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<document id="9ACD3DA1455F98EB5E583A5CAAEA7E13" ID-CLB-Dataset="24540" ID-DOI="10.11646/zootaxa.4708.1.1" ID-GBIF-Dataset="653d580d-e1c7-499c-8604-4b5a7b1920ef" ID-ISSN="1175-5326" ID-Zenodo-Dep="3587181" ID-ZooBank="09F24F0B-5090-4089-A525-8E5150C91EF7" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1576825058981" checkinUser="plazi" docAuthor="Perkins, Philip D." docDate="2019" docId="03A587E7A87BFFFD0CBA1528FE6F962A" docLanguage="en" docName="zootaxa.4708.1.1.pdf" docOrigin="Zootaxa 4708 (1)" docStyle="DocumentStyle:5EBBA59367AD13919D70D935FA04F6A3.14:Zootaxa.2013-.monograph" docStyleId="5EBBA59367AD13919D70D935FA04F6A3" docStyleName="Zootaxa.2013-.monograph" docStyleVersion="14" docTitle="Hydrochus shorti Perkins 2019, new species" docType="treatment" docVersion="7" lastPageNumber="26" masterDocId="FF9CFF9FA863FFE40C2D1244FFE19255" masterDocTitle="Taxonomy of Venezuelan water beetles in the genus Hydrochus Leach, 1817, and an analysis of male genitalia morphology (Coleoptera: Hydrochidae)" masterLastPageNumber="59" masterPageNumber="1" pageNumber="25" updateTime="1698775642300" updateUser="ExternalLinkService">
<mods:mods id="983D31EDB41E8C1C99182DF8D058C0CA" xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo id="B5471D83FE0B9B59E58C9B2F9A496915">
<mods:title id="0D5C7A00C0E46D50904E98E037664A5C">Taxonomy of Venezuelan water beetles in the genus Hydrochus Leach, 1817, and an analysis of male genitalia morphology (Coleoptera: Hydrochidae)</mods:title>
</mods:titleInfo>
<mods:name id="DF9BCD205C2B77DD8818CA1228028A6C" type="personal">
<mods:role id="B16A56C1E6EB433CEE3437DE432BDA7F">
<mods:roleTerm id="43264ABFA8736F1CCE3E0390EA048FF1">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="F0B1F47AB26D28971AA319F84465DBCE">Perkins, Philip D.</mods:namePart>
</mods:name>
<mods:typeOfResource id="B8FB4A17DDFAC29B80AC7D0DF90B46C3">text</mods:typeOfResource>
<mods:relatedItem id="B0E765CD38B82C3684066D6712E90533" type="host">
<mods:titleInfo id="172213CFE1EF12D396F7B935AC5CAD78">
<mods:title id="97BC23AD7334986E91CABAA688565886">Zootaxa</mods:title>
</mods:titleInfo>
<mods:part id="8FB474325344A828012A1BF6F8B73B7A">
<mods:date id="49B6C11BBCED1DE440544B8F507E56DC">2019</mods:date>
<mods:detail id="3EBD1235D74727C33974A6F7A774EE83" type="pubDate">
<mods:number id="5E634BADF78882B2F392C772E4821B75">2019-12-12</mods:number>
</mods:detail>
<mods:detail id="43FEC7BA083A7552CAD90794DADDEF36" type="volume">
<mods:number id="06E411D11CADB6FD4D204D34C5A34E4A">4708</mods:number>
</mods:detail>
<mods:detail id="0577D4196AD2C2AA3A055F6494B4DDED" type="issue">
<mods:number id="A3C3C7D640E96FEA12FE94794CC41F48">1</mods:number>
</mods:detail>
<mods:extent id="77D501EB0FB30E50FD2C703B33F9189B" unit="page">
<mods:start id="C72544C04D00A9F373DDDEC4A3A5BB27">1</mods:start>
<mods:end id="89A4191FC5DD91E2700014B31ED48AD7">59</mods:end>
</mods:extent>
</mods:part>
</mods:relatedItem>
<mods:classification id="604D4FD8F2B4B3AB3600C1A8BCCEEAE7">journal article</mods:classification>
<mods:identifier id="0E5B36470310E168F0BD2E3F0DC1D911" type="CLB-Dataset">24540</mods:identifier>
<mods:identifier id="0D6E37FFA200A1D89DAD12F45F31CEF8" type="DOI">10.11646/zootaxa.4708.1.1</mods:identifier>
<mods:identifier id="C644E038E71CBC85934371B3ACE7F7E2" type="GBIF-Dataset">653d580d-e1c7-499c-8604-4b5a7b1920ef</mods:identifier>
<mods:identifier id="D0DE2DCF83C84DFC18377FE99A7AE6F7" type="ISSN">1175-5326</mods:identifier>
<mods:identifier id="6029F4E3FF2ED28F174ABB6688A88087" type="Zenodo-Dep">3587181</mods:identifier>
<mods:identifier id="A68C9973BEFF403CF898986F391DD722" type="ZooBank">09F24F0B-5090-4089-A525-8E5150C91EF7</mods:identifier>
</mods:mods>
<treatment id="03A587E7A87BFFFD0CBA1528FE6F962A" ID-GBIF-Taxon="160798400" LSID="urn:lsid:plazi:treatment:03A587E7A87BFFFD0CBA1528FE6F962A" httpUri="http://treatment.plazi.org/id/03A587E7A87BFFFD0CBA1528FE6F962A" lastPageId="25" lastPageNumber="26" pageId="24" pageNumber="25">
<subSubSection id="C316657AA87BFFFC0CBA1528FDE095D2" box="[151,513,1900,1927]" pageId="24" pageNumber="25" type="nomenclature">
<paragraph id="8BB336F1A87BFFFC0CBA1528FDE095D2" blockId="24.[151,513,1900,1963]" box="[151,513,1900,1927]" pageId="24" pageNumber="25">
<heading id="D0FB819DA87BFFFC0CBA1528FDE095D2" bold="true" box="[151,513,1900,1927]" fontSize="11" level="1" pageId="24" pageNumber="25" reason="1">
<emphasis id="B978EAE3A87BFFFC0CBA1528FDE095D2" bold="true" box="[151,513,1900,1927]" pageId="24" pageNumber="25">
<taxonomicName id="4C0C4D72A87BFFFC0CBA1528FE8B95D2" ID-CoL="8Y4RF" authority="Perkins, 2019" authorityName="Perkins" authorityYear="2019" box="[151,362,1900,1927]" class="Insecta" family="Hydrochidae" genus="Hydrochus" kingdom="Animalia" order="Coleoptera" pageId="24" pageNumber="25" phylum="Arthropoda" rank="species" species="shorti" status="sp. nov.">
<emphasis id="B978EAE3A87BFFFC0CBA1528FE8B95D2" bold="true" box="[151,362,1900,1927]" italics="true" pageId="24" pageNumber="25">Hydrochus shorti</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A24B5798A87BFFFC0D5F1529FDE095D2" box="[370,513,1901,1927]" pageId="24" pageNumber="25" rank="species">new species</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C316657AA87BFFFC0CBA15D5FEEC95FE" box="[151,269,1937,1963]" pageId="24" pageNumber="25" type="description">
<paragraph id="8BB336F1A87BFFFC0CBA15D5FEEC95FE" blockId="24.[151,513,1900,1963]" box="[151,269,1937,1963]" pageId="24" pageNumber="25">
<figureCitation id="13372A74A87BFFFC0CBA15D5FF0495FE" box="[151,229,1937,1963]" captionStart="FIGURE 2" captionStartId="14.[151,250,2009,2034]" captionTargetBox="[199,1400,435,1940]" captionTargetId="figure@14.[151,1435,417,1984]" captionTargetPageId="14" captionText="FIGURE 2. Dorsal habitus of Hydrochus species." figureDoi="http://doi.org/10.5281/zenodo.3587185" httpUri="https://zenodo.org/record/3587185/files/figure.png" pageId="24" pageNumber="25">Figs. 2</figureCitation>
,
<figureCitation id="13372A74A87BFFFC0CDC15D5FEEC95FE" box="[241,269,1937,1963]" captionStart="FIGURE 16" captionStartId="33.[151,250,1736,1761]" captionTargetBox="[186,1401,210,1683]" captionTargetId="figure@33.[151,1435,181,1710]" captionTargetPageId="33" captionText="FIGURE 16. Hydrochus shorti n. sp., habitus and male genitalia of holotype." figureDoi="http://doi.org/10.5281/zenodo.3587215" httpUri="https://zenodo.org/record/3587215/files/figure.png" pageId="24" pageNumber="25">16</figureCitation>
</paragraph>
</subSubSection>
<subSubSection id="C316657AA87BFFFD0CBA159DFA9893FB" lastPageId="25" lastPageNumber="26" pageId="24" pageNumber="25" type="materials_examined">
<paragraph id="8BB336F1A87BFFFD0CBA159DFB269282" blockId="24.[151,1436,2008,2035]" lastBlockId="25.[151,1437,152,1151]" lastPageId="25" lastPageNumber="26" pageId="24" pageNumber="25">
<emphasis id="B978EAE3A87BFFFC0CBA159DFEA695A6" bold="true" box="[151,327,2008,2035]" pageId="24" pageNumber="25">Type Material.</emphasis>
<materialsCitation id="3B643CACA87BFFFD0D61159DFBB392E6" ID-GBIF-Occurrence="2556427175" country="Venezuela" elevation="72" lastPageId="25" lastPageNumber="26" latitude="5.3416166" location="Amazonas State" longLatPrecision="1" longitude="-67.75597" pageId="24" pageNumber="25" specimenCount="1" stateProvince="Amazonas" typeStatus="holotype">
<emphasis id="B978EAE3A87BFFFC0D61159DFDF595A6" bold="true" box="[332,532,2008,2035]" pageId="24" pageNumber="25">
<typeStatus id="54B78853A87BFFFC0D61159DFE5895A6" box="[332,441,2009,2035]" pageId="24" pageNumber="25" type="holotype">Holotype</typeStatus>
(male):
</emphasis>
“
<collectingCountry id="F31B7661A87BFFFC0E09159DFD7F95A6" box="[548,670,2009,2035]" name="Venezuela" pageId="24" pageNumber="25">Venezuela</collectingCountry>
:
<collectingRegion id="49C8F813A87BFFFC0E8B159DFCBF95A6" box="[678,862,2009,2035]" country="Venezuela" name="Amazonas" pageId="24" pageNumber="25">Amazonas State</collectingRegion>
:
<geoCoordinate id="EE385036A87BFFFC0F4A159CFC1695A6" box="[871,1015,2008,2035]" degrees="5" direction="north" minutes="20.497" orientation="latitude" pageId="24" pageNumber="25" precision="1" value="5.3416166">5° 20.497’ N</geoCoordinate>
,
<geoCoordinate id="EE385036A87BFFFC082C159DFB4295A6" box="[1025,1187,2008,2035]" degrees="67" direction="west" minutes="45.358" orientation="longitude" pageId="24" pageNumber="25" precision="1" value="-67.75597">67° 45.358’ W</geoCoordinate>
,
<quantity id="4CF49B14A87BFFFC0881159DFB0195A6" box="[1196,1248,2009,2035]" metricMagnitude="1" metricUnit="m" metricValue="7.2" pageId="24" pageNumber="25" unit="m" value="72.0">
<elevation id="0021D1C2A87BFFFC0881159DFB0195A6" box="[1196,1248,2009,2035]" metricMagnitude="1" metricUnit="m" metricValue="7.2" pageId="24" pageNumber="25" unit="m" value="72.0">72m</elevation>
</quantity>
, ca.
<quantity id="4CF49B14A87BFFFC093D159DFAB195A6" box="[1296,1360,2009,2035]" metricMagnitude="4" metricUnit="m" metricValue="1.2" pageId="24" pageNumber="25" unit="km" value="12.0">12km</quantity>
N. Sa- mariapo, small stream @ rd x-ing;
<date id="FFB21031A87AFFFD0E0C12DCFD6C92E7" box="[545,653,152,178]" pageId="25" pageNumber="26" value="2006-01-06">06.i.2006</date>
, AS-06-013, leg. A.E.Z. Short” (
<collectionCode id="ED1DAE34A87AFFFD0FD112DDFBAB92E6" box="[1020,1098,153,179]" country="Venezuela" httpUri="http://biocol.org/urn:lsid:biocol.org:col:33834" lsid="urn:lsid:biocol.org:col:33834" name="Museo del Instituto de Zoologia Agricola Francisco Fernandez Yepez" pageId="25" pageNumber="26" type="Museum">MIZA</collectionCode>
)
</materialsCitation>
.
<materialsCitation id="3B643CACA87AFFFD087012DDFB259282" ID-GBIF-Occurrence="2521263826" collectionCode="SEMC, MIZA, MALUZ" country="Venezuela" location="Same" pageId="25" pageNumber="26" specimenCode="VZ09-0115, 03" specimenCount="2" stateProvince="Amazonas" typeStatus="paratype">
<emphasis id="B978EAE3A87AFFFD087012DDFEFC9282" bold="true" pageId="25" pageNumber="26">
<typeStatus id="54B78853A87AFFFD087012DDFB3492E6" box="[1117,1237,153,179]" pageId="25" pageNumber="26" type="paratype">Paratypes</typeStatus>
(31):
<collectingCountry id="F31B7661A87AFFFD093612DDFA7992E6" box="[1307,1432,153,179]" name="Venezuela" pageId="25" pageNumber="26">Venezuela</collectingCountry>
:
<collectingRegion id="49C8F813A87AFFFD0CBA12F8FEF99282" box="[151,280,188,215]" country="Venezuela" name="Amazonas" pageId="25" pageNumber="26">Amazonas</collectingRegion>
:
</emphasis>
<location id="8ED3602AA87AFFFD0D0912F9FE839282" LSID="urn:lsid:plazi:treatment:03A587E7A87BFFFD0CBA1528FE6F962A:8ED3602AA87AFFFD0D0912F9FE839282" box="[292,354,189,215]" country="Venezuela" name="Same" pageId="25" pageNumber="26" stateProvince="Amazonas">Same</location>
data as
<typeStatus id="54B78853A87AFFFD0D9012F8FDFE9283" box="[445,543,188,214]" pageId="25" pageNumber="26" type="holotype">holotype</typeStatus>
(21
<collectionCode id="ED1DAE34A87AFFFD0E7F12F9FD409282" box="[594,673,189,215]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34264" name="University of Kansas - Biodiversity Institute" pageId="25" pageNumber="26" type="Museum">SEMC</collectionCode>
,
<collectionCode id="ED1DAE34A87AFFFD0E8012F9FD169282" box="[685,759,189,215]" country="Venezuela" httpUri="http://biocol.org/urn:lsid:biocol.org:col:33834" lsid="urn:lsid:biocol.org:col:33834" name="Museo del Instituto de Zoologia Agricola Francisco Fernandez Yepez" pageId="25" pageNumber="26" type="Museum">MIZA</collectionCode>
,
<collectionCode id="ED1DAE34A87AFFFD0F2F12F9FC8A9282" box="[770,875,189,215]" pageId="25" pageNumber="26">MALUZ</collectionCode>
);
<specimenCode id="DBAA9E8AA87AFFFD0F5012F8FBE19282" box="[893,1024,188,215]" collectionCode="MALUZ" pageId="25" pageNumber="26">VZ09-0115</specimenCode>
-
<specimenCode id="DBAA9E8AA87AFFFD082A12F8FBC39282" box="[1031,1058,188,215]" collectionCode="MALUZ" pageId="25" pageNumber="26">03</specimenCode>
A (10
<collectionCode id="ED1DAE34A87AFFFD084712F9FB5A9282" box="[1130,1211,189,215]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34264" name="University of Kansas - Biodiversity Institute" pageId="25" pageNumber="26" type="Museum">SEMC</collectionCode>
)
</materialsCitation>
.
</paragraph>
<paragraph id="8BB336F1A87AFFFD0CEA12A4FA9893FB" blockId="25.[151,1437,152,1151]" pageId="25" pageNumber="26">
<emphasis id="B978EAE3A87AFFFD0CEA12A4FE3092AE" bold="true" box="[199,465,224,251]" pageId="25" pageNumber="26">Differential Diagnosis.</emphasis>
Among Venezuelan species, recognized by the combination of moderate size (ca. 3.00–
<quantity id="4CF49B14A87AFFFD0CBA1341FEE3934A" box="[151,258,261,287]" metricMagnitude="-3" metricUnit="m" metricValue="3.39" pageId="25" pageNumber="26" unit="mm" value="3.39">3.39 mm</quantity>
), the relatively high and relatively punctate pronotal reliefs, and the male genitalia (
<figureCitation id="13372A74A87AFFFD08E01340FAC2934B" box="[1229,1315,260,286]" captionStart="FIGURE 16" captionStartId="33.[151,250,1736,1761]" captionTargetBox="[186,1401,210,1683]" captionTargetId="figure@33.[151,1435,181,1710]" captionTargetPageId="33" captionText="FIGURE 16. Hydrochus shorti n. sp., habitus and male genitalia of holotype." figureDoi="http://doi.org/10.5281/zenodo.3587215" httpUri="https://zenodo.org/record/3587215/files/figure.png" pageId="25" pageNumber="26">Fig. 16</figureCitation>
; described below). The general shape of the outline of the male genitalia show some similarity to that of
<taxonomicName id="4C0C4D72A87AFFFD08EB136DFAB19316" authority="Perkins, 2019" authorityName="Perkins" authorityYear="2019" box="[1222,1360,297,323]" class="Insecta" family="Hydrochidae" genus="Hydrochus" kingdom="Animalia" order="Coleoptera" pageId="25" pageNumber="26" phylum="Arthropoda" rank="species" species="formosus" status="sp. nov.">
<emphasis id="B978EAE3A87AFFFD08EB136DFAB19316" box="[1222,1360,297,323]" italics="true" pageId="25" pageNumber="26">H. formosus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A24B5798A87AFFFD0975136DFA769316" box="[1368,1431,297,323]" pageId="25" pageNumber="26" rank="species">n. sp.</taxonomicNameLabel>
, but the shape of the aedeagi markedly differ in the two species, among other genitalic differences.
<taxonomicName id="4C0C4D72A87AFFFD08F31309FAA09333" authority="Perkins, 2019" authorityName="Perkins" authorityYear="2019" box="[1246,1345,332,358]" class="Insecta" family="Hydrochidae" genus="Hydrochus" kingdom="Animalia" order="Coleoptera" pageId="25" pageNumber="26" phylum="Arthropoda" rank="species" species="shorti" status="sp. nov.">
<emphasis id="B978EAE3A87AFFFD08F31309FAA09333" box="[1246,1345,332,358]" italics="true" pageId="25" pageNumber="26">H. shorti</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A24B5798A87AFFFD09651309FA629332" box="[1352,1411,333,359]" pageId="25" pageNumber="26" rank="species">n. sp.</taxonomicNameLabel>
is smaller than
<taxonomicName id="4C0C4D72A87AFFFD0D051335FE5393DE" authority="Perkins, 2019" authorityName="Perkins" authorityYear="2019" box="[296,434,369,395]" class="Insecta" family="Hydrochidae" genus="Hydrochus" kingdom="Animalia" order="Coleoptera" pageId="25" pageNumber="26" phylum="Arthropoda" rank="species" species="formosus" status="sp. nov.">
<emphasis id="B978EAE3A87AFFFD0D051335FE5393DE" box="[296,434,369,395]" italics="true" pageId="25" pageNumber="26">H. formosus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A24B5798A87AFFFD0D951335FE1493DE" box="[440,501,369,395]" pageId="25" pageNumber="26" rank="species">n. sp.</taxonomicNameLabel>
(ca. 3.32 vs.
<quantity id="4CF49B14A87AFFFD0EA61335FD1293DE" box="[651,755,369,395]" metricMagnitude="-3" metricUnit="m" metricValue="4.04" pageId="25" pageNumber="26" unit="mm" value="4.04">4.04 mm</quantity>
), and males lack the abdominal tubercles present in that species. This species has a generalized
<taxonomicName id="4C0C4D72A87AFFFD0E0413D0FD4293FB" authorityName="Leach" authorityYear="1817" box="[553,675,404,430]" class="Insecta" family="Hydrochidae" genus="Hydrochus" kingdom="Animalia" order="Coleoptera" pageId="25" pageNumber="26" phylum="Arthropoda" rank="genus">
<emphasis id="B978EAE3A87AFFFD0E0413D0FD4293FB" box="[553,675,404,430]" italics="true" pageId="25" pageNumber="26">Hydrochus</emphasis>
</taxonomicName>
habitus; reliable determinations will require dissection of males.
</paragraph>
</subSubSection>
<subSubSection id="C316657AA87AFFFD0CEA13FCFDBF9663" pageId="25" pageNumber="26" type="description">
<paragraph id="8BB336F1A87AFFFD0CEA13FCFDB890D3" blockId="25.[151,1437,152,1151]" pageId="25" pageNumber="26">
<emphasis id="B978EAE3A87AFFFD0CEA13FCFEB09386" bold="true" box="[199,337,440,467]" pageId="25" pageNumber="26">Description</emphasis>
. Habitus as illustrated (
<figureCitation id="13372A74A87AFFFD0E7413FCFD4D9386" box="[601,684,440,467]" captionStart="FIGURE 16" captionStartId="33.[151,250,1736,1761]" captionTargetBox="[186,1401,210,1683]" captionTargetId="figure@33.[151,1435,181,1710]" captionTargetPageId="33" captionText="FIGURE 16. Hydrochus shorti n. sp., habitus and male genitalia of holotype." figureDoi="http://doi.org/10.5281/zenodo.3587215" httpUri="https://zenodo.org/record/3587215/files/figure.png" pageId="25" pageNumber="26">Fig. 16</figureCitation>
). Size:
<typeStatus id="54B78853A87AFFFD0ED313FCFC819387" box="[766,864,440,466]" pageId="25" pageNumber="26" type="holotype">holotype</typeStatus>
(length/width, mm): body (length to elytral apices) 3.32/1.23; head width 0.81; pronotum l/w 0.71/0.81; PA 0.81; PB 0.65; elytra 2.06/1.23. Body size moderate (ca. 3.00–
<quantity id="4CF49B14A87AFFFD0CFA1045FEA0904E" box="[215,321,513,539]" metricMagnitude="-3" metricUnit="m" metricValue="3.39" pageId="25" pageNumber="26" unit="mm" value="3.39">3.39 mm</quantity>
). Dorsum grey, with slight to moderately strong iridescence, elytra with black spots. Legs testaceous, with tibiofemoral joints darker. Punctation on elytra only slightly larger that of pronotum. Elytra interstriae ca. 1.5–
<date id="FFB21031A87AFFFD0CC2100CFEEA9036" box="[239,267,584,611]" pageId="25" pageNumber="26">2x</date>
strial puncture diameter. Interstria 9
<superScript id="7C799BB9A87AFFFD0E891003FD509000" attach="left" box="[676,689,583,597]" fontSize="6" pageId="25" pageNumber="26">th</superScript>
more convex than others, overhanging 10
<superScript id="7C799BB9A87AFFFD08A71003FB769000" attach="left" box="[1162,1175,583,597]" fontSize="6" pageId="25" pageNumber="26">th</superScript>
interstria. Callus on 5
<superScript id="7C799BB9A87AFFFD09BD1003FA7C9000" attach="none" box="[1424,1437,583,597]" fontSize="6" pageId="25" pageNumber="26">th</superScript>
moderately raised, moderately elongate.
</paragraph>
<paragraph id="8BB336F1A87AFFFD0CEA10D5FE3290A7" blockId="25.[151,1437,152,1151]" pageId="25" pageNumber="26">Pronotum length less than width (as ca. 71/81), widest slightly behind anterior margin, narrowed at base, sides sinuate, smooth; anterior margin slightly bisinuate; depressions rather deep, slightly more densely punctate than reliefs between depressions.</paragraph>
<paragraph id="8BB336F1A87AFFFD0CEA10B8FC44916F" blockId="25.[151,1437,152,1151]" pageId="25" pageNumber="26">
Elytra with spaces between strial punctures ca. 0.5–
<date id="FFB21031A87AFFFD0F2210B8FCC69142" box="[783,807,764,791]" pageId="25" pageNumber="26">1x</date>
their diameter; apices rather sharply conjointly rounded in dorsal view, in lateral view outer margin with moderate angulation.
</paragraph>
<paragraph id="8BB336F1A87AFFFD0CEA1100FA65910B" blockId="25.[151,1437,152,1151]" box="[199,1412,836,863]" pageId="25" pageNumber="26">Ventral characters: Mentum moderately coarsely punctate, with median fovea. Submentum with two foveae.</paragraph>
<paragraph id="8BB336F1A87AFFFD0CEA112CFB1391F3" blockId="25.[151,1437,152,1151]" pageId="25" pageNumber="26">
Male genitalia (
<figureCitation id="13372A74A87AFFFD0D51112CFE3191D6" box="[380,464,872,899]" captionStart="FIGURE 16" captionStartId="33.[151,250,1736,1761]" captionTargetBox="[186,1401,210,1683]" captionTargetId="figure@33.[151,1435,181,1710]" captionTargetPageId="33" captionText="FIGURE 16. Hydrochus shorti n. sp., habitus and male genitalia of holotype." figureDoi="http://doi.org/10.5281/zenodo.3587215" httpUri="https://zenodo.org/record/3587215/files/figure.png" pageId="25" pageNumber="26">Fig. 16</figureCitation>
) general characters: paramere tips in dorsal/ventral views arcuate toward midline, in lateral view narrow and tapering and arcing slightly ventrad, very lightly sclerotized; alm bisinuate.
</paragraph>
<paragraph id="8BB336F1A87AFFFD0CEA11F4FE6E91BB" blockId="25.[151,1437,152,1151]" pageId="25" pageNumber="26">Dorsal surface: adtl near midlength, not distinctive; pdmm sinuate and at points slightly separated from or beneath alm; adbl small.</paragraph>
<paragraph id="8BB336F1A87AFFFD0CEA11BCFDBF9663" blockId="25.[151,1437,152,1151]" pageId="25" pageNumber="26">Ventral surface: pvmm approximating alm; pseudoparameres extending past midlength of parameres, m rather large in lateral view; avtl very indistinct.</paragraph>
</subSubSection>
<subSubSection id="C316657AA87AFFFD0CEA1605FE6F962A" pageId="25" pageNumber="26" type="etymology">
<paragraph id="8BB336F1A87AFFFD0CEA1605FE6F962A" blockId="25.[151,1437,152,1151]" pageId="25" pageNumber="26">
<emphasis id="B978EAE3A87AFFFD0CEA1605FEAF960E" bold="true" box="[199,334,1089,1115]" pageId="25" pageNumber="26">Etymology.</emphasis>
Named for Andrew E. Z. Short, an outstanding water beetle collector who has netted many new species of
<taxonomicName id="4C0C4D72A87AFFFD0D201620FE66962B" authorityName="Leach" authorityYear="1817" box="[269,391,1124,1150]" class="Insecta" family="Hydrochidae" genus="Hydrochus" kingdom="Animalia" order="Coleoptera" pageId="25" pageNumber="26" phylum="Arthropoda" rank="genus">
<emphasis id="B978EAE3A87AFFFD0D201620FE66962B" box="[269,391,1124,1150]" italics="true" pageId="25" pageNumber="26">Hydrochus</emphasis>
</taxonomicName>
.
</paragraph>
</subSubSection>
</treatment>
</document>
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