treatments-xml/data/03/AC/87/03AC87D0B119FF93FF4690CDFB278BF5.xml
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<document id="4DF7549B2A4ACD474B1657A289E4992E" ID-DOI="10.5281/zenodo.176960" ID-GBIF-Dataset="a67acf3d-f43a-4d16-b437-aacf9f0d09ec" ID-ISSN="1175-5326" ID-Zenodo-Dep="176960" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1459844304451" checkinUser="plazi" docAuthor="Ziegler, Thomas, Hendrix, Ralf, Thanh, Vu Ngoc, Vogt, Martina, Forster, Bernhard &amp; Kien, Dang Ngoc" docDate="2007" docId="03AC87D0B119FF93FF4690CDFB278BF5" docLanguage="en" docName="zt01493p040.pdf" docOrigin="Zootaxa 1493" docStyle="DocumentStyle:6581673A57F01A3145754A1E615EDFF0.4:Zootaxa.2007-2008.journal_article" docStyleId="6581673A57F01A3145754A1E615EDFF0" docStyleName="Zootaxa.2007-2008.journal_article" docStyleVersion="4" docTitle="Lycodon" docType="treatment" docVersion="9" lastPageNumber="14" masterDocId="FF95FFA8B115FF9EFFD19607FFD28827" masterDocTitle="The diversity of a snake community in a karst forest ecosystem in the central Truong Son, Vietnam, with an identification key" masterLastPageNumber="40" masterPageNumber="1" pageNumber="13" updateTime="1698224724941" updateUser="plazi">
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<mods:title id="CAC4D8992B508281AE2A3710E483E71A">The diversity of a snake community in a karst forest ecosystem in the central Truong Son, Vietnam, with an identification key</mods:title>
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<mods:namePart id="CA0DCB8035737916BBD294F69A051E47">Ziegler, Thomas</mods:namePart>
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<mods:namePart id="B91C3B7EB058A9B18A0C5D10ED8D04AB">Hendrix, Ralf</mods:namePart>
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<mods:namePart id="84D70066E2FD60B211EE3BCE312288F5">Thanh, Vu Ngoc</mods:namePart>
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<mods:namePart id="31360186C70C8B907C1226B067D47B65">Vogt, Martina</mods:namePart>
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<mods:namePart id="665A458BFAB2625A3310D55A19CDF3D0">Forster, Bernhard</mods:namePart>
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<mods:namePart id="AFAFA490E91EC4FD52A30C2A5D30FEB3">Kien, Dang Ngoc</mods:namePart>
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<mods:date id="A7778C2D59E8EAF0CAA4B7A4545140C7">2007</mods:date>
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<mods:identifier id="98F19AFD599EADE956C7467991AAF5B3" type="DOI">10.5281/zenodo.176960</mods:identifier>
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<treatment id="03AC87D0B119FF93FF4690CDFB278BF5" ID-DOI="http://doi.org/10.5281/zenodo.6247459" ID-GBIF-Taxon="119361337" ID-Zenodo-Dep="6247459" LSID="urn:lsid:plazi:treatment:03AC87D0B119FF93FF4690CDFB278BF5" httpUri="http://treatment.plazi.org/id/03AC87D0B119FF93FF4690CDFB278BF5" lastPageId="13" lastPageNumber="14" pageId="12" pageNumber="13">
<subSubSection id="C31F654DB119FF93FF4690CDFED18AB5" lastPageId="13" lastPageNumber="14" pageId="12" pageNumber="13" type="nomenclature">
<paragraph id="8BBA36C6B119FF92FF4690CDFEFB8EC4" blockId="12.[151,297,1737,1764]" box="[151,297,1737,1764]" pageId="12" pageNumber="13">
<heading id="D0F281AAB119FF92FF4690CDFEFB8EC4" bold="true" box="[151,297,1737,1764]" fontSize="11" level="1" pageId="12" pageNumber="13" reason="1">
<emphasis id="B971EAD4B119FF92FF4690CDFEFB8EC4" bold="true" box="[151,297,1737,1764]" pageId="12" pageNumber="13">
<taxonomicName id="4C054D45B119FF92FF4690CDFF2C8EC3" box="[151,254,1738,1764]" class="Reptilia" family="Colubridae" genus="Lycodon" kingdom="Animalia" order="Squamata" pageId="12" pageNumber="13" phylum="Chordata" rank="genus">
<emphasis id="B971EAD4B119FF92FF4690CDFF2C8EC3" bold="true" box="[151,254,1738,1764]" italics="true" pageId="12" pageNumber="13">Lycodon</emphasis>
</taxonomicName>
sp.
</emphasis>
</heading>
</paragraph>
<paragraph id="8BBA36C6B119FF93FF46911DFED18AB5" blockId="12.[151,1436,1818,2004]" lastBlockId="13.[151,1437,152,978]" lastPageId="13" lastPageNumber="14" pageId="12" pageNumber="13">
Another female
<taxonomicName id="4C054D45B119FF92FE84911DFE6B8F14" box="[341,441,1818,1843]" class="Reptilia" family="Colubridae" genus="Lycodon" kingdom="Animalia" order="Squamata" pageId="12" pageNumber="13" phylum="Chordata" rank="genus">
<emphasis id="B971EAD4B119FF92FE84911DFE6B8F14" box="[341,441,1818,1843]" italics="true" pageId="12" pageNumber="13">Lycodon</emphasis>
</taxonomicName>
was collected by us about one month after the discovery of the afore mentioned
<taxonomicName id="4C054D45B119FF92FAB8911DFEB78F7C" class="Reptilia" family="Colubridae" genus="Lycodon" kingdom="Animalia" order="Squamata" pageId="12" pageNumber="13" phylum="Chordata" rank="species" species="ruhstrati">
<emphasis id="B971EAD4B119FF92FAB8911DFF008F7C" italics="true" pageId="12" pageNumber="13">Lycodon</emphasis>
cf.
<emphasis id="B971EAD4B119FF92FF2E9145FEB78F7C" box="[255,357,1858,1883]" italics="true" pageId="12" pageNumber="13">ruhstrati</emphasis>
</taxonomicName>
. The discovery took place in the same region at an altitude of about
<quantity id="4CFD9B23B119FF92FB5F9145FB0B8F7B" box="[1166,1241,1858,1884]" metricMagnitude="2" metricUnit="m" metricValue="4.8" pageId="12" pageNumber="13" unit="m" value="480.0">480 m</quantity>
above sea level. This second
<taxonomicName id="4C054D45B119FF92FEFB916DFE5C8FA4" box="[298,398,1898,1923]" class="Reptilia" family="Colubridae" genus="Lycodon" kingdom="Animalia" order="Squamata" pageId="12" pageNumber="13" phylum="Chordata" rank="genus">
<emphasis id="B971EAD4B119FF92FEFB916DFE5C8FA4" box="[298,398,1898,1923]" italics="true" pageId="12" pageNumber="13">Lycodon</emphasis>
</taxonomicName>
(
<figureCitation id="133E2A43B119FF92FE4C916DFDC28FA3" box="[413,528,1898,1924]" captionStart="FIGURE 13" captionStartId="12.[151,255,1663,1687]" captionTargetBox="[151,1434,698,1655]" captionTargetId="figure@12.[151,1436,695,1657]" captionTargetPageId="12" captionText="FIGURE 13. Dorsal view of Lycodon sp. from Phong Nha - Ke Bang (ZFMK 86453). Photo: Ralf Hendrix" httpUri="https://zenodo.org/record/176973/files/figure.png" pageId="12" pageNumber="13">Figure 13</figureCitation>
) was found at night of
<date id="FFBB1006B119FF92FCCE916DFC6B8FA3" box="[799,953,1898,1924]" pageId="12" pageNumber="13" value="2006-07-30">30 July 2006</date>
, at the end of the dry season, crawling on pebbles near a steep primary forest stream cascade in the U Bo region. The specimen (ZFMK 86453, SVL 505, TaL 147, TL
<quantity id="4CFD9B23B119FF92FEA791BDFE098FF3" box="[374,475,1978,2004]" metricMagnitude="-1" metricUnit="m" metricValue="6.52" pageId="12" pageNumber="13" unit="mm" value="652.0">652 mm</quantity>
) differs from the syntopic
<taxonomicName id="4C054D45B119FF92FCCC91BDFBCA8FF4" box="[797,1048,1978,2004]" class="Reptilia" family="Colubridae" genus="Lycodon" kingdom="Animalia" order="Squamata" pageId="12" pageNumber="13" phylum="Chordata" rank="species" species="ruhstrati">
<emphasis id="B971EAD4B119FF92FCCC91BDFC538FF4" box="[797,897,1978,2003]" italics="true" pageId="12" pageNumber="13">Lycodon</emphasis>
cf.
<emphasis id="B971EAD4B119FF92FC6391BDFBCA8FF4" box="[946,1048,1978,2003]" italics="true" pageId="12" pageNumber="13">ruhstrati</emphasis>
</taxonomicName>
among others in having smooth dorsal scale rows, bearing enlarged vertebral scales, and having somewhat lower ventral and subcaudal counts (although within the range of
<taxonomicName id="4C054D45B118FF93FE2E96C7FD5488FE" box="[511,646,192,217]" class="Reptilia" family="Colubridae" genus="Lycodon" kingdom="Animalia" order="Squamata" pageId="13" pageNumber="14" phylum="Chordata" rank="species" species="ruhstrati">
<emphasis id="B971EAD4B118FF93FE2E96C7FD5488FE" box="[511,646,192,217]" italics="true" pageId="13" pageNumber="14">L. ruhstrati</emphasis>
</taxonomicName>
, see above), as well as a differing banded pattern. The remaining scalation features of the specimen are as follows: eight supralabials, of which third to fifth are in contact with eye; 10 infralabials; one loreal, not in contact with eye; one preocular, not in contact with frontal; prefrontal not entering eye; two postoculars; two anterior temporals, followed by three posterior temporals; 17 dorsal scale rows at midbody; one undivided preventral and 208 ventrals; ventrals distinctly angulate laterally; anal entire; 85 divided subcaudals. Concerning coloration and pattern, the specimen has 20 dark transversal crossbands on its body dorsum, which continue on the venter (except for the neck band). The 19 creamy interspaces of the dark body crossbands are marbled by greyish-brown. Tail dorsum with 10 such light crossbands and 9 dark crossbars (including tail tip), underside of tail dark. On the right maxillary bone seven anterior fang-like teeth increasing in size were discernible, separated by a large toothless interspace from four medium-sized teeth, which were followed by three posterior teeth, the first two ones of which being distinctly enlarged.
</paragraph>
</subSubSection>
<subSubSection id="C31F654DB118FF93FF1794A7FB278BF5" pageId="13" pageNumber="14" type="description">
<paragraph id="8BBA36C6B118FF93FF1794A7FB278BF5" blockId="13.[151,1437,152,978]" pageId="13" pageNumber="14">
This
<taxonomicName id="4C054D45B118FF93FED394A7FEB48A9E" box="[258,358,672,697]" class="Reptilia" family="Colubridae" genus="Lycodon" kingdom="Animalia" order="Squamata" pageId="13" pageNumber="14" phylum="Chordata" rank="genus">
<emphasis id="B971EAD4B118FF93FED394A7FEB48A9E" box="[258,358,672,697]" italics="true" pageId="13" pageNumber="14">Lycodon</emphasis>
</taxonomicName>
specimen morphologically resembles the female
<taxonomicName id="4C054D45B118FF93FC7E94A7FBE78A9E" box="[943,1077,672,697]" class="Reptilia" family="Colubridae" genus="Lycodon" kingdom="Animalia" order="Squamata" pageId="13" pageNumber="14" phylum="Chordata" rank="species" species="ruhstrati">
<emphasis id="B971EAD4B118FF93FC7E94A7FBE78A9E" box="[943,1077,672,697]" italics="true" pageId="13" pageNumber="14">L. ruhstrati</emphasis>
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specimen from Ha Tinh province as shown in Ziegler (2002). However, the latter specimen has no enlarged vertebrals and bears minor differences in the banded pattern. The Ha Tinh specimen further shows a distinct light area on the rear head, however, that is still slightly discernible in the larger
<taxonomicName id="4C054D45B118FF93FCE0951FFC478B16" box="[817,917,792,817]" class="Reptilia" family="Colubridae" genus="Lycodon" kingdom="Animalia" order="Squamata" pageId="13" pageNumber="14" phylum="Chordata" rank="genus">
<emphasis id="B971EAD4B118FF93FCE0951FFC478B16" box="[817,917,792,817]" italics="true" pageId="13" pageNumber="14">Lycodon</emphasis>
</taxonomicName>
specimen from Phong Nha - Ke Bang as obscured area.
<collectingCountry id="F3127656B118FF93FE809547FEA38B7D" box="[337,369,832,858]" name="American Samoa" pageId="13" pageNumber="14">As</collectingCountry>
was pointed out above, only future research including currently undertaken molecular methods will prove, whether the mainland &quot;
<taxonomicName id="4C054D45B118FF93FD46956FFCCF8BA6" box="[663,797,872,897]" class="Reptilia" family="Colubridae" genus="Lycodon" kingdom="Animalia" order="Squamata" pageId="13" pageNumber="14" phylum="Chordata" rank="species" species="ruhstrati">
<emphasis id="B971EAD4B118FF93FD46956FFCCF8BA6" box="[663,797,872,897]" italics="true" pageId="13" pageNumber="14">L. ruhstrati</emphasis>
</taxonomicName>
&quot; with smooth dorsal scales in fact represent a separate taxon (cf.
<taxonomicName id="4C054D45B118FF93FEC99597FE148B8E" box="[280,454,912,937]" class="Reptilia" family="Colubridae" genus="Dinodon" kingdom="Animalia" order="Squamata" pageId="13" pageNumber="14" phylum="Chordata" rank="species" species="futsingensis">
<emphasis id="B971EAD4B118FF93FEC99597FE148B8E" box="[280,454,912,937]" italics="true" pageId="13" pageNumber="14">L. futsingensis</emphasis>
</taxonomicName>
) and whether mainland
<taxonomicName id="4C054D45B118FF93FD279597FC798B8E" box="[758,939,912,938]" class="Reptilia" family="Colubridae" genus="Lycodon" kingdom="Animalia" order="Squamata" pageId="13" pageNumber="14" phylum="Chordata" rank="species" species="ruhstrati">
<emphasis id="B971EAD4B118FF93FD279597FCDC8B8E" box="[758,782,912,937]" italics="true" pageId="13" pageNumber="14">L.</emphasis>
cf.
<emphasis id="B971EAD4B118FF93FC959597FC798B8E" box="[836,939,912,937]" italics="true" pageId="13" pageNumber="14">ruhstrati</emphasis>
</taxonomicName>
(with keeled dorsal scales) represent an unnamed taxon indeed, different from the populations from the terra typica in southern
<collectingCountry id="F3127656B118FF93FB4695BFFB238BF5" box="[1175,1265,952,978]" name="Taiwan" pageId="13" pageNumber="14">Taiwan</collectingCountry>
.
</paragraph>
</subSubSection>
</treatment>
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