gridrow="1" pageId="7" pageNumber="403" rowspan-0="1">
<td id="55653412FFDC98FDE5C7FA99FCE1FACE" box="[664,851,1347,1369]" gridcol="1" gridrow="1" pageId="7" pageNumber="403">KAUM–I. 120091</td>
<td id="55653412FFDC98FDE4E0FA99FB83FACE" box="[959,1073,1347,1369]" gridcol="2" gridrow="1" pageId="7" pageNumber="403">
<emphasis id="9AF0833EFFDC6705E4E0FA9EFC7EFACE" box="[959,972,1348,1369]" italics="true" pageId="7" pageNumber="403">n</emphasis>
= 8
</td>
</tr>
<table id="DA84AD8CFFDC98FDE7FAFAB6FA95F83F" box="[165,1319,1388,1960]" gridcols="1" gridrows="14" pageId="7" pageNumber="403">
<tr id="16B45D6EFFDC98FDE7FAFAB6FB83FA15" box="[165,1073,1388,1410]" gridrow="2" pageId="7" pageNumber="403">
<th id="55653412FFDC98FDE7FAFAB6FE1FFA15" box="[165,429,1388,1410]" gridcol="0" gridrow="2" pageId="7" pageNumber="403">Standard length (mm; SL)</th>
<td id="55653412FFDC98FDE5C7FAB6FCE1FA15" box="[664,851,1388,1410]" gridcol="1" gridrow="2" pageId="7" pageNumber="403">130.0</td>
<td id="55653412FFDC98FDE4E0FAB6FB83FA15" box="[959,1073,1388,1410]" gridcol="2" gridrow="2" pageId="7" pageNumber="403">98.1–174.8</td>
</tr>
<paragraph id="A83B5F2CFFDC6705E7FAFA4CFB4DF831" pageId="7" pageNumber="403">
<emphasis id="9AF0833EFFDC6705E7FAFA4CFF40FA3B" bold="true" box="[165,242,1430,1452]" pageId="7" pageNumber="403">Counts</emphasis>
Range 
<tr id="16B45D6EFFDC98FDE7FAFA4CFA95FA3B" box="[165,1319,1430,1452]" gridrow="1" pageId="7" pageNumber="403">
<td id="55653412FFDC98FDE3BAFA4CFA95FA3B" box="[1253,1319,1430,1452]" gridcol="0" gridrow="1" pageId="7" pageNumber="403">Mode</td>
</tr>
Dorsal-fin rays 11 10–11 
<tr id="16B45D6EFFDC98FDE7FAFA65FA95FA42" box="[165,1319,1471,1493]" gridrow="2" pageId="7" pageNumber="403">
<td id="55653412FFDC98FDE3BAFA65FA95FA42" box="[1253,1319,1471,1493]" gridcol="0" gridrow="2" pageId="7" pageNumber="403">11</td>
</tr>
Anal-fin rays 8 9 
<tr id="16B45D6EFFDC98FDE7FAFA32FA95FA69" box="[165,1319,1512,1534]" gridrow="3" pageId="7" pageNumber="403">
<td id="55653412FFDC98FDE3BAFA32FA95FA69" box="[1253,1319,1512,1534]" gridcol="0" gridrow="3" pageId="7" pageNumber="403">9</td>
</tr>
Pectoral-fin rays 15 15–16 
<tr id="16B45D6EFFDC98FDE7FAF9C8FA95F9BF" box="[165,1319,1554,1576]" gridrow="4" pageId="7" pageNumber="403">
<td id="55653412FFDC98FDE3BAF9C8FA95F9BF" box="[1253,1319,1554,1576]" gridcol="0" gridrow="4" pageId="7" pageNumber="403">15</td>
</tr>
Pelvic-fin rays 9 9 
<tr id="16B45D6EFFDC98FDE7FAF9E1FA95F9C6" box="[165,1319,1595,1617]" gridrow="5" pageId="7" pageNumber="403">
<td id="55653412FFDC98FDE3BAF9E1FA95F9C6" box="[1253,1319,1595,1617]" gridcol="0" gridrow="5" pageId="7" pageNumber="403">9</td>
</tr>
Gill rakers 3 + 20 3 + 19–22 
<tr id="16B45D6EFFDC98FDE7FAF9BEFA95F9ED" box="[165,1319,1636,1658]" gridrow="6" pageId="7" pageNumber="403">
<td id="55653412FFDC98FDE3BAF9BEFA95F9ED" box="[1253,1319,1636,1658]" gridcol="0" gridrow="6" pageId="7" pageNumber="403">3 + 19</td>
</tr>
Lateral-line scales 50 49–51a 
<tr id="16B45D6EFFDC98FDE7FAF957FA95F931" box="[165,1319,1677,1702]" gridrow="7" pageId="7" pageNumber="403">
<td id="55653412FFDC98FDE3BAF957FA95F931" box="[1253,1319,1677,1702]" gridcol="0" gridrow="7" pageId="7" pageNumber="403">49–51</td>
</tr>
Scale rows above lateral line 6 6a 
<tr id="16B45D6EFFDC98FDE7FAF963FA95F945" box="[165,1319,1721,1746]" gridrow="8" pageId="7" pageNumber="403">
<td id="55653412FFDC98FDE3BAF963FA95F945" box="[1253,1319,1721,1746]" gridcol="0" gridrow="8" pageId="7" pageNumber="403">6</td>
</tr>
Scale rows below lateral line 5 5–6a 
<tr id="16B45D6EFFDC98FDE7FAF93FFA95F969" box="[165,1319,1765,1790]" gridrow="9" pageId="7" pageNumber="403">
<td id="55653412FFDC98FDE3BAF93FFA95F969" box="[1253,1319,1765,1790]" gridcol="0" gridrow="9" pageId="7" pageNumber="403">5</td>
</tr>
Scales between anus and anal-fin origin 18 18–19 
<tr id="16B45D6EFFDC98FDE7FAF8CBFA95F8B0" box="[165,1319,1809,1831]" gridrow="10" pageId="7" pageNumber="403">
<td id="55653412FFDC98FDE3BAF8CBFA95F8B0" box="[1253,1319,1809,1831]" gridcol="0" gridrow="10" pageId="7" pageNumber="403">19</td>
</tr>
Pre-adipose scales 19 18–19 
<tr id="16B45D6EFFDC98FDE7FAF8E1FA95F8C6" box="[165,1319,1851,1873]" gridrow="11" pageId="7" pageNumber="403">
<td id="55653412FFDC98FDE3BAF8E1FA95F8C6" box="[1253,1319,1851,1873]" gridcol="0" gridrow="11" pageId="7" pageNumber="403">19</td>
</tr>
Pre-dorsal scale 13 12–14b 
<tr id="16B45D6EFFDC98FDE7FAF8BEFA95F8EB" box="[165,1319,1892,1916]" gridrow="12" pageId="7" pageNumber="403">
<td id="55653412FFDC98FDE3BAF8BEFA95F8EB" box="[1253,1319,1892,1916]" gridcol="0" gridrow="12" pageId="7" pageNumber="403">13</td>
</tr>
Pre-pelvic scale 22 23–25b 
<tr id="16B45D6EFFDC98FDE7FAF84AFA95F83F" box="[165,1319,1936,1960]" gridrow="13" pageId="7" pageNumber="403">
<td id="55653412FFDC98FDE3BAF84AFA95F83F" box="[1253,1319,1936,1960]" gridcol="0" gridrow="13" pageId="7" pageNumber="403">22</td>
</tr>
</paragraph>
</table>
</table>
</paragraph>
<paragraph id="A83B5F2CFFDC6705E304F866FA23F845" blockId="7.[1115,1425,1980,2002]" box="[1115,1425,1980,2002]" pageId="7" pageNumber="403">
<tableNote id="55625EA2FFDC6705E304F866FA23F845" box="[1115,1425,1980,2002]" pageId="7" pageNumber="403" targetBox="[165,1319,1388,1960]" targetPageId="7">......continued on the next page</tableNote>
</paragraph>
<caption id="FCFB0FA4FFD3670AE7C0FF7AFE3FFF21" box="[159,397,160,182]" pageId="8" pageNumber="404">
<paragraph id="A83B5F2CFFD3670AE7C0FF7AFE3FFF21" blockId="8.[159,397,160,182]" box="[159,397,160,182]" pageId="8" pageNumber="404">
<emphasis id="9AF0833EFFD3670AE7C0FF7AFEBEFF21" bold="true" box="[159,268,160,182]" pageId="8" pageNumber="404">TABLE 1.</emphasis>
(Continued)
</paragraph>
</caption>
<paragraph id="A83B5F2CFFD3670AE5C7FF13FAA3F967" pageId="8" pageNumber="404">
<table id="DA84AD8CFFD398FDE7FAFF13FAADF967" box="[165,1311,201,1776]" gridcols="4" gridrows="38" pageId="8" pageNumber="404">
<tr id="16B45D6EFFD398FDE7FAFF13FAADFF48" box="[165,1311,201,223]" gridrow="0" pageId="8" pageNumber="404" rowspan-0="1" rowspan-3="1">
<th id="55653412FFD398FDE5C7FF13FCE1FF48" box="[664,851,201,223]" gridcol="1" gridrow="0" pageId="8" pageNumber="404">Holotype</th>
<th id="55653412FFD398FDE4E0FF13FB99FF48" box="[959,1067,201,223]" gridcol="2" gridrow="0" pageId="8" pageNumber="404">Paratypes</th>
</tr>
<tr id="16B45D6EFFD398FDE7FAFF2CFAADFE9B" box="[165,1311,246,268]" gridrow="1" pageId="8" pageNumber="404" rowspan-0="1" rowspan-3="1">
<td id="55653412FFD398FDE5C7FF2CFCE1FE9B" box="[664,851,246,268]" gridcol="1" gridrow="1" pageId="8" pageNumber="404">KAUM–I. 120091</td>
<td id="55653412FFD398FDE4E0FF2CFB99FE9B" box="[959,1067,246,268]" gridcol="2" gridrow="1" pageId="8" pageNumber="404">
<emphasis id="9AF0833EFFD3670AE4E0FF2DFC7EFE9B" box="[959,972,247,268]" italics="true" pageId="8" pageNumber="404">n</emphasis>
= 8
</td>
</tr>
<tr id="16B45D6EFFD398FDE7FAFEC5FAADFEA2" box="[165,1311,287,309]" gridrow="2" pageId="8" pageNumber="404" rowspan-1="1" rowspan-2="1">
<th id="55653412FFD398FDE7FAFEC5FE57FEA2" box="[165,485,287,309]" gridcol="0" gridrow="2" pageId="8" pageNumber="404">
<emphasis id="9AF0833EFFD3670AE7FAFEC5FEF1FEA2" bold="true" box="[165,323,287,309]" pageId="8" pageNumber="404">Measurements</emphasis>
</th>
<td id="55653412FFD398FDE3BAFEC5FAADFEA2" box="[1253,1311,287,309]" gridcol="3" gridrow="2" pageId="8" pageNumber="404">Mean</td>
</tr>
<tr id="16B45D6EFFD398FDE7FAFE92FAADFEC9" box="[165,1311,328,350]" gridrow="3" pageId="8" pageNumber="404">
<th id="55653412FFD398FDE7FAFE92FE57FEC9" box="[165,485,328,350]" gridcol="0" gridrow="3" pageId="8" pageNumber="404">Head length</th>
<td id="55653412FFD398FDE5C7FE92FCE1FEC9" box="[664,851,328,350]" gridcol="1" gridrow="3" pageId="8" pageNumber="404">28.4</td>
<td id="55653412FFD398FDE4E0FE92FB99FEC9" box="[959,1067,328,350]" gridcol="2" gridrow="3" pageId="8" pageNumber="404">26.7–28.9</td>
<td id="55653412FFD398FDE3BAFE92FAADFEC9" box="[1253,1311,328,350]" gridcol="3" gridrow="3" pageId="8" pageNumber="404">27.7</td>
</tr>
<tr id="16B45D6EFFD398FDE7FAFEA8FAADFE1F" box="[165,1311,370,392]" gridrow="4" pageId="8" pageNumber="404">
<th id="55653412FFD398FDE7FAFEA8FE57FE1F" box="[165,485,370,392]" gridcol="0" gridrow="4" pageId="8" pageNumber="404">Head depth</th>
<td id="55653412FFD398FDE5C7FEA8FCE1FE1F" box="[664,851,370,392]" gridcol="1" gridrow="4" pageId="8" pageNumber="404">13.3</td>
<td id="55653412FFD398FDE4E0FEA8FB99FE1F" box="[959,1067,370,392]" gridcol="2" gridrow="4" pageId="8" pageNumber="404">12.5–16.2</td>
<td id="55653412FFD398FDE3BAFEA8FAADFE1F" box="[1253,1311,370,392]" gridcol="3" gridrow="4" pageId="8" pageNumber="404">13.8</td>
</tr>
<tr id="16B45D6EFFD398FDE7FAFE41FAADFE26" box="[165,1311,411,433]" gridrow="5" pageId="8" pageNumber="404">
<th id="55653412FFD398FDE7FAFE41FE57FE26" box="[165,485,411,433]" gridcol="0" gridrow="5" pageId="8" pageNumber="404">Post-orbital length</th>
<td id="55653412FFD398FDE5C7FE41FCE1FE26" box="[664,851,411,433]" gridcol="1" gridrow="5" pageId="8" pageNumber="404">9.2</td>
<td id="55653412FFD398FDE4E0FE41FB99FE26" box="[959,1067,411,433]" gridcol="2" gridrow="5" pageId="8" pageNumber="404">8.0–9.8</td>
<td id="55653412FFD398FDE3BAFE41FAADFE26" box="[1253,1311,411,433]" gridcol="3" gridrow="5" pageId="8" pageNumber="404">9.0</td>
</tr>
<tr id="16B45D6EFFD398FDE7FAFE1EFAADFE4A" box="[165,1311,452,477]" gridrow="6" pageId="8" pageNumber="404">
<th id="55653412FFD398FDE7FAFE1EFE57FE4A" box="[165,485,452,477]" gridcol="0" gridrow="6" pageId="8" pageNumber="404">Body depth at dorsal-fin origin</th>
<td id="55653412FFD398FDE5C7FE1EFCE1FE4A" box="[664,851,452,477]" gridcol="1" gridrow="6" pageId="8" pageNumber="404">15.1</td>
<td id="55653412FFD398FDE4E0FE1EFB99FE4A" box="[959,1067,452,477]" gridcol="2" gridrow="6" pageId="8" pageNumber="404">13.2–17.4b</td>
<td id="55653412FFD398FDE3BAFE1EFAADFE4A" box="[1253,1311,452,477]" gridcol="3" gridrow="6" pageId="8" pageNumber="404">15.1</td>
</tr>
<tr id="16B45D6EFFD398FDE7FAFE2AFAADFD91" box="[165,1311,496,518]" gridrow="7" pageId="8" pageNumber="404">
<th id="55653412FFD398FDE7FAFE2AFE57FD91" box="[165,485,496,518]" gridcol="0" gridrow="7" pageId="8" pageNumber="404">Body depth at anal-fin origin</th>
<td id="55653412FFD398FDE5C7FE2AFCE1FD91" box="[664,851,496,518]" gridcol="1" gridrow="7" pageId="8" pageNumber="404">9.3</td>
<td id="55653412FFD398FDE4E0FE2AFB99FD91" box="[959,1067,496,518]" gridcol="2" gridrow="7" pageId="8" pageNumber="404">8.6–9.8</td>
<td id="55653412FFD398FDE3BAFE2AFAADFD91" box="[1253,1311,496,518]" gridcol="3" gridrow="7" pageId="8" pageNumber="404">9.1</td>
</tr>
<tr id="16B45D6EFFD398FDE7FAFDC0FAADFDA7" box="[165,1311,538,560]" gridrow="8" pageId="8" pageNumber="404">
<th id="55653412FFD398FDE7FAFDC0FE57FDA7" box="[165,485,538,560]" gridcol="0" gridrow="8" pageId="8" pageNumber="404">Body width</th>
<td id="55653412FFD398FDE5C7FDC0FCE1FDA7" box="[664,851,538,560]" gridcol="1" gridrow="8" pageId="8" pageNumber="404">14.3</td>
<td id="55653412FFD398FDE4E0FDC0FB99FDA7" box="[959,1067,538,560]" gridcol="2" gridrow="8" pageId="8" pageNumber="404">12.2–14.8</td>
<td id="55653412FFD398FDE3BAFDC0FAADFDA7" box="[1253,1311,538,560]" gridcol="3" gridrow="8" pageId="8" pageNumber="404">13.5</td>
</tr>
<tr id="16B45D6EFFD398FDE7FAFD99FAADFDCE" box="[165,1311,579,601]" gridrow="9" pageId="8" pageNumber="404">
<th id="55653412FFD398FDE7FAFD99FE57FDCE" box="[165,485,579,601]" gridcol="0" gridrow="9" pageId="8" pageNumber="404">Snout length</th>
<td id="55653412FFD398FDE5C7FD99FCE1FDCE" box="[664,851,579,601]" gridcol="1" gridrow="9" pageId="8" pageNumber="404">7.6</td>
<td id="55653412FFD398FDE4E0FD99FB99FDCE" box="[959,1067,579,601]" gridcol="2" gridrow="9" pageId="8" pageNumber="404">6.9–7.7</td>
<td id="55653412FFD398FDE3BAFD99FAADFDCE" box="[1253,1311,579,601]" gridcol="3" gridrow="9" pageId="8" pageNumber="404">7.3</td>
</tr>
<tr id="16B45D6EFFD398FDE7FAFDB6FAADFD15" box="[165,1311,620,642]" gridrow="10" pageId="8" pageNumber="404">
<th id="55653412FFD398FDE7FAFDB6FE57FD15" box="[165,485,620,642]" gridcol="0" gridrow="10" pageId="8" pageNumber="404">Upper-jaw length</th>
<td id="55653412FFD398FDE5C7FDB6FCE1FD15" box="[664,851,620,642]" gridcol="1" gridrow="10" pageId="8" pageNumber="404">12.7</td>
<td id="55653412FFD398FDE4E0FDB6FB99FD15" box="[959,1067,620,642]" gridcol="2" gridrow="10" pageId="8" pageNumber="404">11.7–13.2</td>
<td id="55653412FFD398FDE3BAFDB6FAADFD15" box="[1253,1311,620,642]" gridcol="3" gridrow="10" pageId="8" pageNumber="404">12.6</td>
</tr>
<tr id="16B45D6EFFD398FDE7FAFD4CFAADFD3B" box="[165,1311,662,684]" gridrow="11" pageId="8" pageNumber="404">
<th id="55653412FFD398FDE7FAFD4CFE57FD3B" box="[165,485,662,684]" gridcol="0" gridrow="11" pageId="8" pageNumber="404">Maxillary depth</th>
<td id="55653412FFD398FDE5C7FD4CFCE1FD3B" box="[664,851,662,684]" gridcol="1" gridrow="11" pageId="8" pageNumber="404">3.1</td>
<td id="55653412FFD398FDE4E0FD4CFB99FD3B" box="[959,1067,662,684]" gridcol="2" gridrow="11" pageId="8" pageNumber="404">3.1–3.8</td>
<td id="55653412FFD398FDE3BAFD4CFAADFD3B" box="[1253,1311,662,684]" gridcol="3" gridrow="11" pageId="8" pageNumber="404">3.4</td>
</tr>
<tr id="16B45D6EFFD398FDE7FAFD65FAADFD42" box="[165,1311,703,725]" gridrow="12" pageId="8" pageNumber="404">
<th id="55653412FFD398FDE7FAFD65FE57FD42" box="[165,485,703,725]" gridcol="0" gridrow="12" pageId="8" pageNumber="404">Horizontal orbit diameter</th>
<td id="55653412FFD398FDE5C7FD65FCE1FD42" box="[664,851,703,725]" gridcol="1" gridrow="12" pageId="8" pageNumber="404">12.9</td>
<td id="55653412FFD398FDE4E0FD65FB99FD42" box="[959,1067,703,725]" gridcol="2" gridrow="12" pageId="8" pageNumber="404">11.9–12.5</td>
<td id="55653412FFD398FDE3BAFD65FAADFD42" box="[1253,1311,703,725]" gridcol="3" gridrow="12" pageId="8" pageNumber="404">12.3</td>
</tr>
<tr id="16B45D6EFFD398FDE7FAFD32FAADFD69" box="[165,1311,744,766]" gridrow="13" pageId="8" pageNumber="404">
<th id="55653412FFD398FDE7FAFD32FE57FD69" box="[165,485,744,766]" gridcol="0" gridrow="13" pageId="8" pageNumber="404">Vertical orbit diameter</th>
<td id="55653412FFD398FDE5C7FD32FCE1FD69" box="[664,851,744,766]" gridcol="1" gridrow="13" pageId="8" pageNumber="404">9.3</td>
<td id="55653412FFD398FDE4E0FD32FB99FD69" box="[959,1067,744,766]" gridcol="2" gridrow="13" pageId="8" pageNumber="404">8.1–10.5</td>
<td id="55653412FFD398FDE3BAFD32FAADFD69" box="[1253,1311,744,766]" gridcol="3" gridrow="13" pageId="8" pageNumber="404">9.1</td>
</tr>
<tr id="16B45D6EFFD398FDE7FAFCC8FAADFCBF" box="[165,1311,786,808]" gridrow="14" pageId="8" pageNumber="404">
<th id="55653412FFD398FDE7FAFCC8FE57FCBF" box="[165,485,786,808]" gridcol="0" gridrow="14" pageId="8" pageNumber="404">Horizontal pupil diameter</th>
<td id="55653412FFD398FDE5C7FCC8FCE1FCBF" box="[664,851,786,808]" gridcol="1" gridrow="14" pageId="8" pageNumber="404">11.0</td>
<td id="55653412FFD398FDE4E0FCC8FB99FCBF" box="[959,1067,786,808]" gridcol="2" gridrow="14" pageId="8" pageNumber="404">9.3–10.8</td>
<td id="55653412FFD398FDE3BAFCC8FAADFCBF" box="[1253,1311,786,808]" gridcol="3" gridrow="14" pageId="8" pageNumber="404">10.3</td>
</tr>
<tr id="16B45D6EFFD398FDE7FAFCE1FAADFCC6" box="[165,1311,827,849]" gridrow="15" pageId="8" pageNumber="404">
<th id="55653412FFD398FDE7FAFCE1FE57FCC6" box="[165,485,827,849]" gridcol="0" gridrow="15" pageId="8" pageNumber="404">Anterior interorbital width</th>
<td id="55653412FFD398FDE5C7FCE1FCE1FCC6" box="[664,851,827,849]" gridcol="1" gridrow="15" pageId="8" pageNumber="404">9.8</td>
<td id="55653412FFD398FDE4E0FCE1FB99FCC6" box="[959,1067,827,849]" gridcol="2" gridrow="15" pageId="8" pageNumber="404">8.7–10.0</td>
<td id="55653412FFD398FDE3BAFCE1FAADFCC6" box="[1253,1311,827,849]" gridcol="3" gridrow="15" pageId="8" pageNumber="404">9.3</td>
</tr>
<tr id="16B45D6EFFD398FDE7FAFCBEFAADFCED" box="[165,1311,868,890]" gridrow="16" pageId="8" pageNumber="404">
<th id="55653412FFD398FDE7FAFCBEFE57FCED" box="[165,485,868,890]" gridcol="0" gridrow="16" pageId="8" pageNumber="404">Posterior interorbital width</th>
<td id="55653412FFD398FDE5C7FCBEFCE1FCED" box="[664,851,868,890]" gridcol="1" gridrow="16" pageId="8" pageNumber="404">13.1</td>
<td id="55653412FFD398FDE4E0FCBEFB99FCED" box="[959,1067,868,890]" gridcol="2" gridrow="16" pageId="8" pageNumber="404">11.0–14.4</td>
<td id="55653412FFD398FDE3BAFCBEFAADFCED" box="[1253,1311,868,890]" gridcol="3" gridrow="16" pageId="8" pageNumber="404">12.4</td>
</tr>
<tr id="16B45D6EFFD398FDE7FAFC54FAADFC33" box="[165,1311,910,932]" gridrow="17" pageId="8" pageNumber="404">
<th id="55653412FFD398FDE7FAFC54FE57FC33" box="[165,485,910,932]" gridcol="0" gridrow="17" pageId="8" pageNumber="404">Least interorbital width</th>
<td id="55653412FFD398FDE5C7FC54FCE1FC33" box="[664,851,910,932]" gridcol="1" gridrow="17" pageId="8" pageNumber="404">2.2</td>
<td id="55653412FFD398FDE4E0FC54FB99FC33" box="[959,1067,910,932]" gridcol="2" gridrow="17" pageId="8" pageNumber="404">2.0–2.9</td>
<td id="55653412FFD398FDE3BAFC54FAADFC33" box="[1253,1311,910,932]" gridcol="3" gridrow="17" pageId="8" pageNumber="404">2.2</td>
</tr>
<tr id="16B45D6EFFD398FDE7FAFC6DFAADFC5A" box="[165,1311,951,973]" gridrow="18" pageId="8" pageNumber="404">
<th id="55653412FFD398FDE7FAFC6DFE57FC5A" box="[165,485,951,973]" gridcol="0" gridrow="18" pageId="8" pageNumber="404">Caudal-peduncle length</th>
<td id="55653412FFD398FDE5C7FC6DFCE1FC5A" box="[664,851,951,973]" gridcol="1" gridrow="18" pageId="8" pageNumber="404">17.2</td>
<td id="55653412FFD398FDE4E0FC6DFB99FC5A" box="[959,1067,951,973]" gridcol="2" gridrow="18" pageId="8" pageNumber="404">15.6–18.7</td>
<td id="55653412FFD398FDE3BAFC6DFAADFC5A" box="[1253,1311,951,973]" gridcol="3" gridrow="18" pageId="8" pageNumber="404">17.4</td>
</tr>
<tr id="16B45D6EFFD398FDE7FAFC3AFAADFC61" box="[165,1311,992,1014]" gridrow="19" pageId="8" pageNumber="404">
<th id="55653412FFD398FDE7FAFC3AFE57FC61" box="[165,485,992,1014]" gridcol="0" gridrow="19" pageId="8" pageNumber="404">Caudal-peduncle depth</th>
<td id="55653412FFD398FDE5C7FC3AFCE1FC61" box="[664,851,992,1014]" gridcol="1" gridrow="19" pageId="8" pageNumber="404">5.6</td>
<td id="55653412FFD398FDE4E0FC3AFB99FC61" box="[959,1067,992,1014]" gridcol="2" gridrow="19" pageId="8" pageNumber="404">5.8–6.3</td>
<td id="55653412FFD398FDE3BAFC3AFAADFC61" box="[1253,1311,992,1014]" gridcol="3" gridrow="19" pageId="8" pageNumber="404">6.0</td>
</tr>
<tr id="16B45D6EFFD398FDE7FAFBD0FAADFBB7" box="[165,1311,1034,1056]" gridrow="20" pageId="8" pageNumber="404">
<th id="55653412FFD398FDE7FAFBD0FE57FBB7" box="[165,485,1034,1056]" gridcol="0" gridrow="20" pageId="8" pageNumber="404">Anus to anal-fin origin length</th>
<td id="55653412FFD398FDE5C7FBD0FCE1FBB7" box="[664,851,1034,1056]" gridcol="1" gridrow="20" pageId="8" pageNumber="404">29.6</td>
<td id="55653412FFD398FDE4E0FBD0FB99FBB7" box="[959,1067,1034,1056]" gridcol="2" gridrow="20" pageId="8" pageNumber="404">27.3–29.9</td>
<td id="55653412FFD398FDE3BAFBD0FAADFBB7" box="[1253,1311,1034,1056]" gridcol="3" gridrow="20" pageId="8" pageNumber="404">28.5</td>
</tr>
<tr id="16B45D6EFFD398FDE7FAFBE9FAADFBDE" box="[165,1311,1075,1097]" gridrow="21" pageId="8" pageNumber="404">
<th id="55653412FFD398FDE7FAFBE9FE57FBDE" box="[165,485,1075,1097]" gridcol="0" gridrow="21" pageId="8" pageNumber="404">Pre-adipose-fin length</th>
<td id="55653412FFD398FDE5C7FBE9FCE1FBDE" box="[664,851,1075,1097]" gridcol="1" gridrow="21" pageId="8" pageNumber="404">77.0</td>
<td id="55653412FFD398FDE4E0FBE9FB99FBDE" box="[959,1067,1075,1097]" gridcol="2" gridrow="21" pageId="8" pageNumber="404">76.5–79.2</td>
<td id="55653412FFD398FDE3BAFBE9FAADFBDE" box="[1253,1311,1075,1097]" gridcol="3" gridrow="21" pageId="8" pageNumber="404">77.8</td>
</tr>
<tr id="16B45D6EFFD398FDE7FAFB86FAADFBE2" box="[165,1311,1116,1141]" gridrow="22" pageId="8" pageNumber="404">
<th id="55653412FFD398FDE7FAFB86FE57FBE2" box="[165,485,1116,1141]" gridcol="0" gridrow="22" pageId="8" pageNumber="404">Dorsal- to adipose-fin length</th>
<td id="55653412FFD398FDE5C7FB86FCE1FBE2" box="[664,851,1116,1141]" gridcol="1" gridrow="22" pageId="8" pageNumber="404">42.3</td>
<td id="55653412FFD398FDE4E0FB86FB99FBE2" box="[959,1067,1116,1141]" gridcol="2" gridrow="22" pageId="8" pageNumber="404">42.2–45.1b</td>
<td id="55653412FFD398FDE3BAFB86FAADFBE2" box="[1253,1311,1116,1141]" gridcol="3" gridrow="22" pageId="8" pageNumber="404">43.4</td>
</tr>
<tr id="16B45D6EFFD398FDE7FAFB52FAADFB36" box="[165,1311,1160,1185]" gridrow="23" pageId="8" pageNumber="404">
<th id="55653412FFD398FDE7FAFB52FE57FB36" box="[165,485,1160,1185]" gridcol="0" gridrow="23" pageId="8" pageNumber="404">Pre-dorsal-fin length</th>
<td id="55653412FFD398FDE5C7FB52FCE1FB36" box="[664,851,1160,1185]" gridcol="1" gridrow="23" pageId="8" pageNumber="404">36.1</td>
<td id="55653412FFD398FDE4E0FB52FB99FB36" box="[959,1067,1160,1185]" gridcol="2" gridrow="23" pageId="8" pageNumber="404">34.2–36.6b</td>
<td id="55653412FFD398FDE3BAFB52FAADFB36" box="[1253,1311,1160,1185]" gridcol="3" gridrow="23" pageId="8" pageNumber="404">35.3</td>
</tr>
<tr id="16B45D6EFFD398FDE7FAFB6EFAADFB5A" box="[165,1311,1204,1229]" gridrow="24" pageId="8" pageNumber="404">
<th id="55653412FFD398FDE7FAFB6EFE57FB5A" box="[165,485,1204,1229]" gridcol="0" gridrow="24" pageId="8" pageNumber="404">Dorsal-caudal length</th>
<td id="55653412FFD398FDE5C7FB6EFCE1FB5A" box="[664,851,1204,1229]" gridcol="1" gridrow="24" pageId="8" pageNumber="404">64.6</td>
<td id="55653412FFD398FDE4E0FB6EFB99FB5A" box="[959,1067,1204,1229]" gridcol="2" gridrow="24" pageId="8" pageNumber="404">64.0–67.4b</td>
<td id="55653412FFD398FDE3BAFB6EFAADFB5A" box="[1253,1311,1204,1229]" gridcol="3" gridrow="24" pageId="8" pageNumber="404">65.6</td>
</tr>
<tr id="16B45D6EFFD398FDE7FAFB3AFAADFB6E" box="[165,1311,1248,1273]" gridrow="25" pageId="8" pageNumber="404">
<th id="55653412FFD398FDE7FAFB3AFE57FB6E" box="[165,485,1248,1273]" gridcol="0" gridrow="25" pageId="8" pageNumber="404">Dorsal-fin base length</th>
<td id="55653412FFD398FDE5C7FB3AFCE1FB6E" box="[664,851,1248,1273]" gridcol="1" gridrow="25" pageId="8" pageNumber="404">12.3</td>
<td id="55653412FFD398FDE4E0FB3AFB99FB6E" box="[959,1067,1248,1273]" gridcol="2" gridrow="25" pageId="8" pageNumber="404">11.1–13.8b</td>
<td id="55653412FFD398FDE3BAFB3AFAADFB6E" box="[1253,1311,1248,1273]" gridcol="3" gridrow="25" pageId="8" pageNumber="404">12.5</td>
</tr>
<tr id="16B45D6EFFD398FDE7FAFAD1FAADFAB3" box="[165,1311,1291,1316]" gridrow="26" pageId="8" pageNumber="404">
<th id="55653412FFD398FDE7FAFAD1FE57FAB3" box="[165,485,1291,1316]" gridcol="0" gridrow="26" pageId="8" pageNumber="404">Longest dorsal-fin ray length</th>
<td id="55653412FFD398FDE5C7FAD1FCE1FAB3" box="[664,851,1291,1316]" gridcol="1" gridrow="26" pageId="8" pageNumber="404">19.5</td>
<td id="55653412FFD398FDE4E0FAD1FB99FAB3" box="[959,1067,1291,1316]" gridcol="2" gridrow="26" pageId="8" pageNumber="404">16.4–20.2b</td>
<td id="55653412FFD398FDE3BAFAD1FAADFAB3" box="[1253,1311,1291,1316]" gridcol="3" gridrow="26" pageId="8" pageNumber="404">18.6</td>
</tr>
<tr id="16B45D6EFFD398FDE7FAFAE2FAADFAD9" box="[165,1311,1336,1358]" gridrow="27" pageId="8" pageNumber="404">
<th id="55653412FFD398FDE7FAFAE2FE57FAD9" box="[165,485,1336,1358]" gridcol="0" gridrow="27" pageId="8" pageNumber="404">Pre-anal-fin length</th>
<td id="55653412FFD398FDE5C7FAE2FCE1FAD9" box="[664,851,1336,1358]" gridcol="1" gridrow="27" pageId="8" pageNumber="404">77.8</td>
<td id="55653412FFD398FDE4E0FAE2FB99FAD9" box="[959,1067,1336,1358]" gridcol="2" gridrow="27" pageId="8" pageNumber="404">75.7–78.1</td>
<td id="55653412FFD398FDE3BAFAE2FAADFAD9" box="[1253,1311,1336,1358]" gridcol="3" gridrow="27" pageId="8" pageNumber="404">76.8</td>
</tr>
<tr id="16B45D6EFFD398FDE7FAFABBFAADFAE0" box="[165,1311,1377,1399]" gridrow="28" pageId="8" pageNumber="404">
<th id="55653412FFD398FDE7FAFABBFE57FAE0" box="[165,485,1377,1399]" gridcol="0" gridrow="28" pageId="8" pageNumber="404">Anal-caudal length</th>
<td id="55653412FFD398FDE5C7FABBFCE1FAE0" box="[664,851,1377,1399]" gridcol="1" gridrow="28" pageId="8" pageNumber="404">24.1</td>
<td id="55653412FFD398FDE4E0FABBFB99FAE0" box="[959,1067,1377,1399]" gridcol="2" gridrow="28" pageId="8" pageNumber="404">23.3–26.1</td>
<td id="55653412FFD398FDE3BAFABBFAADFAE0" box="[1253,1311,1377,1399]" gridcol="3" gridrow="28" pageId="8" pageNumber="404">24.7</td>
</tr>
<tr id="16B45D6EFFD398FDE7FAFA50FAADFA37" box="[165,1311,1418,1440]" gridrow="29" pageId="8" pageNumber="404">
<th id="55653412FFD398FDE7FAFA50FE57FA37" box="[165,485,1418,1440]" gridcol="0" gridrow="29" pageId="8" pageNumber="404">Anal-fin base length</th>
<td id="55653412FFD398FDE5C7FA50FCE1FA37" box="[664,851,1418,1440]" gridcol="1" gridrow="29" pageId="8" pageNumber="404">8.6</td>
<td id="55653412FFD398FDE4E0FA50FB99FA37" box="[959,1067,1418,1440]" gridcol="2" gridrow="29" pageId="8" pageNumber="404">7.2–8.1</td>
<td id="55653412FFD398FDE3BAFA50FAADFA37" box="[1253,1311,1418,1440]" gridcol="3" gridrow="29" pageId="8" pageNumber="404">7.6</td>
</tr>
<tr id="16B45D6EFFD398FDE7FAFA69FAADFA5B" box="[165,1311,1459,1484]" gridrow="30" pageId="8" pageNumber="404">
<th id="55653412FFD398FDE7FAFA69FE57FA5B" box="[165,485,1459,1484]" gridcol="0" gridrow="30" pageId="8" pageNumber="404">Longest anal-fin ray length</th>
<td id="55653412FFD398FDE5C7FA69FCE1FA5B" box="[664,851,1459,1484]" gridcol="1" gridrow="30" pageId="8" pageNumber="404">11.7</td>
<td id="55653412FFD398FDE4E0FA69FB99FA5B" box="[959,1067,1459,1484]" gridcol="2" gridrow="30" pageId="8" pageNumber="404">10.1–12.3b</td>
<td id="55653412FFD398FDE3BAFA69FAADFA5B" box="[1253,1311,1459,1484]" gridcol="3" gridrow="30" pageId="8" pageNumber="404">11.3</td>
</tr>
<tr id="16B45D6EFFD398FDE7FAFA05FAADFA62" box="[165,1311,1503,1525]" gridrow="31" pageId="8" pageNumber="404">
<th id="55653412FFD398FDE7FAFA05FE57FA62" box="[165,485,1503,1525]" gridcol="0" gridrow="31" pageId="8" pageNumber="404">Pre-pectoral-fin length</th>
<td id="55653412FFD398FDE5C7FA05FCE1FA62" box="[664,851,1503,1525]" gridcol="1" gridrow="31" pageId="8" pageNumber="404">28.3</td>
<td id="55653412FFD398FDE4E0FA05FB99FA62" box="[959,1067,1503,1525]" gridcol="2" gridrow="31" pageId="8" pageNumber="404">27.3–29.8</td>
<td id="55653412FFD398FDE3BAFA05FAADFA62" box="[1253,1311,1503,1525]" gridcol="3" gridrow="31" pageId="8" pageNumber="404">27.9</td>
</tr>
<tr id="16B45D6EFFD398FDE7FAF9D3FAADF9B5" box="[165,1311,1545,1570]" gridrow="32" pageId="8" pageNumber="404">
<th id="55653412FFD398FDE7FAF9D3FE57F9B5" box="[165,485,1545,1570]" gridcol="0" gridrow="32" pageId="8" pageNumber="404">Pectoral-fin length</th>
<td id="55653412FFD398FDE5C7F9D3FCE1F9B5" box="[664,851,1545,1570]" gridcol="1" gridrow="32" pageId="8" pageNumber="404">24.1</td>
<td id="55653412FFD398FDE4E0F9D3FB99F9B5" box="[959,1067,1545,1570]" gridcol="2" gridrow="32" pageId="8" pageNumber="404">21.0–24.9b</td>
<td id="55653412FFD398FDE3BAF9D3FAADF9B5" box="[1253,1311,1545,1570]" gridcol="3" gridrow="32" pageId="8" pageNumber="404">22.7</td>
</tr>
<tr id="16B45D6EFFD398FDE7FAF9EFFAADF9DC" box="[165,1311,1589,1611]" gridrow="33" pageId="8" pageNumber="404">
<th id="55653412FFD398FDE7FAF9EFFE57F9DC" box="[165,485,1589,1611]" gridcol="0" gridrow="33" pageId="8" pageNumber="404">Pre-pelvic-fin length</th>
<td id="55653412FFD398FDE5C7F9EFFCE1F9DC" box="[664,851,1589,1611]" gridcol="1" gridrow="33" pageId="8" pageNumber="404">41.0</td>
<td id="55653412FFD398FDE4E0F9EFFB99F9DC" box="[959,1067,1589,1611]" gridcol="2" gridrow="33" pageId="8" pageNumber="404">38.0–43.5</td>
<td id="55653412FFD398FDE3BAF9EFFAADF9DC" box="[1253,1311,1589,1611]" gridcol="3" gridrow="33" pageId="8" pageNumber="404">40.4</td>
</tr>
<tr id="16B45D6EFFD398FDE7FAF984FAADF9E3" box="[165,1311,1630,1652]" gridrow="34" pageId="8" pageNumber="404">
<th id="55653412FFD398FDE7FAF984FE57F9E3" box="[165,485,1630,1652]" gridcol="0" gridrow="34" pageId="8" pageNumber="404">Pelvic-fin length</th>
<td id="55653412FFD398FDE5C7F984FCE1F9E3" box="[664,851,1630,1652]" gridcol="1" gridrow="34" pageId="8" pageNumber="404">18.5</td>
<td id="55653412FFD398FDE4E0F984FB99F9E3" box="[959,1067,1630,1652]" gridcol="2" gridrow="34" pageId="8" pageNumber="404">16.5–18.1</td>
<td id="55653412FFD398FDE3BAF984FAADF9E3" box="[1253,1311,1630,1652]" gridcol="3" gridrow="34" pageId="8" pageNumber="404">17.6</td>
</tr>
<tr id="16B45D6EFFD398FDE7FAF95DFAADF90A" box="[165,1311,1671,1693]" gridrow="35" pageId="8" pageNumber="404">
<th id="55653412FFD398FDE7FAF95DFE57F90A" box="[165,485,1671,1693]" gridcol="0" gridrow="35" pageId="8" pageNumber="404">Pelvic-fin base length</th>
<td id="55653412FFD398FDE5C7F95DFCE1F90A" box="[664,851,1671,1693]" gridcol="1" gridrow="35" pageId="8" pageNumber="404">3.8</td>
<td id="55653412FFD398FDE4E0F95DFB99F90A" box="[959,1067,1671,1693]" gridcol="2" gridrow="35" pageId="8" pageNumber="404">2.9–3.6</td>
<td id="55653412FFD398FDE3BAF95DFAADF90A" box="[1253,1311,1671,1693]" gridcol="3" gridrow="35" pageId="8" pageNumber="404">3.3</td>
</tr>
<tr id="16B45D6EFFD398FDE7FAF96BFAADF950" box="[165,1311,1713,1735]" gridrow="36" pageId="8" pageNumber="404">
<th id="55653412FFD398FDE7FAF96BFE57F950" box="[165,485,1713,1735]" gridcol="0" gridrow="36" pageId="8" pageNumber="404">Pectoral- to pelvic-fin length</th>
<td id="55653412FFD398FDE5C7F96BFCE1F950" box="[664,851,1713,1735]" gridcol="1" gridrow="36" pageId="8" pageNumber="404">14.2</td>
<td id="55653412FFD398FDE4E0F96BFB99F950" box="[959,1067,1713,1735]" gridcol="2" gridrow="36" pageId="8" pageNumber="404">13.0–15.9</td>
<td id="55653412FFD398FDE3BAF96BFAADF950" box="[1253,1311,1713,1735]" gridcol="3" gridrow="36" pageId="8" pageNumber="404">14.7</td>
</tr>
<tr id="16B45D6EFFD398FDE7FAF900FAADF967" box="[165,1311,1754,1776]" gridrow="37" pageId="8" pageNumber="404">
<th id="55653412FFD398FDE7FAF900FE57F967" box="[165,485,1754,1776]" gridcol="0" gridrow="37" pageId="8" pageNumber="404">Pelvic- to anal-fin length</th>
<td id="55653412FFD398FDE5C7F900FCE1F967" box="[664,851,1754,1776]" gridcol="1" gridrow="37" pageId="8" pageNumber="404">39.3</td>
<td id="55653412FFD398FDE4E0F900FB99F967" box="[959,1067,1754,1776]" gridcol="2" gridrow="37" pageId="8" pageNumber="404">36.8–40.2</td>
<td id="55653412FFD398FDE3BAF900FAADF967" box="[1253,1311,1754,1776]" gridcol="3" gridrow="37" pageId="8" pageNumber="404">38.1</td>
</tr>
</table>
</paragraph>
<paragraph id="A83B5F2CFFD3670AE7C8F8D6FCABF8B2" blockId="8.[151,793,1804,1829]" box="[151,793,1804,1829]" pageId="8" pageNumber="404">
<tableNote id="55625EA2FFD3670AE7C8F8D6FCABF8B2" box="[151,793,1804,1829]" pageId="8" pageNumber="404" targetBox="[165,1311,201,1776]" targetPageId="8">
<superScript id="5FF1F264FFD3670AE7C8F8D6FF2FF88F" attach="left" box="[151,157,1804,1816]" fontSize="5" pageId="8" pageNumber="404">a</superScript>
4 specimens counted on right side; 
<superScript id="5FF1F264FFD3670AE543F8D6FD91F88F" attach="right" box="[540,547,1804,1816]" fontSize="5" pageId="8" pageNumber="404">b</superScript>
based on 7 specimens.
</tableNote>
</paragraph>
<paragraph id="A83B5F2CFFD3670AE798F896FC9AF862" blockId="8.[151,1437,1868,2038]" pageId="8" pageNumber="404">
<taxonomicName id="6F8424AFFFD3670AE798F896FDB6F8F2" authorityName="Gunther" authorityYear="1887" box="[199,516,1868,1893]" class="Actinopterygii" family="Chlorophthalmidae" genus="Chlorophthalmus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Aulopiformes" pageId="8" pageNumber="404" phylum="Chordata" rank="species" species="productus">
<emphasis id="9AF0833EFFD3670AE798F896FDB6F8F2" box="[199,516,1868,1893]" italics="true" pageId="8" pageNumber="404">Chlorophthalmus productus</emphasis>
</taxonomicName>
differs from the new species in lacking enlarge teeth on the periphery of the dentary (M. Gomon, personal communication) and in having 58 lateral-line scales and 2 + 15 gill rakers. The remaining nine species 
<taxonomicName id="6F8424AFFFD3670AE6FAF84CFD90F83A" authorityName="Bonaparte" authorityYear="1840" box="[421,546,1942,1965]" class="Actinopterygii" family="Chlorophthalmidae" genus="Chlorophthalmus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Aulopiformes" pageId="8" pageNumber="404" phylum="Chordata" rank="species" species="agassizi">
<emphasis id="9AF0833EFFD3670AE6FAF84CFD90F83A" box="[421,546,1942,1965]" italics="true" pageId="8" pageNumber="404">C. agassizi</emphasis>
</taxonomicName>
, 
<taxonomicName id="6F8424AFFFD3670AE56CF84CFD7CF839" baseAuthorityName="Goode" baseAuthorityYear="1881" box="[563,718,1941,1966]" class="Actinopterygii" family="Chlorophthalmidae" genus="Chlorophthalmus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Aulopiformes" pageId="8" pageNumber="404" phylum="Chordata" rank="species" species="chalybeius">
<emphasis id="9AF0833EFFD3670AE56CF84CFD7CF839" box="[563,718,1941,1966]" italics="true" pageId="8" pageNumber="404">C. chalybeius</emphasis>
</taxonomicName>
, 
<taxonomicName id="6F8424AFFFD3670AE581F84CFCF7F83A" authorityName="Garman" authorityYear="1899" box="[734,837,1942,1965]" class="Actinopterygii" family="Chlorophthalmidae" genus="Chlorophthalmus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Aulopiformes" pageId="8" pageNumber="404" phylum="Chordata" rank="species" species="mento">
<emphasis id="9AF0833EFFD3670AE581F84CFCF7F83A" box="[734,837,1942,1965]" italics="true" pageId="8" pageNumber="404">C. mento</emphasis>
</taxonomicName>
, 
<taxonomicName id="6F8424AFFFD3670AE409F84CFC51F83A" authorityName="Hiyama" authorityYear="1940" box="[854,995,1941,1965]" class="Actinopterygii" family="Chlorophthalmidae" genus="Chlorophthalmus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Aulopiformes" pageId="8" pageNumber="404" phylum="Chordata" rank="species" species="acutifrons">
<emphasis id="9AF0833EFFD3670AE409F84CFC51F83A" box="[854,995,1941,1965]" italics="true" pageId="8" pageNumber="404">C acutifrons</emphasis>
</taxonomicName>
, 
<taxonomicName id="6F8424AFFFD3670AE4ACF84CFB36F839" authorityName="Poll" authorityYear="1953" box="[1011,1156,1941,1966]" class="Actinopterygii" family="Chlorophthalmidae" genus="Chlorophthalmus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Aulopiformes" pageId="8" pageNumber="404" phylum="Chordata" rank="species" species="atlanticus">
<emphasis id="9AF0833EFFD3670AE4ACF84CFB36F839" box="[1011,1156,1941,1966]" italics="true" pageId="8" pageNumber="404">C. atlanticus</emphasis>
</taxonomicName>
, 
<taxonomicName id="6F8424AFFFD3670AE3CBF84CFAC3F83A" authorityName="Kamohara" authorityYear="1953" box="[1172,1393,1942,1965]" class="Actinopterygii" family="Chlorophthalmidae" genus="Chlorophthalmus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Aulopiformes" pageId="8" pageNumber="404" phylum="Chordata" rank="species" species="nigromarginatus">
<emphasis id="9AF0833EFFD3670AE3CBF84CFAC3F83A" box="[1172,1393,1942,1965]" italics="true" pageId="8" pageNumber="404">C. nigromarginatus</emphasis>
</taxonomicName>
, 
<taxonomicName id="6F8424AFFFD3670AE2DDF84CFEAAF845" authorityName="Mead" authorityYear="1958" class="Actinopterygii" family="Chlorophthalmidae" genus="Chlorophthalmus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Aulopiformes" pageId="8" pageNumber="404" phylum="Chordata" rank="species" species="brasiliensis">
<emphasis id="9AF0833EFFD3670AE2DDF84CFEAAF845" italics="true" pageId="8" pageNumber="404">C. brasiliensis</emphasis>
</taxonomicName>
, 
<taxonomicName id="6F8424AFFFD3670AE673F860FE19F845" authorityName="Kotlyar &amp; Parin" authorityYear="1986" box="[300,427,1977,2002]" class="Actinopterygii" family="Chlorophthalmidae" genus="Chlorophthalmus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Aulopiformes" pageId="8" pageNumber="404" phylum="Chordata" rank="species" species="zvezdae">
<emphasis id="9AF0833EFFD3670AE673F860FE19F845" box="[300,427,1977,2002]" italics="true" pageId="8" pageNumber="404">C. zvezdae</emphasis>
</taxonomicName>
and 
<taxonomicName id="6F8424AFFFD3670AE6B1F860FDECF846" authorityName="Kobylianskii" authorityYear="2013" box="[494,606,1978,2001]" class="Actinopterygii" family="Chlorophthalmidae" genus="Chlorophthalmus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Aulopiformes" pageId="8" pageNumber="404" phylum="Chordata" rank="species" species="vityazi">
<emphasis id="9AF0833EFFD3670AE6B1F860FDECF846" box="[494,606,1978,2001]" italics="true" pageId="8" pageNumber="404">C. vityazi</emphasis>
</taxonomicName>
all have a smooth lower-jaw profile without tips of lower-jaw teeth extending in advance of the lower-jaw profile (
<figureCitation id="30BF43A9FFD3670AE5FAF807FCA9F862" box="[677,795,2013,2038]" captionStart="FIGURE 5" captionStartId="6.[151,250,1882,1904]" captionTargetBox="[340,1247,772,1860]" captionTargetId="figure@6.[340,1247,772,1861]" captionTargetPageId="6" captionText="FIGURE 5. Dorsal view of lower-jaw tips of (A–E) Chlorophthalmus proridens, (F) C. acutifrons, (G) C. nigromarginatus and (H, I) C. albatrossis. (A, C), USNM 47715 (paratypes); (B, D), CAS-SU 4916 (paratypes); (E), USNM 47715 (holotype); (F), KAUM–I. 151740; (G), KAUM–I. 111668; (H), KAUM–I. 151705; (I), KAUM–I. 151739." httpUri="https://zenodo.org/record/2624540/files/figure.png" pageId="8" pageNumber="404">Fig. 5F, G</figureCitation>
).
</paragraph>
</subSubSection>
</treatment>
</document>