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<document id="C312810AC0751002B6E1C49837508954" ID="10.11646/zootaxa.4169.2.1" ID-DOI="10.11646/zootaxa.4169.2.1" ID-GBIF-Dataset="d1fbe4b3-de10-4d72-a117-aa6bfedf210d" ID-ISSN="1175-5326" ID-Zenodo-Dep="262585" ID-ZooBank="B8B067BF-F6E6-4122-B884-AA385FF04421" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1474351831659" checkinUser="plazi" docAuthor="Andrew Hamilton, K. G." docDate="2016" docId="03A5742DFFC7AE19FF76AB3BCC2FFECD" docLanguage="en" docName="zootaxa.4169.2.1.pdf" docOrigin="Zootaxa 4169 (2)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Menytes Hamilton, 2016, gen.nov." docType="treatment" docVersion="7" lastPageNumber="220" masterDocId="FF9C0C55FFD5AE0AFFE1A955CD70FFD1" masterDocTitle="Neotropical spittlebugs related to Neaenini (Hemiptera, Cercopidae) and the origins of subfamily Cercopinae" masterLastPageNumber="250" masterPageNumber="201" pageNumber="219" updateTime="1698386375821" updateUser="plazi">
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<mods:title id="715810CFAD560CE6BDD0FBA9E60D385A">Neotropical spittlebugs related to Neaenini (Hemiptera, Cercopidae) and the origins of subfamily Cercopinae</mods:title>
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<mods:namePart id="9DB406025215CBA062E8CCF35413397D">Andrew Hamilton, K. G.</mods:namePart>
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<mods:part id="ADC1EA64030C90B2BA50075F602A8E39">
<mods:date id="60A621DBABD7599EAC2D66EE21DD6A1D">2016</mods:date>
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<mods:number id="C149496B073DBB8E9FBDAA09EF7E049B">4169</mods:number>
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<mods:identifier id="CBC57FBA93F8F50E618EAF6DA00F0BF4" type="DOI">10.11646/zootaxa.4169.2.1</mods:identifier>
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<treatment id="03A5742DFFC7AE19FF76AB3BCC2FFECD" ID-DOI="http://doi.org/10.5281/zenodo.5613347" ID-GBIF-Taxon="124552103" ID-Zenodo-Dep="5613347" LSID="urn:lsid:plazi:treatment:03A5742DFFC7AE19FF76AB3BCC2FFECD" httpUri="http://treatment.plazi.org/id/03A5742DFFC7AE19FF76AB3BCC2FFECD" lastPageId="19" lastPageNumber="220" pageId="18" pageNumber="219">
<subSubSection id="C31696B0FFC7AE18FF76AB3BCF50FD1A" pageId="18" pageNumber="219" type="nomenclature">
<paragraph id="8BB3C53BFFC7AE18FF76AB3BCC19FD59" blockId="18.[151,361,622,648]" box="[151,361,622,648]" pageId="18" pageNumber="219">
<heading id="D0FB7257FFC7AE18FF76AB3BCC19FD59" bold="true" box="[151,361,622,648]" fontSize="11" level="1" pageId="18" pageNumber="219" reason="1">
<emphasis id="B9781929FFC7AE18FF76AB3BCC19FD59" bold="true" box="[151,361,622,648]" pageId="18" pageNumber="219">
<taxonomicName id="4C0CBEB8FFC7AE18FF76AB3BCD8CFD59" box="[151,252,622,648]" class="Insecta" family="Cercopidae" genus="Menytes" kingdom="Animalia" order="Hemiptera" pageId="18" pageNumber="219" phylum="Arthropoda" rank="genus" status="gen. nov.">
<emphasis id="B9781929FFC7AE18FF76AB3BCD8CFD59" bold="true" box="[151,252,622,648]" italics="true" pageId="18" pageNumber="219">Menytes</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A24BA452FFC7AE18FEE2AB3BCC19FD59" box="[259,361,622,648]" pageId="18" pageNumber="219" rank="genus">gen.nov.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
<paragraph id="8BB3C53BFFC7AE18FF76ABE0CF50FD1A" blockId="18.[151,544,693,715]" box="[151,544,693,715]" pageId="18" pageNumber="219">
Type-species:
<taxonomicName id="4C0CBEB8FFC7AE18FEC9ABE3CCA2FD1A" box="[296,466,693,715]" class="Insecta" family="Cercopidae" genus="Menytes" kingdom="Animalia" order="Hemiptera" pageId="18" pageNumber="219" phylum="Arthropoda" rank="species" species="lathrius" status="sp. nov.">
<emphasis id="B9781929FFC7AE18FEC9ABE3CCA2FD1A" box="[296,466,693,715]" italics="true" pageId="18" pageNumber="219">Menytes lathrius</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A24BA452FFC7AE18FE39ABE3CF50FD1A" box="[472,544,694,715]" pageId="18" pageNumber="219" rank="species">sp.nov.</taxonomicNameLabel>
</paragraph>
</subSubSection>
<subSubSection id="C31696B0FFC7AE18FF76ABADCF54FCC0" box="[151,548,760,785]" pageId="18" pageNumber="219" type="etymology">
<paragraph id="8BB3C53BFFC7AE18FF76ABADCF54FCC0" blockId="18.[151,1437,760,1793]" box="[151,548,760,785]" pageId="18" pageNumber="219">
<emphasis id="B9781929FFC7AE18FF76ABADCC6EFCC0" bold="true" box="[151,286,760,785]" pageId="18" pageNumber="219">Etymology.</emphasis>
<emphasis id="B9781929FFC7AE18FEC5ABACCC0EFCC1" box="[292,382,761,784]" italics="true" pageId="18" pageNumber="219">menytes</emphasis>
(m), informer.
</paragraph>
</subSubSection>
<subSubSection id="C31696B0FFC7AE18FF26AA49CFE1FCE4" box="[199,657,796,821]" pageId="18" pageNumber="219" type="distribution">
<paragraph id="8BB3C53BFFC7AE18FF26AA49CFE1FCE4" blockId="18.[151,1437,760,1793]" box="[199,657,796,821]" pageId="18" pageNumber="219">
<emphasis id="B9781929FFC7AE18FF26AA49CC10FCE4" bold="true" box="[199,352,796,821]" pageId="18" pageNumber="219">Distribution.</emphasis>
Amazon basin of
<collectingCountry id="F31B85ABFFC7AE18FDCCAA49CFFDFCE4" box="[557,653,796,821]" name="Ecuador" pageId="18" pageNumber="219">Ecuador</collectingCountry>
.
</paragraph>
</subSubSection>
<subSubSection id="C31696B0FFC7AE18FF26AA15CC71FCAD" pageId="18" pageNumber="219" type="diagnosis">
<paragraph id="8BB3C53BFFC7AE18FF26AA15CC71FCAD" blockId="18.[151,1437,760,1793]" pageId="18" pageNumber="219">
<emphasis id="B9781929FFC7AE18FF26AA15CC31FC88" bold="true" box="[199,321,832,857]" pageId="18" pageNumber="219">Diagnosis.</emphasis>
Intermediate between the very different genera
<taxonomicName id="4C0CBEB8FFC7AE18FC95AA15CE98FC88" box="[884,1000,832,857]" class="Insecta" family="Cercopidae" genus="Simorhina" kingdom="Animalia" order="Hemiptera" pageId="18" pageNumber="219" phylum="Arthropoda" rank="genus">
<emphasis id="B9781929FFC7AE18FC95AA15CE98FC88" box="[884,1000,832,857]" italics="true" pageId="18" pageNumber="219">Simorhina</emphasis>
</taxonomicName>
and
<taxonomicName id="4C0CBEB8FFC7AE18FBC7AA15C9BFFC88" box="[1062,1231,832,857]" class="Insecta" family="Cercopidae" genus="Tomaspisinella" kingdom="Animalia" order="Hemiptera" pageId="18" pageNumber="219" phylum="Arthropoda" rank="genus">
<emphasis id="B9781929FFC7AE18FBC7AA15C9BFFC88" box="[1062,1231,832,857]" italics="true" pageId="18" pageNumber="219">Tomaspisinella</emphasis>
</taxonomicName>
, and with unique antennae.
</paragraph>
</subSubSection>
<subSubSection id="C31696B0FFC7AE18FF26AADDCF49FA49" pageId="18" pageNumber="219" type="description">
<paragraph id="8BB3C53BFFC7AE18FF26AADDCF49FA49" blockId="18.[151,1437,760,1793]" pageId="18" pageNumber="219">
<emphasis id="B9781929FFC7AE18FF26AADDCC29FC70" bold="true" box="[199,345,904,929]" pageId="18" pageNumber="219">Description.</emphasis>
Superficially similar to a small
<taxonomicName id="4C0CBEB8FFC7AE18FD3CAADDCE15FC70" box="[733,869,904,929]" class="Insecta" family="Aphrophoridae" genus="Aphrophora" kingdom="Animalia" order="Hemiptera" pageId="18" pageNumber="219" phylum="Arthropoda" rank="genus">
<emphasis id="B9781929FFC7AE18FD3CAADDCE15FC70" box="[733,869,904,929]" italics="true" pageId="18" pageNumber="219">Aphrophora</emphasis>
</taxonomicName>
but strongly hirsute (
<figureCitation id="1337D9BEFFC7AE18FB88AADDC9B0FC71" box="[1129,1216,904,929]" captionStart="FIGURES 14 15" captionStartId="32.[151,264,1884,1906]" captionTargetBox="[237,1347,193,1862]" captionTargetId="figure@32.[237,1347,193,1862]" captionTargetPageId="32" captionText="FIGURES 14 15. Menytes gen. nov. (14), compared to Simorhina Jacobi (15). A, habitus; B, profile; C, hind wing; D, venter (restoration) with photograph of actual specimen (inset); E, left half of head, anterior aspect; F, enlargement of tip of antenna (scale bar: 100 F), widest aspect, showing length of basiconic sensillum (b) and width compared to arista (a); G, tip of postpedicel, with arista to left and basiconic sensillum to right; H, enlargement of antennal pits (scale bar: 10 F) showing spiral base of each vertically pleated coeloconic sensillum (dark centre). Original photograph (inset in 12 D) provided by J. Boone." httpUri="https://zenodo.org/record/262590/files/figure.png" pageId="18" pageNumber="219">Fig. 14</figureCitation>
B). Head distinctly narrower than pronotum; eyes transverse; crown moderately short, with tylus and crown depressed and strongly elevated antennal ledge (
<figureCitation id="1337D9BEFFC7AE18FE4EAA85CF71FC39" box="[431,513,976,1001]" captionStart="FIGURES 14 15" captionStartId="32.[151,264,1884,1906]" captionTargetBox="[237,1347,193,1862]" captionTargetId="figure@32.[237,1347,193,1862]" captionTargetPageId="32" captionText="FIGURES 14 15. Menytes gen. nov. (14), compared to Simorhina Jacobi (15). A, habitus; B, profile; C, hind wing; D, venter (restoration) with photograph of actual specimen (inset); E, left half of head, anterior aspect; F, enlargement of tip of antenna (scale bar: 100 F), widest aspect, showing length of basiconic sensillum (b) and width compared to arista (a); G, tip of postpedicel, with arista to left and basiconic sensillum to right; H, enlargement of antennal pits (scale bar: 10 F) showing spiral base of each vertically pleated coeloconic sensillum (dark centre). Original photograph (inset in 12 D) provided by J. Boone." httpUri="https://zenodo.org/record/262590/files/figure.png" pageId="18" pageNumber="219">Fig. 14</figureCitation>
E); facial part of frons medially sulcate (
<figureCitation id="1337D9BEFFC7AE18FC28AA85C96BFC39" box="[969,1051,976,1001]" captionStart="FIGURES 14 15" captionStartId="32.[151,264,1884,1906]" captionTargetBox="[237,1347,193,1862]" captionTargetId="figure@32.[237,1347,193,1862]" captionTargetPageId="32" captionText="FIGURES 14 15. Menytes gen. nov. (14), compared to Simorhina Jacobi (15). A, habitus; B, profile; C, hind wing; D, venter (restoration) with photograph of actual specimen (inset); E, left half of head, anterior aspect; F, enlargement of tip of antenna (scale bar: 100 F), widest aspect, showing length of basiconic sensillum (b) and width compared to arista (a); G, tip of postpedicel, with arista to left and basiconic sensillum to right; H, enlargement of antennal pits (scale bar: 10 F) showing spiral base of each vertically pleated coeloconic sensillum (dark centre). Original photograph (inset in 12 D) provided by J. Boone." httpUri="https://zenodo.org/record/262590/files/figure.png" pageId="18" pageNumber="219">Fig. 14</figureCitation>
D); antenna with slender, elongate sensillum (
<figureCitation id="1337D9BEFFC7AE18FEF3AAA1CC13FBDD" box="[274,355,1012,1037]" captionStart="FIGURES 14 15" captionStartId="32.[151,264,1884,1906]" captionTargetBox="[237,1347,193,1862]" captionTargetId="figure@32.[237,1347,193,1862]" captionTargetPageId="32" captionText="FIGURES 14 15. Menytes gen. nov. (14), compared to Simorhina Jacobi (15). A, habitus; B, profile; C, hind wing; D, venter (restoration) with photograph of actual specimen (inset); E, left half of head, anterior aspect; F, enlargement of tip of antenna (scale bar: 100 F), widest aspect, showing length of basiconic sensillum (b) and width compared to arista (a); G, tip of postpedicel, with arista to left and basiconic sensillum to right; H, enlargement of antennal pits (scale bar: 10 F) showing spiral base of each vertically pleated coeloconic sensillum (dark centre). Original photograph (inset in 12 D) provided by J. Boone." httpUri="https://zenodo.org/record/262590/files/figure.png" pageId="18" pageNumber="219">Fig. 14</figureCitation>
F) as in
<taxonomicName id="4C0CBEB8FFC7AE18FE5EAAA1CF43FBDC" box="[447,563,1012,1037]" class="Insecta" family="Cercopidae" genus="Simorhina" kingdom="Animalia" order="Hemiptera" pageId="18" pageNumber="219" phylum="Arthropoda" rank="genus">
<emphasis id="B9781929FFC7AE18FE5EAAA1CF43FBDC" box="[447,563,1012,1037]" italics="true" pageId="18" pageNumber="219">Simorhina</emphasis>
</taxonomicName>
and
<taxonomicName id="4C0CBEB8FFC7AE18FD8AAAA1CE64FBDC" box="[619,788,1012,1037]" class="Insecta" family="Cercopidae" genus="Tomaspisinella" kingdom="Animalia" order="Hemiptera" pageId="18" pageNumber="219" phylum="Arthropoda" rank="genus">
<emphasis id="B9781929FFC7AE18FD8AAAA1CE64FBDC" box="[619,788,1012,1037]" italics="true" pageId="18" pageNumber="219">Tomaspisinella</emphasis>
</taxonomicName>
, but with the postpedicel large, almost as wide as pedicel (
<figureCitation id="1337D9BEFFC7AE18FF7EAD4DCD82FBE1" box="[159,242,1048,1073]" captionStart="FIGURES 14 15" captionStartId="32.[151,264,1884,1906]" captionTargetBox="[237,1347,193,1862]" captionTargetId="figure@32.[237,1347,193,1862]" captionTargetPageId="32" captionText="FIGURES 14 15. Menytes gen. nov. (14), compared to Simorhina Jacobi (15). A, habitus; B, profile; C, hind wing; D, venter (restoration) with photograph of actual specimen (inset); E, left half of head, anterior aspect; F, enlargement of tip of antenna (scale bar: 100 F), widest aspect, showing length of basiconic sensillum (b) and width compared to arista (a); G, tip of postpedicel, with arista to left and basiconic sensillum to right; H, enlargement of antennal pits (scale bar: 10 F) showing spiral base of each vertically pleated coeloconic sensillum (dark centre). Original photograph (inset in 12 D) provided by J. Boone." httpUri="https://zenodo.org/record/262590/files/figure.png" pageId="18" pageNumber="219">Fig. 14</figureCitation>
G), entirely exposed and with very many large pits occupying the whole surface with coeloconic sensilla having unique spiral bases (
<figureCitation id="1337D9BEFFC7AE18FE2FAD69CF6FFB85" box="[462,543,1084,1109]" captionStart="FIGURES 14 15" captionStartId="32.[151,264,1884,1906]" captionTargetBox="[237,1347,193,1862]" captionTargetId="figure@32.[237,1347,193,1862]" captionTargetPageId="32" captionText="FIGURES 14 15. Menytes gen. nov. (14), compared to Simorhina Jacobi (15). A, habitus; B, profile; C, hind wing; D, venter (restoration) with photograph of actual specimen (inset); E, left half of head, anterior aspect; F, enlargement of tip of antenna (scale bar: 100 F), widest aspect, showing length of basiconic sensillum (b) and width compared to arista (a); G, tip of postpedicel, with arista to left and basiconic sensillum to right; H, enlargement of antennal pits (scale bar: 10 F) showing spiral base of each vertically pleated coeloconic sensillum (dark centre). Original photograph (inset in 12 D) provided by J. Boone." httpUri="https://zenodo.org/record/262590/files/figure.png" pageId="18" pageNumber="219">Fig. 14</figureCitation>
H). Pronotum with anterior margin steeply declivous (
<figureCitation id="1337D9BEFFC7AE18FB9AAD69C9BDFB85" box="[1147,1229,1084,1109]" captionStart="FIGURES 14 15" captionStartId="32.[151,264,1884,1906]" captionTargetBox="[237,1347,193,1862]" captionTargetId="figure@32.[237,1347,193,1862]" captionTargetPageId="32" captionText="FIGURES 14 15. Menytes gen. nov. (14), compared to Simorhina Jacobi (15). A, habitus; B, profile; C, hind wing; D, venter (restoration) with photograph of actual specimen (inset); E, left half of head, anterior aspect; F, enlargement of tip of antenna (scale bar: 100 F), widest aspect, showing length of basiconic sensillum (b) and width compared to arista (a); G, tip of postpedicel, with arista to left and basiconic sensillum to right; H, enlargement of antennal pits (scale bar: 10 F) showing spiral base of each vertically pleated coeloconic sensillum (dark centre). Original photograph (inset in 12 D) provided by J. Boone." httpUri="https://zenodo.org/record/262590/files/figure.png" pageId="18" pageNumber="219">Fig. 14</figureCitation>
B), anterior border weakly bowed; lateral margins more than half length of eyes. Tegmina elongate, hirsute, with obscure veins. Hind wing with triangular basal lobe bearing an erect hook at tip and 2 recumbent smaller hooks on distal edge; straight veins defining 4 apical cells; appendix narrowest around tip of wing (
<figureCitation id="1337D9BEFFC7AE18FC25ADFDC96AFB11" box="[964,1050,1192,1217]" captionStart="FIGURES 14 15" captionStartId="32.[151,264,1884,1906]" captionTargetBox="[237,1347,193,1862]" captionTargetId="figure@32.[237,1347,193,1862]" captionTargetPageId="32" captionText="FIGURES 14 15. Menytes gen. nov. (14), compared to Simorhina Jacobi (15). A, habitus; B, profile; C, hind wing; D, venter (restoration) with photograph of actual specimen (inset); E, left half of head, anterior aspect; F, enlargement of tip of antenna (scale bar: 100 F), widest aspect, showing length of basiconic sensillum (b) and width compared to arista (a); G, tip of postpedicel, with arista to left and basiconic sensillum to right; H, enlargement of antennal pits (scale bar: 10 F) showing spiral base of each vertically pleated coeloconic sensillum (dark centre). Original photograph (inset in 12 D) provided by J. Boone." httpUri="https://zenodo.org/record/262590/files/figure.png" pageId="18" pageNumber="219">Fig. 14</figureCitation>
C). Fore femora subequal to hind femora. Hind tibiae with 2 almost equally long spines on outer edge (
<figureCitation id="1337D9BEFFC7AE18FC7CAD99CE9EFB35" box="[925,1006,1228,1253]" captionStart="FIGURES 14 15" captionStartId="32.[151,264,1884,1906]" captionTargetBox="[237,1347,193,1862]" captionTargetId="figure@32.[237,1347,193,1862]" captionTargetPageId="32" captionText="FIGURES 14 15. Menytes gen. nov. (14), compared to Simorhina Jacobi (15). A, habitus; B, profile; C, hind wing; D, venter (restoration) with photograph of actual specimen (inset); E, left half of head, anterior aspect; F, enlargement of tip of antenna (scale bar: 100 F), widest aspect, showing length of basiconic sensillum (b) and width compared to arista (a); G, tip of postpedicel, with arista to left and basiconic sensillum to right; H, enlargement of antennal pits (scale bar: 10 F) showing spiral base of each vertically pleated coeloconic sensillum (dark centre). Original photograph (inset in 12 D) provided by J. Boone." httpUri="https://zenodo.org/record/262590/files/figure.png" pageId="18" pageNumber="219">Fig. 14</figureCitation>
D), basal one 0.2 × and distal one 0.25 × as long as tibia; hind basitarsal pecten of 6 black-tipped spines, that of second tarsomere with 11 such spines. Male pygofer elongate, deeply notched dorsally on each side of anal tube; subgenital plates oblique, with decurved tips (
<figureCitation id="1337D9BEFFC7AE18FF31AC6DCC52FA81" box="[208,290,1336,1361]" captionStart="FIGURES 25 35" captionStartId="38.[151,264,1734,1756]" captionTargetBox="[161,1433,223,1706]" captionTargetId="figure@38.[151,1436,211,1711]" captionTargetPageId="38" captionText="FIGURES 25 35. Neaenini, Menytes (25) and Tomaspisinella (26 35): 25 A B, genital capsule and genitalia of Menytes lathrius, lateral aspect; 25 C, same, tip of theca in caudal aspect; 25 D, subgenital plates, ventral aspect; 26, head and thorax of Tomaspisinella (s. s.) caligata; 27, same, of T. (Ovotomaspis) minuscula; 28 35, Tomaspisinella spp., with genital capsule and style (A), aedeagus in lateral aspect (B), tip of aedeagus in caudal aspect (C), subgenital plates in ventral aspect (D): 28, T. (s. s.) parva; 29, T. (s. s.) lucifer; 30, T. (Meretricula) punctatissima; 31, T. (Meretricula) oliveirai; 32, T. (Meretricula) pallidiceps; 33, T. (Ovotomaspis) diabolus sp. nov.; 34, T. (Merinx) jocosa; 35, T. (Merinx) bolivari. Only genital illustrations to scale, with 28 C and 35 at larger scale than others, as indicated by scale lines." httpUri="https://zenodo.org/record/262595/files/figure.png" pageId="18" pageNumber="219">Fig. 27</figureCitation>
AB) separated by V-shaped notch (
<figureCitation id="1337D9BEFFC7AE18FD21AC6DCE64FA81" box="[704,788,1336,1361]" captionStart="FIGURES 25 35" captionStartId="38.[151,264,1734,1756]" captionTargetBox="[161,1433,223,1706]" captionTargetId="figure@38.[151,1436,211,1711]" captionTargetPageId="38" captionText="FIGURES 25 35. Neaenini, Menytes (25) and Tomaspisinella (26 35): 25 A B, genital capsule and genitalia of Menytes lathrius, lateral aspect; 25 C, same, tip of theca in caudal aspect; 25 D, subgenital plates, ventral aspect; 26, head and thorax of Tomaspisinella (s. s.) caligata; 27, same, of T. (Ovotomaspis) minuscula; 28 35, Tomaspisinella spp., with genital capsule and style (A), aedeagus in lateral aspect (B), tip of aedeagus in caudal aspect (C), subgenital plates in ventral aspect (D): 28, T. (s. s.) parva; 29, T. (s. s.) lucifer; 30, T. (Meretricula) punctatissima; 31, T. (Meretricula) oliveirai; 32, T. (Meretricula) pallidiceps; 33, T. (Ovotomaspis) diabolus sp. nov.; 34, T. (Merinx) jocosa; 35, T. (Merinx) bolivari. Only genital illustrations to scale, with 28 C and 35 at larger scale than others, as indicated by scale lines." httpUri="https://zenodo.org/record/262595/files/figure.png" pageId="18" pageNumber="219">Fig. 27</figureCitation>
D); styles truncate at base, tapered to sharp tip extending nearly as far as subgenital plates; phallobase tubular; theca laterally compressed except for bulbous tip (
<figureCitation id="1337D9BEFFC7AE18FAC9AC09C80BFAA5" box="[1320,1403,1372,1397]" captionStart="FIGURES 25 35" captionStartId="38.[151,264,1734,1756]" captionTargetBox="[161,1433,223,1706]" captionTargetId="figure@38.[151,1436,211,1711]" captionTargetPageId="38" captionText="FIGURES 25 35. Neaenini, Menytes (25) and Tomaspisinella (26 35): 25 A B, genital capsule and genitalia of Menytes lathrius, lateral aspect; 25 C, same, tip of theca in caudal aspect; 25 D, subgenital plates, ventral aspect; 26, head and thorax of Tomaspisinella (s. s.) caligata; 27, same, of T. (Ovotomaspis) minuscula; 28 35, Tomaspisinella spp., with genital capsule and style (A), aedeagus in lateral aspect (B), tip of aedeagus in caudal aspect (C), subgenital plates in ventral aspect (D): 28, T. (s. s.) parva; 29, T. (s. s.) lucifer; 30, T. (Meretricula) punctatissima; 31, T. (Meretricula) oliveirai; 32, T. (Meretricula) pallidiceps; 33, T. (Ovotomaspis) diabolus sp. nov.; 34, T. (Merinx) jocosa; 35, T. (Merinx) bolivari. Only genital illustrations to scale, with 28 C and 35 at larger scale than others, as indicated by scale lines." httpUri="https://zenodo.org/record/262595/files/figure.png" pageId="18" pageNumber="219">Fig. 27</figureCitation>
C), shaft sinuate, unarmed (
<figureCitation id="1337D9BEFFC7AE18FE42ACD5CC84FA49" box="[419,500,1408,1433]" captionStart="FIGURES 25 35" captionStartId="38.[151,264,1734,1756]" captionTargetBox="[161,1433,223,1706]" captionTargetId="figure@38.[151,1436,211,1711]" captionTargetPageId="38" captionText="FIGURES 25 35. Neaenini, Menytes (25) and Tomaspisinella (26 35): 25 A B, genital capsule and genitalia of Menytes lathrius, lateral aspect; 25 C, same, tip of theca in caudal aspect; 25 D, subgenital plates, ventral aspect; 26, head and thorax of Tomaspisinella (s. s.) caligata; 27, same, of T. (Ovotomaspis) minuscula; 28 35, Tomaspisinella spp., with genital capsule and style (A), aedeagus in lateral aspect (B), tip of aedeagus in caudal aspect (C), subgenital plates in ventral aspect (D): 28, T. (s. s.) parva; 29, T. (s. s.) lucifer; 30, T. (Meretricula) punctatissima; 31, T. (Meretricula) oliveirai; 32, T. (Meretricula) pallidiceps; 33, T. (Ovotomaspis) diabolus sp. nov.; 34, T. (Merinx) jocosa; 35, T. (Merinx) bolivari. Only genital illustrations to scale, with 28 C and 35 at larger scale than others, as indicated by scale lines." httpUri="https://zenodo.org/record/262595/files/figure.png" pageId="18" pageNumber="219">Fig. 27</figureCitation>
AB).
</paragraph>
</subSubSection>
<subSubSection id="C31696B0FFC7AE18FF26ACF1CCB3F8BC" pageId="18" pageNumber="219" type="discussion">
<paragraph id="8BB3C53BFFC7AE18FF26ACF1CC3DF90D" blockId="18.[151,1437,760,1793]" pageId="18" pageNumber="219">
<emphasis id="B9781929FFC7AE18FF26ACF1CC4BFA6C" bold="true" box="[199,315,1444,1469]" pageId="18" pageNumber="219">Remarks.</emphasis>
The spiral bases of the coeloconic sensilla set in large pits are unique. The sulcate face, concave crown and arched pronotum of
<taxonomicName id="4C0CBEB8FFC7AE18FDE9AC9CCF16FA31" box="[520,614,1481,1504]" class="Insecta" family="Cercopidae" genus="Menytes" kingdom="Animalia" order="Hemiptera" pageId="18" pageNumber="247" phylum="Arthropoda" rank="genus" status="gen. nov.">
<emphasis id="B9781929FFC7AE18FDE9AC9CCF16FA31" box="[520,614,1481,1504]" italics="true" pageId="18" pageNumber="219">Menytes</emphasis>
</taxonomicName>
are synapomorphies with
<taxonomicName id="4C0CBEB8FFC7AE18FC7AAC9DC97FFA30" box="[923,1039,1480,1505]" class="Insecta" family="Cercopidae" genus="Simorhina" kingdom="Animalia" order="Hemiptera" pageId="18" pageNumber="219" phylum="Arthropoda" rank="genus">
<emphasis id="B9781929FFC7AE18FC7AAC9DC97FFA30" box="[923,1039,1480,1505]" italics="true" pageId="18" pageNumber="219">Simorhina</emphasis>
</taxonomicName>
(
<figureCitation id="1337D9BEFFC7AE18FBC3AC9DC907FA30" box="[1058,1143,1480,1505]" captionStart="FIGURES 14 15" captionStartId="32.[151,264,1884,1906]" captionTargetBox="[237,1347,193,1862]" captionTargetId="figure@32.[237,1347,193,1862]" captionTargetPageId="32" captionText="FIGURES 14 15. Menytes gen. nov. (14), compared to Simorhina Jacobi (15). A, habitus; B, profile; C, hind wing; D, venter (restoration) with photograph of actual specimen (inset); E, left half of head, anterior aspect; F, enlargement of tip of antenna (scale bar: 100 F), widest aspect, showing length of basiconic sensillum (b) and width compared to arista (a); G, tip of postpedicel, with arista to left and basiconic sensillum to right; H, enlargement of antennal pits (scale bar: 10 F) showing spiral base of each vertically pleated coeloconic sensillum (dark centre). Original photograph (inset in 12 D) provided by J. Boone." httpUri="https://zenodo.org/record/262590/files/figure.png" pageId="18" pageNumber="219">Fig. 15</figureCitation>
B). Synapomorphies with
<taxonomicName id="4C0CBEB8FFC7AE18FF76ACB9CC30F9D4" box="[151,320,1516,1541]" class="Insecta" family="Cercopidae" genus="Tomaspisinella" kingdom="Animalia" order="Hemiptera" pageId="18" pageNumber="219" phylum="Arthropoda" rank="genus">
<emphasis id="B9781929FFC7AE18FF76ACB9CC30F9D4" box="[151,320,1516,1541]" italics="true" pageId="18" pageNumber="219">Tomaspisinella</emphasis>
</taxonomicName>
include strongly hirsute body (
<figureCitation id="1337D9BEFFC7AE18FD41ACB9CF81F9D5" box="[672,753,1516,1541]" captionStart="FIGURES 17 19" captionStartId="34.[151,264,1878,1900]" captionTargetBox="[180,1422,193,1855]" captionTargetId="figure@34.[180,1427,193,1856]" captionTargetPageId="34" captionText="FIGURES 17 19. Antennal characters of apomorphic new-world Cercopinae: Olcotomaspis (Phymatostethini, 17), Zuata (Neaenini, 18) and Tomaspisella (Neaenini, 19): 17 A, Olcotomaspis versicolor (Lallemand); 17 B, same, from another angle, showing paired basiconic sensillae on inner side; 17 C, O. laterinotata (Fowler); 17 D, O. (= Hyalotomaspis) clarissa; 18 A, Zuata infecta sp. nov.; 18 B, same, detail (note peglike seta surrounded by large socket and tiny secondary seta behind it); 19 A, Tomaspisella parva Lallemand, male; 19 B, same, female; 19 C, same, detail of pits." httpUri="https://zenodo.org/record/262592/files/figure.png" pageId="18" pageNumber="219">Fig. 18</figureCitation>
B) and elongate pygofers (
<bibRefCitation id="EF9DB8CAFFC7AE18FBFCACB9CDD7F9F9" author="Carvalho" pageId="18" pageNumber="219" refString="Carvalho, G. S. &amp; Webb, M. D. (2005) Cercopid Spittle Bugs of the New World (Hemiptera, Auchenorrhyncha, Cercopidae). Pensoft, Sofia-Moscow, 271 pp." type="book" year="2005">Carvalho &amp; Webb 2005, figs 4, 5, 8</bibRefCitation>
).
<taxonomicName id="4C0CBEB8FFC7AE18FF20AF45CC1AF9F8" box="[193,362,1552,1577]" class="Insecta" family="Cercopidae" genus="Tomaspisinella" kingdom="Animalia" order="Hemiptera" pageId="18" pageNumber="219" phylum="Arthropoda" rank="genus">
<emphasis id="B9781929FFC7AE18FF20AF45CC1AF9F8" box="[193,362,1552,1577]" italics="true" pageId="18" pageNumber="219">Tomaspisinella</emphasis>
</taxonomicName>
have the postpedicel retracted into the apical cup of the pedicel, while the postpedicel of
<taxonomicName id="4C0CBEB8FFC7AE18FF76AF61CC7BF99C" box="[151,267,1588,1613]" class="Insecta" family="Cercopidae" genus="Simorhina" kingdom="Animalia" order="Hemiptera" pageId="18" pageNumber="219" phylum="Arthropoda" rank="genus">
<emphasis id="B9781929FFC7AE18FF76AF61CC7BF99C" box="[151,267,1588,1613]" italics="true" pageId="18" pageNumber="219">Simorhina</emphasis>
</taxonomicName>
represents a more plesiomorphic condition (
<figureCitation id="1337D9BEFFC7AE18FCA0AF61CEECF99C" box="[833,924,1588,1613]" captionStart="FIGURES 14 15" captionStartId="32.[151,264,1884,1906]" captionTargetBox="[237,1347,193,1862]" captionTargetId="figure@32.[237,1347,193,1862]" captionTargetPageId="32" captionText="FIGURES 14 15. Menytes gen. nov. (14), compared to Simorhina Jacobi (15). A, habitus; B, profile; C, hind wing; D, venter (restoration) with photograph of actual specimen (inset); E, left half of head, anterior aspect; F, enlargement of tip of antenna (scale bar: 100 F), widest aspect, showing length of basiconic sensillum (b) and width compared to arista (a); G, tip of postpedicel, with arista to left and basiconic sensillum to right; H, enlargement of antennal pits (scale bar: 10 F) showing spiral base of each vertically pleated coeloconic sensillum (dark centre). Original photograph (inset in 12 D) provided by J. Boone." httpUri="https://zenodo.org/record/262590/files/figure.png" pageId="18" pageNumber="219">Fig. 15</figureCitation>
G). Some other distinctive characters are synapomorphic with allied genera or homoplastic with unrelated spittlebugs (for example, the depressed crown is also characteristic of
<taxonomicName id="4C0CBEB8FFC7AE18FE66AF29CF74F944" box="[391,516,1660,1685]" class="Insecta" family="Cercopidae" genus="Liorhinella" kingdom="Animalia" order="Hemiptera" pageId="18" pageNumber="219" phylum="Arthropoda" rank="genus">
<emphasis id="B9781929FFC7AE18FE66AF29CF74F944" box="[391,516,1660,1685]" italics="true" pageId="18" pageNumber="219">Liorhinella</emphasis>
</taxonomicName>
, and the wedge-shaped style is similar to one found in an undescribed Epipygid). Phylogenetically,
<taxonomicName id="4C0CBEB8FFC7AE18FE88AFF4CCB7F969" box="[361,455,1697,1720]" class="Insecta" family="Cercopidae" genus="Menytes" kingdom="Animalia" order="Hemiptera" pageId="18" pageNumber="247" phylum="Arthropoda" rank="genus" status="gen. nov.">
<emphasis id="B9781929FFC7AE18FE88AFF4CCB7F969" box="[361,455,1697,1720]" italics="true" pageId="18" pageNumber="219">Menytes</emphasis>
</taxonomicName>
represents a sister-lineage of
<taxonomicName id="4C0CBEB8FFC7AE18FCA6AFF5CECBF968" box="[839,955,1696,1721]" class="Insecta" family="Cercopidae" genus="Simorhina" kingdom="Animalia" order="Hemiptera" pageId="18" pageNumber="219" phylum="Arthropoda" rank="genus">
<emphasis id="B9781929FFC7AE18FCA6AFF5CECBF968" box="[839,955,1696,1721]" italics="true" pageId="18" pageNumber="219">Simorhina</emphasis>
</taxonomicName>
linking that autapomorphous genus to
<taxonomicName id="4C0CBEB8FFC7AE18FF76AF91CC30F90C" box="[151,320,1732,1757]" class="Insecta" family="Cercopidae" genus="Tomaspisinella" kingdom="Animalia" order="Hemiptera" pageId="18" pageNumber="219" phylum="Arthropoda" rank="genus">
<emphasis id="B9781929FFC7AE18FF76AF91CC30F90C" box="[151,320,1732,1757]" italics="true" pageId="18" pageNumber="219">Tomaspisinella</emphasis>
</taxonomicName>
.
</paragraph>
<paragraph id="8BB3C53BFFC7AE18FF26AFBDCFA1F8D0" blockId="18.[151,1437,760,1793]" box="[199,721,1768,1793]" pageId="18" pageNumber="219">There is only one known species in this genus.</paragraph>
<paragraph id="8BB3C53BFFC7AE18FF76AE01CCB3F8BC" blockId="18.[151,451,1876,1902]" box="[151,451,1876,1902]" pageId="18" pageNumber="219">
<taxonomicName id="4C0CBEB8FFC7AE18FF76AE01CC12F8BF" box="[151,354,1876,1902]" class="Insecta" family="Cercopidae" genus="Menytes" kingdom="Animalia" order="Hemiptera" pageId="18" pageNumber="219" phylum="Arthropoda" rank="species" species="lathrius" status="sp. nov.">
<emphasis id="B9781929FFC7AE18FF76AE01CC12F8BF" bold="true" box="[151,354,1876,1902]" italics="true" pageId="18" pageNumber="219">Menytes lathrius</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A24BA452FFC7AE18FE88AE01CCB3F8BC" box="[361,451,1876,1901]" pageId="18" pageNumber="219" rank="species">
<emphasis id="B9781929FFC7AE18FE88AE01CCB3F8BC" bold="true" box="[361,451,1876,1901]" pageId="18" pageNumber="219">sp. nov.</emphasis>
</taxonomicNameLabel>
</paragraph>
</subSubSection>
<subSubSection id="C31696B0FFC7AE18FF76AECFCF19F862" box="[151,617,1946,1971]" pageId="18" pageNumber="219" type="etymology">
<paragraph id="8BB3C53BFFC7AE18FF76AECFCF19F862" blockId="18.[151,1436,1946,2008]" box="[151,617,1946,1971]" pageId="18" pageNumber="219">
<emphasis id="B9781929FFC7AE18FF76AECFCC6EF862" bold="true" box="[151,286,1946,1971]" pageId="18" pageNumber="219">Etymology.</emphasis>
<emphasis id="B9781929FFC7AE18FEC5AECFCC0BF862" box="[292,379,1946,1971]" italics="true" pageId="18" pageNumber="219">lathrios</emphasis>
(adj.) secret, hidden.
</paragraph>
</subSubSection>
<subSubSection id="C31696B0FFC7AE19FF26AEEBC932FF04" lastPageId="19" lastPageNumber="220" pageId="18" pageNumber="219" type="description">
<paragraph id="8BB3C53BFFC7AE19FF26AEEBC932FF04" blockId="18.[151,1436,1946,2008]" lastBlockId="19.[151,1436,151,285]" lastPageId="19" lastPageNumber="220" pageId="18" pageNumber="219">
<emphasis id="B9781929FFC7AE18FF26AEEBCC29F806" bold="true" box="[199,345,1982,2007]" pageId="18" pageNumber="219">Description.</emphasis>
Brown, antennal ledges tawny; venter marked with yellow on tip of frons, black on rest of face, pro- and mesopleura and tarsi; tegmina with broken, ivory band across apical quarter extending from costal dash to a chain of 23 small spots on each tegmen. Length: male
<quantity id="4CF468DEFFC6AE19FCFAA9E9CE04FF05" box="[795,884,188,213]" metricMagnitude="-3" metricUnit="m" metricValue="6.7" pageId="19" pageNumber="220" unit="mm" value="6.7">6.7 mm</quantity>
; female unknown.
</paragraph>
</subSubSection>
<subSubSection id="C31696B0FFC6AE19FF26A98ACC2FFECD" pageId="19" pageNumber="220" type="materials_examined">
<paragraph id="8BB3C53BFFC6AE19FF26A98ACC2FFECD" blockId="19.[151,1436,151,285]" pageId="19" pageNumber="220">
<materialsCitation id="3B64CF66FFC6AE19FF26A98ACC2FFECD" ID-GBIF-Occurrence="1313797421" collectingDate="1971-05-15" collectingDateMax="1971-05-31" collectingDateMin="1971-05-15" collectionCode="FMC" collectorName="B. Malkin" country="Ecuador" location="Rio Cusuimi" municipality="Cusuimi" pageId="19" pageNumber="220" specimenCount="2" stateProvince="Pastaza" typeStatus="holotype">
<emphasis id="B9781929FFC6AE19FF26A98ACC78FF29" bold="true" box="[199,264,223,248]" pageId="19" pageNumber="220">
<typeStatus id="54B77B99FFC6AE19FF26A98ACC74FF29" box="[199,260,223,248]" pageId="19" pageNumber="220">Type</typeStatus>
.
</emphasis>
<typeStatus id="54B77B99FFC6AE19FEF0A98ACC09FF29" box="[273,377,223,248]" pageId="19" pageNumber="220" type="holotype">Holotype</typeStatus>
male,
<collectingCountry id="F31B85ABFFC6AE19FE27A9B5CF22FF29" box="[454,594,224,248]" name="Ecuador" pageId="19" pageNumber="220">ECUADOR</collectingCountry>
:
<collectingRegion id="49C80BD9FFC6AE19FDBFA9B4CFC7FF29" box="[606,695,225,248]" country="Ecuador" name="Pastaza" pageId="19" pageNumber="220">
<emphasis id="B9781929FFC6AE19FDBFA9B4CFC7FF29" box="[606,695,225,248]" italics="true" pageId="19" pageNumber="220">Pastaza</emphasis>
</collectingRegion>
<collectingMunicipality id="6BD75F41FFC6AE19FD35A98ACE45FF29" box="[724,821,223,248]" pageId="19" pageNumber="220">Cusuimi</collectingMunicipality>
,
<location id="8ED393E0FFC6AE19FCA0A98ACEA3FF29" LSID="urn:lsid:plazi:treatment:03A5742DFFC7AE19FF76AB3BCC2FFECD:8ED393E0FFC6AE19FCA0A98ACEA3FF29" box="[833,979,223,248]" country="Ecuador" municipality="Cusuimi" name="Rio Cusuimi" pageId="19" pageNumber="220" stateProvince="Pastaza">Rio Cusuimi</location>
,
<locationDeviation id="9A01DC19FFC6AE19FC3EA98AC9D2FF29" box="[991,1186,223,248]" pageId="19" pageNumber="220">
<quantity id="4CF468DEFFC6AE19FC3EA98AC946FF29" box="[991,1078,223,248]" metricMagnitude="5" metricUnit="m" metricValue="1.5" pageId="19" pageNumber="220" unit="km" value="150.0">150 km</quantity>
SE Puyo
</locationDeviation>
,
<collectingDate id="EFF61A13FFC6AE19FB4EA98AC801FF29" box="[1199,1393,223,248]" pageId="19" pageNumber="220" value="1971-05-15" valueMax="1971-05-31" valueMin="1971-05-15">
<date id="FFB2E3FBFFC6AE19FB4EA98AC801FF29" box="[1199,1393,223,248]" pageId="19" pageNumber="220" value="1971-05-15" valueMax="1971-05-31" valueMin="1971-05-15">1531 May 1971</date>
</collectingDate>
(
<collectorName id="26F9A0EDFFC6AE19FA62A9B5CD9DFECC" pageId="19" pageNumber="220">B. Malkin</collectorName>
); in
<collectionCode id="ED1D5DFEFFC6AE19FEFCA851CC2CFECD" box="[285,348,260,284]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:13060" name="North Museum of Natural History and Science" pageId="19" pageNumber="220">FMC</collectionCode>
.
</materialsCitation>
</paragraph>
</subSubSection>
</treatment>
</document>