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treatments-xml/data/03/A5/87/03A587E7A869FFE80CBA1789FBEE9142.xml
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<document id="6EE753A0CD82B7BD581B902CA8D3FFCD" ID-CLB-Dataset="24540" ID-DOI="10.11646/zootaxa.4708.1.1" ID-GBIF-Dataset="653d580d-e1c7-499c-8604-4b5a7b1920ef" ID-ISSN="1175-5326" ID-Zenodo-Dep="3587181" ID-ZooBank="09F24F0B-5090-4089-A525-8E5150C91EF7" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1576825058981" checkinUser="plazi" docAuthor="Perkins, Philip D." docDate="2019" docId="03A587E7A869FFE80CBA1789FBEE9142" docLanguage="en" docName="zootaxa.4708.1.1.pdf" docOrigin="Zootaxa 4708 (1)" docStyle="DocumentStyle:5EBBA59367AD13919D70D935FA04F6A3.14:Zootaxa.2013-.monograph" docStyleId="5EBBA59367AD13919D70D935FA04F6A3" docStyleName="Zootaxa.2013-.monograph" docStyleVersion="14" docTitle="Hydrochus ducalis Knisch" docType="treatment" docVersion="7" lastPageNumber="13" masterDocId="FF9CFF9FA863FFE40C2D1244FFE19255" masterDocTitle="Taxonomy of Venezuelan water beetles in the genus Hydrochus Leach, 1817, and an analysis of male genitalia morphology (Coleoptera: Hydrochidae)" masterLastPageNumber="59" masterPageNumber="1" pageNumber="11" updateTime="1698775642300" updateUser="ExternalLinkService">
<mods:mods id="3B669B935190F0D13AEF55B314D0EDD7" xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo id="79E8E23B04E77623E09318D490C7CA79">
<mods:title id="DE74E327EB2C3440025AF9CA31E66C7D">Taxonomy of Venezuelan water beetles in the genus Hydrochus Leach, 1817, and an analysis of male genitalia morphology (Coleoptera: Hydrochidae)</mods:title>
</mods:titleInfo>
<mods:name id="3F12F2A2D722B499D65197F525AD90EB" type="personal">
<mods:role id="8A83EEA838CE7FF5192F58FEBFE526D6">
<mods:roleTerm id="63CE9176F113066CB1444348233C12CE">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="484344305AE34AB04FA5A1644D4C63FC">Perkins, Philip D.</mods:namePart>
</mods:name>
<mods:typeOfResource id="7EC41078D571A8E821FDE875DE08731E">text</mods:typeOfResource>
<mods:relatedItem id="5077980F5E16335ABCD3E7BB29EB69F9" type="host">
<mods:titleInfo id="343709C433CA49A6E32C1BAF056E3FEF">
<mods:title id="A75DE34D11624160B5D31B6C3F824EC5">Zootaxa</mods:title>
</mods:titleInfo>
<mods:part id="67BCEFDA46F9313321B17B4FF03140DE">
<mods:date id="A3EB5B6EDE2B31EF1A002C0597954062">2019</mods:date>
<mods:detail id="803BE35E0016B3234841517588C8FD20" type="pubDate">
<mods:number id="FFE32476073FD31E6140C93A437D5523">2019-12-12</mods:number>
</mods:detail>
<mods:detail id="FB51C7A602D104893B981CA52E662A3C" type="volume">
<mods:number id="4DB1D63DF3D3ADED1969666A9B9E3513">4708</mods:number>
</mods:detail>
<mods:detail id="CCF9CF0AB849D9CD544DC8781FAD7462" type="issue">
<mods:number id="F12B2DF96B6B8037BFB46AAB0AE22D81">1</mods:number>
</mods:detail>
<mods:extent id="4891F7F2FD4F6CEBA32C9F4144205167" unit="page">
<mods:start id="F81A8518E7C1477C2349B655DB577FB2">1</mods:start>
<mods:end id="903033B23126CD104D2B3DBF693FCF6F">59</mods:end>
</mods:extent>
</mods:part>
</mods:relatedItem>
<mods:classification id="6F9BE6546198DD97BDDAE716B910FBB1">journal article</mods:classification>
<mods:identifier id="C753301DA9AD98A60BF211CD485C9029" type="CLB-Dataset">24540</mods:identifier>
<mods:identifier id="1354DA123D826854AF580385E030384B" type="DOI">10.11646/zootaxa.4708.1.1</mods:identifier>
<mods:identifier id="85D8FCEED20882B61FF82D22EA2861C1" type="GBIF-Dataset">653d580d-e1c7-499c-8604-4b5a7b1920ef</mods:identifier>
<mods:identifier id="737F43023DA51CE949B9773E2B2BF92A" type="ISSN">1175-5326</mods:identifier>
<mods:identifier id="485605E178B344BA413275023CE7266B" type="Zenodo-Dep">3587181</mods:identifier>
<mods:identifier id="EEC3E4049D63185C59CD887D76419414" type="ZooBank">09F24F0B-5090-4089-A525-8E5150C91EF7</mods:identifier>
</mods:mods>
<treatment id="03A587E7A869FFE80CBA1789FBEE9142" ID-GBIF-Taxon="160798409" LSID="urn:lsid:plazi:treatment:03A587E7A869FFE80CBA1789FBEE9142" httpUri="http://treatment.plazi.org/id/03A587E7A869FFE80CBA1789FBEE9142" lastPageId="12" lastPageNumber="13" pageId="10" pageNumber="11">
<subSubSection id="C316657AA869FFEE0CBA1789FE3B97B2" box="[151,474,1485,1512]" pageId="10" pageNumber="11" type="nomenclature">
<paragraph id="8BB336F1A869FFEE0CBA1789FE3B97B2" blockId="10.[151,474,1485,1547]" box="[151,474,1485,1512]" pageId="10" pageNumber="11">
<heading id="D0FB819DA869FFEE0CBA1789FE3B97B2" bold="true" box="[151,474,1485,1512]" fontSize="11" level="1" pageId="10" pageNumber="11" reason="1">
<taxonomicName id="4C0C4D72A869FFEE0CBA1789FE3B97B2" ID-CoL="3N67D" authority="Knisch" authorityName="Knisch" box="[151,474,1485,1512]" class="Insecta" family="Hydrochidae" genus="Hydrochus" kingdom="Animalia" order="Coleoptera" pageId="10" pageNumber="11" phylum="Arthropoda" rank="species" species="ducalis">
<emphasis id="B978EAE3A869FFEE0CBA1789FE3B97B2" bold="true" box="[151,474,1485,1512]" pageId="10" pageNumber="11">
<emphasis id="B978EAE3A869FFEE0CBA1789FE9B97BD" bold="true" box="[151,378,1485,1512]" italics="true" pageId="10" pageNumber="11">Hydrochus ducalis</emphasis>
Knisch
</emphasis>
</taxonomicName>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C316657AA869FFEE0CBA17B5FED1945E" box="[151,304,1521,1547]" pageId="10" pageNumber="11" type="description">
<paragraph id="8BB336F1A869FFEE0CBA17B5FED1945E" blockId="10.[151,474,1485,1547]" box="[151,304,1521,1547]" pageId="10" pageNumber="11">
<figureCitation id="13372A74A869FFEE0CBA17B5FF02945E" box="[151,227,1521,1547]" captionStart="FIGURE 1" captionStartId="11.[151,250,1894,1919]" captionTargetBox="[156,1422,330,1824]" captionTargetId="figure@11.[151,1435,306,1866]" captionTargetPageId="11" captionText="FIGURE 1. Dorsal habitus of Hydrochus species." figureDoi="http://doi.org/10.5281/zenodo.3587183" httpUri="https://zenodo.org/record/3587183/files/figure.png" pageId="10" pageNumber="11">Figs. 1</figureCitation>
,
<figureCitation id="13372A74A869FFEE0CDC17B5FEF6945E" box="[241,279,1521,1547]" captionStart="FIGURES 6" captionStartId="19.[151,264,1522,1547]" captionTargetBox="[151,1435,181,1496]" captionTargetId="figure@19.[151,1435,181,1496]" captionTargetPageId="19" captionText="FIGURES 6A–C. (A) Male genitalia of H. ducalis Knisch, with parameres spread, showing shape of aedeagus; (B) Male genitalia of H. variabilis Knisch and (C) H. n. sp. from Peru, showing the articulation point of the paramere dorsal basal lobe with the aedeagus dorsal basal lobe." figureDoi="http://doi.org/10.5281/zenodo.3587193" httpUri="https://zenodo.org/record/3587193/files/figure.png" pageId="10" pageNumber="11">6A</figureCitation>
,
<figureCitation id="13372A74A869FFEE0D0F17B5FED1945E" box="[290,304,1521,1547]" captionStart="FIGURE 8" captionStartId="23.[151,250,1965,1990]" captionTargetBox="[190,1388,466,1915]" captionTargetId="figure@23.[151,1435,453,1940]" captionTargetPageId="23" captionText="FIGURE 8. Hydrochus ducalis Knisch, habitus and male genitalia." figureDoi="http://doi.org/10.5281/zenodo.3587197" httpUri="https://zenodo.org/record/3587197/files/figure.png" pageId="10" pageNumber="11">8</figureCitation>
</paragraph>
</subSubSection>
<subSubSection id="C316657AA869FFEE0CBA1472FE1D941B" box="[151,508,1589,1614]" pageId="10" pageNumber="11" type="reference_group">
<paragraph id="8BB336F1A869FFEE0CBA1472FE1D941B" blockId="10.[151,508,1589,1614]" box="[151,508,1589,1614]" pageId="10" pageNumber="11">
<treatmentCitationGroup id="AB1C11DFA869FFEE0CBA1472FE1D941B" box="[151,508,1589,1614]" pageId="10" pageNumber="11">
<taxonomicName id="4C0C4D72A869FFEE0CBA1472FE16941B" ID-CoL="3N67D" authority="Knisch 1921: 1" authorityName="Knisch" authorityPageNumber="1" authorityYear="1921" box="[151,503,1589,1614]" class="Insecta" family="Hydrochidae" genus="Hydrochus" kingdom="Animalia" order="Coleoptera" pageId="10" pageNumber="11" phylum="Arthropoda" rank="species" species="ducalis">
<emphasis id="B978EAE3A869FFEE0CBA1472FEB4941B" box="[151,341,1590,1614]" italics="true" pageId="10" pageNumber="11">Hydrochus ducalis</emphasis>
<treatmentCitation id="0AAD10E0A869FFEE0D761471FE16941B" author="Knisch, A." box="[347,503,1589,1614]" page="1" pageId="10" pageNumber="11" year="1921">
<bibRefCitation id="EF9D4B00A869FFEE0D761471FE16941B" author="Knisch, A." box="[347,503,1589,1614]" pageId="10" pageNumber="11" pagination="1 - 24" refId="ref30148" refString="Knisch, A. (1921) Hydrophiliden aus Matto Grosso (Erster Teil). Archiv fur Naturegeschichte, 87 A, 1 - 24." type="journal article" year="1921">Knisch 1921: 1</bibRefCitation>
</treatmentCitation>
</taxonomicName>
.
</treatmentCitationGroup>
</paragraph>
</subSubSection>
<subSubSection id="C316657AA869FFEF0CBA143FFF01934A" lastPageId="11" lastPageNumber="12" pageId="10" pageNumber="11" type="materials_examined">
<paragraph id="8BB336F1A869FFEE0CBA143FFE3595C7" blockId="10.[151,1437,1659,2011]" pageId="10" pageNumber="11">
<emphasis id="B978EAE3A869FFEE0CBA143FFE5594C3" bold="true" box="[151,436,1659,1686]" pageId="10" pageNumber="11">Material examined (28).</emphasis>
<materialsCitation id="3B643CACA869FFEE0D941438FBB994C3" ID-GBIF-Occurrence="2521263694" box="[441,1112,1659,1686]" collectionCode="SEMC, MIZA, MALUZ" country="Venezuela" location="Apure" pageId="10" pageNumber="11" specimenCount="1" stateProvince="Apure">
<emphasis id="B978EAE3A869FFEE0D941438FD7794C3" bold="true" box="[441,662,1659,1686]" pageId="10" pageNumber="11">
<collectingCountry id="F31B7661A869FFEE0D941438FDD794C3" box="[441,566,1660,1686]" name="Venezuela" pageId="10" pageNumber="11">Venezuela</collectingCountry>
:
<collectingRegion id="49C8F813A869FFEE0E6C143FFD7094C3" box="[577,657,1659,1686]" country="Venezuela" name="Apure" pageId="10" pageNumber="11">Apure</collectingRegion>
:
</emphasis>
AS-06-009 (8
<collectionCode id="ED1DAE34A869FFEE0F171438FC6894C3" box="[826,905,1660,1686]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34264" name="University of Kansas - Biodiversity Institute" pageId="10" pageNumber="11" type="Museum">SEMC</collectionCode>
,
<collectionCode id="ED1DAE34A869FFEE0FBE1438FC3C94C3" box="[915,989,1660,1686]" country="Venezuela" httpUri="http://biocol.org/urn:lsid:biocol.org:col:33834" lsid="urn:lsid:biocol.org:col:33834" name="Museo del Instituto de Zoologia Agricola Francisco Fernandez Yepez" pageId="10" pageNumber="11" type="Museum">MIZA</collectionCode>
,
<collectionCode id="ED1DAE34A869FFEE0FCA1438FBB194C3" box="[999,1104,1660,1686]" pageId="10" pageNumber="11">MALUZ</collectionCode>
)
</materialsCitation>
;
<materialsCitation id="3B643CACA869FFEE084C1438FA7894C3" ID-GBIF-Occurrence="2521263690" box="[1121,1433,1660,1686]" collectingDate="2001-10" collectionCode="SEMC" country="Venezuela" pageId="10" pageNumber="11" specimenCode="VZ09-0117" specimenCount="1" stateProvince="Apure">
<specimenCode id="DBAA9E8AA869FFEE084C1438FB0794C3" box="[1121,1254,1660,1686]" collectionCode="MALUZ" pageId="10" pageNumber="11">VZ09-0117</specimenCode>
-
<date id="FFB21031A869FFEE08C01438FAFC94C3" box="[1261,1309,1660,1686]" pageId="10" pageNumber="11" value="2001-10">
<collectingDate id="EFF6E9D9A869FFEE08C01438FAFC94C3" box="[1261,1309,1660,1686]" pageId="10" pageNumber="11" value="2001-10">01X</collectingDate>
</date>
(1
<collectionCode id="ED1DAE34A869FFEE096D1438FA7094C3" box="[1344,1425,1660,1686]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34264" name="University of Kansas - Biodiversity Institute" pageId="10" pageNumber="11" type="Museum">SEMC</collectionCode>
)
</materialsCitation>
.
<materialsCitation id="3B643CACA869FFEE0CBA14DBFDA094EF" ID-GBIF-Occurrence="2521263695" box="[151,577,1695,1722]" collectionCode="SEMC" country="Venezuela" location="Bolivar" pageId="10" pageNumber="11" specimenCount="1" stateProvince="Bolivar">
<emphasis id="B978EAE3A869FFEE0CBA14DBFF1894EF" bold="true" box="[151,249,1695,1722]" pageId="10" pageNumber="11">
<collectingRegion id="49C8F813A869FFEE0CBA14DBFF1594EF" box="[151,244,1695,1722]" country="Venezuela" name="Bolivar" pageId="10" pageNumber="11">Bolivar</collectingRegion>
:
</emphasis>
VZ09-0112-01B (1
<collectionCode id="ED1DAE34A869FFEE0DC514E4FDD894EF" box="[488,569,1696,1722]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34264" name="University of Kansas - Biodiversity Institute" pageId="10" pageNumber="11" type="Museum">SEMC</collectionCode>
)
</materialsCitation>
;
<materialsCitation id="3B643CACA869FFEE0E6214DBFAAE94EF" ID-GBIF-Occurrence="2521263697" box="[591,1359,1695,1723]" collectingDate="1996-06-11" collectionCode="CMN" collectorName="H. &amp; A. Howden" country="Venezuela" location="Caicara" pageId="10" pageNumber="11" specimenCount="1" stateProvince="Bolivar">
<quantity id="4CF49B14A869FFEE0E6214DBFD6A94EF" box="[591,651,1695,1722]" metricMagnitude="3" metricUnit="m" metricValue="5.0" pageId="10" pageNumber="11" unit="km" value="5.0">5 km</quantity>
E.
<location id="8ED3602AA869FFEE0E9A14E4FCEE94EF" LSID="urn:lsid:plazi:treatment:03A587E7A869FFE80CBA1789FBEE9142:8ED3602AA869FFEE0E9A14E4FCEE94EF" box="[695,783,1696,1722]" country="Venezuela" name="Caicara" pageId="10" pageNumber="11" stateProvince="Bolivar">Caicara</location>
,
<date id="FFB21031A869FFEE0F3014E4FC7F94EF" box="[797,926,1696,1722]" pageId="10" pageNumber="11" value="1996-06-11">
<collectingDate id="EFF6E9D9A869FFEE0F3014E4FC7F94EF" box="[797,926,1696,1722]" pageId="10" pageNumber="11" value="1996-06-11">11.VI.1996</collectingDate>
</date>
, at light,
<collectorName id="26F95327A869FFEE082214E5FB3694EF" box="[1039,1239,1696,1723]" pageId="10" pageNumber="11">H. &amp; A. Howden</collectorName>
(2
<collectionCode id="ED1DAE34A869FFEE092F14E4FAA694EF" box="[1282,1351,1696,1722]" country="Canada" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34935" lsid="urn:lsid:biocol.org:col:34935" name="Canadian Museum of Nature" pageId="10" pageNumber="11" type="Museum">CMN</collectionCode>
)
</materialsCitation>
.
<materialsCitation id="3B643CACA869FFEE097114E4FDB7948B" ID-GBIF-Occurrence="2521263816" collectionCode="SEMC" country="Venezuela" location="Delta Amacuro" pageId="10" pageNumber="11" specimenCount="1" stateProvince="Delta Amacuro">
<emphasis id="B978EAE3A869FFEE097114E4FEF1948B" bold="true" pageId="10" pageNumber="11">
<collectingRegion id="49C8F813A869FFEE097114E4FEEA948B" country="Venezuela" name="Delta Amacuro" pageId="10" pageNumber="11">Delta Amacuro</collectingRegion>
:
</emphasis>
VZ10-0203-02A (2
<collectionCode id="ED1DAE34A869FFEE0DD01480FDAF948B" box="[509,590,1732,1758]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34264" name="University of Kansas - Biodiversity Institute" pageId="10" pageNumber="11" type="Museum">SEMC</collectionCode>
)
</materialsCitation>
.
<materialsCitation id="3B643CACA869FFEE0E4E1487FDE69556" ID-GBIF-Occurrence="2521263741" collectingDate="1984-06-17" collectingDateMax="1984-06-18" collectingDateMin="1984-06-17" collectionCode="NMNH" collectorName="F. W. Eiland &amp; V. Linares" country="Venezuela" location="Corozo Pando" pageId="10" pageNumber="11" specimenCount="1" stateProvince="Guarico">
<emphasis id="B978EAE3A869FFEE0E4E1487FD2F948B" bold="true" box="[611,718,1731,1758]" pageId="10" pageNumber="11">
<collectingRegion id="49C8F813A869FFEE0E4E1487FD28948B" box="[611,713,1731,1758]" country="Venezuela" name="Guarico" pageId="10" pageNumber="11">Guarico</collectingRegion>
:
</emphasis>
<location id="8ED3602AA869FFEE0EF51480FC9D948B" LSID="urn:lsid:plazi:treatment:03A587E7A869FFE80CBA1789FBEE9142:8ED3602AA869FFEE0EF51480FC9D948B" box="[728,892,1732,1758]" country="Venezuela" name="Corozo Pando" pageId="10" pageNumber="11" stateProvince="Guarico">Corozo Pando</location>
, (
<quantity id="4CF49B14A869FFEE0FBE1480FC2F948B" box="[915,974,1732,1758]" metricMagnitude="3" metricUnit="m" metricValue="8.0" pageId="10" pageNumber="11" unit="km" value="8.0">8 km</quantity>
N.),
<date id="FFB21031A869FFEE08211480FB32948B" box="[1036,1235,1732,1759]" pageId="10" pageNumber="11" value="1984-06-17" valueMax="1984-06-18" valueMin="1984-06-17">
<collectingDate id="EFF6E9D9A869FFEE08211480FB32948B" box="[1036,1235,1732,1759]" pageId="10" pageNumber="11" value="1984-06-17" valueMax="1984-06-18" valueMin="1984-06-17">17–18 June 1984</collectingDate>
</date>
, blacklight,
<collectorName id="26F95327A869FFEE094D1481FF019557" pageId="10" pageNumber="11">F. W. Eiland</collectorName>
&amp;
<collectorName id="26F95327A869FFEE0D2A14ADFE9A9557" box="[263,379,1768,1795]" pageId="10" pageNumber="11">V. Linares</collectorName>
(2
<collectionCode id="ED1DAE34A869FFEE0D8914ADFE1E9556" box="[420,511,1769,1795]" country="USA" httpUri="http://grbio.org/cool/142r-0w94" name="Smithsonian Institution, National Museum of Natural History" pageId="10" pageNumber="11" type="Museum">NMNH</collectionCode>
)
</materialsCitation>
;
<materialsCitation id="3B643CACA869FFEE0E3914ACFEB19572" ID-GBIF-Occurrence="2521263710" collectingDate="1984-06-20" collectingDateMax="1984-06-21" collectingDateMin="1984-06-20" collectionCode="NMNH" collectorName="F. W. Eiland &amp; V. Linares" country="Venezuela" location="Corozo Pando" pageId="10" pageNumber="11" specimenCount="1" stateProvince="Guarico">
<location id="8ED3602AA869FFEE0E3914ACFD599557" LSID="urn:lsid:plazi:treatment:03A587E7A869FFE80CBA1789FBEE9142:8ED3602AA869FFEE0E3914ACFD599557" box="[532,696,1768,1794]" country="Venezuela" name="Corozo Pando" pageId="10" pageNumber="11" stateProvince="Guarico">Corozo Pando</location>
, (
<quantity id="4CF49B14A869FFEE0EE314ACFCE89557" box="[718,777,1768,1794]" metricMagnitude="3" metricUnit="m" metricValue="8.0" pageId="10" pageNumber="11" unit="km" value="8.0">8 km</quantity>
N.),
<date id="FFB21031A869FFEE0F6814ACFBEA9557" box="[837,1035,1768,1795]" pageId="10" pageNumber="11" value="1984-06-20" valueMax="1984-06-21" valueMin="1984-06-20">
<collectingDate id="EFF6E9D9A869FFEE0F6814ACFBEA9557" box="[837,1035,1768,1795]" pageId="10" pageNumber="11" value="1984-06-20" valueMax="1984-06-21" valueMin="1984-06-20">20–21 June 1984</collectingDate>
</date>
, blacklight,
<collectorName id="26F95327A869FFEE08B514ADFAC49557" box="[1176,1317,1768,1795]" pageId="10" pageNumber="11">F. W. Eiland</collectorName>
&amp;
<collectorName id="26F95327A869FFEE096114ADFF259572" pageId="10" pageNumber="11">V. Linares</collectorName>
(3
<collectionCode id="ED1DAE34A869FFEE0CC01549FEA99572" box="[237,328,1805,1831]" country="USA" httpUri="http://grbio.org/cool/142r-0w94" name="Smithsonian Institution, National Museum of Natural History" pageId="10" pageNumber="11" type="Museum">NMNH</collectionCode>
)
</materialsCitation>
;
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<quantity id="4CF49B14A869FFEE0D70154FFE479573" box="[349,422,1803,1830]" metricMagnitude="4" metricUnit="m" metricValue="1.5" pageId="10" pageNumber="11" unit="km" value="15.0">15 km</quantity>
<collectorName id="26F95327A869FFEE0D831548FDD99573" box="[430,568,1804,1830]" pageId="10" pageNumber="11">S. Calabozo</collectorName>
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</date>
, collected in
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,
<collectorName id="26F95327A869FFEE087F1549FAEF9573" box="[1106,1294,1804,1831]" pageId="10" pageNumber="11">P. &amp; P. Spangler</collectorName>
(1
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)
</materialsCitation>
.
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<emphasis id="B978EAE3A869FFEE0CBA156BFEEB951F" bold="true" box="[151,266,1839,1866]" pageId="10" pageNumber="11">
<collectingRegion id="49C8F813A869FFEE0CBA156BFEE4951F" box="[151,261,1839,1866]" country="Venezuela" name="Miranda" pageId="10" pageNumber="11">Miranda</collectingRegion>
:
</emphasis>
<collectorName id="26F95327A869FFEE0D3E1574FE41951E" box="[275,416,1840,1867]" pageId="10" pageNumber="11">Guatopo NP</collectorName>
,
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,
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<quantity id="4CF49B14A869FFEE0E031574FD96951F" box="[558,631,1840,1866]" metricMagnitude="4" metricUnit="m" metricValue="2.8" pageId="10" pageNumber="11" unit="km" value="28.0">28 km</quantity>
N Altagracia
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,
<quantity id="4CF49B14A869FFEE0F321574FC83951F" box="[799,866,1840,1866]" metricMagnitude="2" metricUnit="m" metricValue="6.2" pageId="10" pageNumber="11" unit="m" value="620.0">
<elevation id="0021D1C2A869FFEE0F321574FC83951F" box="[799,866,1840,1866]" metricMagnitude="2" metricUnit="m" metricValue="6.2" pageId="10" pageNumber="11" unit="m" value="620.0">620m</elevation>
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,
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<collectingDate id="EFF6E9D9A869FFEE0F421574FBEC951F" box="[879,1037,1840,1866]" pageId="10" pageNumber="11" value="1987-06-01" valueMax="1987-06-14" valueMin="1987-06-01">1–14.VI.1987</collectingDate>
</date>
,
<collectorName id="26F95327A869FFEE08371574FB6D951F" box="[1050,1164,1840,1866]" pageId="10" pageNumber="11">S&amp;J Peck</collectorName>
,
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87-17 (1
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)
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.
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<emphasis id="B978EAE3A869FFEE0CC31510FE84953B" bold="true" box="[238,357,1876,1902]" pageId="10" pageNumber="11">
<collectingRegion id="49C8F813A869FFEE0CC31510FEBE953B" box="[238,351,1876,1902]" country="Venezuela" name="Monagas" pageId="10" pageNumber="11">Monagas</collectingRegion>
:
</emphasis>
VZ10-0202-01B (1
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)
</materialsCitation>
;
<materialsCitation id="3B643CACA869FFEE0E821510FC08953B" ID-GBIF-Occurrence="2521263714" box="[687,1001,1876,1902]" collectionCode="SEMC" country="Venezuela" pageId="10" pageNumber="11" specimenCode="VZ10-0202, 03" specimenCount="1" stateProvince="Monagas">
<specimenCode id="DBAA9E8AA869FFEE0E821510FCD2953B" box="[687,819,1876,1902]" collectionCode="SEMC" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34264" name="University of Kansas - Biodiversity Institute" pageId="10" pageNumber="11" type="Museum">VZ10-0202</specimenCode>
-
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A (3
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<emphasis id="B978EAE3A869FFEE0FDE1517FBB9953B" bold="true" box="[1011,1112,1875,1902]" pageId="10" pageNumber="11">
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:
</emphasis>
VZ09-0716-07A (1
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)
</materialsCitation>
;
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<specimenCode id="DBAA9E8AA869FFEE0CBA153CFEF895C7" box="[151,281,1912,1938]" collectionCode="SEMC" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34264" name="University of Kansas - Biodiversity Institute" pageId="10" pageNumber="11" type="Museum">VZ12-0126</specimenCode>
-
<specimenCode id="DBAA9E8AA869FFEE0D0C153CFEDC95C7" box="[289,317,1912,1938]" collectionCode="SEMC" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34264" name="University of Kansas - Biodiversity Institute" pageId="10" pageNumber="11" type="Museum">02</specimenCode>
A (2
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.
</paragraph>
<paragraph id="8BB336F1A869FFEE0CEA15DFFA7C958E" blockId="10.[151,1437,1659,2011]" pageId="10" pageNumber="11">
<emphasis id="B978EAE3A869FFEE0CEA15DFFE3095E3" bold="true" box="[199,465,1947,1974]" pageId="10" pageNumber="11">Differential Diagnosis.</emphasis>
Among Venezuelan species, recognized by the combination of large size (ca.
<quantity id="4CF49B14A869FFEE090115DFFF26958E" metricMagnitude="-3" metricUnit="m" metricValue="5.77" metricValueMax="6.31" metricValueMin="5.23" pageId="10" pageNumber="11" unit="mm" value="5.77" valueMax="6.31" valueMin="5.23">5.23–6.31 mm</quantity>
), the only moderately coarse dorsal punctation, and the distinctive male genitalia (
<figureCitation id="13372A74A869FFEE08741584FB5C958F" box="[1113,1213,1984,2010]" captionStart="FIGURES 6" captionStartId="19.[151,264,1522,1547]" captionTargetBox="[151,1435,181,1496]" captionTargetId="figure@19.[151,1435,181,1496]" captionTargetPageId="19" captionText="FIGURES 6A–C. (A) Male genitalia of H. ducalis Knisch, with parameres spread, showing shape of aedeagus; (B) Male genitalia of H. variabilis Knisch and (C) H. n. sp. from Peru, showing the articulation point of the paramere dorsal basal lobe with the aedeagus dorsal basal lobe." figureDoi="http://doi.org/10.5281/zenodo.3587193" httpUri="https://zenodo.org/record/3587193/files/figure.png" pageId="10" pageNumber="11">Figs. 6A</figureCitation>
,
<figureCitation id="13372A74A869FFEE08E51584FB39958F" box="[1224,1240,1984,2010]" captionStart="FIGURE 8" captionStartId="23.[151,250,1965,1990]" captionTargetBox="[190,1388,466,1915]" captionTargetId="figure@23.[151,1435,453,1940]" captionTargetPageId="23" captionText="FIGURE 8. Hydrochus ducalis Knisch, habitus and male genitalia." figureDoi="http://doi.org/10.5281/zenodo.3587197" httpUri="https://zenodo.org/record/3587197/files/figure.png" pageId="10" pageNumber="11">8</figureCitation>
; describe below).
</paragraph>
<paragraph id="8BB336F1A868FFEF0CBA12DDFF01934A" blockId="11.[151,1436,153,287]" pageId="11" pageNumber="12">
It is the largest Venezuelan
<taxonomicName id="4C0C4D72A868FFEF0DE412DDFDA292E6" authorityName="Leach" authorityYear="1817" box="[457,579,153,179]" class="Insecta" family="Hydrochidae" genus="Hydrochus" kingdom="Animalia" order="Coleoptera" pageId="11" pageNumber="12" phylum="Arthropoda" rank="genus">
<emphasis id="B978EAE3A868FFEF0DE412DDFDA292E6" box="[457,579,153,179]" italics="true" pageId="11" pageNumber="12">Hydrochus</emphasis>
</taxonomicName>
, and compared with other large species is not very coarsely densely punctate as is
<taxonomicName id="4C0C4D72A868FFEF0CE212FAFE7A9282" authorityName="Oliva" authorityYear="1996" box="[207,411,189,215]" class="Insecta" family="Hydrochidae" genus="Hydrochus" kingdom="Animalia" order="Coleoptera" pageId="11" pageNumber="12" phylum="Arthropoda" rank="species" species="pseudosecretus">
<emphasis id="B978EAE3A868FFEF0CE212FAFE7A9282" box="[207,411,189,215]" italics="true" pageId="11" pageNumber="12">H. pseudosecretus</emphasis>
</taxonomicName>
, and is generally larger than
<taxonomicName id="4C0C4D72A868FFEF0EF312FAFCB59282" box="[734,852,189,215]" class="Insecta" family="Hydrochidae" genus="Hydrochus" kingdom="Animalia" order="Coleoptera" pageId="11" pageNumber="12" phylum="Arthropoda" rank="species" species="tarsalis">
<emphasis id="B978EAE3A868FFEF0EF312FAFCB59282" box="[734,852,189,215]" italics="true" pageId="11" pageNumber="12">H. tarsalis</emphasis>
</taxonomicName>
, and slightly broader (ca. 5.65 vs.
<quantity id="4CF49B14A868FFEF08FB12F9FADC9282" box="[1238,1341,189,215]" metricMagnitude="-3" metricUnit="m" metricValue="4.77" pageId="11" pageNumber="12" unit="mm" value="4.77">4.77 mm</quantity>
; TL/EW ca. 2.97 vs. 3.01). However, reliably separating specimens of
<taxonomicName id="4C0C4D72A868FFEF0F7912A6FC2D92AE" box="[852,972,225,251]" class="Insecta" family="Hydrochidae" genus="Hydrochus" kingdom="Animalia" order="Coleoptera" pageId="11" pageNumber="12" phylum="Arthropoda" rank="species" species="tarsalis">
<emphasis id="B978EAE3A868FFEF0F7912A6FC2D92AE" box="[852,972,225,251]" italics="true" pageId="11" pageNumber="12">H. tarsalis</emphasis>
</taxonomicName>
and
<taxonomicName id="4C0C4D72A868FFEF082812A6FB9892AE" box="[1029,1145,225,251]" class="Insecta" family="Hydrochidae" genus="Hydrochus" kingdom="Animalia" order="Coleoptera" pageId="11" pageNumber="12" phylum="Arthropoda" rank="species" species="ducalis">
<emphasis id="B978EAE3A868FFEF082812A6FB9892AE" box="[1029,1145,225,251]" italics="true" pageId="11" pageNumber="12">H. ducalis</emphasis>
</taxonomicName>
will require dissection of males.
</paragraph>
</subSubSection>
<caption id="DF736679A868FFEF0CBA1522FD72952A" ID-DOI="http://doi.org/10.5281/zenodo.3587183" ID-Zenodo-Dep="3587183" box="[151,659,1894,1919]" httpUri="https://zenodo.org/record/3587183/files/figure.png" pageId="11" pageNumber="12" startId="11.[151,250,1894,1919]" targetBox="[156,1422,330,1824]" targetPageId="11">
<paragraph id="8BB336F1A868FFEF0CBA1522FD72952A" blockId="11.[151,659,1894,1919]" box="[151,659,1894,1919]" pageId="11" pageNumber="12">
<emphasis id="B978EAE3A868FFEF0CBA1522FEF2952A" bold="true" box="[151,275,1894,1919]" pageId="11" pageNumber="12">FIGURE 1.</emphasis>
Dorsal habitus of
<taxonomicName id="4C0C4D72A868FFEF0DE31522FDDD952B" authorityName="Leach" authorityYear="1817" box="[462,572,1894,1918]" class="Insecta" family="Hydrochidae" genus="Hydrochus" kingdom="Animalia" order="Coleoptera" pageId="11" pageNumber="12" phylum="Arthropoda" rank="genus">
<emphasis id="B978EAE3A868FFEF0DE31522FDDD952B" box="[462,572,1894,1918]" italics="true" pageId="11" pageNumber="12">Hydrochus</emphasis>
</taxonomicName>
species.
</paragraph>
</caption>
<subSubSection id="C316657AA868FFE80CEA15E8FF0C90A6" lastPageId="12" lastPageNumber="13" pageId="11" pageNumber="12" type="description">
<paragraph id="8BB336F1A868FFE80CEA15E8FAE5934A" blockId="11.[151,1437,1964,2028]" lastBlockId="12.[151,1437,152,791]" lastPageId="12" lastPageNumber="13" pageId="11" pageNumber="12">
<emphasis id="B978EAE3A868FFEF0CEA15E8FEB09592" bold="true" box="[199,337,1964,1991]" pageId="11" pageNumber="12">Description</emphasis>
. Habitus as illustrated (
<figureCitation id="13372A74A868FFEF0E4F15E9FD499592" box="[610,680,1965,1991]" captionStart="FIGURE 8" captionStartId="23.[151,250,1965,1990]" captionTargetBox="[190,1388,466,1915]" captionTargetId="figure@23.[151,1435,453,1940]" captionTargetPageId="23" captionText="FIGURE 8. Hydrochus ducalis Knisch, habitus and male genitalia." figureDoi="http://doi.org/10.5281/zenodo.3587197" httpUri="https://zenodo.org/record/3587197/files/figure.png" pageId="11" pageNumber="12">Fig. 8</figureCitation>
). Size of imaged male (length/width, mm): body (length to elytral apices) body (length to elytral apices) 5.65 /1.90; head width 1.20; pronotum l/w 1.15 /1.25; PA 1.15; PB 1.10; elytra 3.60 /1.90. Large (ca.
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, Venezuelan specimens), moderately narrow. Dorsum grey with slight iridescence, elytra with black spots. Legs dark brown to black. Dorsal punctation small to moderately large, moderately dense, distinctly larger on elytra than pronotum. Elytra interstriae slightly wider than strial punctures. Interstria 9
<superScript id="7C799BB9A86FFFE809BD12A4FA7C92BB" attach="none" box="[1424,1437,224,238]" fontSize="6" pageId="12" pageNumber="13">th</superScript>
more convex than others, overhanging 10
<superScript id="7C799BB9A86FFFE80E451340FD949347" attach="left" box="[616,629,260,274]" fontSize="6" pageId="12" pageNumber="13">th</superScript>
interstria. Callus on 5
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interstria small and low, indistinct.
</paragraph>
<paragraph id="8BB336F1A86FFFE80CEA136DFF3A9332" blockId="12.[151,1437,152,791]" pageId="12" pageNumber="13">Pronotum slightly narrowed at base, sides arcuate or very weakly sinuate, smooth; depressions moderately small.</paragraph>
<paragraph id="8BB336F1A86FFFE80CEA1335FBCA93FA" blockId="12.[151,1437,152,791]" pageId="12" pageNumber="13">Elytral strial punctures ca. twice the size pronotal punctures, the spaces between them about half their diameter; apices rounded in dorsal view, in lateral view outer margin with slight angulation.</paragraph>
<paragraph id="8BB336F1A86FFFE80CEA13FDFB1E9386" blockId="12.[151,1437,152,791]" box="[199,1279,441,467]" pageId="12" pageNumber="13">Ventral characters: Mentum coarsely punctate, with median fovea. Submentum with two foveae.</paragraph>
<paragraph id="8BB336F1A86FFFE80CEA1399FE9A90D2" blockId="12.[151,1437,152,791]" pageId="12" pageNumber="13">
Male genitalia (
<figureCitation id="13372A74A86FFFE80D551399FE3C93A2" box="[376,477,477,503]" captionStart="FIGURES 6" captionStartId="19.[151,264,1522,1547]" captionTargetBox="[151,1435,181,1496]" captionTargetId="figure@19.[151,1435,181,1496]" captionTargetPageId="19" captionText="FIGURES 6A–C. (A) Male genitalia of H. ducalis Knisch, with parameres spread, showing shape of aedeagus; (B) Male genitalia of H. variabilis Knisch and (C) H. n. sp. from Peru, showing the articulation point of the paramere dorsal basal lobe with the aedeagus dorsal basal lobe." figureDoi="http://doi.org/10.5281/zenodo.3587193" httpUri="https://zenodo.org/record/3587193/files/figure.png" pageId="12" pageNumber="13">Figs. 6A</figureCitation>
,
<figureCitation id="13372A74A86FFFE80DC41399FE1693A2" box="[489,503,477,503]" captionStart="FIGURE 8" captionStartId="23.[151,250,1965,1990]" captionTargetBox="[190,1388,466,1915]" captionTargetId="figure@23.[151,1435,453,1940]" captionTargetPageId="23" captionText="FIGURE 8. Hydrochus ducalis Knisch, habitus and male genitalia." figureDoi="http://doi.org/10.5281/zenodo.3587197" httpUri="https://zenodo.org/record/3587197/files/figure.png" pageId="12" pageNumber="13">8</figureCitation>
) general characters: paramere tips slender and pointed, with medial margins parallel or nearly so; basal piece slightly longer than parameres; aedeagus sinuate in dorsal and ventral views, slender in distal 2/3, widened basally (
<figureCitation id="13372A74A86FFFE80DC31061FDA99015" box="[494,584,549,576]" captionStart="FIGURES 6" captionStartId="19.[151,264,1522,1547]" captionTargetBox="[151,1435,181,1496]" captionTargetId="figure@19.[151,1435,181,1496]" captionTargetPageId="19" captionText="FIGURES 6A–C. (A) Male genitalia of H. ducalis Knisch, with parameres spread, showing shape of aedeagus; (B) Male genitalia of H. variabilis Knisch and (C) H. n. sp. from Peru, showing the articulation point of the paramere dorsal basal lobe with the aedeagus dorsal basal lobe." figureDoi="http://doi.org/10.5281/zenodo.3587193" httpUri="https://zenodo.org/record/3587193/files/figure.png" pageId="12" pageNumber="13">Fig. 6A</figureCitation>
); in lateral view much wider, and produced in a prominent lobe (adm), what appears to be another lobe near the base is actually the dorsal margin of the paramere; lateral margins of aedeagus strongly sclerotized.
</paragraph>
<paragraph id="8BB336F1A86FFFE80CEA10D5FB6890FE" blockId="12.[151,1437,152,791]" box="[199,1161,657,683]" pageId="12" pageNumber="13">Dorsal surface: pdmm slightly sinuate; adbl very large, oval; adtl very near distal end.</paragraph>
<paragraph id="8BB336F1A86FFFE80CEA10F1FF0C90A6" blockId="12.[151,1437,152,791]" pageId="12" pageNumber="13">Ventral surface: pvmm more strongly sinuate than pdmm, widened part accommodating widened part of aedeagus.</paragraph>
</subSubSection>
<subSubSection id="C316657AA86FFFE80CEA10B9FBEE9142" box="[199,1039,765,791]" pageId="12" pageNumber="13" type="discussion">
<paragraph id="8BB336F1A86FFFE80CEA10B9FBEE9142" blockId="12.[151,1437,152,791]" box="[199,1039,765,791]" pageId="12" pageNumber="13">
<emphasis id="B978EAE3A86FFFE80CEA10B9FEDA9142" bold="true" box="[199,315,765,791]" pageId="12" pageNumber="13">Remarks.</emphasis>
The specimen imaged (
<figureCitation id="13372A74A86FFFE80E6A10B9FD689142" box="[583,649,765,791]" captionStart="FIGURE 8" captionStartId="23.[151,250,1965,1990]" captionTargetBox="[190,1388,466,1915]" captionTargetId="figure@23.[151,1435,453,1940]" captionTargetPageId="23" captionText="FIGURE 8. Hydrochus ducalis Knisch, habitus and male genitalia." figureDoi="http://doi.org/10.5281/zenodo.3587197" httpUri="https://zenodo.org/record/3587197/files/figure.png" pageId="12" pageNumber="13">Fig. 8</figureCitation>
) is from locality VZ10-0202-03A.
</paragraph>
</subSubSection>
</treatment>
</document>
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