treatments-xml/data/03/A5/87/03A587843914582B8D83FD11FC500655.xml
2024-06-21 12:22:17 +02:00

190 lines
20 KiB
XML
Raw Blame History

This file contains ambiguous Unicode characters

This file contains Unicode characters that might be confused with other characters. If you think that this is intentional, you can safely ignore this warning. Use the Escape button to reveal them.

<document id="B35AD38A5B84D361A4055883531B8599" ID-CLB-Dataset="57167" ID-DOI="10.31610/zsr/2020.29.1.11" ID-GBIF-Dataset="e387fc4b-0c64-494e-a2ad-d3b07f69e985" ID-ISSN="2410-0226" ID-Zenodo-Dep="10124222" ID-ZooBank="urn:lsid:zoobank.org:pub:0BAEA1B3-2FE7-427E-A6BA-718F5800CFDB" IM.bibliography_approvedBy="juliana" IM.illustrations_approvedBy="juliana" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_approvedBy="juliana" IM.taxonomicNames_approvedBy="felipe" IM.treatments_approvedBy="juliana" checkinTime="1688480079307" checkinUser="felipe" docAuthor="Tsalolikhin, S. Ya." docDate="2020" docId="03A587843914582B8D83FD11FC500655" docLanguage="en" docName="ZoosystRoss.29.1.11-16.pdf" docOrigin="Zoosystematica Rossica (Zoosyst. Rossica) 29 (1)" docSource="http://dx.doi.org/10.31610/zsr/2020.29.1.11" docStyle="DocumentStyle:8B7E50B51175314E898B46063364B751.1:ZoosystRoss.2019-.journal_article" docStyleId="8B7E50B51175314E898B46063364B751" docStyleName="ZoosystRoss.2019-.journal_article" docStyleVersion="1" docTitle="Monhystera vietnamica Tsalolikhin 2020, sp. nov." docType="treatment" docUuid="C86F7C84-18C0-4255-8E75-E90CD1F88F38" docUuidSource="ZooBank" docVersion="8" lastPageNumber="12" masterDocId="FF9CFFFC3915582A8D13FFBFFFD70447" masterDocTitle="Two new species of free-living nematodes (Nematoda) from Vietnam" masterLastPageNumber="16" masterPageNumber="11" pageNumber="12" updateTime="1700050034748" updateUser="ExternalLinkService" zenodo-license-document="CC-BY-4.0">
<mods:mods id="9F4C6A5602283707D9FCD7E77D6AAAC3" xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo id="757E22FFAB72D683C4193D0605CB4D87">
<mods:title id="F2F6AE4BF6B5D1D84F341BAAB23BF05E">Two new species of free-living nematodes (Nematoda) from Vietnam</mods:title>
</mods:titleInfo>
<mods:name id="B42F67A46830C5865DC4D1940E26E179" type="personal">
<mods:role id="E4D6564F240812A3414B7B3A3E251700">
<mods:roleTerm id="94FBC36D94A96CDA743BE05B8029ACE0">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="F0A969CB7B20E0A1264B655A89B85BC5">Tsalolikhin, S. Ya.</mods:namePart>
</mods:name>
<mods:typeOfResource id="A2B3721885DE74DAB5B2DAF196CD7F8B">text</mods:typeOfResource>
<mods:relatedItem id="7E22A03FDFC6F3BFFB663D000EA80A07" type="host">
<mods:titleInfo id="86D27AC262450ED2C2409B0587555653">
<mods:title id="F46DF52C5E253FE47110F48DB9D266B3">Zoosystematica Rossica</mods:title>
</mods:titleInfo>
<mods:part id="3DB03E080FA7507FE69920D632D63D87">
<mods:date id="263E6803FE17A0DFAB6AD61A3B150158">2020</mods:date>
<mods:detail id="CAFA8EB0C6021518DB068186D69BA82B" type="series">
<mods:title id="E84132A885E9A44ADF8949C7859090DE">Zoosyst. Rossica</mods:title>
</mods:detail>
<mods:detail id="5A34A73BC3EFD5639F6DC61BDD2E52A2" type="pubDate">
<mods:number id="B1A176668E99EE04B6B525836715F199">2020-03-31</mods:number>
</mods:detail>
<mods:detail id="EE0587A975B1E8B8DE07DF97B6631871" type="volume">
<mods:number id="550407CD7CA67AD87BCCD4EFE7A62E4A">29</mods:number>
</mods:detail>
<mods:detail id="C055DB5177A81EA547CE9845B2187310" type="issue">
<mods:number id="393EF91FFDFB3B3BEEBDA2A2304509BA">1</mods:number>
</mods:detail>
<mods:extent id="CB67FFB402AF50ED86C94C6B90AA49F2" unit="page">
<mods:start id="FDB66A8B765C28D792B180176D563F14">11</mods:start>
<mods:end id="C04D435E205239307ED0E3FD856F4C51">16</mods:end>
</mods:extent>
</mods:part>
</mods:relatedItem>
<mods:location id="9E2B71EDB52D76C201DBA7188B62C13F">
<mods:url id="EA174B22CD248CBE5591114C404E77A8">http://dx.doi.org/10.31610/zsr/2020.29.1.11</mods:url>
</mods:location>
<mods:classification id="A44CA46DAB045E14239B3F5BD8B1F9EB">journal article</mods:classification>
<mods:identifier id="D7FA0615208B5D15018A876A67A0C0A2" type="CLB-Dataset">57167</mods:identifier>
<mods:identifier id="B37B0EF07482162B3A96F2B9A83EC859" type="DOI">10.31610/zsr/2020.29.1.11</mods:identifier>
<mods:identifier id="D56B216E9D3D4EA994DB7712E0F4B7AA" type="GBIF-Dataset">e387fc4b-0c64-494e-a2ad-d3b07f69e985</mods:identifier>
<mods:identifier id="B4195500580E156EDE57D3F9B2945D26" type="ISSN">2410-0226</mods:identifier>
<mods:identifier id="7ED8F80EA4CA1B548BC22065739D4E93" type="Zenodo-Dep">10124222</mods:identifier>
<mods:identifier id="B46A16F7795391E2A457700FCEACD3AA" type="ZooBank">urn:lsid:zoobank.org:pub:0BAEA1B3-2FE7-427E-A6BA-718F5800CFDB</mods:identifier>
</mods:mods>
<treatment id="03A587843914582B8D83FD11FC500655" ID-DOI="http://doi.org/10.5281/zenodo.8113945" ID-GBIF-Taxon="210811757" ID-Zenodo-Dep="8113945" LSID="urn:lsid:zoobank.org:act:C86F7C84-18C0-4255-8E75-E90CD1F88F38" httpUri="http://treatment.plazi.org/id/03A587843914582B8D83FD11FC500655" lastPageNumber="12" pageId="1" pageNumber="12">
<subSubSection id="C31665193914582B8D83FD11FDEF068E" box="[144,568,686,713]" pageId="1" pageNumber="12" type="nomenclature">
<paragraph id="8BB336923914582B8D83FD11FDEF068E" blockId="1.[144,568,686,713]" box="[144,568,686,713]" pageId="1" pageNumber="12">
<heading id="D0FB81FE3914582B8D83FD11FDEF068E" box="[144,568,686,713]" fontSize="11" level="2" pageId="1" pageNumber="12" reason="2">
<emphasis id="B978EA803914582B8D83FD11FDEF068E" bold="true" box="[144,568,686,713]" pageId="1" pageNumber="12">
<taxonomicName id="4C0C4D113914582B8D83FD11FE1A068E" authority="Tsalolikhin, 2020" authorityName="Tsalolikhin" authorityYear="2020" box="[144,461,686,713]" class="Adenophorea" family="Monhysteridae" genus="Monhystera" kingdom="Animalia" order="Monhysterida" pageId="1" pageNumber="12" phylum="Nematoda" rank="species" species="vietnamica" status="sp. nov.">
<emphasis id="B978EA803914582B8D83FD11FE1A068E" bold="true" box="[144,461,686,713]" italics="true" pageId="1" pageNumber="12">Monhystera vietnamica</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A24B57FB3914582B8CC7FD10FDEF068E" box="[468,568,687,713]" pageId="1" pageNumber="12" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C31665193914582B8D83FD6DFEC306AA" box="[144,276,722,749]" pageId="1" pageNumber="12" type="description">
<paragraph id="8BB336923914582B8D83FD6DFEC306AA" blockId="1.[144,276,722,749]" box="[144,276,722,749]" pageId="1" pageNumber="12">
(
<figureCitation id="13372A173914582B8D8AFD6DFEDD06AA" box="[153,266,722,749]" captionStart="Figs 19" captionStartId="2.[159,210,1837,1862]" captionTargetBox="[174,1431,189,1767]" captionTargetId="figure-21@2.[159,1444,181,1812]" captionTargetPageId="2" captionText="Figs 19. Morphology of Monhistera vietnamica sp. nov. (15) and Brevitobrilus larae sp. nov. (69). 1, 6, entire body of female; 2, 7, head; 3, esophagus; 4, 8, vulval section; 5, 9, tail. Scale bars: 2, 7 5 µm; 3, 4, 5, 8, 9 20 µm; 1, 6 50 µm." figureDoi="http://doi.org/10.5281/zenodo.10135599" httpUri="https://zenodo.org/record/10135599/files/figure.png" pageId="1" pageNumber="12">Figs 14</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="C31665193914582B8DA6FCB4FD0607E2" pageId="1" pageNumber="12" type="materials_examined">
<paragraph id="8BB336923914582B8DA6FCB4FD2407C2" blockId="1.[144,764,779,1927]" pageId="1" pageNumber="12">
<materialsCitation id="3B643CCF3914582B8DA6FCB4FD2407C2" ID-GBIF-Occurrence="4144866301" collectingDate="2019-06-16" collectionCode="ZISP" collectorName="L. Iogansen" country="Vietnam" county="Tung Vui Commune" elevation="400" latitude="22.329445" location="Tung Vui Commune" longLatPrecision="21" longitude="105.42972" municipality="Na Hang Distr." pageId="1" pageNumber="12" specimenCode="A-9238" specimenCount="1" stateProvince="Tuyn Quang Prov" typeStatus="holotype">
<emphasis id="B978EA803914582B8DA6FCB4FECC0763" box="[181,283,779,804]" italics="true" pageId="1" pageNumber="12">
<typeStatus id="54B788303914582B8DA6FCB4FECC0763" box="[181,283,779,804]" pageId="1" pageNumber="12" type="holotype">Holotype</typeStatus>
</emphasis>
.
<specimenCount id="9D0AFD1B3914582B8C22FCB4FE510763" box="[305,390,779,804]" pageId="1" pageNumber="12" type="female">Female</specimenCount>
,
<emphasis id="B978EA803914582B8C8AFCB4FDD70763" bold="true" box="[409,512,779,804]" pageId="1" pageNumber="12">
<collectingCountry id="F31B76023914582B8C8AFCB4FDD70763" box="[409,512,779,804]" name="Vietnam" pageId="1" pageNumber="12">Vietnam</collectingCountry>
</emphasis>
,
<emphasis id="B978EA803914582B8F05FCB3FD210764" box="[534,758,780,803]" italics="true" pageId="1" pageNumber="12">
<collectingRegion id="49C8F8703914582B8F05FCB3FD260764" box="[534,753,780,803]" pageId="1" pageNumber="12">Tuyn Quang Prov</collectingRegion>
.
</emphasis>
,
<collectingMunicipality id="6BD7ACE83914582B8D83FC93FE900703" box="[144,327,811,836]" pageId="1" pageNumber="12">Na Hang Distr.</collectingMunicipality>
,
<location id="8ED360493914582B8C47FC93FD920703" LSID="urn:lsid:plazi:treatment:03A587843914582B8D83FD11FC500655:8ED360493914582B8C47FC93FD920703" box="[340,581,811,836]" country="Vietnam" county="Tung Vui Commune" latitude="22.329445" longLatPrecision="21" longitude="105.42972" municipality="Na Hang Distr." name="Tung Vui Commune" pageId="1" pageNumber="12" stateProvince="Tuyn Quang Prov">Tung Vui Commune</location>
, pond in forest,
<elevation id="0021D1A13914582B8D83FCF4FECF0723" box="[144,280,843,868]" metricMagnitude="2" metricUnit="m" metricValue="4.0" pageId="1" pageNumber="12" unit="m" value="400.0">
<quantity id="4CF49B773914582B8D83FCF4FF0D0724" box="[144,218,843,868]" metricMagnitude="2" metricUnit="m" metricValue="4.0" pageId="1" pageNumber="12" unit="m" value="400.0">400 m</quantity>
a.s.l.
</elevation>
,
<geoCoordinate id="EE3850553914582B8C3BFCF4FE7A0724" box="[296,429,842,868]" degrees="22" direction="north" minutes="19" orientation="latitude" pageId="1" pageNumber="12" precision="15" seconds="46" value="22.329445">22°1946″N</geoCoordinate>
,
<geoCoordinate id="EE3850553914582B8CA8FCF4FD9C0724" box="[443,587,842,868]" degrees="105" direction="east" minutes="25" orientation="longitude" pageId="1" pageNumber="12" precision="15" seconds="47" value="105.42972">105°2547″E</geoCoordinate>
,
<date id="FFB210523914582B8F49FCF4FD2F0723" box="[602,760,843,868]" pageId="1" pageNumber="12" value="2019-06-16">
<collectingDate id="EFF6E9BA3914582B8F49FCF4FD2F0723" box="[602,760,843,868]" pageId="1" pageNumber="12" value="2019-06-16">16 June 2019</collectingDate>
</date>
, coll.
<collectorName id="26F953443914582B8DD5FCD2FE0807C2" box="[198,479,876,901]" pageId="1" pageNumber="12">L. Iogansen. Microscope</collectorName>
slide No.
<specimenCode id="DBAA9EE93914582B8F5DFCD3FD7107C2" box="[590,678,876,901]" pageId="1" pageNumber="12">A-9238</specimenCode>
,
<collectionCode id="ED1DAE573914582B8FA3FCD3FD3907C2" box="[688,750,876,901]" country="Russia" httpUri="http://grbio.org/cool/vmfx-g997" name="Zoological Institute, Russian Academy of Sciences" pageId="1" pageNumber="12">ZISP</collectionCode>
.
</materialsCitation>
</paragraph>
<paragraph id="8BB336923914582B8DA6FC32FD0607E2" blockId="1.[144,764,779,1927]" box="[181,721,908,933]" pageId="1" pageNumber="12">
<materialsCitation id="3B643CCF3914582B8DA6FC32FD1A07E2" ID-GBIF-Occurrence="4144866303" box="[181,717,908,933]" collectingDate="2019-06-16" collectionCode="ZISP" collectorName="L. Iogansen" country="Vietnam" county="Tung Vui Commune" elevation="400" latitude="22.329445" longLatPrecision="21" longitude="105.42972" municipality="Na Hang Distr." pageId="1" pageNumber="12" specimenCode="A-9238" specimenCount="9" specimenCount-female="9" stateProvince="Tuyn Quang Prov" typeStatus="paratype">
<emphasis id="B978EA803914582B8DA6FC32FEF007E3" box="[181,295,909,932]" italics="true" pageId="1" pageNumber="12">
<typeStatus id="54B788303914582B8DA6FC32FEF007E3" box="[181,295,909,932]" pageId="1" pageNumber="12" type="paratype">Paratypes</typeStatus>
</emphasis>
.
<specimenCount id="9D0AFD1B3914582B8C27FC33FE4807E2" box="[308,415,908,933]" pageId="1" pageNumber="12" type="female">9 females</specimenCount>
, same data as for holotype
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="C31665193914582B8DA6FC10FEAA02C9" pageId="1" pageNumber="12" type="description">
<paragraph id="8BB336923914582B8DA6FC10FD37022C" blockId="1.[144,764,779,1927]" pageId="1" pageNumber="12">
<emphasis id="B978EA803914582B8DA6FC10FE9B078F" box="[181,332,943,968]" italics="true" pageId="1" pageNumber="12">Description.</emphasis>
Females. Morphometric description of the
<typeStatus id="54B788303914582B8C3FFC6EFE4D07AB" box="[300,410,977,1004]" pageId="1" pageNumber="12" type="holotype">holotype</typeStatus>
and
<typeStatus id="54B788303914582B8CF6FC6DFDB707AB" box="[485,608,978,1004]" pageId="1" pageNumber="12" type="paratype">paratypes</typeStatus>
is given in
<tableCitation id="C68E03293914582B8D83FC4BFF3B0048" box="[144,236,1012,1039]" captionStart="Table 1" captionStartId="3.[115,176,441,463]" captionTargetId="graphics-38@3.[115,1851,484,1645]" captionTargetPageId="3" captionText="Table 1. Comparative morphometry of Monhystera vietnamica sp. nov., M. rolandi and M. uncigubernaculum (females)." httpUri="http://table.plazi.org/id/DF73661A391658298D60FE06FA940588" pageId="1" pageNumber="12" tableUuid="DF73661A391658298D60FE06FA940588">Table 1</tableCitation>
. Cuticle thin, about 1 µm. Space between internal organs with small cristalloid bodies of various sizes. Head not offset from body outline. Labial papillae undetectable. Diameter of amphidial fovea corresponding to 30% of body width. Ocellus present at about 2026 (23) µm from anterior end. Cheilostome medium-sized, with thin walls. Esophagus muscular, gradually dilating towards its base. Cardium mushroom-shaped. Renette more or less oval, located somewhat below cardium; canal of renette and excretory pore not visible. Reproductive system monodelphic, prodelphic. Oocytes arranged in several rows in germinal zone. Uterus usually containing one or
<specimenCount id="9D0AFD1B3914582B8D83FA59FED40247" box="[144,259,1510,1536]" pageId="1" pageNumber="12" type="egg">two eggs</specimenCount>
: 2936 x 1423 µm. Vagina very long, cuticularised, noticeably longer than one vulval body diameter. Paravulval gland present. Rectum longer than anal diameter. Tail slender, filiform.
</paragraph>
<paragraph id="8BB336923914582B8DA6F9CCFEAA02C9" blockId="1.[144,764,779,1927]" box="[181,381,1651,1678]" pageId="1" pageNumber="12">Male not found.</paragraph>
</subSubSection>
<subSubSection id="C31665193914582B8DA6F927FC5A058C" pageId="1" pageNumber="12" type="diagnosis">
<paragraph id="8BB336923914582B8DA6F927FC5A058C" blockId="1.[144,764,779,1927]" lastBlockId="1.[809,1429,184,530]" pageId="1" pageNumber="12">
<emphasis id="B978EA803914582B8DA6F927FE8302F5" box="[181,340,1688,1714]" italics="true" pageId="1" pageNumber="12">Comparision</emphasis>
. The new species
<taxonomicName id="4C0C4D113914582B8F7BF928FEC80292" authority="Tsalolikhin, 2020" authorityName="Tsalolikhin" authorityYear="2020" class="Adenophorea" family="Monhysteridae" genus="Monhystera" kingdom="Animalia" order="Monhysterida" pageId="1" pageNumber="12" phylum="Nematoda" rank="species" species="vietnamica" status="sp. nov.">
<emphasis id="B978EA803914582B8F7BF928FEC80292" italics="true" pageId="1" pageNumber="12">Monhystera vietnamica</emphasis>
</taxonomicName>
<emphasis id="B978EA803914582B8C0CF904FE530292" bold="true" box="[287,388,1723,1749]" pageId="1" pageNumber="12">
<taxonomicNameLabel id="A24B57FB3914582B8C0CF904FE530292" box="[287,388,1723,1749]" pageId="1" pageNumber="12" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
most closely resembles
<taxonomicName id="4C0C4D113914582B8FBAF903FD2002BF" authority="Zeidan, Jacobs et Geraert, 1989" authorityName="Zeidan, Jacobs et Geraert" authorityYear="1989" class="Adenophorea" family="Monhysteridae" genus="Monhystera" kingdom="Animalia" order="Monhysterida" pageId="1" pageNumber="12" phylum="Nematoda" rank="species" species="uncigubernaculum">
<emphasis id="B978EA803914582B8FBAF903FE8102BE" italics="true" pageId="1" pageNumber="12">M. uncigubernaculum</emphasis>
Zeidan, Jacobs et Geraert, 1989
</taxonomicName>
, which was described from Africa (Zeidan, Jacobs &amp; Geraert, 1989), and
<taxonomicName id="4C0C4D113914582B8CAFF899FEC00324" authority="Khan et Tahseen, 2006" authorityName="Khan et Tahseen" authorityYear="2006" class="Adenophorea" family="Monhysteridae" genus="Monhystera" kingdom="Animalia" order="Monhysterida" pageId="1" pageNumber="12" phylum="Nematoda" rank="species" species="rolandi">
<emphasis id="B978EA803914582B8CAFF899FD960307" box="[444,577,1829,1856]" italics="true" pageId="1" pageNumber="12">M. rolandi</emphasis>
<bibRefCitation id="EF9D4B633914582B8F5FF89AFEC00324" author="Khan R. &amp; Tahseen Q." pageId="1" pageNumber="12" pagination="91 - 101" refId="ref2689" refString="Khan R. &amp; Tahseen Q. 2006. Description of two new monhysterid species (Nematoda: Monhysterida) from sewage in India. Journal of Nematode Morphology and Systematics, (2005), 8 (2): 91 - 101." type="journal article" year="2006">Khan et Tahseen, 2006</bibRefCitation>
</taxonomicName>
from
<collectingCountry id="F31B76023914582B8C75F8F7FE7E0324" box="[358,425,1864,1891]" name="India" pageId="1" pageNumber="12">India</collectingCountry>
(Khan &amp; Tahseen, 2005). The main differences of the new species from
<taxonomicName id="4C0C4D113914582B8E3AFF05FBE90494" authorityName="Zeidan, Jacobs et Geraert" authorityYear="1989" box="[809,1086,184,211]" class="Adenophorea" family="Monhysteridae" genus="Monhystera" kingdom="Animalia" order="Monhysterida" pageId="1" pageNumber="12" phylum="Nematoda" rank="species" species="uncigubernaculum">
<emphasis id="B978EA803914582B8E3AFF05FBE90494" box="[809,1086,184,211]" italics="true" pageId="1" pageNumber="12">M. uncigubernaculum</emphasis>
</taxonomicName>
are the following: vagina 4552 µm (
<emphasis id="B978EA803914582B8ED2FF62FC0D04B1" box="[961,986,221,246]" italics="true" pageId="1" pageNumber="12">vs</emphasis>
2537 µm), rectum/anal body diameter 0.9 (
<emphasis id="B978EA803914582B8ED0FEBEFC0B055D" box="[963,988,257,282]" italics="true" pageId="1" pageNumber="12">vs</emphasis>
0.7), and tail filiform (
<emphasis id="B978EA803914582B8817FEBEFACA055D" box="[1284,1309,257,282]" italics="true" pageId="1" pageNumber="12">vs</emphasis>
tail with slightly swollen spinneret). The main differences of the new species from
<taxonomicName id="4C0C4D113914582B896EFEF8FAD70526" authorityName="Khan et Tahseen" authorityYear="2006" box="[1149,1280,326,353]" class="Adenophorea" family="Monhysteridae" genus="Monhystera" kingdom="Animalia" order="Monhysterida" pageId="1" pageNumber="12" phylum="Nematoda" rank="species" species="rolandi">
<emphasis id="B978EA803914582B896EFEF8FAD70526" box="[1149,1280,326,353]" italics="true" pageId="1" pageNumber="12">M. rolandi</emphasis>
</taxonomicName>
are: ocellus present (
<emphasis id="B978EA803914582B8E87FED4FC7A05C3" box="[916,941,363,388]" italics="true" pageId="1" pageNumber="12">vs</emphasis>
absence of ocellus), cephalic setae 5 µm (
<emphasis id="B978EA803914582B8E20FE31FC9B05E0" box="[819,844,398,423]" italics="true" pageId="1" pageNumber="12">vs</emphasis>
34 µm), and tail filiform (
<emphasis id="B978EA803914582B89B0FE31FB6B05E0" box="[1187,1212,398,423]" italics="true" pageId="1" pageNumber="12">vs</emphasis>
tail gradually tapering).
</paragraph>
</subSubSection>
<subSubSection id="C31665193914582B8E5CFE6BFC500655" pageId="1" pageNumber="12" type="etymology">
<paragraph id="8BB336923914582B8E5CFE6BFC500655" blockId="1.[809,1429,184,530]" pageId="1" pageNumber="12">
<emphasis id="B978EA803914582B8E5CFE6BFC0605A8" box="[847,977,468,495]" italics="true" pageId="1" pageNumber="12">Etymology</emphasis>
. This species name is a Latinized toponym.
</paragraph>
</subSubSection>
</treatment>
</document>