787 lines
100 KiB
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787 lines
100 KiB
XML
<document id="734F93DDED1F111CAC956BE710903612" ID="10.11646/zootaxa.4232.3.8SLASH10211" ID-CLB-Dataset="36515" ID-DOI="10.11646/zootaxa.4232.3.8" ID-GBIF-Dataset="7cff4fc5-f588-4d9e-b590-00b0642e58e5" ID-ISSN="1175-5326" ID-Zenodo-Dep="312543" ID-ZooBank="FBF3B84C-AE4E-4266-9C44-2057F8C1EBDC" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1487322540372" checkinUser="plazi" docAuthor="Seiter, Michael & Wolff, Jonas O." docDate="2017" docId="AC03BD57FFB3FFC7BBF09DF2B725FE20" docLanguage="en" docName="zootaxa.4232.3.8SLASH10211.pdf" docOrigin="Zootaxa 4232 (3)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Stygophrynus orientalis Seiter & Wolff, 2017, sp. nov." docType="treatment" docVersion="9" lastPageNumber="406" masterDocId="503AC52FFFB1FFCEBB679D65B608FFD0" masterDocTitle="Stygophrynus orientalis sp. nov. (Amblypygi: Charontidae) from Indonesia with the description of a remarkable spermatophore" masterLastPageNumber="408" masterPageNumber="397" pageNumber="399" updateTime="1698437049029" updateUser="ExternalLinkService">
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<mods:mods id="8EDB5C238A2186D22D6868A569A8647F" xmlns:mods="http://www.loc.gov/mods/v3">
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<mods:titleInfo id="93D332290E1455905D918642117F2BD0">
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<mods:title id="E3A311EF493C4FA2D87362BFD44DDCB1">Stygophrynus orientalis sp. nov. (Amblypygi: Charontidae) from Indonesia with the description of a remarkable spermatophore</mods:title>
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<mods:name id="2EA1C2F1D30B20F1FE5BEBEDFB90EF35" type="personal">
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<mods:roleTerm id="FA00B058773E5EF39A825D04FF89DC0B">Author</mods:roleTerm>
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<mods:namePart id="730C17FE11C188945449D602081F0F70">Seiter, Michael</mods:namePart>
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<mods:name id="1EB9CBD335A91252A150364E738CB0EA" type="personal">
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<mods:roleTerm id="A3BB41717A642824133ACD0EE9411EF1">Author</mods:roleTerm>
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<mods:namePart id="6C275FE53429987E9165681B0257BD3E">Wolff, Jonas O.</mods:namePart>
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<mods:typeOfResource id="F0801C7C55EB0BE09B0BBD375383DDF3">text</mods:typeOfResource>
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<mods:title id="C7336450BBC42AC4FEAC927EB66EBF6A">Zootaxa</mods:title>
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<mods:part id="18F39A3A3CE781A398BF7D4F5D5E07D6">
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<mods:date id="C6289C8DBEDC600A189BE29CC4E1879B">2017</mods:date>
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<mods:number id="3B469F06D3352A01BAF63E8D0CA5B59F">4232</mods:number>
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<mods:number id="05DC560A7722301FD8C17F33F4DCA170">3</mods:number>
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<mods:identifier id="C18FBB85B521B4E22B584B6F41CFF15A" type="CLB-Dataset">36515</mods:identifier>
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<mods:identifier id="73300AC7E0E7EADEB0763162E5262758" type="DOI">10.11646/zootaxa.4232.3.8</mods:identifier>
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<treatment id="AC03BD57FFB3FFC7BBF09DF2B725FE20" ID-DOI="http://doi.org/10.5281/zenodo.5416836" ID-GBIF-Taxon="127696572" ID-Zenodo-Dep="5416836" LSID="urn:lsid:plazi:treatment:AC03BD57FFB3FFC7BBF09DF2B725FE20" httpUri="http://treatment.plazi.org/id/AC03BD57FFB3FFC7BBF09DF2B725FE20" lastPageId="9" lastPageNumber="406" pageId="2" pageNumber="399">
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<subSubSection id="6CB05FCAFFB3FFCCBBF09DF2B70FFF03" pageId="2" pageNumber="399" type="nomenclature">
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<paragraph id="24150C41FFB3FFCCBBF09DF2B411FF62" blockId="2.[151,537,151,211]" box="[151,537,151,178]" pageId="2" pageNumber="399">
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<heading id="7F5DBB2DFFB3FFCCBBF09DF2B411FF62" bold="true" box="[151,537,151,178]" fontSize="11" level="1" pageId="2" pageNumber="399" reason="1">
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<emphasis id="16DED053FFB3FFCCBBF09DF2B411FF62" bold="true" box="[151,537,151,178]" pageId="2" pageNumber="399">
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<taxonomicName id="E3AA77C2FFB3FFCCBBF09DF2B7BDFF61" ID-CoL="6ZZNZ" box="[151,437,151,177]" class="Arachnida" family="Charontidae" genus="Stygophrynus" kingdom="Animalia" order="Amblypygi" pageId="2" pageNumber="399" phylum="Arthropoda" rank="species" species="orientalis" status="sp. nov.">
|
||
<emphasis id="16DED053FFB3FFCCBBF09DF2B7BDFF61" bold="true" box="[151,437,151,177]" italics="true" pageId="2" pageNumber="399">Stygophrynus orientalis</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="0DED6D28FFB3FFCCBADB9DFDB411FF62" box="[444,537,152,178]" pageId="2" pageNumber="399" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
</heading>
|
||
</paragraph>
|
||
<paragraph id="24150C41FFB3FFCCBBF09DDFB70FFF03" blockId="2.[151,537,151,211]" box="[151,263,186,211]" pageId="2" pageNumber="399">
|
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(
|
||
<figureCitation id="BC9110C4FFB3FFCCBBF89DDFB6F5FF03" box="[159,253,186,211]" captionStart-0="FIGURE 1" captionStart-1="FIGURE 2" captionStart-2="FIGURE 3" captionStart-3="FIGURE 4" captionStart-4="FIGURE 5" captionStartId-0="3.[151,250,1857,1879]" captionStartId-1="4.[151,250,1866,1888]" captionStartId-2="5.[151,250,1141,1163]" captionStartId-3="6.[151,250,1539,1561]" captionStartId-4="7.[151,250,1922,1944]" captionTargetBox-0="[252,1295,248,1759]" captionTargetBox-1="[200,1385,232,1845]" captionTargetBox-2="[151,1436,195,1120]" captionTargetBox-3="[151,1436,193,1517]" captionTargetBox-4="[151,1391,896,1878]" captionTargetId-0="figure@3.[199,1344,193,1813]" captionTargetId-1="figure@4.[200,1386,193,1845]" captionTargetId-2="figure@5.[151,1436,187,1120]" captionTargetId-3="figure@6.[151,1436,193,1517]" captionTargetId-4="figure@7.[151,1436,877,1936]" captionTargetPageId-0="3" captionTargetPageId-1="4" captionTargetPageId-2="5" captionTargetPageId-3="6" captionTargetPageId-4="7" captionText-0="FIGURE 1. Stygophrynus orientalis sp. nov. A Habitus of female holotype, dorsal and ventral view, respectively. B – D Female chelicera, denticles enhanced with white lines (B internal view; C external view; D external view, close-up on external teeth row). E Pedipalp (left ventral, right dorsal view), spines of femur and tibia numbered. F Distitibia and distal basitibia of walking leg IV, dorsal; trichobothria marked as follows: basitibia (bt = basitibial), distitibia (bf: basofrontal; bc: basocaudal; sbf: subbasofrontal; sbc-x: series caudal trichobothria; stf 1 x: series frontal trichobothria). Note: not all trichobothria are visible on this figure. Scale bars: A 20 mm, B – C 2 mm, D 0.25 mm E 2 mm, F 1 mm." captionText-1="FIGURE 2. Stygophrynus orientalis sp. nov. Variability of pedipalpal spination. Distal part of pedipalp, pro-lateral view. A Holotype female (21944), tarsus of right pedipalp, left drawing, right original photograph. B Paratype male (# 2), tarsus of left pedipalp. C Paratype male (# 2), tarsus of right pedipalp. D Paratype male (# 3), tibia and tarsus of left pedipalp. Labels: see text. Scale bars: 1 mm." captionText-2="FIGURE 3. Reproductive structures. A – B Detail of male gonopods, right gonopod enhanced by white lines (A ventral view, B dorsal view). C Female gonopods, dorsal view, gonopod enhanced by white lines. D Intact spermatophore, frontal view. E Intact spermatophore, lateral view. Arrow indicates the direction from which the female approaches the spermatophore. F Emptied spermatophore, frontal view. G Emptied spermatophore, lateral view. Arrowheads indicate former placement of the seminal packages. Scale bars: A – C 0.2 mm, D – G 0.5 mm." captionText-3="FIGURE 4. Photographs of living adult Stygophrynus orientalis sp. nov. individuals in captivity. A female (above) and male (below). B Two adult males exhibiting agonistic behavior. C Adult female with protonymphs (shortly after moulting)." captionText-4="FIGURE 5. Cerotegument of Charinidae and Charontidae (after Wolff et al. 2017). A – C Charon cf. grayi (Negros Island, Philippines), D – F Stygophrynus orientalis sp. nov., G – I Sarax curioi (Pany Island, Philippines). SEM of air dried exuvia of carapace. A, B, D, E, G, H globular surface. C, F, I detail of colloidal particles. For interpretations see text. Scale bars: see figure." httpUri-0="https://zenodo.org/record/312558/files/figure.png" httpUri-1="https://zenodo.org/record/312577/files/figure.png" httpUri-2="https://zenodo.org/record/312600/files/figure.png" httpUri-3="https://zenodo.org/record/312616/files/figure.png" httpUri-4="https://zenodo.org/record/312679/files/figure.png" pageId="2" pageNumber="399">Fig. 1–5</figureCitation>
|
||
)
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="6CB05FCAFFB3FFCCBBF09C67B46CFE57" pageId="2" pageNumber="399" type="materials_examined">
|
||
<paragraph id="24150C41FFB3FFCCBBF09C67B46CFE57" blockId="2.[151,1437,258,1795]" pageId="2" pageNumber="399">
|
||
<emphasis id="16DED053FFB3FFCCBBF09C67B743FECB" bold="true" box="[151,331,258,283]" pageId="2" pageNumber="399">Type material.</emphasis>
|
||
Holotype: Adult female (NHMW 21944). Paratypes: Adult male (NHMW 21945); 5 Adult females, 5 adult males, 8 subadult specimens (NHMW 21946);
|
||
<materialsCitation id="94C2061CFFB3FFCCB8069C42B434FEB3" ID-GBIF-Occurrence="1434655566" collectionCode="SMNS" country="Indonesia" location="Central Sulawesi province" pageId="2" pageNumber="399" specimenCount="6" specimenCount-adult="3" specimenCount-juvenile="3" stateProvince="Central Sulawesi province">
|
||
<specimenCount id="32ACC7C8FFB3FFCCB8069C42B5A7FEEF" box="[865,943,294,319]" pageId="2" pageNumber="399" type="adult">2 adult</specimenCount>
|
||
females,
|
||
<specimenCount id="32ACC7C8FFB3FFCCBF7C9C42B260FEEF" box="[1051,1128,294,319]" pageId="2" pageNumber="399" type="adult">1 adult</specimenCount>
|
||
male,
|
||
<specimenCount id="32ACC7C8FFB3FFCCBFD39C42B326FEEF" box="[1204,1326,294,319]" pageId="2" pageNumber="399" type="juvenile">3 juveniles</specimenCount>
|
||
(
|
||
<collectionCode id="42BB9484FFB3FFCCBE599C42B399FEEF" box="[1342,1425,295,319]" country="Germany" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34840" name="Staatliches Museum fuer Naturkund Stuttgart" pageId="2" pageNumber="399">SMNS</collectionCode>
|
||
):
|
||
<collectingCountry id="5CBD4CD1FFB3FFCCBBF09C2FB70FFEB3" box="[151,263,330,355]" name="Indonesia" pageId="2" pageNumber="399">Indonesia</collectingCountry>
|
||
,
|
||
<collectingRegion id="E66EC2A3FFB3FFCCBA759C2FB434FEB3" box="[274,572,330,355]" country="Indonesia" name="Sulawesi Tengah" pageId="2" pageNumber="399">Central Sulawesi province</collectingRegion>
|
||
</materialsCitation>
|
||
,
|
||
<materialsCitation id="94C2061CFFB3FFCCB9209C2FB46CFE57" ID-GBIF-Occurrence="1434655567" collectingDate="2013-10" collectorName="P. Grabowitz." country="Iceland" location="Island" pageId="2" pageNumber="399" specimenCount="1">
|
||
<collectingCountry id="5CBD4CD1FFB3FFCCB9209C2FB483FEB3" box="[583,651,330,355]" name="Iceland" pageId="2" pageNumber="399">Island</collectingCountry>
|
||
Banggai (Pulau Banggai), all individuals were collected close to each other in
|
||
<date id="50142A81FFB3FFCCBB939C0BB798FE57" box="[244,400,366,391]" pageId="2" pageNumber="399" value="2013-10">
|
||
<collectingDate id="4050D369FFB3FFCCBB939C0BB798FE57" box="[244,400,366,391]" pageId="2" pageNumber="399" value="2013-10">October 2013</collectingDate>
|
||
</date>
|
||
,
|
||
<emphasis id="16DED053FFB3FFCCBAFC9C0BB7CDFE57" box="[411,453,366,391]" italics="true" pageId="2" pageNumber="399">leg.</emphasis>
|
||
<collectorName id="895F6997FFB3FFCCBAAB9C0AB46CFE57" box="[460,612,366,391]" pageId="2" pageNumber="399">P. Grabowitz.</collectorName>
|
||
</materialsCitation>
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="6CB05FCAFFB3FFCCBBA09CF7B777FBE3" pageId="2" pageNumber="399" type="diagnosis">
|
||
<paragraph id="24150C41FFB3FFCCBBA09CF7B777FBE3" blockId="2.[151,1437,258,1795]" pageId="2" pageNumber="399">
|
||
<emphasis id="16DED053FFB3FFCCBBA09CF7B7ECFE7B" bold="true" box="[199,484,402,427]" pageId="2" pageNumber="399">Comparative diagnosis.</emphasis>
|
||
<taxonomicName id="E3AA77C2FFB3FFCCBA8A9CF7B4F2FE7B" box="[493,762,402,427]" class="Arachnida" family="Charontidae" genus="Stygophrynus" kingdom="Animalia" order="Amblypygi" pageId="2" pageNumber="399" phylum="Arthropoda" rank="species" species="orientalis" status="sp. nov.">
|
||
<emphasis id="16DED053FFB3FFCCBA8A9CF7B4F2FE7B" box="[493,762,402,427]" italics="true" pageId="2" pageNumber="399">Stygophrynus orientalis</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="16DED053FFB3FFCCB8639CF7B569FE7B" bold="true" box="[772,865,402,427]" pageId="2" pageNumber="399">
|
||
<taxonomicNameLabel id="0DED6D28FFB3FFCCB8639CF7B569FE7B" box="[772,865,402,427]" pageId="2" pageNumber="399" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
is easily distinguishable from
|
||
<taxonomicName id="E3AA77C2FFB3FFCCBFA09CF6B342FE7B" box="[1223,1354,402,427]" class="Arachnida" family="Charontidae" genus="Stygophrynus" kingdom="Animalia" order="Amblypygi" pageId="2" pageNumber="399" phylum="Arthropoda" rank="species" species="moultoni">
|
||
<emphasis id="16DED053FFB3FFCCBFA09CF6B342FE7B" box="[1223,1354,402,427]" italics="true" pageId="2" pageNumber="399">S. moultoni</emphasis>
|
||
</taxonomicName>
|
||
by the presence of only one large spine with a smaller subsidiary basal spine on the antero-dorsal margin of the pedipalp basitarsus. Two big and well separated spines of equal size can be found antero-dorsal margin of the basitarsus in
|
||
<taxonomicName id="E3AA77C2FFB3FFCCBEE09CBEB6F3FDC7" class="Arachnida" family="Charontidae" genus="Stygophrynus" kingdom="Animalia" order="Amblypygi" pageId="2" pageNumber="399" phylum="Arthropoda" rank="species" species="moultoni">
|
||
<emphasis id="16DED053FFB3FFCCBEE09CBEB6F3FDC7" italics="true" pageId="2" pageNumber="399">S. moultoni</emphasis>
|
||
</taxonomicName>
|
||
.
|
||
<taxonomicName id="E3AA77C2FFB3FFCCBA6F9C9BB419FDC7" box="[264,529,510,535]" class="Arachnida" family="Charontidae" genus="Stygophrynus" kingdom="Animalia" order="Amblypygi" pageId="2" pageNumber="399" phylum="Arthropoda" rank="species" species="orientalis" status="sp. nov.">
|
||
<emphasis id="16DED053FFB3FFCCBA6F9C9BB419FDC7" box="[264,529,510,535]" italics="true" pageId="2" pageNumber="399">Stygophrynus orientalis</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="16DED053FFB3FFCCB97F9C9BB479FDC7" bold="true" box="[536,625,510,535]" pageId="2" pageNumber="399">
|
||
<taxonomicNameLabel id="0DED6D28FFB3FFCCB97F9C9BB479FDC7" box="[536,625,510,535]" pageId="2" pageNumber="399" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
differs from
|
||
<taxonomicName id="E3AA77C2FFB3FFCCB8639C9BB2C7FDC7" authority="Weygoldt, 2002" authorityName="Weygoldt" authorityYear="2002" box="[772,1231,510,535]" class="Arachnida" family="Charontidae" genus="Stygophrynus" kingdom="Animalia" order="Amblypygi" pageId="2" pageNumber="399" phylum="Arthropoda" rank="species" species="brevispina">
|
||
<emphasis id="16DED053FFB3FFCCB8639C9BB21FFDC7" box="[772,1047,510,535]" italics="true" pageId="2" pageNumber="399">Stygophrynus brevispina</emphasis>
|
||
<bibRefCitation id="403B71B0FFB3FFCCBF7A9C9BB2C7FDC7" author="Weygoldt" box="[1053,1231,510,535]" pageId="2" pageNumber="399" refString="Weygoldt, P. (2002) Sperm transfer and spermatophore morphology of the whip spiders Sarax buxtoni, S. brachydactylus (Charinidae), Charon cf. grayi, and Stygophrynus brevispina nov. spec. (Charontidae) (Chelicerata, Amblypygi). Zoologischer Anzeiger, 241 (2), 131 - 148." type="journal article" year="2002">Weygoldt, 2002</bibRefCitation>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="E3AA77C2FFB3FFCCBE639C9BB799FDEB" authority="Simon, 1901" authorityName="Simon" authorityYear="1901" class="Arachnida" family="Charontidae" genus="Stygophrynus" kingdom="Animalia" order="Amblypygi" pageId="2" pageNumber="399" phylum="Arthropoda" rank="species" species="cerberus">
|
||
<emphasis id="16DED053FFB3FFCCBE639C9BB6F1FDEB" italics="true" pageId="2" pageNumber="399">Stygophrynus cerberus</emphasis>
|
||
<bibRefCitation id="403B71B0FFB3FFCCBA669F47B799FDEB" author="Simon" box="[257,401,546,571]" pageId="2" pageNumber="399" refString="Simon, E. (1901) On the Arachnida collected during the " Skeat Expedition " to the Malay Peninsula, 1899 - 1900. Proceedings of the Zoological Society of London, 71, 45 - 84." type="journal article" year="1901">
|
||
Simon,
|
||
<quantity id="E352A1A4FFB3FFCCBA3E9F46B799FDEB" box="[345,401,547,571]" metricMagnitude="1" metricUnit="m" metricValue="4.828539999999999" pageId="2" pageNumber="399" unit="in" value="1901.0">1901</quantity>
|
||
</bibRefCitation>
|
||
</taxonomicName>
|
||
in the count of dorsal and ventral spines of the pedipalp femur.
|
||
<taxonomicName id="E3AA77C2FFB3FFCCBF0F9F47B37BFDEB" box="[1128,1395,546,571]" class="Arachnida" family="Charontidae" genus="Stygophrynus" kingdom="Animalia" order="Amblypygi" pageId="2" pageNumber="399" phylum="Arthropoda" rank="species" species="orientalis" status="sp. nov.">
|
||
<emphasis id="16DED053FFB3FFCCBF0F9F47B37BFDEB" box="[1128,1395,546,571]" italics="true" pageId="2" pageNumber="399">Stygophrynus orientalis</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="16DED053FFB3FFCCBE1D9F47B6C0FD8F" bold="true" pageId="2" pageNumber="399">
|
||
<taxonomicNameLabel id="0DED6D28FFB3FFCCBE1D9F47B6C0FD8F" pageId="2" pageNumber="399" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
has four of these spines while females of
|
||
<taxonomicName id="E3AA77C2FFB3FFCCB9FB9F22B524FD8F" box="[668,812,582,607]" class="Arachnida" family="Charontidae" genus="Stygophrynus" kingdom="Animalia" order="Amblypygi" pageId="2" pageNumber="399" phylum="Arthropoda" rank="species" species="brevispina">
|
||
<emphasis id="16DED053FFB3FFCCB9FB9F22B524FD8F" box="[668,812,582,607]" italics="true" pageId="2" pageNumber="399">S. brevispina</emphasis>
|
||
</taxonomicName>
|
||
have five dorsal and five ventral spines and
|
||
<taxonomicName id="E3AA77C2FFB3FFCCBE799F22B394FD8F" box="[1310,1436,582,607]" class="Arachnida" family="Charontidae" genus="Stygophrynus" kingdom="Animalia" order="Amblypygi" pageId="2" pageNumber="399" phylum="Arthropoda" rank="species" species="cerberus">
|
||
<emphasis id="16DED053FFB3FFCCBE799F22B394FD8F" box="[1310,1436,582,607]" italics="true" pageId="2" pageNumber="399">S. cerberus</emphasis>
|
||
</taxonomicName>
|
||
harbors five dorsal spines. Furthermore, in adult males of
|
||
<taxonomicName id="E3AA77C2FFB3FFCCB8439F0EB5BEFD53" box="[804,950,618,643]" class="Arachnida" family="Charontidae" genus="Stygophrynus" kingdom="Animalia" order="Amblypygi" pageId="2" pageNumber="399" phylum="Arthropoda" rank="species" species="brevispina">
|
||
<emphasis id="16DED053FFB3FFCCB8439F0EB5BEFD53" box="[804,950,618,643]" italics="true" pageId="2" pageNumber="399">S. brevispina</emphasis>
|
||
</taxonomicName>
|
||
not only
|
||
<emphasis id="16DED053FFB3FFCCBF459F0FB24BFD53" bold="true" box="[1058,1091,618,643]" pageId="2" pageNumber="399">T4</emphasis>
|
||
is strongly reduced in size but also
|
||
<emphasis id="16DED053FFB3FFCCBBAC9FEBB6E3FD77" bold="true" box="[203,235,654,679]" pageId="2" pageNumber="399">T3</emphasis>
|
||
. The cheliceral dentition allows distinguishing
|
||
<taxonomicName id="E3AA77C2FFB3FFCCB9989FEAB58FFD77" box="[767,903,654,679]" class="Arachnida" family="Charontidae" genus="Stygophrynus" kingdom="Animalia" order="Amblypygi" pageId="2" pageNumber="399" phylum="Arthropoda" rank="species" species="orientalis" status="sp. nov.">
|
||
<emphasis id="16DED053FFB3FFCCB9989FEAB58FFD77" box="[767,903,654,679]" italics="true" pageId="2" pageNumber="399">S. orientalis</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="16DED053FFB3FFCCB8EA9FEBB5EEFD77" bold="true" box="[909,998,654,679]" pageId="2" pageNumber="399">
|
||
<taxonomicNameLabel id="0DED6D28FFB3FFCCB8EA9FEBB5EEFD77" box="[909,998,654,679]" pageId="2" pageNumber="399" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
from
|
||
<taxonomicName id="E3AA77C2FFB3FFCCBF4D9FEAB281FD77" box="[1066,1161,654,679]" class="Arachnida" family="Charontidae" genus="Stygophrynus" kingdom="Animalia" order="Amblypygi" pageId="2" pageNumber="399" phylum="Arthropoda" rank="species" species="sunda">
|
||
<emphasis id="16DED053FFB3FFCCBF4D9FEAB281FD77" box="[1066,1161,654,679]" italics="true" pageId="2" pageNumber="399">S. sunda</emphasis>
|
||
</taxonomicName>
|
||
. In
|
||
<taxonomicName id="E3AA77C2FFB3FFCCBFD39FEAB334FD77" box="[1204,1340,654,679]" class="Arachnida" family="Charontidae" genus="Stygophrynus" kingdom="Animalia" order="Amblypygi" pageId="2" pageNumber="399" phylum="Arthropoda" rank="species" species="orientalis" status="sp. nov.">
|
||
<emphasis id="16DED053FFB3FFCCBFD39FEAB334FD77" box="[1204,1340,654,679]" italics="true" pageId="2" pageNumber="399">S. orientalis</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="16DED053FFB3FFCCBE249FEBB394FD77" bold="true" box="[1347,1436,654,679]" pageId="2" pageNumber="399">
|
||
<taxonomicNameLabel id="0DED6D28FFB3FFCCBE249FEBB394FD77" box="[1347,1436,654,679]" pageId="2" pageNumber="399" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
the uppermost tooth of the external row of teeth is bicuspid while it is not in
|
||
<taxonomicName id="E3AA77C2FFB3FFCCBF769FD6B27AFD1B" box="[1041,1138,690,715]" class="Arachnida" family="Charontidae" genus="Stygophrynus" kingdom="Animalia" order="Amblypygi" pageId="2" pageNumber="399" phylum="Arthropoda" rank="species" species="sunda">
|
||
<emphasis id="16DED053FFB3FFCCBF769FD6B27AFD1B" box="[1041,1138,690,715]" italics="true" pageId="2" pageNumber="399">S. sunda</emphasis>
|
||
</taxonomicName>
|
||
, furthermore the movable cheliceral finger harbors five teeth in
|
||
<taxonomicName id="E3AA77C2FFB3FFCCB95E9FB2B4C8FD3F" box="[569,704,726,751]" class="Arachnida" family="Charontidae" genus="Stygophrynus" kingdom="Animalia" order="Amblypygi" pageId="2" pageNumber="399" phylum="Arthropoda" rank="species" species="orientalis" status="sp. nov.">
|
||
<emphasis id="16DED053FFB3FFCCB95E9FB2B4C8FD3F" box="[569,704,726,751]" italics="true" pageId="2" pageNumber="399">S. orientalis</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="16DED053FFB3FFCCB9A19FB3B517FD3F" bold="true" box="[710,799,726,751]" pageId="2" pageNumber="399">
|
||
<taxonomicNameLabel id="0DED6D28FFB3FFCCB9A19FB3B517FD3F" box="[710,799,726,751]" pageId="2" pageNumber="399" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
and six in
|
||
<taxonomicName id="E3AA77C2FFB3FFCCB8FF9FB2B5FFFD3F" box="[920,1015,726,751]" class="Arachnida" family="Charontidae" genus="Stygophrynus" kingdom="Animalia" order="Amblypygi" pageId="2" pageNumber="399" phylum="Arthropoda" rank="species" species="sunda">
|
||
<emphasis id="16DED053FFB3FFCCB8FF9FB2B5FFFD3F" box="[920,1015,726,751]" italics="true" pageId="2" pageNumber="399">S. sunda</emphasis>
|
||
</taxonomicName>
|
||
. In
|
||
<taxonomicName id="E3AA77C2FFB3FFCCBF459FB2B2D9FD3F" box="[1058,1233,726,751]" class="Arachnida" family="Charontidae" genus="Stygophrynus" kingdom="Animalia" order="Amblypygi" pageId="2" pageNumber="399" phylum="Arthropoda" rank="species" species="dammermani">
|
||
<emphasis id="16DED053FFB3FFCCBF459FB2B2D9FD3F" box="[1058,1233,726,751]" italics="true" pageId="2" pageNumber="399">S. dammermani</emphasis>
|
||
</taxonomicName>
|
||
the dorsal surface of the chelicera is smooth, and in
|
||
<taxonomicName id="E3AA77C2FFB3FFCCB9569F9FB5E2FCC3" authority="Gravely, 1915" authorityName="Gravely" authorityYear="1915" box="[561,1002,762,787]" class="Arachnida" family="Charontidae" genus="Stygophrynus" kingdom="Animalia" order="Amblypygi" pageId="2" pageNumber="399" phylum="Arthropoda" rank="species" species="berkeleyi">
|
||
<emphasis id="16DED053FFB3FFCCB9569F9FB534FCC3" box="[561,828,762,787]" italics="true" pageId="2" pageNumber="399">Stygophrynus berkeleyi</emphasis>
|
||
<bibRefCitation id="403B71B0FFB3FFCCB8209F9FB5E2FCC3" author="Gravely" box="[839,1002,762,787]" pageId="2" pageNumber="399" refString="Gravely, F. H. (1915) A revision of the Oriental subfamilies of Tarantulidae (order Pedipalpi). Records of the Indian Museum, 11, 433 - 455." type="journal article" year="1915">Gravely, 1915</bibRefCitation>
|
||
</taxonomicName>
|
||
it is roughened with small tubercles whereas, it is finely roughened with and harbors eight big tubercles in
|
||
<taxonomicName id="E3AA77C2FFB3FFCCB89C9E7AB282FCE7" box="[1019,1162,798,823]" class="Arachnida" family="Charontidae" genus="Stygophrynus" kingdom="Animalia" order="Amblypygi" pageId="2" pageNumber="399" phylum="Arthropoda" rank="species" species="orientalis" status="sp. nov.">
|
||
<emphasis id="16DED053FFB3FFCCB89C9E7AB282FCE7" box="[1019,1162,798,823]" italics="true" pageId="2" pageNumber="399">S. orientalis</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="16DED053FFB3FFCCBFF09E7BB2FFFCE7" bold="true" box="[1175,1271,798,823]" pageId="2" pageNumber="399">
|
||
<taxonomicNameLabel id="0DED6D28FFB3FFCCBFF09E7BB2FFFCE7" box="[1175,1271,798,823]" pageId="2" pageNumber="399" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
<taxonomicName id="E3AA77C2FFB3FFCCBE639E7BB7DEFC8B" authority="Thorell, 1889" authorityName="Thorell" authorityYear="1889" class="Arachnida" family="Charontidae" genus="Stygophrynus" kingdom="Animalia" order="Amblypygi" pageId="2" pageNumber="399" phylum="Arthropoda" rank="species" species="cavernicola">
|
||
<emphasis id="16DED053FFB3FFCCBE639E7BB713FC8B" italics="true" pageId="2" pageNumber="399">Stygophrynus cavernicola</emphasis>
|
||
(
|
||
<bibRefCitation id="403B71B0FFB3FFCCBA499E27B7C7FC8B" author="Thorell" box="[302,463,834,859]" pageId="2" pageNumber="399" refString="Thorell, T. (1889) Aracnidi Artrogastri Birmani raccolti da L. Fea nel 1885 - 1887. Annali del Museo Civico di Storia Naturale di Genova, Series 2, 7, 521 - 729." type="journal article" year="1889">Thorell, 1889</bibRefCitation>
|
||
)
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="E3AA77C2FFB3FFCCB9719E27B5D5FC8B" authority="Gravely, 1915" authorityName="Gravely" authorityYear="1915" box="[534,989,834,859]" class="Arachnida" family="Charontidae" genus="Stygophrynus" kingdom="Animalia" order="Amblypygi" pageId="2" pageNumber="399" phylum="Arthropoda" rank="species" species="longispina">
|
||
<emphasis id="16DED053FFB3FFCCB9719E27B527FC8B" box="[534,815,834,859]" italics="true" pageId="2" pageNumber="399">Stygophrynus longispina</emphasis>
|
||
<bibRefCitation id="403B71B0FFB3FFCCB85C9E27B5D5FC8B" author="Gravely" box="[827,989,834,859]" pageId="2" pageNumber="399" refString="Gravely, F. H. (1915) A revision of the Oriental subfamilies of Tarantulidae (order Pedipalpi). Records of the Indian Museum, 11, 433 - 455." type="journal article" year="1915">Gravely, 1915</bibRefCitation>
|
||
</taxonomicName>
|
||
exhibit five to six contiguous ventral spines distally of the large submedial ventral spine. Four contiguous spines can be found, respectively, on the dorso- and the ventrodistal margin of the pedipalp basitarsus of
|
||
<taxonomicName id="E3AA77C2FFB3FFCCB8FE9EEEB246FC73" box="[921,1102,906,931]" class="Arachnida" family="Charontidae" genus="Stygophrynus" kingdom="Animalia" order="Amblypygi" pageId="2" pageNumber="399" phylum="Arthropoda" rank="species" species="dammermani">
|
||
<emphasis id="16DED053FFB3FFCCB8FE9EEEB246FC73" box="[921,1102,906,931]" italics="true" pageId="2" pageNumber="399">S. dammermani</emphasis>
|
||
</taxonomicName>
|
||
. In contrast none of the 26 examined specimens of
|
||
<taxonomicName id="E3AA77C2FFB3FFCCBAC89ECAB432FC17" box="[431,570,942,967]" class="Arachnida" family="Charontidae" genus="Stygophrynus" kingdom="Animalia" order="Amblypygi" pageId="2" pageNumber="399" phylum="Arthropoda" rank="species" species="orientalis" status="sp. nov.">
|
||
<emphasis id="16DED053FFB3FFCCBAC89ECAB432FC17" box="[431,570,942,967]" italics="true" pageId="2" pageNumber="399">S. orientalis</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="16DED053FFB3FFCCB9229ECBB4AAFC17" bold="true" box="[581,674,942,967]" pageId="2" pageNumber="399">
|
||
<taxonomicNameLabel id="0DED6D28FFB3FFCCB9229ECBB4AAFC17" box="[581,674,942,967]" pageId="2" pageNumber="399" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
harbored more than three contiguous spines on the dorso- or the ventrodistal margin.
|
||
<taxonomicName id="E3AA77C2FFB3FFCCBA189EB7B4BAFC3B" box="[383,690,978,1003]" class="Arachnida" family="Charontidae" genus="Stygophrynus" kingdom="Animalia" order="Amblypygi" pageId="2" pageNumber="399" phylum="Arthropoda" rank="species" species="dammermani">
|
||
<emphasis id="16DED053FFB3FFCCBA189EB7B4BAFC3B" box="[383,690,978,1003]" italics="true" pageId="2" pageNumber="399">Stygophrynus dammermani</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="E3AA77C2FFB3FFCCB98E9EB6B540FC3B" box="[745,840,978,1003]" class="Arachnida" family="Charontidae" genus="Stygophrynus" kingdom="Animalia" order="Amblypygi" pageId="2" pageNumber="399" phylum="Arthropoda" rank="species" species="sunda">
|
||
<emphasis id="16DED053FFB3FFCCB98E9EB6B540FC3B" box="[745,840,978,1003]" italics="true" pageId="2" pageNumber="399">S. sunda</emphasis>
|
||
</taxonomicName>
|
||
can both be distinguished from
|
||
<taxonomicName id="E3AA77C2FFB3FFCCBFD49EB6B333FC3B" box="[1203,1339,978,1003]" class="Arachnida" family="Charontidae" genus="Stygophrynus" kingdom="Animalia" order="Amblypygi" pageId="2" pageNumber="399" phylum="Arthropoda" rank="species" species="orientalis" status="sp. nov.">
|
||
<emphasis id="16DED053FFB3FFCCBFD49EB6B333FC3B" box="[1203,1339,978,1003]" italics="true" pageId="2" pageNumber="399">S. orientalis</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="16DED053FFB3FFCCBE259EB7B394FC3B" bold="true" box="[1346,1436,978,1003]" pageId="2" pageNumber="399">
|
||
<taxonomicNameLabel id="0DED6D28FFB3FFCCBE259EB7B394FC3B" box="[1346,1436,978,1003]" pageId="2" pageNumber="399" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
by the count of trichobothria on the distitibia of leg IV, which is
|
||
<quantity id="E352A1A4FFB3FFCCB80E9E92B5AAFBDF" box="[873,930,1014,1039]" metricMagnitude="-1" metricUnit="m" metricValue="5.334" pageId="2" pageNumber="399" unit="in" value="21.0">21 in</quantity>
|
||
<taxonomicName id="E3AA77C2FFB3FFCCB8CE9E92B251FBDF" box="[937,1113,1014,1039]" class="Arachnida" family="Charontidae" genus="Stygophrynus" kingdom="Animalia" order="Amblypygi" pageId="2" pageNumber="399" phylum="Arthropoda" rank="species" species="dammermani">
|
||
<emphasis id="16DED053FFB3FFCCB8CE9E92B251FBDF" box="[937,1113,1014,1039]" italics="true" pageId="2" pageNumber="399">S. dammermani</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<quantity id="E352A1A4FFB3FFCCBF009E92B2A8FBDF" box="[1127,1184,1014,1039]" metricMagnitude="-1" metricUnit="m" metricValue="5.842" pageId="2" pageNumber="399" unit="in" value="23.0">23 in</quantity>
|
||
<taxonomicName id="E3AA77C2FFB3FFCCBFC19E92B30EFBDF" box="[1190,1286,1014,1039]" class="Arachnida" family="Charontidae" genus="Stygophrynus" kingdom="Animalia" order="Amblypygi" pageId="2" pageNumber="399" phylum="Arthropoda" rank="species" species="sunda">
|
||
<emphasis id="16DED053FFB3FFCCBFC19E92B30EFBDF" box="[1190,1286,1014,1039]" italics="true" pageId="2" pageNumber="399">S. sunda</emphasis>
|
||
</taxonomicName>
|
||
against
|
||
<quantity id="E352A1A4FFB3FFCCBE049E92B394FBDF" box="[1379,1436,1014,1039]" metricMagnitude="-1" metricUnit="m" metricValue="5.588" pageId="2" pageNumber="399" unit="in" value="22.0">22 in</quantity>
|
||
<taxonomicName id="E3AA77C2FFB3FFCCBBF0997EB716FBE3" box="[151,286,1050,1075]" class="Arachnida" family="Charontidae" genus="Stygophrynus" kingdom="Animalia" order="Amblypygi" pageId="2" pageNumber="399" phylum="Arthropoda" rank="species" species="orientalis" status="sp. nov.">
|
||
<emphasis id="16DED053FFB3FFCCBBF0997EB716FBE3" box="[151,286,1050,1075]" italics="true" pageId="2" pageNumber="399">S. orientalis</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="16DED053FFB3FFCCBA42997FB777FBE3" bold="true" box="[293,383,1050,1075]" pageId="2" pageNumber="399">
|
||
<taxonomicNameLabel id="0DED6D28FFB3FFCCBA42997FB777FBE3" box="[293,383,1050,1075]" pageId="2" pageNumber="399" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="6CB05FCAFFB3FFC9BBA0995BB47CFF28" lastPageId="7" lastPageNumber="404" pageId="2" pageNumber="399" type="description">
|
||
<paragraph id="24150C41FFB3FFCCBBA0995BB210FBAB" blockId="2.[151,1437,258,1795]" pageId="2" pageNumber="399">
|
||
<emphasis id="16DED053FFB3FFCCBBA0995BB437FB87" bold="true" box="[199,575,1086,1111]" pageId="2" pageNumber="399">
|
||
Description of female
|
||
<typeStatus id="FB11B2E3FFB3FFCCBAB6995BB432FB87" box="[465,570,1086,1111]" pageId="2" pageNumber="399" type="holotype">holotype</typeStatus>
|
||
.
|
||
</emphasis>
|
||
<emphasis id="16DED053FFB3FFCCB920995BB4DAFB87" box="[583,722,1086,1111]" italics="true" pageId="2" pageNumber="399">Color in life</emphasis>
|
||
(
|
||
<figureCitation id="BC9110C4FFB3FFCCB984995BB522FB87" box="[739,810,1086,1111]" captionStart="FIGURE 4" captionStartId="6.[151,250,1539,1561]" captionTargetBox="[151,1436,193,1517]" captionTargetId="figure@6.[151,1436,193,1517]" captionTargetPageId="6" captionText="FIGURE 4. Photographs of living adult Stygophrynus orientalis sp. nov. individuals in captivity. A female (above) and male (below). B Two adult males exhibiting agonistic behavior. C Adult female with protonymphs (shortly after moulting)." httpUri="https://zenodo.org/record/312616/files/figure.png" pageId="2" pageNumber="399">Fig. 4</figureCitation>
|
||
): Carapace, opisthosoma and pedipalps uniformly dark brown to blackish; walking leg segments distal to the patella are lighter colored.
|
||
</paragraph>
|
||
<paragraph id="24150C41FFB3FFCCBBA099E2B778FAFF" blockId="2.[151,1437,258,1795]" pageId="2" pageNumber="399">
|
||
<emphasis id="16DED053FFB3FFCCBBA099E2B73DFB4E" box="[199,309,1159,1182]" italics="true" pageId="2" pageNumber="399">Carapace</emphasis>
|
||
(
|
||
<figureCitation id="BC9110C4FFB3FFCCBA2F99E3B781FB4F" box="[328,393,1158,1183]" captionStart="FIGURE 1" captionStartId="3.[151,250,1857,1879]" captionTargetBox="[252,1295,248,1759]" captionTargetId="figure@3.[199,1344,193,1813]" captionTargetPageId="3" captionText="FIGURE 1. Stygophrynus orientalis sp. nov. A Habitus of female holotype, dorsal and ventral view, respectively. B – D Female chelicera, denticles enhanced with white lines (B internal view; C external view; D external view, close-up on external teeth row). E Pedipalp (left ventral, right dorsal view), spines of femur and tibia numbered. F Distitibia and distal basitibia of walking leg IV, dorsal; trichobothria marked as follows: basitibia (bt = basitibial), distitibia (bf: basofrontal; bc: basocaudal; sbf: subbasofrontal; sbc-x: series caudal trichobothria; stf 1 x: series frontal trichobothria). Note: not all trichobothria are visible on this figure. Scale bars: A 20 mm, B – C 2 mm, D 0.25 mm E 2 mm, F 1 mm." httpUri="https://zenodo.org/record/312558/files/figure.png" pageId="2" pageNumber="399">Fig. 1</figureCitation>
|
||
A, 4, 5D–F): 1.36 times wider than long, finely granular, with setiferous tubercles on the anterior margin. Cerotegument with regular, globular granules with a rough surface structure, based on crystal- and fiber-like particles. Short connecting fibers between granules and the underlying cuticle. Eyes well developed, elevated and large, more pronounced in median eyes; median eyes rounded with small triangular frontal process, visible from above.
|
||
</paragraph>
|
||
<paragraph id="24150C41FFB3FFCCBBA0985FB7D5FA33" blockId="2.[151,1437,258,1795]" pageId="2" pageNumber="399">
|
||
<emphasis id="16DED053FFB3FFCCBBA0985FB73EFA83" box="[199,310,1338,1363]" italics="true" pageId="2" pageNumber="399">Chelicera</emphasis>
|
||
(
|
||
<figureCitation id="BC9110C4FFB3FFCCBA21985FB78EFA83" box="[326,390,1338,1363]" captionStart="FIGURE 1" captionStartId="3.[151,250,1857,1879]" captionTargetBox="[252,1295,248,1759]" captionTargetId="figure@3.[199,1344,193,1813]" captionTargetPageId="3" captionText="FIGURE 1. Stygophrynus orientalis sp. nov. A Habitus of female holotype, dorsal and ventral view, respectively. B – D Female chelicera, denticles enhanced with white lines (B internal view; C external view; D external view, close-up on external teeth row). E Pedipalp (left ventral, right dorsal view), spines of femur and tibia numbered. F Distitibia and distal basitibia of walking leg IV, dorsal; trichobothria marked as follows: basitibia (bt = basitibial), distitibia (bf: basofrontal; bc: basocaudal; sbf: subbasofrontal; sbc-x: series caudal trichobothria; stf 1 x: series frontal trichobothria). Note: not all trichobothria are visible on this figure. Scale bars: A 20 mm, B – C 2 mm, D 0.25 mm E 2 mm, F 1 mm." httpUri="https://zenodo.org/record/312558/files/figure.png" pageId="2" pageNumber="399">Fig. 1</figureCitation>
|
||
B–D): Dorsal surface finely roughened with eight big tubercles, each with a setum, internal teeth row with four denticles, the lower tooth the biggest, the most dorsal tooth bicuspid of which the lower cusp is the larger one; external row with two teeth, the uppermost tooth bicuspid. Claw (movable finger) with five teeth, the proximal tooth the biggest, following teeth decreasing in size distally; teeth flattened and blunt. Cheliceral surface with many fine hairs.
|
||
</paragraph>
|
||
<paragraph id="24150C41FFB3FFCCBBA0988AB571F99E" blockId="2.[151,1437,258,1795]" pageId="2" pageNumber="399">
|
||
<emphasis id="16DED053FFB3FFCCBBA0988AB72CF9D6" box="[199,292,1519,1542]" italics="true" pageId="2" pageNumber="399">Sternum</emphasis>
|
||
(
|
||
<figureCitation id="BC9110C4FFB3FFCCBA50988BB773F9D7" box="[311,379,1518,1543]" captionStart="FIGURE 1" captionStartId="3.[151,250,1857,1879]" captionTargetBox="[252,1295,248,1759]" captionTargetId="figure@3.[199,1344,193,1813]" captionTargetPageId="3" captionText="FIGURE 1. Stygophrynus orientalis sp. nov. A Habitus of female holotype, dorsal and ventral view, respectively. B – D Female chelicera, denticles enhanced with white lines (B internal view; C external view; D external view, close-up on external teeth row). E Pedipalp (left ventral, right dorsal view), spines of femur and tibia numbered. F Distitibia and distal basitibia of walking leg IV, dorsal; trichobothria marked as follows: basitibia (bt = basitibial), distitibia (bf: basofrontal; bc: basocaudal; sbf: subbasofrontal; sbc-x: series caudal trichobothria; stf 1 x: series frontal trichobothria). Note: not all trichobothria are visible on this figure. Scale bars: A 20 mm, B – C 2 mm, D 0.25 mm E 2 mm, F 1 mm." httpUri="https://zenodo.org/record/312558/files/figure.png" pageId="2" pageNumber="399">Fig. 1</figureCitation>
|
||
A): The first sternite (tritosternum) elongated with paired apical setae, stretching between pedipalp coxae and several other lateral setae facing distally; second and third sternites small and rounded with small setae; fourth sternite (metasternum) unpaired with six setae.
|
||
</paragraph>
|
||
<paragraph id="24150C41FFB3FFCBBBA09B3FB716F8EE" blockId="2.[151,1437,258,1795]" lastBlockId="5.[151,1437,1325,2034]" lastPageId="5" lastPageNumber="402" pageId="2" pageNumber="399">
|
||
<emphasis id="16DED053FFB3FFCCBBA09B3FB73FF9A3" box="[199,311,1626,1651]" italics="true" pageId="2" pageNumber="399">Pedipalps</emphasis>
|
||
(
|
||
<figureCitation id="BC9110C4FFB3FFCCBA2E9B3FB781F9A3" box="[329,393,1626,1651]" captionStart="FIGURE 1" captionStartId="3.[151,250,1857,1879]" captionTargetBox="[252,1295,248,1759]" captionTargetId="figure@3.[199,1344,193,1813]" captionTargetPageId="3" captionText="FIGURE 1. Stygophrynus orientalis sp. nov. A Habitus of female holotype, dorsal and ventral view, respectively. B – D Female chelicera, denticles enhanced with white lines (B internal view; C external view; D external view, close-up on external teeth row). E Pedipalp (left ventral, right dorsal view), spines of femur and tibia numbered. F Distitibia and distal basitibia of walking leg IV, dorsal; trichobothria marked as follows: basitibia (bt = basitibial), distitibia (bf: basofrontal; bc: basocaudal; sbf: subbasofrontal; sbc-x: series caudal trichobothria; stf 1 x: series frontal trichobothria). Note: not all trichobothria are visible on this figure. Scale bars: A 20 mm, B – C 2 mm, D 0.25 mm E 2 mm, F 1 mm." httpUri="https://zenodo.org/record/312558/files/figure.png" pageId="2" pageNumber="399">Fig. 1</figureCitation>
|
||
A, E, 2A):
|
||
<emphasis id="16DED053FFB3FFCCB9779B3FB484F9A3" box="[528,652,1626,1651]" italics="true" pageId="2" pageNumber="399">Trochanter</emphasis>
|
||
with twelve spines and several setiferous tubercles on antero-dorsal margin, one prominent large spine on the antero-ventral margin.
|
||
<emphasis id="16DED053FFB3FFCCB8E49B1AB5C6F946" box="[899,974,1663,1686]" italics="true" pageId="2" pageNumber="399">Femur</emphasis>
|
||
with four major spines and many small denticles on antero-dorsal margin,
|
||
<emphasis id="16DED053FFB3FFCCB97E9BC7B430F96B" bold="true" box="[537,568,1698,1723]" pageId="2" pageNumber="399">F2</emphasis>
|
||
largest one (
|
||
<emphasis id="16DED053FFB3FFCCB9A09BC7B592F96B" bold="true" box="[711,922,1698,1723]" pageId="2" pageNumber="399">F2> F3> F1> F4</emphasis>
|
||
), one small spine between
|
||
<emphasis id="16DED053FFB3FFCCBFA09BC7B31CF96B" bold="true" box="[1223,1300,1698,1723]" pageId="2" pageNumber="399">F2–F3</emphasis>
|
||
and
|
||
<emphasis id="16DED053FFB3FFCCBE2E9BC7B39DF96B" bold="true" box="[1353,1429,1698,1723]" pageId="2" pageNumber="399">F3–F4</emphasis>
|
||
, two smaller spines internally close to the margin of trochanter, four major spines and four small spines on the anteroventral margin (
|
||
<emphasis id="16DED053FFB3FFCCBAF69B8FB491F8D3" bold="true" box="[401,665,1770,1795]" pageId="2" pageNumber="399">FIII> FII> FIV> FI)</emphasis>
|
||
, between
|
||
<emphasis id="16DED053FFB3FFCCB86C9B8FB56FF8D3" bold="true" box="[779,871,1770,1795]" pageId="2" pageNumber="399">FII–FII</emphasis>
|
||
and
|
||
<emphasis id="16DED053FFB3FFCCB8FA9B8FB205F8D3" bold="true" box="[925,1037,1770,1795]" pageId="2" pageNumber="399">FIII–FIV</emphasis>
|
||
one smaller spine more internally, the rest with several small spines in between,
|
||
<emphasis id="16DED053FFB4FFCBB9FC9848B4C3FA96" bold="true" box="[667,715,1325,1350]" pageId="5" pageNumber="402">FIV</emphasis>
|
||
closer to the ventral surface than the rest.
|
||
<emphasis id="16DED053FFB4FFCBBFCD9848B2EBFA96" box="[1194,1251,1325,1350]" italics="true" pageId="5" pageNumber="402">Tibia</emphasis>
|
||
with four major dorsal spines,
|
||
<emphasis id="16DED053FFB4FFCBBA529834B75EFABA" bold="true" box="[309,342,1361,1386]" pageId="5" pageNumber="402">T2</emphasis>
|
||
largest one (
|
||
<emphasis id="16DED053FFB4FFCBBA829834B4B6FABA" bold="true" box="[485,702,1361,1386]" pageId="5" pageNumber="402">T2> T1> T3> T4</emphasis>
|
||
), one small spine between
|
||
<emphasis id="16DED053FFB4FFCBB88B9834B233FABA" bold="true" box="[1004,1083,1361,1386]" pageId="5" pageNumber="402">T1–T2</emphasis>
|
||
, two small spines between
|
||
<emphasis id="16DED053FFB4FFCBBE0A9834B6BFFA5E" bold="true" pageId="5" pageNumber="402">T3– T4</emphasis>
|
||
, three small spines between
|
||
<emphasis id="16DED053FFB4FFCBB9669810B42AFA5E" bold="true" box="[513,546,1397,1422]" pageId="5" pageNumber="402">T1</emphasis>
|
||
and the distal margin (some specimen only harbors two small spines between
|
||
<emphasis id="16DED053FFB4FFCBBBF098FCB6B0FA62" bold="true" box="[151,184,1433,1458]" pageId="5" pageNumber="402">T1</emphasis>
|
||
and the distal margin on one pedipalp hand; compare
|
||
<tableCitation id="692839FAFFB4FFCBB84698FCB57CFA62" box="[801,884,1433,1458]" captionStart="TABLE 1" captionStartId="8.[119,204,554,576]" captionTargetBox="[119,1909,643,1636]" captionTargetPageId="8" captionText="TABLE 1. Measurements (mm) of female holotype anđ selecteđ paratype specimens. Abbreviations: Peđipalp tibia (T 1 T ³, T 1 the most đistal one); * Number of spines between anđ đistal margin of peđipalp tibia; ** Number of trichobothria on the basitibia anđ đistitibia of leg IV." httpUri="http://table.plazi.org/id/70D55CC9FFB9FFC6BB109F4FB2ABFDBC" pageId="5" pageNumber="402" tableUuid="70D55CC9FFB9FFC6BB109F4FB2ABFDBC">Table 1</tableCitation>
|
||
), most distal one the smallest, four major spines on the antero-ventral margin,
|
||
<emphasis id="16DED053FFB4FFCBBA8198D8B406FA06" bold="true" box="[486,526,1469,1494]" pageId="5" pageNumber="402">TII</emphasis>
|
||
largest one (
|
||
<emphasis id="16DED053FFB4FFCBB9C798D8B5A0FA06" bold="true" box="[672,936,1469,1494]" pageId="5" pageNumber="402">TII> TIII> TIV> TI</emphasis>
|
||
), one small spine between
|
||
<emphasis id="16DED053FFB4FFCBBFBB98D8B338FA06" bold="true" box="[1244,1328,1469,1494]" pageId="5" pageNumber="402">TI–TII</emphasis>
|
||
and
|
||
<emphasis id="16DED053FFB4FFCBBE0098D8B6CDFA2A" bold="true" pageId="5" pageNumber="402">TV– TIV</emphasis>
|
||
,
|
||
<emphasis id="16DED053FFB4FFCBBBB59884B6F8FA2A" bold="true" box="[210,240,1505,1530]" pageId="5" pageNumber="402">TI</emphasis>
|
||
markedly curved.
|
||
<emphasis id="16DED053FFB4FFCBBAA79886B43EFA2A" box="[448,566,1507,1530]" italics="true" pageId="5" pageNumber="402">Basitarsus</emphasis>
|
||
with one large submedial spine on antero-dorsal margin (lsmd-s), which has a subsidiary proximal basal spine (spb-s), one small setiferous tubercle (set) followed by three contiguous spines (cod-s) on the distal margin, these three spines decreasing in size proximally, the most distal the largest one, anteroventral margin with one large spine (lsmv-s) followed by three smaller spines (cov-s) on the distal margin, these three spines decreasing in size proximally, the most distal the largest one, a row of two to three long setae with thickened tips similar to those at proximal end of cleaning organ of the distitarsus at dorsal distal margin.
|
||
<emphasis id="16DED053FFB4FFCBBBF09BDEB71AF902" box="[151,274,1723,1746]" italics="true" pageId="5" pageNumber="402">Distitarsus</emphasis>
|
||
spineless, with large well developed cleaning organ (co), with a row of short dorsal setae (co-ss) and 27 long setae (co-ls) ventrally, cleaning organ with five small cuspid tubercles (co-ct), a row of two to three long setae with thickened tips near proximal end of cleaning organ (rs).
|
||
<emphasis id="16DED053FFB4FFCBB80C9A67B216F8CA" box="[875,1054,1793,1818]" italics="true" pageId="5" pageNumber="402">Pretarsus/ claw</emphasis>
|
||
separated from distitarsus by an articulation.
|
||
</paragraph>
|
||
<caption id="70D55CC9FFB2FFCDBBF09A24B5B9F820" httpUri="https://zenodo.org/record/312558/files/figure.png" pageId="3" pageNumber="400" targetBox="[252,1295,248,1759]" targetPageId="3">
|
||
<paragraph id="24150C41FFB2FFCDBBF09A24B5B9F820" blockId="3.[151,1436,1857,2032]" pageId="3" pageNumber="400">
|
||
<emphasis id="16DED053FFB2FFCDBBF09A24B71BF886" bold="true" box="[151,275,1857,1879]" pageId="3" pageNumber="400">FIGURE 1.</emphasis>
|
||
<taxonomicName id="E3AA77C2FFB2FFCDBA7E9A24B401F886" box="[281,521,1857,1878]" class="Arachnida" family="Charontidae" genus="Stygophrynus" kingdom="Animalia" order="Amblypygi" pageId="3" pageNumber="400" phylum="Arthropoda" rank="species" species="orientalis" status="sp. nov.">
|
||
<emphasis id="16DED053FFB2FFCDBA7E9A24B401F886" box="[281,521,1857,1878]" italics="true" pageId="3" pageNumber="400">Stygophrynus orientalis</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="16DED053FFB2FFCDB9689A24B470F886" bold="true" box="[527,632,1857,1878]" pageId="3" pageNumber="400">
|
||
<taxonomicNameLabel id="0DED6D28FFB2FFCDB9689A24B457F886" box="[527,607,1857,1878]" pageId="3" pageNumber="400" rank="species">sp. nov.</taxonomicNameLabel>
|
||
A
|
||
</emphasis>
|
||
Habitus of female holotype, dorsal and ventral view, respectively.
|
||
<emphasis id="16DED053FFB2FFCDBE7C9A24B343F886" bold="true" box="[1307,1355,1857,1878]" pageId="3" pageNumber="400">B–D</emphasis>
|
||
Female chelicera, denticles enhanced with white lines (
|
||
<emphasis id="16DED053FFB2FFCDB9189A05B498F8A5" bold="true" box="[639,656,1888,1909]" pageId="3" pageNumber="400">B</emphasis>
|
||
internal view;
|
||
<emphasis id="16DED053FFB2FFCDB84A9A3AB537F8A5" bold="true" box="[813,831,1887,1909]" pageId="3" pageNumber="400">C</emphasis>
|
||
external view;
|
||
<emphasis id="16DED053FFB2FFCDB8869A05B5FBF8A5" bold="true" box="[993,1011,1888,1909]" pageId="3" pageNumber="400">D</emphasis>
|
||
external view, close-up on external teeth row).
|
||
<emphasis id="16DED053FFB2FFCDBBB09A1AB6E0F844" bold="true" box="[215,232,1919,1940]" pageId="3" pageNumber="400">E</emphasis>
|
||
Pedipalp (left ventral, right dorsal view), spines of femur and tibia numbered.
|
||
<emphasis id="16DED053FFB2FFCDBF529A1AB24CF844" bold="true" box="[1077,1092,1919,1940]" pageId="3" pageNumber="400">F</emphasis>
|
||
Distitibia and distal basitibia of walking leg IV, dorsal; trichobothria marked as follows: basitibia (bt = basitibial), distitibia (bf: basofrontal; bc: basocaudal; sbf: subbasofrontal; sbc-x: series caudal trichobothria; stf1x: series frontal trichobothria). Note: not all trichobothria are visible on this figure.Scale bars:
|
||
<emphasis id="16DED053FFB2FFCDBAFE9ABFB7A3F83F" bold="true" box="[409,427,2010,2031]" pageId="3" pageNumber="400">A</emphasis>
|
||
20 mm,
|
||
<emphasis id="16DED053FFB2FFCDB9629ABFB43CF820" bold="true" box="[517,564,2010,2032]" pageId="3" pageNumber="400">B–C</emphasis>
|
||
2 mm,
|
||
<emphasis id="16DED053FFB2FFCDB9E59ABFB49CF83F" bold="true" box="[642,660,2010,2031]" pageId="3" pageNumber="400">D</emphasis>
|
||
0.25 mm
|
||
<emphasis id="16DED053FFB2FFCDB99C9ABFB504F83F" bold="true" box="[763,780,2010,2031]" pageId="3" pageNumber="400">E</emphasis>
|
||
2 mm,
|
||
<emphasis id="16DED053FFB2FFCDB83E9ABFB560F83F" bold="true" box="[857,872,2010,2031]" pageId="3" pageNumber="400">F</emphasis>
|
||
1 mm.
|
||
</paragraph>
|
||
</caption>
|
||
<caption id="70D55CC9FFB5FFCABBF09A2FB745F86C" httpUri="https://zenodo.org/record/312577/files/figure.png" pageId="4" pageNumber="401" targetBox="[200,1385,232,1845]" targetPageId="4">
|
||
<paragraph id="24150C41FFB5FFCABBF09A2FB745F86C" blockId="4.[151,1437,1866,1980]" pageId="4" pageNumber="401">
|
||
<emphasis id="16DED053FFB5FFCABBF09A2FB71DF88F" bold="true" box="[151,277,1866,1888]" pageId="4" pageNumber="401">FIGURE 2.</emphasis>
|
||
<taxonomicName id="E3AA77C2FFB5FFCABA7A9A2EB407F8B0" box="[285,527,1867,1888]" class="Arachnida" family="Charontidae" genus="Stygophrynus" kingdom="Animalia" order="Amblypygi" pageId="4" pageNumber="401" phylum="Arthropoda" rank="species" species="orientalis" status="sp. nov.">
|
||
<emphasis id="16DED053FFB5FFCABA7A9A2EB407F8B0" box="[285,527,1867,1888]" italics="true" pageId="4" pageNumber="401">Stygophrynus orientalis</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="16DED053FFB5FFCAB97F9A2EB462F8B0" bold="true" box="[536,618,1867,1888]" pageId="4" pageNumber="401">
|
||
<taxonomicNameLabel id="0DED6D28FFB5FFCAB97F9A2EB462F8B0" box="[536,618,1867,1888]" pageId="4" pageNumber="401" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
Variability of pedipalpal spination. Distal part of pedipalp, pro-lateral view.
|
||
<emphasis id="16DED053FFB5FFCABEED9A2FB394F88F" bold="true" box="[1418,1436,1866,1887]" pageId="4" pageNumber="401">A</emphasis>
|
||
Holotype female (21944), tarsus of right pedipalp, left drawing, right original photograph.
|
||
<emphasis id="16DED053FFB5FFCABF529A0CB24EF8AE" bold="true" box="[1077,1094,1897,1918]" pageId="4" pageNumber="401">B</emphasis>
|
||
Paratype male (#2), tarsus of left pedipalp.
|
||
<emphasis id="16DED053FFB5FFCABB9D9AE2B704F84D" bold="true" box="[250,268,1927,1949]" pageId="4" pageNumber="401">C</emphasis>
|
||
Paratype male (#2), tarsus of right pedipalp.
|
||
<emphasis id="16DED053FFB5FFCAB9B59AEDB4ECF84D" bold="true" box="[722,740,1928,1949]" pageId="4" pageNumber="401">D</emphasis>
|
||
Paratype male (#3), tibia and tarsus of left pedipalp. Labels: see text. Scale bars: 1 mm.
|
||
</paragraph>
|
||
</caption>
|
||
<caption id="70D55CC9FFB4FFCBBBF09910B4E1FAD6" httpUri="https://zenodo.org/record/312600/files/figure.png" pageId="5" pageNumber="402" targetBox="[151,1436,195,1120]" targetPageId="5">
|
||
<paragraph id="24150C41FFB4FFCBBBF09910B4E1FAD6" blockId="5.[151,1436,1141,1286]" pageId="5" pageNumber="402">
|
||
<emphasis id="16DED053FFB4FFCBBBF09910B71AFB5B" bold="true" box="[151,274,1141,1163]" pageId="5" pageNumber="402">FIGURE 3.</emphasis>
|
||
Reproductive structures.
|
||
<emphasis id="16DED053FFB4FFCBB9719913B44EFB5B" bold="true" box="[534,582,1142,1163]" pageId="5" pageNumber="402">A–B</emphasis>
|
||
Detail of male gonopods, right gonopod enhanced by white lines (
|
||
<emphasis id="16DED053FFB4FFCBBF8D9913B2F4FB5B" bold="true" box="[1258,1276,1142,1163]" pageId="5" pageNumber="402">A</emphasis>
|
||
ventral view,
|
||
<emphasis id="16DED053FFB4FFCBBEEC9913B394FB5B" bold="true" box="[1419,1436,1142,1163]" pageId="5" pageNumber="402">B</emphasis>
|
||
dorsal view).
|
||
<emphasis id="16DED053FFB4FFCBBA4099F1B731FB7A" bold="true" box="[295,313,1172,1194]" pageId="5" pageNumber="402">C</emphasis>
|
||
Female gonopods, dorsal view, gonopod enhanced by white lines.
|
||
<emphasis id="16DED053FFB4FFCBB89C99F0B205FB7A" bold="true" box="[1019,1037,1173,1194]" pageId="5" pageNumber="402">D</emphasis>
|
||
Intact spermatophore, frontal view.
|
||
<emphasis id="16DED053FFB4FFCBBEEC99F0B394FB7A" bold="true" box="[1419,1436,1173,1194]" pageId="5" pageNumber="402">E</emphasis>
|
||
Intact spermatophore, lateral view. Arrow indicates the direction from which the female approaches the spermatophore.
|
||
<emphasis id="16DED053FFB4FFCBBEEB99D6B393FB18" bold="true" box="[1420,1435,1203,1224]" pageId="5" pageNumber="402">F</emphasis>
|
||
Emptied spermatophore, frontal view.
|
||
<emphasis id="16DED053FFB4FFCBB94499B4B430FB37" bold="true" box="[547,568,1233,1255]" pageId="5" pageNumber="402">G</emphasis>
|
||
Emptied spermatophore, lateral view. Arrowheads indicate former placement of the seminal packages. Scale bars:
|
||
<emphasis id="16DED053FFB4FFCBBAAD9995B7F3FAD6" bold="true" box="[458,507,1264,1286]" pageId="5" pageNumber="402">A–C</emphasis>
|
||
0.2 mm,
|
||
<emphasis id="16DED053FFB4FFCBB93B9995B486FAD6" bold="true" box="[604,654,1264,1286]" pageId="5" pageNumber="402">D–G</emphasis>
|
||
0.5 mm.
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="24150C41FFB4FFCBBBA09A2EB746F821" blockId="5.[151,1437,1325,2034]" pageId="5" pageNumber="402">
|
||
<emphasis id="16DED053FFB4FFCBBBA09A2EB6F4F8B2" box="[199,252,1867,1890]" italics="true" pageId="5" pageNumber="402">Legs</emphasis>
|
||
(
|
||
<figureCitation id="BC9110C4FFB4FFCBBA6D9A2CB740F8B2" box="[266,328,1865,1890]" captionStart="FIGURE 1" captionStartId="3.[151,250,1857,1879]" captionTargetBox="[252,1295,248,1759]" captionTargetId="figure@3.[199,1344,193,1813]" captionTargetPageId="3" captionText="FIGURE 1. Stygophrynus orientalis sp. nov. A Habitus of female holotype, dorsal and ventral view, respectively. B – D Female chelicera, denticles enhanced with white lines (B internal view; C external view; D external view, close-up on external teeth row). E Pedipalp (left ventral, right dorsal view), spines of femur and tibia numbered. F Distitibia and distal basitibia of walking leg IV, dorsal; trichobothria marked as follows: basitibia (bt = basitibial), distitibia (bf: basofrontal; bc: basocaudal; sbf: subbasofrontal; sbc-x: series caudal trichobothria; stf 1 x: series frontal trichobothria). Note: not all trichobothria are visible on this figure. Scale bars: A 20 mm, B – C 2 mm, D 0.25 mm E 2 mm, F 1 mm." httpUri="https://zenodo.org/record/312558/files/figure.png" pageId="5" pageNumber="402">Fig. 1</figureCitation>
|
||
A, F):
|
||
<emphasis id="16DED053FFB4FFCBBAF29A2FB45CF8B2" box="[405,596,1865,1890]" italics="true" pageId="5" pageNumber="402">Antenniform legs</emphasis>
|
||
with 25 tibial and 46 tarsal articles;
|
||
<emphasis id="16DED053FFB4FFCBB88F9A2CB27CF8B2" box="[1000,1140,1865,1890]" italics="true" pageId="5" pageNumber="402">walking legs</emphasis>
|
||
elongated, basitibia II and III with one segment, basitibia IV divided into four segments, fourth segment with single trichobothrium
|
||
<emphasis id="16DED053FFB4FFCBBE4D9A08B348F856" box="[1322,1344,1901,1926]" italics="true" pageId="5" pageNumber="402">bt</emphasis>
|
||
close to the distal margin, 22 trichobothria on distitibia,
|
||
<emphasis id="16DED053FFB4FFCBB9D79AF4B4C3F87A" box="[688,715,1937,1962]" italics="true" pageId="5" pageNumber="402">bc</emphasis>
|
||
closer to
|
||
<emphasis id="16DED053FFB4FFCBB85E9AF4B552F87A" box="[825,858,1937,1962]" italics="true" pageId="5" pageNumber="402">sbf</emphasis>
|
||
than
|
||
<emphasis id="16DED053FFB4FFCBB8FF9AF4B5A6F87A" box="[920,942,1937,1962]" italics="true" pageId="5" pageNumber="402">bf</emphasis>
|
||
(distance: 1.06 to 0.38 to 0.14 to 0.78,
|
||
<emphasis id="16DED053FFB4FFCBBE0E9AF4B377F87A" box="[1385,1407,1937,1962]" italics="true" pageId="5" pageNumber="402">bt</emphasis>
|
||
to
|
||
<emphasis id="16DED053FFB4FFCBBBF09AD0B6A5F81E" box="[151,173,1973,1998]" italics="true" pageId="5" pageNumber="402">bf</emphasis>
|
||
to
|
||
<emphasis id="16DED053FFB4FFCBBBB39AD0B6E6F81E" box="[212,238,1973,1998]" italics="true" pageId="5" pageNumber="402">bc</emphasis>
|
||
to
|
||
<emphasis id="16DED053FFB4FFCBBA709AD0B730F81E" box="[279,312,1973,1998]" italics="true" pageId="5" pageNumber="402">sbf</emphasis>
|
||
to
|
||
<emphasis id="16DED053FFB4FFCBBA079AD3B780F81E" box="[352,392,1974,1998]" italics="true" pageId="5" pageNumber="402">stf1</emphasis>
|
||
; mm), tarsi with well pronounced light transverse line, tarsal segment without oblique slit, arolium present.
|
||
</paragraph>
|
||
<caption id="70D55CC9FFB7FFC8BBF09B66B349F9E7" httpUri="https://zenodo.org/record/312616/files/figure.png" pageId="6" pageNumber="403" targetBox="[151,1436,193,1517]" targetPageId="6">
|
||
<paragraph id="24150C41FFB7FFC8BBF09B66B349F9E7" blockId="6.[151,1435,1539,1591]" pageId="6" pageNumber="403">
|
||
<emphasis id="16DED053FFB7FFC8BBF09B66B71CF9C8" bold="true" box="[151,276,1539,1561]" pageId="6" pageNumber="403">FIGURE 4.</emphasis>
|
||
Photographs of living adult
|
||
<taxonomicName id="E3AA77C2FFB7FFC8B95D9B66B524F9C8" box="[570,812,1539,1560]" class="Arachnida" family="Charontidae" genus="Stygophrynus" kingdom="Animalia" order="Amblypygi" pageId="6" pageNumber="403" phylum="Arthropoda" rank="species" species="orientalis" status="sp. nov.">
|
||
<emphasis id="16DED053FFB7FFC8B95D9B66B524F9C8" box="[570,812,1539,1560]" italics="true" pageId="6" pageNumber="403">Stygophrynus orientalis</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="0DED6D28FFB7FFC8B8549B66B58CF9C9" box="[819,900,1539,1561]" pageId="6" pageNumber="403" rank="species">
|
||
<emphasis id="16DED053FFB7FFC8B8549B66B577F9C8" bold="true" box="[819,895,1539,1560]" pageId="6" pageNumber="403">sp. nov</emphasis>
|
||
.
|
||
</taxonomicNameLabel>
|
||
individuals in captivity.
|
||
<emphasis id="16DED053FFB7FFC8BFE39B66B29EF9C8" bold="true" box="[1156,1174,1539,1560]" pageId="6" pageNumber="403">A</emphasis>
|
||
female (above) and male (below).
|
||
<emphasis id="16DED053FFB7FFC8BB959B47B70BF9E7" bold="true" box="[242,259,1570,1591]" pageId="6" pageNumber="403">B</emphasis>
|
||
Two adult males exhibiting agonistic behavior.
|
||
<emphasis id="16DED053FFB7FFC8B98E9B44B4F3F9E7" bold="true" box="[745,763,1569,1591]" pageId="6" pageNumber="403">C</emphasis>
|
||
Adult female with protonymphs (shortly after moulting).
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="24150C41FFB7FFC8BBA09B3BB4F6F94B" blockId="6.[151,1437,1630,2015]" pageId="6" pageNumber="403">
|
||
<emphasis id="16DED053FFB7FFC8BBA09B3BB732F9A7" box="[199,314,1630,1655]" italics="true" pageId="6" pageNumber="403">Gonopods</emphasis>
|
||
(
|
||
<figureCitation id="BC9110C4FFB7FFC8BA2D9B3BB785F9A7" box="[330,397,1630,1655]" captionStart="FIGURE 3" captionStartId="5.[151,250,1141,1163]" captionTargetBox="[151,1436,195,1120]" captionTargetId="figure@5.[151,1436,187,1120]" captionTargetPageId="5" captionText="FIGURE 3. Reproductive structures. A – B Detail of male gonopods, right gonopod enhanced by white lines (A ventral view, B dorsal view). C Female gonopods, dorsal view, gonopod enhanced by white lines. D Intact spermatophore, frontal view. E Intact spermatophore, lateral view. Arrow indicates the direction from which the female approaches the spermatophore. F Emptied spermatophore, frontal view. G Emptied spermatophore, lateral view. Arrowheads indicate former placement of the seminal packages. Scale bars: A – C 0.2 mm, D – G 0.5 mm." httpUri="https://zenodo.org/record/312600/files/figure.png" pageId="6" pageNumber="403">Fig. 3</figureCitation>
|
||
C): Soft, cone-shaped with paired finger-like apically pointed projections. Ventrally covered by genital operculum harboring many setae on margin.
|
||
</paragraph>
|
||
<paragraph id="24150C41FFB7FFC8BBA09BC3B2CDF8D7" blockId="6.[151,1437,1630,2015]" pageId="6" pageNumber="403">
|
||
<emphasis id="16DED053FFB7FFC8BBA09BC3B423F96F" bold="true" box="[199,555,1702,1727]" pageId="6" pageNumber="403">
|
||
Description of male
|
||
<typeStatus id="FB11B2E3FFB7FFC8BADE9BC3B42EF96F" box="[441,550,1702,1727]" pageId="6" pageNumber="403" type="paratype">paratype</typeStatus>
|
||
.
|
||
</emphasis>
|
||
<emphasis id="16DED053FFB7FFC8B9559BC3B4D0F96F" box="[562,728,1702,1727]" italics="true" pageId="6" pageNumber="403">Male genitalia</emphasis>
|
||
(
|
||
<figureCitation id="BC9110C4FFB7FFC8B9809BC3B520F96F" box="[743,808,1702,1727]" captionStart="FIGURE 3" captionStartId="5.[151,250,1141,1163]" captionTargetBox="[151,1436,195,1120]" captionTargetId="figure@5.[151,1436,187,1120]" captionTargetPageId="5" captionText="FIGURE 3. Reproductive structures. A – B Detail of male gonopods, right gonopod enhanced by white lines (A ventral view, B dorsal view). C Female gonopods, dorsal view, gonopod enhanced by white lines. D Intact spermatophore, frontal view. E Intact spermatophore, lateral view. Arrow indicates the direction from which the female approaches the spermatophore. F Emptied spermatophore, frontal view. G Emptied spermatophore, lateral view. Arrowheads indicate former placement of the seminal packages. Scale bars: A – C 0.2 mm, D – G 0.5 mm." httpUri="https://zenodo.org/record/312600/files/figure.png" pageId="6" pageNumber="403">Fig. 3</figureCitation>
|
||
A–B): Dark brown sclerotized regions on the distal end of the fistula. The LoL1 and the LoL2 blunt, the LoL1 being one third of the size of the LoL2, posterodorsal LoD smaller than LoL1 and blunt. Spermatophore organ covered ventrally by the genital operculum.
|
||
</paragraph>
|
||
<paragraph id="24150C41FFB7FFC8BBA09A77B395F80F" blockId="6.[151,1437,1630,2015]" pageId="6" pageNumber="403">
|
||
<emphasis id="16DED053FFB7FFC8BBA09A77B77BF8FB" box="[199,371,1810,1835]" italics="true" pageId="6" pageNumber="403">Spermatophore</emphasis>
|
||
(
|
||
<figureCitation id="BC9110C4FFB7FFC8BAE29A77B7C0F8FB" box="[389,456,1810,1835]" captionStart="FIGURE 3" captionStartId="5.[151,250,1141,1163]" captionTargetBox="[151,1436,195,1120]" captionTargetId="figure@5.[151,1436,187,1120]" captionTargetPageId="5" captionText="FIGURE 3. Reproductive structures. A – B Detail of male gonopods, right gonopod enhanced by white lines (A ventral view, B dorsal view). C Female gonopods, dorsal view, gonopod enhanced by white lines. D Intact spermatophore, frontal view. E Intact spermatophore, lateral view. Arrow indicates the direction from which the female approaches the spermatophore. F Emptied spermatophore, frontal view. G Emptied spermatophore, lateral view. Arrowheads indicate former placement of the seminal packages. Scale bars: A – C 0.2 mm, D – G 0.5 mm." httpUri="https://zenodo.org/record/312600/files/figure.png" pageId="6" pageNumber="403">Fig. 3</figureCitation>
|
||
D–G): The spermatophore consists of the stalk, the head and the sperm masses. The complete structure is about
|
||
<quantity id="E352A1A4FFB7FFC8BAB99A53B44EF89F" box="[478,582,1846,1871]" metricMagnitude="-3" metricUnit="m" metricValue="2.68" pageId="6" pageNumber="403" unit="mm" value="2.68">2.68 mm</quantity>
|
||
in height. It bears two protruding sperm masses (
|
||
<emphasis id="16DED053FFB7FFC8BFFF9A5DB2B9F89F" box="[1176,1201,1848,1871]" italics="true" pageId="6" pageNumber="403">sp</emphasis>
|
||
) apically which are connected by a broad strip (
|
||
<emphasis id="16DED053FFB7FFC8BA879A3FB7F1F8A3" box="[480,505,1882,1907]" italics="true" pageId="6" pageNumber="403">bs</emphasis>
|
||
). Two small upwardly directed processes emanate apically of the sperm masses (upper wings;
|
||
<emphasis id="16DED053FFB7FFC8BA599AE5B756F847" box="[318,350,1920,1943]" italics="true" pageId="6" pageNumber="403">uw</emphasis>
|
||
). Lateral horns (
|
||
<emphasis id="16DED053FFB7FFC8B9449A1BB431F847" box="[547,569,1918,1943]" italics="true" pageId="6" pageNumber="403">lh</emphasis>
|
||
) are well developed. Posterior extensions (
|
||
<emphasis id="16DED053FFB7FFC8BF4A9AE5B24FF847" box="[1069,1095,1920,1943]" italics="true" pageId="6" pageNumber="403">pe</emphasis>
|
||
) present but stout, connected with notched structures (
|
||
<emphasis id="16DED053FFB7FFC8BAAE9AC1B7EAF86B" box="[457,482,1956,1979]" italics="true" pageId="6" pageNumber="403">ns</emphasis>
|
||
), connected basally to the stalk.
|
||
<emphasis id="16DED053FFB7FFC8B8EE9AC1B5ACF86B" box="[905,932,1956,1979]" italics="true" pageId="6" pageNumber="403">pe</emphasis>
|
||
and
|
||
<emphasis id="16DED053FFB7FFC8B88A9AC1B20EF86B" box="[1005,1030,1956,1979]" italics="true" pageId="6" pageNumber="403">ns</emphasis>
|
||
linked like a hinge. The empty spermatophore consists of the same parts. One lateral wing is missing in the depictured emptied spermatophore.
|
||
</paragraph>
|
||
<paragraph id="24150C41FFB6FFC9BBF09DF2B297FF05" blockId="7.[151,1437,151,824]" pageId="7" pageNumber="404">The inner part is destroyed and the notched structures are dissolved. The region of the spermatophore head formerly bearing the sperm packages is lifted up nearly at an angle of 90 degree to the stalk.</paragraph>
|
||
<paragraph id="24150C41FFB6FFC9BBA09DBAB47CFF28" blockId="7.[151,1437,151,824]" box="[199,628,223,248]" pageId="7" pageNumber="404">
|
||
<emphasis id="16DED053FFB6FFC9BBA09DBAB7EBFF28" bold="true" box="[199,483,223,248]" pageId="7" pageNumber="404">Measurements (in mm).</emphasis>
|
||
See
|
||
<tableCitation id="692839FAFFB6FFC9B97D9DBAB466FF28" box="[538,622,223,248]" captionStart="TABLE 1" captionStartId="8.[119,204,554,576]" captionTargetBox="[119,1909,643,1636]" captionTargetPageId="8" captionText="TABLE 1. Measurements (mm) of female holotype anđ selecteđ paratype specimens. Abbreviations: Peđipalp tibia (T 1 T ³, T 1 the most đistal one); * Number of spines between anđ đistal margin of peđipalp tibia; ** Number of trichobothria on the basitibia anđ đistitibia of leg IV." httpUri="http://table.plazi.org/id/70D55CC9FFB9FFC6BB109F4FB2ABFDBC" pageId="7" pageNumber="404" tableUuid="70D55CC9FFB9FFC6BB109F4FB2ABFDBC">Table 1</tableCitation>
|
||
.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="6CB05FCAFFB6FFC9BBA09C61B38BFECD" box="[199,1411,260,285]" pageId="7" pageNumber="404" type="etymology">
|
||
<paragraph id="24150C41FFB6FFC9BBA09C61B38BFECD" blockId="7.[151,1437,151,824]" box="[199,1411,260,285]" pageId="7" pageNumber="404">
|
||
<emphasis id="16DED053FFB6FFC9BBA09C61B740FECD" bold="true" box="[199,328,260,285]" pageId="7" pageNumber="404">Etymology</emphasis>
|
||
: The name refers to this species representing the easternmost record of the genus known to date.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="6CB05FCAFFB6FFC7BBA09C42B725FE20" lastPageId="9" lastPageNumber="406" pageId="7" pageNumber="404" type="materials_examined">
|
||
<paragraph id="24150C41FFB6FFC9BBA09C42B516FD78" blockId="7.[151,1437,151,824]" pageId="7" pageNumber="404">
|
||
<emphasis id="16DED053FFB6FFC9BBA09C42B4A1FE90" bold="true" box="[199,681,295,320]" pageId="7" pageNumber="404">Variability between analyzed specimens.</emphasis>
|
||
The description above is based on the female
|
||
<typeStatus id="FB11B2E3FFB6FFC9BFD69C42B31EFE90" box="[1201,1302,295,320]" pageId="7" pageNumber="404" type="holotype">holotype</typeStatus>
|
||
, but several important morphological variations within the 26 studied specimens were observed. In the female
|
||
<typeStatus id="FB11B2E3FFB6FFC9BE699C29B378FEB5" box="[1294,1392,332,357]" pageId="7" pageNumber="404" type="holotype">holotype</typeStatus>
|
||
the dorsal margin of the basitarsus harbors three contiguous spines (cod-s) distal of the large submedial spine (lsmd-s) which decreases in in size proximally. Differences in this pattern were found in several specimens. Sometimes the two distal spines are fused on their bases as it is the case in
|
||
<typeStatus id="FB11B2E3FFB6FFC9B85E9CDDB591FE00" box="[825,921,440,464]" pageId="7" pageNumber="404" type="paratype">paratype</typeStatus>
|
||
male specimens (
|
||
<figureCitation id="BC9110C4FFB6FFC9BF049CD2B2AFFE00" box="[1123,1191,439,464]" captionStart="FIGURE 2" captionStartId="4.[151,250,1866,1888]" captionTargetBox="[200,1385,232,1845]" captionTargetId="figure@4.[200,1386,193,1845]" captionTargetPageId="4" captionText="FIGURE 2. Stygophrynus orientalis sp. nov. Variability of pedipalpal spination. Distal part of pedipalp, pro-lateral view. A Holotype female (21944), tarsus of right pedipalp, left drawing, right original photograph. B Paratype male (# 2), tarsus of left pedipalp. C Paratype male (# 2), tarsus of right pedipalp. D Paratype male (# 3), tibia and tarsus of left pedipalp. Labels: see text. Scale bars: 1 mm." httpUri="https://zenodo.org/record/312577/files/figure.png" pageId="7" pageNumber="404">Fig. 2</figureCitation>
|
||
B, D). In one case the spination of the basitarsi differs from one another in the same specimen (compare
|
||
<figureCitation id="BC9110C4FFB6FFC9BF5B9CB9B276FE24" box="[1084,1150,476,501]" captionStart="FIGURE 2" captionStartId="4.[151,250,1866,1888]" captionTargetBox="[200,1385,232,1845]" captionTargetId="figure@4.[200,1386,193,1845]" captionTargetPageId="4" captionText="FIGURE 2. Stygophrynus orientalis sp. nov. Variability of pedipalpal spination. Distal part of pedipalp, pro-lateral view. A Holotype female (21944), tarsus of right pedipalp, left drawing, right original photograph. B Paratype male (# 2), tarsus of left pedipalp. C Paratype male (# 2), tarsus of right pedipalp. D Paratype male (# 3), tibia and tarsus of left pedipalp. Labels: see text. Scale bars: 1 mm." httpUri="https://zenodo.org/record/312577/files/figure.png" pageId="7" pageNumber="404">Fig. 2</figureCitation>
|
||
B and 2C). The spination of the disto-ventral margin of the basitarsus also exhibits variation. In the
|
||
<typeStatus id="FB11B2E3FFB6FFC9B8B09C9AB231FDC8" box="[983,1081,511,536]" pageId="7" pageNumber="404" type="holotype">holotype</typeStatus>
|
||
female three contiguous spines proximally decreasing in size (cov-s) can be found distally of the large spine. In some specimens these can be reduced or only visible as setiferous tubercles (
|
||
<figureCitation id="BC9110C4FFB6FFC9B9B79F22B512FDB0" box="[720,794,583,608]" captionStart="FIGURE 2" captionStartId="4.[151,250,1866,1888]" captionTargetBox="[200,1385,232,1845]" captionTargetId="figure@4.[200,1386,193,1845]" captionTargetPageId="4" captionText="FIGURE 2. Stygophrynus orientalis sp. nov. Variability of pedipalpal spination. Distal part of pedipalp, pro-lateral view. A Holotype female (21944), tarsus of right pedipalp, left drawing, right original photograph. B Paratype male (# 2), tarsus of left pedipalp. C Paratype male (# 2), tarsus of right pedipalp. D Paratype male (# 3), tibia and tarsus of left pedipalp. Labels: see text. Scale bars: 1 mm." httpUri="https://zenodo.org/record/312577/files/figure.png" pageId="7" pageNumber="404">Fig. 2</figureCitation>
|
||
B), or enlarged and well pronounced (
|
||
<figureCitation id="BC9110C4FFB6FFC9BF819F22B326FDB0" box="[1254,1326,583,608]" captionStart="FIGURE 2" captionStartId="4.[151,250,1866,1888]" captionTargetBox="[200,1385,232,1845]" captionTargetId="figure@4.[200,1386,193,1845]" captionTargetPageId="4" captionText="FIGURE 2. Stygophrynus orientalis sp. nov. Variability of pedipalpal spination. Distal part of pedipalp, pro-lateral view. A Holotype female (21944), tarsus of right pedipalp, left drawing, right original photograph. B Paratype male (# 2), tarsus of left pedipalp. C Paratype male (# 2), tarsus of right pedipalp. D Paratype male (# 3), tibia and tarsus of left pedipalp. Labels: see text. Scale bars: 1 mm." httpUri="https://zenodo.org/record/312577/files/figure.png" pageId="7" pageNumber="404">Fig. 2</figureCitation>
|
||
A, C–D). Furthermore, the form and shape of the subsidiary proximal basal spine on the pedipalp basitarsus can vary in size and the tip can be curved proximally or distally (
|
||
<figureCitation id="BC9110C4FFB6FFC9B9DF9FEAB4F5FD78" box="[696,765,655,680]" captionStart="FIGURE 2" captionStartId="4.[151,250,1866,1888]" captionTargetBox="[200,1385,232,1845]" captionTargetId="figure@4.[200,1386,193,1845]" captionTargetPageId="4" captionText="FIGURE 2. Stygophrynus orientalis sp. nov. Variability of pedipalpal spination. Distal part of pedipalp, pro-lateral view. A Holotype female (21944), tarsus of right pedipalp, left drawing, right original photograph. B Paratype male (# 2), tarsus of left pedipalp. C Paratype male (# 2), tarsus of right pedipalp. D Paratype male (# 3), tibia and tarsus of left pedipalp. Labels: see text. Scale bars: 1 mm." httpUri="https://zenodo.org/record/312577/files/figure.png" pageId="7" pageNumber="404">Fig. 2</figureCitation>
|
||
B).
|
||
</paragraph>
|
||
<paragraph id="24150C41FFB6FFC9BBA09FD1B2A3FCE8" blockId="7.[151,1437,151,824]" pageId="7" pageNumber="404">
|
||
Variation was also observed in the region between spine
|
||
<emphasis id="16DED053FFB6FFC9B82D9FD1B563FD1D" bold="true" box="[842,875,692,717]" pageId="7" pageNumber="404">T1</emphasis>
|
||
and the distal margin of the pedipalp tibia. In the
|
||
<typeStatus id="FB11B2E3FFB6FFC9BBF09FB2B6F1FD20" box="[151,249,727,752]" pageId="7" pageNumber="404" type="holotype">holotype</typeStatus>
|
||
female three small spines are situated in this area (
|
||
<figureCitation id="BC9110C4FFB6FFC9B8399FB2B5A8FD20" box="[862,928,727,752]" captionStart="FIGURE 1" captionStartId="3.[151,250,1857,1879]" captionTargetBox="[252,1295,248,1759]" captionTargetId="figure@3.[199,1344,193,1813]" captionTargetPageId="3" captionText="FIGURE 1. Stygophrynus orientalis sp. nov. A Habitus of female holotype, dorsal and ventral view, respectively. B – D Female chelicera, denticles enhanced with white lines (B internal view; C external view; D external view, close-up on external teeth row). E Pedipalp (left ventral, right dorsal view), spines of femur and tibia numbered. F Distitibia and distal basitibia of walking leg IV, dorsal; trichobothria marked as follows: basitibia (bt = basitibial), distitibia (bf: basofrontal; bc: basocaudal; sbf: subbasofrontal; sbc-x: series caudal trichobothria; stf 1 x: series frontal trichobothria). Note: not all trichobothria are visible on this figure. Scale bars: A 20 mm, B – C 2 mm, D 0.25 mm E 2 mm, F 1 mm." httpUri="https://zenodo.org/record/312558/files/figure.png" pageId="7" pageNumber="404">Fig. 1</figureCitation>
|
||
A, E). Out of 26 analyzed specimens eight showed an aberrant spination pattern (either on both sides or only on one side). These specimens harbor two or one spine with another “fused” second and third one in this region of the pedipalp tibia (
|
||
<figureCitation id="BC9110C4FFB6FFC9BF239E7AB28FFCE8" box="[1092,1159,799,824]" captionStart="FIGURE 2" captionStartId="4.[151,250,1866,1888]" captionTargetBox="[200,1385,232,1845]" captionTargetId="figure@4.[200,1386,193,1845]" captionTargetPageId="4" captionText="FIGURE 2. Stygophrynus orientalis sp. nov. Variability of pedipalpal spination. Distal part of pedipalp, pro-lateral view. A Holotype female (21944), tarsus of right pedipalp, left drawing, right original photograph. B Paratype male (# 2), tarsus of left pedipalp. C Paratype male (# 2), tarsus of right pedipalp. D Paratype male (# 3), tibia and tarsus of left pedipalp. Labels: see text. Scale bars: 1 mm." httpUri="https://zenodo.org/record/312577/files/figure.png" pageId="7" pageNumber="404">Fig. 2</figureCitation>
|
||
D).
|
||
</paragraph>
|
||
<caption id="70D55CC9FFB6FFC9BBF09AE7B6D3F823" httpUri="https://zenodo.org/record/312679/files/figure.png" pageId="7" pageNumber="404" targetBox="[151,1391,896,1878]" targetPageId="7">
|
||
<paragraph id="24150C41FFB6FFC9BBF09AE7B6D3F823" blockId="7.[151,1436,1922,2035]" pageId="7" pageNumber="404">
|
||
<emphasis id="16DED053FFB6FFC9BBF09AE7B71DF847" bold="true" box="[151,277,1922,1944]" pageId="7" pageNumber="404">FIGURE 5.</emphasis>
|
||
Cerotegument of
|
||
<taxonomicName id="E3AA77C2FFB6FFC9BAB29AE7B44CF848" box="[469,580,1922,1944]" class="Arachnida" family="Charinidae" kingdom="Animalia" order="Amblypygi" pageId="7" pageNumber="404" phylum="Arthropoda" rank="family">Charinidae</taxonomicName>
|
||
and
|
||
<taxonomicName id="E3AA77C2FFB6FFC9B91C9AE7B4FFF848" box="[635,759,1922,1944]" class="Arachnida" family="Charontidae" kingdom="Animalia" order="Amblypygi" pageId="7" pageNumber="404" phylum="Arthropoda" rank="family">Charontidae</taxonomicName>
|
||
(after Wolff
|
||
<emphasis id="16DED053FFB6FFC9B8E49AE6B5B1F847" box="[899,953,1922,1944]" italics="true" pageId="7" pageNumber="404">et al.</emphasis>
|
||
2017).
|
||
<emphasis id="16DED053FFB6FFC9BF6B9AE7B234F848" bold="true" box="[1036,1084,1922,1944]" pageId="7" pageNumber="404">A–C</emphasis>
|
||
<taxonomicName id="E3AA77C2FFB6FFC9BF219AE7B2FBF848" box="[1094,1267,1922,1944]" class="Arachnida" family="Charontidae" genus="Charon" kingdom="Animalia" order="Amblypygi" pageId="7" pageNumber="404" phylum="Arthropoda" rank="species" species="grayi">
|
||
<emphasis id="16DED053FFB6FFC9BF219AE7B29AF847" box="[1094,1170,1922,1943]" italics="true" pageId="7" pageNumber="404">Charon</emphasis>
|
||
cf.
|
||
<emphasis id="16DED053FFB6FFC9BFD99AE6B2FBF848" box="[1214,1267,1923,1944]" italics="true" pageId="7" pageNumber="404">grayi</emphasis>
|
||
</taxonomicName>
|
||
(Negros Island, Philippines),
|
||
<emphasis id="16DED053FFB6FFC9BA789AC4B745F866" bold="true" box="[287,333,1953,1974]" pageId="7" pageNumber="404">D–F</emphasis>
|
||
<taxonomicName id="E3AA77C2FFB6FFC9BA319AC4B440F866" box="[342,584,1953,1974]" class="Arachnida" family="Charontidae" genus="Stygophrynus" kingdom="Animalia" order="Amblypygi" pageId="7" pageNumber="404" phylum="Arthropoda" rank="species" species="orientalis" status="sp. nov.">
|
||
<emphasis id="16DED053FFB6FFC9BA319AC4B440F866" box="[342,584,1953,1974]" italics="true" pageId="7" pageNumber="404">Stygophrynus orientalis</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="16DED053FFB6FFC9B9379AC4B4AAF866" bold="true" box="[592,674,1953,1974]" pageId="7" pageNumber="404">
|
||
<taxonomicNameLabel id="0DED6D28FFB6FFC9B9379AC4B4AAF866" box="[592,674,1953,1974]" pageId="7" pageNumber="404" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
,
|
||
<emphasis id="16DED053FFB6FFC9B9D69AC5B4D3F866" bold="true" box="[689,731,1952,1974]" pageId="7" pageNumber="404">G–I</emphasis>
|
||
<taxonomicName id="E3AA77C2FFB6FFC9B9849AC4B56AF866" box="[739,866,1953,1974]" class="Arachnida" family="Charinidae" genus="Sarax" kingdom="Animalia" order="Amblypygi" pageId="7" pageNumber="404" phylum="Arthropoda" rank="species" species="curioi">
|
||
<emphasis id="16DED053FFB6FFC9B9849AC4B56AF866" box="[739,866,1953,1974]" italics="true" pageId="7" pageNumber="404">Sarax curioi</emphasis>
|
||
</taxonomicName>
|
||
(Pany Island, Philippines). SEM of air dried exuvia of carapace.
|
||
<emphasis id="16DED053FFB6FFC9BB999ADAB7BAF804" bold="true" box="[254,434,1983,2005]" pageId="7" pageNumber="404">A, B, D, E, G, H</emphasis>
|
||
globular surface.
|
||
<emphasis id="16DED053FFB6FFC9B9099ADAB4BDF804" bold="true" box="[622,693,1983,2005]" pageId="7" pageNumber="404">C, F, I</emphasis>
|
||
detail of colloidal particles. For interpretations see text. Scale bars: see figure.
|
||
</paragraph>
|
||
</caption>
|
||
<caption id="70D55CC9FFB9FFC6BB109F4FB2ABFDBC" ID-Table-UUID="70D55CC9FFB9FFC6BB109F4FB2ABFDBC" httpUri="http://table.plazi.org/id/70D55CC9FFB9FFC6BB109F4FB2ABFDBC" pageId="8" pageNumber="405" targetBox="[119,1909,643,1636]" targetIsTable="true" targetPageId="8">
|
||
<paragraph id="24150C41FFB9FFC6BB109F4FB2ABFDBC" blockId="8.[119,1909,554,620]" pageId="8" pageNumber="405">
|
||
<emphasis id="16DED053FFB9FFC6BB109F4FB6EEFD90" bold="true" box="[119,230,554,576]" pageId="8" pageNumber="405">TABLE 1.</emphasis>
|
||
Measurements (mm) of female holotype anđ selecteđ paratype specimens. Abbreviations: Peđipalp tibia (T1 T³, T1 the most đistal one); * Number of spines between anđ đistal margin of peđipalp tibia; ** Number of trichobothria on the basitibia anđ đistitibia of leg IV.
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="24150C41FFB9FFC6BBE49FDBB13EF9B3" pageId="8" pageNumber="405">
|
||
<table id="56AAFEE1FFB90031BB109FE6B17DF9B4" box="[119,1909,643,1636]" gridcols="10" gridrows="6" pageId="8" pageNumber="405">
|
||
<tr id="9A9A0E03FFB90031BB109FE6B17DFC6F" box="[119,1909,643,959]" gridrow="0" pageId="8" pageNumber="405">
|
||
<th id="D94B677FFFB90031BB109FE6B76BFC6F" box="[119,355,643,959]" gridcol="0" gridrow="0" pageId="8" pageNumber="405">
|
||
<emphasis id="16DED053FFB9FFC6BBE49FDBB6E9FCDE" bold="true" pageId="8" pageNumber="405">Carapace (length / width) Pedipalp</emphasis>
|
||
Femur Tibia Tibia Spine T1 Tibia Spine Tƻ Tibia Spine T³
|
||
</th>
|
||
<th id="D94B677FFFB90031BAE69FE6B7FEFC6F" box="[385,502,643,959]" gridcol="1" gridrow="0" pageId="8" pageNumber="405">
|
||
<emphasis id="16DED053FFB9FFC6BAE69FE6B7D8FD66" bold="true" pageId="8" pageNumber="405">♀ holotype (21944)</emphasis>
|
||
6.84 / 9.³1 6.8ƻ 7.11 ƻ.1³ ƻ.³5 ƻ.04
|
||
</th>
|
||
<th id="D94B677FFFB90031B94C9FE6B4ACFC6F" box="[555,676,643,959]" gridcol="2" gridrow="0" pageId="8" pageNumber="405">
|
||
<emphasis id="16DED053FFB9FFC6B94C9FE6B49CFD66" bold="true" pageId="8" pageNumber="405">♀ paratype (21946_7)</emphasis>
|
||
5.7ƻ / 7.76 4.87 5.74 ƻ.³ ƻ.54 ƻ.06
|
||
</th>
|
||
<th id="D94B677FFFB90031B9B09FE6B558FC6F" box="[727,848,643,959]" gridcol="3" gridrow="0" pageId="8" pageNumber="405">
|
||
<emphasis id="16DED053FFB9FFC6B9B09FE6B537FD66" bold="true" pageId="8" pageNumber="405">♀ paratype (21946_9)</emphasis>
|
||
6.1ƻ / 8.07 5.75 6.59 ƻ.17 ƻ.45 ƻ.11
|
||
</th>
|
||
<th id="D94B677FFFB90031B8E29FE6B5F6FC6F" box="[901,1022,643,959]" gridcol="4" gridrow="0" pageId="8" pageNumber="405">
|
||
<emphasis id="16DED053FFB9FFC6B8E29FE1B5DDFD66" bold="true" pageId="8" pageNumber="405">Ƌ paratype (21945)</emphasis>
|
||
7.64 / 10.01 9.³5 10.³8 ƻ.71 ³.06 ƻ.9ƻ
|
||
</th>
|
||
<th id="D94B677FFFB90031BF569FE6B2A2FC6F" box="[1073,1194,643,959]" gridcol="5" gridrow="0" pageId="8" pageNumber="405">
|
||
<emphasis id="16DED053FFB9FFC6BF569FE1B291FD66" bold="true" pageId="8" pageNumber="405">Ƌ paratype (21946_³)</emphasis>
|
||
6.³1 / 8.98 7.³³ 8.98 ƻ.³7 ƻ.8³ ƻ.49
|
||
</th>
|
||
<th id="D94B677FFFB90031BFBA9FE6B35EFC6F" box="[1245,1366,643,959]" gridcol="6" gridrow="0" pageId="8" pageNumber="405">
|
||
<emphasis id="16DED053FFB9FFC6BFBA9FE1B34EFD66" bold="true" pageId="8" pageNumber="405">Ƌ paratype (21946_4)</emphasis>
|
||
6.4³ / 8.99 6.99 7.5 ƻ.41 ƻ.81 ƻ.49
|
||
</th>
|
||
<th id="D94B677FFFB90031BEEE9FE6B00AFC6F" box="[1417,1538,643,959]" gridcol="7" gridrow="0" pageId="8" pageNumber="405">
|
||
<emphasis id="16DED053FFB9FFC6BEEE9FE1B3F9FD66" bold="true" pageId="8" pageNumber="405">Ƌ paratype (21946_5)</emphasis>
|
||
6.15 / 8.77 6.³6 8.ƻƻ ƻ.59 ³ ƻ.87
|
||
</th>
|
||
<th id="D94B677FFFB90031BD529FE6B0A6FC6F" box="[1589,1710,643,959]" gridcol="8" gridrow="0" pageId="8" pageNumber="405">
|
||
<emphasis id="16DED053FFB9FFC6BD529FE1B096FD66" bold="true" pageId="8" pageNumber="405">Ƌ paratype (21946_6)</emphasis>
|
||
5.86 / 8.ƻ³ 5.76 6.ƻƻ ƻ.45 ƻ.58 1.98
|
||
</th>
|
||
<th id="D94B677FFFB90031BD849FE6B17DFC6F" box="[1763,1909,643,959]" gridcol="9" gridrow="0" pageId="8" pageNumber="405">
|
||
<emphasis id="16DED053FFB9FFC6BD849FE1B143FD66" bold="true" pageId="8" pageNumber="405">Ƌ paratype (21946_8)</emphasis>
|
||
6.44 / 8.ƻƻ 6.³ƻ 7.1³ ƻ.³6 ƻ.59 ƻ.18
|
||
</th>
|
||
</tr>
|
||
<tr id="9A9A0E03FFB90031BB109EA8B17DFBFA" box="[119,1909,973,1066]" gridrow="1" pageId="8" pageNumber="405">
|
||
<th id="D94B677FFFB90031BB109EA8B76BFBFA" box="[119,355,973,1066]" gridcol="0" gridrow="1" pageId="8" pageNumber="405">Spines to margin* Basitarsus Distitarsus</th>
|
||
<td id="D94B677FFFB90031BAE69EA8B7FEFBFA" box="[385,502,973,1066]" gridcol="1" gridrow="1" pageId="8" pageNumber="405">³ ³.07 ƻ.49</td>
|
||
<td id="D94B677FFFB90031B94C9EA8B4ACFBFA" box="[555,676,973,1066]" gridcol="2" gridrow="1" pageId="8" pageNumber="405">³ ƻ.55 ƻ.³7</td>
|
||
<td id="D94B677FFFB90031B9B09EA8B558FBFA" box="[727,848,973,1066]" gridcol="3" gridrow="1" pageId="8" pageNumber="405">³ ƻ.69 ƻ.³8</td>
|
||
<td id="D94B677FFFB90031B8E29EA8B5F6FBFA" box="[901,1022,973,1066]" gridcol="4" gridrow="1" pageId="8" pageNumber="405">ƻ ³.7 ³.48</td>
|
||
<td id="D94B677FFFB90031BF569EA8B2A2FBFA" box="[1073,1194,973,1066]" gridcol="5" gridrow="1" pageId="8" pageNumber="405">ƻ‒³ ³.57 ³.³4</td>
|
||
<td id="D94B677FFFB90031BFBA9EA8B35EFBFA" box="[1245,1366,973,1066]" gridcol="6" gridrow="1" pageId="8" pageNumber="405">³ ƻ.99 ƻ.65</td>
|
||
<td id="D94B677FFFB90031BEEE9EA8B00AFBFA" box="[1417,1538,973,1066]" gridcol="7" gridrow="1" pageId="8" pageNumber="405">³ ³.16 ³.ƻ</td>
|
||
<td id="D94B677FFFB90031BD529EA8B0A6FBFA" box="[1589,1710,973,1066]" gridcol="8" gridrow="1" pageId="8" pageNumber="405">ƻ-³ ƻ.³ ƻ.17</td>
|
||
<td id="D94B677FFFB90031BD849EA8B17DFBFA" box="[1763,1909,973,1066]" gridcol="9" gridrow="1" pageId="8" pageNumber="405">2 ³.ƻ4 ƻ.9ƻ</td>
|
||
</tr>
|
||
<tr id="9A9A0E03FFB90031BB10995DB17DFB45" box="[119,1909,1080,1173]" gridrow="2" pageId="8" pageNumber="405">
|
||
<th id="D94B677FFFB90031BB10995DB76BFB45" box="[119,355,1080,1173]" gridcol="0" gridrow="2" pageId="8" pageNumber="405">
|
||
<emphasis id="16DED053FFB9FFC6BBE4995DB715FB9E" bold="true" box="[131,285,1080,1102]" pageId="8" pageNumber="405">Walking leg II</emphasis>
|
||
Femur Patella
|
||
</th>
|
||
<td id="D94B677FFFB90031BAE6995DB7FEFB45" box="[385,502,1080,1173]" gridcol="1" gridrow="2" pageId="8" pageNumber="405">9.95 1.58</td>
|
||
<td id="D94B677FFFB90031B94C995DB4ACFB45" box="[555,676,1080,1173]" gridcol="2" gridrow="2" pageId="8" pageNumber="405">8.7 1.5</td>
|
||
<td id="D94B677FFFB90031B9B0995DB558FB45" box="[727,848,1080,1173]" gridcol="3" gridrow="2" pageId="8" pageNumber="405">10.8 1.7</td>
|
||
<td id="D94B677FFFB90031B8E2995DB5F6FB45" box="[901,1022,1080,1173]" gridcol="4" gridrow="2" pageId="8" pageNumber="405">1³.64 ƻ.16</td>
|
||
<td id="D94B677FFFB90031BF56995DB2A2FB45" box="[1073,1194,1080,1173]" gridcol="5" gridrow="2" pageId="8" pageNumber="405">11 1.9</td>
|
||
<td id="D94B677FFFB90031BFBA995DB35EFB45" box="[1245,1366,1080,1173]" gridcol="6" gridrow="2" pageId="8" pageNumber="405">10.5 1.4</td>
|
||
<td id="D94B677FFFB90031BEEE995DB00AFB45" box="[1417,1538,1080,1173]" gridcol="7" gridrow="2" pageId="8" pageNumber="405">11.ƻ 1.5</td>
|
||
<td id="D94B677FFFB90031BD52995DB0A6FB45" box="[1589,1710,1080,1173]" gridcol="8" gridrow="2" pageId="8" pageNumber="405">9.1 1.6</td>
|
||
<td id="D94B677FFFB90031BD84995DB17DFB45" box="[1763,1909,1080,1173]" gridcol="9" gridrow="2" pageId="8" pageNumber="405">1³.5 ƻ</td>
|
||
</tr>
|
||
<tr id="9A9A0E03FFB90031BB1099C6B17DFA61" box="[119,1909,1187,1457]" gridrow="3" pageId="8" pageNumber="405">
|
||
<th id="D94B677FFFB90031BB1099C6B76BFA61" box="[119,355,1187,1457]" gridcol="0" gridrow="3" pageId="8" pageNumber="405">
|
||
<emphasis id="16DED053FFB9FFC6BBE499C6B72FFB69" bold="true" box="[131,295,1187,1209]" pageId="8" pageNumber="405">Walking leg III</emphasis>
|
||
Femur Patella Basitibia Distitibia Basitarsus + claw
|
||
<emphasis id="16DED053FFB9FFC6BBE4981DB72EFA5E" bold="true" box="[131,294,1400,1422]" pageId="8" pageNumber="405">Walking leg IV</emphasis>
|
||
Femur
|
||
</th>
|
||
<td id="D94B677FFFB90031BAE699C6B7FEFA61" box="[385,502,1187,1457]" gridcol="1" gridrow="3" pageId="8" pageNumber="405">11.57 ƻ.³ƻ 11.77 4.68 4.16 11.ƻ³</td>
|
||
<td id="D94B677FFFB90031B94C99C6B4ACFA61" box="[555,676,1187,1457]" gridcol="2" gridrow="3" pageId="8" pageNumber="405">9.7 1.6 9.5 4.ƻ ³.ƻ 9</td>
|
||
<td id="D94B677FFFB90031B9B099C6B558FA61" box="[727,848,1187,1457]" gridcol="3" gridrow="3" pageId="8" pageNumber="405">11.6 ƻ.ƻ 11.7 4.8 4 10.6</td>
|
||
<td id="D94B677FFFB90031B8E299C6B5F6FA61" box="[901,1022,1187,1457]" gridcol="4" gridrow="3" pageId="8" pageNumber="405">14.9³ ƻ.³9 14.91 5.88 4.54 1³.³5</td>
|
||
<td id="D94B677FFFB90031BF5699C6B2A2FA61" box="[1073,1194,1187,1457]" gridcol="5" gridrow="3" pageId="8" pageNumber="405">1ƻ.6 1.9 1ƻ.³ 5.ƻ 4 11.5</td>
|
||
<td id="D94B677FFFB90031BFBA99C6B35EFA61" box="[1245,1366,1187,1457]" gridcol="6" gridrow="3" pageId="8" pageNumber="405">1ƻ.1 1.8 11.7 4.6 4 10.9</td>
|
||
<td id="D94B677FFFB90031BEEE99C6B00AFA61" box="[1417,1538,1187,1457]" gridcol="7" gridrow="3" pageId="8" pageNumber="405">1ƻ.ƻ 1.7 1ƻ.ƻ 5.1 4 11.³</td>
|
||
<td id="D94B677FFFB90031BD5299C6B0A6FA61" box="[1589,1710,1187,1457]" gridcol="8" gridrow="3" pageId="8" pageNumber="405">9.6 1.9 10.5 4.1 ³.5 9.7</td>
|
||
<td id="D94B677FFFB90031BD8499C6B17DFA61" box="[1763,1909,1187,1457]" gridcol="9" gridrow="3" pageId="8" pageNumber="405">14.³ ƻ.1 14.4 6.1 4.³ 1³.1</td>
|
||
</tr>
|
||
<tr id="9A9A0E03FFB90031BB1098DAB17DFA05" box="[119,1909,1471,1493]" gridrow="4" pageId="8" pageNumber="405">
|
||
<th id="D94B677FFFB90031BB1098DAB76BFA05" box="[119,355,1471,1493]" gridcol="0" gridrow="4" pageId="8" pageNumber="405">Patella</th>
|
||
<td id="D94B677FFFB90031BAE698DAB7FEFA05" box="[385,502,1471,1493]" gridcol="1" gridrow="4" pageId="8" pageNumber="405">1.71</td>
|
||
<td id="D94B677FFFB90031B94C98DAB4ACFA05" box="[555,676,1471,1493]" gridcol="2" gridrow="4" pageId="8" pageNumber="405">1.6</td>
|
||
<td id="D94B677FFFB90031B9B098DAB558FA05" box="[727,848,1471,1493]" gridcol="3" gridrow="4" pageId="8" pageNumber="405">1.8</td>
|
||
<td id="D94B677FFFB90031B8E298DAB5F6FA05" box="[901,1022,1471,1493]" gridcol="4" gridrow="4" pageId="8" pageNumber="405">1.98</td>
|
||
<td id="D94B677FFFB90031BF5698DAB2A2FA05" box="[1073,1194,1471,1493]" gridcol="5" gridrow="4" pageId="8" pageNumber="405">1.5</td>
|
||
<td id="D94B677FFFB90031BFBA98DAB35EFA05" box="[1245,1366,1471,1493]" gridcol="6" gridrow="4" pageId="8" pageNumber="405">1.6</td>
|
||
<td id="D94B677FFFB90031BEEE98DAB00AFA05" box="[1417,1538,1471,1493]" gridcol="7" gridrow="4" pageId="8" pageNumber="405">1.8</td>
|
||
<td id="D94B677FFFB90031BD5298DAB0A6FA05" box="[1589,1710,1471,1493]" gridcol="8" gridrow="4" pageId="8" pageNumber="405">1.5</td>
|
||
<td id="D94B677FFFB90031BD8498DAB17DFA05" box="[1763,1909,1471,1493]" gridcol="9" gridrow="4" pageId="8" pageNumber="405">1.9</td>
|
||
</tr>
|
||
<tr id="9A9A0E03FFB90031BB109886B17DF9B4" box="[119,1909,1507,1636]" gridrow="5" pageId="8" pageNumber="405">
|
||
<th id="D94B677FFFB90031BB109886B76BF9B4" box="[119,355,1507,1636]" gridcol="0" gridrow="5" pageId="8" pageNumber="405">Basitibia Distitibia Basitarsus + claw Trichobothria**</th>
|
||
<td id="D94B677FFFB90031BAE69886B7FEF9B4" box="[385,502,1507,1636]" gridcol="1" gridrow="5" pageId="8" pageNumber="405">1ƻ.³1 4.ƻ1 4.14 ƻ³</td>
|
||
<td id="D94B677FFFB90031B94C9886B4ACF9B4" box="[555,676,1507,1636]" gridcol="2" gridrow="5" pageId="8" pageNumber="405">10.1 ³.6 ³.³ ƻ³</td>
|
||
<td id="D94B677FFFB90031B9B09886B558F9B4" box="[727,848,1507,1636]" gridcol="3" gridrow="5" pageId="8" pageNumber="405">1ƻ.1 4.1 4.1 ƻ³</td>
|
||
<td id="D94B677FFFB90031B8E29886B5F6F9B4" box="[901,1022,1507,1636]" gridcol="4" gridrow="5" pageId="8" pageNumber="405">14.56 5.ƻ 4.41 ƻ³</td>
|
||
<td id="D94B677FFFB90031BF569886B2A2F9B4" box="[1073,1194,1507,1636]" gridcol="5" gridrow="5" pageId="8" pageNumber="405">1ƻ.9 4.4 ³.9 ƻ³</td>
|
||
<td id="D94B677FFFB90031BFBA9886B35EF9B4" box="[1245,1366,1507,1636]" gridcol="6" gridrow="5" pageId="8" pageNumber="405">1ƻ.1 4.1 ³.9 ƻ³</td>
|
||
<td id="D94B677FFFB90031BEEE9886B00AF9B4" box="[1417,1538,1507,1636]" gridcol="7" gridrow="5" pageId="8" pageNumber="405">1ƻ.9 4.ƻ ³.8 23</td>
|
||
<td id="D94B677FFFB90031BD529886B0A6F9B4" box="[1589,1710,1507,1636]" gridcol="8" gridrow="5" pageId="8" pageNumber="405">11 ³.6 ³.5 ƻ³</td>
|
||
<td id="D94B677FFFB90031BD849886B17DF9B4" box="[1763,1909,1507,1636]" gridcol="9" gridrow="5" pageId="8" pageNumber="405">15.5 4.9 4.7 2³</td>
|
||
</tr>
|
||
</table>
|
||
</paragraph>
|
||
<paragraph id="24150C41FFB8FFC7BBA09DF2B725FE20" blockId="9.[151,1436,151,496]" pageId="9" pageNumber="406">
|
||
<emphasis id="16DED053FFB8FFC7BBA09DF2B541FF60" bold="true" box="[199,841,151,176]" pageId="9" pageNumber="406">Comparative morphology of cerotegument structure.</emphasis>
|
||
The cerotegument of
|
||
<taxonomicName id="E3AA77C2FFB8FFC7BF2C9DFDB2DCFF60" box="[1099,1236,151,176]" class="Arachnida" family="Charontidae" genus="Stygophrynus" kingdom="Animalia" order="Amblypygi" pageId="9" pageNumber="406" phylum="Arthropoda" rank="species" species="orientalis" status="sp. nov.">
|
||
<emphasis id="16DED053FFB8FFC7BF2C9DFDB2DCFF60" box="[1099,1236,151,176]" italics="true" pageId="9" pageNumber="406">S. orientalis</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="16DED053FFB8FFC7BFB99DF2B332FF60" bold="true" box="[1246,1338,151,176]" pageId="9" pageNumber="406">
|
||
<taxonomicNameLabel id="0DED6D28FFB8FFC7BFB99DF2B332FF60" box="[1246,1338,151,176]" pageId="9" pageNumber="406" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
exhibits regular, globular granules with a rough surface structure. There are short connecting fibers between granules and the underlying cuticle (
|
||
<figureCitation id="BC9110C4FFB8FFC7BAFD9DBBB7D4FF27" box="[410,476,222,247]" captionStart="FIGURE 5" captionStartId="7.[151,250,1922,1944]" captionTargetBox="[151,1391,896,1878]" captionTargetId="figure@7.[151,1436,877,1936]" captionTargetPageId="7" captionText="FIGURE 5. Cerotegument of Charinidae and Charontidae (after Wolff et al. 2017). A – C Charon cf. grayi (Negros Island, Philippines), D – F Stygophrynus orientalis sp. nov., G – I Sarax curioi (Pany Island, Philippines). SEM of air dried exuvia of carapace. A, B, D, E, G, H globular surface. C, F, I detail of colloidal particles. For interpretations see text. Scale bars: see figure." httpUri="https://zenodo.org/record/312679/files/figure.png" pageId="9" pageNumber="406">Fig. 5</figureCitation>
|
||
D). The granule surface is built by both crystal- and fiber-like nano-particles (
|
||
<figureCitation id="BC9110C4FFB8FFC7BE259DBBB38CFF27" box="[1346,1412,222,247]" captionStart="FIGURE 5" captionStartId="7.[151,250,1922,1944]" captionTargetBox="[151,1391,896,1878]" captionTargetId="figure@7.[151,1436,877,1936]" captionTargetPageId="7" captionText="FIGURE 5. Cerotegument of Charinidae and Charontidae (after Wolff et al. 2017). A – C Charon cf. grayi (Negros Island, Philippines), D – F Stygophrynus orientalis sp. nov., G – I Sarax curioi (Pany Island, Philippines). SEM of air dried exuvia of carapace. A, B, D, E, G, H globular surface. C, F, I detail of colloidal particles. For interpretations see text. Scale bars: see figure." httpUri="https://zenodo.org/record/312679/files/figure.png" pageId="9" pageNumber="406">Fig. 5</figureCitation>
|
||
E, F). In
|
||
<taxonomicName id="E3AA77C2FFB8FFC7BB8D9C66B565FECB" authority="Giupponi & Miranda, 2012" authorityName="Giupponi & Miranda" authorityYear="2012" box="[234,877,258,283]" class="Arachnida" family="Charinidae" genus="Sarax" kingdom="Animalia" order="Amblypygi" pageId="9" pageNumber="406" phylum="Arthropoda" rank="species" species="curioi">
|
||
<emphasis id="16DED053FFB8FFC7BB8D9C66B774FECA" box="[234,380,259,282]" italics="true" pageId="9" pageNumber="406">Sarax curioi</emphasis>
|
||
Giupponi & Miranda, 2012 (Charinidae)
|
||
</taxonomicName>
|
||
the globule surface is fine granular, which is comparable to the structure found in
|
||
<taxonomicName id="E3AA77C2FFB8FFC7B9249C40B4A3FEEE" box="[579,683,293,318]" class="Arachnida" family="Charinidae" genus="Charinus" kingdom="Animalia" order="Amblypygi" pageId="9" pageNumber="406" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="16DED053FFB8FFC7B9249C40B4A3FEEE" box="[579,683,293,318]" italics="true" pageId="9" pageNumber="406">Charinus</emphasis>
|
||
</taxonomicName>
|
||
spp. (
|
||
<taxonomicName id="E3AA77C2FFB8FFC7B9939C40B57AFEEE" box="[756,882,293,318]" class="Arachnida" family="Charinidae" kingdom="Animalia" order="Amblypygi" pageId="9" pageNumber="406" phylum="Arthropoda" rank="family">Charinidae</taxonomicName>
|
||
) (
|
||
<bibRefCitation id="403B71B0FFB8FFC7B8ED9C40B25FFEEE" author="Wolff" box="[906,1111,293,318]" pageId="9" pageNumber="406" refString="Wolff, J. O., Seiter, M. & Gorb, S. N. (2017) The water-repellent cerotegument of whip-spiders (Arachnida: Amblypygi). Arthropod Structure & Development. 46, 116 - 129." type="journal article" year="2017">
|
||
Wolff
|
||
<emphasis id="16DED053FFB8FFC7B8B19C42B202FEEE" box="[982,1034,293,318]" italics="true" pageId="9" pageNumber="406">et al</emphasis>
|
||
. 2017
|
||
</bibRefCitation>
|
||
). Similar to
|
||
<taxonomicName id="E3AA77C2FFB8FFC7BF809C43B379FEEE" box="[1255,1393,293,318]" class="Arachnida" family="Charontidae" genus="Stygophrynus" kingdom="Animalia" order="Amblypygi" pageId="9" pageNumber="406" phylum="Arthropoda" rank="species" species="orientalis" status="sp. nov.">
|
||
<emphasis id="16DED053FFB8FFC7BF809C43B379FEEE" box="[1255,1393,293,318]" italics="true" pageId="9" pageNumber="406">S. orientalis</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="16DED053FFB8FFC7BE1D9C40B6C0FEB2" bold="true" pageId="9" pageNumber="406">
|
||
<taxonomicNameLabel id="0DED6D28FFB8FFC7BE1D9C40B6C0FEB2" pageId="9" pageNumber="406" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
there are connecting fibers between granules and the underlying cuticle (
|
||
<figureCitation id="BC9110C4FFB8FFC7BF739C2CB25FFEB2" box="[1044,1111,329,354]" captionStart="FIGURE 5" captionStartId="7.[151,250,1922,1944]" captionTargetBox="[151,1391,896,1878]" captionTargetId="figure@7.[151,1436,877,1936]" captionTargetPageId="7" captionText="FIGURE 5. Cerotegument of Charinidae and Charontidae (after Wolff et al. 2017). A – C Charon cf. grayi (Negros Island, Philippines), D – F Stygophrynus orientalis sp. nov., G – I Sarax curioi (Pany Island, Philippines). SEM of air dried exuvia of carapace. A, B, D, E, G, H globular surface. C, F, I detail of colloidal particles. For interpretations see text. Scale bars: see figure." httpUri="https://zenodo.org/record/312679/files/figure.png" pageId="9" pageNumber="406">Fig. 5</figureCitation>
|
||
G, H), and the cerotegument surface is built by both crystal- and fiber-like nano-particles (
|
||
<figureCitation id="BC9110C4FFB8FFC7B8239C09B58FFE55" box="[836,903,364,389]" captionStart="FIGURE 5" captionStartId="7.[151,250,1922,1944]" captionTargetBox="[151,1391,896,1878]" captionTargetId="figure@7.[151,1436,877,1936]" captionTargetPageId="7" captionText="FIGURE 5. Cerotegument of Charinidae and Charontidae (after Wolff et al. 2017). A – C Charon cf. grayi (Negros Island, Philippines), D – F Stygophrynus orientalis sp. nov., G – I Sarax curioi (Pany Island, Philippines). SEM of air dried exuvia of carapace. A, B, D, E, G, H globular surface. C, F, I detail of colloidal particles. For interpretations see text. Scale bars: see figure." httpUri="https://zenodo.org/record/312679/files/figure.png" pageId="9" pageNumber="406">
|
||
Fig.
|
||
<date id="50142A81FFB8FFC7B81F9C09B58FFE55" box="[888,903,364,389]" pageId="9" pageNumber="406">5</date>
|
||
</figureCitation>
|
||
I). In contrast,
|
||
<taxonomicName id="E3AA77C2FFB8FFC7BF4C9C09B73AFE79" authority="Gervais, 1842" authorityName="Gervais" authorityYear="1842" class="Arachnida" family="Charontidae" genus="Charon" kingdom="Animalia" order="Amblypygi" pageId="9" pageNumber="406" phylum="Arthropoda" rank="species" species="grayi">
|
||
<emphasis id="16DED053FFB8FFC7BF4C9C09B277FE55" box="[1067,1151,364,389]" italics="true" pageId="9" pageNumber="406">Charon</emphasis>
|
||
cf.
|
||
<emphasis id="16DED053FFB8FFC7BFCD9C08B2EDFE54" box="[1194,1253,365,388]" italics="true" pageId="9" pageNumber="406">grayi</emphasis>
|
||
(Gervais, 1842) (Charontidae)
|
||
</taxonomicName>
|
||
exhibits small globular crystals (
|
||
<figureCitation id="BC9110C4FFB8FFC7B9C99CF5B4FAFE79" box="[686,754,400,425]" captionStart="FIGURE 5" captionStartId="7.[151,250,1922,1944]" captionTargetBox="[151,1391,896,1878]" captionTargetId="figure@7.[151,1436,877,1936]" captionTargetPageId="7" captionText="FIGURE 5. Cerotegument of Charinidae and Charontidae (after Wolff et al. 2017). A – C Charon cf. grayi (Negros Island, Philippines), D – F Stygophrynus orientalis sp. nov., G – I Sarax curioi (Pany Island, Philippines). SEM of air dried exuvia of carapace. A, B, D, E, G, H globular surface. C, F, I detail of colloidal particles. For interpretations see text. Scale bars: see figure." httpUri="https://zenodo.org/record/312679/files/figure.png" pageId="9" pageNumber="406">Fig. 5</figureCitation>
|
||
C), forming the regular nubby granule surface (
|
||
<figureCitation id="BC9110C4FFB8FFC7BE769CF5B35DFE79" box="[1297,1365,400,425]" captionStart="FIGURE 5" captionStartId="7.[151,250,1922,1944]" captionTargetBox="[151,1391,896,1878]" captionTargetId="figure@7.[151,1436,877,1936]" captionTargetPageId="7" captionText="FIGURE 5. Cerotegument of Charinidae and Charontidae (after Wolff et al. 2017). A – C Charon cf. grayi (Negros Island, Philippines), D – F Stygophrynus orientalis sp. nov., G – I Sarax curioi (Pany Island, Philippines). SEM of air dried exuvia of carapace. A, B, D, E, G, H globular surface. C, F, I detail of colloidal particles. For interpretations see text. Scale bars: see figure." httpUri="https://zenodo.org/record/312679/files/figure.png" pageId="9" pageNumber="406">Fig. 5</figureCitation>
|
||
A, B), that has some similarity with that of
|
||
<taxonomicName id="E3AA77C2FFB8FFC7B9519CD1B4B6FE1C" box="[566,702,435,460]" class="Arachnida" family="Charontidae" genus="Stygophrynus" kingdom="Animalia" order="Amblypygi" pageId="9" pageNumber="406" phylum="Arthropoda" rank="species" species="orientalis" status="sp. nov.">
|
||
<emphasis id="16DED053FFB8FFC7B9519CD1B4B6FE1C" box="[566,702,435,460]" italics="true" pageId="9" pageNumber="406">S. orientalis</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="16DED053FFB8FFC7B9A19CD6B529FE1C" bold="true" box="[710,801,435,460]" pageId="9" pageNumber="406">
|
||
<taxonomicNameLabel id="0DED6D28FFB8FFC7B9A19CD6B529FE1C" box="[710,801,435,460]" pageId="9" pageNumber="406" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
However, fibers are lacking, and the overall structure is more regular.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |