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<document id="886B41C303B8A1F4A8D075E0DEB67473" ID-CLB-Dataset="4012" ID-DOI="10.11646/zootaxa.5051.1.16" ID-GBIF-Dataset="dffdc4cc-d108-4e27-a6c9-35042eb69b0e" ID-ISSN="1175-5326" ID-Zenodo-Dep="5563793" ID-ZooBank="BDE521C5-BB26-4445-8826-9CA5B2A00A03" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1634022355770" checkinUser="plazi" docAuthor="Kim, Jong Guk, Lee, Jimin &amp; Huys, Rony" docDate="2021" docId="03A587F5FFB21A66FF4FFEEC857CFC72" docLanguage="en" docName="zootaxa.5051.1.16.pdf" docOrigin="Zootaxa 5051 (1)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Psammis Sars 1910" docType="treatment" docVersion="6" lastPageNumber="389" masterDocId="FF9CFF8DFFB01A64FFD8FFCB841CFF9F" masterDocTitle="New records of Pseudotachidiidae from South Korea, including a key to species of the Pseudomesochrinae (Copepoda: Harpacticoida)" masterLastPageNumber="422" masterPageNumber="387" pageNumber="389" updateTime="1699459546377" updateUser="ExternalLinkService" zenodo-license-document="CLOSED">
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<mods:title id="1B6532DC43C7C85CB6C178EC59B93B9B">New records of Pseudotachidiidae from South Korea, including a key to species of the Pseudomesochrinae (Copepoda: Harpacticoida)</mods:title>
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<mods:namePart id="EE7C02B2D58D64357184513FD54C8338">Kim, Jong Guk</mods:namePart>
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<mods:affiliation id="EADE2BC6120169A9249A20BA80F9AEA6">Marine Ecosystem Research Center, Korea Institute of Ocean Science &amp; Technology, Busan 49111, Korea. &amp; jgkim @ kiost. ac. kr; https: // orcid. org / 0000 - 0001 - 5299 - 9838</mods:affiliation>
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<mods:namePart id="AEF1AC4BD14BB5D62ACF125C3D12745C">Lee, Jimin</mods:namePart>
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<mods:affiliation id="198B2FB81A0687F6DB6D40D1354FDB1F">Marine Ecosystem Research Center, Korea Institute of Ocean Science &amp; Technology, Busan 49111, Korea. &amp; leejm @ kiost. ac. kr; https: // orcid. org / 0000 - 0001 - 9004 - 8275</mods:affiliation>
<mods:nameIdentifier id="FF77586954FC1924885A2C144AFE8DF3" type="email">leejm@kiost.ac.kr</mods:nameIdentifier>
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<mods:namePart id="97A75D2257371521DC777AA4E7678F10">Huys, Rony</mods:namePart>
<mods:affiliation id="2F331836578A461433A1889D27FAD08D">Department of Life Sciences, The Natural History Museum, Cromwell Road, London SW 7 5 BD, U. K.</mods:affiliation>
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<mods:date id="EBED2E58ACCB8CACBF7D3D9E3228D4CF">2021</mods:date>
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<mods:number id="5B5982F98EE870A14F30F645A4C0343E">5051</mods:number>
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Genus
<taxonomicName id="4C0C4D60FFB21A66FF36FEEC85F8FEDE" authority="Sars, 1910" authorityName="Sars" authorityYear="1910" box="[238,484,295,321]" class="Hexanauplia" family="Pseudotachidiidae" genus="Psammis" kingdom="Animalia" order="Harpacticoida" pageId="2" pageNumber="389" phylum="Arthropoda" rank="genus">
<emphasis id="B978EAF1FFB21A66FF36FEEC8547FEDE" bold="true" box="[238,347,295,321]" italics="true" pageId="2" pageNumber="389">Psammis</emphasis>
<bibRefCitation id="EF9D4B12FFB21A66FEBBFEEC85F8FEDE" author="Sars, G. O." box="[355,484,295,321]" pageId="2" pageNumber="389" pagination="337 - 368" refId="ref24819" refString="Sars, G. O. (1910) Copepoda Harpacticoida. Parts XXIX &amp; XXX. Tachidiidae (concluded), Metidae, Balaenophilidae, supplement (part). An Account of the Crustacea of Norway, with Short Descriptions and Figures of all the Species, 5, 337 - 368, pls. CCXXV - CCXXX, supplement pls. 1 - 10." type="journal article" year="1910">Sars, 1910</bibRefCitation>
</taxonomicName>
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<subSubSection id="C3166568FFB21A66FF4FFEA4857CFC72" pageId="2" pageNumber="389" type="discussion">
<paragraph id="8BB336E3FFB21A66FF4FFEA4857CFC72" blockId="2.[151,1437,367,1005]" pageId="2" pageNumber="389">
<bibRefCitation id="EF9D4B12FFB21A66FF4FFEA4854DFE16" author="Sars, G. O." box="[151,337,367,393]" pageId="2" pageNumber="389" pagination="337 - 368" refId="ref24819" refString="Sars, G. O. (1910) Copepoda Harpacticoida. Parts XXIX &amp; XXX. Tachidiidae (concluded), Metidae, Balaenophilidae, supplement (part). An Account of the Crustacea of Norway, with Short Descriptions and Figures of all the Species, 5, 337 - 368, pls. CCXXV - CCXXX, supplement pls. 1 - 10." type="journal article" year="1910">Sars (1910: 339)</bibRefCitation>
proposed
<taxonomicName id="4C0C4D60FFB21A66FE1CFEBB8635FE16" authorityName="Sars" authorityYear="1910" box="[452,553,368,393]" class="Hexanauplia" family="Pseudotachidiidae" genus="Psammis" kingdom="Animalia" order="Harpacticoida" pageId="2" pageNumber="389" phylum="Arthropoda" rank="genus">
<emphasis id="B978EAF1FFB21A66FE1CFEBB8635FE16" box="[452,553,368,393]" italics="true" pageId="2" pageNumber="389">Psammis</emphasis>
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for its
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(by monotypy) and only species
<taxonomicName id="4C0C4D60FFB21A66FBC2FEBB812DFE16" authority="Sars, 1910" authorityName="Sars" authorityYear="1910" box="[1050,1329,367,393]" class="Hexanauplia" family="Pseudotachidiidae" genus="Psammis" kingdom="Animalia" order="Harpacticoida" pageId="2" pageNumber="389" phylum="Arthropoda" rank="species" species="longisetosa">
<emphasis id="B978EAF1FFB21A66FBC2FEBB80A9FE16" box="[1050,1205,367,393]" italics="true" pageId="2" pageNumber="389">P. longisetosa</emphasis>
<bibRefCitation id="EF9D4B12FFB21A66FB63FEA4812DFE16" author="Sars, G. O." box="[1211,1329,367,393]" pageId="2" pageNumber="389" pagination="337 - 368" refId="ref24819" refString="Sars, G. O. (1910) Copepoda Harpacticoida. Parts XXIX &amp; XXX. Tachidiidae (concluded), Metidae, Balaenophilidae, supplement (part). An Account of the Crustacea of Norway, with Short Descriptions and Figures of all the Species, 5, 337 - 368, pls. CCXXV - CCXXX, supplement pls. 1 - 10." type="journal article" year="1910">Sars, 1910</bibRefCitation>
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collected from Farsund in southern
<collectingCountry id="F31B7673FFB21A66FE62FE5F860AFE31" box="[442,534,404,430]" name="Norway" pageId="2" pageNumber="389">Norway</collectingCountry>
. The genus has seen the addition of four species since,
<emphasis id="B978EAF1FFB21A66FBA7FE5F80BEFE32" box="[1151,1186,404,429]" italics="true" pageId="2" pageNumber="389">i.e.</emphasis>
<taxonomicName id="4C0C4D60FFB21A66FB71FE5F8180FE32" authority="Klie, 1939" authorityName="Klie" authorityYear="1939" box="[1193,1436,403,429]" class="Hexanauplia" family="Pseudotachidiidae" genus="Psammis" kingdom="Animalia" order="Harpacticoida" pageId="2" pageNumber="389" phylum="Arthropoda" rank="species" species="borealis">
<emphasis id="B978EAF1FFB21A66FB71FE5F8103FE32" box="[1193,1311,403,429]" italics="true" pageId="2" pageNumber="389">P. borealis</emphasis>
<bibRefCitation id="EF9D4B12FFB21A66FAFEFE588180FE32" author="Klie, W." box="[1318,1436,403,429]" pageId="2" pageNumber="389" pagination="223 - 226" refId="ref23007" refString="Klie, W. (1939) Diagnosen neuer Harpacticoiden aus den Gewassern um Island. Zoologischer Anzeiger, 126, 223 - 226." type="journal article" year="1939">Klie, 1939</bibRefCitation>
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collected north of
<collectingCountry id="F31B7673FFB21A66FEB8FE7C85A9FE4E" box="[352,437,439,465]" name="Iceland" pageId="2" pageNumber="389">Iceland</collectingCountry>
,
<taxonomicName id="4C0C4D60FFB21A66FE67FE7386AEFE4E" authority="Smirnov, 1946" authorityName="Smirnov" authorityYear="1946" box="[447,690,439,465]" class="Hexanauplia" family="Pseudotachidiidae" genus="Psammis" kingdom="Animalia" order="Harpacticoida" pageId="2" pageNumber="389" phylum="Arthropoda" rank="species" species="kliei">
<emphasis id="B978EAF1FFB21A66FE67FE738615FE4E" box="[447,521,439,465]" italics="true" pageId="2" pageNumber="389">P. kliei</emphasis>
<bibRefCitation id="EF9D4B12FFB21A66FDD7FE7C86AEFE4E" author="Smirnov, S. S." box="[527,690,439,465]" pageId="2" pageNumber="389" pagination="231 - 263" refId="ref25667" refString="Smirnov, S. S. (1946) Novye vidy Copepoda Harpacticoida iz severnogo ledovitogo okeana. [New species of Copepoda-Harpacticoida from the northern Arctic Ocean]. Trudy Dreifuyuschei Ekspeditsii Glavsevmorputi na Ledokol'nom Parokhode ' G. Sedov', 1937 - 1940 gg, 3, 231 - 263. [in Russian with English summary]" type="journal article" year="1946">Smirnov, 1946</bibRefCitation>
</taxonomicName>
from the East Siberian Sea,
<taxonomicName id="4C0C4D60FFB21A66FC31FE73811AFE4E" authority="Bodin, 1968" authorityName="Bodin" authorityYear="1968" box="[1001,1286,439,465]" class="Hexanauplia" family="Pseudotachidiidae" genus="Psammis" kingdom="Animalia" order="Harpacticoida" pageId="2" pageNumber="389" phylum="Arthropoda" rank="species" species="longifurca">
<emphasis id="B978EAF1FFB21A66FC31FE73806BFE4E" box="[1001,1143,439,465]" italics="true" pageId="2" pageNumber="389">P. longifurca</emphasis>
<bibRefCitation id="EF9D4B12FFB21A66FBA5FE7C811AFE4E" author="Bodin, P." box="[1149,1286,439,465]" pageId="2" pageNumber="389" pagination="1 - 107" refId="ref19241" refString="Bodin, P. (1968) Copepodes Harpacticoides des etages bathyal et abyssal du golfe de Gascogne. Memoires du Museum national d'Histoire naturelle, Paris, Nouvelle Serie (A), 55 (1), 1 - 107." type="journal article" year="1968">Bodin, 1968</bibRefCitation>
</taxonomicName>
from the Bay of Biscay and
<taxonomicName id="4C0C4D60FFB21A66FEE4FE17864AFE6A" authority="Becker, 1974" authorityName="Becker" authorityYear="1974" box="[316,598,475,501]" class="Hexanauplia" family="Pseudotachidiidae" genus="Psammis" kingdom="Animalia" order="Harpacticoida" pageId="2" pageNumber="389" phylum="Arthropoda" rank="species" species="longipes">
<emphasis id="B978EAF1FFB21A66FEE4FE1785A4FE6A" box="[316,440,475,501]" italics="true" pageId="2" pageNumber="389">P. longipes</emphasis>
<bibRefCitation id="EF9D4B12FFB21A66FE18FE10864AFE6A" author="Becker, K. - H." box="[448,598,475,501]" pageId="2" pageNumber="389" pagination="1 - 28" refId="ref19142" refString="Becker, K. - H. (1974) Eidonomie und Taxonomie abyssaler Harpacticoidea (Crustacea, Copepoda). Teil 1. Cerviniidae - Ameiridae. Meteor Forschungs-Ergebnisse (D), 18, 1 - 28." type="journal article" year="1974">Becker, 1974</bibRefCitation>
</taxonomicName>
from the
<collectingCountry id="F31B7673FFB21A66FD1FFE1086E7FE6A" box="[711,763,475,501]" name="Peru" pageId="2" pageNumber="389">Peru</collectingCountry>
–
<collectingCountry id="F31B7673FFB21A66FCD2FE10875AFE6A" box="[778,838,475,501]" name="Chile" pageId="2" pageNumber="389">Chile</collectingCountry>
(
<collectingRegion id="49C8F801FFB21A66FC8DFE1087A0FE6A" box="[853,956,475,501]" country="Chile" name="Atacama" pageId="2" pageNumber="389">Atacama</collectingRegion>
) Trench (
<bibRefCitation id="EF9D4B12FFB21A66FBF2FE108083FE6A" author="Klie, W." box="[1066,1183,475,501]" pageId="2" pageNumber="389" pagination="223 - 226" refId="ref23007" refString="Klie, W. (1939) Diagnosen neuer Harpacticoiden aus den Gewassern um Island. Zoologischer Anzeiger, 126, 223 - 226." type="journal article" year="1939">Klie 1939</bibRefCitation>
,
<bibRefCitation id="EF9D4B12FFB21A66FB74FE1080FBFE6A" author="Klie, W." box="[1196,1255,475,501]" pageId="2" pageNumber="389" pagination="1 - 44" refId="ref23032" refString="Klie, W. (1941) Marine Harpacticoiden von Island. Kieler Meeresforschungen, 5, 1 - 44." type="journal article" year="1941">1941</bibRefCitation>
;
<bibRefCitation id="EF9D4B12FFB21A66FB2CFE10818BFE6A" author="Smirnov, S. S." box="[1268,1431,475,501]" pageId="2" pageNumber="389" pagination="231 - 263" refId="ref25667" refString="Smirnov, S. S. (1946) Novye vidy Copepoda Harpacticoida iz severnogo ledovitogo okeana. [New species of Copepoda-Harpacticoida from the northern Arctic Ocean]. Trudy Dreifuyuschei Ekspeditsii Glavsevmorputi na Ledokol'nom Parokhode ' G. Sedov', 1937 - 1940 gg, 3, 231 - 263. [in Russian with English summary]" type="journal article" year="1946">Smirnov 1946</bibRefCitation>
;
<bibRefCitation id="EF9D4B12FFB21A66FF4FFE348502FD86" author="Bodin, P." box="[151,286,511,537]" pageId="2" pageNumber="389" pagination="1 - 107" refId="ref19241" refString="Bodin, P. (1968) Copepodes Harpacticoides des etages bathyal et abyssal du golfe de Gascogne. Memoires du Museum national d'Histoire naturelle, Paris, Nouvelle Serie (A), 55 (1), 1 - 107." type="journal article" year="1968">Bodin 1968</bibRefCitation>
;
<bibRefCitation id="EF9D4B12FFB21A66FEF3FE3485A2FD86" author="Becker, K. - H." box="[299,446,511,537]" pageId="2" pageNumber="389" pagination="1 - 28" refId="ref19142" refString="Becker, K. - H. (1974) Eidonomie und Taxonomie abyssaler Harpacticoidea (Crustacea, Copepoda). Teil 1. Cerviniidae - Ameiridae. Meteor Forschungs-Ergebnisse (D), 18, 1 - 28." type="journal article" year="1974">Becker 1974</bibRefCitation>
).
<bibRefCitation id="EF9D4B12FFB21A66FE09FE34867FFD85" author="Wells, J. B. J." box="[465,611,511,538]" pageId="2" pageNumber="389" pagination="189 - 358" refId="ref26534" refString="Wells, J. B. J. (1967) The littoral Copepoda (Crustacea) of Inhaca Island, Mozambique. Transactions of the Royal Society of Edinburgh, 67, 189 - 358. https: // doi. org / 10.1017 / s 0080456800024017" type="journal article" year="1967">Wells (1967)</bibRefCitation>
had previously expressed doubts about the validity of the genus, arguing that the only character that separates it from
<taxonomicName id="4C0C4D60FFB21A66FD5CFDE8870AFDA2" authorityName="Boeck" authorityYear="1873" box="[644,790,547,573]" class="Hexanauplia" family="Pseudotachidiidae" genus="Danielssenia" kingdom="Animalia" order="Harpacticoida" pageId="2" pageNumber="389" phylum="Arthropoda" rank="genus">
<emphasis id="B978EAF1FFB21A66FD5CFDE8870AFDA2" box="[644,790,547,573]" italics="true" pageId="2" pageNumber="389">Danielssenia</emphasis>
</taxonomicName>
is the number of setae on the exopod of P5; however,
<bibRefCitation id="EF9D4B12FFB21A66FAB7FDE884FEFDFE" author="Gee, J. M." pageId="2" pageNumber="389" pagination="181 - 194" refId="ref20921" refString="Gee, J. M. (1988) Some harpacticoid copepods (Crustacea) of the family Tachidiidae from sublittoral soft sediments in Norway, the Celtic Sea and Gulf of Mexico. Zoologica Scripta, 17, 181 - 194. https: // doi. org / 10.1111 / j. 1463 - 6409.1988. tb 00094. x" type="journal article" year="1988">Gee (1988)</bibRefCitation>
pointed out that species of both genera are probably trophically isolated as indicated by differences in mandibular gnathobase morphology and that this—in conjunction with the fused rami of P5—constitutes sufficient evidence for maintaining the genera separate.
<bibRefCitation id="EF9D4B12FFB21A66FDA9FD5B8757FD36" author="Huys, R. &amp; Gee, J. M." box="[625,843,655,682]" pageId="2" pageNumber="389" pagination="45 - 81" refId="ref22117" refString="Huys, R. &amp; Gee, J. M. (1993) A revision of Danielssenia Boeck and Psammis Sars with the establishment of two new genera Archisenia and Bathypsammis (Harpacticoida: Paranannopidae). Bulletin of the Natural History Museum, Zoology, 59, 45 - 81." type="journal article" year="1993">Huys &amp; Gee (1993)</bibRefCitation>
redefined the boundaries of
<taxonomicName id="4C0C4D60FFB21A66FB5EFD5B80F7FD36" authorityName="Sars" authorityYear="1910" box="[1158,1259,656,681]" class="Hexanauplia" family="Pseudotachidiidae" genus="Psammis" kingdom="Animalia" order="Harpacticoida" pageId="2" pageNumber="389" phylum="Arthropoda" rank="genus">
<emphasis id="B978EAF1FFB21A66FB5EFD5B80F7FD36" box="[1158,1259,656,681]" italics="true" pageId="2" pageNumber="389">Psammis</emphasis>
</taxonomicName>
and restricted it to its
<typeStatus id="54B78841FFB21A66FF0BFD7F851FFD51" box="[211,259,692,718]" pageId="2" pageNumber="389">type</typeStatus>
species and
<taxonomicName id="4C0C4D60FFB21A66FE56FD7F861BFD52" authorityName="Becker" authorityYear="1974" box="[398,519,691,717]" class="Hexanauplia" family="Pseudotachidiidae" genus="Psammis" kingdom="Animalia" order="Harpacticoida" pageId="2" pageNumber="389" phylum="Arthropoda" rank="species" species="longipes">
<emphasis id="B978EAF1FFB21A66FE56FD7F861BFD52" box="[398,519,691,717]" italics="true" pageId="2" pageNumber="389">P. longipes</emphasis>
</taxonomicName>
. While
<taxonomicName id="4C0C4D60FFB21A66FD85FD7F86F7FD52" authorityName="Bodin" authorityYear="1968" box="[605,747,691,717]" class="Hexanauplia" family="Pseudotachidiidae" genus="Psammis" kingdom="Animalia" order="Harpacticoida" pageId="2" pageNumber="389" phylum="Arthropoda" rank="species" species="longifurca">
<emphasis id="B978EAF1FFB21A66FD85FD7F86F7FD52" box="[605,747,691,717]" italics="true" pageId="2" pageNumber="389">P. longifurca</emphasis>
</taxonomicName>
was fixed as the
<typeStatus id="54B78841FFB21A66FC7EFD7F87CAFD51" box="[934,982,692,718]" pageId="2" pageNumber="389">type</typeStatus>
of the new genus
<taxonomicName id="4C0C4D60FFB21A66FB45FD788511FD6E" authority="Huys &amp; Gee, 1993" authorityName="Huys &amp; Gee" authorityYear="1993" class="Hexanauplia" family="Pseudotachidiidae" genus="Bathypsammis" kingdom="Animalia" order="Harpacticoida" pageId="2" pageNumber="389" phylum="Arthropoda" rank="genus">
<emphasis id="B978EAF1FFB21A66FB45FD78815CFD52" box="[1181,1344,691,717]" italics="true" pageId="2" pageNumber="389">Bathypsammis</emphasis>
<bibRefCitation id="EF9D4B12FFB21A66FA9EFD7F8511FD6E" author="Huys, R. &amp; Gee, J. M." pageId="2" pageNumber="389" pagination="45 - 81" refId="ref22117" refString="Huys, R. &amp; Gee, J. M. (1993) A revision of Danielssenia Boeck and Psammis Sars with the establishment of two new genera Archisenia and Bathypsammis (Harpacticoida: Paranannopidae). Bulletin of the Natural History Museum, Zoology, 59, 45 - 81." type="journal article" year="1993">Huys &amp; Gee, 1993</bibRefCitation>
</taxonomicName>
, both
<taxonomicName id="4C0C4D60FFB21A66FE89FD1385DBFD6E" authorityName="Klie" authorityYear="1939" box="[337,455,727,753]" class="Hexanauplia" family="Pseudotachidiidae" genus="Psammis" kingdom="Animalia" order="Harpacticoida" pageId="2" pageNumber="389" phylum="Arthropoda" rank="species" species="borealis">
<emphasis id="B978EAF1FFB21A66FE89FD1385DBFD6E" box="[337,455,727,753]" italics="true" pageId="2" pageNumber="389">P. borealis</emphasis>
</taxonomicName>
and
<taxonomicName id="4C0C4D60FFB21A66FE25FD138654FD6E" authorityName="Smirnov" authorityYear="1946" box="[509,584,727,753]" class="Hexanauplia" family="Pseudotachidiidae" genus="Psammis" kingdom="Animalia" order="Harpacticoida" pageId="2" pageNumber="389" phylum="Arthropoda" rank="species" species="kliei">
<emphasis id="B978EAF1FFB21A66FE25FD138654FD6E" box="[509,584,727,753]" italics="true" pageId="2" pageNumber="389">P. kliei</emphasis>
</taxonomicName>
were tentatively maintained as
<emphasis id="B978EAF1FFB21A66FC73FD138081FD6E" box="[939,1181,727,753]" italics="true" pageId="2" pageNumber="389">species incertae sedis</emphasis>
in the
<taxonomicName id="4C0C4D60FFB21A66FB31FD1C8180FD6E" authorityName="Por" authorityYear="1986" box="[1257,1436,727,753]" class="Hexanauplia" family="Paranannopidae" kingdom="Animalia" order="Harpacticoida" pageId="2" pageNumber="389" phylum="Arthropoda" rank="family">Paranannopidae</taxonomicName>
(=
<taxonomicName id="4C0C4D60FFB21A66FF6EFD308577FC8A" box="[182,363,763,789]" class="Hexanauplia" family="Pseudotachidiidae" kingdom="Animalia" order="Harpacticoida" pageId="2" pageNumber="418" phylum="Arthropoda" rank="subFamily" subFamily="Danielsseniinae">Danielsseniinae</taxonomicName>
;
<emphasis id="B978EAF1FFB21A66FEADFD30858EFC8A" box="[373,402,763,789]" italics="true" pageId="2" pageNumber="389">cf.</emphasis>
<bibRefCitation id="EF9D4B12FFB21A66FE4FFD37860EFC8A" author="Huys, R." box="[407,530,763,790]" pageId="2" pageNumber="389" pagination="1 - 99" refId="ref21962" refString="Huys, R. (2009) Unresolved cases of type fixation, synonymy and homonymy in harpacticoid copepod nomenclature (Crustacea: Copepoda). Zootaxa, 2183 (1), 1 - 99. https: // doi. org / 10.11646 / zootaxa. 2183.1.1" type="journal article" year="2009">Huys 2009</bibRefCitation>
) pending further re-examination. In subsequent contributions
<bibRefCitation id="EF9D4B12FFB21A66FB1AFD308180FC8A" author="Gee, J. M. &amp; Huys, R." box="[1218,1436,763,790]" pageId="2" pageNumber="389" pagination="1007 - 1046" refId="ref21049" refString="Gee, J. M. &amp; Huys, R. (1994) Paranannopidae (Copepoda: Harpacticoida) from sublittoral soft sediments in Spitsbergen. Journal of natural History, 28, 1007 - 1046. https: // doi. org / 10.1080 / 00222939400770541" type="journal article" year="1994">Gee &amp; Huys (1994)</bibRefCitation>
assigned
<taxonomicName id="4C0C4D60FFB21A66FED8FCEB8552FCA6" authorityName="Smirnov" authorityYear="1946" box="[256,334,799,825]" class="Hexanauplia" family="Pseudotachidiidae" genus="Psammis" kingdom="Animalia" order="Harpacticoida" pageId="2" pageNumber="389" phylum="Arthropoda" rank="species" species="kliei">
<emphasis id="B978EAF1FFB21A66FED8FCEB8552FCA6" box="[256,334,799,825]" italics="true" pageId="2" pageNumber="389">P. kliei</emphasis>
</taxonomicName>
to the genus
<taxonomicName id="4C0C4D60FFB21A66FE32FCEB8752FCA6" authority="Gee &amp; Huys, 1994" authorityName="Gee &amp; Huys" authorityYear="1994" box="[490,846,799,826]" class="Hexanauplia" family="Pseudotachidiidae" genus="Mucrosenia" kingdom="Animalia" order="Harpacticoida" pageId="2" pageNumber="389" phylum="Arthropoda" rank="genus">
<emphasis id="B978EAF1FFB21A66FE32FCEB8673FCA6" box="[490,623,800,825]" italics="true" pageId="2" pageNumber="389">Mucrosenia</emphasis>
<bibRefCitation id="EF9D4B12FFB21A66FDAFFCD48752FCA6" author="Gee, J. M. &amp; Huys, R." box="[631,846,799,826]" pageId="2" pageNumber="389" pagination="1007 - 1046" refId="ref21049" refString="Gee, J. M. &amp; Huys, R. (1994) Paranannopidae (Copepoda: Harpacticoida) from sublittoral soft sediments in Spitsbergen. Journal of natural History, 28, 1007 - 1046. https: // doi. org / 10.1080 / 00222939400770541" type="journal article" year="1994">Gee &amp; Huys, 1994</bibRefCitation>
</taxonomicName>
as
<emphasis id="B978EAF1FFB21A66FCAFFCEB8071FCA6" box="[887,1133,799,825]" italics="true" pageId="2" pageNumber="389">species incertae sedis</emphasis>
and Huys and Gee (1996) designated
<taxonomicName id="4C0C4D60FFB21A66FEC0FC8F858DFCC2" authorityName="Klie" authorityYear="1939" box="[280,401,835,861]" class="Hexanauplia" family="Pseudotachidiidae" genus="Psammis" kingdom="Animalia" order="Harpacticoida" pageId="2" pageNumber="389" phylum="Arthropoda" rank="species" species="borealis">
<emphasis id="B978EAF1FFB21A66FEC0FC8F858DFCC2" box="[280,401,835,861]" italics="true" pageId="2" pageNumber="389">P. borealis</emphasis>
</taxonomicName>
as the
<typeStatus id="54B78841FFB21A66FE3EFC8F860AFCC1" box="[486,534,836,862]" pageId="2" pageNumber="389">type</typeStatus>
(by original designation) of their new genus
<taxonomicName id="4C0C4D60FFB21A66FBF9FC888184FCC2" authority="Huys &amp; Gee, 1996" authorityName="Huys &amp; Gee" authorityYear="1996" box="[1057,1432,835,862]" class="Hexanauplia" family="Pseudotachidiidae" genus="Anapophysia" kingdom="Animalia" order="Harpacticoida" pageId="2" pageNumber="389" phylum="Arthropoda" rank="genus">
<emphasis id="B978EAF1FFB21A66FBF9FC8880AFFCC2" box="[1057,1203,835,861]" italics="true" pageId="2" pageNumber="389">Anapophysia</emphasis>
Huys &amp; Gee, 1996
</taxonomicName>
. The original descriptions of both sexes of
<taxonomicName id="4C0C4D60FFB21A66FDB2FCA38719FC1E" authorityName="Sars" authorityYear="1910" box="[618,773,871,897]" class="Hexanauplia" family="Pseudotachidiidae" genus="Psammis" kingdom="Animalia" order="Harpacticoida" pageId="2" pageNumber="389" phylum="Arthropoda" rank="species" species="longisetosa">
<emphasis id="B978EAF1FFB21A66FDB2FCA38719FC1E" box="[618,773,871,897]" italics="true" pageId="2" pageNumber="389">P. longisetosa</emphasis>
</taxonomicName>
by
<bibRefCitation id="EF9D4B12FFB21A66FCF5FCAC800BFC1E" author="Sars, G. O." box="[813,1047,871,897]" pageId="2" pageNumber="389" pagination="337 - 368" refId="ref24819" refString="Sars, G. O. (1910) Copepoda Harpacticoida. Parts XXIX &amp; XXX. Tachidiidae (concluded), Metidae, Balaenophilidae, supplement (part). An Account of the Crustacea of Norway, with Short Descriptions and Figures of all the Species, 5, 337 - 368, pls. CCXXV - CCXXX, supplement pls. 1 - 10." type="journal article" year="1910">Sars (1910: 339–340</bibRefCitation>
, plate CCXXV; 1921: 94–95, plate LXV.) were supplemented and largely superseded by a number of subsequent re-examinations including those by
<bibRefCitation id="EF9D4B12FFB21A66FF4FFC64859DFC56" author="Gee, J. M." box="[151,385,943,969]" pageId="2" pageNumber="389" pagination="181 - 194" refId="ref20921" refString="Gee, J. M. (1988) Some harpacticoid copepods (Crustacea) of the family Tachidiidae from sublittoral soft sediments in Norway, the Celtic Sea and Gulf of Mexico. Zoologica Scripta, 17, 181 - 194. https: // doi. org / 10.1111 / j. 1463 - 6409.1988. tb 00094. x" type="journal article" year="1988">Gee (1988: 181–187</bibRefCitation>
,
<figureCitation id="13372A66FFB21A66FE55FC6485E4FC56" box="[397,504,943,969]" captionStart-0="FIGURE 1" captionStart-1="FIGURE 2" captionStart-2="FIGURE 3" captionStart-3="FIGURE 4" captionStartId-0="2.[151,250,1992,2016]" captionStartId-1="8.[151,250,1999,2023]" captionStartId-2="9.[151,250,1943,1967]" captionStartId-3="10.[151,250,1893,1917]" captionTargetBox-0="[151,1433,1034,1955]" captionTargetBox-1="[162,1425,238,1975]" captionTargetBox-2="[178,1409,181,1913]" captionTargetBox-3="[192,1395,181,1869]" captionTargetId-0="figure-356@2.[151,1436,1027,1967]" captionTargetId-1="figure-51@8.[162,1425,237,1975]" captionTargetId-2="figure-19@9.[178,1409,181,1919]" captionTargetId-3="figure-19@10.[192,1395,181,1869]" captionTargetPageId-0="2" captionTargetPageId-1="8" captionTargetPageId-2="9" captionTargetPageId-3="10" captionText-0="FIGURE 1. Map showing sampling stations in the Southern Sea." captionText-1="FIGURE 2. Psammis wellsi sp. nov. (♂): A, habitus, lateral; B, prosome, dorsal; C, urosome, dorsal." captionText-2="FIGURE 3. Psammis wellsi sp. nov., (♂): A, urosome, ventral; B, left caudal ramus, ventral; C, antennule (armature omitted); C’, antennule (disarticulated; segments shown in posterior view indicated by asterisks)." captionText-3="FIGURE 4. Psammis wellsi sp. nov. (♂): A, antenna (inset showing disarticulated exopod with complete armature); B, mandi- ble with disarticulated palp; C, maxillule with disarticulated palp (exopod shown in separate inset); D, maxilla (praecoxal endite and endopod shown in separate insets); E, maxilliped (minute seta on syncoxa indicated by arrowhead). Arrowheads in C–D indicate tubular setae or extensions." figureDoi-0="http://doi.org/10.5281/zenodo.5563795" figureDoi-1="http://doi.org/10.5281/zenodo.5563797" figureDoi-2="http://doi.org/10.5281/zenodo.5563801" figureDoi-3="http://doi.org/10.5281/zenodo.5563805" httpUri-0="https://zenodo.org/record/5563795/files/figure.png" httpUri-1="https://zenodo.org/record/5563797/files/figure.png" httpUri-2="https://zenodo.org/record/5563801/files/figure.png" httpUri-3="https://zenodo.org/record/5563805/files/figure.png" pageId="2" pageNumber="389">Figs. 1–4</figureCitation>
),
<bibRefCitation id="EF9D4B12FFB21A66FDD6FC7B8739FC56" author="Huys, R. &amp; Gee, J. M." box="[526,805,943,970]" pageId="2" pageNumber="389" pagination="45 - 81" refId="ref22117" refString="Huys, R. &amp; Gee, J. M. (1993) A revision of Danielssenia Boeck and Psammis Sars with the establishment of two new genera Archisenia and Bathypsammis (Harpacticoida: Paranannopidae). Bulletin of the Natural History Museum, Zoology, 59, 45 - 81." type="journal article" year="1993">Huys and Gee (1993: 72</bibRefCitation>
, 75, Figs. 21–22) and Huys
<emphasis id="B978EAF1FFB21A66FBBCFC7B8089FC56" box="[1124,1173,943,969]" italics="true" pageId="2" pageNumber="389">et al</emphasis>
. (1996: 248–249, Figs. 93B, 98, Table 4).
</paragraph>
</subSubSection>
</treatment>
</document>
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