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<document id="534875C82EC3A4F88EEB8AF5AA449F81" ID-CLB-Dataset="198089" ID-DOI="10.11646/zootaxa.5213.5.5" ID-GBIF-Dataset="4cb73371-a794-43fb-8bd6-d3e7561b4041" ID-ISSN="1175-5326" ID-Zenodo-Dep="7381712" ID-ZooBank="648F79BD-36F1-4B4D-978B-13903BCE5F0F" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1669810691679" checkinUser="plazi" docAuthor="Yu, Xin, Li, Linyan &amp; Gu, Haijun" docDate="2022" docId="03A587BAFFB98A71FF51821CFE3CF848" docLanguage="en" docName="zootaxa.5213.5.5.pdf" docOrigin="Zootaxa 5213 (5)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Ophiogomphus tibeticus Yu &amp; Li &amp; Gu 2022, sp. nov." docType="treatment" docVersion="9" lastPageNumber="576" masterDocId="FF9CFFC2FFBB8A76FFC6853EFA39FFB9" masterDocTitle="Ophiogomphus tibeticus sp. nov. from Sichuan, China (Anisoptera: Gomphidae)" masterLastPageNumber="577" masterPageNumber="569" pageNumber="571" updateTime="1699412604890" updateUser="ExternalLinkService" zenodo-license-document="CLOSED">
<mods:mods id="4AB2876274E4AC9FEAC853B6FC0E0F6F" xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo id="42913EED8849F14F9DC32202866AECE1">
<mods:title id="B576FEBFDD8BE7FE0E83B3142859CB50">Ophiogomphus tibeticus sp. nov. from Sichuan, China (Anisoptera: Gomphidae)</mods:title>
</mods:titleInfo>
<mods:name id="8779A6B7F2F7CCCB60F698457C103E36" type="personal">
<mods:role id="EA964382D04D85C6FA59B82997046B47">
<mods:roleTerm id="132B29F43E49C45D6B0163EED77CC485">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="085E0C115C8E7DA9C3BC8E27FD671AFD">Yu, Xin</mods:namePart>
<mods:affiliation id="D268EEA2BCC4ECFF76F55D901061CA34">College of Life Sciences, Chongqing Normal University, Chongqing, 401331, China</mods:affiliation>
</mods:name>
<mods:name id="C708277171F1976707F76B7B2120F38E" type="personal">
<mods:role id="BAA3B585937231F7F4FA7ADBC8B4A03A">
<mods:roleTerm id="6ACC55F08B15E0277ACA5A37644E276D">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="8D3FB11E2629FA910B43EFEB708865F7">Li, Linyan</mods:namePart>
<mods:nameIdentifier id="8C89DDA1C9A271CD91E92585134E2B78" type="ORCID">0000-0002-3030-8335</mods:nameIdentifier>
<mods:affiliation id="21E73EF3598EEC47EC5A8429D7EFC0C5">College of Life Sciences, Chongqing Normal University, Chongqing, 401331, China &amp; 1160010021 @ qq. com; https: // orcid. org / 0000 - 0002 - 3030 - 8335</mods:affiliation>
<mods:nameIdentifier id="AA1677AE5C88F13F4E5FD93CDCFA9242" type="email">1160010021@qq.com</mods:nameIdentifier>
</mods:name>
<mods:name id="491BC028A3DECE61A3D827CC37DDD83D" type="personal">
<mods:role id="AF40737C4994689964D3807C1E768E94">
<mods:roleTerm id="97D30E6B2583D40BBA7B16D97B0F2C30">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="E85A2CA2F33B626245B81067EB70521D">Gu, Haijun</mods:namePart>
<mods:affiliation id="AD4C4A3F4260D1D952A816CDCFF71950">Wildlife Resources Investigation and Protection Station of Sichuan Province, Chengdu, 610081, China</mods:affiliation>
</mods:name>
<mods:typeOfResource id="0F8330BBF3B06AA8C655ABEBFBF4BD58">text</mods:typeOfResource>
<mods:relatedItem id="C3F8FF8874EB23C49E459BCACCAB2194" type="host">
<mods:titleInfo id="6582B201263F5B176BC1AD9EE8BC15A8">
<mods:title id="9E95DD8054064899CF05E262ADD94D4B">Zootaxa</mods:title>
</mods:titleInfo>
<mods:part id="075223C5F7E4170098B024B15F04B6A6">
<mods:date id="225F7EEF049F77DBE20F4AF279BF39E3">2022</mods:date>
<mods:detail id="F1481DB858349C5C1F37B1F534E7F3A2" type="pubDate">
<mods:number id="2F72FFC09012B71FE80683911026AB69">2022-11-29</mods:number>
</mods:detail>
<mods:detail id="60E1E307E3778EB7F765F1F487379E5B" type="volume">
<mods:number id="88475CEAD84E33CFAA824D987061B5B5">5213</mods:number>
</mods:detail>
<mods:detail id="86352A491F1BA1191BCAE12549E6E7E8" type="issue">
<mods:number id="D958DBA522C4E672CFE0EBB2BE52D24F">5</mods:number>
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<mods:extent id="B610034BE15883E0CCDBE722F3E490D6" unit="page">
<mods:start id="31E9F4FF730FC45BDE7D8590CF71780C">569</mods:start>
<mods:end id="8DA6F7A13D283B88BC3F760B2F8E9D7D">577</mods:end>
</mods:extent>
</mods:part>
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<mods:classification id="7EF482BD2E6C04CEC0510E2248D70953">journal article</mods:classification>
<mods:identifier id="B8F6C3188B740573E0FC6AE6B89A0384" type="CLB-Dataset">198089</mods:identifier>
<mods:identifier id="D93A2A90C0EF3EA3A7F47971974D2E23" type="DOI">10.11646/zootaxa.5213.5.5</mods:identifier>
<mods:identifier id="4CCB40F36A942F9AF58F9C21D0860599" type="GBIF-Dataset">4cb73371-a794-43fb-8bd6-d3e7561b4041</mods:identifier>
<mods:identifier id="B77344A03D7C634C77FD4F014A62EEC7" type="ISSN">1175-5326</mods:identifier>
<mods:identifier id="E55F6B05CD058839E0DC4323F2923267" type="Zenodo-Dep">7381712</mods:identifier>
<mods:identifier id="55ADBA6C3187DC5E0DD3E5D39B802438" type="ZooBank">648F79BD-36F1-4B4D-978B-13903BCE5F0F</mods:identifier>
</mods:mods>
<treatment id="03A587BAFFB98A71FF51821CFE3CF848" ID-DOI="http://doi.org/10.5281/zenodo.7383633" ID-GBIF-Taxon="205238263" ID-Zenodo-Dep="7383633" LSID="urn:lsid:plazi:treatment:03A587BAFFB98A71FF51821CFE3CF848" httpUri="http://treatment.plazi.org/id/03A587BAFFB98A71FF51821CFE3CF848" lastPageId="7" lastPageNumber="576" pageId="2" pageNumber="571">
<subSubSection id="C3166527FFB98A74FF51821CF89BF884" box="[151,674,1826,1853]" pageId="2" pageNumber="571" type="nomenclature">
<paragraph id="8BB336ACFFB98A74FF51821CF89BF884" blockId="2.[151,674,1826,1889]" box="[151,674,1826,1853]" pageId="2" pageNumber="571">
<heading id="D0FB81C0FFB98A74FF51821CF89BF884" bold="true" box="[151,674,1826,1853]" fontSize="11" level="1" pageId="2" pageNumber="571" reason="1">
<emphasis id="B978EABEFFB98A74FF51821CF89BF884" bold="true" box="[151,674,1826,1853]" pageId="2" pageNumber="571">
<taxonomicName id="4C0C4D2FFFB98A74FF51821CF801F885" ID-CoL="9X7GQ" authority="Yu &amp; Gu" authorityName="Yu &amp; Li &amp; Gu" authorityYear="2022" box="[151,568,1826,1853]" class="Insecta" family="Gomphidae" genus="Ophiogomphus" kingdom="Animalia" order="Odonata" pageId="2" pageNumber="571" phylum="Arthropoda" rank="species" species="tibeticus" status="sp. nov.">
<emphasis id="B978EABEFFB98A74FF51821CFB84F884" bold="true" box="[151,445,1826,1853]" italics="true" pageId="2" pageNumber="571">Ophiogomphus tibeticus</emphasis>
Yu &amp; Gu
</taxonomicName>
,
<taxonomicNameLabel id="A24B57C5FFB98A74FD82821DF89BF884" box="[580,674,1827,1853]" pageId="2" pageNumber="571" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C3166527FFB98A74FF518278FB19F8D8" box="[151,288,1862,1889]" pageId="2" pageNumber="571" type="description">
<paragraph id="8BB336ACFFB98A74FF518278FB19F8D8" blockId="2.[151,674,1826,1889]" box="[151,288,1862,1889]" pageId="2" pageNumber="571">
(
<figureCitation id="13372A29FFB98A74FF598278FB21F8D8" box="[159,280,1862,1889]" captionStart-0="FIGURE 2" captionStart-1="FIGURE 3" captionStart-2="FIGURE 4" captionStart-3="FIGURE 5" captionStart-4="FIGURE 6" captionStart-5="FIGURE 7" captionStart-6="FIGURE 8" captionStart-7="FIGURE 9" captionStart-8="FIGURE 10" captionStartId-0="3.[152,255,1343,1367]" captionStartId-1="3.[152,255,1995,2019]" captionStartId-2="4.[152,255,1416,1440]" captionStartId-3="5.[152,255,1111,1135]" captionStartId-4="5.[152,255,1893,1917]" captionStartId-5="6.[152,255,864,888]" captionStartId-6="6.[152,255,2004,2028]" captionStartId-7="7.[152,255,1172,1196]" captionStartId-8="7.[152,255,1792,1816]" captionTargetBox-2="[151,1402,541,1378]" captionTargetBox-3="[151,1364,181,1040]" captionTargetBox-4="[151,1436,1247,1839]" captionTargetBox-7="[243,1343,210,1132]" captionTargetBox-8="[237,1351,1221,1769]" captionTargetId-0="figure-95@3.[436,1150,380,1319]" captionTargetId-1="figure-113@3.[439,1148,1392,1971]" captionTargetId-2="figure-197@4.[151,1436,525,1393]" captionTargetId-3="figure-17@5.[151,1436,181,1087]" captionTargetId-4="figure-33@5.[151,1436,1160,1870]" captionTargetId-5="figure-18@6.[305,1283,181,840]" captionTargetId-6="figure-32@6.[346,1240,912,1980]" captionTargetId-7="figure-17@7.[221,1367,181,1149]" captionTargetId-8="figure-39@7.[237,1351,1221,1769]" captionTargetPageId-0="3" captionTargetPageId-1="3" captionTargetPageId-2="4" captionTargetPageId-3="5" captionTargetPageId-4="5" captionTargetPageId-5="6" captionTargetPageId-6="6" captionTargetPageId-7="7" captionTargetPageId-8="7" captionText-0="FIGURE 2. Living photos of the holotype of Ophiogomphus tibeticus." captionText-1="FIGURE 3. Face of the holotype." captionText-2="FIGURE 4. Habitus photos of the holotype." captionText-3="FIGURE 5. Caudal appendages of the holotype, dorsal view." captionText-4="FIGURE 6. Caudal appendages of the holotype, lateral view." captionText-5="FIGURE 7. Anterior hamule of the holotype." captionText-6="FIGURE 8. Posterior hamule of the holotype, removed from the body." captionText-7="FIGURE 9. Vesica spermalis of the holotype, lateral view, blue arrow indicating the hook-like projection." captionText-8="FIGURE 10. Vesica spermalis of the holotype, ventral view." figureDoi-0="http://doi.org/10.5281/zenodo.7386835" figureDoi-1="http://doi.org/10.5281/zenodo.7386837" figureDoi-2="http://doi.org/10.5281/zenodo.7381716" figureDoi-3="http://doi.org/10.5281/zenodo.7381718" figureDoi-4="http://doi.org/10.5281/zenodo.7381720" figureDoi-5="http://doi.org/10.5281/zenodo.7386841" figureDoi-6="http://doi.org/10.5281/zenodo.7386843" figureDoi-7="http://doi.org/10.5281/zenodo.7381722" figureDoi-8="http://doi.org/10.5281/zenodo.7381724" httpUri-0="https://zenodo.org/record/7386835/files/figure.png" httpUri-1="https://zenodo.org/record/7386837/files/figure.png" httpUri-2="https://zenodo.org/record/7381716/files/figure.png" httpUri-3="https://zenodo.org/record/7381718/files/figure.png" httpUri-4="https://zenodo.org/record/7381720/files/figure.png" httpUri-5="https://zenodo.org/record/7386841/files/figure.png" httpUri-6="https://zenodo.org/record/7386843/files/figure.png" httpUri-7="https://zenodo.org/record/7381722/files/figure.png" httpUri-8="https://zenodo.org/record/7381724/files/figure.png" pageId="2" pageNumber="571">Figs. 2–10</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="C3166527FFB98A74FF5182B1FB2DF849" pageId="2" pageNumber="571" type="materials_examined">
<paragraph id="8BB336ACFFB98A74FF5182B1FB2DF849" blockId="2.[151,1436,1934,2032]" pageId="2" pageNumber="571">
<materialsCitation id="3B643CF1FFB98A74FF5182B1FB45F875" ID-GBIF-Occurrence="3978607302" collectingDate="2019-06-25" collectorName="Xiu Yong" country="China" county="Hongyuan County" elevation="3463" latitude="32.722786" location="Hongyuan County" longLatPrecision="1" longitude="102.39523" pageId="2" pageNumber="571" specimenCount="1" specimenCount-male="1" stateProvince="Sichuan" typeStatus="holotype">
<emphasis id="B978EABEFFB98A74FF5182B1FB34F810" bold="true" box="[151,269,1935,1961]" pageId="2" pageNumber="571">
<typeStatus id="54B7880EFFB98A74FF5182B1FB31F810" box="[151,264,1935,1961]" pageId="2" pageNumber="571" type="holotype">Holotype</typeStatus>
:
</emphasis>
<specimenCount id="9D0AFD25FFB98A74FED382B1FB15F810" box="[277,300,1935,1961]" pageId="2" pageNumber="571" type="male">♂</specimenCount>
,
<collectingCountry id="F31B763CFFB98A74FEFF82B0FB46F811" box="[313,383,1934,1960]" name="China" pageId="2" pageNumber="571">China</collectingCountry>
,
<collectingRegion id="49C8F84EFFB98A74FE4D82B0F86AF811" box="[395,595,1934,1960]" country="China" name="Sichuan" pageId="2" pageNumber="571">Sichuan Province</collectingRegion>
,
<collectingCounty id="62D24E20FFB98A74FD9982B1F909F810" box="[607,816,1935,1961]" pageId="2" pageNumber="571">Hongyuan County</collectingCounty>
, (
<geoCoordinate id="EE38506BFFB98A74FC8582B0F9E2F810" box="[835,987,1934,1961]" degrees="32.722786" direction="north" orientation="latitude" pageId="2" pageNumber="571" precision="1" value="32.722786">32.722786 N</geoCoordinate>
,
<geoCoordinate id="EE38506BFFB98A74FC2182B0FEBFF810" box="[999,1158,1934,1961]" degrees="102.395236" direction="east" orientation="longitude" pageId="2" pageNumber="571" precision="1" value="102.39523">102.395236 E</geoCoordinate>
,
<elevation id="0021D19FFFB98A74FB5582B0FF14F811" box="[1171,1325,1934,1961]" metricMagnitude="3" metricUnit="m" metricValue="3.463" pageId="2" pageNumber="571" unit="m" value="3463.0">
<quantity id="4CF49B49FFB98A74FB5582B0FED0F810" box="[1171,1257,1934,1961]" metricMagnitude="3" metricUnit="m" metricValue="3.463" pageId="2" pageNumber="571" unit="m" value="3463.0">3463 m</quantity>
a.s. l.
</elevation>
),
<date id="FFB2106CFFB98A74FA8382B0FAEBF874" pageId="2" pageNumber="571" value="2019-06-25">
<collectingDate id="EFF6E984FFB98A74FA8382B0FAEBF874" pageId="2" pageNumber="571" value="2019-06-25">25 June 2019</collectingDate>
</date>
,
<collectorName id="26F9537AFFB98A74FF18828CFB73F874" box="[222,330,1970,1997]" pageId="2" pageNumber="571">Xiu Yong</collectorName>
leg.
</materialsCitation>
<materialsCitation id="3B643CF1FFB98A74FE42828CFB29F849" ID-GBIF-Occurrence="3978607301" country="China" location="College of Life Sciences" municipality="Normal University" pageId="2" pageNumber="571" specimenCount="1" stateProvince="Chongqing" typeStatus="holotype">
<typeStatus id="54B7880EFFB98A74FE42828CFBD5F875" box="[388,492,1970,1996]" pageId="2" pageNumber="571" type="holotype">Holotype</typeStatus>
deposited in the
<location id="8ED36077FFB98A74FD69828CF9F0F875" LSID="urn:lsid:plazi:treatment:03A587BAFFB98A71FF51821CFE3CF848:8ED36077FFB98A74FD69828CF9F0F875" box="[687,969,1970,1996]" country="China" municipality="Normal University" name="College of Life Sciences" pageId="2" pageNumber="571" stateProvince="Chongqing">College of Life Sciences</location>
,
<collectingRegion id="49C8F84EFFB98A74FC13828CFE68F875" box="[981,1105,1970,1996]" country="China" name="Chongqing" pageId="2" pageNumber="571">Chongqing</collectingRegion>
<collectingMunicipality id="6BD7ACD6FFB98A74FB9F828CFF17F875" box="[1113,1326,1970,1996]" pageId="2" pageNumber="571">Normal University</collectingMunicipality>
,
<collectingRegion id="49C8F84EFFB98A74FAFC828CFA84F849" country="China" name="Chongqing" pageId="2" pageNumber="571">Chongqing</collectingRegion>
,
<collectingCountry id="F31B763CFFB98A74FF0F82E8FB29F849" box="[201,272,2006,2032]" name="China" pageId="2" pageNumber="571">China</collectingCountry>
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="C3166527FFB88A75FF0185A8F9D4FF08" box="[199,1005,150,177]" pageId="3" pageNumber="572" type="etymology">
<paragraph id="8BB336ACFFB88A75FF0185A8F9D4FF08" blockId="3.[151,1436,150,357]" box="[199,1005,150,177]" pageId="3" pageNumber="572">
<emphasis id="B978EABEFFB88A75FF0185A8FB71FF09" bold="true" box="[199,328,150,176]" pageId="3" pageNumber="572">Etymology</emphasis>
. The name indicates that this species occurs in Tibetan area.
</paragraph>
</subSubSection>
<subSubSection id="C3166527FFB88A72FF018584FEB1F804" lastPageId="4" lastPageNumber="573" pageId="3" pageNumber="572" type="description">
<paragraph id="8BB336ACFFB88A75FF018584FE08FF6D" blockId="3.[151,1436,150,357]" box="[199,1073,186,212]" pageId="3" pageNumber="572">
<emphasis id="B978EABEFFB88A75FF018584FBDCFF6D" bold="true" box="[199,485,186,212]" pageId="3" pageNumber="572">
Description of
<typeStatus id="54B7880EFFB88A75FEB18584FBD9FF6D" box="[375,480,186,212]" pageId="3" pageNumber="572" type="holotype">holotype</typeStatus>
.
</emphasis>
Body greenish-yellow with black markings (
<figureCitation id="13372A29FFB88A75FC198584FE1DFF6D" box="[991,1060,186,212]" captionStart="FIGURE 2" captionStartId="3.[152,255,1343,1367]" captionTargetId="figure-95@3.[436,1150,380,1319]" captionTargetPageId="3" captionText="FIGURE 2. Living photos of the holotype of Ophiogomphus tibeticus." figureDoi="http://doi.org/10.5281/zenodo.7386835" httpUri="https://zenodo.org/record/7386835/files/figure.png" pageId="3" pageNumber="572">Fig. 2</figureCitation>
).
</paragraph>
<paragraph id="8BB336ACFFB88A75FF0185E0FB6CFEDD" blockId="3.[151,1436,150,357]" pageId="3" pageNumber="572">
<emphasis id="B978EABEFFB88A75FF0185E0FB3DFF41" box="[199,260,222,248]" italics="true" pageId="3" pageNumber="572">Head</emphasis>
. Eyes green in life (
<figureCitation id="13372A29FFB88A75FE3485E0F803FF41" box="[498,570,222,248]" captionStart="FIGURE 2" captionStartId="3.[152,255,1343,1367]" captionTargetId="figure-95@3.[436,1150,380,1319]" captionTargetPageId="3" captionText="FIGURE 2. Living photos of the holotype of Ophiogomphus tibeticus." figureDoi="http://doi.org/10.5281/zenodo.7386835" httpUri="https://zenodo.org/record/7386835/files/figure.png" pageId="3" pageNumber="572">Fig. 2</figureCitation>
). Labium yellow. Face (
<figureCitation id="13372A29FFB88A75FC9485E0F9A0FF41" box="[850,921,222,248]" captionStart="FIGURE 3" captionStartId="3.[152,255,1995,2019]" captionTargetId="figure-113@3.[439,1148,1392,1971]" captionTargetPageId="3" captionText="FIGURE 3. Face of the holotype." figureDoi="http://doi.org/10.5281/zenodo.7386837" httpUri="https://zenodo.org/record/7386837/files/figure.png" pageId="3" pageNumber="572">Fig. 3</figureCitation>
): mandible bases, labrum, anteclypeus, postclypeus and frons yellow with narrow black stripes across fronto-clypeal suture, along ventolateral margins of postcyleus and base and margin of labrum. Vertex black, antennae black. Top of head black. Occipital plate yellow, even and simple.
</paragraph>
<caption id="DF736624FFB88A75FF5E8001F952FAEE" ID-DOI="http://doi.org/10.5281/zenodo.7386835" ID-Zenodo-Dep="7386835" box="[152,875,1343,1367]" httpUri="https://zenodo.org/record/7386835/files/figure.png" pageId="3" pageNumber="572" startId="3.[152,255,1343,1367]" targetBox="[436,1150,380,1319]" targetPageId="3">
<paragraph id="8BB336ACFFB88A75FF5E8001F952FAEE" blockId="3.[152,875,1343,1367]" box="[152,875,1343,1367]" pageId="3" pageNumber="572">
<emphasis id="B978EABEFFB88A75FF5E8001FB21FAEE" bold="true" box="[152,280,1343,1367]" pageId="3" pageNumber="572">FIGURE 2.</emphasis>
Living photos of the holotype of
<taxonomicName id="4C0C4D2FFFB88A75FDA88001F95CFAEE" authorityName="Yu &amp; Li &amp; Gu" authorityYear="2022" box="[622,869,1343,1367]" class="Insecta" family="Gomphidae" genus="Ophiogomphus" kingdom="Animalia" order="Odonata" pageId="3" pageNumber="572" phylum="Arthropoda" rank="species" species="tibeticus">
<emphasis id="B978EABEFFB88A75FDA88001F95CFAEE" box="[622,869,1343,1367]" italics="true" pageId="3" pageNumber="572">Ophiogomphus tibeticus</emphasis>
</taxonomicName>
.
</paragraph>
</caption>
<caption id="DF736624FFB88A75FF5E82F5FBCFF85A" ID-DOI="http://doi.org/10.5281/zenodo.7386837" ID-Zenodo-Dep="7386837" box="[152,502,1995,2019]" httpUri="https://zenodo.org/record/7386837/files/figure.png" pageId="3" pageNumber="572" startId="3.[152,255,1995,2019]" targetBox="[439,1148,1392,1971]" targetPageId="3">
<paragraph id="8BB336ACFFB88A75FF5E82F5FBCFF85A" blockId="3.[152,502,1995,2019]" box="[152,502,1995,2019]" pageId="3" pageNumber="572">
<emphasis id="B978EABEFFB88A75FF5E82F5FB21F85A" bold="true" box="[152,280,1995,2019]" pageId="3" pageNumber="572">FIGURE 3.</emphasis>
Face of the holotype.
</paragraph>
</caption>
<paragraph id="8BB336ACFFBF8A72FF0185A9FB41FEF8" blockId="4.[151,1437,151,501]" pageId="4" pageNumber="573">
<emphasis id="B978EABEFFBF8A72FF0185A9FB21FF08" box="[199,280,151,177]" italics="true" pageId="4" pageNumber="573">Thorax</emphasis>
greenish yellow with black pattern (
<figureCitation id="13372A29FFBF8A72FD7585A9F8C0FF08" box="[691,761,151,177]" captionStart="FIGURE 2" captionStartId="3.[152,255,1343,1367]" captionTargetId="figure-95@3.[436,1150,380,1319]" captionTargetPageId="3" captionText="FIGURE 2. Living photos of the holotype of Ophiogomphus tibeticus." figureDoi="http://doi.org/10.5281/zenodo.7386835" httpUri="https://zenodo.org/record/7386835/files/figure.png" pageId="4" pageNumber="573">Fig. 2</figureCitation>
). Anterior lobe of prothorax black with a broad yellow margin, middle lobe black with a pair of twin adjacent yellow spots dorsally and a pair of larger yellow spots laterally, posterior lobe black; synthorax greenish yellow with narrow black stripes along dorsal carina, collar, antealar ridge, humeral suture, and metapleural suture. The stripe on humeral suture relatively broad with a short, narrow yellow stripe in the middle.
</paragraph>
<paragraph id="8BB336ACFFBF8A72FF018475FF63FE4C" blockId="4.[151,1437,151,501]" pageId="4" pageNumber="573">
<emphasis id="B978EABEFFBF8A72FF018475FB32FEDD" box="[199,267,331,356]" italics="true" pageId="4" pageNumber="573">Wings</emphasis>
(
<figureCitation id="13372A29FFBF8A72FEDC8475FB64FEDC" box="[282,349,331,357]" captionStart="FIGURE 4" captionStartId="4.[152,255,1416,1440]" captionTargetBox="[151,1402,541,1378]" captionTargetId="figure-197@4.[151,1436,525,1393]" captionTargetPageId="4" captionText="FIGURE 4. Habitus photos of the holotype." figureDoi="http://doi.org/10.5281/zenodo.7381716" httpUri="https://zenodo.org/record/7381716/files/figure.png" pageId="4" pageNumber="573">Fig. 4</figureCitation>
) hyaline, venation blackish brown. Arculus at 2nd antenodal. Antenodals 12 (1st and 5th primary) in both FW; 9 (1st and 5th primary) in both HW; postnodals 11 (right)–12 (left) in FW, 12 (left)–11 (right) in HW. One cubito-anal vein in all wings except left FW (two). Triangles not crossed. Anal loop three-celled in left HW, two in right HW. Anal triangle 3-celled in left wing,
<quantity id="4CF49B49FFBF8A72FD6D8489F8EFFE68" box="[683,726,439,465]" metricMagnitude="-1" metricUnit="m" metricValue="1.016" pageId="4" pageNumber="573" unit="in" value="4.0">4 in</quantity>
right wing. Pt pale brown with black bordering veins (considering this may be not a fully matured individual, the color could be darker later), overlying 5 cells in all wings.
</paragraph>
<caption id="DF736624FFBF8A72FF5E80B6F867FA19" ID-DOI="http://doi.org/10.5281/zenodo.7381716" ID-Zenodo-Dep="7381716" box="[152,606,1416,1440]" httpUri="https://zenodo.org/record/7381716/files/figure.png" pageId="4" pageNumber="573" startId="4.[152,255,1416,1440]" targetBox="[151,1402,541,1378]" targetPageId="4">
<paragraph id="8BB336ACFFBF8A72FF5E80B6F867FA19" blockId="4.[152,606,1416,1440]" box="[152,606,1416,1440]" pageId="4" pageNumber="573">
<emphasis id="B978EABEFFBF8A72FF5E80B6FB21FA19" bold="true" box="[152,280,1416,1440]" pageId="4" pageNumber="573">FIGURE 4.</emphasis>
Habitus photos of the holotype.
</paragraph>
</caption>
<paragraph id="8BB336ACFFBF8A72FF0180F0FE8CF9ED" blockId="4.[151,1437,1486,1981]" pageId="4" pageNumber="573">
<emphasis id="B978EABEFFBF8A72FF0180F0FB08FA51" box="[199,305,1486,1512]" italics="true" pageId="4" pageNumber="573">Abdomen</emphasis>
black with light-yellow markings (
<figureCitation id="13372A29FFBF8A72FD0580F0F92AFA50" box="[707,787,1486,1513]" captionStart="FIGURE 2" captionStartId="3.[152,255,1343,1367]" captionTargetId="figure-95@3.[436,1150,380,1319]" captionTargetPageId="3" captionText="FIGURE 2. Living photos of the holotype of Ophiogomphus tibeticus." figureDoi="http://doi.org/10.5281/zenodo.7386835" httpUri="https://zenodo.org/record/7386835/files/figure.png" pageId="4" pageNumber="573">Figs. 2</figureCitation>
,
<figureCitation id="13372A29FFBF8A72FCE780F1F90BFA50" box="[801,818,1487,1513]" captionStart="FIGURE 4" captionStartId="4.[152,255,1416,1440]" captionTargetBox="[151,1402,541,1378]" captionTargetId="figure-197@4.[151,1436,525,1393]" captionTargetPageId="4" captionText="FIGURE 4. Habitus photos of the holotype." figureDoi="http://doi.org/10.5281/zenodo.7381716" httpUri="https://zenodo.org/record/7381716/files/figure.png" pageId="4" pageNumber="573">4</figureCitation>
). S1 almost wholly light yellow. S2 with a huge triangular dorsal light-yellow spot; auricles light yellow; S2 light yellow laterally with a narrow crossed black marking in the middle. S3–9 each with a large subtriangular light-yellow middorsal spot and a pair of irregular light-yellow spots laterally. S10 with a pentagonal light-yellow middorsal spot, light yellow spots on sides.
</paragraph>
<paragraph id="8BB336ACFFBF8A72FF018360F83BF95D" blockId="4.[151,1437,1486,1981]" pageId="4" pageNumber="573">
<emphasis id="B978EABEFFBF8A72FF018360FB9EF9C1" box="[199,423,1630,1656]" italics="true" pageId="4" pageNumber="573">Caudal appendages</emphasis>
relatively simple and robust (
<figureCitation id="13372A29FFBF8A72FD338360F95BF9C0" box="[757,866,1630,1657]" captionStart-0="FIGURE 5" captionStart-1="FIGURE 6" captionStartId-0="5.[152,255,1111,1135]" captionStartId-1="5.[152,255,1893,1917]" captionTargetBox-0="[151,1364,181,1040]" captionTargetBox-1="[151,1436,1247,1839]" captionTargetId-0="figure-17@5.[151,1436,181,1087]" captionTargetId-1="figure-33@5.[151,1436,1160,1870]" captionTargetPageId-0="5" captionTargetPageId-1="5" captionText-0="FIGURE 5. Caudal appendages of the holotype, dorsal view." captionText-1="FIGURE 6. Caudal appendages of the holotype, lateral view." figureDoi-0="http://doi.org/10.5281/zenodo.7381718" figureDoi-1="http://doi.org/10.5281/zenodo.7381720" httpUri-0="https://zenodo.org/record/7381718/files/figure.png" httpUri-1="https://zenodo.org/record/7381720/files/figure.png" pageId="4" pageNumber="573">Figs. 5–6</figureCitation>
). Cerci dark brown, paler laterally, forcipate, twice as long as S10, almost touching to each other, in lateral view decurved distally. Epiproct dark brown, widely divaricate, shorter than cerci, in lateral view bending upward with a distinct swelling at basal third followed distally by a slightly expanded apex. (
<figureCitation id="13372A29FFBF8A72FE7683F4FBCDF95D" box="[432,500,1738,1764]" captionStart="FIGURE 6" captionStartId="5.[152,255,1893,1917]" captionTargetBox="[151,1436,1247,1839]" captionTargetId="figure-33@5.[151,1436,1160,1870]" captionTargetPageId="5" captionText="FIGURE 6. Caudal appendages of the holotype, lateral view." figureDoi="http://doi.org/10.5281/zenodo.7381720" httpUri="https://zenodo.org/record/7381720/files/figure.png" pageId="4" pageNumber="573">Fig. 6</figureCitation>
).
</paragraph>
<paragraph id="8BB336ACFFBF8A72FF0183D0F958F820" blockId="4.[151,1437,1486,1981]" pageId="4" pageNumber="573">
<emphasis id="B978EABEFFBF8A72FF0183D0FB90F8B1" box="[199,425,1774,1800]" italics="true" pageId="4" pageNumber="573">Secondary genitalia</emphasis>
(
<figureCitation id="13372A29FFBF8A72FE7E83D0F80CF8B0" box="[440,565,1774,1801]" captionStart-0="FIGURE 7" captionStart-1="FIGURE 8" captionStart-2="FIGURE 9" captionStart-3="FIGURE 10" captionStartId-0="6.[152,255,864,888]" captionStartId-1="6.[152,255,2004,2028]" captionStartId-2="7.[152,255,1172,1196]" captionStartId-3="7.[152,255,1792,1816]" captionTargetBox-2="[243,1343,210,1132]" captionTargetBox-3="[237,1351,1221,1769]" captionTargetId-0="figure-18@6.[305,1283,181,840]" captionTargetId-1="figure-32@6.[346,1240,912,1980]" captionTargetId-2="figure-17@7.[221,1367,181,1149]" captionTargetId-3="figure-39@7.[237,1351,1221,1769]" captionTargetPageId-0="6" captionTargetPageId-1="6" captionTargetPageId-2="7" captionTargetPageId-3="7" captionText-0="FIGURE 7. Anterior hamule of the holotype." captionText-1="FIGURE 8. Posterior hamule of the holotype, removed from the body." captionText-2="FIGURE 9. Vesica spermalis of the holotype, lateral view, blue arrow indicating the hook-like projection." captionText-3="FIGURE 10. Vesica spermalis of the holotype, ventral view." figureDoi-0="http://doi.org/10.5281/zenodo.7386841" figureDoi-1="http://doi.org/10.5281/zenodo.7386843" figureDoi-2="http://doi.org/10.5281/zenodo.7381722" figureDoi-3="http://doi.org/10.5281/zenodo.7381724" httpUri-0="https://zenodo.org/record/7386841/files/figure.png" httpUri-1="https://zenodo.org/record/7386843/files/figure.png" httpUri-2="https://zenodo.org/record/7381722/files/figure.png" httpUri-3="https://zenodo.org/record/7381724/files/figure.png" pageId="4" pageNumber="573">Figs. 7–10</figureCitation>
). Blackish brown. Anterior hamule with apex strongly hooked in lateral view (
<figureCitation id="13372A29FFBF8A72FF59822CFADBF894" box="[159,226,1810,1837]" captionStart="FIGURE 7" captionStartId="6.[152,255,864,888]" captionTargetId="figure-18@6.[305,1283,181,840]" captionTargetPageId="6" captionText="FIGURE 7. Anterior hamule of the holotype." figureDoi="http://doi.org/10.5281/zenodo.7386841" httpUri="https://zenodo.org/record/7386841/files/figure.png" pageId="4" pageNumber="573">Fig. 7</figureCitation>
) with long, robust hairs medially; posterior hamule simple and robust, conical, with long hairs, and terminating in a strong spine (
<figureCitation id="13372A29FFBF8A72FE428208FBF1F8E8" box="[388,456,1846,1873]" captionStart="FIGURE 8" captionStartId="6.[152,255,2004,2028]" captionTargetId="figure-32@6.[346,1240,912,1980]" captionTargetPageId="6" captionText="FIGURE 8. Posterior hamule of the holotype, removed from the body." figureDoi="http://doi.org/10.5281/zenodo.7386843" httpUri="https://zenodo.org/record/7386843/files/figure.png" pageId="4" pageNumber="573">Fig. 8</figureCitation>
). Vesica spermalis with a large basal segment, triangular in lateral view; second segment broad, recurved, gently expanding apically; third segment robust, prepuse linear, spine-like; apical segment with a pair of flagella and a hook-like projection apically (
<figureCitation id="13372A29FFBF8A72FD1F8240F96DF820" box="[729,852,1918,1945]" captionStart-0="FIGURE 9" captionStart-1="FIGURE 10" captionStartId-0="7.[152,255,1172,1196]" captionStartId-1="7.[152,255,1792,1816]" captionTargetBox-0="[243,1343,210,1132]" captionTargetBox-1="[237,1351,1221,1769]" captionTargetId-0="figure-17@7.[221,1367,181,1149]" captionTargetId-1="figure-39@7.[237,1351,1221,1769]" captionTargetPageId-0="7" captionTargetPageId-1="7" captionText-0="FIGURE 9. Vesica spermalis of the holotype, lateral view, blue arrow indicating the hook-like projection." captionText-1="FIGURE 10. Vesica spermalis of the holotype, ventral view." figureDoi-0="http://doi.org/10.5281/zenodo.7381722" figureDoi-1="http://doi.org/10.5281/zenodo.7381724" httpUri-0="https://zenodo.org/record/7381722/files/figure.png" httpUri-1="https://zenodo.org/record/7381724/files/figure.png" pageId="4" pageNumber="573">Figs. 9–10</figureCitation>
).
</paragraph>
<paragraph id="8BB336ACFFBF8A72FF01829DFEB1F804" blockId="4.[151,1437,1486,1981]" box="[199,1160,1954,1981]" pageId="4" pageNumber="573">
<emphasis id="B978EABEFFBF8A72FF01829DFB50F805" box="[199,361,1955,1980]" italics="true" pageId="4" pageNumber="573">Measurements</emphasis>
(mm): Total length 50.0; abdomen + appendages 36.0; hind wing 33.0.
</paragraph>
</subSubSection>
<caption id="DF736624FFBE8A73FF5E8169F933FBD6" ID-DOI="http://doi.org/10.5281/zenodo.7381718" ID-Zenodo-Dep="7381718" box="[152,778,1111,1135]" httpUri="https://zenodo.org/record/7381718/files/figure.png" pageId="5" pageNumber="574" startId="5.[152,255,1111,1135]" targetBox="[151,1364,181,1040]" targetPageId="5">
<paragraph id="8BB336ACFFBE8A73FF5E8169F933FBD6" blockId="5.[152,778,1111,1135]" box="[152,778,1111,1135]" pageId="5" pageNumber="574">
<emphasis id="B978EABEFFBE8A73FF5E8169FB21FBD6" bold="true" box="[152,280,1111,1135]" pageId="5" pageNumber="574">FIGURE 5.</emphasis>
Caudal appendages of the holotype, dorsal view.
</paragraph>
</caption>
<caption id="DF736624FFBE8A73FF5E825BF935F8C4" ID-DOI="http://doi.org/10.5281/zenodo.7381720" ID-Zenodo-Dep="7381720" box="[152,780,1893,1917]" httpUri="https://zenodo.org/record/7381720/files/figure.png" pageId="5" pageNumber="574" startId="5.[152,255,1893,1917]" targetBox="[151,1436,1247,1839]" targetPageId="5">
<paragraph id="8BB336ACFFBE8A73FF5E825BF935F8C4" blockId="5.[152,780,1893,1917]" box="[152,780,1893,1917]" pageId="5" pageNumber="574">
<emphasis id="B978EABEFFBE8A73FF5E825BFB21F8C4" bold="true" box="[152,280,1893,1917]" pageId="5" pageNumber="574">FIGURE 6.</emphasis>
Caudal appendages of the holotype, lateral view.
</paragraph>
</caption>
<caption id="DF736624FFBD8A70FF5E865EF854FCC1" ID-DOI="http://doi.org/10.5281/zenodo.7386841" ID-Zenodo-Dep="7386841" box="[152,621,864,888]" httpUri="https://zenodo.org/record/7386841/files/figure.png" pageId="6" pageNumber="575" startId="6.[152,255,864,888]" targetBox="[305,1283,181,840]" targetPageId="6">
<paragraph id="8BB336ACFFBD8A70FF5E865EF854FCC1" blockId="6.[152,621,864,888]" box="[152,621,864,888]" pageId="6" pageNumber="575">
<emphasis id="B978EABEFFBD8A70FF5E865EFB21FCC1" bold="true" box="[152,280,864,888]" pageId="6" pageNumber="575">FIGURE 7.</emphasis>
Anterior hamule of the holotype.
</paragraph>
</caption>
<caption id="DF736624FFBD8A70FF5E82EAF955F855" ID-DOI="http://doi.org/10.5281/zenodo.7386843" ID-Zenodo-Dep="7386843" box="[152,876,2004,2028]" httpUri="https://zenodo.org/record/7386843/files/figure.png" pageId="6" pageNumber="575" startId="6.[152,255,2004,2028]" targetBox="[346,1203,950,1961]" targetPageId="6">
<paragraph id="8BB336ACFFBD8A70FF5E82EAF955F855" blockId="6.[152,876,2004,2028]" box="[152,876,2004,2028]" pageId="6" pageNumber="575">
<emphasis id="B978EABEFFBD8A70FF5E82EAFB21F855" bold="true" box="[152,280,2004,2028]" pageId="6" pageNumber="575">FIGURE 8.</emphasis>
Posterior hamule of the holotype, removed from the body.
</paragraph>
</caption>
<caption id="DF736624FFBC8A71FF5E81AAFEE9FB14" ID-DOI="http://doi.org/10.5281/zenodo.7381722" ID-Zenodo-Dep="7381722" box="[152,1232,1172,1197]" httpUri="https://zenodo.org/record/7381722/files/figure.png" pageId="7" pageNumber="576" startId="7.[152,255,1172,1196]" targetBox="[243,1343,210,1132]" targetPageId="7">
<paragraph id="8BB336ACFFBC8A71FF5E81AAFEE9FB14" blockId="7.[152,1232,1172,1197]" box="[152,1232,1172,1197]" pageId="7" pageNumber="576">
<emphasis id="B978EABEFFBC8A71FF5E81AAFB21FB14" bold="true" box="[152,280,1172,1197]" pageId="7" pageNumber="576">FIGURE 9.</emphasis>
Vesica spermalis of the holotype, lateral view, blue arrow indicating the hook-like projection.
</paragraph>
</caption>
<caption id="DF736624FFBC8A71FF5E823EF93DF8A0" ID-DOI="http://doi.org/10.5281/zenodo.7381724" ID-Zenodo-Dep="7381724" box="[152,772,1792,1817]" httpUri="https://zenodo.org/record/7381724/files/figure.png" pageId="7" pageNumber="576" startId="7.[152,255,1792,1816]" targetBox="[237,1351,1221,1769]" targetPageId="7">
<paragraph id="8BB336ACFFBC8A71FF5E823EF93DF8A0" blockId="7.[152,772,1792,1817]" box="[152,772,1792,1817]" pageId="7" pageNumber="576">
<emphasis id="B978EABEFFBC8A71FF5E823EFB1CF8A1" bold="true" box="[152,293,1792,1816]" pageId="7" pageNumber="576">FIGURE 10.</emphasis>
Vesica spermalis of the holotype, ventral view.
</paragraph>
</caption>
<subSubSection id="C3166527FFBC8A71FF018279FE3CF848" pageId="7" pageNumber="576" type="diagnosis">
<paragraph id="8BB336ACFFBC8A71FF018279FE3CF848" blockId="7.[151,1437,1863,2033]" pageId="7" pageNumber="576">
<emphasis id="B978EABEFFBC8A71FF018279FB7BF8D8" bold="true" box="[199,322,1863,1889]" pageId="7" pageNumber="576">Diagnosis.</emphasis>
The body colour pattern, penile organ and hamuli of the new species most closely resembles
<taxonomicName id="4C0C4D2FFFBC8A71FA478279FB2AF83C" authorityName="Selys" authorityYear="1878" class="Insecta" family="Gomphidae" genus="Ophiogomphus" kingdom="Animalia" order="Odonata" pageId="7" pageNumber="576" phylum="Arthropoda" rank="species" species="spinicornis">
<emphasis id="B978EABEFFBC8A71FA478279FB2AF83C" italics="true" pageId="7" pageNumber="576">O. spinicornis</emphasis>
</taxonomicName>
, but differs by the more slender cerci and the divaricate epiproct. The colour pattern is similar other congeners such as
<taxonomicName id="4C0C4D2FFFBC8A71FEAB82B1FBCCF810" authorityName="Bartenev" authorityYear="1909" box="[365,501,1935,1961]" class="Insecta" family="Gomphidae" genus="Ophiogomphus" kingdom="Animalia" order="Odonata" pageId="7" pageNumber="576" phylum="Arthropoda" rank="species" species="obscurus">
<emphasis id="B978EABEFFBC8A71FEAB82B1FBCCF810" box="[365,501,1935,1961]" italics="true" pageId="7" pageNumber="576">O. obscurus</emphasis>
</taxonomicName>
,
<taxonomicName id="4C0C4D2FFFBC8A71FDC382B1F904F813" authority="(Fourcroy, 1785)" baseAuthorityName="Fourcroy" baseAuthorityYear="1785" box="[517,829,1935,1962]" class="Insecta" family="Gomphidae" genus="Ophiogomphus" kingdom="Animalia" order="Odonata" pageId="7" pageNumber="576" phylum="Arthropoda" rank="species" species="cecilia">
<emphasis id="B978EABEFFBC8A71FDC382B1F84DF810" box="[517,628,1935,1961]" italics="true" pageId="7" pageNumber="576">O. cecilia</emphasis>
(Fourcroy, 1785)
</taxonomicName>
, and some American species like
<taxonomicName id="4C0C4D2FFFBC8A71FB0682B1FAEBF877" authority="Morse, 1895" authorityName="Morse" authorityYear="1895" class="Insecta" family="Gomphidae" genus="Ophiogomphus" kingdom="Animalia" order="Odonata" pageId="7" pageNumber="576" phylum="Arthropoda" rank="species" species="aspersus">
<emphasis id="B978EABEFFBC8A71FB0682B1FF7CF810" box="[1216,1349,1935,1961]" italics="true" pageId="7" pageNumber="576">O. aspersus</emphasis>
Morse, 1895
</taxonomicName>
,
<taxonomicName id="4C0C4D2FFFBC8A71FF1A828DF832F874" authority="Needham, 1897" authorityName="Needham" authorityYear="1897" box="[220,523,1971,1997]" class="Insecta" family="Gomphidae" genus="Ophiogomphus" kingdom="Animalia" order="Odonata" pageId="7" pageNumber="576" phylum="Arthropoda" rank="species" species="carolus">
<emphasis id="B978EABEFFBC8A71FF1A828DFB68F874" box="[220,337,1971,1997]" italics="true" pageId="7" pageNumber="576">O. carolus</emphasis>
Needham, 1897
</taxonomicName>
,
<taxonomicName id="4C0C4D2FFFBC8A71FDD3828DF93EF874" authority="Packard" authorityName="Packard" authorityYear="1863" box="[533,775,1971,1997]" class="Insecta" family="Gomphidae" genus="Ophiogomphus" kingdom="Animalia" order="Odonata" pageId="7" pageNumber="576" phylum="Arthropoda" rank="species" species="mainensis">
<emphasis id="B978EABEFFBC8A71FDD3828DF89EF874" box="[533,679,1971,1997]" italics="true" pageId="7" pageNumber="576">O. mainensis</emphasis>
Packard
</taxonomicName>
<emphasis id="B978EABEFFBC8A71FCCB828AF91AF874" box="[781,803,1972,1997]" italics="true" pageId="7" pageNumber="576">in</emphasis>
Walsh, 1863,
<taxonomicName id="4C0C4D2FFFBC8A71FC07828DFEC4F877" authority="(Hagen, 1885)" baseAuthorityName="Hagen" baseAuthorityYear="1885" box="[961,1277,1971,1998]" class="Insecta" family="Gomphidae" genus="Ophiogomphus" kingdom="Animalia" order="Odonata" pageId="7" pageNumber="576" phylum="Arthropoda" rank="species" species="occidentis">
<emphasis id="B978EABEFFBC8A71FC07828DFE6DF874" box="[961,1108,1971,1997]" italics="true" pageId="7" pageNumber="576">O. occidentis</emphasis>
(Hagen, 1885)
</taxonomicName>
, and so on but the more slender cerci and divaricate epiproct differs from all of these species.
</paragraph>
</subSubSection>
</treatment>
</document>
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