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<document id="A0043882AC9D9AFA10B53B0124730391" ID-CLB-Dataset="20629" ID-DOI="10.11646/zootaxa.5099.2.4" ID-GBIF-Dataset="1b46d215-475d-471b-ac35-cc9490be53f4" ID-ISSN="1175-5326" ID-Zenodo-Dep="6048528" ID-ZooBank="442E337A-E7A1-4B30-AB87-D252B8E938B2" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1644581927792" checkinUser="plazi" docAuthor="Marques, Mariana P., Parrinha, Diogo, Santos, Bruna S., Bandeira, Suzana, Butler, Brett O., Sousa, Ana Carolina A., Bauer, Aaron M. &amp; Wagner, Philipp" docDate="2022" docId="03A587EA7926FFC3FF6F5550FD41FE06" docLanguage="en" docName="zootaxa.5099.2.4.pdf" docOrigin="Zootaxa 5099 (2)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Acanthocercus ceriacoi Marques &amp; Parrinha &amp; Santos &amp; Bandeira &amp; Butler &amp; Sousa &amp; Bauer &amp; Wagner 2022, sp. nov." docType="treatment" docVersion="12" lastPageNumber="235" masterDocId="FF9CFF927922FFCDFFF8527CFFF7FF8E" masterDocTitle="All in all its just another branch in the tree: A new species of Acanthocercus Fitzinger, 1843 (Squamata: Agamidae), from Angola" masterLastPageNumber="243" masterPageNumber="221" pageNumber="225" updateTime="1722559247796" updateUser="ExternalLinkService" zenodo-license-document="CLOSED">
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<mods:title id="1A00A0F30DDD3ED4C3444113E5AA7FAF">All in all its just another branch in the tree: A new species of Acanthocercus Fitzinger, 1843 (Squamata: Agamidae), from Angola</mods:title>
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<mods:affiliation id="2EBDA6A69F9C2D9D042F363B42A92704">Departamento de Biologia, Faculdade de Ciências da Universidade do Porto, Rua do Campo Alegre 1021, 4169 - 007 Porto, Portugal BIOPOLIS Program in Genomics, Biodiversity and Land Planning, CIBIO, Campus de Vairão, 4485 - 661, Vairão, Portugal Departamento de Zoologia e Antropologia (Museu Bocage), Museu Nacional de História Natural e da Ciência, Rua da Escola Politécnica 56 - 58, 1250 - 102 Lisboa, Portugal</mods:affiliation>
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<mods:namePart id="E5A1EA484AB5433C977EE0B57702CE30">Parrinha, Diogo</mods:namePart>
<mods:affiliation id="6635D6E5B5FC25F61909297421DBD860">Departamento de Biologia, Faculdade de Ciências da Universidade do Porto, Rua do Campo Alegre 1021, 4169 - 007 Porto, Portugal BIOPOLIS Program in Genomics, Biodiversity and Land Planning, CIBIO, Campus de Vairão, 4485 - 661, Vairão, Portugal Departamento de Zoologia e Antropologia (Museu Bocage), Museu Nacional de História Natural e da Ciência, Rua da Escola Politécnica 56 - 58, 1250 - 102 Lisboa, Portugal</mods:affiliation>
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<mods:affiliation id="2635D898FF65485A83447523F24FFE9F">Departamento de Biologia, Faculdade de Ciências da Universidade do Porto, Rua do Campo Alegre 1021, 4169 - 007 Porto, Portugal BIOPOLIS Program in Genomics, Biodiversity and Land Planning, CIBIO, Campus de Vairão, 4485 - 661, Vairão, Portugal Departamento de Zoologia e Antropologia (Museu Bocage), Museu Nacional de História Natural e da Ciência, Rua da Escola Politécnica 56 - 58, 1250 - 102 Lisboa, Portugal</mods:affiliation>
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<mods:affiliation id="06055E882C8A85274E23131011657A32">Holísticos, Serviços, Estudos e Consultoria, Lda., Rua 60, Casa 560, Lar do Patriota, Luanda, Angola &amp; suzanaadelino 22 @ hotmail. com; https: // orcid. org / 0000 - 0002 - 7496 - 9119</mods:affiliation>
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<mods:affiliation id="41FF977B6A9CE777F87E0F39CD85117F">Department of Biology and Center for Biodiversity and Ecosystem Stewardship, Villanova University, 800 Lancaster Avenue, Villanova, Pennsylvania 19085, USA &amp; Museo de Zoología “ Alfonso L. Herrera ”, Departamento de Biología Evolutiva, Facultad de Ciencias, Universidad Nacional Autónoma de México, A. P. 70 - 399, Ciudad de México CP 04510, México</mods:affiliation>
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<mods:namePart id="8CF8D318085E6AA1C84A2C97AB2D3326">Sousa, Ana Carolina A.</mods:namePart>
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<mods:affiliation id="053ED2E4FF7D57D68DA235AB9110715F">Grutas da Moeda e Fátima, Lda, São Mamede, Batalha, 2495 - 028, Portugal &amp; sousa _ 334 @ hotmail. com; https: // orcid. org / 0000 - 0003 - 3435 - 8775</mods:affiliation>
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<mods:namePart id="6812D85C8E043F42689A56F0AEE16CD4">Bauer, Aaron M.</mods:namePart>
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<mods:affiliation id="54778CBC510C7ACCE720CC8AF53A3C62">Department of Biology and Center for Biodiversity and Ecosystem Stewardship, Villanova University, 800 Lancaster Avenue, Villanova, Pennsylvania 19085, USA &amp; aaron. bauer @ villanova. edu; https: // orcid. org / 0000 - 0001 - 6839 - 8025</mods:affiliation>
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<mods:namePart id="C336B38DE8A247F86D6D446F582D9071">Wagner, Philipp</mods:namePart>
<mods:affiliation id="9B03E721CEFA286943BB83881B75D031">Department of Biology and Center for Biodiversity and Ecosystem Stewardship, Villanova University, 800 Lancaster Avenue, Villanova, Pennsylvania 19085, USA &amp; Allwetterzoo, Sentruper Str. 315, D 48161 Münster, Germany</mods:affiliation>
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<emphasis id="B978EAEE7926FFC9FF6F5550FE44F8C9" bold="true" box="[151,435,1836,1863]" italics="true" pageId="4" pageNumber="225">Acanthocercus ceriacoi</emphasis>
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(
<figureCitation id="13372A797926FFC9FF67552DFEFDF8E5" box="[159,266,1873,1899]" captionStart-0="FIGURE 2" captionStart-1="FIGURE 3" captionStart-2="FIGURE 4" captionStart-3="FIGURE 5" captionStart-4="FIGURE 6" captionStart-5="FIGURE 7" captionStart-6="FIGURE 8" captionStartId-0="5.[151,250,1986,2010]" captionStartId-1="6.[151,250,1752,1776]" captionStartId-2="7.[151,250,1963,1987]" captionStartId-3="8.[151,250,1806,1830]" captionStartId-4="12.[151,250,1507,1531]" captionStartId-5="13.[151,250,992,1016]" captionStartId-6="13.[151,250,1967,1991]" captionTargetBox-0="[190,1399,444,1960]" captionTargetBox-1="[255,1333,181,1729]" captionTargetBox-2="[153,1435,319,1939]" captionTargetBox-3="[151,1436,181,1782]" captionTargetBox-4="[240,1347,463,1474]" captionTargetBox-5="[203,1384,181,968]" captionTargetBox-6="[215,1371,1077,1944]" captionTargetId-0="figure-110@5.[189,1399,435,1962]" captionTargetId-1="figure-23@6.[255,1333,181,1729]" captionTargetId-2="figure-76@7.[151,1435,309,1940]" captionTargetId-3="figure-23@8.[151,1436,181,1782]" captionTargetId-4="figure-149@12.[230,1357,453,1484]" captionTargetId-5="figure-18@13.[203,1384,181,968]" captionTargetId-6="figure-47@13.[215,1371,1077,1944]" captionTargetPageId-0="5" captionTargetPageId-1="6" captionTargetPageId-2="7" captionTargetPageId-3="8" captionTargetPageId-4="12" captionTargetPageId-5="13" captionTargetPageId-6="13" captionText-0="FIGURE 2. Holotype of Acanthocercus ceriacoi sp. nov. (CAS 258430). Photos by Erica Ely." captionText-1="FIGURE 3. Life photo of the holotype (CAS 258430), an adult male of Acanthocercus ceriacoi sp. nov. from Cangandala National Park. Photo by Luis M.P. Ceríaco." captionText-2="FIGURE 4. Paratype of Acanthocercus ceriacoi sp. nov. (Paratype, MHNCUP/REP 860), an adult female. Photos by Luis M.P. Ceríaco." captionText-3="FIGURE 5. Life photos of an adult female of Acanthocercus ceriacoi sp. nov. (Paratype, MHNCUP/REP 860) from Uíge Province. Photos by Luis M.P. Ceríaco." captionText-4="FIGURE 6. Geographic distribution of Angolan Acanthocercus. Green triangles: A. cyanocephalus; Pink squares: A. margaritae; Blue circles: A. ceriacoi sp. nov.; Black star: Type locality of A. ceriacoi sp. nov.. A. cyanocephalus and A. margaritae records from Marques et al. (2018), Wagner et al. (2018, 2021)." captionText-5="FIGURE 7. Habitat of the type locality of Acanthocercus ceriacoi sp. nov. in Cangandala National Park, Malanje Province, Angola. Photo by Luis M.P. Ceríaco." captionText-6="FIGURE 8. Habitat of Acanthocercus ceriacoi sp. nov. at the paratype locality of Zulumongo village, Uíge Province, northern Angola. Photo by Luis M.P. Ceríaco." figureDoi-0="http://doi.org/10.5281/zenodo.6048532" figureDoi-1="http://doi.org/10.5281/zenodo.6048534" figureDoi-2="http://doi.org/10.5281/zenodo.6048538" figureDoi-3="http://doi.org/10.5281/zenodo.6048540" figureDoi-4="http://doi.org/10.5281/zenodo.6048542" figureDoi-5="http://doi.org/10.5281/zenodo.6048546" figureDoi-6="http://doi.org/10.5281/zenodo.6048550" httpUri-0="https://zenodo.org/record/6048532/files/figure.png" httpUri-1="https://zenodo.org/record/6048534/files/figure.png" httpUri-2="https://zenodo.org/record/6048538/files/figure.png" httpUri-3="https://zenodo.org/record/6048540/files/figure.png" httpUri-4="https://zenodo.org/record/6048542/files/figure.png" httpUri-5="https://zenodo.org/record/6048546/files/figure.png" httpUri-6="https://zenodo.org/record/6048550/files/figure.png" pageId="4" pageNumber="225">Figs. 28</figureCitation>
)
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<treatmentCitationGroup id="AB1C11D27926FFC9FF6F55E9FB88F820" box="[151,1151,1941,1966]" pageId="4" pageNumber="225">
<treatmentCitation id="0AAD10ED7926FFC9FF6F55E9FDE7F820" author="Bocage, J. V. B." box="[151,528,1941,1966]" page=")" pageId="4" pageNumber="225" year="1866">
<taxonomicName id="4C0C4D7F7926FFC9FF6F55E9FDE7F820" ID-CoL="CGFCC" authority=": Bocage (1866: 43)" authorityName=": Bocage" authorityPageNumber="43" authorityYear="1866" box="[151,528,1941,1966]" class="Reptilia" family="Agamidae" genus="Stellio" higherTaxonomySource="GBIF" kingdom="Animalia" order="Squamata" pageId="4" pageNumber="225" phylum="Chordata" rank="species" species="nigricollis">
<emphasis id="B978EAEE7926FFC9FF6F55E9FEB1F823" box="[151,326,1941,1965]" italics="true" pageId="4" pageNumber="225">Stellio nigricollis</emphasis>
:
<bibRefCitation id="EF9D4B0D7926FFC9FEAC55EAFDE7F820" author="Bocage, J. V. B." box="[340,528,1941,1966]" pageId="4" pageNumber="225" pagination="37 - 56" refId="ref9333" refString="Bocage, J. V. B. (1866) Lista de reptis das possessoes potuguezas d'Africa ocidental que existem no Museu de Lisboa. Jornal de Sciencias Mathematicas, Physicas e Naturaes, 1, 37 - 56." type="journal article" year="1866">Bocage (1866: 43)</bibRefCitation>
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[
<emphasis id="B978EAEE7926FFC9FDE755E9FD5CF823" box="[543,683,1941,1965]" italics="true" pageId="4" pageNumber="225">lapsus calami</emphasis>
; see
<bibRefCitation id="EF9D4B0D7926FFC9FD2755EAFC82F823" author="Bocage, J. V. B." box="[735,885,1941,1966]" pageId="4" pageNumber="225" refId="ref9421" refString="Bocage, J. V. B. (1895) Herpetologie d'Angola et du Congo. Ministerio da Marinha e das Colonias, Lisbonne, 203 pp., 20 pls." type="book" year="1895">Bocage (1895)</bibRefCitation>
and
<bibRefCitation id="EF9D4B0D7926FFC9FC5E55EAFB8EF820" author="Wagner, P. &amp; Greenbaum, E. &amp; Bauer, A. M. &amp; Kusamba, C. &amp; Leache, A. D." box="[934,1145,1941,1966]" pageId="4" pageNumber="225" pagination="771 - 817" refId="ref12867" refString="Wagner, P., Greenbaum, E., Bauer, A. M., Kusamba, C. &amp; Leache, A. D. (2018) Lifting the blue-headed veil - integrative taxonomy of the Acanthocercus atricollis species complex (Squamata: Agamidae). Journal of Natural History, 52, 771 - 817. https: // doi. org / 10.1080 / 00222933.2018.1435833" type="journal article" year="2018">
Wagner
<emphasis id="B978EAEE7926FFC9FC0255EAFBD9F823" box="[1018,1070,1941,1965]" italics="true" pageId="4" pageNumber="225">et al.</emphasis>
(2018)
</bibRefCitation>
]
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</paragraph>
<paragraph id="8BB336FC7926FFC9FF6F55C8FCFDF842" blockId="4.[151,1152,1941,2027]" box="[151,778,1972,1997]" pageId="4" pageNumber="225">
<treatmentCitationGroup id="AB1C11D27926FFC9FF6F55C8FCFDF842" box="[151,778,1972,1997]" pageId="4" pageNumber="225">
<taxonomicName id="4C0C4D7F7926FFC9FF6F55C8FECEF842" authorityName="Lanza" authorityYear="1990" box="[151,313,1972,1996]" class="Reptilia" family="Agamidae" genus="Stellio" higherTaxonomySource="GBIF" kingdom="Animalia" order="Squamata" pageId="4" pageNumber="225" phylum="Chordata" rank="species" species="atricollis">
<emphasis id="B978EAEE7926FFC9FF6F55C8FECEF842" box="[151,313,1972,1996]" italics="true" pageId="4" pageNumber="225">Stellio atricollis</emphasis>
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: [part]
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<bibRefCitation id="EF9D4B0D7926FFC9FE7E55C9FDB4F842" author="Peters, W. C. H." box="[390,579,1972,1996]" pageId="4" pageNumber="225" pagination="147 - 150" refId="ref12379" refString="Peters, W. C. H. (1881) Zwei neue von Herrn Major von Mechow wahrend seiner letzten Expedition nach west-Afrika entdeckte Schlangen und eine ubersicht der von ihm mitgebrachten herpetologischen Sammlung. Sitzungsberichte der Gesellschaft Naturforschender Freunde zu Berlin, 1881, 147 - 150." type="journal article" year="1881">Peters (1881: 147)</bibRefCitation>
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<emphasis id="B978EAEE7926FFC9FF6F55AFFEB7F865" box="[151,320,2003,2027]" italics="true" pageId="4" pageNumber="225">Agama atricollis</emphasis>
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: [part]
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),
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<emphasis id="B978EAEE7927FFC8FF6F52EBFE96FF20" box="[151,353,151,174]" italics="true" pageId="5" pageNumber="226">Agama cyanogaster</emphasis>
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: [part]
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<emphasis id="B978EAEE7927FFC8FF6F52C9FE3DFF43" box="[151,458,181,205]" italics="true" pageId="5" pageNumber="226">Acanthocercus cyanocephalus</emphasis>
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: [part]
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Ceríaco
<emphasis id="B978EAEE7927FFC8FD9152CAFD6BFF43" box="[617,668,181,205]" italics="true" pageId="5" pageNumber="226">et al.</emphasis>
(2014: 670
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; 2016: 75; 2018b: 422),
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<bibRefCitation id="EF9D4B0D7927FFC8FBF152CAFAFBFF43" author="Marques, M. P. &amp; Ceriaco, L. M. P. &amp; Blackburn, D. C. &amp; Bauer, A. M." box="[1033,1292,181,205]" pageId="5" pageNumber="226" pagination="1 - 501" refId="ref11862" refString="Marques, M. P., Ceriaco, L. M. P., Blackburn, D. C. &amp; Bauer, A. M. (2018) Diversity and distribution of the amphibians and terrestrial reptiles of Angola: Atlas of historical and bibliographic records (1850 - 2017). Proceedings of the California Academy of Sciences, 65 (Supplement 2), 1 - 501." type="journal article" year="2018">
Marques
<emphasis id="B978EAEE7927FFC8FB9052CAFB6CFF43" box="[1128,1179,181,205]" italics="true" pageId="5" pageNumber="226">et al.</emphasis>
(2018: 27)
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<treatmentCitation id="0AAD10ED7927FFC8FAEE52CAFEC3FF62" author="Wagner, P. &amp; Greenbaum, E. &amp; Bauer, A. M. &amp; Kusamba, C. &amp; Leache, A. D." httpUri="http://treatment.plazi.org/id/03BFD058FF80FF8B14DB8B66BA1BFD01" page="27" pageId="5" pageNumber="226" year="2018">
<bibRefCitation id="EF9D4B0D7927FFC8FAEE52CAFEC3FF62" author="Wagner, P. &amp; Greenbaum, E. &amp; Bauer, A. M. &amp; Kusamba, C. &amp; Leache, A. D." pageId="5" pageNumber="226" pagination="771 - 817" refId="ref12867" refString="Wagner, P., Greenbaum, E., Bauer, A. M., Kusamba, C. &amp; Leache, A. D. (2018) Lifting the blue-headed veil - integrative taxonomy of the Acanthocercus atricollis species complex (Squamata: Agamidae). Journal of Natural History, 52, 771 - 817. https: // doi. org / 10.1080 / 00222933.2018.1435833" type="journal article" year="2018">
Wagner
<emphasis id="B978EAEE7927FFC8FA9252CAFA6AFF43" box="[1386,1437,181,205]" italics="true" pageId="5" pageNumber="226">et al.</emphasis>
(2018: 27)
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<bibRefCitation id="EF9D4B0D7927FFC8FEC652A8FDBDFF62" author="Branch, W. R. &amp; Vaz Pinto, P. &amp; Baptista, N. &amp; Conradie, W." box="[318,586,212,236]" pageId="5" pageNumber="226" pagination="283 - 334" refId="ref9703" refString="Branch, W. R., Vaz Pinto, P., Baptista, N. &amp; Conradie, W. (2019 b) The reptiles of Angola: History, diversity, endemism and hotspots. In: Huntley, B. J., Russo, V., Lages, F. &amp; Ferrand, N. (Eds.), Biodiversity of Angola. Science &amp; Conservation: A Modern Synthesis. Springer, Cham, pp. 283 - 334. https: // doi. org / 10.1007 / 978 - 3 - 030 - 03083 - 4 _ 13" type="book chapter" year="2019">
Branch
<emphasis id="B978EAEE7927FFC8FE7652A9FE36FF62" box="[398,449,212,236]" italics="true" pageId="5" pageNumber="226">et al.</emphasis>
(2019b: 313)
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<emphasis id="B978EAEE7927FFC8FF6F528FFEDAFE85" box="[151,301,243,267]" italics="true" pageId="5" pageNumber="226">Acanthocercus</emphasis>
sp.
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:
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Wagner
<emphasis id="B978EAEE7927FFC8FDF3528FFDC9FE85" box="[523,574,243,267]" italics="true" pageId="5" pageNumber="226">et al.</emphasis>
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), adult male from
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headquarters [-
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a.s.l.
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],
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,
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,
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and
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.
<collectorName id="26F9532A7927FFC8FC8753FDFC2EFE15" box="[895,985,385,411]" pageId="5" pageNumber="226">Vindum</collectorName>
on
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<collectingDate id="EFF6E9D47927FFC8FBFB53FDFB17FE12" box="[1027,1248,385,412]" pageId="5" pageNumber="226" value="2015-09-14">14 September 2015</collectingDate>
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<caption id="DF7366747927FFC8FF6F55BEFBA0F854" ID-DOI="http://doi.org/10.5281/zenodo.6048532" ID-Zenodo-Dep="6048532" box="[151,1111,1986,2010]" httpUri="https://zenodo.org/record/6048532/files/figure.png" pageId="5" pageNumber="226" startId="5.[151,250,1986,2010]" targetBox="[190,1399,444,1960]" targetPageId="5">
<paragraph id="8BB336FC7927FFC8FF6F55BEFBA0F854" blockId="5.[151,1111,1986,2010]" box="[151,1111,1986,2010]" pageId="5" pageNumber="226">
<emphasis id="B978EAEE7927FFC8FF6F55BEFEE4F854" bold="true" box="[151,275,1986,2010]" pageId="5" pageNumber="226">FIGURE 2.</emphasis>
Holotype of
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<emphasis id="B978EAEE7927FFC8FE6155BEFD7FF854" box="[409,648,1986,2010]" italics="true" pageId="5" pageNumber="226">Acanthocercus ceriacoi</emphasis>
</taxonomicName>
<emphasis id="B978EAEE7927FFC8FD7755BEFD28F854" bold="true" box="[655,735,1986,2010]" pageId="5" pageNumber="226">
<taxonomicNameLabel id="A24B57957927FFC8FD7755BEFD28F854" box="[655,735,1986,2010]" pageId="5" pageNumber="226" rank="species">sp. nov.</taxonomicNameLabel>
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(CAS 258430). Photos by Erica Ely.
</paragraph>
</caption>
<caption id="DF7366747924FFCBFF6F54A4FDB8F89B" ID-DOI="http://doi.org/10.5281/zenodo.6048534" ID-Zenodo-Dep="6048534" httpUri="https://zenodo.org/record/6048534/files/figure.png" pageId="6" pageNumber="227" startId="6.[151,250,1752,1776]" targetBox="[255,1333,181,1729]" targetPageId="6">
<paragraph id="8BB336FC7924FFCBFF6F54A4FDB8F89B" blockId="6.[151,1437,1752,1813]" pageId="6" pageNumber="227">
<emphasis id="B978EAEE7924FFCBFF6F54A4FEE2F97E" bold="true" box="[151,277,1752,1776]" pageId="6" pageNumber="227">FIGURE 3.</emphasis>
Life photo of the holotype (CAS 258430), an adult male of
<taxonomicName id="4C0C4D7F7924FFCBFC6C54A4FB71F97E" authority="Marques &amp; Parrinha &amp; Santos &amp; Bandeira &amp; Butler &amp; Sousa &amp; Bauer &amp; Wagner, 2022" authorityName="Marques &amp; Parrinha &amp; Santos &amp; Bandeira &amp; Butler &amp; Sousa &amp; Bauer &amp; Wagner" authorityYear="2022" box="[916,1158,1752,1776]" class="Reptilia" family="Agamidae" genus="Acanthocercus" kingdom="Animalia" order="Squamata" pageId="6" pageNumber="227" phylum="Chordata" rank="species" species="ceriacoi" status="sp. nov.">
<emphasis id="B978EAEE7924FFCBFC6C54A4FB71F97E" box="[916,1158,1752,1776]" italics="true" pageId="6" pageNumber="227">Acanthocercus ceriacoi</emphasis>
</taxonomicName>
<emphasis id="B978EAEE7924FFCBFB7754A5FB16F97F" bold="true" box="[1167,1249,1753,1777]" pageId="6" pageNumber="227">
<taxonomicNameLabel id="A24B57957924FFCBFB7754A5FB16F97F" box="[1167,1249,1753,1777]" pageId="6" pageNumber="227" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
from Cangandala National Park. Photo by Luis M.P. Ceríaco.
</paragraph>
</caption>
<paragraph id="8BB336FC7924FFCAFF3F553FFDB0FE93" blockId="6.[151,1437,1859,2030]" lastBlockId="7.[151,1437,150,285]" lastPageId="7" lastPageNumber="228" pageId="6" pageNumber="227">
<materialsCitation id="3B643CA17924FFCBFF3F553FFD39F828" ID-GBIF-Occurrence="3467090304" collectingDate="2015-09-13" collectionCode="CAS" country="Angola" elevation="1106" latitude="9.818917" location="Cangandala National Park" longLatPrecision="5" longitude="16.6541" municipality="Five" pageId="6" pageNumber="227" specimenCode="CAS 258429" specimenCount="2" stateProvince="Malanje" typeStatus="paratype">
<emphasis id="B978EAEE7924FFCBFF3F553FFEB1F8D3" bold="true" box="[199,326,1859,1885]" pageId="6" pageNumber="227">
<typeStatus id="54B7885E7924FFCBFF3F553FFEB6F8D3" box="[199,321,1859,1885]" pageId="6" pageNumber="227" type="paratype">Paratypes</typeStatus>
.
</emphasis>
<collectingMunicipality id="6BD7AC867924FFCBFEB4553FFE89F8D3" box="[332,382,1859,1885]" pageId="6" pageNumber="227">Five</collectingMunicipality>
specimens, all from
<collectingCountry id="F31B766C7924FFCBFD9C553FFD4CF8D3" box="[612,699,1859,1885]" name="Angola" pageId="6" pageNumber="227">Angola</collectingCountry>
:
<specimenCode id="DBAA9E877924FFCBFD3C553FFCA2F8D0" box="[708,853,1859,1886]" collectionCode="CAS" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:15690" lsid="urn:lsid:biocol.org:col:34700" name="California Academy of Sciences" pageId="6" pageNumber="227" type="Museum">CAS 258429</specimenCode>
(field number JVV 9099) adult male from the vicinity of
<location id="8ED360277924FFCBFF24551BFDF3F80F" LSID="urn:lsid:plazi:treatment:03A587EA7926FFC3FF6F5550FD41FE06:8ED360277924FFCBFF24551BFDF3F80F" box="[220,516,1895,1921]" country="Angola" latitude="9.818917" longLatPrecision="5" longitude="16.6541" municipality="Five" name="Cangandala National Park" pageId="6" pageNumber="227" stateProvince="Malanje">Cangandala National Park</location>
headquarters [-
<geoCoordinate id="EE38503B7924FFCBFD49551BFCBEF80F" box="[689,841,1895,1921]" degrees="9.818917" direction="north" orientation="latitude" pageId="6" pageNumber="227" precision="1" value="9.818917">9.818917° N</geoCoordinate>
,
<geoCoordinate id="EE38503B7924FFCBFCAD551BFC21F80F" box="[853,982,1895,1922]" degrees="16.6541" direction="east" orientation="longitude" pageId="6" pageNumber="227" precision="5" value="16.6541">16.6541° E</geoCoordinate>
,
<elevation id="0021D1CF7924FFCBFC1B551BFB81F80F" box="[995,1142,1894,1921]" metricMagnitude="3" metricUnit="m" metricValue="1.1059999999999999" pageId="6" pageNumber="227" unit="m" value="1106.0">
<quantity id="4CF49B197924FFCBFC1B551BFBCFF80F" box="[995,1080,1895,1921]" metricMagnitude="3" metricUnit="m" metricValue="1.1059999999999999" pageId="6" pageNumber="227" unit="m" value="1106.0">1106 m</quantity>
a.s.l.
</elevation>
],
<collectingRegion id="49C8F81E7924FFCBFB74551BFAA1F80F" box="[1164,1366,1895,1921]" country="Angola" name="Malanje" pageId="6" pageNumber="227">Malanje Province</collectingRegion>
, same collectors as the
<typeStatus id="54B7885E7924FFCBFEA355F7FE4AF82B" box="[347,445,1931,1957]" pageId="6" pageNumber="227" type="holotype">holotype</typeStatus>
on
<date id="FFB2103C7924FFCBFE1455F7FD39F828" box="[492,718,1931,1958]" pageId="6" pageNumber="227" value="2015-09-13">
<collectingDate id="EFF6E9D47924FFCBFE1455F7FD39F828" box="[492,718,1931,1958]" pageId="6" pageNumber="227" value="2015-09-13">13 September 2015</collectingDate>
</date>
</materialsCitation>
;
<materialsCitation id="3B643CA17924FFCBFD2455F7FDC3F847" ID-GBIF-Occurrence="3467090301" collectingDate="2015-09-14" collectionCode="CAS" collectorName="Mariana P. Marques &amp; Luis M. P. Ceriaco &amp; Suzana A. Bandeira &amp; Edward L. Stanley &amp; Jens &amp; Vindum" country="Angola" elevation="1089" latitude="9.819417" location="Cangandala National Park" longLatPrecision="1" longitude="16.65386" pageId="6" pageNumber="227" specimenCode="CAS 258431" specimenCount="1" stateProvince="Malanje" typeStatus="paratype">
<specimenCode id="DBAA9E877924FFCBFD2455F7FC87F828" box="[732,880,1931,1958]" collectionCode="CAS" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:15690" lsid="urn:lsid:biocol.org:col:34700" name="California Academy of Sciences" pageId="6" pageNumber="227" type="Museum">CAS 258431</specimenCode>
(field number JVV 9460) adult female with the same collecting data as the holotype
</materialsCitation>
;
<materialsCitation id="3B643CA17924FFCAFDC755D3FAC3FF3F" ID-GBIF-Occurrence="3467090305" collectingDate="2015-09-18" collectionCode="CAS" country="Angola" elevation="1089" lastPageId="7" lastPageNumber="228" latitude="9.818583" location="Cangandala National Park" longLatPrecision="1" longitude="16.654028" pageId="6" pageNumber="227" specimenCode="CAS 258433, CAS 258434" specimenCount="1" specimenCount-adult="1" stateProvince="Malanje" typeStatus="holotype">
<specimenCode id="DBAA9E877924FFCBFDC755D3FD26F844" box="[575,721,1967,1994]" collectionCode="CAS" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:15690" lsid="urn:lsid:biocol.org:col:34700" name="California Academy of Sciences" pageId="6" pageNumber="227" type="Museum">CAS 258433</specimenCode>
(field number JVV 9261) and
<specimenCode id="DBAA9E877924FFCBFBD655D3FB37F844" box="[1070,1216,1967,1994]" collectionCode="CAS" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:15690" lsid="urn:lsid:biocol.org:col:34700" name="California Academy of Sciences" pageId="6" pageNumber="227" type="Museum">CAS 258434</specimenCode>
(field number JVV 9262),
<specimenCount id="9D0AFD757924FFCBFF1E55AFFEB8F863" box="[230,335,2003,2029]" pageId="6" pageNumber="227" type="adult">two adult</specimenCount>
males, respectively, both from the vicinity of
<location id="8ED360277924FFCBFCAB55AFFB8FF863" LSID="urn:lsid:plazi:treatment:03A587EA7926FFC3FF6F5550FD41FE06:8ED360277924FFCBFCAB55AFFB8FF863" box="[851,1144,2003,2029]" country="Angola" latitude="9.818583" longLatPrecision="1" longitude="16.654028" name="Cangandala National Park" pageId="6" pageNumber="227" stateProvince="Malanje">Cangandala National Park</location>
headquarters [-
<geoCoordinate id="EE38503B7924FFCAFADB55AFFF59FF3F" degrees="9.818583" direction="north" lastPageId="7" lastPageNumber="228" orientation="latitude" pageId="6" pageNumber="227" precision="1" value="9.818583">9.818583° N</geoCoordinate>
,
<geoCoordinate id="EE38503B7925FFCAFF4152EAFEA3FF3F" box="[185,340,150,177]" degrees="16.654028" direction="east" orientation="longitude" pageId="7" pageNumber="228" precision="1" value="16.654028">16.654028° E</geoCoordinate>
,
<elevation id="0021D1CF7925FFCAFEA752EBFE18FF3F" box="[351,495,150,177]" metricMagnitude="3" metricUnit="m" metricValue="1.089" pageId="7" pageNumber="228" unit="m" value="1089.0">
<quantity id="4CF49B197925FFCAFEA752EBFE44FF3F" box="[351,435,151,177]" metricMagnitude="3" metricUnit="m" metricValue="1.089" pageId="7" pageNumber="228" unit="m" value="1089.0">1089 m</quantity>
a.s.l.
</elevation>
],
<collectingRegion id="49C8F81E7925FFCAFDFC52EBFD3BFF3F" box="[516,716,151,177]" country="Angola" name="Malanje" pageId="7" pageNumber="228">Malanje Province</collectingRegion>
, same collectors as the
<typeStatus id="54B7885E7925FFCAFC3752EBFBC6FF3F" box="[975,1073,151,177]" pageId="7" pageNumber="228" type="holotype">holotype</typeStatus>
on
<date id="FFB2103C7925FFCAFBA152EBFAC3FF3F" box="[1113,1332,150,177]" pageId="7" pageNumber="228" value="2015-09-18">
<collectingDate id="EFF6E9D47925FFCAFBA152EBFAC3FF3F" box="[1113,1332,150,177]" pageId="7" pageNumber="228" value="2015-09-18">18 September 2015</collectingDate>
</date>
</materialsCitation>
;
<materialsCitation id="3B643CA17925FFCAFAC752EBFDB4FE93" ID-GBIF-Occurrence="3467090302" collectingDate="2019-11-28" collectionCode="UP" collectorName="Luis M. P. Ceriaco &amp; Gregorius Jongsma &amp; Stuart Nielsen &amp; Ilola Jorge" country="Angola" elevation="1184" latitude="7.207669" location="Zulumongo village" longLatPrecision="1" longitude="15.167342" pageId="7" pageNumber="228" specimenCode="REP 860" specimenCount="1" stateProvince="Uige" typeStatus="paratype">
MHNC-
<collectionCode id="ED1DAE397925FFCAFF6F52C7FF4DFF5B" box="[151,186,187,213]" country="Papua New Guinea" name="University of Papua and New Guinea" pageId="7" pageNumber="228">UP</collectionCode>
/
<specimenCode id="DBAA9E877925FFCAFF3A52C7FEDEFF5B" box="[194,297,187,213]" collectionCode="UP" country="Papua New Guinea" name="University of Papua and New Guinea" pageId="7" pageNumber="228">REP 860</specimenCode>
(field number GJ 04564,
<figureCitation id="13372A797925FFCAFDAE52C7FD33FF5B" box="[598,708,186,213]" captionStart-0="FIGURE 4" captionStart-1="FIGURE 5" captionStartId-0="7.[151,250,1963,1987]" captionStartId-1="8.[151,250,1806,1830]" captionTargetBox-0="[153,1435,319,1939]" captionTargetBox-1="[151,1436,181,1782]" captionTargetId-0="figure-76@7.[151,1435,309,1940]" captionTargetId-1="figure-23@8.[151,1436,181,1782]" captionTargetPageId-0="7" captionTargetPageId-1="8" captionText-0="FIGURE 4. Paratype of Acanthocercus ceriacoi sp. nov. (Paratype, MHNCUP/REP 860), an adult female. Photos by Luis M.P. Ceríaco." captionText-1="FIGURE 5. Life photos of an adult female of Acanthocercus ceriacoi sp. nov. (Paratype, MHNCUP/REP 860) from Uíge Province. Photos by Luis M.P. Ceríaco." figureDoi-0="http://doi.org/10.5281/zenodo.6048538" figureDoi-1="http://doi.org/10.5281/zenodo.6048540" httpUri-0="https://zenodo.org/record/6048538/files/figure.png" httpUri-1="https://zenodo.org/record/6048540/files/figure.png" pageId="7" pageNumber="228">Figs. 45</figureCitation>
), an adult female from near
<location id="8ED360277925FFCAFBF752C7FB10FF5B" LSID="urn:lsid:plazi:treatment:03A587EA7926FFC3FF6F5550FD41FE06:8ED360277925FFCAFBF752C7FB10FF5B" box="[1039,1255,187,213]" country="Angola" latitude="7.207669" longLatPrecision="1" longitude="15.167342" name="Zulumongo village" pageId="7" pageNumber="228" stateProvince="Uige">Zulumongo village</location>
[-
<geoCoordinate id="EE38503B7925FFCAFB0752C7FA6FFF5B" box="[1279,1432,187,213]" degrees="7.207669" direction="north" orientation="latitude" pageId="7" pageNumber="228" precision="1" value="7.207669">7.207669° N</geoCoordinate>
,
<geoCoordinate id="EE38503B7925FFCAFF6F52A2FEC2FF77" box="[151,309,222,249]" degrees="15.167342" direction="east" orientation="longitude" pageId="7" pageNumber="228" precision="1" value="15.167342">15.167342° E</geoCoordinate>
,
<elevation id="0021D1CF7925FFCAFEBB52A3FE20FF77" box="[323,471,222,249]" metricMagnitude="3" metricUnit="m" metricValue="1.184" pageId="7" pageNumber="228" unit="m" value="1184.0">
<quantity id="4CF49B197925FFCAFEBB52A3FE6EFF77" box="[323,409,223,249]" metricMagnitude="3" metricUnit="m" metricValue="1.184" pageId="7" pageNumber="228" unit="m" value="1184.0">1184 m</quantity>
a.s.l.
</elevation>
],
<collectingRegion id="49C8F81E7925FFCAFE1652A3FD63FF77" box="[494,660,223,249]" country="Angola" name="Uige" pageId="7" pageNumber="228">Uíge Province</collectingRegion>
, collected by
<collectorName id="26F9532A7925FFCAFCCB52A3FBF3FF77" box="[819,1028,223,249]" pageId="7" pageNumber="228">Luis M.P. Ceríaco</collectorName>
,
<collectorName id="26F9532A7925FFCAFBE952A3FB19FF77" box="[1041,1262,223,249]" pageId="7" pageNumber="228">Gregorius Jongsma</collectorName>
,
<collectorName id="26F9532A7925FFCAFB0352A3FA6AFF77" box="[1275,1437,223,249]" pageId="7" pageNumber="228">Stuart Nielsen</collectorName>
and
<collectorName id="26F9532A7925FFCAFF3F537FFECAFE93" box="[199,317,259,285]" pageId="7" pageNumber="228">Ilola Jorge</collectorName>
on
<date id="FFB2103C7925FFCAFE9F537FFDB4FE93" box="[359,579,259,285]" pageId="7" pageNumber="228" value="2019-11-28">
<collectingDate id="EFF6E9D47925FFCAFE9F537FFDB4FE93" box="[359,579,259,285]" pageId="7" pageNumber="228" value="2019-11-28">28 November 2019</collectingDate>
</date>
</materialsCitation>
.
</paragraph>
<caption id="DF7366747925FFCAFF6F55D7FF1AF869" ID-DOI="http://doi.org/10.5281/zenodo.6048538" ID-Zenodo-Dep="6048538" httpUri="https://zenodo.org/record/6048538/files/figure.png" pageId="7" pageNumber="228" startId="7.[151,250,1963,1987]" targetBox="[153,1435,319,1939]" targetPageId="7">
<paragraph id="8BB336FC7925FFCAFF6F55D7FF1AF869" blockId="7.[151,1437,1963,2023]" pageId="7" pageNumber="228">
<emphasis id="B978EAEE7925FFCAFF6F55D7FEE5F84D" bold="true" box="[151,274,1963,1987]" pageId="7" pageNumber="228">FIGURE 4.</emphasis>
Paratype of
<taxonomicName id="4C0C4D7F7925FFCAFE6955D7FD77F84D" authority="Marques &amp; Parrinha &amp; Santos &amp; Bandeira &amp; Butler &amp; Sousa &amp; Bauer &amp; Wagner, 2022" authorityName="Marques &amp; Parrinha &amp; Santos &amp; Bandeira &amp; Butler &amp; Sousa &amp; Bauer &amp; Wagner" authorityYear="2022" box="[401,640,1963,1987]" class="Reptilia" family="Agamidae" genus="Acanthocercus" kingdom="Animalia" order="Squamata" pageId="7" pageNumber="228" phylum="Chordata" rank="species" species="ceriacoi" status="sp. nov.">
<emphasis id="B978EAEE7925FFCAFE6955D7FD77F84D" box="[401,640,1963,1987]" italics="true" pageId="7" pageNumber="228">Acanthocercus ceriacoi</emphasis>
</taxonomicName>
<emphasis id="B978EAEE7925FFCAFD7E55D7FD22F84D" bold="true" box="[646,725,1963,1987]" pageId="7" pageNumber="228">
<taxonomicNameLabel id="A24B57957925FFCAFD7E55D7FD22F84D" box="[646,725,1963,1987]" pageId="7" pageNumber="228" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
(Paratype, MHNCUP/REP 860), an adult female. Photos by Luis M.P. Ceríaco.
</paragraph>
</caption>
<caption id="DF736674792AFFC5FF6F5572FDD0F8C5" ID-DOI="http://doi.org/10.5281/zenodo.6048540" ID-Zenodo-Dep="6048540" httpUri="https://zenodo.org/record/6048540/files/figure.png" pageId="8" pageNumber="229" startId="8.[151,250,1806,1830]" targetBox="[151,1436,181,1782]" targetPageId="8">
<paragraph id="8BB336FC792AFFC5FF6F5572FDD0F8C5" blockId="8.[151,1436,1806,1867]" pageId="8" pageNumber="229">
<emphasis id="B978EAEE792AFFC5FF6F5572FEE1F8A9" bold="true" box="[151,278,1806,1831]" pageId="8" pageNumber="229">FIGURE 5.</emphasis>
Life photos of an adult female of
<taxonomicName id="4C0C4D7F792AFFC5FD7E5573FC8FF8A8" authority="Marques &amp; Parrinha &amp; Santos &amp; Bandeira &amp; Butler &amp; Sousa &amp; Bauer &amp; Wagner, 2022" authorityName="Marques &amp; Parrinha &amp; Santos &amp; Bandeira &amp; Butler &amp; Sousa &amp; Bauer &amp; Wagner" authorityYear="2022" box="[646,888,1807,1831]" class="Reptilia" family="Agamidae" genus="Acanthocercus" kingdom="Animalia" order="Squamata" pageId="8" pageNumber="229" phylum="Chordata" rank="species" species="ceriacoi" status="sp. nov.">
<emphasis id="B978EAEE792AFFC5FD7E5573FC8FF8A8" box="[646,888,1807,1831]" italics="true" pageId="8" pageNumber="229">Acanthocercus ceriacoi</emphasis>
</taxonomicName>
<emphasis id="B978EAEE792AFFC5FC795573FC22F8A9" bold="true" box="[897,981,1807,1831]" pageId="8" pageNumber="229">
<taxonomicNameLabel id="A24B5795792AFFC5FC795573FC22F8A9" box="[897,981,1807,1831]" pageId="8" pageNumber="229" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
(Paratype, MHNCUP/REP 860) from Uíge Province. Photos by Luis M.P. Ceríaco.
</paragraph>
</caption>
<paragraph id="8BB336FC792AFFC4FF3F5504FEC6FD27" blockId="8.[151,1436,1912,2012]" lastBlockId="9.[151,1437,150,1329]" lastPageId="9" lastPageNumber="230" pageId="8" pageNumber="229">
<emphasis id="B978EAEE792AFFC5FF3F5504FE4AF81C" bold="true" box="[199,445,1912,1938]" pageId="8" pageNumber="229">Additional material.</emphasis>
<specimenCount id="9D0AFD75792AFFC5FE305505FD90F81D" box="[456,615,1913,1939]" pageId="8" pageNumber="229" type="generic">42 specimens</specimenCount>
:
<emphasis id="B978EAEE792AFFC5FD8D5504FCA2F81C" bold="true" box="[629,853,1912,1938]" pageId="8" pageNumber="229">
<collectingRegion id="49C8F81E792AFFC5FD8D5504FCA6F81C" box="[629,849,1912,1938]" country="Angola" name="Malanje" pageId="8" pageNumber="229">Malanje Province</collectingRegion>
:
</emphasis>
<collectionCode id="ED1DAE39792AFFC5FC985505FC61F81D" box="[864,918,1913,1939]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:15690" lsid="urn:lsid:biocol.org:col:34700" name="California Academy of Sciences" pageId="8" pageNumber="229" type="Museum">CAS</collectionCode>
258428, 258432 258435 (field number JVV 9098, 9218, 9440)
<specimenCount id="9D0AFD75792AFFC5FE9355E1FE15F839" box="[363,482,1949,1975]" pageId="8" pageNumber="229" type="adult">three adult</specimenCount>
males from the same locality and collectors as
<typeStatus id="54B7885E792AFFC5FC0B55E1FBAFF839" box="[1011,1112,1949,1975]" pageId="8" pageNumber="229" type="holotype">holotype</typeStatus>
, on 13, 17 and 22 September 2015, respectively;
<collectionCode id="ED1DAE39792AFFC5FE8C55BDFE5DF855" box="[372,426,1985,2011]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:15690" lsid="urn:lsid:biocol.org:col:34700" name="California Academy of Sciences" pageId="8" pageNumber="229" type="Museum">CAS</collectionCode>
258436 (field number JVV 9298), an unsexed juvenile from
<quantity id="4CF49B19792AFFC5FBA655BDFB3FF855" box="[1118,1224,1985,2011]" metricMagnitude="4" metricUnit="m" metricValue="1.367" pageId="8" pageNumber="229" unit="km" value="13.67">13.67 km</quantity>
southwest by road of Cangandala National Park headquarters [-
<geoCoordinate id="EE38503B792BFFC4FD7E52EBFCEAFF3F" box="[646,797,151,177]" degrees="9.819222" direction="north" orientation="latitude" pageId="9" pageNumber="230" precision="1" value="9.819222">9.819222° N</geoCoordinate>
,
<geoCoordinate id="EE38503B792BFFC4FCD052EAFC32FF3F" box="[808,965,150,177]" degrees="16.654139" direction="east" orientation="longitude" pageId="9" pageNumber="230" precision="1" value="16.654139">16.654139° E</geoCoordinate>
,
<quantity id="4CF49B19792BFFC4FC2952EBFBD1FF3F" box="[977,1062,151,177]" metricMagnitude="3" metricUnit="m" metricValue="1.0859999999999999" pageId="9" pageNumber="230" unit="m" value="1086.0">1086 m</quantity>
a.s.l.], collected by the same collectors as the
<typeStatus id="54B7885E792BFFC4FED552C7FE78FF5B" box="[301,399,187,213]" pageId="9" pageNumber="230" type="holotype">holotype</typeStatus>
on
<date id="FFB2103C792BFFC4FE4F52C7FD65FF5B" box="[439,658,186,213]" pageId="9" pageNumber="230" value="2015-09-20">20 September 2015</date>
;
<collectionCode id="ED1DAE39792BFFC4FD6552C7FCF0FF5B" box="[669,775,187,213]" country="United Kingdom" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34665" lsid="urn:lsid:biocol.org:col:34665" name="Natural History Museum, London" pageId="9" pageNumber="230" type="Museum">NHMUK</collectionCode>
<date id="FFB2103C792BFFC4FCF452C7FC71FF5B" box="[780,902,187,213]" pageId="9" pageNumber="230" value="1866-06-11">1866.6.11.</date>
<specimenCount id="9D0AFD75792BFFC4FC7E52C7FC39FF5B" box="[902,974,187,213]" pageId="9" pageNumber="230" type="adult">1 adult</specimenCount>
male from Duque de Bragança (currently Kalandula) [-
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,
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,
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a.s.l.], collected by Francisco Pinheiro Bayão in 1866 [specimen noted by
<bibRefCitation id="EF9D4B0D792BFFC4FF44537FFE94FE90" author="Bocage, J. V. B." box="[188,355,259,286]" pageId="9" pageNumber="230" pagination="37 - 56" refId="ref9333" refString="Bocage, J. V. B. (1866) Lista de reptis das possessoes potuguezas d'Africa ocidental que existem no Museu de Lisboa. Jornal de Sciencias Mathematicas, Physicas e Naturaes, 1, 37 - 56." type="journal article" year="1866">Bocage (1866)</bibRefCitation>
and
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];
<collectionCode id="ED1DAE39792BFFC4FD78537FFD1DFE93" box="[640,746,259,285]" country="United Kingdom" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34665" lsid="urn:lsid:biocol.org:col:34665" name="Natural History Museum, London" pageId="9" pageNumber="230" type="Museum">NHMUK</collectionCode>
<date id="FFB2103C792BFFC4FD0A537EFCAFFE93" box="[754,856,258,285]" pageId="9" pageNumber="230" value="1904-05-02">1904.5.2.</date>
<specimenCount id="9D0AFD75792BFFC4FCA0537EFC4FFE93" box="[856,952,258,285]" pageId="9" pageNumber="230" type="adult">26 adult</specimenCount>
male from Pungo Andongo [-
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,
<geoCoordinate id="EE38503B792BFFC4FF6F535AFED1FECF" box="[151,294,294,321]" degrees="15.58333" direction="east" orientation="longitude" pageId="9" pageNumber="230" precision="1" value="15.58333">15.58333° E</geoCoordinate>
,
<quantity id="4CF49B19792BFFC4FECC535BFE7DFECF" box="[308,394,295,321]" metricMagnitude="3" metricUnit="m" metricValue="1.22" pageId="9" pageNumber="230" unit="m" value="1220.0">1220 m</quantity>
a.s.l.], collected by William John Ansorge in June/
<date id="FFB2103C792BFFC4FC34535BFBCDFECF" box="[972,1082,295,321]" pageId="9" pageNumber="230" value="1903-07">July 1903</date>
[specimen noted by
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];
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<date id="FFB2103C792BFFC4FE925336FE26FEEB" box="[362,465,330,357]" pageId="9" pageNumber="230" value="1904-05-02">1904.5.2.</date>
<specimenCount id="9D0AFD75792BFFC4FE285336FDDAFEEB" box="[464,557,330,357]" pageId="9" pageNumber="230" type="adult">25 adult</specimenCount>
male from Bange NGola [-
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,
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,
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a.s.l.], collected by William John Ansorge in the end of 1903 [specimen noted by
<bibRefCitation id="EF9D4B0D792BFFC4FC9B5313FBD9FE07" author="Boulenger, G. A." box="[867,1070,366,393]" pageId="9" pageNumber="230" pagination="105 - 115" refId="ref9509" refString="Boulenger, G. A. (1905) A list of the batrachians and reptiles collected by Dr. W. J. Ansorge in Angola, with descriptions of new species. Annals and Magazine of Natural History, Series 7 (16), 105 - 115, pl. 4. https: // doi. org / 10.1080 / 03745480509443656" type="journal article" year="1905">Boulenger (1905)</bibRefCitation>
];
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1904.5.2.1724
<specimenCount id="9D0AFD75792BFFC4FA9C5313FEFBFE23" pageId="9" pageNumber="230" type="generic">eight specimens</specimenCount>
from Quanza River (most likely Duque de Bragança), collected by William John Ansorge at the end of 1903 [specimen noted by
<bibRefCitation id="EF9D4B0D792BFFC4FE4753CBFD7BFE5F" author="Boulenger, G. A." box="[447,652,438,465]" pageId="9" pageNumber="230" pagination="105 - 115" refId="ref9509" refString="Boulenger, G. A. (1905) A list of the batrachians and reptiles collected by Dr. W. J. Ansorge in Angola, with descriptions of new species. Annals and Magazine of Natural History, Series 7 (16), 105 - 115, pl. 4. https: // doi. org / 10.1080 / 03745480509443656" type="journal article" year="1905">Boulenger (1905)</bibRefCitation>
];
<collectionCode id="ED1DAE39792BFFC4FD5953CBFD29FE5F" box="[673,734,439,465]" country="Germany" httpUri="http://grbio.org/cool/8syf-d21i" name="Museum für Naturkunde Berlin (Zoological Collections)" pageId="9" pageNumber="230" type="Museum">ZMB</collectionCode>
10027 adult male from
<collectingRegion id="49C8F81E792BFFC4FC0B53CBFBB8FE5F" box="[1011,1103,439,465]" country="Angola" name="Malanje" pageId="9" pageNumber="230">Malanje</collectingRegion>
[-
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,
<geoCoordinate id="EE38503B792BFFC4FB2B53CAFACEFE5F" box="[1235,1337,438,465]" degrees="16.35" direction="east" orientation="longitude" pageId="9" pageNumber="230" precision="555" value="16.35">16.35° E</geoCoordinate>
,
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a.s.l.], collected by Friedrich Wilhelm Alexander von Mechow between 1879 and 1881 [specimen noted by
<bibRefCitation id="EF9D4B0D792BFFC4FAA053A7FF10FD97" author="Peters, W. C. H." pageId="9" pageNumber="230" pagination="147 - 150" refId="ref12379" refString="Peters, W. C. H. (1881) Zwei neue von Herrn Major von Mechow wahrend seiner letzten Expedition nach west-Afrika entdeckte Schlangen und eine ubersicht der von ihm mitgebrachten herpetologischen Sammlung. Sitzungsberichte der Gesellschaft Naturforschender Freunde zu Berlin, 1881, 147 - 150." type="journal article" year="1881">Peters (1881)</bibRefCitation>
];
<collectionCode id="ED1DAE39792BFFC4FF035383FE95FD97" box="[251,354,511,537]" pageId="9" pageNumber="230">MNHNL</collectionCode>
/Rep/A/Ag 108126,
<specimenCount id="9D0AFD75792BFFC4FDAE5382FD18FD97" box="[598,751,510,537]" pageId="9" pageNumber="230" type="generic">25 specimens</specimenCount>
from Capanda Dam area [-
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,
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,
<quantity id="4CF49B19792BFFC4FAAC5382FA6BFD97" box="[1364,1436,510,537]" metricMagnitude="2" metricUnit="m" metricValue="8.59" pageId="9" pageNumber="230" unit="m" value="859.0">859 m</quantity>
a.s.l.], collected by Ana Lavres between
<date id="FFB2103C792BFFC4FDA3505FFC96FDB3" box="[603,865,547,573]" pageId="9" pageNumber="230" value="2003-01" valueMax="2003-04-31" valueMin="2003-01-00">January and April 2003</date>
[specimens noted by
<bibRefCitation id="EF9D4B0D792BFFC4FBAC505FFAB7FDB3" author="Ceriaco, L. M. P. &amp; Bauer, A. M. &amp; Blackburn, D. C. &amp; Lavres, A. C. F. C." box="[1108,1344,546,573]" pageId="9" pageNumber="230" pagination="667 - 674" refId="ref9975" refString="Ceriaco, L. M. P., Bauer, A. M., Blackburn, D. C. &amp; Lavres, A. C. F. C. (2014) The herpetofauna of the Capanda Dam region, Malanje, Angola. Herpetological Review, 45 (4), 667 - 674." type="journal article" year="2014">
Ceríaco
<emphasis id="B978EAEE792BFFC4FB49505FFB1EFDB3" box="[1201,1257,547,573]" italics="true" pageId="9" pageNumber="230">et al.</emphasis>
(2014)
</bibRefCitation>
].
<emphasis id="B978EAEE792BFFC4FAAA505FFEFDFDEE" bold="true" pageId="9" pageNumber="230">
<collectingRegion id="49C8F81E792BFFC4FAAA505FFEF2FDEE" country="Angola" name="Bengo" pageId="9" pageNumber="230">Bengo Province</collectingRegion>
:
</emphasis>
<collectionCode id="ED1DAE39792BFFC4FEF7503BFE92FDEF" box="[271,357,583,609]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34925" lsid="urn:lsid:biocol.org:col:34925" name="American Museum of Natural History" pageId="9" pageNumber="230">AMNH</collectionCode>
<collectionCode id="ED1DAE39792BFFC4FE94503BFE88FDEF" box="[364,383,583,609]" country="Chile" name="Departamento de Geologia, Universidad de Chile" pageId="9" pageNumber="230">R</collectionCode>
48179 adult female from Dande [-
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,
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,
<quantity id="4CF49B19792BFFC4FBC4503AFB82FDEF" box="[1084,1141,582,609]" metricMagnitude="1" metricUnit="m" metricValue="4.5" pageId="9" pageNumber="230" unit="m" value="45.0">45 m</quantity>
a.s.l.], collected by Rudyerd Boulton and Herbert Lang (Vernay-Lang expedition to
<collectingCountry id="F31B766C792BFFC4FCD75017FC74FD08" box="[815,899,619,646]" name="Angola" pageId="9" pageNumber="230">Angola</collectingCountry>
) on
<date id="FFB2103C792BFFC4FC4E5017FBACFD0B" box="[950,1115,618,645]" pageId="9" pageNumber="230" value="1925-08-04">4 August 1925</date>
[specimen noted by
<bibRefCitation id="EF9D4B0D792BFFC4FABE5017FED1FD27" author="Wagner, P. &amp; Greenbaum, E. &amp; Bauer, A. M. &amp; Kusamba, C. &amp; Leache, A. D." pageId="9" pageNumber="230" pagination="771 - 817" refId="ref12867" refString="Wagner, P., Greenbaum, E., Bauer, A. M., Kusamba, C. &amp; Leache, A. D. (2018) Lifting the blue-headed veil - integrative taxonomy of the Acanthocercus atricollis species complex (Squamata: Agamidae). Journal of Natural History, 52, 771 - 817. https: // doi. org / 10.1080 / 00222933.2018.1435833" type="journal article" year="2018">
Wagner
<emphasis id="B978EAEE792BFFC4FF6F50F3FF38FD27" box="[151,207,655,681]" italics="true" pageId="9" pageNumber="230">et al.</emphasis>
(2018)
</bibRefCitation>
].
</paragraph>
<paragraph id="8BB336FC792BFFC4FF3F50CFFE5AFCD3" blockId="9.[151,1437,150,1329]" pageId="9" pageNumber="230">
The records from Cassange [-
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,
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,
<quantity id="4CF49B19792BFFC4FC9F50CEFC47FD43" box="[871,944,690,717]" metricMagnitude="2" metricUnit="m" metricValue="9.55" pageId="9" pageNumber="230" unit="m" value="955.0">955 m</quantity>
a.s.l.],
<collectingRegion id="49C8F81E792BFFC4FBFD50CFFB38FD43" box="[1029,1231,691,717]" country="Angola" name="Malanje" pageId="9" pageNumber="230">Malanje Province</collectingRegion>
, noted by
<bibRefCitation id="EF9D4B0D792BFFC4FAB050CFFF13FD7F" author="Bocage, J. V. B." pageId="9" pageNumber="230" refId="ref9421" refString="Bocage, J. V. B. (1895) Herpetologie d'Angola et du Congo. Ministerio da Marinha e das Colonias, Lisbonne, 203 pp., 20 pls." type="book" year="1895">Bocage (1895)</bibRefCitation>
, as well as those from Duque de
<collectingRegion id="49C8F81E792BFFC4FDB550ABFD40FD7F" box="[589,695,727,753]" country="Portugal" name="Braganca" pageId="9" pageNumber="230">Bragança</collectingRegion>
[-
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,
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,
<quantity id="4CF49B19792BFFC4FC6A50ABFC11FD7F" box="[914,998,727,753]" metricMagnitude="3" metricUnit="m" metricValue="1.01" pageId="9" pageNumber="230" unit="m" value="1010.0">1010 m</quantity>
a.s.l.],
<collectingRegion id="49C8F81E792BFFC4FBCE50ABFB09FD7F" box="[1078,1278,727,753]" country="Angola" name="Malanje" pageId="9" pageNumber="230">Malanje Province</collectingRegion>
, noted by
<bibRefCitation id="EF9D4B0D792BFFC4FA9650ABFEEAFC9B" author="Ferreira, J. B." pageId="9" pageNumber="230" pagination="9 - 16" refId="ref11027" refString="Ferreira, J. B. (1903) Reptis de Angola da regiao norte do Quanza da colleccao Pereira do Nascimento (1902). Jornal de Sciencias Mathematicas, Physicas e Naturaes, Segunda Serie, 7, 9 - 16." type="journal article" year="1903">Ferreira (1903)</bibRefCitation>
cannot be confirmed, as these specimens were destroyed in the fire that engulfed the
<collectingRegion id="49C8F81E792BFFC4FB1D5087FAC4FC9B" box="[1253,1331,763,789]" country="Portugal" name="Lisboa" pageId="9" pageNumber="230">Lisbon</collectingRegion>
Museum in 1978. However, due to their geographic proximity and similar habitat, they are here tentatively identified as the newly described species.
</paragraph>
</subSubSection>
<subSubSection id="C3166577792BFFC4FF3F511AFDBEFB67" pageId="9" pageNumber="230" type="diagnosis">
<paragraph id="8BB336FC792BFFC4FF3F511AFDBEFB67" blockId="9.[151,1437,150,1329]" pageId="9" pageNumber="230">
<emphasis id="B978EAEE792BFFC4FF3F511AFEB5FC0E" bold="true" box="[199,322,870,896]" pageId="9" pageNumber="230">Diagnosis.</emphasis>
Segmentation of the tail into distinct whorls places the new species in the genus
<emphasis id="B978EAEE792BFFC4FB08511BFA6AFC0F" box="[1264,1437,871,897]" italics="true" pageId="9" pageNumber="230">
<taxonomicName id="4C0C4D7F792BFFC4FB08511BFA6EFC0F" authorityName="Fitzinger" authorityYear="1843" box="[1264,1433,871,897]" class="Reptilia" family="Agamidae" genus="Acanthocercus" kingdom="Animalia" order="Squamata" pageId="9" pageNumber="230" phylum="Chordata" rank="genus">Acanthocercus</taxonomicName>
.
</emphasis>
Within this genus, heterogeneous body scalation refers it to the
<taxonomicName id="4C0C4D7F792BFFC4FCA251F7FC14FC2A" authorityName="MVZ" baseAuthorityName="Ruppell" baseAuthorityYear="1835" box="[858,995,907,932]" class="Reptilia" family="Agamidae" genus="Agama" kingdom="Animalia" order="Squamata" pageId="9" pageNumber="230" phylum="Chordata" rank="species" species="cyanogaster">
<emphasis id="B978EAEE792BFFC4FCA251F7FC14FC2A" box="[858,995,907,932]" italics="true" pageId="9" pageNumber="230">cyanogaster</emphasis>
</taxonomicName>
/
<taxonomicName id="4C0C4D7F792BFFC4FC1251F7FBB8FC2B" authorityName="Smith" authorityYear="1849" box="[1002,1103,907,933]" class="Reptilia" family="Agamidae" genus="Agama" kingdom="Animalia" order="Squamata" pageId="9" pageNumber="230" phylum="Chordata" rank="species" species="atricollis">
<emphasis id="B978EAEE792BFFC4FC1251F7FBB8FC2B" box="[1002,1103,907,933]" italics="true" pageId="9" pageNumber="230">atricollis</emphasis>
</taxonomicName>
group and the high density of enlarged trunk scales, in combination with a black patch on the shoulder, place it in the
<taxonomicName id="4C0C4D7F792BFFC4FB9A51D3FB12FC47" authorityName="Smith" authorityYear="1849" box="[1122,1253,943,969]" class="Reptilia" family="Agamidae" genus="Agama" kingdom="Animalia" order="Squamata" pageId="9" pageNumber="230" phylum="Chordata" rank="species" species="atricollis">
<emphasis id="B978EAEE792BFFC4FB9A51D3FB12FC47" box="[1122,1253,943,969]" italics="true" pageId="9" pageNumber="230">A. atricollis</emphasis>
</taxonomicName>
complex.
<taxonomicName id="4C0C4D7F792BFFC4FAA151D3FE9FFC62" authority="Marques &amp; Parrinha &amp; Santos &amp; Bandeira &amp; Butler &amp; Sousa &amp; Bauer &amp; Wagner, 2022" authorityName="Marques &amp; Parrinha &amp; Santos &amp; Bandeira &amp; Butler &amp; Sousa &amp; Bauer &amp; Wagner" authorityYear="2022" class="Reptilia" family="Agamidae" genus="Acanthocercus" kingdom="Animalia" order="Squamata" pageId="9" pageNumber="230" phylum="Chordata" rank="species" species="ceriacoi" status="sp. nov.">
<emphasis id="B978EAEE792BFFC4FAA151D3FE9FFC62" italics="true" pageId="9" pageNumber="230">Acanthocercus ceriacoi</emphasis>
</taxonomicName>
<emphasis id="B978EAEE792BFFC4FE8851AFFE3CFC63" bold="true" box="[368,459,979,1005]" pageId="9" pageNumber="230">
<taxonomicNameLabel id="A24B5795792BFFC4FE8851AFFE3CFC63" box="[368,459,979,1005]" pageId="9" pageNumber="230" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
is a medium sized lizard (mean SVL
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[
<quantity id="4CF49B19792BFFC4FBF851AFFB67FC63" box="[1024,1168,978,1005]" metricMagnitude="-1" metricUnit="m" metricValue="1.21" metricValueMax="1.43" metricValueMin="0.99" pageId="9" pageNumber="230" unit="mm" value="121.0" valueMax="143.0" valueMin="99.0">99143 mm</quantity>
]; mean total length
<quantity id="4CF49B19792BFFC4FA8A51AEFF34FB9F" metricMagnitude="-1" metricUnit="m" metricValue="2.85" pageId="9" pageNumber="230" unit="mm" value="285.0">285 mm</quantity>
[
<quantity id="4CF49B19792BFFC4FF28518BFE9AFB9F" box="[208,365,1014,1041]" metricMagnitude="-1" metricUnit="m" metricValue="2.77" metricValueMax="3.3" metricValueMin="2.2399999999999998" pageId="9" pageNumber="230" unit="mm" value="277.0" valueMax="330.0" valueMin="224.0">224330 mm</quantity>
]) for its genus and overall is very similar in morphology to
<emphasis id="B978EAEE792BFFC4FBF5518BFB2DFB9F" box="[1037,1242,1015,1041]" italics="true" pageId="9" pageNumber="230">
<taxonomicName id="4C0C4D7F792BFFC4FBF5518BFB21FB9F" authorityName="HF" baseAuthorityName="Falk" baseAuthorityYear="1925" box="[1037,1238,1015,1041]" class="Reptilia" family="Agamidae" genus="Acanthocercus" kingdom="Animalia" order="Squamata" pageId="9" pageNumber="230" phylum="Chordata" rank="species" species="cyanocephalus">A. cyanocephalus</taxonomicName>
.
</emphasis>
However, it may be distinguished from all other species of the complex by its meristic characters. The new species has lower scale counts when compared to other species of the
<taxonomicName id="4C0C4D7F792BFFC4FD5A5643FCD0FBD7" authorityName="Smith" authorityYear="1849" box="[674,807,1087,1113]" class="Reptilia" family="Agamidae" genus="Agama" kingdom="Animalia" order="Squamata" pageId="9" pageNumber="230" phylum="Chordata" rank="species" species="atricollis">
<emphasis id="B978EAEE792BFFC4FD5A5643FCD0FBD7" box="[674,807,1087,1113]" italics="true" pageId="9" pageNumber="230">A. atricollis</emphasis>
</taxonomicName>
complex group (mean scale rows around midbody 108, mean dorsal scale rows 64, mean ventral scale rows 85). In life, displaying males are characterized by a blue coloration of the head; neck, shoulders, body and limbs brownish with a pattern of white and black stripes and speckles; and a pale vertebral band from the neck to the tail, distinct anteriorly between the flanks, becoming more indistinct along a brown background posteriorly.
</paragraph>
</subSubSection>
<subSubSection id="C3166577792BFFC7FF3F568EFEE5F81B" lastPageId="10" lastPageNumber="231" pageId="9" pageNumber="230" type="discussion">
<paragraph id="8BB336FC792BFFC4FF3F568EFB44FABF" blockId="9.[151,1437,150,1329]" pageId="9" pageNumber="230">
<emphasis id="B978EAEE792BFFC4FF3F568EFC7AFA82" bold="true" box="[199,909,1266,1293]" pageId="9" pageNumber="230">
Comparison with other close related
<taxonomicName id="4C0C4D7F792BFFC4FD78568EFC7AFA82" box="[640,909,1266,1293]" class="Reptilia" family="Agamidae" genus="Acanthocercus" kingdom="Animalia" order="Squamata" pageId="9" pageNumber="230" phylum="Chordata" rank="species" species="undetermined">
<emphasis id="B978EAEE792BFFC4FD78568EFCDAFA83" bold="true" box="[640,813,1266,1293]" italics="true" pageId="9" pageNumber="230">Acanthocercus</emphasis>
species.
</taxonomicName>
</emphasis>
<taxonomicName id="4C0C4D7F792BFFC4FC6D568FFB6AFA82" authority="Marques &amp; Parrinha &amp; Santos &amp; Bandeira &amp; Butler &amp; Sousa &amp; Bauer &amp; Wagner, 2022" authorityName="Marques &amp; Parrinha &amp; Santos &amp; Bandeira &amp; Butler &amp; Sousa &amp; Bauer &amp; Wagner" authorityYear="2022" box="[917,1181,1267,1293]" class="Reptilia" family="Agamidae" genus="Acanthocercus" kingdom="Animalia" order="Squamata" pageId="9" pageNumber="230" phylum="Chordata" rank="species" species="ceriacoi" status="sp. nov.">
<emphasis id="B978EAEE792BFFC4FC6D568FFB6AFA82" box="[917,1181,1267,1293]" italics="true" pageId="9" pageNumber="230">Acanthocercus ceriacoi</emphasis>
</taxonomicName>
<emphasis id="B978EAEE792BFFC4FB5C568FFB09FA83" bold="true" box="[1188,1278,1267,1293]" pageId="9" pageNumber="230">
<taxonomicNameLabel id="A24B5795792BFFC4FB5C568FFB09FA83" box="[1188,1278,1267,1293]" pageId="9" pageNumber="230" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
can be distinguished from other taxa of the
<taxonomicName id="4C0C4D7F792BFFC4FE08576BFD82FABF" authorityName="Smith" authorityYear="1849" box="[496,629,1303,1329]" class="Reptilia" family="Agamidae" genus="Agama" kingdom="Animalia" order="Squamata" pageId="9" pageNumber="230" phylum="Chordata" rank="species" species="atricollis">
<emphasis id="B978EAEE792BFFC4FE08576BFD82FABF" box="[496,629,1303,1329]" italics="true" pageId="9" pageNumber="230">A. atricollis</emphasis>
</taxonomicName>
complex by the following characters (see
<tableCitation id="C68E0347792BFFC4FBB7576BFB52FABF" box="[1103,1189,1303,1329]" captionStart="TABLE 2" captionStartId="9.[151,239,1376,1400]" captionTargetPageId="9" captionText="TABLE 2. Data from all taxa (except A. ceriacoi sp. nov.) from Wagner et al. (2021). Data presented as: minmax (mean). Abbreviations: Dorsal scale rows at midbody (SaM) Dorsal scale numbers (D); Ventral scale numbers (V); Total number of precloacal pores (PP)." httpUri="http://table.plazi.org/id/DF736674792BFFC4FF6F571CFE00FA4F" pageId="9" pageNumber="230" tableUuid="DF736674792BFFC4FF6F571CFE00FA4F">Table 2</tableCitation>
):
</paragraph>
<caption id="DF736674792BFFC4FF6F571CFE00FA4F" ID-Table-UUID="DF736674792BFFC4FF6F571CFE00FA4F" httpUri="http://table.plazi.org/id/DF736674792BFFC4FF6F571CFE00FA4F" pageId="9" pageNumber="230" startId="9.[151,239,1376,1400]" targetBox="[166,1421,1488,2023]" targetIsTable="true" targetPageId="9">
<paragraph id="8BB336FC792BFFC4FF6F571CFE00FA4F" blockId="9.[151,1437,1376,1473]" pageId="9" pageNumber="230">
<emphasis id="B978EAEE792BFFC4FF6F571CFEFAFAF6" bold="true" box="[151,269,1376,1400]" pageId="9" pageNumber="230">TABLE 2.</emphasis>
Data from all taxa (except
<taxonomicName id="4C0C4D7F792BFFC4FDBC571DFD4BFAF6" authority="Marques &amp; Parrinha &amp; Santos &amp; Bandeira &amp; Butler &amp; Sousa &amp; Bauer &amp; Wagner, 2022" authorityName="Marques &amp; Parrinha &amp; Santos &amp; Bandeira &amp; Butler &amp; Sousa &amp; Bauer &amp; Wagner" authorityYear="2022" box="[580,700,1377,1400]" class="Reptilia" family="Agamidae" genus="Acanthocercus" kingdom="Animalia" order="Squamata" pageId="9" pageNumber="230" phylum="Chordata" rank="species" species="ceriacoi" status="sp. nov.">
<emphasis id="B978EAEE792BFFC4FDBC571DFD4BFAF6" box="[580,700,1377,1400]" italics="true" pageId="9" pageNumber="230">A. ceriacoi</emphasis>
</taxonomicName>
<emphasis id="B978EAEE792BFFC4FD3E571DFCE8FAF7" bold="true" box="[710,799,1377,1401]" pageId="9" pageNumber="230">
<taxonomicNameLabel id="A24B5795792BFFC4FD3E571DFCE8FAF7" box="[710,799,1377,1401]" pageId="9" pageNumber="230" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
) from
<bibRefCitation id="EF9D4B0D792BFFC4FC96571DFBA0FAF7" author="Wagner, P. &amp; Butler, B. &amp; Ceriaco, L. M. P. &amp; Bauer, A. M." box="[878,1111,1376,1401]" pageId="9" pageNumber="230" pagination="449 - 463" refId="ref12938" refString="Wagner, P., Butler, B., Ceriaco, L. M. P. &amp; Bauer, A. M. (2021) A new species of the Acanthocercus atricollis (Smith, 1849) complex (Squamata: Agamidae). Salamandra, 57 (4), 449 - 463." type="journal article" year="2021">
Wagner
<emphasis id="B978EAEE792BFFC4FC32571DFBF4FAF6" box="[970,1027,1376,1400]" italics="true" pageId="9" pageNumber="230">et al.</emphasis>
(2021)
</bibRefCitation>
. Data presented as: minmax (mean). Abbreviations: Dorsal scale rows at midbody (SaM) Dorsal scale numbers (D); Ventral scale numbers (V); Total number of precloacal pores (PP).
</paragraph>
</caption>
<paragraph id="8BB336FC792BFFC4FE6357ACFAA7F869" pageId="9" pageNumber="230">
<table id="F90CC45C792B0032FF5E57ACFA7AF869" box="[166,1421,1488,2023]" gridcols="7" gridrows="11" pageId="9" pageNumber="230">
<tr id="353C34BE792B0032FF5E57ACFA7AFA66" box="[166,1421,1488,1512]" gridrow="0" pageId="9" pageNumber="230" rowspan-0="1">
<th id="76ED5DC2792B0032FE6357ACFDC6FA66" box="[411,561,1488,1512]" gridcol="1" gridrow="0" pageId="9" pageNumber="230">
<emphasis id="B978EAEE792BFFC4FE6357ACFE0CFA66" bold="true" box="[411,507,1488,1512]" pageId="9" pageNumber="230">SVL mm</emphasis>
</th>
<th id="76ED5DC2792B0032FDAB57ACFD1EFA66" box="[595,745,1488,1512]" gridcol="2" gridrow="0" pageId="9" pageNumber="230">
<emphasis id="B978EAEE792BFFC4FDAB57ACFD2EFA66" bold="true" box="[595,729,1488,1512]" pageId="9" pageNumber="230">TOTAL mm</emphasis>
</th>
<th id="76ED5DC2792B0032FCF457ACFC54FA66" box="[780,931,1488,1512]" gridcol="3" gridrow="0" pageId="9" pageNumber="230">
<emphasis id="B978EAEE792BFFC4FCF457ACFCC8FA66" bold="true" box="[780,831,1488,1512]" pageId="9" pageNumber="230">SaM</emphasis>
</th>
<th id="76ED5DC2792B0032FC3F57ACFBA7FA66" box="[967,1104,1488,1512]" gridcol="4" gridrow="0" pageId="9" pageNumber="230">
<emphasis id="B978EAEE792BFFC4FC3F57ACFC2EFA66" bold="true" box="[967,985,1488,1512]" pageId="9" pageNumber="230">D</emphasis>
</th>
<th id="76ED5DC2792B0032FB9557ACFB01FA66" box="[1133,1270,1488,1512]" gridcol="5" gridrow="0" pageId="9" pageNumber="230">
<emphasis id="B978EAEE792BFFC4FB9557ACFB88FA66" bold="true" box="[1133,1151,1488,1512]" pageId="9" pageNumber="230">V</emphasis>
</th>
<th id="76ED5DC2792B0032FAE057ACFA7AFA66" box="[1304,1421,1488,1512]" gridcol="6" gridrow="0" pageId="9" pageNumber="230">
<emphasis id="B978EAEE792BFFC4FAE057ACFAC0FA66" bold="true" box="[1304,1335,1488,1512]" pageId="9" pageNumber="230">PP</emphasis>
</th>
</tr>
<tr id="353C34BE792B0032FF5E5785FA7AF99F" box="[166,1421,1529,1553]" gridrow="1" pageId="9" pageNumber="230">
<th id="76ED5DC2792B0032FF5E5785FE8AF99F" box="[166,381,1529,1553]" gridcol="0" gridrow="1" pageId="9" pageNumber="230">
<taxonomicName id="4C0C4D7F792BFFC4FF5E5785FEE9F99F" authorityName="Smith" authorityYear="1849" box="[166,286,1529,1553]" class="Reptilia" family="Agamidae" genus="Agama" kingdom="Animalia" order="Squamata" pageId="9" pageNumber="230" phylum="Chordata" rank="species" species="atricollis">
<emphasis id="B978EAEE792BFFC4FF5E5785FEE9F99F" box="[166,286,1529,1553]" italics="true" pageId="9" pageNumber="230">A. atricollis</emphasis>
</taxonomicName>
(n=21)
</th>
<td id="76ED5DC2792B0032FE635785FDC6F99F" box="[411,561,1529,1553]" gridcol="1" gridrow="1" pageId="9" pageNumber="230">107151 (129)</td>
<td id="76ED5DC2792B0032FDAB5785FD1EF99F" box="[595,745,1529,1553]" gridcol="2" gridrow="1" pageId="9" pageNumber="230">248347 (302)</td>
<td id="76ED5DC2792B0032FCF45785FC54F99F" box="[780,931,1529,1553]" gridcol="3" gridrow="1" pageId="9" pageNumber="230">103125 (112)</td>
<td id="76ED5DC2792B0032FC3F5785FBA7F99F" box="[967,1104,1529,1553]" gridcol="4" gridrow="1" pageId="9" pageNumber="230">5283 (67)</td>
<td id="76ED5DC2792B0032FB955785FB01F99F" box="[1133,1270,1529,1553]" gridcol="5" gridrow="1" pageId="9" pageNumber="230">76 102 (87)</td>
<td id="76ED5DC2792B0032FAE05785FA7AF99F" box="[1304,1421,1529,1553]" gridcol="6" gridrow="1" pageId="9" pageNumber="230">935 (22)</td>
</tr>
<tr id="353C34BE792B0032FF5E545DFA7AF9B7" box="[166,1421,1569,1593]" gridrow="2" pageId="9" pageNumber="230">
<th id="76ED5DC2792B0032FF5E545DFE8AF9B7" box="[166,381,1569,1593]" gridcol="0" gridrow="2" pageId="9" pageNumber="230">
<taxonomicName id="4C0C4D7F792BFFC4FF5E545EFEE6F9B7" box="[166,273,1569,1593]" class="Reptilia" family="Agamidae" genus="Acanthocercus" kingdom="Animalia" order="Squamata" pageId="9" pageNumber="230" phylum="Chordata" rank="species" species="branchi">
<emphasis id="B978EAEE792BFFC4FF5E545EFEE6F9B7" box="[166,273,1569,1593]" italics="true" pageId="9" pageNumber="230">A. branchi</emphasis>
</taxonomicName>
(n=6)
</th>
<td id="76ED5DC2792B0032FE63545DFDC6F9B7" box="[411,561,1569,1593]" gridcol="1" gridrow="2" pageId="9" pageNumber="230">83131 (109)</td>
<td id="76ED5DC2792B0032FDAB545DFD1EF9B7" box="[595,745,1569,1593]" gridcol="2" gridrow="2" pageId="9" pageNumber="230">194313 (262)</td>
<td id="76ED5DC2792B0032FCF4545DFC54F9B7" box="[780,931,1569,1593]" gridcol="3" gridrow="2" pageId="9" pageNumber="230">103130 (117)</td>
<td id="76ED5DC2792B0032FC3F545DFBA7F9B7" box="[967,1104,1569,1593]" gridcol="4" gridrow="2" pageId="9" pageNumber="230">6781 (73)</td>
<td id="76ED5DC2792B0032FB95545DFB01F9B7" box="[1133,1270,1569,1593]" gridcol="5" gridrow="2" pageId="9" pageNumber="230">7995 (88)</td>
<td id="76ED5DC2792B0032FAE0545DFA7AF9B7" box="[1304,1421,1569,1593]" gridcol="6" gridrow="2" pageId="9" pageNumber="230">1723 (20)</td>
</tr>
<tr id="353C34BE792B0032FF5E5435FA7AF908" box="[166,1421,1609,1670]" gridrow="3" pageId="9" pageNumber="230">
<th id="76ED5DC2792B0032FF5E5435FE8AF908" box="[166,381,1609,1670]" gridcol="0" gridrow="3" pageId="9" pageNumber="230">
<taxonomicName id="4C0C4D7F792BFFC4FF5E5436FEAEF9EF" authorityName="HF" baseAuthorityName="Falk" baseAuthorityYear="1925" box="[166,345,1609,1633]" class="Reptilia" family="Agamidae" genus="Acanthocercus" kingdom="Animalia" order="Squamata" pageId="9" pageNumber="230" phylum="Chordata" rank="species" species="cyanocephalus">
<emphasis id="B978EAEE792BFFC4FF5E5436FEAEF9EF" box="[166,345,1609,1633]" italics="true" pageId="9" pageNumber="230">A. cyanocephalus</emphasis>
</taxonomicName>
(n=13)
</th>
<td id="76ED5DC2792B0032FE635435FDC6F908" box="[411,561,1609,1670]" gridcol="1" gridrow="3" pageId="9" pageNumber="230">113149 (138)</td>
<td id="76ED5DC2792B0032FDAB5435FD1EF908" box="[595,745,1609,1670]" gridcol="2" gridrow="3" pageId="9" pageNumber="230">276372 (334)</td>
<td id="76ED5DC2792B0032FCF45435FC54F908" box="[780,931,1609,1670]" gridcol="3" gridrow="3" pageId="9" pageNumber="230">96119 (108)</td>
<td id="76ED5DC2792B0032FC3F5435FBA7F908" box="[967,1104,1609,1670]" gridcol="4" gridrow="3" pageId="9" pageNumber="230">5878 (68)</td>
<td id="76ED5DC2792B0032FB955435FB01F908" box="[1133,1270,1609,1670]" gridcol="5" gridrow="3" pageId="9" pageNumber="230">6994 (80)</td>
<td id="76ED5DC2792B0032FAE05435FA7AF908" box="[1304,1421,1609,1670]" gridcol="6" gridrow="3" pageId="9" pageNumber="230">1533 (22)</td>
</tr>
<tr id="353C34BE792B0032FF5E54EAFA7AF920" box="[166,1421,1686,1710]" gridrow="4" pageId="9" pageNumber="230">
<th id="76ED5DC2792B0032FF5E54EAFE8AF920" box="[166,381,1686,1710]" gridcol="0" gridrow="4" pageId="9" pageNumber="230">
<taxonomicName id="4C0C4D7F792BFFC4FF5E54EAFEC9F923" authorityName="MVZ" baseAuthorityName="Ruppell" baseAuthorityYear="1835" box="[166,318,1686,1709]" class="Reptilia" family="Agamidae" genus="Agama" kingdom="Animalia" order="Squamata" pageId="9" pageNumber="230" phylum="Chordata" rank="species" species="cyanogaster">
<emphasis id="B978EAEE792BFFC4FF5E54EAFEC9F923" box="[166,318,1686,1709]" italics="true" pageId="9" pageNumber="230">A. cyanogaster</emphasis>
</taxonomicName>
(n=8)
</th>
<td id="76ED5DC2792B0032FE6354EAFDC6F920" box="[411,561,1686,1710]" gridcol="1" gridrow="4" pageId="9" pageNumber="230">80118 (101)</td>
<td id="76ED5DC2792B0032FDAB54EAFD1EF920" box="[595,745,1686,1710]" gridcol="2" gridrow="4" pageId="9" pageNumber="230">176303 (255)</td>
<td id="76ED5DC2792B0032FCF454EAFC54F920" box="[780,931,1686,1710]" gridcol="3" gridrow="4" pageId="9" pageNumber="230">104115 (109)</td>
<td id="76ED5DC2792B0032FC3F54EAFBA7F920" box="[967,1104,1686,1710]" gridcol="4" gridrow="4" pageId="9" pageNumber="230">5975 (65)</td>
<td id="76ED5DC2792B0032FB9554EAFB01F920" box="[1133,1270,1686,1710]" gridcol="5" gridrow="4" pageId="9" pageNumber="230">6982 (76)</td>
<td id="76ED5DC2792B0032FAE054EAFA7AF920" box="[1304,1421,1686,1710]" gridcol="6" gridrow="4" pageId="9" pageNumber="230">2654 (38)</td>
</tr>
<tr id="353C34BE792B0032FF5E54C2FA7AF958" box="[166,1421,1726,1750]" gridrow="5" pageId="9" pageNumber="230">
<th id="76ED5DC2792B0032FF5E54C2FE8AF958" box="[166,381,1726,1750]" gridcol="0" gridrow="5" pageId="9" pageNumber="230">
<taxonomicName id="4C0C4D7F792BFFC4FF5E54C2FEE3F958" authorityName="PEMZAF" baseAuthorityName="Gunther" baseAuthorityYear="1894" box="[166,276,1726,1750]" class="Reptilia" family="Agamidae" genus="Acanthocercus" kingdom="Animalia" order="Squamata" pageId="9" pageNumber="230" phylum="Chordata" rank="species" species="gregorii">
<emphasis id="B978EAEE792BFFC4FF5E54C2FEE3F958" box="[166,276,1726,1750]" italics="true" pageId="9" pageNumber="230">A. gregorii</emphasis>
</taxonomicName>
(n=52)
</th>
<td id="76ED5DC2792B0032FE6354C2FDC6F958" box="[411,561,1726,1750]" gridcol="1" gridrow="5" pageId="9" pageNumber="230">74138 (119)</td>
<td id="76ED5DC2792B0032FDAB54C2FD1EF958" box="[595,745,1726,1750]" gridcol="2" gridrow="5" pageId="9" pageNumber="230">189360 (291)</td>
<td id="76ED5DC2792B0032FCF454C2FC54F958" box="[780,931,1726,1750]" gridcol="3" gridrow="5" pageId="9" pageNumber="230">88119 (104)</td>
<td id="76ED5DC2792B0032FC3F54C2FBA7F958" box="[967,1104,1726,1750]" gridcol="4" gridrow="5" pageId="9" pageNumber="230">5285 (64)</td>
<td id="76ED5DC2792B0032FB9554C2FB01F958" box="[1133,1270,1726,1750]" gridcol="5" gridrow="5" pageId="9" pageNumber="230">7096 (81)</td>
<td id="76ED5DC2792B0032FAE054C2FA7AF958" box="[1304,1421,1726,1750]" gridcol="6" gridrow="5" pageId="9" pageNumber="230">737 (22)</td>
</tr>
<tr id="353C34BE792B0032FF5E549AFA7AF970" box="[166,1421,1766,1790]" gridrow="6" pageId="9" pageNumber="230">
<th id="76ED5DC2792B0032FF5E549AFE8AF970" box="[166,381,1766,1790]" gridcol="0" gridrow="6" pageId="9" pageNumber="230">
<taxonomicName id="4C0C4D7F792BFFC4FF5E549BFED2F970" authorityName="ZFMK" baseAuthorityName="Klausewitz" baseAuthorityYear="1957" box="[166,293,1766,1790]" class="Reptilia" family="Agamidae" genus="Acanthocercus" kingdom="Animalia" order="Squamata" pageId="9" pageNumber="230" phylum="Chordata" rank="species" species="kiwuensis">
<emphasis id="B978EAEE792BFFC4FF5E549BFED2F970" box="[166,293,1766,1790]" italics="true" pageId="9" pageNumber="230">A. kiwuensis</emphasis>
</taxonomicName>
(n=19)
</th>
<td id="76ED5DC2792B0032FE63549AFDC6F970" box="[411,561,1766,1790]" gridcol="1" gridrow="6" pageId="9" pageNumber="230">78130 (111)</td>
<td id="76ED5DC2792B0032FDAB549AFD1EF970" box="[595,745,1766,1790]" gridcol="2" gridrow="6" pageId="9" pageNumber="230">200334 (290)</td>
<td id="76ED5DC2792B0032FCF4549AFC54F970" box="[780,931,1766,1790]" gridcol="3" gridrow="6" pageId="9" pageNumber="230">76108 (87)</td>
<td id="76ED5DC2792B0032FC3F549AFBA7F970" box="[967,1104,1766,1790]" gridcol="4" gridrow="6" pageId="9" pageNumber="230">5577 (63)</td>
<td id="76ED5DC2792B0032FB95549AFB01F970" box="[1133,1270,1766,1790]" gridcol="5" gridrow="6" pageId="9" pageNumber="230">5788 (67)</td>
<td id="76ED5DC2792B0032FAE0549AFA7AF970" box="[1304,1421,1766,1790]" gridcol="6" gridrow="6" pageId="9" pageNumber="230">812 (10)</td>
</tr>
<tr id="353C34BE792B0032FF5E5572FA7AF8A9" box="[166,1421,1806,1831]" gridrow="7" pageId="9" pageNumber="230">
<th id="76ED5DC2792B0032FF5E5572FE8AF8A9" box="[166,381,1806,1831]" gridcol="0" gridrow="7" pageId="9" pageNumber="230">
<taxonomicName id="4C0C4D7F792BFFC4FF5E5573FEE5F8A8" baseAuthorityName="Klausewitz" baseAuthorityYear="1957" box="[166,274,1807,1830]" class="Reptilia" family="Agamidae" genus="Acanthocercus" kingdom="Animalia" order="Squamata" pageId="9" pageNumber="230" phylum="Chordata" rank="species" species="minutus">
<emphasis id="B978EAEE792BFFC4FF5E5573FEE5F8A8" box="[166,274,1807,1830]" italics="true" pageId="9" pageNumber="230">A. minutus</emphasis>
</taxonomicName>
(n=20)
</th>
<td id="76ED5DC2792B0032FE635572FDC6F8A9" box="[411,561,1806,1831]" gridcol="1" gridrow="7" pageId="9" pageNumber="230">85120 (101)</td>
<td id="76ED5DC2792B0032FDAB5572FD1EF8A9" box="[595,745,1806,1831]" gridcol="2" gridrow="7" pageId="9" pageNumber="230">189288 (234)</td>
<td id="76ED5DC2792B0032FCF45572FC54F8A9" box="[780,931,1806,1831]" gridcol="3" gridrow="7" pageId="9" pageNumber="230">100120 (108)</td>
<td id="76ED5DC2792B0032FC3F5572FBA7F8A9" box="[967,1104,1806,1831]" gridcol="4" gridrow="7" pageId="9" pageNumber="230">6988 (79)</td>
<td id="76ED5DC2792B0032FB955572FB01F8A9" box="[1133,1270,1806,1831]" gridcol="5" gridrow="7" pageId="9" pageNumber="230">7188 (78)</td>
<td id="76ED5DC2792B0032FAE05572FA7AF8A9" box="[1304,1421,1806,1831]" gridcol="6" gridrow="7" pageId="9" pageNumber="230">934 (26)</td>
</tr>
<tr id="353C34BE792B0032FF5E554BFA7AF8FD" box="[166,1421,1847,1907]" gridrow="8" pageId="9" pageNumber="230">
<th id="76ED5DC2792B0032FF5E554BFE8AF8FD" box="[166,381,1847,1907]" gridcol="0" gridrow="8" pageId="9" pageNumber="230">
<taxonomicName id="4C0C4D7F792BFFC4FF5E554BFEB6F8C1" authorityName="JB" baseAuthorityName="Klausewitz" baseAuthorityYear="1957" box="[166,321,1847,1871]" class="Reptilia" family="Agamidae" genus="Acanthocercus" kingdom="Animalia" order="Squamata" pageId="9" pageNumber="230" phylum="Chordata" rank="species" species="ugandaensis">
<emphasis id="B978EAEE792BFFC4FF5E554BFEB6F8C1" box="[166,321,1847,1871]" italics="true" pageId="9" pageNumber="230">A. ugandaensis</emphasis>
</taxonomicName>
(n=25)
</th>
<td id="76ED5DC2792B0032FE63554BFDC6F8FD" box="[411,561,1847,1907]" gridcol="1" gridrow="8" pageId="9" pageNumber="230">91126 (103)</td>
<td id="76ED5DC2792B0032FDAB554BFD1EF8FD" box="[595,745,1847,1907]" gridcol="2" gridrow="8" pageId="9" pageNumber="230">221351 (275)</td>
<td id="76ED5DC2792B0032FCF4554BFC54F8FD" box="[780,931,1847,1907]" gridcol="3" gridrow="8" pageId="9" pageNumber="230">75117 (94)</td>
<td id="76ED5DC2792B0032FC3F554BFBA7F8FD" box="[967,1104,1847,1907]" gridcol="4" gridrow="8" pageId="9" pageNumber="230">5474 (63)</td>
<td id="76ED5DC2792B0032FB95554BFB01F8FD" box="[1133,1270,1847,1907]" gridcol="5" gridrow="8" pageId="9" pageNumber="230">5995 (73)</td>
<td id="76ED5DC2792B0032FAE0554BFA7AF8FD" box="[1304,1421,1847,1907]" gridcol="6" gridrow="8" pageId="9" pageNumber="230">18 35 (24)</td>
</tr>
<tr id="353C34BE792B0032FF5E55FFFA7AF815" box="[166,1421,1923,1947]" gridrow="9" pageId="9" pageNumber="230">
<th id="76ED5DC2792B0032FF5E55FFFE8AF815" box="[166,381,1923,1947]" gridcol="0" gridrow="9" pageId="9" pageNumber="230">
<taxonomicName id="4C0C4D7F792BFFC4FF5E55FFFEC4F815" authorityName="Wagner, Butler, Ceriaco &amp; Bauer" authorityYear="2021" box="[166,307,1923,1947]" class="Reptilia" family="Agamidae" genus="Acanthocercus" kingdom="Animalia" order="Squamata" pageId="9" pageNumber="230" phylum="Chordata" rank="species" species="margaritae">
<emphasis id="B978EAEE792BFFC4FF5E55FFFEC4F815" box="[166,307,1923,1947]" italics="true" pageId="9" pageNumber="230">A. margaritae</emphasis>
</taxonomicName>
(n=4)
</th>
<td id="76ED5DC2792B0032FE6355FFFDC6F815" box="[411,561,1923,1947]" gridcol="1" gridrow="9" pageId="9" pageNumber="230">96129 (112)</td>
<td id="76ED5DC2792B0032FDAB55FFFD1EF815" box="[595,745,1923,1947]" gridcol="2" gridrow="9" pageId="9" pageNumber="230">217299 (258)</td>
<td id="76ED5DC2792B0032FCF455FFFC54F815" box="[780,931,1923,1947]" gridcol="3" gridrow="9" pageId="9" pageNumber="230">131143 (136)</td>
<td id="76ED5DC2792B0032FC3F55FFFBA7F815" box="[967,1104,1923,1947]" gridcol="4" gridrow="9" pageId="9" pageNumber="230">81133 (103)</td>
<td id="76ED5DC2792B0032FB9555FFFB01F815" box="[1133,1270,1923,1947]" gridcol="5" gridrow="9" pageId="9" pageNumber="230">94107 (101)</td>
<td id="76ED5DC2792B0032FAE055FFFA7AF815" box="[1304,1421,1923,1947]" gridcol="6" gridrow="9" pageId="9" pageNumber="230">919 (14)</td>
</tr>
<tr id="353C34BE792B0032FF5E55D7FA7AF869" box="[166,1421,1963,2023]" gridrow="10" pageId="9" pageNumber="230">
<th id="76ED5DC2792B0032FF5E55D7FE8AF869" box="[166,381,1963,2023]" gridcol="0" gridrow="10" pageId="9" pageNumber="230">
<taxonomicName id="4C0C4D7F792BFFC4FF5E55D0FEE2F84D" authority="Marques &amp; Parrinha &amp; Santos &amp; Bandeira &amp; Butler &amp; Sousa &amp; Bauer &amp; Wagner, 2022" authorityName="Marques &amp; Parrinha &amp; Santos &amp; Bandeira &amp; Butler &amp; Sousa &amp; Bauer &amp; Wagner" authorityYear="2022" box="[166,277,1964,1987]" class="Reptilia" family="Agamidae" genus="Acanthocercus" kingdom="Animalia" order="Squamata" pageId="9" pageNumber="230" phylum="Chordata" rank="species" species="ceriacoi" status="sp. nov.">
<emphasis id="B978EAEE792BFFC4FF5E55D0FEE2F84D" box="[166,277,1964,1987]" italics="true" pageId="9" pageNumber="230">A. ceriacoi</emphasis>
</taxonomicName>
<emphasis id="B978EAEE792BFFC4FEE355D7FE9BF84D" bold="true" box="[283,364,1963,1987]" pageId="9" pageNumber="230">
<taxonomicNameLabel id="A24B5795792BFFC4FEE355D7FE9BF84D" box="[283,364,1963,1987]" pageId="9" pageNumber="230" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
(n=6)
</th>
<td id="76ED5DC2792B0032FE6355D7FDC6F869" box="[411,561,1963,2023]" gridcol="1" gridrow="10" pageId="9" pageNumber="230">99143 (123)</td>
<td id="76ED5DC2792B0032FDAB55D7FD1EF869" box="[595,745,1963,2023]" gridcol="2" gridrow="10" pageId="9" pageNumber="230">224330 (285)</td>
<td id="76ED5DC2792B0032FCF455D7FC54F869" box="[780,931,1963,2023]" gridcol="3" gridrow="10" pageId="9" pageNumber="230">103114 (108)</td>
<td id="76ED5DC2792B0032FC3F55D7FBA7F869" box="[967,1104,1963,2023]" gridcol="4" gridrow="10" pageId="9" pageNumber="230">6065 (63)</td>
<td id="76ED5DC2792B0032FB9555D7FB01F869" box="[1133,1270,1963,2023]" gridcol="5" gridrow="10" pageId="9" pageNumber="230">8089 (85)</td>
<td id="76ED5DC2792B0032FAE055D7FA7AF869" box="[1304,1421,1963,2023]" gridcol="6" gridrow="10" pageId="9" pageNumber="230">16-20 (18) (n=4)</td>
</tr>
</table>
</paragraph>
<paragraph id="8BB336FC7928FFC7FF6F52EBFA8BFECF" blockId="10.[151,1437,150,1941]" pageId="10" pageNumber="231">
(a)
<taxonomicName id="4C0C4D7F7928FFC7FF3E52EBFE20FF3F" authorityName="ZFMK" baseAuthorityName="Smith" baseAuthorityYear="1849" box="[198,471,151,177]" class="Reptilia" family="Agamidae" genus="Acanthocercus" kingdom="Animalia" order="Squamata" pageId="10" pageNumber="231" phylum="Chordata" rank="species" species="atricollis">
<emphasis id="B978EAEE7928FFC7FF3E52EBFE20FF3F" box="[198,471,151,177]" italics="true" pageId="10" pageNumber="231">Acanthocercus atricollis</emphasis>
</taxonomicName>
has a larger SVL (mean
<quantity id="4CF49B197928FFC7FD1652EBFCBCFF3F" box="[750,843,151,177]" metricMagnitude="-1" metricUnit="m" metricValue="1.29" pageId="10" pageNumber="231" unit="mm" value="129.0">129 mm</quantity>
vs.
<quantity id="4CF49B197928FFC7FC8052EBFC20FF3F" box="[888,983,151,177]" metricMagnitude="-1" metricUnit="m" metricValue="1.23" pageId="10" pageNumber="231" unit="mm" value="123.0">123 mm</quantity>
) and total length (mean
<quantity id="4CF49B197928FFC7FB1052EBFAB2FF3F" box="[1256,1349,151,177]" metricMagnitude="-1" metricUnit="m" metricValue="3.02" pageId="10" pageNumber="231" unit="mm" value="302.0">302 mm</quantity>
vs.
<quantity id="4CF49B197928FFC7FA8A52EAFF01FF5B" metricMagnitude="-1" metricUnit="m" metricValue="2.85" pageId="10" pageNumber="231" unit="mm" value="285.0">285 mm</quantity>
). It has higher mean counts of scale rows around midbody (112 vs. 108), dorsal scales (67 vs. 63), and ventral scales (87 vs. 85) than
<taxonomicName id="4C0C4D7F7928FFC7FE0D52A3FD86FF76" authority="Marques &amp; Parrinha &amp; Santos &amp; Bandeira &amp; Butler &amp; Sousa &amp; Bauer &amp; Wagner, 2022" authorityName="Marques &amp; Parrinha &amp; Santos &amp; Bandeira &amp; Butler &amp; Sousa &amp; Bauer &amp; Wagner" authorityYear="2022" box="[501,625,223,249]" class="Reptilia" family="Agamidae" genus="Acanthocercus" kingdom="Animalia" order="Squamata" pageId="10" pageNumber="231" phylum="Chordata" rank="species" species="ceriacoi" status="sp. nov.">
<emphasis id="B978EAEE7928FFC7FE0D52A3FD86FF76" box="[501,625,223,249]" italics="true" pageId="10" pageNumber="231">A. ceriacoi</emphasis>
</taxonomicName>
<emphasis id="B978EAEE7928FFC7FD8052A3FD25FF77" bold="true" box="[632,722,223,249]" pageId="10" pageNumber="231">
<taxonomicNameLabel id="A24B57957928FFC7FD8052A3FD25FF77" box="[632,722,223,249]" pageId="10" pageNumber="231" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
. The mean number of precloacal pores is higher in
<taxonomicName id="4C0C4D7F7928FFC7FAE052A3FA6BFF77" authorityName="ZFMK" baseAuthorityName="Smith" baseAuthorityYear="1849" box="[1304,1436,223,249]" class="Reptilia" family="Agamidae" genus="Acanthocercus" kingdom="Animalia" order="Squamata" pageId="10" pageNumber="231" phylum="Chordata" rank="species" species="atricollis">
<emphasis id="B978EAEE7928FFC7FAE052A3FA6BFF77" box="[1304,1436,223,249]" italics="true" pageId="10" pageNumber="231">A. atricollis</emphasis>
</taxonomicName>
than in
<taxonomicName id="4C0C4D7F7928FFC7FEE3537FFE60FE92" authority="Marques &amp; Parrinha &amp; Santos &amp; Bandeira &amp; Butler &amp; Sousa &amp; Bauer &amp; Wagner, 2022" authorityName="Marques &amp; Parrinha &amp; Santos &amp; Bandeira &amp; Butler &amp; Sousa &amp; Bauer &amp; Wagner" authorityYear="2022" box="[283,407,259,285]" class="Reptilia" family="Agamidae" genus="Acanthocercus" kingdom="Animalia" order="Squamata" pageId="10" pageNumber="231" phylum="Chordata" rank="species" species="ceriacoi" status="sp. nov.">
<emphasis id="B978EAEE7928FFC7FEE3537FFE60FE92" box="[283,407,259,285]" italics="true" pageId="10" pageNumber="231">A. ceriacoi</emphasis>
</taxonomicName>
<emphasis id="B978EAEE7928FFC7FE66537FFE0FFE93" bold="true" box="[414,504,259,285]" pageId="10" pageNumber="231">
<taxonomicNameLabel id="A24B57957928FFC7FE66537FFE0FFE93" box="[414,504,259,285]" pageId="10" pageNumber="231" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
(22 vs 18).
<taxonomicName id="4C0C4D7F7928FFC7FD79537FFC64FE93" authorityName="ZFMK" baseAuthorityName="Smith" baseAuthorityYear="1849" box="[641,915,259,285]" class="Reptilia" family="Agamidae" genus="Acanthocercus" kingdom="Animalia" order="Squamata" pageId="10" pageNumber="231" phylum="Chordata" rank="species" species="atricollis">
<emphasis id="B978EAEE7928FFC7FD79537FFC64FE93" box="[641,915,259,285]" italics="true" pageId="10" pageNumber="231">Acanthocercus atricollis</emphasis>
</taxonomicName>
presents a blue or green head and ventral surface, in some individuals extending to the body, while in
<taxonomicName id="4C0C4D7F7928FFC7FCBE535BFC35FECE" authority="Marques &amp; Parrinha &amp; Santos &amp; Bandeira &amp; Butler &amp; Sousa &amp; Bauer &amp; Wagner, 2022" authorityName="Marques &amp; Parrinha &amp; Santos &amp; Bandeira &amp; Butler &amp; Sousa &amp; Bauer &amp; Wagner" authorityYear="2022" box="[838,962,295,321]" class="Reptilia" family="Agamidae" genus="Acanthocercus" kingdom="Animalia" order="Squamata" pageId="10" pageNumber="231" phylum="Chordata" rank="species" species="ceriacoi" status="sp. nov.">
<emphasis id="B978EAEE7928FFC7FCBE535BFC35FECE" box="[838,962,295,321]" italics="true" pageId="10" pageNumber="231">A. ceriacoi</emphasis>
</taxonomicName>
<emphasis id="B978EAEE7928FFC7FC30535BFBD5FECF" bold="true" box="[968,1058,295,321]" pageId="10" pageNumber="231">
<taxonomicNameLabel id="A24B57957928FFC7FC30535BFBD5FECF" box="[968,1058,295,321]" pageId="10" pageNumber="231" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
the blue is limited to the head.
</paragraph>
<paragraph id="8BB336FC7928FFC7FF6F5337FC80FE7B" blockId="10.[151,1437,150,1941]" pageId="10" pageNumber="231">
(b)
<taxonomicName id="4C0C4D7F7928FFC7FF3E5337FE3EFEEB" box="[198,457,331,357]" class="Reptilia" family="Agamidae" genus="Acanthocercus" kingdom="Animalia" order="Squamata" pageId="10" pageNumber="231" phylum="Chordata" rank="species" species="branchi">
<emphasis id="B978EAEE7928FFC7FF3E5337FE3EFEEB" box="[198,457,331,357]" italics="true" pageId="10" pageNumber="231">Acanthocercus branchi</emphasis>
</taxonomicName>
has a smaller SVL (mean
<quantity id="4CF49B197928FFC7FD085337FCBAFEEB" box="[752,845,331,357]" metricMagnitude="-1" metricUnit="m" metricValue="1.09" pageId="10" pageNumber="231" unit="mm" value="109.0">109 mm</quantity>
vs.
<quantity id="4CF49B197928FFC7FC825337FC2FFEEB" box="[890,984,331,357]" metricMagnitude="-1" metricUnit="m" metricValue="1.23" pageId="10" pageNumber="231" unit="mm" value="123.0">123 mm</quantity>
) and total length (mean
<quantity id="4CF49B197928FFC7FB105337FAB2FEEB" box="[1256,1349,331,357]" metricMagnitude="-1" metricUnit="m" metricValue="2.62" pageId="10" pageNumber="231" unit="mm" value="262.0">262 mm</quantity>
vs.
<quantity id="4CF49B197928FFC7FA8A5336FF01FE07" metricMagnitude="-1" metricUnit="m" metricValue="2.85" pageId="10" pageNumber="231" unit="mm" value="285.0">285 mm</quantity>
). It has higher mean counts of scale rows around midbody (117 vs. 108), dorsal scales (73 vs. 63), and ventral scales (88 vs. 85) than
<taxonomicName id="4C0C4D7F7928FFC7FDE753EFFD6DFE22" authority="Marques &amp; Parrinha &amp; Santos &amp; Bandeira &amp; Butler &amp; Sousa &amp; Bauer &amp; Wagner, 2022" authorityName="Marques &amp; Parrinha &amp; Santos &amp; Bandeira &amp; Butler &amp; Sousa &amp; Bauer &amp; Wagner" authorityYear="2022" box="[543,666,403,429]" class="Reptilia" family="Agamidae" genus="Acanthocercus" kingdom="Animalia" order="Squamata" pageId="10" pageNumber="231" phylum="Chordata" rank="species" species="ceriacoi" status="sp. nov.">
<emphasis id="B978EAEE7928FFC7FDE753EFFD6DFE22" box="[543,666,403,429]" italics="true" pageId="10" pageNumber="231">A. ceriacoi</emphasis>
</taxonomicName>
<emphasis id="B978EAEE7928FFC7FD5A53EFFD0BFE23" bold="true" box="[674,764,403,429]" pageId="10" pageNumber="231">
<taxonomicNameLabel id="A24B57957928FFC7FD5A53EFFD0BFE23" box="[674,764,403,429]" pageId="10" pageNumber="231" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
. The ranges of these characters largely overlap in both species, although they are higher in
<taxonomicName id="4C0C4D7F7928FFC7FDCE53CBFD5AFE5F" box="[566,685,439,465]" class="Reptilia" family="Agamidae" genus="Acanthocercus" kingdom="Animalia" order="Squamata" pageId="10" pageNumber="231" phylum="Chordata" rank="species" species="branchi">
<emphasis id="B978EAEE7928FFC7FDCE53CBFD5AFE5F" box="[566,685,439,465]" italics="true" pageId="10" pageNumber="231">A. branchi</emphasis>
</taxonomicName>
than in
<taxonomicName id="4C0C4D7F7928FFC7FCF153CBFC73FE5E" authority="Marques &amp; Parrinha &amp; Santos &amp; Bandeira &amp; Butler &amp; Sousa &amp; Bauer &amp; Wagner, 2022" authorityName="Marques &amp; Parrinha &amp; Santos &amp; Bandeira &amp; Butler &amp; Sousa &amp; Bauer &amp; Wagner" authorityYear="2022" box="[777,900,439,465]" class="Reptilia" family="Agamidae" genus="Acanthocercus" kingdom="Animalia" order="Squamata" pageId="10" pageNumber="231" phylum="Chordata" rank="species" species="ceriacoi" status="sp. nov.">
<emphasis id="B978EAEE7928FFC7FCF153CBFC73FE5E" box="[777,900,439,465]" italics="true" pageId="10" pageNumber="231">A. ceriacoi</emphasis>
</taxonomicName>
<emphasis id="B978EAEE7928FFC7FC7353CBFC12FE5F" bold="true" box="[907,997,439,465]" pageId="10" pageNumber="231">
<taxonomicNameLabel id="A24B57957928FFC7FC7353CBFC12FE5F" box="[907,997,439,465]" pageId="10" pageNumber="231" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
. The mean number of precloacal pores is higher in
<taxonomicName id="4C0C4D7F7928FFC7FEB353A7FE36FE7B" box="[331,449,475,501]" class="Reptilia" family="Agamidae" genus="Acanthocercus" kingdom="Animalia" order="Squamata" pageId="10" pageNumber="231" phylum="Chordata" rank="species" species="branchi">
<emphasis id="B978EAEE7928FFC7FEB353A7FE36FE7B" box="[331,449,475,501]" italics="true" pageId="10" pageNumber="231">A. branchi</emphasis>
</taxonomicName>
than in
<taxonomicName id="4C0C4D7F7928FFC7FDE453A7FD60FE7A" authority="Marques &amp; Parrinha &amp; Santos &amp; Bandeira &amp; Butler &amp; Sousa &amp; Bauer &amp; Wagner, 2022" authorityName="Marques &amp; Parrinha &amp; Santos &amp; Bandeira &amp; Butler &amp; Sousa &amp; Bauer &amp; Wagner" authorityYear="2022" box="[540,663,475,501]" class="Reptilia" family="Agamidae" genus="Acanthocercus" kingdom="Animalia" order="Squamata" pageId="10" pageNumber="231" phylum="Chordata" rank="species" species="ceriacoi" status="sp. nov.">
<emphasis id="B978EAEE7928FFC7FDE453A7FD60FE7A" box="[540,663,475,501]" italics="true" pageId="10" pageNumber="231">A. ceriacoi</emphasis>
</taxonomicName>
<emphasis id="B978EAEE7928FFC7FD6653A7FD00FE7B" bold="true" box="[670,759,475,501]" pageId="10" pageNumber="231">
<taxonomicNameLabel id="A24B57957928FFC7FD6653A7FD00FE7B" box="[670,759,475,501]" pageId="10" pageNumber="231" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
(20 vs 18).
</paragraph>
<paragraph id="8BB336FC7928FFC7FF6F5383FBA8FD7F" blockId="10.[151,1437,150,1941]" pageId="10" pageNumber="231">
(c)
<taxonomicName id="4C0C4D7F7928FFC7FF3E5383FDE0FD97" authorityName="HF" baseAuthorityName="Falk" baseAuthorityYear="1925" box="[198,535,511,537]" class="Reptilia" family="Agamidae" genus="Acanthocercus" kingdom="Animalia" order="Squamata" pageId="10" pageNumber="231" phylum="Chordata" rank="species" species="cyanocephalus">
<emphasis id="B978EAEE7928FFC7FF3E5383FDE0FD97" box="[198,535,511,537]" italics="true" pageId="10" pageNumber="231">Acanthocercus cyanocephalus</emphasis>
</taxonomicName>
has a larger SVL (mean
<quantity id="4CF49B197928FFC7FCD15383FC72FD97" box="[809,901,511,537]" metricMagnitude="-1" metricUnit="m" metricValue="1.38" pageId="10" pageNumber="231" unit="mm" value="138.0">138 mm</quantity>
vs.
<quantity id="4CF49B197928FFC7FC495383FBF9FD97" box="[945,1038,511,537]" metricMagnitude="-1" metricUnit="m" metricValue="1.23" pageId="10" pageNumber="231" unit="mm" value="123.0">123 mm</quantity>
) and total length (mean
<quantity id="4CF49B197928FFC7FAE35383FA80FD97" box="[1307,1399,511,537]" metricMagnitude="-1" metricUnit="m" metricValue="3.34" pageId="10" pageNumber="231" unit="mm" value="334.0">334 mm</quantity>
vs.
<quantity id="4CF49B197928FFC7FF3E505EFED1FDB3" box="[198,294,546,573]" metricMagnitude="-1" metricUnit="m" metricValue="2.85" pageId="10" pageNumber="231" unit="mm" value="285.0">285 mm</quantity>
). It is morphologically very similar to
<taxonomicName id="4C0C4D7F7928FFC7FD28505FFCBDFDB2" authority="Marques &amp; Parrinha &amp; Santos &amp; Bandeira &amp; Butler &amp; Sousa &amp; Bauer &amp; Wagner, 2022" authorityName="Marques &amp; Parrinha &amp; Santos &amp; Bandeira &amp; Butler &amp; Sousa &amp; Bauer &amp; Wagner" authorityYear="2022" box="[720,842,547,573]" class="Reptilia" family="Agamidae" genus="Acanthocercus" kingdom="Animalia" order="Squamata" pageId="10" pageNumber="231" phylum="Chordata" rank="species" species="ceriacoi" status="sp. nov.">
<emphasis id="B978EAEE7928FFC7FD28505FFCBDFDB2" box="[720,842,547,573]" italics="true" pageId="10" pageNumber="231">A. ceriacoi</emphasis>
</taxonomicName>
<emphasis id="B978EAEE7928FFC7FCA8505FFC5EFDB3" bold="true" box="[848,937,547,573]" pageId="10" pageNumber="231">
<taxonomicNameLabel id="A24B57957928FFC7FCA8505FFC5EFDB3" box="[848,937,547,573]" pageId="10" pageNumber="231" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
and its meristic differences are subtle.
<taxonomicName id="4C0C4D7F7928FFC7FAA1505FFE29FDEF" authorityName="HF" baseAuthorityName="Falk" baseAuthorityYear="1925" class="Reptilia" family="Agamidae" genus="Acanthocercus" kingdom="Animalia" order="Squamata" pageId="10" pageNumber="231" phylum="Chordata" rank="species" species="cyanocephalus">
<emphasis id="B978EAEE7928FFC7FAA1505FFE29FDEF" italics="true" pageId="10" pageNumber="231">Acanthocercus cyanocephalus</emphasis>
</taxonomicName>
has a higher mean dorsal scale count (68 vs. 63), and a lower mean number of ventral scales (80 vs. 85) than
<taxonomicName id="4C0C4D7F7928FFC7FE3B5017FDC9FD0A" authority="Marques &amp; Parrinha &amp; Santos &amp; Bandeira &amp; Butler &amp; Sousa &amp; Bauer &amp; Wagner, 2022" authorityName="Marques &amp; Parrinha &amp; Santos &amp; Bandeira &amp; Butler &amp; Sousa &amp; Bauer &amp; Wagner" authorityYear="2022" box="[451,574,619,645]" class="Reptilia" family="Agamidae" genus="Acanthocercus" kingdom="Animalia" order="Squamata" pageId="10" pageNumber="231" phylum="Chordata" rank="species" species="ceriacoi" status="sp. nov.">
<emphasis id="B978EAEE7928FFC7FE3B5017FDC9FD0A" box="[451,574,619,645]" italics="true" pageId="10" pageNumber="231">A. ceriacoi</emphasis>
</taxonomicName>
<emphasis id="B978EAEE7928FFC7FDBC5017FD6AFD0B" bold="true" box="[580,669,619,645]" pageId="10" pageNumber="231">
<taxonomicNameLabel id="A24B57957928FFC7FDBC5017FD6AFD0B" box="[580,669,619,645]" pageId="10" pageNumber="231" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
. The mean number of precloacal pores is higher in
<taxonomicName id="4C0C4D7F7928FFC7FB2F5017FA6BFD0B" authorityName="HF" baseAuthorityName="Falk" baseAuthorityYear="1925" box="[1239,1436,619,645]" class="Reptilia" family="Agamidae" genus="Acanthocercus" kingdom="Animalia" order="Squamata" pageId="10" pageNumber="231" phylum="Chordata" rank="species" species="cyanocephalus">
<emphasis id="B978EAEE7928FFC7FB2F5017FA6BFD0B" box="[1239,1436,619,645]" italics="true" pageId="10" pageNumber="231">A. cyanocephalus</emphasis>
</taxonomicName>
than in
<taxonomicName id="4C0C4D7F7928FFC7FEE250F3FE63FD26" authority="Marques &amp; Parrinha &amp; Santos &amp; Bandeira &amp; Butler &amp; Sousa &amp; Bauer &amp; Wagner, 2022" authorityName="Marques &amp; Parrinha &amp; Santos &amp; Bandeira &amp; Butler &amp; Sousa &amp; Bauer &amp; Wagner" authorityYear="2022" box="[282,404,655,681]" class="Reptilia" family="Agamidae" genus="Acanthocercus" kingdom="Animalia" order="Squamata" pageId="10" pageNumber="231" phylum="Chordata" rank="species" species="ceriacoi" status="sp. nov.">
<emphasis id="B978EAEE7928FFC7FEE250F3FE63FD26" box="[282,404,655,681]" italics="true" pageId="10" pageNumber="231">A. ceriacoi</emphasis>
</taxonomicName>
<emphasis id="B978EAEE7928FFC7FE6350F3FE03FD27" bold="true" box="[411,500,655,681]" pageId="10" pageNumber="231">
<taxonomicNameLabel id="A24B57957928FFC7FE6350F3FE03FD27" box="[411,500,655,681]" pageId="10" pageNumber="231" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
(22 vs 18). The major difference between these two species is the coloration of displaying males, which in
<taxonomicName id="4C0C4D7F7928FFC7FE2450CFFD55FD43" authorityName="HF" baseAuthorityName="Falk" baseAuthorityYear="1925" box="[476,674,691,717]" class="Reptilia" family="Agamidae" genus="Acanthocercus" kingdom="Animalia" order="Squamata" pageId="10" pageNumber="231" phylum="Chordata" rank="species" species="cyanocephalus">
<emphasis id="B978EAEE7928FFC7FE2450CFFD55FD43" box="[476,674,691,717]" italics="true" pageId="10" pageNumber="231">A. cyanocephalus</emphasis>
</taxonomicName>
is characterized by the blue coloration of the head extending to the neck and shoulders, whereas in
<taxonomicName id="4C0C4D7F7928FFC7FDD250ABFD52FD7E" authority="Marques &amp; Parrinha &amp; Santos &amp; Bandeira &amp; Butler &amp; Sousa &amp; Bauer &amp; Wagner, 2022" authorityName="Marques &amp; Parrinha &amp; Santos &amp; Bandeira &amp; Butler &amp; Sousa &amp; Bauer &amp; Wagner" authorityYear="2022" box="[554,677,727,753]" class="Reptilia" family="Agamidae" genus="Acanthocercus" kingdom="Animalia" order="Squamata" pageId="10" pageNumber="231" phylum="Chordata" rank="species" species="ceriacoi" status="sp. nov.">
<emphasis id="B978EAEE7928FFC7FDD250ABFD52FD7E" box="[554,677,727,753]" italics="true" pageId="10" pageNumber="231">A. ceriacoi</emphasis>
</taxonomicName>
<emphasis id="B978EAEE7928FFC7FD5450ABFCF2FD7F" bold="true" box="[684,773,727,753]" pageId="10" pageNumber="231">
<taxonomicNameLabel id="A24B57957928FFC7FD5450ABFCF2FD7F" box="[684,773,727,753]" pageId="10" pageNumber="231" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
the blue is limited to the head.
</paragraph>
<paragraph id="8BB336FC7928FFC7FF6F5087FC0AFC47" blockId="10.[151,1437,150,1941]" pageId="10" pageNumber="231">
(d)
<taxonomicName id="4C0C4D7F7928FFC7FF3E5087FE0CFC9A" baseAuthorityName="Ruppell" baseAuthorityYear="1835" box="[198,507,763,789]" class="Reptilia" family="Agamidae" genus="Acanthocercus" kingdom="Animalia" order="Squamata" pageId="10" pageNumber="231" phylum="Chordata" rank="species" species="cyanogaster">
<emphasis id="B978EAEE7928FFC7FF3E5087FE0CFC9A" box="[198,507,763,789]" italics="true" pageId="10" pageNumber="231">Acanthocercus cyanogaster</emphasis>
</taxonomicName>
has a smaller SVL (mean
<quantity id="4CF49B197928FFC7FCDA5087FC89FC9B" box="[802,894,763,789]" metricMagnitude="-1" metricUnit="m" metricValue="1.01" pageId="10" pageNumber="231" unit="mm" value="101.0">101 mm</quantity>
vs.
<quantity id="4CF49B197928FFC7FC535087FBFEFC9B" box="[939,1033,763,789]" metricMagnitude="-1" metricUnit="m" metricValue="1.23" pageId="10" pageNumber="231" unit="mm" value="123.0">123 mm</quantity>
) and total length (mean
<quantity id="4CF49B197928FFC7FAE15086FA81FC9B" box="[1305,1398,762,789]" metricMagnitude="-1" metricUnit="m" metricValue="2.55" pageId="10" pageNumber="231" unit="mm" value="255.0">255 mm</quantity>
vs.
<quantity id="4CF49B197928FFC7FF3E5162FEDFFCB7" box="[198,296,798,825]" metricMagnitude="-1" metricUnit="m" metricValue="2.85" pageId="10" pageNumber="231" unit="mm" value="285.0">285 mm</quantity>
). It has lower mean counts of ventral scales (76 vs. 85) than
<taxonomicName id="4C0C4D7F7928FFC7FC185163FBAAFCB6" authority="Marques &amp; Parrinha &amp; Santos &amp; Bandeira &amp; Butler &amp; Sousa &amp; Bauer &amp; Wagner, 2022" authorityName="Marques &amp; Parrinha &amp; Santos &amp; Bandeira &amp; Butler &amp; Sousa &amp; Bauer &amp; Wagner" authorityYear="2022" box="[992,1117,799,825]" class="Reptilia" family="Agamidae" genus="Acanthocercus" kingdom="Animalia" order="Squamata" pageId="10" pageNumber="231" phylum="Chordata" rank="species" species="ceriacoi" status="sp. nov.">
<emphasis id="B978EAEE7928FFC7FC185163FBAAFCB6" box="[992,1117,799,825]" italics="true" pageId="10" pageNumber="231">A. ceriacoi</emphasis>
</taxonomicName>
<emphasis id="B978EAEE7928FFC7FB9D5163FB37FCB7" bold="true" box="[1125,1216,799,825]" pageId="10" pageNumber="231">
<taxonomicNameLabel id="A24B57957928FFC7FB9D5163FB37FCB7" box="[1125,1216,799,825]" pageId="10" pageNumber="231" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
. The ranges of this character slightly overlap in the two species, although they are higher in
<taxonomicName id="4C0C4D7F7928FFC7FC00513FFB83FCD2" authority="Marques &amp; Parrinha &amp; Santos &amp; Bandeira &amp; Butler &amp; Sousa &amp; Bauer &amp; Wagner, 2022" authorityName="Marques &amp; Parrinha &amp; Santos &amp; Bandeira &amp; Butler &amp; Sousa &amp; Bauer &amp; Wagner" authorityYear="2022" box="[1016,1140,835,861]" class="Reptilia" family="Agamidae" genus="Acanthocercus" kingdom="Animalia" order="Squamata" pageId="10" pageNumber="231" phylum="Chordata" rank="species" species="ceriacoi" status="sp. nov.">
<emphasis id="B978EAEE7928FFC7FC00513FFB83FCD2" box="[1016,1140,835,861]" italics="true" pageId="10" pageNumber="231">A. ceriacoi</emphasis>
</taxonomicName>
<emphasis id="B978EAEE7928FFC7FB83513FFB22FCD3" bold="true" box="[1147,1237,835,861]" pageId="10" pageNumber="231">
<taxonomicNameLabel id="A24B57957928FFC7FB83513FFB22FCD3" box="[1147,1237,835,861]" pageId="10" pageNumber="231" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
than in
<taxonomicName id="4C0C4D7F7928FFC7FAC9513FFEFBFC0E" baseAuthorityName="Ruppell" baseAuthorityYear="1835" class="Reptilia" family="Agamidae" genus="Acanthocercus" kingdom="Animalia" order="Squamata" pageId="10" pageNumber="231" phylum="Chordata" rank="species" species="cyanogaster">
<emphasis id="B978EAEE7928FFC7FAC9513FFEFBFC0E" italics="true" pageId="10" pageNumber="231">A. cyanogaster</emphasis>
</taxonomicName>
. The mean number of precloacal pores is much higher in
<taxonomicName id="4C0C4D7F7928FFC7FC6A511BFBCDFC0E" baseAuthorityName="Ruppell" baseAuthorityYear="1835" box="[914,1082,871,897]" class="Reptilia" family="Agamidae" genus="Acanthocercus" kingdom="Animalia" order="Squamata" pageId="10" pageNumber="231" phylum="Chordata" rank="species" species="cyanogaster">
<emphasis id="B978EAEE7928FFC7FC6A511BFBCDFC0E" box="[914,1082,871,897]" italics="true" pageId="10" pageNumber="231">A. cyanogaster</emphasis>
</taxonomicName>
than in
<taxonomicName id="4C0C4D7F7928FFC7FB6D511BFAE7FC0E" authority="Marques &amp; Parrinha &amp; Santos &amp; Bandeira &amp; Butler &amp; Sousa &amp; Bauer &amp; Wagner, 2022" authorityName="Marques &amp; Parrinha &amp; Santos &amp; Bandeira &amp; Butler &amp; Sousa &amp; Bauer &amp; Wagner" authorityYear="2022" box="[1173,1296,871,897]" class="Reptilia" family="Agamidae" genus="Acanthocercus" kingdom="Animalia" order="Squamata" pageId="10" pageNumber="231" phylum="Chordata" rank="species" species="ceriacoi" status="sp. nov.">
<emphasis id="B978EAEE7928FFC7FB6D511BFAE7FC0E" box="[1173,1296,871,897]" italics="true" pageId="10" pageNumber="231">A. ceriacoi</emphasis>
</taxonomicName>
<emphasis id="B978EAEE7928FFC7FAEE511BFA87FC0F" bold="true" box="[1302,1392,871,897]" pageId="10" pageNumber="231">
<taxonomicNameLabel id="A24B57957928FFC7FAEE511BFA87FC0F" box="[1302,1392,871,897]" pageId="10" pageNumber="231" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
(38 vs 18). [
<taxonomicName id="4C0C4D7F7928FFC7FED851F7FE30FC2A" baseAuthorityName="Ruppell" baseAuthorityYear="1835" box="[288,455,907,933]" class="Reptilia" family="Agamidae" genus="Acanthocercus" kingdom="Animalia" order="Squamata" pageId="10" pageNumber="231" phylum="Chordata" rank="species" species="cyanogaster">
<emphasis id="B978EAEE7928FFC7FED851F7FE30FC2A" box="[288,455,907,933]" italics="true" pageId="10" pageNumber="231">A. cyanogaster</emphasis>
</taxonomicName>
is not a part of
<taxonomicName id="4C0C4D7F7928FFC7FD9751F7FD05FC2B" authorityName="ZFMK" baseAuthorityName="Smith" baseAuthorityYear="1849" box="[623,754,907,933]" class="Reptilia" family="Agamidae" genus="Acanthocercus" kingdom="Animalia" order="Squamata" pageId="10" pageNumber="231" phylum="Chordata" rank="species" species="atricollis">
<emphasis id="B978EAEE7928FFC7FD9751F7FD05FC2B" box="[623,754,907,933]" italics="true" pageId="10" pageNumber="231">A. atricollis</emphasis>
</taxonomicName>
group but is included here due to its morphological similarity and because of the historical use of the name to describe this population].
</paragraph>
<paragraph id="8BB336FC7928FFC7FF6F51AFFEDBFB4B" blockId="10.[151,1437,150,1941]" pageId="10" pageNumber="231">
(e)
<taxonomicName id="4C0C4D7F7928FFC7FF3E51AFFE3AFC62" authorityName="PEMZAF" baseAuthorityName="Gunther" baseAuthorityYear="1894" box="[198,461,979,1005]" class="Reptilia" family="Agamidae" genus="Acanthocercus" kingdom="Animalia" order="Squamata" pageId="10" pageNumber="231" phylum="Chordata" rank="species" species="gregorii">
<emphasis id="B978EAEE7928FFC7FF3E51AFFE3AFC62" box="[198,461,979,1005]" italics="true" pageId="10" pageNumber="231">Acanthocercus gregorii</emphasis>
</taxonomicName>
has a smaller SVL (mean
<quantity id="4CF49B197928FFC7FD0E51AFFCA5FC63" box="[758,850,979,1005]" metricMagnitude="-1" metricUnit="m" metricValue="1.19" pageId="10" pageNumber="231" unit="mm" value="119.0">119 mm</quantity>
vs.
<quantity id="4CF49B197928FFC7FC7851AFFC29FC63" box="[896,990,979,1005]" metricMagnitude="-1" metricUnit="m" metricValue="1.23" pageId="10" pageNumber="231" unit="mm" value="123.0">123 mm</quantity>
) but greater total length (mean
<quantity id="4CF49B197928FFC7FAC751AFFA6AFC63" box="[1343,1437,979,1005]" metricMagnitude="-1" metricUnit="m" metricValue="2.91" pageId="10" pageNumber="231" unit="mm" value="291.0">291 mm</quantity>
vs.
<quantity id="4CF49B197928FFC7FF15518AFEB8FB9F" box="[237,335,1014,1041]" metricMagnitude="-1" metricUnit="m" metricValue="2.85" pageId="10" pageNumber="231" unit="mm" value="285.0">285 mm</quantity>
).
<taxonomicName id="4C0C4D7F7928FFC7FE9B518BFE2AFB9E" authorityName="PEMZAF" baseAuthorityName="Gunther" baseAuthorityYear="1894" box="[355,477,1015,1041]" class="Reptilia" family="Agamidae" genus="Acanthocercus" kingdom="Animalia" order="Squamata" pageId="10" pageNumber="231" phylum="Chordata" rank="species" species="gregorii">
<emphasis id="B978EAEE7928FFC7FE9B518BFE2AFB9E" box="[355,477,1015,1041]" italics="true" pageId="10" pageNumber="231">A. gregorii</emphasis>
</taxonomicName>
has lower mean counts of scale rows around midbody (104 vs. 108) than
<taxonomicName id="4C0C4D7F7928FFC7FAD9518BFA6BFB9E" authority="Marques &amp; Parrinha &amp; Santos &amp; Bandeira &amp; Butler &amp; Sousa &amp; Bauer &amp; Wagner, 2022" authorityName="Marques &amp; Parrinha &amp; Santos &amp; Bandeira &amp; Butler &amp; Sousa &amp; Bauer &amp; Wagner" authorityYear="2022" box="[1313,1436,1015,1041]" class="Reptilia" family="Agamidae" genus="Acanthocercus" kingdom="Animalia" order="Squamata" pageId="10" pageNumber="231" phylum="Chordata" rank="species" species="ceriacoi" status="sp. nov.">
<emphasis id="B978EAEE7928FFC7FAD9518BFA6BFB9E" box="[1313,1436,1015,1041]" italics="true" pageId="10" pageNumber="231">A. ceriacoi</emphasis>
</taxonomicName>
<emphasis id="B978EAEE7928FFC7FF3E5667FEE8FBBB" bold="true" box="[198,287,1051,1077]" pageId="10" pageNumber="231">
<taxonomicNameLabel id="A24B57957928FFC7FF3E5667FEE8FBBB" box="[198,287,1051,1077]" pageId="10" pageNumber="231" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
. The ranges of these characters are higher in
<taxonomicName id="4C0C4D7F7928FFC7FCEF5667FC65FBBA" authority="Marques &amp; Parrinha &amp; Santos &amp; Bandeira &amp; Butler &amp; Sousa &amp; Bauer &amp; Wagner, 2022" authorityName="Marques &amp; Parrinha &amp; Santos &amp; Bandeira &amp; Butler &amp; Sousa &amp; Bauer &amp; Wagner" authorityYear="2022" box="[791,914,1051,1077]" class="Reptilia" family="Agamidae" genus="Acanthocercus" kingdom="Animalia" order="Squamata" pageId="10" pageNumber="231" phylum="Chordata" rank="species" species="ceriacoi" status="sp. nov.">
<emphasis id="B978EAEE7928FFC7FCEF5667FC65FBBA" box="[791,914,1051,1077]" italics="true" pageId="10" pageNumber="231">A. ceriacoi</emphasis>
</taxonomicName>
<emphasis id="B978EAEE7928FFC7FC605667FC06FBBB" bold="true" box="[920,1009,1051,1077]" pageId="10" pageNumber="231">
<taxonomicNameLabel id="A24B57957928FFC7FC605667FC06FBBB" box="[920,1009,1051,1077]" pageId="10" pageNumber="231" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
than in
<taxonomicName id="4C0C4D7F7928FFC7FBB35667FB32FBBA" authorityName="PEMZAF" baseAuthorityName="Gunther" baseAuthorityYear="1894" box="[1099,1221,1051,1077]" class="Reptilia" family="Agamidae" genus="Acanthocercus" kingdom="Animalia" order="Squamata" pageId="10" pageNumber="231" phylum="Chordata" rank="species" species="gregorii">
<emphasis id="B978EAEE7928FFC7FBB35667FB32FBBA" box="[1099,1221,1051,1077]" italics="true" pageId="10" pageNumber="231">A. gregorii</emphasis>
</taxonomicName>
. The mean number of precloacal pores is higher in
<taxonomicName id="4C0C4D7F7928FFC7FDDE5643FD57FBD6" authorityName="PEMZAF" baseAuthorityName="Gunther" baseAuthorityYear="1894" box="[550,672,1087,1113]" class="Reptilia" family="Agamidae" genus="Acanthocercus" kingdom="Animalia" order="Squamata" pageId="10" pageNumber="231" phylum="Chordata" rank="species" species="gregorii">
<emphasis id="B978EAEE7928FFC7FDDE5643FD57FBD6" box="[550,672,1087,1113]" italics="true" pageId="10" pageNumber="231">A. gregorii</emphasis>
</taxonomicName>
than in
<taxonomicName id="4C0C4D7F7928FFC7FD035643FC81FBD6" authority="Marques &amp; Parrinha &amp; Santos &amp; Bandeira &amp; Butler &amp; Sousa &amp; Bauer &amp; Wagner, 2022" authorityName="Marques &amp; Parrinha &amp; Santos &amp; Bandeira &amp; Butler &amp; Sousa &amp; Bauer &amp; Wagner" authorityYear="2022" box="[763,886,1087,1113]" class="Reptilia" family="Agamidae" genus="Acanthocercus" kingdom="Animalia" order="Squamata" pageId="10" pageNumber="231" phylum="Chordata" rank="species" species="ceriacoi" status="sp. nov.">
<emphasis id="B978EAEE7928FFC7FD035643FC81FBD6" box="[763,886,1087,1113]" italics="true" pageId="10" pageNumber="231">A. ceriacoi</emphasis>
</taxonomicName>
<emphasis id="B978EAEE7928FFC7FC855643FC21FBD7" bold="true" box="[893,982,1087,1113]" pageId="10" pageNumber="231">
<taxonomicNameLabel id="A24B57957928FFC7FC855643FC21FBD7" box="[893,982,1087,1113]" pageId="10" pageNumber="231" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
(22 vs 18). In displaying adult males,
<taxonomicName id="4C0C4D7F7928FFC7FA7C5643FED6FBF2" authorityName="PEMZAF" baseAuthorityName="Gunther" baseAuthorityYear="1894" class="Reptilia" family="Agamidae" genus="Acanthocercus" kingdom="Animalia" order="Squamata" pageId="10" pageNumber="231" phylum="Chordata" rank="species" species="gregorii">
<emphasis id="B978EAEE7928FFC7FA7C5643FED6FBF2" italics="true" pageId="10" pageNumber="231">A. gregorii</emphasis>
</taxonomicName>
are characterized by their bluish-green head and a uniform greenish-blue throat, with a uniform blue body (
<bibRefCitation id="EF9D4B0D7928FFC7FEF556FBFE38FB2F" author="Klausewitz, W." box="[269,463,1158,1185]" pageId="10" pageNumber="231" pagination="157 - 174" refId="ref11407" refString="Klausewitz, W. (1957) Eidonomische Untersuchungen uber die Rassenkreise Agama cyanogaster und Agama atricollis. 2. Die Unterarten von Agama atricollis. Senckenbergiana biologica, 38, 157 - 174." type="journal article" year="1957">Klausewitz 1957</bibRefCitation>
;
<bibRefCitation id="EF9D4B0D7928FFC7FE2356FBFD42FB2F" author="Wagner, P. &amp; Greenbaum, E. &amp; Bauer, A. M. &amp; Kusamba, C. &amp; Leache, A. D." box="[475,693,1159,1185]" pageId="10" pageNumber="231" pagination="771 - 817" refId="ref12867" refString="Wagner, P., Greenbaum, E., Bauer, A. M., Kusamba, C. &amp; Leache, A. D. (2018) Lifting the blue-headed veil - integrative taxonomy of the Acanthocercus atricollis species complex (Squamata: Agamidae). Journal of Natural History, 52, 771 - 817. https: // doi. org / 10.1080 / 00222933.2018.1435833" type="journal article" year="2018">
Wagner
<emphasis id="B978EAEE7928FFC7FDC156FBFD85FB2F" box="[569,626,1159,1185]" italics="true" pageId="10" pageNumber="231">et al.</emphasis>
2018
</bibRefCitation>
), whereas in
<taxonomicName id="4C0C4D7F7928FFC7FCB156FBFC32FB2E" authority="Marques &amp; Parrinha &amp; Santos &amp; Bandeira &amp; Butler &amp; Sousa &amp; Bauer &amp; Wagner, 2022" authorityName="Marques &amp; Parrinha &amp; Santos &amp; Bandeira &amp; Butler &amp; Sousa &amp; Bauer &amp; Wagner" authorityYear="2022" box="[841,965,1159,1185]" class="Reptilia" family="Agamidae" genus="Acanthocercus" kingdom="Animalia" order="Squamata" pageId="10" pageNumber="231" phylum="Chordata" rank="species" species="ceriacoi" status="sp. nov.">
<emphasis id="B978EAEE7928FFC7FCB156FBFC32FB2E" box="[841,965,1159,1185]" italics="true" pageId="10" pageNumber="231">A. ceriacoi</emphasis>
</taxonomicName>
<emphasis id="B978EAEE7928FFC7FC3456FBFBD1FB2F" bold="true" box="[972,1062,1159,1185]" pageId="10" pageNumber="231">
<taxonomicNameLabel id="A24B57957928FFC7FC3456FBFBD1FB2F" box="[972,1062,1159,1185]" pageId="10" pageNumber="231" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
the blue of the male is limited to the head.
</paragraph>
<paragraph id="8BB336FC7928FFC7FF6F56B3FDCAFA13" blockId="10.[151,1437,150,1941]" pageId="10" pageNumber="231">
(f)
<taxonomicName id="4C0C4D7F7928FFC7FF3E56B3FE29FB67" authorityName="ZFMK" baseAuthorityName="Klausewitz" baseAuthorityYear="1957" box="[198,478,1231,1257]" class="Reptilia" family="Agamidae" genus="Acanthocercus" kingdom="Animalia" order="Squamata" pageId="10" pageNumber="231" phylum="Chordata" rank="species" species="kiwuensis">
<emphasis id="B978EAEE7928FFC7FF3E56B3FE29FB67" box="[198,478,1231,1257]" italics="true" pageId="10" pageNumber="231">Acanthocercus kiwuensis</emphasis>
</taxonomicName>
has a smaller SVL (mean
<quantity id="4CF49B197928FFC7FCF956B3FCACFB67" box="[769,859,1231,1257]" metricMagnitude="-1" metricUnit="m" metricValue="1.1099999999999999" pageId="10" pageNumber="231" unit="mm" value="111.0">111 mm</quantity>
vs.
<quantity id="4CF49B197928FFC7FC7F56B3FC13FB67" box="[903,996,1231,1257]" metricMagnitude="-1" metricUnit="m" metricValue="1.23" pageId="10" pageNumber="231" unit="mm" value="123.0">123 mm</quantity>
) but greater total length (mean
<quantity id="4CF49B197928FFC7FAB856B3FA6AFB67" box="[1344,1437,1231,1257]" metricMagnitude="-1" metricUnit="m" metricValue="2.9" pageId="10" pageNumber="231" unit="mm" value="290.0">290 mm</quantity>
vs.
<quantity id="4CF49B197928FFC7FF14568EFEBAFA83" box="[236,333,1266,1293]" metricMagnitude="-1" metricUnit="m" metricValue="2.85" pageId="10" pageNumber="231" unit="mm" value="285.0">285 mm</quantity>
).
<taxonomicName id="4C0C4D7F7928FFC7FEA6568FFE1DFA83" authorityName="ZFMK" baseAuthorityName="Klausewitz" baseAuthorityYear="1957" box="[350,490,1267,1293]" class="Reptilia" family="Agamidae" genus="Acanthocercus" kingdom="Animalia" order="Squamata" pageId="10" pageNumber="231" phylum="Chordata" rank="species" species="kiwuensis">
<emphasis id="B978EAEE7928FFC7FEA6568FFE1DFA83" box="[350,490,1267,1293]" italics="true" pageId="10" pageNumber="231">A. kiwuensis</emphasis>
</taxonomicName>
has lower mean counts of scale rows around midbody (87 vs. 108) and ventral scales (67 vs. 85) than
<taxonomicName id="4C0C4D7F7928FFC7FE78576BFE0BFABE" authority="Marques &amp; Parrinha &amp; Santos &amp; Bandeira &amp; Butler &amp; Sousa &amp; Bauer &amp; Wagner, 2022" authorityName="Marques &amp; Parrinha &amp; Santos &amp; Bandeira &amp; Butler &amp; Sousa &amp; Bauer &amp; Wagner" authorityYear="2022" box="[384,508,1303,1329]" class="Reptilia" family="Agamidae" genus="Acanthocercus" kingdom="Animalia" order="Squamata" pageId="10" pageNumber="231" phylum="Chordata" rank="species" species="ceriacoi" status="sp. nov.">
<emphasis id="B978EAEE7928FFC7FE78576BFE0BFABE" box="[384,508,1303,1329]" italics="true" pageId="10" pageNumber="231">A. ceriacoi</emphasis>
</taxonomicName>
<emphasis id="B978EAEE7928FFC7FDFC576BFDA8FABF" bold="true" box="[516,607,1303,1329]" pageId="10" pageNumber="231">
<taxonomicNameLabel id="A24B57957928FFC7FDFC576BFDA8FABF" box="[516,607,1303,1329]" pageId="10" pageNumber="231" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
. The ranges of this character largely overlap in the two species, although they are higher in
<taxonomicName id="4C0C4D7F7928FFC7FE695747FDFBFADA" authority="Marques &amp; Parrinha &amp; Santos &amp; Bandeira &amp; Butler &amp; Sousa &amp; Bauer &amp; Wagner, 2022" authorityName="Marques &amp; Parrinha &amp; Santos &amp; Bandeira &amp; Butler &amp; Sousa &amp; Bauer &amp; Wagner" authorityYear="2022" box="[401,524,1339,1365]" class="Reptilia" family="Agamidae" genus="Acanthocercus" kingdom="Animalia" order="Squamata" pageId="10" pageNumber="231" phylum="Chordata" rank="species" species="ceriacoi" status="sp. nov.">
<emphasis id="B978EAEE7928FFC7FE695747FDFBFADA" box="[401,524,1339,1365]" italics="true" pageId="10" pageNumber="231">A. ceriacoi</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A24B57957928FFC7FDEB5747FD99FADB" box="[531,622,1339,1365]" pageId="10" pageNumber="231" rank="species">
<emphasis id="B978EAEE7928FFC7FDEB5747FD90FADB" bold="true" box="[531,615,1339,1365]" pageId="10" pageNumber="231">sp. nov</emphasis>
.
</taxonomicNameLabel>
than in
<taxonomicName id="4C0C4D7F7928FFC7FD315747FCA2FADB" authorityName="ZFMK" baseAuthorityName="Klausewitz" baseAuthorityYear="1957" box="[713,853,1339,1365]" class="Reptilia" family="Agamidae" genus="Acanthocercus" kingdom="Animalia" order="Squamata" pageId="10" pageNumber="231" phylum="Chordata" rank="species" species="kiwuensis">
<emphasis id="B978EAEE7928FFC7FD315747FCA2FADB" box="[713,853,1339,1365]" italics="true" pageId="10" pageNumber="231">A. kiwuensis</emphasis>
</taxonomicName>
. Moreover
<taxonomicName id="4C0C4D7F7928FFC7FC205747FB93FADB" authorityName="ZFMK" baseAuthorityName="Klausewitz" baseAuthorityYear="1957" box="[984,1124,1339,1365]" class="Reptilia" family="Agamidae" genus="Acanthocercus" kingdom="Animalia" order="Squamata" pageId="10" pageNumber="231" phylum="Chordata" rank="species" species="kiwuensis">
<emphasis id="B978EAEE7928FFC7FC205747FB93FADB" box="[984,1124,1339,1365]" italics="true" pageId="10" pageNumber="231">A. kiwuensis</emphasis>
</taxonomicName>
does not possess transverse rows of enlarged scales (
<bibRefCitation id="EF9D4B0D7928FFC7FE2C5723FD5BFAF7" author="Wagner, P. &amp; Greenbaum, E. &amp; Bauer, A. M. &amp; Kusamba, C. &amp; Leache, A. D." box="[468,684,1375,1401]" pageId="10" pageNumber="231" pagination="771 - 817" refId="ref12867" refString="Wagner, P., Greenbaum, E., Bauer, A. M., Kusamba, C. &amp; Leache, A. D. (2018) Lifting the blue-headed veil - integrative taxonomy of the Acanthocercus atricollis species complex (Squamata: Agamidae). Journal of Natural History, 52, 771 - 817. https: // doi. org / 10.1080 / 00222933.2018.1435833" type="journal article" year="2018">
Wagner
<emphasis id="B978EAEE7928FFC7FDCA5723FD9EFAF7" box="[562,617,1375,1401]" italics="true" pageId="10" pageNumber="231">et al.</emphasis>
2018
</bibRefCitation>
). The mean number of precloacal pores is lower in
<taxonomicName id="4C0C4D7F7928FFC7FB245723FA91FAF7" authorityName="ZFMK" baseAuthorityName="Klausewitz" baseAuthorityYear="1957" box="[1244,1382,1375,1401]" class="Reptilia" family="Agamidae" genus="Acanthocercus" kingdom="Animalia" order="Squamata" pageId="10" pageNumber="231" phylum="Chordata" rank="species" species="kiwuensis">
<emphasis id="B978EAEE7928FFC7FB245723FA91FAF7" box="[1244,1382,1375,1401]" italics="true" pageId="10" pageNumber="231">A. kiwuensis</emphasis>
</taxonomicName>
than in
<taxonomicName id="4C0C4D7F7928FFC7FF1B57FFFEA9FA12" authority="Marques &amp; Parrinha &amp; Santos &amp; Bandeira &amp; Butler &amp; Sousa &amp; Bauer &amp; Wagner, 2022" authorityName="Marques &amp; Parrinha &amp; Santos &amp; Bandeira &amp; Butler &amp; Sousa &amp; Bauer &amp; Wagner" authorityYear="2022" box="[227,350,1411,1437]" class="Reptilia" family="Agamidae" genus="Acanthocercus" kingdom="Animalia" order="Squamata" pageId="10" pageNumber="231" phylum="Chordata" rank="species" species="ceriacoi" status="sp. nov.">
<emphasis id="B978EAEE7928FFC7FF1B57FFFEA9FA12" box="[227,350,1411,1437]" italics="true" pageId="10" pageNumber="231">A. ceriacoi</emphasis>
</taxonomicName>
<emphasis id="B978EAEE7928FFC7FE9D57FFFE49FA13" bold="true" box="[357,446,1411,1437]" pageId="10" pageNumber="231">
<taxonomicNameLabel id="A24B57957928FFC7FE9D57FFFE49FA13" box="[357,446,1411,1437]" pageId="10" pageNumber="231" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
(10 vs 18).
</paragraph>
<paragraph id="8BB336FC7928FFC7FF6F57DBFE68F9DF" blockId="10.[151,1437,150,1941]" pageId="10" pageNumber="231">
(g)
<taxonomicName id="4C0C4D7F7928FFC7FF3E57DBFE07FA4E" authorityName="Wagner, Butler, Ceriaco &amp; Bauer" authorityYear="2021" box="[198,496,1447,1473]" class="Reptilia" family="Agamidae" genus="Acanthocercus" kingdom="Animalia" order="Squamata" pageId="10" pageNumber="231" phylum="Chordata" rank="species" species="margaritae">
<emphasis id="B978EAEE7928FFC7FF3E57DBFE07FA4E" box="[198,496,1447,1473]" italics="true" pageId="10" pageNumber="231">Acanthocercus margaritae</emphasis>
</taxonomicName>
has a smaller SVL (mean
<quantity id="4CF49B197928FFC7FCE357DBFC80FA4F" box="[795,887,1447,1473]" metricMagnitude="-1" metricUnit="m" metricValue="1.1199999999999999" pageId="10" pageNumber="231" unit="mm" value="112.0">112 mm</quantity>
vs.
<quantity id="4CF49B197928FFC7FC5E57DBFBF3FA4F" box="[934,1028,1447,1473]" metricMagnitude="-1" metricUnit="m" metricValue="1.23" pageId="10" pageNumber="231" unit="mm" value="123.0">123 mm</quantity>
) and total length (mean
<quantity id="4CF49B197928FFC7FAE057DAFA82FA4F" box="[1304,1397,1446,1473]" metricMagnitude="-1" metricUnit="m" metricValue="2.58" pageId="10" pageNumber="231" unit="mm" value="258.0">258 mm</quantity>
vs.
<quantity id="4CF49B197928FFC7FF3E57B6FED1FA6B" box="[198,294,1482,1509]" metricMagnitude="-1" metricUnit="m" metricValue="2.85" pageId="10" pageNumber="231" unit="mm" value="285.0">285 mm</quantity>
). It has higher mean counts of scale rows around midbody (136 vs. 108), dorsal scales (103 vs. 63), and ventral scales (101 vs. 85) than
<taxonomicName id="4C0C4D7F7928FFC7FDD15793FD53F986" authority="Marques &amp; Parrinha &amp; Santos &amp; Bandeira &amp; Butler &amp; Sousa &amp; Bauer &amp; Wagner, 2022" authorityName="Marques &amp; Parrinha &amp; Santos &amp; Bandeira &amp; Butler &amp; Sousa &amp; Bauer &amp; Wagner" authorityYear="2022" box="[553,676,1519,1545]" class="Reptilia" family="Agamidae" genus="Acanthocercus" kingdom="Animalia" order="Squamata" pageId="10" pageNumber="231" phylum="Chordata" rank="species" species="ceriacoi" status="sp. nov.">
<emphasis id="B978EAEE7928FFC7FDD15793FD53F986" box="[553,676,1519,1545]" italics="true" pageId="10" pageNumber="231">A. ceriacoi</emphasis>
</taxonomicName>
<emphasis id="B978EAEE7928FFC7FD535793FCF2F987" bold="true" box="[683,773,1519,1545]" pageId="10" pageNumber="231">
<taxonomicNameLabel id="A24B57957928FFC7FD535793FCF2F987" box="[683,773,1519,1545]" pageId="10" pageNumber="231" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
. The ranges of these characters are lower in
<taxonomicName id="4C0C4D7F7928FFC7FB015793FA83F986" authority="Marques &amp; Parrinha &amp; Santos &amp; Bandeira &amp; Butler &amp; Sousa &amp; Bauer &amp; Wagner, 2022" authorityName="Marques &amp; Parrinha &amp; Santos &amp; Bandeira &amp; Butler &amp; Sousa &amp; Bauer &amp; Wagner" authorityYear="2022" box="[1273,1396,1519,1545]" class="Reptilia" family="Agamidae" genus="Acanthocercus" kingdom="Animalia" order="Squamata" pageId="10" pageNumber="231" phylum="Chordata" rank="species" species="ceriacoi" status="sp. nov.">
<emphasis id="B978EAEE7928FFC7FB015793FA83F986" box="[1273,1396,1519,1545]" italics="true" pageId="10" pageNumber="231">A. ceriacoi</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A24B57957928FFC7FA835793FF0FF9A3" pageId="10" pageNumber="231" rank="species">
<emphasis id="B978EAEE7928FFC7FA835793FF05F9A3" bold="true" pageId="10" pageNumber="231">sp. nov</emphasis>
.
</taxonomicNameLabel>
than in
<emphasis id="B978EAEE7928FFC7FEAC546FFE00F9A2" box="[340,503,1555,1581]" italics="true" pageId="10" pageNumber="231">
<taxonomicName id="4C0C4D7F7928FFC7FEAC546FFE03F9A2" authorityName="Wagner, Butler, Ceriaco &amp; Bauer" authorityYear="2021" box="[340,500,1555,1581]" class="Reptilia" family="Agamidae" genus="Acanthocercus" kingdom="Animalia" order="Squamata" pageId="10" pageNumber="231" phylum="Chordata" rank="species" species="margaritae">A. margaritae</taxonomicName>
.
</emphasis>
The mean number of precloacal pores is lower in
<taxonomicName id="4C0C4D7F7928FFC7FBD2546FFB31F9A2" authorityName="Wagner, Butler, Ceriaco &amp; Bauer" authorityYear="2021" box="[1066,1222,1555,1581]" class="Reptilia" family="Agamidae" genus="Acanthocercus" kingdom="Animalia" order="Squamata" pageId="10" pageNumber="231" phylum="Chordata" rank="species" species="margaritae">
<emphasis id="B978EAEE7928FFC7FBD2546FFB31F9A2" box="[1066,1222,1555,1581]" italics="true" pageId="10" pageNumber="231">A. margaritae</emphasis>
</taxonomicName>
than in
<taxonomicName id="4C0C4D7F7928FFC7FAD9546FFA6BF9A2" authority="Marques &amp; Parrinha &amp; Santos &amp; Bandeira &amp; Butler &amp; Sousa &amp; Bauer &amp; Wagner, 2022" authorityName="Marques &amp; Parrinha &amp; Santos &amp; Bandeira &amp; Butler &amp; Sousa &amp; Bauer &amp; Wagner" authorityYear="2022" box="[1313,1436,1555,1581]" class="Reptilia" family="Agamidae" genus="Acanthocercus" kingdom="Animalia" order="Squamata" pageId="10" pageNumber="231" phylum="Chordata" rank="species" species="ceriacoi" status="sp. nov.">
<emphasis id="B978EAEE7928FFC7FAD9546FFA6BF9A2" box="[1313,1436,1555,1581]" italics="true" pageId="10" pageNumber="231">A. ceriacoi</emphasis>
</taxonomicName>
<emphasis id="B978EAEE7928FFC7FF3E544BFEE8F9DF" bold="true" box="[198,287,1591,1617]" pageId="10" pageNumber="231">
<taxonomicNameLabel id="A24B57957928FFC7FF3E544BFEE8F9DF" box="[198,287,1591,1617]" pageId="10" pageNumber="231" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
(14 vs 18).
</paragraph>
<paragraph id="8BB336FC7928FFC7FF6F5427FE81F96F" blockId="10.[151,1437,150,1941]" pageId="10" pageNumber="231">
(h)
<taxonomicName id="4C0C4D7F7928FFC7FF3E5427FE3CF9FA" baseAuthorityName="Klausewitz" baseAuthorityYear="1957" box="[198,459,1627,1653]" class="Reptilia" family="Agamidae" genus="Acanthocercus" kingdom="Animalia" order="Squamata" pageId="10" pageNumber="231" phylum="Chordata" rank="species" species="minutus">
<emphasis id="B978EAEE7928FFC7FF3E5427FE3CF9FA" box="[198,459,1627,1653]" italics="true" pageId="10" pageNumber="231">Acanthocercus minutus</emphasis>
</taxonomicName>
has a smaller SVL (mean
<quantity id="4CF49B197928FFC7FD095427FCB9F9FB" box="[753,846,1627,1653]" metricMagnitude="-1" metricUnit="m" metricValue="1.01" pageId="10" pageNumber="231" unit="mm" value="101.0">101 mm</quantity>
vs.
<quantity id="4CF49B197928FFC7FC835427FC2EF9FB" box="[891,985,1627,1653]" metricMagnitude="-1" metricUnit="m" metricValue="1.23" pageId="10" pageNumber="231" unit="mm" value="123.0">123 mm</quantity>
) and total length (mean
<quantity id="4CF49B197928FFC7FB105427FAB2F9FB" box="[1256,1349,1627,1653]" metricMagnitude="-1" metricUnit="m" metricValue="2.34" pageId="10" pageNumber="231" unit="mm" value="234.0">234 mm</quantity>
vs.
<quantity id="4CF49B197928FFC7FA8A5426FF01F917" metricMagnitude="-1" metricUnit="m" metricValue="2.85" pageId="10" pageNumber="231" unit="mm" value="285.0">285 mm</quantity>
). It has higher mean counts of dorsal scales (79 vs. 63), and lower mean counts of ventral scales (78 vs. 85) than
<taxonomicName id="4C0C4D7F7928FFC7FF0654DFFE8EF932" authority="Marques &amp; Parrinha &amp; Santos &amp; Bandeira &amp; Butler &amp; Sousa &amp; Bauer &amp; Wagner, 2022" authorityName="Marques &amp; Parrinha &amp; Santos &amp; Bandeira &amp; Butler &amp; Sousa &amp; Bauer &amp; Wagner" authorityYear="2022" box="[254,377,1699,1725]" class="Reptilia" family="Agamidae" genus="Acanthocercus" kingdom="Animalia" order="Squamata" pageId="10" pageNumber="231" phylum="Chordata" rank="species" species="ceriacoi" status="sp. nov.">
<emphasis id="B978EAEE7928FFC7FF0654DFFE8EF932" box="[254,377,1699,1725]" italics="true" pageId="10" pageNumber="231">A. ceriacoi</emphasis>
</taxonomicName>
<emphasis id="B978EAEE7928FFC7FE7954DFFE2CF933" bold="true" box="[385,475,1699,1725]" pageId="10" pageNumber="231">
<taxonomicNameLabel id="A24B57957928FFC7FE7954DFFE2CF933" box="[385,475,1699,1725]" pageId="10" pageNumber="231" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
. The mean number of precloacal pores is higher in
<taxonomicName id="4C0C4D7F7928FFC7FBDA54DFFB6CF932" baseAuthorityName="Klausewitz" baseAuthorityYear="1957" box="[1058,1179,1699,1725]" class="Reptilia" family="Agamidae" genus="Acanthocercus" kingdom="Animalia" order="Squamata" pageId="10" pageNumber="231" phylum="Chordata" rank="species" species="minutus">
<emphasis id="B978EAEE7928FFC7FBDA54DFFB6CF932" box="[1058,1179,1699,1725]" italics="true" pageId="10" pageNumber="231">A. minutus</emphasis>
</taxonomicName>
than in
<taxonomicName id="4C0C4D7F7928FFC7FB0054DFFA84F932" authority="Marques &amp; Parrinha &amp; Santos &amp; Bandeira &amp; Butler &amp; Sousa &amp; Bauer &amp; Wagner, 2022" authorityName="Marques &amp; Parrinha &amp; Santos &amp; Bandeira &amp; Butler &amp; Sousa &amp; Bauer &amp; Wagner" authorityYear="2022" box="[1272,1395,1699,1725]" class="Reptilia" family="Agamidae" genus="Acanthocercus" kingdom="Animalia" order="Squamata" pageId="10" pageNumber="231" phylum="Chordata" rank="species" species="ceriacoi" status="sp. nov.">
<emphasis id="B978EAEE7928FFC7FB0054DFFA84F932" box="[1272,1395,1699,1725]" italics="true" pageId="10" pageNumber="231">A. ceriacoi</emphasis>
</taxonomicName>
<emphasis id="B978EAEE7928FFC7FA8354DFFF00F96F" bold="true" pageId="10" pageNumber="231">
<taxonomicNameLabel id="A24B57957928FFC7FA8354DFFF00F96F" pageId="10" pageNumber="231" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
(26 vs 18).
</paragraph>
<paragraph id="8BB336FC7928FFC7FF6F5497FEE5F81B" blockId="10.[151,1437,150,1941]" pageId="10" pageNumber="231">
(i)
<taxonomicName id="4C0C4D7F7928FFC7FF3E5497FE09F88B" authorityName="JB" baseAuthorityName="Klausewitz" baseAuthorityYear="1957" box="[198,510,1771,1797]" class="Reptilia" family="Agamidae" genus="Acanthocercus" kingdom="Animalia" order="Squamata" pageId="10" pageNumber="231" phylum="Chordata" rank="species" species="ugandaensis">
<emphasis id="B978EAEE7928FFC7FF3E5497FE09F88B" box="[198,510,1771,1797]" italics="true" pageId="10" pageNumber="231">Acanthocercus ugandaensis</emphasis>
</taxonomicName>
has a smaller SVL (mean
<quantity id="4CF49B197928FFC7FCDC5497FC77F88B" box="[804,896,1771,1797]" metricMagnitude="-1" metricUnit="m" metricValue="1.03" pageId="10" pageNumber="231" unit="mm" value="103.0">103 mm</quantity>
vs.
<quantity id="4CF49B197928FFC7FC555497FBFDF88B" box="[941,1034,1771,1797]" metricMagnitude="-1" metricUnit="m" metricValue="1.23" pageId="10" pageNumber="231" unit="mm" value="123.0">123 mm</quantity>
) and total length (mean
<quantity id="4CF49B197928FFC7FAE25496FA81F88B" box="[1306,1398,1770,1797]" metricMagnitude="-1" metricUnit="m" metricValue="2.75" pageId="10" pageNumber="231" unit="mm" value="275.0">275 mm</quantity>
vs.
<quantity id="4CF49B197928FFC7FF3E5572FED0F8A7" box="[198,295,1806,1833]" metricMagnitude="-1" metricUnit="m" metricValue="2.85" pageId="10" pageNumber="231" unit="mm" value="285.0">285 mm</quantity>
).
<taxonomicName id="4C0C4D7F7928FFC7FEC15573FE13F8A7" authorityName="JB" baseAuthorityName="Klausewitz" baseAuthorityYear="1957" box="[313,484,1807,1833]" class="Reptilia" family="Agamidae" genus="Acanthocercus" kingdom="Animalia" order="Squamata" pageId="10" pageNumber="231" phylum="Chordata" rank="species" species="ugandaensis">
<emphasis id="B978EAEE7928FFC7FEC15573FE13F8A7" box="[313,484,1807,1833]" italics="true" pageId="10" pageNumber="231">A. ugandaensis</emphasis>
</taxonomicName>
has lower mean counts of scale rows around midbody (94 vs. 108) and ventral scales (73 vs. 85) than
<taxonomicName id="4C0C4D7F7928FFC7FE84554FFE01F8C2" authority="Marques &amp; Parrinha &amp; Santos &amp; Bandeira &amp; Butler &amp; Sousa &amp; Bauer &amp; Wagner, 2022" authorityName="Marques &amp; Parrinha &amp; Santos &amp; Bandeira &amp; Butler &amp; Sousa &amp; Bauer &amp; Wagner" authorityYear="2022" box="[380,502,1843,1869]" class="Reptilia" family="Agamidae" genus="Acanthocercus" kingdom="Animalia" order="Squamata" pageId="10" pageNumber="231" phylum="Chordata" rank="species" species="ceriacoi" status="sp. nov.">
<emphasis id="B978EAEE7928FFC7FE84554FFE01F8C2" box="[380,502,1843,1869]" italics="true" pageId="10" pageNumber="231">A. ceriacoi</emphasis>
</taxonomicName>
<emphasis id="B978EAEE7928FFC7FE05554FFDA1F8C3" bold="true" box="[509,598,1843,1869]" pageId="10" pageNumber="231">
<taxonomicNameLabel id="A24B57957928FFC7FE05554FFDA1F8C3" box="[509,598,1843,1869]" pageId="10" pageNumber="231" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
. The ranges of these characters are higher in
<taxonomicName id="4C0C4D7F7928FFC7FBB7554FFB3DF8C2" authority="Marques &amp; Parrinha &amp; Santos &amp; Bandeira &amp; Butler &amp; Sousa &amp; Bauer &amp; Wagner, 2022" authorityName="Marques &amp; Parrinha &amp; Santos &amp; Bandeira &amp; Butler &amp; Sousa &amp; Bauer &amp; Wagner" authorityYear="2022" box="[1103,1226,1843,1869]" class="Reptilia" family="Agamidae" genus="Acanthocercus" kingdom="Animalia" order="Squamata" pageId="10" pageNumber="231" phylum="Chordata" rank="species" species="ceriacoi" status="sp. nov.">
<emphasis id="B978EAEE7928FFC7FBB7554FFB3DF8C2" box="[1103,1226,1843,1869]" italics="true" pageId="10" pageNumber="231">A. ceriacoi</emphasis>
</taxonomicName>
<emphasis id="B978EAEE7928FFC7FB29554FFADDF8C3" bold="true" box="[1233,1322,1843,1869]" pageId="10" pageNumber="231">
<taxonomicNameLabel id="A24B57957928FFC7FB29554FFADDF8C3" box="[1233,1322,1843,1869]" pageId="10" pageNumber="231" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
than in
<taxonomicName id="4C0C4D7F7928FFC7FA7C554FFEA5F8FF" authorityName="JB" baseAuthorityName="Klausewitz" baseAuthorityYear="1957" class="Reptilia" family="Agamidae" genus="Acanthocercus" kingdom="Animalia" order="Squamata" pageId="10" pageNumber="231" phylum="Chordata" rank="species" species="ugandaensis">
<emphasis id="B978EAEE7928FFC7FA7C554FFEA5F8FF" italics="true" pageId="10" pageNumber="231">A. ugandaensis</emphasis>
</taxonomicName>
. The mean number of precloacal pores is higher in
<taxonomicName id="4C0C4D7F7928FFC7FC69552BFBCCF8FF" authorityName="JB" baseAuthorityName="Klausewitz" baseAuthorityYear="1957" box="[913,1083,1879,1905]" class="Reptilia" family="Agamidae" genus="Acanthocercus" kingdom="Animalia" order="Squamata" pageId="10" pageNumber="231" phylum="Chordata" rank="species" species="ugandaensis">
<emphasis id="B978EAEE7928FFC7FC69552BFBCCF8FF" box="[913,1083,1879,1905]" italics="true" pageId="10" pageNumber="231">A. ugandaensis</emphasis>
</taxonomicName>
than in
<taxonomicName id="4C0C4D7F7928FFC7FB6E552BFAE7F8FE" authority="Marques &amp; Parrinha &amp; Santos &amp; Bandeira &amp; Butler &amp; Sousa &amp; Bauer &amp; Wagner, 2022" authorityName="Marques &amp; Parrinha &amp; Santos &amp; Bandeira &amp; Butler &amp; Sousa &amp; Bauer &amp; Wagner" authorityYear="2022" box="[1174,1296,1879,1905]" class="Reptilia" family="Agamidae" genus="Acanthocercus" kingdom="Animalia" order="Squamata" pageId="10" pageNumber="231" phylum="Chordata" rank="species" species="ceriacoi" status="sp. nov.">
<emphasis id="B978EAEE7928FFC7FB6E552BFAE7F8FE" box="[1174,1296,1879,1905]" italics="true" pageId="10" pageNumber="231">A. ceriacoi</emphasis>
</taxonomicName>
<emphasis id="B978EAEE7928FFC7FAEF552BFA87F8FF" bold="true" box="[1303,1392,1879,1905]" pageId="10" pageNumber="231">
<taxonomicNameLabel id="A24B57957928FFC7FAEF552BFA87F8FF" box="[1303,1392,1879,1905]" pageId="10" pageNumber="231" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
(24 vs 18).
</paragraph>
</subSubSection>
<subSubSection id="C31665777929FFC1FF3F52EAFDECFE93" lastPageId="12" lastPageNumber="233" pageId="11" pageNumber="232" type="description">
<paragraph id="8BB336FC7929FFC6FF3F52EAFE6DFA19" blockId="11.[151,1437,150,2040]" pageId="11" pageNumber="232">
<emphasis id="B978EAEE7929FFC6FF3F52EAFDF8FF3F" bold="true" box="[199,527,150,177]" pageId="11" pageNumber="232">
Description of the
<typeStatus id="54B7885E7929FFC6FE5952EBFDFDFF3F" box="[417,522,151,177]" pageId="11" pageNumber="232" type="holotype">holotype</typeStatus>
.
</emphasis>
Adult male (
<figureCitation id="13372A797929FFC6FD6652EBFD14FF3F" box="[670,739,151,177]" captionStart="FIGURE 2" captionStartId="5.[151,250,1986,2010]" captionTargetBox="[190,1399,444,1960]" captionTargetId="figure-110@5.[189,1399,435,1962]" captionTargetPageId="5" captionText="FIGURE 2. Holotype of Acanthocercus ceriacoi sp. nov. (CAS 258430). Photos by Erica Ely." figureDoi="http://doi.org/10.5281/zenodo.6048532" httpUri="https://zenodo.org/record/6048532/files/figure.png" pageId="11" pageNumber="232">Fig. 2</figureCitation>
). Measurements: snoutvent length (SVL)
<quantity id="4CF49B197929FFC6FB3B52EBFAD6FF3F" box="[1219,1313,151,177]" metricMagnitude="-1" metricUnit="m" metricValue="1.43" pageId="11" pageNumber="232" unit="mm" value="143.0">143 mm</quantity>
, tail length (TL)
<quantity id="4CF49B197929FFC6FF2B52C7FEC5FF5B" box="[211,306,187,213]" metricMagnitude="-1" metricUnit="m" metricValue="1.8699999999999999" pageId="11" pageNumber="232" unit="mm" value="187.0">187 mm</quantity>
, head length (HL)
<quantity id="4CF49B197929FFC6FDFE52C7FDAFFF5B" box="[518,600,187,213]" metricMagnitude="-2" metricUnit="m" metricValue="4.8" pageId="11" pageNumber="232" unit="mm" value="48.0">48 mm</quantity>
, head width (HW)
<quantity id="4CF49B197929FFC6FCD752C7FC77FF5B" box="[815,896,187,213]" metricMagnitude="-2" metricUnit="m" metricValue="3.8" pageId="11" pageNumber="232" unit="mm" value="38.0">38 mm</quantity>
, head height (
<collectingRegion id="49C8F81E7929FFC6FBE552C7FBBFFF5B" box="[1053,1096,187,213]" country="Swaziland" name="Hhohho" pageId="11" pageNumber="232">HH</collectingRegion>
)
<quantity id="4CF49B197929FFC6FBAF52C7FB5EFF5B" box="[1111,1193,187,213]" metricMagnitude="-2" metricUnit="m" metricValue="3.4" pageId="11" pageNumber="232" unit="mm" value="34.0">34 mm</quantity>
. Habitus stout, with a large triangular head distinct from body; nasal scale flat to faintly convex, smooth, oval and situated slightly below canthus rostralis, pierced by round nostril in posterior part of scale, directed laterally. Scales on anterior, lateral, and central parts of head large, but smaller from behind ear opening, half the size of large head scales; head scales unequal in size, not oriented in the same direction, usually smooth, sporadically heavily keeled or with a rough surface, with free anterior margins and only sporadically with sensory pits. Fourteen supralabial scales and twelve infralabial scales on both sides; supraocular scales smooth, twelve scales along the canthus rostralis from nasal to behind the eye; six scales from nasal to anterior margin of eye; parietal shield quadrangular; pineal organ small, visible in posterior part of parietal shield. Ear opening large, but smaller than eye, posterior margin lacking semicircle of spinose mucronate scales; tympanum superficial. No nuchal crest present. Gular scales smooth, flat to faintly convex in the anterior part of the gular area; mucronate and spinose in the posterior part, becoming more spiny and smaller towards gular fold. Dorsal body scales a mix of small and smooth matrix scales, sometimes giving the impression of granular scales, and scattered, pale white-edged, enlarged scales. Large scales keeled, sometimes mucronate or spinose, not arranged in clusters, but rather in seven to eight, very indistinct, transverse rows between limbs. Sixty-four dorsal scales along vertebral column from midpoint of pectoral region to midpoint of pelvic region. Vertebral region covered by a mix of small and large scales, forming a pattern distinct from rest of lateral parts of body, with a large set of vertebral scales strongly keeled to smooth or feebly keeled. Ventral body scales smooth, slightly imbricate at posterior margins, in 89 rows from midpoint of pectoral region to precloacal pores. One hundred twenty-six scale rows around body behind forelimbs; 105 around midbody and 80 around body in front of hind limbs. Precloacal scales in two rows (the more anterior weakly developed), twelve pores in the posterior row and seven pores in the row above. Scales on upper side of forelimb unequal in size and strongly keeled, smooth on underside, upper arm scales somewhat larger than largest dorsal body scales. Relative length of manual digits 4=3&gt;2&gt;5&gt;1, subdigital lamellae keeled and mucronate, 21 under left 4
<superScript id="7C799BB47929FFC6FD7251ACFD60FC50" attach="left" box="[650,663,976,990]" fontSize="6" pageId="11" pageNumber="232">th</superScript>
finger. Scales on upper side of hind limb keeled to smooth, becoming completely smooth on underside; scales on upper thighs unequal in size, a mix of small and scattered enlarged scales that are as large as the enlarged dorsal body scales. Fourth toe longest, relative length of pedal digits 4&gt;3&gt;2&gt;5&gt;1, subdigital lamellae keeled and mucronate, 23 under left 4
<superScript id="7C799BB47929FFC6FCDA5647FCD8FBC7" attach="left" box="[802,815,1083,1097]" fontSize="6" pageId="11" pageNumber="232">th</superScript>
toe. Tail with indistinct whorls of five scale rings (one ring consists of distinctly smaller scales) in its basal portion, caudal scales keeled and mucronate. First third of tail slightly swollen, scales larger than in other parts of the body, strongly robust and keeled. Distal portion of the tail much thinner and slightly depressed. After formalin fixation and six years of preservation in ethanol (
<figureCitation id="13372A797929FFC6FAEB56DBFAA0FB4F" box="[1299,1367,1191,1217]" captionStart="FIGURE 2" captionStartId="5.[151,250,1986,2010]" captionTargetBox="[190,1399,444,1960]" captionTargetId="figure-110@5.[189,1399,435,1962]" captionTargetPageId="5" captionText="FIGURE 2. Holotype of Acanthocercus ceriacoi sp. nov. (CAS 258430). Photos by Erica Ely." figureDoi="http://doi.org/10.5281/zenodo.6048532" httpUri="https://zenodo.org/record/6048532/files/figure.png" pageId="11" pageNumber="232">Fig. 2</figureCitation>
) head dark with some brown to orange scales. Gular region bluish and dark from the mental through the throat, whitish in lateral parts. Black patch on shoulders. Ground color of dorsum grayish to brownish, enlarged scales on upper and lateral parts of body and limbs speckled dirty white. Four to five brownish areola-like markings delimited by white pale circles are visible through the vertebral region. Ventral side of body, limbs and tail dirty grayish, speckled with dark spots. Enlarged scales on upper side of tail base distinctly whitish. Scales becoming more brownish-dirty white striped towards tail tip.
</paragraph>
<paragraph id="8BB336FC7929FFC6FF3757DCFE74F83E" blockId="11.[151,1437,150,2040]" pageId="11" pageNumber="232">
<emphasis id="B978EAEE7929FFC6FF3757DCFEA0FA34" bold="true" box="[207,343,1440,1466]" pageId="11" pageNumber="232">Coloration.</emphasis>
In displaying males (see
<figureCitation id="13372A797929FFC6FD8257DDFD4AFA35" box="[634,701,1441,1467]" captionStart="FIGURE 3" captionStartId="6.[151,250,1752,1776]" captionTargetBox="[255,1333,181,1729]" captionTargetId="figure-23@6.[255,1333,181,1729]" captionTargetPageId="6" captionText="FIGURE 3. Life photo of the holotype (CAS 258430), an adult male of Acanthocercus ceriacoi sp. nov. from Cangandala National Park. Photo by Luis M.P. Ceríaco." figureDoi="http://doi.org/10.5281/zenodo.6048534" httpUri="https://zenodo.org/record/6048534/files/figure.png" pageId="11" pageNumber="232">Fig. 3</figureCitation>
), head vibrantly blue; neck, shoulders, body and limbs brownish with a pattern of white and black stripes and speckles; a pale vertebral band from the neck to the tail is present, distinct anteriorly between the flanks, becoming more indistinct and on a brown background posteriorly; enlarged dorsal scales of the lower part of the body yellowish, matrix scales greyish becoming brownish on the hind limbs; first half of the tail yellowish, second half of the tail with dark and lighter blue bands. Ventral parts of the body, limbs and tail dirty whitish with a brown patterning coloration on females in life (see
<figureCitation id="13372A797929FFC6FB9A542FFB51F9E0" box="[1122,1190,1619,1646]" captionStart="FIGURE 5" captionStartId="8.[151,250,1806,1830]" captionTargetBox="[151,1436,181,1782]" captionTargetId="figure-23@8.[151,1436,181,1782]" captionTargetPageId="8" captionText="FIGURE 5. Life photos of an adult female of Acanthocercus ceriacoi sp. nov. (Paratype, MHNCUP/REP 860) from Uíge Province. Photos by Luis M.P. Ceríaco." figureDoi="http://doi.org/10.5281/zenodo.6048540" httpUri="https://zenodo.org/record/6048540/files/figure.png" pageId="11" pageNumber="232">Fig. 5</figureCitation>
). Shoulders brownish with a black patch, some individuals present a pale blue head. Four dark brown, broad transverse bands between limbs, each interspersed by transverse bands composed of a single row of enlarged yellow scales. Upper and lateral parts of tail with brown and whitish/yellowish bands. Ventral parts of the body, limbs and tail dirty whitish with an extensive dark brown pattern. Gular region of a vibrant blue. Regarding its Angolan congeners, the coloration of displaying males (
<figureCitation id="13372A797929FFC6FE9B557AFE51F8AE" box="[355,422,1798,1824]" captionStart="FIGURE 3" captionStartId="6.[151,250,1752,1776]" captionTargetBox="[255,1333,181,1729]" captionTargetId="figure-23@6.[255,1333,181,1729]" captionTargetPageId="6" captionText="FIGURE 3. Life photo of the holotype (CAS 258430), an adult male of Acanthocercus ceriacoi sp. nov. from Cangandala National Park. Photo by Luis M.P. Ceríaco." figureDoi="http://doi.org/10.5281/zenodo.6048534" httpUri="https://zenodo.org/record/6048534/files/figure.png" pageId="11" pageNumber="232">Fig. 3</figureCitation>
) is overall more similar to
<taxonomicName id="4C0C4D7F7929FFC6FD2E557AFC6CF8AE" authorityName="HF" baseAuthorityName="Falk" baseAuthorityYear="1925" box="[726,923,1798,1824]" class="Reptilia" family="Agamidae" genus="Acanthocercus" kingdom="Animalia" order="Squamata" pageId="11" pageNumber="232" phylum="Chordata" rank="species" species="cyanocephalus">
<emphasis id="B978EAEE7929FFC6FD2E557AFC6CF8AE" box="[726,923,1798,1824]" italics="true" pageId="11" pageNumber="232">A. cyanocephalus</emphasis>
</taxonomicName>
than to
<taxonomicName id="4C0C4D7F7929FFC6FC0F557AFB64F8AE" authorityName="Wagner, Butler, Ceriaco &amp; Bauer" authorityYear="2021" box="[1015,1171,1798,1824]" class="Reptilia" family="Agamidae" genus="Acanthocercus" kingdom="Animalia" order="Squamata" pageId="11" pageNumber="232" phylum="Chordata" rank="species" species="margaritae">
<emphasis id="B978EAEE7929FFC6FC0F557AFB64F8AE" box="[1015,1171,1798,1824]" italics="true" pageId="11" pageNumber="232">A. margaritae</emphasis>
</taxonomicName>
. However, in
<taxonomicName id="4C0C4D7F7929FFC6FAC9557AFF0CF8CA" authorityName="HF" baseAuthorityName="Falk" baseAuthorityYear="1925" class="Reptilia" family="Agamidae" genus="Acanthocercus" kingdom="Animalia" order="Squamata" pageId="11" pageNumber="232" phylum="Chordata" rank="species" species="cyanocephalus">
<emphasis id="B978EAEE7929FFC6FAC9557AFF0CF8CA" italics="true" pageId="11" pageNumber="232">A. cyanocephalus</emphasis>
</taxonomicName>
, the blue coloration of the head extends to the neck and shoulders, while in
<taxonomicName id="4C0C4D7F7929FFC6FBA45556FB2FF8CA" authority="Marques &amp; Parrinha &amp; Santos &amp; Bandeira &amp; Butler &amp; Sousa &amp; Bauer &amp; Wagner, 2022" authorityName="Marques &amp; Parrinha &amp; Santos &amp; Bandeira &amp; Butler &amp; Sousa &amp; Bauer &amp; Wagner" authorityYear="2022" box="[1116,1240,1834,1860]" class="Reptilia" family="Agamidae" genus="Acanthocercus" kingdom="Animalia" order="Squamata" pageId="11" pageNumber="232" phylum="Chordata" rank="species" species="ceriacoi" status="sp. nov.">
<emphasis id="B978EAEE7929FFC6FBA45556FB2FF8CA" box="[1116,1240,1834,1860]" italics="true" pageId="11" pageNumber="232">A. ceriacoi</emphasis>
</taxonomicName>
<emphasis id="B978EAEE7929FFC6FB185556FACDF8CA" bold="true" box="[1248,1338,1834,1860]" pageId="11" pageNumber="232">
<taxonomicNameLabel id="A24B57957929FFC6FB185556FACDF8CA" box="[1248,1338,1834,1860]" pageId="11" pageNumber="232" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
the blue is limited to the head. The new species is clearly distinct from
<taxonomicName id="4C0C4D7F7929FFC6FC9D5532FBF4F8E9" authorityName="Wagner, Butler, Ceriaco &amp; Bauer" authorityYear="2021" box="[869,1027,1870,1896]" class="Reptilia" family="Agamidae" genus="Acanthocercus" kingdom="Animalia" order="Squamata" pageId="11" pageNumber="232" phylum="Chordata" rank="species" species="margaritae">
<emphasis id="B978EAEE7929FFC6FC9D5532FBF4F8E9" box="[869,1027,1870,1896]" italics="true" pageId="11" pageNumber="232">A. margaritae</emphasis>
</taxonomicName>
, in that the latter species has a blue coloration of the head, forelimbs, flanks of the upper part of the body and second half of the tail (not banded as in
<taxonomicName id="4C0C4D7F7929FFC6FF6F55E9FEE5F821" authority="Marques &amp; Parrinha &amp; Santos &amp; Bandeira &amp; Butler &amp; Sousa &amp; Bauer &amp; Wagner, 2022" authorityName="Marques &amp; Parrinha &amp; Santos &amp; Bandeira &amp; Butler &amp; Sousa &amp; Bauer &amp; Wagner" authorityYear="2022" box="[151,274,1941,1967]" class="Reptilia" family="Agamidae" genus="Acanthocercus" kingdom="Animalia" order="Squamata" pageId="11" pageNumber="232" phylum="Chordata" rank="species" species="ceriacoi" status="sp. nov.">
<emphasis id="B978EAEE7929FFC6FF6F55E9FEE5F821" box="[151,274,1941,1967]" italics="true" pageId="11" pageNumber="232">A. ceriacoi</emphasis>
</taxonomicName>
<emphasis id="B978EAEE7929FFC6FEE155E9FE84F821" bold="true" box="[281,371,1941,1967]" pageId="11" pageNumber="232">
<taxonomicNameLabel id="A24B57957929FFC6FEE155E9FE84F821" box="[281,371,1941,1967]" pageId="11" pageNumber="232" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
).
</paragraph>
<paragraph id="8BB336FC7929FFC1FF3F55C5FDECFE93" blockId="11.[151,1437,150,2040]" lastBlockId="12.[151,1437,151,429]" lastPageId="12" lastPageNumber="233" pageId="11" pageNumber="232">
<emphasis id="B978EAEE7929FFC6FF3F55C5FEC8F85D" bold="true" box="[199,319,1977,2003]" pageId="11" pageNumber="232">Variation.</emphasis>
Some variations of the
<typeStatus id="54B7885E7929FFC6FDB555C6FD8AF85A" box="[589,637,1978,2004]" pageId="11" pageNumber="232">type</typeStatus>
series are shown in
<tableCitation id="C68E03477929FFC6FC9B55C5FC4DF85D" box="[867,954,1977,2003]" captionStart="TABLE 2" captionStartId="9.[151,239,1376,1400]" captionTargetPageId="9" captionText="TABLE 2. Data from all taxa (except A. ceriacoi sp. nov.) from Wagner et al. (2021). Data presented as: minmax (mean). Abbreviations: Dorsal scale rows at midbody (SaM) Dorsal scale numbers (D); Ventral scale numbers (V); Total number of precloacal pores (PP)." httpUri="http://table.plazi.org/id/DF736674792BFFC4FF6F571CFE00FA4F" pageId="11" pageNumber="232" tableUuid="DF736674792BFFC4FF6F571CFE00FA4F">Table 2</tableCitation>
. Snout-vent length
<quantity id="4CF49B197929FFC6FB6E55C5FAEAF85A" box="[1174,1309,1977,2004]" metricMagnitude="-1" metricUnit="m" metricValue="1.21" metricValueMax="1.43" metricValueMin="0.99" pageId="11" pageNumber="232" unit="mm" value="121.0" valueMax="143.0" valueMin="99.0">99143 mm</quantity>
(mean
<quantity id="4CF49B197929FFC6FA8A55C5FF30F876" metricMagnitude="-1" metricUnit="m" metricValue="1.23" pageId="11" pageNumber="232" unit="mm" value="123.0">123 mm</quantity>
); Tail length
<quantity id="4CF49B197929FFC6FEA355A1FE06F876" box="[347,497,2013,2040]" metricMagnitude="-1" metricUnit="m" metricValue="1.56" metricValueMax="1.8699999999999999" metricValueMin="1.25" pageId="11" pageNumber="232" unit="mm" value="156.0" valueMax="187.0" valueMin="125.0">125187 mm</quantity>
(mean
<quantity id="4CF49B197929FFC6FDBD55A1FD50F876" box="[581,679,2013,2040]" metricMagnitude="-1" metricUnit="m" metricValue="1.6199999999999999" pageId="11" pageNumber="232" unit="mm" value="162.0">162 mm</quantity>
), with the ratio TL/SVL 1.261.31 (mean 1.32); head length
<quantity id="4CF49B197929FFC1FAAE55A1FF34FF3F" lastPageId="12" lastPageNumber="233" metricMagnitude="-2" metricUnit="m" metricValue="3.8" metricValueMax="4.8" metricValueMin="2.8" pageId="11" pageNumber="232" unit="mm" value="38.0" valueMax="48.0" valueMin="28.0">2848 mm</quantity>
(mean
<quantity id="4CF49B19792EFFC1FEED52EBFE90FF3F" box="[277,359,151,177]" metricMagnitude="-2" metricUnit="m" metricValue="3.9" pageId="12" pageNumber="233" unit="mm" value="39.0">39 mm</quantity>
); supralabial scales on the left side 1214 (mean 13); supraocular scales on the left side 6; temporal scales on the left side 7; scale rows around fore body 121128 (mean 126); scale rows around hind body 7487 (mean 80); subdigital lamellae under Fi4 2123 (mean 21); subdigital lamellae under TOE4 2123 (mean 22); precloacal pores 1720 (mean 18).
</paragraph>
</subSubSection>
<subSubSection id="C3166577792EFFC1FF3F535AFD79FE23" pageId="12" pageNumber="233" type="distribution">
<paragraph id="8BB336FC792EFFC1FF3F535AFD79FE23" blockId="12.[151,1437,151,429]" pageId="12" pageNumber="233">
<emphasis id="B978EAEE792EFFC1FF3F535AFE96FECE" bold="true" box="[199,353,294,320]" pageId="12" pageNumber="233">Distribution.</emphasis>
Currently,
<taxonomicName id="4C0C4D7F792EFFC1FE1E535BFD05FECE" authority="Marques &amp; Parrinha &amp; Santos &amp; Bandeira &amp; Butler &amp; Sousa &amp; Bauer &amp; Wagner, 2022" authorityName="Marques &amp; Parrinha &amp; Santos &amp; Bandeira &amp; Butler &amp; Sousa &amp; Bauer &amp; Wagner" authorityYear="2022" box="[486,754,295,321]" class="Reptilia" family="Agamidae" genus="Acanthocercus" kingdom="Animalia" order="Squamata" pageId="12" pageNumber="233" phylum="Chordata" rank="species" species="ceriacoi" status="sp. nov.">
<emphasis id="B978EAEE792EFFC1FE1E535BFD05FECE" box="[486,754,295,321]" italics="true" pageId="12" pageNumber="233">Acanthocercus ceriacoi</emphasis>
</taxonomicName>
<emphasis id="B978EAEE792EFFC1FD04535BFCAEFECF" bold="true" box="[764,857,295,321]" pageId="12" pageNumber="233">
<taxonomicNameLabel id="A24B5795792EFFC1FD04535BFCAEFECF" box="[764,857,295,321]" pageId="12" pageNumber="233" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
is only known from the central and northwestern regions of
<collectingCountry id="F31B766C792EFFC1FEF75337FE93FEEB" box="[271,356,331,357]" name="Angola" pageId="12" pageNumber="233">Angola</collectingCountry>
, from central
<collectingRegion id="49C8F81E792EFFC1FDFB5337FDA8FEEB" box="[515,607,331,357]" country="Angola" name="Malanje" pageId="12" pageNumber="233">Malanje</collectingRegion>
to
<collectingRegion id="49C8F81E792EFFC1FD7C5337FCCEFEEB" box="[644,825,331,357]" country="Angola" name="Bengo" pageId="12" pageNumber="233">Bengo Province</collectingRegion>
in the west and
<collectingRegion id="49C8F81E792EFFC1FC0B5337FB61FEEB" box="[1011,1174,331,357]" country="Angola" name="Uige" pageId="12" pageNumber="233">Uíge Province</collectingRegion>
in the north (
<figureCitation id="13372A79792EFFC1FAC85337FA82FEE8" box="[1328,1397,331,358]" captionStart="FIGURE 6" captionStartId="12.[151,250,1507,1531]" captionTargetBox="[240,1347,463,1474]" captionTargetId="figure-149@12.[230,1357,453,1484]" captionTargetPageId="12" captionText="FIGURE 6. Geographic distribution of Angolan Acanthocercus. Green triangles: A. cyanocephalus; Pink squares: A. margaritae; Blue circles: A. ceriacoi sp. nov.; Black star: Type locality of A. ceriacoi sp. nov.. A. cyanocephalus and A. margaritae records from Marques et al. (2018), Wagner et al. (2018, 2021)." figureDoi="http://doi.org/10.5281/zenodo.6048542" httpUri="https://zenodo.org/record/6048542/files/figure.png" pageId="12" pageNumber="233">Fig. 6</figureCitation>
). It is likely that the species may also be present in the neighboring provinces of
<collectingRegion id="49C8F81E792EFFC1FC085313FB62FE07" box="[1008,1173,367,393]" country="Angola" name="Kwanza Norte" pageId="12" pageNumber="233">Kwanza-Norte</collectingRegion>
and
<collectingRegion id="49C8F81E792EFFC1FB335313FAFEFE07" box="[1227,1289,367,393]" country="Angola" name="Zaire" pageId="12" pageNumber="233">Zaire</collectingRegion>
, as well as in southern
<collectingCountry id="F31B766C792EFFC1FF0753EFFD7DFE23" box="[255,650,403,429]" name="Democratic Republic of the Congo" pageId="12" pageNumber="233">Democratic Republic of the Congo</collectingCountry>
.
</paragraph>
</subSubSection>
<caption id="DF736674792EFFC1FF6F579FFCE8F9CA" ID-DOI="http://doi.org/10.5281/zenodo.6048542" ID-Zenodo-Dep="6048542" httpUri="https://zenodo.org/record/6048542/files/figure.png" pageId="12" pageNumber="233" startId="12.[151,250,1507,1531]" targetBox="[240,1347,463,1474]" targetPageId="12">
<paragraph id="8BB336FC792EFFC1FF6F579FFCE8F9CA" blockId="12.[151,1437,1507,1604]" pageId="12" pageNumber="233">
<emphasis id="B978EAEE792EFFC1FF6F579FFEE3FA72" bold="true" box="[151,276,1507,1532]" pageId="12" pageNumber="233">FIGURE 6.</emphasis>
Geographic distribution of Angolan
<taxonomicName id="4C0C4D7F792EFFC1FD695798FCD0FA72" authorityName="Fitzinger" authorityYear="1843" box="[657,807,1508,1532]" class="Reptilia" family="Agamidae" genus="Acanthocercus" kingdom="Animalia" order="Squamata" pageId="12" pageNumber="233" phylum="Chordata" rank="genus">
<emphasis id="B978EAEE792EFFC1FD695798FCD0FA72" box="[657,807,1508,1532]" italics="true" pageId="12" pageNumber="233">Acanthocercus</emphasis>
</taxonomicName>
. Green triangles:
<taxonomicName id="4C0C4D7F792EFFC1FC195798FB61FA72" authorityName="HF" baseAuthorityName="Falk" baseAuthorityYear="1925" box="[993,1174,1508,1532]" class="Reptilia" family="Agamidae" genus="Acanthocercus" kingdom="Animalia" order="Squamata" pageId="12" pageNumber="233" phylum="Chordata" rank="species" species="cyanocephalus">
<emphasis id="B978EAEE792EFFC1FC195798FB61FA72" box="[993,1174,1508,1532]" italics="true" pageId="12" pageNumber="233">A. cyanocephalus</emphasis>
</taxonomicName>
; Pink squares:
<taxonomicName id="4C0C4D7F792EFFC1FACD5798FF30F991" authorityName="Wagner, Butler, Ceriaco &amp; Bauer" authorityYear="2021" class="Reptilia" family="Agamidae" genus="Acanthocercus" kingdom="Animalia" order="Squamata" pageId="12" pageNumber="233" phylum="Chordata" rank="species" species="margaritae">
<emphasis id="B978EAEE792EFFC1FACD5798FF30F991" italics="true" pageId="12" pageNumber="233">A. margaritae</emphasis>
</taxonomicName>
; Blue circles:
<taxonomicName id="4C0C4D7F792EFFC1FEA55474FE3AF991" authority="Marques &amp; Parrinha &amp; Santos &amp; Bandeira &amp; Butler &amp; Sousa &amp; Bauer &amp; Wagner, 2022" authorityName="Marques &amp; Parrinha &amp; Santos &amp; Bandeira &amp; Butler &amp; Sousa &amp; Bauer &amp; Wagner" authorityYear="2022" box="[349,461,1544,1567]" class="Reptilia" family="Agamidae" genus="Acanthocercus" kingdom="Animalia" order="Squamata" pageId="12" pageNumber="233" phylum="Chordata" rank="species" species="ceriacoi" status="sp. nov.">
<emphasis id="B978EAEE792EFFC1FEA55474FE3AF991" box="[349,461,1544,1567]" italics="true" pageId="12" pageNumber="233">A. ceriacoi</emphasis>
</taxonomicName>
<emphasis id="B978EAEE792EFFC1FE2C5474FDD2F9AE" bold="true" box="[468,549,1544,1568]" pageId="12" pageNumber="233">
<taxonomicNameLabel id="A24B5795792EFFC1FE2C5474FDD2F9AE" box="[468,549,1544,1568]" pageId="12" pageNumber="233" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
; Black star: Type locality of
<taxonomicName id="4C0C4D7F792EFFC1FCA95474FC36F991" authority="Marques &amp; Parrinha &amp; Santos &amp; Bandeira &amp; Butler &amp; Sousa &amp; Bauer &amp; Wagner, 2022" authorityName="Marques &amp; Parrinha &amp; Santos &amp; Bandeira &amp; Butler &amp; Sousa &amp; Bauer &amp; Wagner" authorityYear="2022" box="[849,961,1544,1567]" class="Reptilia" family="Agamidae" genus="Acanthocercus" kingdom="Animalia" order="Squamata" pageId="12" pageNumber="233" phylum="Chordata" rank="species" species="ceriacoi" status="sp. nov.">
<emphasis id="B978EAEE792EFFC1FCA95474FC36F991" box="[849,961,1544,1567]" italics="true" pageId="12" pageNumber="233">A. ceriacoi</emphasis>
</taxonomicName>
<emphasis id="B978EAEE792EFFC1FC305474FBEDF9AE" bold="true" box="[968,1050,1544,1568]" pageId="12" pageNumber="233">
<taxonomicNameLabel id="A24B5795792EFFC1FC305474FBEDF9AE" box="[968,1050,1544,1568]" pageId="12" pageNumber="233" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
.
<taxonomicName id="4C0C4D7F792EFFC1FBD05474FB2BF9AE" authorityName="HF" baseAuthorityName="Falk" baseAuthorityYear="1925" box="[1064,1244,1544,1568]" class="Reptilia" family="Agamidae" genus="Acanthocercus" kingdom="Animalia" order="Squamata" pageId="12" pageNumber="233" phylum="Chordata" rank="species" species="cyanocephalus">
<emphasis id="B978EAEE792EFFC1FBD05474FB2BF9AE" box="[1064,1244,1544,1568]" italics="true" pageId="12" pageNumber="233">A. cyanocephalus</emphasis>
</taxonomicName>
and
<taxonomicName id="4C0C4D7F792EFFC1FAF65474FA6BF991" authorityName="Wagner, Butler, Ceriaco &amp; Bauer" authorityYear="2021" box="[1294,1436,1544,1567]" class="Reptilia" family="Agamidae" genus="Acanthocercus" kingdom="Animalia" order="Squamata" pageId="12" pageNumber="233" phylum="Chordata" rank="species" species="margaritae">
<emphasis id="B978EAEE792EFFC1FAF65474FA6BF991" box="[1294,1436,1544,1567]" italics="true" pageId="12" pageNumber="233">A. margaritae</emphasis>
</taxonomicName>
records from
<bibRefCitation id="EF9D4B0D792EFFC1FED85450FE0AF9CA" author="Marques, M. P. &amp; Ceriaco, L. M. P. &amp; Blackburn, D. C. &amp; Bauer, A. M." box="[288,509,1580,1604]" pageId="12" pageNumber="233" pagination="1 - 501" refId="ref11862" refString="Marques, M. P., Ceriaco, L. M. P., Blackburn, D. C. &amp; Bauer, A. M. (2018) Diversity and distribution of the amphibians and terrestrial reptiles of Angola: Atlas of historical and bibliographic records (1850 - 2017). Proceedings of the California Academy of Sciences, 65 (Supplement 2), 1 - 501." type="journal article" year="2018">
Marques
<emphasis id="B978EAEE792EFFC1FE875450FE45F9CA" box="[383,434,1580,1604]" italics="true" pageId="12" pageNumber="233">et al.</emphasis>
(2018)
</bibRefCitation>
,
<bibRefCitation id="EF9D4B0D792EFFC1FDFF5450FD25F9CA" author="Wagner, P. &amp; Greenbaum, E. &amp; Bauer, A. M. &amp; Kusamba, C. &amp; Leache, A. D." box="[519,722,1580,1604]" pageId="12" pageNumber="233" pagination="771 - 817" refId="ref12867" refString="Wagner, P., Greenbaum, E., Bauer, A. M., Kusamba, C. &amp; Leache, A. D. (2018) Lifting the blue-headed veil - integrative taxonomy of the Acanthocercus atricollis species complex (Squamata: Agamidae). Journal of Natural History, 52, 771 - 817. https: // doi. org / 10.1080 / 00222933.2018.1435833" type="journal article" year="2018">
Wagner
<emphasis id="B978EAEE792EFFC1FDA45450FD78F9CA" box="[604,655,1580,1604]" italics="true" pageId="12" pageNumber="233">et al.</emphasis>
(2018
</bibRefCitation>
,
<bibRefCitation id="EF9D4B0D792EFFC1FD255450FCE3F9CA" author="Wagner, P. &amp; Butler, B. &amp; Ceriaco, L. M. P. &amp; Bauer, A. M." box="[733,788,1580,1604]" pageId="12" pageNumber="233" pagination="449 - 463" refId="ref12938" refString="Wagner, P., Butler, B., Ceriaco, L. M. P. &amp; Bauer, A. M. (2021) A new species of the Acanthocercus atricollis (Smith, 1849) complex (Squamata: Agamidae). Salamandra, 57 (4), 449 - 463." type="journal article" year="2021">2021</bibRefCitation>
).
</paragraph>
</caption>
<subSubSection id="C3166577792EFFC1FF3F540EFB0CF87A" pageId="12" pageNumber="233" type="biology_ecology">
<paragraph id="8BB336FC792EFFC1FF3F540EFB0CF87A" blockId="12.[151,1437,1650,2036]" pageId="12" pageNumber="233">
<emphasis id="B978EAEE792EFFC1FF3F540EFD9CF902" bold="true" box="[199,619,1650,1676]" pageId="12" pageNumber="233">Habitat and Natural History notes.</emphasis>
In Cangandala National Park and surrounding areas, the species is commonly found dwelling in trees, several meters above ground. The park lies at an approximate elevation of
<quantity id="4CF49B19792EFFC1FA9C54EAFF5AF95A" metricMagnitude="3" metricUnit="m" metricValue="1.0" pageId="12" pageNumber="233" unit="m" value="1000.0">1000 m</quantity>
and its climate is similar to that of the Angolan central plateau, with vegetation mostly dominated by miombo woodlands (
<figureCitation id="13372A79792EFFC1FEE754A2FE95F976" box="[287,354,1758,1784]" captionStart="FIGURE 7" captionStartId="13.[151,250,992,1016]" captionTargetBox="[203,1384,181,968]" captionTargetId="figure-18@13.[203,1384,181,968]" captionTargetPageId="13" captionText="FIGURE 7. Habitat of the type locality of Acanthocercus ceriacoi sp. nov. in Cangandala National Park, Malanje Province, Angola. Photo by Luis M.P. Ceríaco." figureDoi="http://doi.org/10.5281/zenodo.6048546" httpUri="https://zenodo.org/record/6048546/files/figure.png" pageId="12" pageNumber="233">Fig. 7</figureCitation>
), with
<taxonomicName id="4C0C4D7F792EFFC1FE5654A2FD05F976" box="[430,754,1758,1784]" class="Magnoliopsida" family="Fabaceae" genus="Brachystegia" kingdom="Plantae" order="Fabales" pageId="12" pageNumber="233" phylum="Tracheophyta" rank="species" species="wangermeeana">
<emphasis id="B978EAEE792EFFC1FE5654A2FD05F976" box="[430,754,1758,1784]" italics="true" pageId="12" pageNumber="233">Brachystegia wangermeeana</emphasis>
</taxonomicName>
,
<taxonomicName id="4C0C4D7F792EFFC1FD0754A3FC64F976" box="[767,915,1758,1784]" class="Magnoliopsida" family="Fabaceae" genus="Brachystegia" kingdom="Plantae" order="Fabales" pageId="12" pageNumber="233" phylum="Tracheophyta" rank="species" species="floribunda">
<emphasis id="B978EAEE792EFFC1FD0754A3FC64F976" box="[767,915,1758,1784]" italics="true" pageId="12" pageNumber="233">B. floribunda</emphasis>
</taxonomicName>
,
<taxonomicName id="4C0C4D7F792EFFC1FC5854A2FB58F976" box="[928,1199,1758,1784]" class="Magnoliopsida" family="Fabaceae" genus="Julbernardia" kingdom="Plantae" order="Fabales" pageId="12" pageNumber="233" phylum="Tracheophyta" rank="species" species="paniculata">
<emphasis id="B978EAEE792EFFC1FC5854A2FB58F976" box="[928,1199,1758,1784]" italics="true" pageId="12" pageNumber="233">Julbernardia paniculata</emphasis>
</taxonomicName>
,
<taxonomicName id="4C0C4D7F792EFFC1FB4454A2FF15F892" class="Magnoliopsida" family="Fabaceae" genus="Erythrophleum" kingdom="Plantae" order="Fabales" pageId="12" pageNumber="233" phylum="Tracheophyta" rank="species" species="africanum">
<emphasis id="B978EAEE792EFFC1FB4454A2FF15F892" italics="true" pageId="12" pageNumber="233">Erythrophleum africanum</emphasis>
</taxonomicName>
,
<taxonomicName id="4C0C4D7F792EFFC1FF17557EFE5EF892" box="[239,425,1794,1820]" class="Magnoliopsida" family="Combretaceae" genus="Combretum" kingdom="Plantae" order="Myrtales" pageId="12" pageNumber="233" phylum="Tracheophyta" rank="species" species="undetermined">
<emphasis id="B978EAEE792EFFC1FF17557EFE85F892" box="[239,370,1794,1820]" italics="true" pageId="12" pageNumber="233">Combretum</emphasis>
spp.
</taxonomicName>
, and
<taxonomicName id="4C0C4D7F792EFFC1FE1B557EFDA7F892" box="[483,592,1794,1820]" class="Magnoliopsida" family="Anacardiaceae" genus="Rhus" kingdom="Plantae" order="Sapindales" pageId="12" pageNumber="233" phylum="Tracheophyta" rank="species" species="undetermined">
<emphasis id="B978EAEE792EFFC1FE1B557EFDECF892" box="[483,539,1794,1820]" italics="true" pageId="12" pageNumber="233">Rhus</emphasis>
spp.
</taxonomicName>
growing on the red clay soils of the park (
<bibRefCitation id="EF9D4B0D792EFFC1FBDC557EFABDF892" author="Grandvaux-Barbosa, L. A." box="[1060,1354,1794,1820]" pageId="12" pageNumber="233" refId="ref11104" refString="Grandvaux-Barbosa, L. A. (1970) Carta Fitogeografica de Angola. Instituto de Investigacao Cientifica de Angola, Luanda, 323 pp." type="book" year="1970">Grandvaux-Barbosa 1970</bibRefCitation>
). Some small riverine gallery forests also occur in the park. In
<collectingRegion id="49C8F81E792EFFC1FD02555AFC6AF8CE" box="[762,925,1830,1856]" country="Angola" name="Uige" pageId="12" pageNumber="233">Uíge Province</collectingRegion>
, the species was found basking on the ground in open-savannah habitat, in an area particularly devoid of trees (
<figureCitation id="13372A79792EFFC1FC9C5536FC50F8EA" box="[868,935,1866,1892]" captionStart="FIGURE 8" captionStartId="13.[151,250,1967,1991]" captionTargetBox="[215,1371,1077,1944]" captionTargetId="figure-47@13.[215,1371,1077,1944]" captionTargetPageId="13" captionText="FIGURE 8. Habitat of Acanthocercus ceriacoi sp. nov. at the paratype locality of Zulumongo village, Uíge Province, northern Angola. Photo by Luis M.P. Ceríaco." figureDoi="http://doi.org/10.5281/zenodo.6048550" httpUri="https://zenodo.org/record/6048550/files/figure.png" pageId="12" pageNumber="233">Fig. 8</figureCitation>
). The habitat of this region mainly comprises a mosaic of Zambezian savannah and coffee plantation forests, dominated by the plant genera
<taxonomicName id="4C0C4D7F792EFFC1FB4E5512FAB1F806" box="[1206,1350,1902,1928]" class="Liliopsida" family="Poaceae" genus="Hyparrhenia" kingdom="Plantae" order="Poales" pageId="12" pageNumber="233" phylum="Tracheophyta" rank="genus">
<emphasis id="B978EAEE792EFFC1FB4E5512FAB1F806" box="[1206,1350,1902,1928]" italics="true" pageId="12" pageNumber="233">Hyparrhenia</emphasis>
</taxonomicName>
,
<taxonomicName id="4C0C4D7F792EFFC1FAAC5512FA62F806" box="[1364,1429,1902,1928]" class="Magnoliopsida" family="Cannabaceae" genus="Celtis" kingdom="Plantae" order="Rosales" pageId="12" pageNumber="233" phylum="Tracheophyta" rank="genus">
<emphasis id="B978EAEE792EFFC1FAAC5512FA62F806" box="[1364,1429,1902,1928]" italics="true" pageId="12" pageNumber="233">Celtis</emphasis>
</taxonomicName>
,
<taxonomicName id="4C0C4D7F792EFFC1FF6F55EEFF11F822" box="[151,230,1938,1964]" class="Magnoliopsida" family="Fabaceae" genus="Albizia" kingdom="Plantae" order="Fabales" pageId="12" pageNumber="233" phylum="Tracheophyta" rank="genus">
<emphasis id="B978EAEE792EFFC1FF6F55EEFF11F822" box="[151,230,1938,1964]" italics="true" pageId="12" pageNumber="233">Albizia</emphasis>
</taxonomicName>
,
<taxonomicName id="4C0C4D7F792EFFC1FF0C55EFFECAF822" box="[244,317,1939,1964]" class="Magnoliopsida" family="Moraceae" genus="Morus" kingdom="Plantae" order="Rosales" pageId="12" pageNumber="233" phylum="Tracheophyta" rank="genus">
<emphasis id="B978EAEE792EFFC1FF0C55EFFECAF822" box="[244,317,1939,1964]" italics="true" pageId="12" pageNumber="233">Morus</emphasis>
</taxonomicName>
and
<taxonomicName id="4C0C4D7F792EFFC1FE8C55EFFD18F822" authority="(Grandvaux-Barbosa 1970)" baseAuthorityName="Grandvaux-Barbosa" baseAuthorityYear="1970" box="[372,751,1938,1964]" class="Magnoliopsida" family="Moraceae" genus="Ficus" kingdom="Plantae" order="Rosales" pageId="12" pageNumber="233" phylum="Tracheophyta" rank="genus">
<emphasis id="B978EAEE792EFFC1FE8C55EFFE45F822" box="[372,434,1939,1964]" italics="true" pageId="12" pageNumber="233">Ficus</emphasis>
(
<bibRefCitation id="EF9D4B0D792EFFC1FE3955EEFD10F822" author="Grandvaux-Barbosa, L. A." box="[449,743,1938,1964]" pageId="12" pageNumber="233" refId="ref11104" refString="Grandvaux-Barbosa, L. A. (1970) Carta Fitogeografica de Angola. Instituto de Investigacao Cientifica de Angola, Luanda, 323 pp." type="book" year="1970">Grandvaux-Barbosa 1970</bibRefCitation>
)
</taxonomicName>
. The female
<typeStatus id="54B7885E792EFFC1FC8755EEFC28F822" box="[895,991,1938,1964]" pageId="12" pageNumber="233" type="paratype">paratype</typeStatus>
(MHNCUP/REP 860) from
<collectingRegion id="49C8F81E792EFFC1FAD955EEFF3FF85E" country="Angola" name="Uige" pageId="12" pageNumber="233">Uíge Province</collectingRegion>
, collected in November, was active on the ground around midday. After euthanasia and while preparing the specimen for fixing, three well-developed eggs were found in the abdominal cavity of the specimen.
</paragraph>
</subSubSection>
<caption id="DF736674792FFFC0FF6F519CFDF9FB92" ID-DOI="http://doi.org/10.5281/zenodo.6048546" ID-Zenodo-Dep="6048546" httpUri="https://zenodo.org/record/6048546/files/figure.png" pageId="13" pageNumber="234" startId="13.[151,250,992,1016]" targetBox="[203,1384,181,968]" targetPageId="13">
<paragraph id="8BB336FC792FFFC0FF6F519CFDF9FB92" blockId="13.[151,1437,992,1052]" pageId="13" pageNumber="234">
<emphasis id="B978EAEE792FFFC0FF6F519CFEE2FC76" bold="true" box="[151,277,992,1016]" pageId="13" pageNumber="234">FIGURE 7.</emphasis>
Habitat of the type locality of
<taxonomicName id="4C0C4D7F792FFFC0FDAF519CFCBFFC76" authority="Marques &amp; Parrinha &amp; Santos &amp; Bandeira &amp; Butler &amp; Sousa &amp; Bauer &amp; Wagner, 2022" authorityName="Marques &amp; Parrinha &amp; Santos &amp; Bandeira &amp; Butler &amp; Sousa &amp; Bauer &amp; Wagner" authorityYear="2022" box="[599,840,992,1016]" class="Reptilia" family="Agamidae" genus="Acanthocercus" kingdom="Animalia" order="Squamata" pageId="13" pageNumber="234" phylum="Chordata" rank="species" species="ceriacoi" status="sp. nov.">
<emphasis id="B978EAEE792FFFC0FDAF519CFCBFFC76" box="[599,840,992,1016]" italics="true" pageId="13" pageNumber="234">Acanthocercus ceriacoi</emphasis>
</taxonomicName>
<emphasis id="B978EAEE792FFFC0FCA8519CFC55FC76" bold="true" box="[848,930,992,1016]" pageId="13" pageNumber="234">
<taxonomicNameLabel id="A24B5795792FFFC0FCA8519CFC55FC76" box="[848,930,992,1016]" pageId="13" pageNumber="234" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
in Cangandala National Park, Malanje Province, Angola. Photo by Luis M.P. Ceríaco.
</paragraph>
</caption>
<caption id="DF736674792FFFC0FF6F55D3FDF9F862" ID-DOI="http://doi.org/10.5281/zenodo.6048550" ID-Zenodo-Dep="6048550" httpUri="https://zenodo.org/record/6048550/files/figure.png" pageId="13" pageNumber="234" startId="13.[151,250,1967,1991]" targetBox="[215,1371,1077,1944]" targetPageId="13">
<paragraph id="8BB336FC792FFFC0FF6F55D3FDF9F862" blockId="13.[151,1436,1967,2028]" pageId="13" pageNumber="234">
<emphasis id="B978EAEE792FFFC0FF6F55D3FEE4F846" bold="true" box="[151,275,1967,1992]" pageId="13" pageNumber="234">FIGURE 8.</emphasis>
Habitat of
<taxonomicName id="4C0C4D7F792FFFC0FE7D55CCFD83F849" authority="Marques &amp; Parrinha &amp; Santos &amp; Bandeira &amp; Butler &amp; Sousa &amp; Bauer &amp; Wagner, 2022" authorityName="Marques &amp; Parrinha &amp; Santos &amp; Bandeira &amp; Butler &amp; Sousa &amp; Bauer &amp; Wagner" authorityYear="2022" box="[389,628,1968,1992]" class="Reptilia" family="Agamidae" genus="Acanthocercus" kingdom="Animalia" order="Squamata" pageId="13" pageNumber="234" phylum="Chordata" rank="species" species="ceriacoi" status="sp. nov.">
<emphasis id="B978EAEE792FFFC0FE7D55CCFD83F849" box="[389,628,1968,1992]" italics="true" pageId="13" pageNumber="234">Acanthocercus ceriacoi</emphasis>
</taxonomicName>
<emphasis id="B978EAEE792FFFC0FD8255CCFD3CF846" bold="true" box="[634,715,1968,1992]" pageId="13" pageNumber="234">
<taxonomicNameLabel id="A24B5795792FFFC0FD8255CCFD3CF846" box="[634,715,1968,1992]" pageId="13" pageNumber="234" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
at the paratype locality of Zulumongo village, Uíge Province, northern Angola. Photo by Luis M.P. Ceríaco.
</paragraph>
</caption>
<subSubSection id="C3166577792CFFC3FF3F52EAFD41FE06" pageId="14" pageNumber="235" type="etymology">
<paragraph id="8BB336FC792CFFC3FF3F52EAFD41FE06" blockId="14.[151,1437,150,393]" pageId="14" pageNumber="235">
<emphasis id="B978EAEE792CFFC3FF3F52EAFEB9FF3E" bold="true" box="[199,334,150,176]" pageId="14" pageNumber="235">Etymology.</emphasis>
The species is named after the Portuguese zoologist Luis Miguel Pires Ceríaco (1987present), head of collections and research at the Museu de História Natural e da Ciência da Universidade do
<collectingRegion id="49C8F81E792CFFC3FAF452C7FABBFF5B" box="[1292,1356,187,213]" country="Portugal" name="Porto" pageId="14" pageNumber="235">Porto</collectingRegion>
,
<collectingRegion id="49C8F81E792CFFC3FAA152C7FA6EFF5B" box="[1369,1433,187,213]" country="Portugal" name="Porto" pageId="14" pageNumber="235">Porto</collectingRegion>
,
<collectingCountry id="F31B766C792CFFC3FF6F52A2FF0EFF76" box="[151,249,222,248]" name="Portugal" pageId="14" pageNumber="235">Portugal</collectingCountry>
, and invited curator of herpetology at the Museu Nacional de História Natural e da Ciência,
<collectingRegion id="49C8F81E792CFFC3FAE852A2FA96FF76" box="[1296,1377,222,248]" country="Portugal" name="Lisboa" pageId="14" pageNumber="235">Lisbon</collectingRegion>
,
<collectingCountry id="F31B766C792CFFC3FA9552A3FF24FE92" name="Portugal" pageId="14" pageNumber="235">Portugal</collectingCountry>
. Luis M.P. Ceríaco has been a major force comprising the new wave of Angolan herpetological studies and a dedicated supervisor to a new generation of Portuguese and Angolan herpetologists. The name is formed in the genitive masculine singular. We propose the English common name of Ceríacos Tree
<taxonomicName id="4C0C4D7F792CFFC3FB9E5336FB4DFEEA" box="[1126,1210,330,356]" class="Reptilia" family="Agamidae" genus="Agama" kingdom="Animalia" order="Squamata" pageId="14" pageNumber="235" phylum="Chordata" rank="genus">Agama</taxonomicName>
, and the Portuguese common name of Sardão Arborícola de Ceríaco.
</paragraph>
</subSubSection>
</treatment>
</document>