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<document id="FDE0BF8492DC27E1692596EE173BC4D9" ID-CLB-Dataset="7420" ID-DOI="10.11646/zootaxa.4940.1.1" ID-GBIF-Dataset="a09df80c-5d8b-44da-8f25-723f107c6c49" ID-ISSN="1175-5326" ID-Zenodo-Dep="4580917" ID-ZooBank="5F073867-F6EA-4222-A190-77F68076BE9C" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1614844133957" checkinUser="plazi" docAuthor="Booysen, Ruan &amp; Haddad, Charles R." docDate="2021" docId="03AC33149C13FFE27CBDF9ACFB60FE89" docLanguage="en" docName="zootaxa.4940.1.1.pdf" docOrigin="Zootaxa 4940 (1)" docStyle="DocumentStyle:5EBBA59367AD13919D70D935FA04F6A3.14:Zootaxa.2013-.monograph" docStyleId="5EBBA59367AD13919D70D935FA04F6A3" docStyleName="Zootaxa.2013-.monograph" docStyleVersion="14" docTitle="Micaria bimaculata Booysen &amp; Haddad 2021, sp. nov." docType="treatment" docVersion="9" lastPageNumber="37" masterDocId="FF954B6C9C32FFC67C2AFFFCFFCEFFEC" masterDocTitle="Revision and molecular phylogeny of the spider genus Micaria Westring, 1851 (Araneae: Gnaphosidae) in the Afrotropical Region" masterLastPageNumber="82" masterPageNumber="1" pageNumber="34" updateTime="1698932415381" updateUser="ExternalLinkService">
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<mods:title id="0D2EB96235FDEC4CF8CA49E3BA28988C">Revision and molecular phylogeny of the spider genus Micaria Westring, 1851 (Araneae: Gnaphosidae) in the Afrotropical Region</mods:title>
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<mods:namePart id="00AA9E4738842E56BC56FC001E924604">Booysen, Ruan</mods:namePart>
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<mods:namePart id="D385D41AA171ED4DD48C02E2C6EF5D1F">Haddad, Charles R.</mods:namePart>
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<treatment id="03AC33149C13FFE27CBDF9ACFB60FE89" ID-DOI="http://doi.org/10.5281/zenodo.4681000" ID-GBIF-Taxon="180991653" ID-Zenodo-Dep="4681000" LSID="urn:lsid:plazi:treatment:03AC33149C13FFE27CBDF9ACFB60FE89" httpUri="http://treatment.plazi.org/id/03AC33149C13FFE27CBDF9ACFB60FE89" lastPageId="36" lastPageNumber="37" pageId="33" pageNumber="34">
<subSubSection id="C31FD1899C13FFE77CBDF9ACFE25F987" box="[151,491,1616,1643]" pageId="33" pageNumber="34" type="nomenclature">
<paragraph id="8BBA82029C13FFE77CBDF9ACFE25F987" blockId="33.[151,491,1616,1679]" box="[151,491,1616,1643]" pageId="33" pageNumber="34">
<heading id="D0F2356E9C13FFE77CBDF9ACFE25F987" bold="true" box="[151,491,1616,1643]" fontSize="11" level="2" pageId="33" pageNumber="34" reason="4">
<emphasis id="B9715E109C13FFE77CBDF9ACFE25F987" bold="true" box="[151,491,1616,1643]" pageId="33" pageNumber="34">
<taxonomicName id="4C05F9819C13FFE77CBDF9ACFE48F987" ID-CoL="98Z5R" authority="Booysen &amp; Haddad, 2021" authorityName="Booysen &amp; Haddad" authorityYear="2021" box="[151,390,1616,1643]" class="Arachnida" family="Gnaphosidae" genus="Micaria" kingdom="Animalia" order="Araneae" pageId="33" pageNumber="34" phylum="Arthropoda" rank="species" species="bimaculata" status="sp. nov.">
<emphasis id="B9715E109C13FFE77CBDF9ACFE48F987" bold="true" box="[151,390,1616,1643]" italics="true" pageId="33" pageNumber="34">Micariabimaculata</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A242E36B9C13FFE77DA7F9ADFE25F987" box="[397,491,1617,1643]" pageId="33" pageNumber="34" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C31FD1899C13FFE77CBDF988FE4DF963" box="[151,387,1652,1679]" pageId="33" pageNumber="34" type="description">
<paragraph id="8BBA82029C13FFE77CBDF988FE4DF963" blockId="33.[151,491,1616,1679]" box="[151,387,1652,1679]" pageId="33" pageNumber="34">
<figureCitation id="133E9E879C13FFE77CBDF988FED9F963" box="[151,279,1652,1679]" captionStart="FIGURES 917" captionStartId="11.[151,264,1943,1967]" captionTargetBox="[173,1415,181,1920]" captionTargetId="figure-17@11.[173,1415,181,1920]" captionTargetPageId="11" captionText="FIGURES 917. Stereomicroscope photographs of the dorsal habitus ofAfrotropical Micaria species: 9 M. basaliducta sp. nov. female and 10 male; 11 M. beaufortia (Tucker, 1923) female and 12 male; 13 M. bimaculata sp. nov. female and 14 male; 15 M. bispicula sp. nov. female and 16 male; 17 M. chrysis (Simon, 1910) female. Scale bars: 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.4580929" httpUri="https://zenodo.org/record/4580929/files/figure.png" pageId="33" pageNumber="34">Figs 1314</figureCitation>
,
<figureCitation id="133E9E879C13FFE77D08F989FE4DF963" box="[290,387,1653,1679]" captionStart="FIGURES 117120" captionStartId="34.[151,264,1244,1268]" captionTargetBox="[172,1406,462,1215]" captionTargetId="figure-156@34.[164,1423,453,1221]" captionTargetPageId="34" captionText="FIGURES 117120. Genital morphology of Micaria bimaculata sp. nov.: 117 female epigyne, ventral view; 118 female epigyne, dorsal view; 119 male palp, ventral view; 120 same, retrolateral view. Scale bars: 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.4580966" httpUri="https://zenodo.org/record/4580966/files/figure.png" pageId="33" pageNumber="34">117120</figureCitation>
</paragraph>
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<subSubSection id="C31FD1899C13FFE77CBDF94BFCBAF8F6" pageId="33" pageNumber="34" type="materials_examined">
<paragraph id="8BBA82029C13FFE77CBDF94BFB6CF91A" blockId="33.[151,1437,1719,2034]" pageId="33" pageNumber="34">
<emphasis id="B9715E109C13FFE77CBDF94BFE88F93D" bold="true" box="[151,326,1719,1745]" pageId="33" pageNumber="34">Type material:</emphasis>
<materialsCitation id="3B6D885F9C13FFE77D67F94BFB6CF91A" ID-GBIF-Occurrence="3071034393" collectingDate="1994-11" collectorName="F. Borgato" country="Mauritania" latitude="18.081034" location="South of Nouachott" longLatPrecision="1" longitude="-15.96625" pageId="33" pageNumber="34" specimenCount="1" specimenCount-female="1" stateProvince="Nouakchott" typeStatus="holotype">
<emphasis id="B9715E109C13FFE77D67F94BFE74F93D" bold="true" box="[333,442,1719,1745]" pageId="33" pageNumber="34">
<typeStatus id="54BE3CA09C13FFE77D67F94BFE74F93D" box="[333,442,1719,1745]" pageId="33" pageNumber="34" type="holotype">Holotype</typeStatus>
</emphasis>
:
<specimenCount id="9D03498B9C13FFE77DEDF944FE12F93E" box="[455,476,1720,1746]" pageId="33" pageNumber="34" type="female"></specimenCount>
,
<emphasis id="B9715E109C13FFE77DC0F94BFC52F93D" bold="true" box="[490,924,1719,1746]" pageId="33" pageNumber="34">
<collectingCountry id="F312C2929C13FFE77DC0F94BFD6AF93D" box="[490,676,1719,1745]" name="Mauritania" pageId="33" pageNumber="34">MAURITANIA</collectingCountry>
:
<emphasis id="B9715E109C13FFE77E99F94BFC5DF93D" bold="true" box="[691,915,1719,1746]" italics="true" pageId="33" pageNumber="34">
<collectingRegion id="49C14CE09C13FFE77E99F94BFC5DF93D" box="[691,915,1719,1746]" country="Mauritania" name="Nouakchott" pageId="33" pageNumber="34">Nouakchott Region</collectingRegion>
</emphasis>
:
</emphasis>
<collectingRegion id="49C14CE09C13FFE77F89F94BFBE7F93D" box="[931,1065,1719,1745]" country="Mauritania" name="Nouakchott" pageId="33" pageNumber="34">Nouakchott</collectingRegion>
,
<quantity id="4CFD2FE79C13FFE77819F94BFBB7F93D" box="[1075,1145,1719,1745]" metricMagnitude="4" metricUnit="m" metricValue="3.1" pageId="33" pageNumber="34" unit="km" value="31.0">31 km</quantity>
<location id="8EDAD4D99C13FFE77855F94BFA92F93D" LSID="urn:lsid:plazi:treatment:03AC33149C13FFE27CBDF9ACFB60FE89:8EDAD4D99C13FFE77855F94BFA92F93D" box="[1151,1372,1719,1745]" country="Mauritania" latitude="18.081034" longLatPrecision="1" longitude="-15.96625" name="South of Nouachott" pageId="33" pageNumber="34" stateProvince="Nouakchott">South of Nouachott</location>
, hand collecting,
<geoCoordinate id="EE31E4C59C13FFE77D3EF927FE66F919" box="[276,424,1755,1781]" degrees="18" direction="north" minutes="04.862" orientation="latitude" pageId="33" pageNumber="34" precision="1" value="18.081034">18°04.862N</geoCoordinate>
,
<geoCoordinate id="EE31E4C59C13FFE77D99F927FD85F91A" box="[435,587,1755,1782]" degrees="15" direction="west" minutes="57.975" orientation="longitude" pageId="33" pageNumber="34" precision="1" value="-15.96625">15°57.975W</geoCoordinate>
, leg.
<collectorName id="26F0E7D49C13FFE77EADF920FD31F91A" box="[647,767,1756,1782]" pageId="33" pageNumber="34">F. Borgato</collectorName>
,
<date id="FFBBA4C29C13FFE77F20F927FCA9F919" box="[778,871,1755,1781]" pageId="33" pageNumber="34" value="1994-11">
<collectingDate id="EFFF5D2A9C13FFE77F20F927FCA9F919" box="[778,871,1755,1781]" pageId="33" pageNumber="34" value="1994-11">XI.1994</collectingDate>
</date>
(CAS, CASENT 9078556).
</materialsCitation>
</paragraph>
<paragraph id="8BBA82029C13FFE77CEDF903FCBAF8F6" blockId="33.[151,1437,1719,2034]" box="[199,884,1791,1818]" pageId="33" pageNumber="34">
<emphasis id="B9715E109C13FFE77CEDF903FE86F8F5" bold="true" box="[199,328,1791,1817]" pageId="33" pageNumber="34">
<typeStatus id="54BE3CA09C13FFE77CEDF903FE8DF8F5" box="[199,323,1791,1817]" pageId="33" pageNumber="34" type="paratype">Paratypes</typeStatus>
:
</emphasis>
1 ♂, same data as holotype (CASENT 9078556).
</paragraph>
</subSubSection>
<subSubSection id="C31FD1899C13FFE77CEDF8DFFABCF88D" pageId="33" pageNumber="34" type="etymology">
<paragraph id="8BBA82029C13FFE77CEDF8DFFABCF88D" blockId="33.[151,1437,1719,2034]" pageId="33" pageNumber="34">
<emphasis id="B9715E109C13FFE77CEDF8DFFE80F8D1" bold="true" box="[199,334,1827,1853]" pageId="33" pageNumber="34">Etymology.</emphasis>
The name of this species is derived from the two Latin words
<emphasis id="B9715E109C13FFE77834F8DFFBFAF8D1" box="[1054,1076,1827,1853]" italics="true" pageId="33" pageNumber="34">bi</emphasis>
(two) and
<emphasis id="B9715E109C13FFE77898F8DFFAD4F8D1" box="[1202,1306,1827,1853]" italics="true" pageId="33" pageNumber="34">maculata</emphasis>
(spot), and refers to the two large white spots on the midline of the abdomen in both the males and females of this species.
</paragraph>
</subSubSection>
<subSubSection id="C31FD1899C13FFE47CEDF897FDBEFF39" lastPageId="34" lastPageNumber="35" pageId="33" pageNumber="34" type="diagnosis">
<paragraph id="8BBA82029C13FFE47CEDF897FDBEFF39" blockId="33.[151,1437,1719,2034]" lastBlockId="34.[151,1437,151,429]" lastPageId="34" lastPageNumber="35" pageId="33" pageNumber="34">
<emphasis id="B9715E109C13FFE77CEDF897FE8CF869" bold="true" box="[199,322,1899,1925]" pageId="33" pageNumber="34">Diagnosis.</emphasis>
<taxonomicName id="4C05F9819C13FFE77D60F890FDE4F869" authority="Booysen &amp; Haddad, 2021" authorityName="Booysen &amp; Haddad" authorityYear="2021" box="[330,554,1899,1925]" class="Arachnida" family="Gnaphosidae" genus="Micaria" kingdom="Animalia" order="Araneae" pageId="33" pageNumber="34" phylum="Arthropoda" rank="species" species="bimaculata" status="sp. nov.">
<emphasis id="B9715E109C13FFE77D60F890FDE4F869" box="[330,554,1899,1925]" italics="true" pageId="33" pageNumber="34">Micaria bimaculata</emphasis>
</taxonomicName>
<emphasis id="B9715E109C13FFE77E18F897FD43F869" bold="true" box="[562,653,1899,1925]" pageId="33" pageNumber="34">
<taxonomicNameLabel id="A242E36B9C13FFE77E18F897FD43F869" box="[562,653,1899,1925]" pageId="33" pageNumber="34" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
can be distinguished from other Afrotropical
<taxonomicName id="4C05F9819C13FFE778BCF890FB3EF869" authorityName="Westring" authorityYear="1851" box="[1174,1264,1900,1925]" class="Arachnida" family="Gnaphosidae" genus="Micaria" kingdom="Animalia" order="Araneae" pageId="33" pageNumber="34" phylum="Arthropoda" rank="genus">
<emphasis id="B9715E109C13FFE778BCF890FB3EF869" box="[1174,1264,1900,1925]" italics="true" pageId="33" pageNumber="34">Micaria</emphasis>
</taxonomicName>
in bearing two large white spots on the abdomen in both sexes. Furthermore, this species females can be distinguished from
<taxonomicName id="4C05F9819C13FFE77954F86CFE5CF821" authority="Strand, 1911" authorityName="Strand" authorityYear="1911" class="Arachnida" family="Gnaphosidae" genus="Micaria" kingdom="Animalia" order="Araneae" pageId="33" pageNumber="34" phylum="Arthropoda" rank="species" species="gomerae">
<emphasis id="B9715E109C13FFE77954F86CFF37F821" italics="true" pageId="33" pageNumber="34">M. gomerae</emphasis>
Strand,
<quantity id="4CFD2FE79C13FFE77D71F84FFE5CF821" box="[347,402,1971,1997]" metricMagnitude="1" metricUnit="m" metricValue="4.85394" pageId="33" pageNumber="34" unit="in" value="1911.0">1911</quantity>
</taxonomicName>
in having copulatory ducts that are thicker and separated by their width. The males of
<taxonomicName id="4C05F9819C13FFE77954F848FEDBF81D" authority="Booysen &amp; Haddad, 2021" authorityName="Booysen &amp; Haddad" authorityYear="2021" class="Arachnida" family="Gnaphosidae" genus="Micaria" kingdom="Animalia" order="Araneae" pageId="33" pageNumber="34" phylum="Arthropoda" rank="species" species="bimaculata" status="sp. nov.">
<emphasis id="B9715E109C13FFE77954F848FEDBF81D" italics="true" pageId="33" pageNumber="34">M. bimaculata</emphasis>
</taxonomicName>
<emphasis id="B9715E109C13FFE77D34F82BFEB7F81D" bold="true" box="[286,377,2007,2033]" pageId="33" pageNumber="34">
<taxonomicNameLabel id="A242E36B9C13FFE77D34F82BFEB7F81D" box="[286,377,2007,2033]" pageId="33" pageNumber="34" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
differ from
<taxonomicName id="4C05F9819C13FFE77E2DF824FD5EF81D" authorityName="Strand" authorityYear="1911" box="[519,656,2008,2033]" class="Arachnida" family="Gnaphosidae" genus="Micaria" kingdom="Animalia" order="Araneae" pageId="33" pageNumber="34" phylum="Arthropoda" rank="species" species="gomerae">
<emphasis id="B9715E109C13FFE77E2DF824FD5EF81D" box="[519,656,2008,2033]" italics="true" pageId="33" pageNumber="34">M. gomerae</emphasis>
</taxonomicName>
males in have a shorter embolus that curves retrolaterally, at a right angle, at half the length of the embolus. Furthermore, the median apophysis is shorter and thicker than that of
<taxonomicName id="4C05F9819C10FFE47954FF6BFF37FF38" authorityName="Strand" authorityYear="1911" class="Arachnida" family="Gnaphosidae" genus="Micaria" kingdom="Animalia" order="Araneae" pageId="34" pageNumber="35" phylum="Arthropoda" rank="species" species="gomerae">
<emphasis id="B9715E109C10FFE47954FF6BFF37FF38" italics="true" pageId="34" pageNumber="35">M. gomerae</emphasis>
</taxonomicName>
(figs 63a-gin
<bibRefCitation id="EF94FFF39C10FFE47DB4FF47FDAAFF39" author="Wunderlich, V. J." box="[414,612,187,213]" journalOrPublisher="Zoologische Beitrage" pageId="34" pageNumber="35" pagination="233 - 341" part="25" refId="ref60449" refString="Wunderlich, V. J. (1980) Revision der europaischen Arten der Gattung Micaria Westring 1851, mit Anmerkungen zu den ubrigen palaarktischen Arten (Arachnida: Araneida: Gnaphosidae). Zoologische Beitrage, 25, 233 - 341." title="Revision der europaischen Arten der Gattung Micaria Westring 1851, mit Anmerkungen zu den ubrigen palaarktischen Arten (Arachnida: Araneida: Gnaphosidae)" type="journal article" year="1980">Wunderlich 1980</bibRefCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="C31FD1899C10FFE27CEDFF22FCE0FEAD" lastPageId="36" lastPageNumber="37" pageId="34" pageNumber="35" type="description">
<paragraph id="8BBA82029C10FFE47CEDFF22FE69FE41" blockId="34.[151,1437,151,429]" pageId="34" pageNumber="35">
<emphasis id="B9715E109C10FFE47CEDFF22FB09FF15" bold="true" box="[199,1223,222,249]" pageId="34" pageNumber="35">
Description. Female (
<typeStatus id="54BE3CA09C10FFE47DE2FF22FDFEFF14" box="[456,560,222,248]" pageId="34" pageNumber="35" type="holotype">holotype</typeStatus>
from
<collectingRegion id="49C14CE09C10FFE47E56FF23FCC4FF15" box="[636,778,223,249]" country="Mauritania" name="Nouakchott" pageId="34" pageNumber="35">Nouakchott</collectingRegion>
, CASENT 9078556): Measurements:
</emphasis>
TL 3.75, CL 1.35, CW 0.95, CL:CW 1.42, CLH 0.10, CLH:AME 1.20. Eyes: ALE 0.06, AME 0.08, PLE 0.06, PME 0.06, AMEAME 0.04, PMEPME 0.07, MOQAW 0.18, MOQPW 0.19, MOQL 0.19. SL 0.78, SW 0.57, AL 2.15, AW 1.45.
<emphasis id="B9715E109C10FFE47945FEDBFE83FE89" bold="true" pageId="34" pageNumber="35">
<collectingRegion id="49C14CE09C10FFE47945FEDBFA52FEAD" box="[1391,1436,295,321]" country="South Sudan" name="Lakes" pageId="34" pageNumber="35">Leg</collectingRegion>
measurements:
</emphasis>
Palpal segment lengths: 0.44, 0.19, 0.25, 0.43.
<collectingRegion id="49C14CE09C10FFE47F45FEB7FC55FE89" box="[879,923,331,357]" country="South Sudan" name="Lakes" pageId="34" pageNumber="35">Leg</collectingRegion>
I: 0.97, 0.30, 0.49, 0.41, 0.64. II: 0.95, 0.29, 0.50, 0.40, 0.64. III: 0.87, 0.38, 0.65, 0.73, 0.59. IV: 1.40, 0.46, 0.98, 1.19, 0.87. TLoflegs (IIV): 2.81, 2.78, 3.22, 4.90; leg formula: 4312.
</paragraph>
<caption id="DF7AD28A9C10FFE47CBDFB20FBA9FAF4" ID-DOI="http://doi.org/10.5281/zenodo.4580966" ID-Zenodo-Dep="4580966" httpUri="https://zenodo.org/record/4580966/files/figure.png" pageId="34" pageNumber="35" startId="34.[151,264,1244,1268]" targetBox="[172,1406,462,1215]" targetPageId="34">
<paragraph id="8BBA82029C10FFE47CBDFB20FBA9FAF4" blockId="34.[151,1436,1244,1305]" pageId="34" pageNumber="35">
<emphasis id="B9715E109C10FFE47CBDFB20FEA3FB18" bold="true" box="[151,365,1244,1268]" pageId="34" pageNumber="35">FIGURES 117120.</emphasis>
Genital morphology of
<taxonomicName id="4C05F9819C10FFE47E4CFB21FCFFFB18" authority="Booysen &amp; Haddad, 2021" authorityName="Booysen &amp; Haddad" authorityYear="2021" box="[614,817,1244,1268]" class="Arachnida" family="Gnaphosidae" genus="Micaria" kingdom="Animalia" order="Araneae" pageId="34" pageNumber="35" phylum="Arthropoda" rank="species" species="bimaculata" status="sp. nov.">
<emphasis id="B9715E109C10FFE47E4CFB21FCFFFB18" box="[614,817,1244,1268]" italics="true" pageId="34" pageNumber="35">Micaria bimaculata</emphasis>
</taxonomicName>
<emphasis id="B9715E109C10FFE47F12FB21FC47FB19" bold="true" box="[824,905,1245,1269]" pageId="34" pageNumber="35">
<taxonomicNameLabel id="A242E36B9C10FFE47F12FB21FC47FB19" box="[824,905,1245,1269]" pageId="34" pageNumber="35" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
: 117 female epigyne, ventral view; 118 female epigyne, dorsal view; 119 male palp, ventral view; 120 same, retrolateral view. Scale bars: 0.1 mm.
</paragraph>
</caption>
<paragraph id="8BBA82029C10FFE47CEDFABDFC4FF877" blockId="34.[151,1437,1345,2020]" pageId="34" pageNumber="35">
<emphasis id="B9715E109C10FFE47CEDFABDFEAFFAB7" bold="true" box="[199,353,1345,1371]" pageId="34" pageNumber="35">Colouration:</emphasis>
carapace brown; abdomen brown on anterior half and dark brown on posterior half; legs uniform in colour (femora may be darker in live specimens); sternum, endites, labium and chelicerae similar in colour to carapace (
<figureCitation id="133E9E879C10FFE47D22FA75FE92FA4F" box="[264,348,1417,1443]" captionStart="FIGURES 917" captionStartId="11.[151,264,1943,1967]" captionTargetBox="[173,1415,181,1920]" captionTargetId="figure-17@11.[173,1415,181,1920]" captionTargetPageId="11" captionText="FIGURES 917. Stereomicroscope photographs of the dorsal habitus ofAfrotropical Micaria species: 9 M. basaliducta sp. nov. female and 10 male; 11 M. beaufortia (Tucker, 1923) female and 12 male; 13 M. bimaculata sp. nov. female and 14 male; 15 M. bispicula sp. nov. female and 16 male; 17 M. chrysis (Simon, 1910) female. Scale bars: 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.4580929" httpUri="https://zenodo.org/record/4580929/files/figure.png" pageId="34" pageNumber="35">Fig. 13</figureCitation>
).
<emphasis id="B9715E109C10FFE47D5AFA75FE22FA4F" bold="true" box="[368,492,1417,1443]" pageId="34" pageNumber="35">Carapace:</emphasis>
smooth in texture; decorated with squamose setae; pattern faded but seems to be radiating from middle; posterior margin indented medially; anterior margin of carapace narrowing slightly from coxa I; PER procurved in dorsal view, AER recurved in anterior view; fovea absent.
<emphasis id="B9715E109C10FFE47829FA2DFB42FA07" bold="true" box="[1027,1164,1489,1515]" pageId="34" pageNumber="35">Chelicerae:</emphasis>
decorated with plumose setae; apical margin slightly thickened.
<emphasis id="B9715E109C10FFE47E70FA09FD0CF9E3" bold="true" box="[602,706,1525,1551]" pageId="34" pageNumber="35">Labium:</emphasis>
dark spots in posterior corners; triangular in shape; tuft of setae present at tip.
<emphasis id="B9715E109C10FFE47D16F9E5FE51F9DF" bold="true" box="[316,415,1561,1587]" pageId="34" pageNumber="35">Endites:</emphasis>
slanting inwards, constricted just above halfway; cluster of setae present on innerapical margin.
<emphasis id="B9715E109C10FFE47CD9F9C1FEABF9BB" bold="true" box="[243,357,1597,1623]" pageId="34" pageNumber="35">Sternum:</emphasis>
darkened ridges; shield-like; broadest between coxa Iand II; posterior margin truncated between coxa IV.
<emphasis id="B9715E109C10FFE47CD0F99DFEF5F997" bold="true" box="[250,315,1633,1659]" pageId="34" pageNumber="35">Legs:</emphasis>
femora laterally flattened, highest at base, leg Imore pronounced in this fashion; rest of legs normal, thin.
<collectingRegion id="49C14CE09C10FFE47CFAF97AFF32F94C" box="[208,252,1670,1696]" country="South Sudan" name="Lakes" pageId="34" pageNumber="35">Leg</collectingRegion>
macrosetae: Palps: 1do, 1pl, 1do, 1pl, 1pl, 2plv, 4vt.
<collectingRegion id="49C14CE09C10FFE47F66F97AFCB6F94C" box="[844,888,1670,1696]" country="South Sudan" name="Lakes" pageId="34" pageNumber="35">Leg</collectingRegion>
I: I 1do, 1pl, 2plv, 2vt. II: 2do, 1pl, 1rl, 2plv, 2vt. III: 2do, 1pl, 1rl, 2pl, 1rl, 2plv, 1rlv, 2vt, 2pl, 1rl, 2plv, 2rlv, 5vt. IV: 2do, 2pl, 2rl, 2plv, 4vt, 1pl, 1rl, 2plv, 2rlv, 4vt. Legs IIV with two rows of scopulate setae on metatarsi and four rows on tarsi.
<emphasis id="B9715E109C10FFE4781AF931FB60F90B" bold="true" box="[1072,1198,1741,1767]" pageId="34" pageNumber="35">Abdomen:</emphasis>
cylindrical in shape; decorated with squamose setae; dorsal pattern forming two large spots on transverse midline of abdomen (
<figureCitation id="133E9E879C10FFE47911F90DFA43F8E7" box="[1339,1421,1777,1803]" captionStart="FIGURES 917" captionStartId="11.[151,264,1943,1967]" captionTargetBox="[173,1415,181,1920]" captionTargetId="figure-17@11.[173,1415,181,1920]" captionTargetPageId="11" captionText="FIGURES 917. Stereomicroscope photographs of the dorsal habitus ofAfrotropical Micaria species: 9 M. basaliducta sp. nov. female and 10 male; 11 M. beaufortia (Tucker, 1923) female and 12 male; 13 M. bimaculata sp. nov. female and 14 male; 15 M. bispicula sp. nov. female and 16 male; 17 M. chrysis (Simon, 1910) female. Scale bars: 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.4580929" httpUri="https://zenodo.org/record/4580929/files/figure.png" pageId="34" pageNumber="35">Fig. 13</figureCitation>
); venter similar in colour to dorsum; sigilla not visible.
<emphasis id="B9715E109C10FFE47F24F8E9FCB7F8C3" bold="true" box="[782,889,1813,1839]" pageId="34" pageNumber="35">Epigyne:</emphasis>
in ventral view, anterior hood divided, curved; posterior pockets medially between spermathecae; copulatory openings globular. In dorsal view, copulatory ducts present medially on inner apical margin of spermathecae; fertilisation ducts present on basal interior margin of spermathecae, curved laterally; spermathecae globular (
<figureCitation id="133E9E879C10FFE47EF2F87DFCBBF877" box="[728,885,1921,1947]" captionStart="FIGURES 117120" captionStartId="34.[151,264,1244,1268]" captionTargetBox="[172,1406,462,1215]" captionTargetId="figure-156@34.[164,1423,453,1221]" captionTargetPageId="34" captionText="FIGURES 117120. Genital morphology of Micaria bimaculata sp. nov.: 117 female epigyne, ventral view; 118 female epigyne, dorsal view; 119 male palp, ventral view; 120 same, retrolateral view. Scale bars: 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.4580966" httpUri="https://zenodo.org/record/4580966/files/figure.png" pageId="34" pageNumber="35">Figs 117, 118</figureCitation>
).
</paragraph>
<paragraph id="8BBA82029C10FFE57CEDF859FF19FEF1" blockId="34.[151,1437,1345,2020]" lastBlockId="35.[151,1437,150,285]" lastPageId="35" lastPageNumber="36" pageId="34" pageNumber="35">
<emphasis id="B9715E109C10FFE47CEDF859FC93F853" bold="true" box="[199,861,1957,1983]" pageId="34" pageNumber="35">
Male. (
<typeStatus id="54BE3CA09C10FFE47D31F859FE46F853" box="[283,392,1957,1983]" pageId="34" pageNumber="35" type="paratype">paratype</typeStatus>
from
<collectingRegion id="49C14CE09C10FFE47DF9F859FDAFF853" box="[467,609,1957,1983]" country="Mauritania" name="Nouakchott" pageId="34" pageNumber="35">Nouakchott</collectingRegion>
, CASENT 9078556):
</emphasis>
<materialsCitation id="3B6D885F9C10FFE57F4FF859FF1DFEF1" ID-GBIF-Occurrence="3071034304" collectionCode="MOQAW" lastPageId="35" lastPageNumber="36" location="Palpal" pageId="34" pageNumber="35" specimenCount="1" stateProvince="Lakes">
<emphasis id="B9715E109C10FFE47F4FF859FBD0F853" bold="true" box="[869,1054,1957,1983]" pageId="34" pageNumber="35">Measurements:</emphasis>
TL 3.25, CL 1.45, CW 1.05, CL: CW 1.38, CLH 0.13, CLH:AME 1.65. Eyes: ALE 0.06, AME 0.08, PLE 0.06, PME 0.06, AMEAME 0.02, PME PME 0.10,
<collectionCode id="ED141AC79C11FFE57D3FFF6BFE4CFF5D" box="[277,386,151,177]" pageId="35" pageNumber="36">MOQAW</collectionCode>
0.19, MOQPW 0.21, MOQL 0.22. SL 0.74, SW 0.59, AL 1.60, AW 1.10.
<emphasis id="B9715E109C11FFE57899FF6BFA52FF5D" bold="true" box="[1203,1436,151,177]" pageId="35" pageNumber="36">
<collectingRegion id="49C14CE09C11FFE57899FF6BFB2EFF5D" box="[1203,1248,151,177]" country="South Sudan" name="Lakes" pageId="35" pageNumber="36">Leg</collectingRegion>
measurements:
</emphasis>
<location id="8EDAD4D99C11FFE57CBDFF47FF13FF39" LSID="urn:lsid:plazi:treatment:03AC33149C13FFE27CBDF9ACFB60FE89:8EDAD4D99C11FFE57CBDFF47FF13FF39" box="[151,221,187,213]" name="Palpal" pageId="35" pageNumber="36" stateProvince="Lakes">Palpal</location>
segment lengths: 0.49, 0.29, 0.30, 0.38.
<collectingRegion id="49C14CE09C11FFE57EB5FF47FD05FF39" box="[671,715,187,213]" country="South Sudan" name="Lakes" pageId="35" pageNumber="36">Leg</collectingRegion>
I: 1.48, 0.49, 0.95, 0.81, 0.78. II: 1.13, 0.48, 0.81, 0.73, 0.78. III: 0.94, 0.44, 0.83, 0.92, 0.76. IV: 1.46, 0.46, 1.16, 1.19, 0.94. TLoflegs (IIV): 4.51, 3.93, 3.89, 5.21; leg formula: 4123
</materialsCitation>
.
</paragraph>
<caption id="DF7AD28A9C11FFE57CBDFA6EFF26FA22" ID-DOI="http://doi.org/10.5281/zenodo.4737271" ID-Zenodo-Dep="4737271" httpUri="https://zenodo.org/record/4737271/files/figure.png" pageId="35" pageNumber="36" startId="35.[151,209,1426,1450]" targetBox="[151,1435,309,1402]" targetPageId="35">
<paragraph id="8BBA82029C11FFE57CBDFA6EFF26FA22" blockId="35.[151,1436,1426,1486]" pageId="35" pageNumber="36">
<emphasis id="B9715E109C11FFE57CBDFA6EFF27FA46" bold="true" box="[151,233,1426,1450]" pageId="35" pageNumber="36">MAP 2.</emphasis>
The distribution of
<taxonomicName id="4C05F9819C11FFE57D9FFA6FFDB1FA46" authority="Booysen &amp; Haddad, 2021" authorityName="Booysen &amp; Haddad" authorityYear="2021" box="[437,639,1426,1450]" class="Arachnida" family="Gnaphosidae" genus="Micaria" kingdom="Animalia" order="Araneae" pageId="35" pageNumber="36" phylum="Arthropoda" rank="species" species="bimaculata" status="sp. nov.">
<emphasis id="B9715E109C11FFE57D9FFA6FFDB1FA46" box="[437,639,1426,1450]" italics="true" pageId="35" pageNumber="36">Micariabimaculata</emphasis>
</taxonomicName>
<emphasis id="B9715E109C11FFE57EAFFA6EFD18FA46" bold="true" box="[645,726,1426,1450]" pageId="35" pageNumber="36">
<taxonomicNameLabel id="A242E36B9C11FFE57EAFFA6EFD18FA46" box="[645,726,1426,1450]" pageId="35" pageNumber="36" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
(black circle) and
<taxonomicName id="4C05F9819C11FFE57FBCFA6FFBDBFA46" authority="Booysen &amp; Haddad, 2021" authorityName="Booysen &amp; Haddad" authorityYear="2021" box="[918,1045,1426,1450]" class="Arachnida" family="Gnaphosidae" genus="Micaria" kingdom="Animalia" order="Araneae" pageId="35" pageNumber="36" phylum="Arthropoda" rank="species" species="bispicula" status="sp. nov.">
<emphasis id="B9715E109C11FFE57FBCFA6FFBDBFA46" box="[918,1045,1426,1450]" italics="true" pageId="35" pageNumber="36">M. bispicula</emphasis>
</taxonomicName>
<emphasis id="B9715E109C11FFE57836FA6EFBA2FA46" bold="true" box="[1052,1132,1426,1450]" pageId="35" pageNumber="36">
<taxonomicNameLabel id="A242E36B9C11FFE57836FA6EFBA2FA46" box="[1052,1132,1426,1450]" pageId="35" pageNumber="36" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
(red stars) in the Afrotropical Region.
</paragraph>
</caption>
<paragraph id="8BBA82029C11FFE27CEDFA00FCE0FEAD" blockId="35.[151,1437,1532,2027]" lastBlockId="36.[151,1436,151,357]" lastPageId="36" lastPageNumber="37" pageId="35" pageNumber="36">
<emphasis id="B9715E109C11FFE57CEDFA00FEAFF9FA" bold="true" box="[199,353,1532,1558]" pageId="35" pageNumber="36">Colouration:</emphasis>
carapace dark brown; abdomen brown on anterior half and dark brown on posterior half; femora darkened partially, rest of legs normal in colour; sternum, endites, labium and chelicerae similar in colour to carapace (
<figureCitation id="133E9E879C11FFE57CF3F9B8FEE2F9B3" box="[217,300,1604,1631]" captionStart="FIGURES 917" captionStartId="11.[151,264,1943,1967]" captionTargetBox="[173,1415,181,1920]" captionTargetId="figure-17@11.[173,1415,181,1920]" captionTargetPageId="11" captionText="FIGURES 917. Stereomicroscope photographs of the dorsal habitus ofAfrotropical Micaria species: 9 M. basaliducta sp. nov. female and 10 male; 11 M. beaufortia (Tucker, 1923) female and 12 male; 13 M. bimaculata sp. nov. female and 14 male; 15 M. bispicula sp. nov. female and 16 male; 17 M. chrysis (Simon, 1910) female. Scale bars: 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.4580929" httpUri="https://zenodo.org/record/4580929/files/figure.png" pageId="35" pageNumber="36">Fig. 14</figureCitation>
).
<emphasis id="B9715E109C11FFE57D15F9B8FE72F9B2" bold="true" box="[319,444,1604,1630]" pageId="35" pageNumber="36">Carapace:</emphasis>
smooth in texture, decorated with squamose setae; pattern faded, but radiating from middle towards coxae, posterior margin similar to female; fovea absent; anterior margin of carapace narrowing slightly from coxa I; PER procurved in dorsal view, AER recurved in anterior view.
<emphasis id="B9715E109C11FFE57FD7F970FB4BF94A" bold="true" box="[1021,1157,1676,1702]" pageId="35" pageNumber="36">Chelicerae:</emphasis>
decorated with plumose setae; apical margin slightly thickened.
<emphasis id="B9715E109C11FFE57E70F94CFD0CF926" bold="true" box="[602,706,1712,1738]" pageId="35" pageNumber="36">Labium:</emphasis>
dark spots in posterior corners; triangular in shape; tuft of setae present at tip.
<emphasis id="B9715E109C11FFE57D16F928FE51F902" bold="true" box="[316,415,1748,1774]" pageId="35" pageNumber="36">Endites:</emphasis>
slanting inwards, constricted just above halfway; cluster of setae present on innerapical margin; apical margin flattened; posterior margin bulbous.
<emphasis id="B9715E109C11FFE57F6EF904FC7BF8FE" bold="true" box="[836,949,1784,1810]" pageId="35" pageNumber="36">Sternum:</emphasis>
darkened ridges; shield-like; broadest between coxa Iand II; posterior margin narrowly truncated between coxa IV.
<emphasis id="B9715E109C11FFE57FF6F8E1FBD3F8DB" bold="true" box="[988,1053,1821,1847]" pageId="35" pageNumber="36">Legs:</emphasis>
femora laterally flattened, highest at base, leg Imore pronounced in this fashion; rest of legs normal, thin.
<collectingRegion id="49C14CE09C11FFE57FE9F8BDFC21F8B7" box="[963,1007,1857,1883]" country="South Sudan" name="Lakes" pageId="35" pageNumber="36">Leg</collectingRegion>
macrosetae: Palps: 1do, 1pl, 1do, 1pl, 1pl, 1plv, 4vt.
<collectingRegion id="49C14CE09C11FFE57D13F899FEABF893" box="[313,357,1893,1919]" country="South Sudan" name="Lakes" pageId="35" pageNumber="36">Leg</collectingRegion>
I: 1do, 1pl. II: 2do, 2pl, 2plv, 2vt. III: 2do, 2plv, 1rl, 2pl, 2rl, 2plv, 2rlv, 2vt, 1pl, 1rl, 2l, 1rl, 4vt. IV: 2do, 1rl, 2pl, 2rl, 2plv, 2rlv, 2vt, 1pl, 1rl, 2plv, 2rlv, 4vt. Legs IIV with two rows of scopulate setae on metatarsi and four rows on tarsi.
<emphasis id="B9715E109C11FFE57DB0F850FDD6F82A" bold="true" box="[410,536,1964,1990]" pageId="35" pageNumber="36">Abdomen:</emphasis>
cylindrical in shape; decorated with squamose setae; dorsal pattern forming two large spots on transverse midline of abdomen (
<figureCitation id="133E9E879C11FFE57EB5F82CFD3FF807" box="[671,753,2000,2027]" captionStart="FIGURES 917" captionStartId="11.[151,264,1943,1967]" captionTargetBox="[173,1415,181,1920]" captionTargetId="figure-17@11.[173,1415,181,1920]" captionTargetPageId="11" captionText="FIGURES 917. Stereomicroscope photographs of the dorsal habitus ofAfrotropical Micaria species: 9 M. basaliducta sp. nov. female and 10 male; 11 M. beaufortia (Tucker, 1923) female and 12 male; 13 M. bimaculata sp. nov. female and 14 male; 15 M. bispicula sp. nov. female and 16 male; 17 M. chrysis (Simon, 1910) female. Scale bars: 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.4580929" httpUri="https://zenodo.org/record/4580929/files/figure.png" pageId="35" pageNumber="36">Fig. 14</figureCitation>
); venter similar in colour to dorsum; sigilla not visible.
<emphasis id="B9715E109C11FFE57976F82DFA52F807" bold="true" box="[1372,1436,2001,2027]" pageId="35" pageNumber="36">Palp:</emphasis>
retrolateral tibial apophysis present dorsally; median apophysis present next to embolus; median apophysis broad, elongate, bending prolaterally, skewed diagonally; sperm duct present on retrolateral surface of tegulum, curved basally; sperm duct “S” shaped before ending at embolus; embolus broad at base, strongly curved retrolaterally, tip elongate; cymbium elongate, widest near middle; cymbial tip rounded, two small spines ventrally on tip; three ventralterminalspinesinfrontof embolus (
<figureCitation id="133E9E879C16FFE27EAEFEDBFCEFFEAD" box="[644,801,295,321]" captionStart="FIGURES 117120" captionStartId="34.[151,264,1244,1268]" captionTargetBox="[172,1406,462,1215]" captionTargetId="figure-156@34.[164,1423,453,1221]" captionTargetPageId="34" captionText="FIGURES 117120. Genital morphology of Micaria bimaculata sp. nov.: 117 female epigyne, ventral view; 118 female epigyne, dorsal view; 119 male palp, ventral view; 120 same, retrolateral view. Scale bars: 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.4580966" httpUri="https://zenodo.org/record/4580966/files/figure.png" pageId="36" pageNumber="37">Figs 119, 120</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="C31FD1899C16FFE27CEDFEB6FB60FE89" box="[199,1198,330,357]" pageId="36" pageNumber="37" type="distribution">
<paragraph id="8BBA82029C16FFE27CEDFEB6FB60FE89" blockId="36.[151,1436,151,357]" box="[199,1198,330,357]" pageId="36" pageNumber="37">
<emphasis id="B9715E109C16FFE27CEDFEB6FEAFFE88" bold="true" box="[199,353,330,356]" pageId="36" pageNumber="37">Distribution.</emphasis>
<taxonomicName id="4C05F9819C16FFE27D4DFEB7FD89FE89" authority="Booysen &amp; Haddad, 2021" authorityName="Booysen &amp; Haddad" authorityYear="2021" box="[359,583,331,357]" class="Arachnida" family="Gnaphosidae" genus="Micaria" kingdom="Animalia" order="Araneae" pageId="36" pageNumber="37" phylum="Arthropoda" rank="species" species="bimaculata" status="sp. nov.">
<emphasis id="B9715E109C16FFE27D4DFEB7FD89FE89" box="[359,583,331,357]" italics="true" pageId="36" pageNumber="37">Micaria bimaculata</emphasis>
</taxonomicName>
<emphasis id="B9715E109C16FFE27E64FEB7FD69FE89" bold="true" box="[590,679,331,357]" pageId="36" pageNumber="37">
<taxonomicNameLabel id="A242E36B9C16FFE27E64FEB7FD69FE89" box="[590,679,331,357]" pageId="36" pageNumber="37" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
is only known from the
<typeStatus id="54BE3CA09C16FFE27F90FEB7FC24FE89" box="[954,1002,331,357]" pageId="36" pageNumber="37">type</typeStatus>
locality (
<figureCitation id="133E9E879C16FFE2787EFEB7FB6FFE89" box="[1108,1185,331,357]" captionStart="MAP 2" captionStartId="35.[151,209,1426,1450]" captionTargetBox="[151,1435,309,1402]" captionTargetId="figure-79@35.[151,1435,309,1402]" captionTargetPageId="35" captionText="MAP 2. The distribution of Micariabimaculata sp. nov. (black circle) and M. bispicula sp. nov. (red stars) in the Afrotropical Region." figureDoi="http://doi.org/10.5281/zenodo.4737271" httpUri="https://zenodo.org/record/4737271/files/figure.png" pageId="36" pageNumber="37">Map 2</figureCitation>
).
</paragraph>
</subSubSection>
</treatment>
</document>