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<document ID-CLB-Dataset="55906" ID-DOI="http://dx.doi.org/10.3897/zookeys.1097.81792" ID-GBIF-Dataset="77c4a984-d3fb-4ec3-a1fb-acf366ecce2f" ID-PMC="PMC9042997" ID-Pensoft-Pub="1313-2970-1097-47" ID-Pensoft-UUID="F648624DC6F95540BA3A31A28DC3FD4F" ID-PubMed="35837585" ID-ZooBank="18F467AF6AC6441297945A004A671F5B" ModsDocID="1313-2970-1097-47" checkinTime="1650371384403" checkinUser="pensoft" docAuthor="Evsyukov, Aleksandr P., Vagalinski, Boyan, Zabiyaka, Igor Y. &amp; Sadyrin, Evgeniy V." docDate="2022" docId="0D955BFA370F5C2DB82F8D8EDE4F6E00" docLanguage="en" docName="ZooKeys 1097: 47-63" docOrigin="ZooKeys 1097" docPubDate="2022-04-19" docSource="http://dx.doi.org/10.3897/zookeys.1097.81792" docTitle="Bellatoiulus golovatchi gen. et 2022, sp. nov." docType="treatment" docUuid="A8A39314-2E0B-4E99-A65A-EBB595D16EF2" docUuidSource="ZooBank" docVersion="6" id="F648624DC6F95540BA3A31A28DC3FD4F" lastPageNumber="47" masterDocId="F648624DC6F95540BA3A31A28DC3FD4F" masterDocTitle="A new millipede genus and species of the tribe Pachyiulini from the Caucasus (Diplopoda, Julida, Julidae)" masterLastPageNumber="63" masterPageNumber="47" pageNumber="47" updateTime="1732849202006" updateUser="ExternalLinkService">
<mods:mods id="57CDC0D408CBE40D143714D196B6335A" xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo id="930629282549E5E2CD114ADCE10450D5">
<mods:title id="9F11CBAC27A9B657CBBB76255B439AE9">A new millipede genus and species of the tribe Pachyiulini from the Caucasus (Diplopoda, Julida, Julidae)</mods:title>
</mods:titleInfo>
<mods:name id="AE3CC8783AC0537CE0668CB173D0A4F2" type="personal">
<mods:role id="F3AEE21349302544D6AB9563B08263C5">
<mods:roleTerm id="06F953C419EB3A5F30A6A0F033B3EA37">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="0DC8C517DB9E3C9AF201140620F711E5">Evsyukov, Aleksandr P.</mods:namePart>
<mods:nameIdentifier id="6B57E55AB319ED9343C06FFF73A6D4C2" type="ORCID">https://orcid.org/0000-0001-5521-7563</mods:nameIdentifier>
<mods:affiliation id="39B709139EF27DEA41E04AD33B93711A">Don State Technical University, Department of Biology and General Pathology, Rostov-on-Don, Russia</mods:affiliation>
<mods:nameIdentifier id="9D009ACB48DE7A3B941AD2369E9A2F10" type="email">aevsukov@mail.ru</mods:nameIdentifier>
</mods:name>
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<mods:roleTerm id="93D8306C93505D5CC3B7B9708B134AA5">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="000ED94475DD3AF5A6402AE0D56ABE40">Vagalinski, Boyan</mods:namePart>
<mods:nameIdentifier id="99E9E5480EBAA17BCD6C5297D6F692B0" type="ORCID">https://orcid.org/0000-0002-4661-8952</mods:nameIdentifier>
<mods:affiliation id="39DF510EDE90FCDD74F21EC403549997">Institute of Biodiversity and Ecosystem Research at the Bulgarian Academy of Sciences, Sofia, Bulgaria</mods:affiliation>
</mods:name>
<mods:name id="AC6F0F8EE63472A47D18B6B21BCC3606" type="personal">
<mods:role id="A9FAF9A496193C137A2B1A08C4462EC0">
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</mods:role>
<mods:namePart id="31965C6F1B414BC698D777BEFD282B1B">Zabiyaka, Igor Y.</mods:namePart>
<mods:nameIdentifier id="AC78820504BAEDCEF145BB21B0CB8C27" type="ORCID">https://orcid.org/0000-0001-6759-549X</mods:nameIdentifier>
<mods:affiliation id="F5F03C59D4FCB9C00EC516E0DEA2B1D8">Don State Technical University, Research and Education Center &quot; Materials &quot;, Rostov-on-Don, Russia</mods:affiliation>
</mods:name>
<mods:name id="06EEBD6F4757143EDC3BDE2756B60CA1" type="personal">
<mods:role id="E4B45AB32E1439A0339421925CF18E0C">
<mods:roleTerm id="7AC25BC8E8AF7649C628714368FF7B36">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="53DF8B5421EA1467075E62D487245A5E">Sadyrin, Evgeniy V.</mods:namePart>
<mods:nameIdentifier id="DA9FAE34607C5FC0972A7E2A8EE78BC3" type="ORCID">https://orcid.org/0000-0002-5395-3310</mods:nameIdentifier>
<mods:affiliation id="A6284CF0CB64D95D0A31AFAC5EB52B49">Don State Technical University, Research and Education Center &quot; Materials &quot;, Rostov-on-Don, Russia</mods:affiliation>
</mods:name>
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<mods:part id="549A63CDC39B5AE0CFF49B17DB05B60D">
<mods:date id="88FA95B088DED81992367900D78FFD1C">2022</mods:date>
<mods:detail id="4A22CA2BD579868AE46A9F850527DF7A" type="pubDate">
<mods:number id="2CD909B5228FD97F63EB5464C8304CB0">2022-04-19</mods:number>
</mods:detail>
<mods:detail id="29A1809F0734E9E0AEE6B9F7FED9A054" type="volume">
<mods:number id="44EA1A5CCF5E4197CC37A2C5F5BD0BBF">1097</mods:number>
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<mods:start id="A7D1C78CF6941849700CCCD613FF174E">47</mods:start>
<mods:end id="E53D856F4783CBAA3A43A9B8142E077C">63</mods:end>
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<mods:classification id="2B7C0F0F0CEE5CF7E337F98B81C0FF94">journal article</mods:classification>
<mods:identifier id="FA457794B16C2AD05D15C58154E350B2" type="DOI">http://dx.doi.org/10.3897/zookeys.1097.81792</mods:identifier>
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<mods:identifier id="C753DFB155FA3898CCD0CDE6E0D1E674" type="Pensoft-UUID">F648624DC6F95540BA3A31A28DC3FD4F</mods:identifier>
</mods:mods>
<subSection id="5A819D0E6A233883D6D3679C0EF96A2F" pageId="0" pageNumber="47" type="taxonomic part">
<treatment id="0D955BFA370F5C2DB82F8D8EDE4F6E00" ID-GBIF-Taxon="194660808" LSID="urn:lsid:zoobank.org:act:A8A39314-2E0B-4E99-A65A-EBB595D16EF2" httpUri="http://treatment.plazi.org/id/0D955BFA370F5C2DB82F8D8EDE4F6E00" lastPageNumber="47" pageId="0" pageNumber="47" scope_class="Diplopoda" scope_family="Julidae" scope_order="Julida">
<subSubSection id="A27B24C0AB1639000263AA268986FF12" pageId="0" pageNumber="47" type="nomenclature">
<paragraph id="1FB409A78293931EFAE9C19C2F160CEE" pageId="0" pageNumber="47">
<taxonomicName id="BC391AD93604A1540B80772859390067" LSID="http://zoobank.org/A8A39314-2E0B-4E99-A65A-EBB595D16EF2" authority="gen. et" authorityName="gen. et" authorityYear="2022" class="Diplopoda" family="Julidae" genus="Bellatoiulus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Bellatoiulus golovatchi" order="Julida" pageId="0" pageNumber="47" phylum="Arthropoda" rank="species" species="golovatchi" status="sp. nov.">Bellatoiulus golovatchi gen. et</taxonomicName>
<taxonomicNameLabel id="DD7116BC2DCBB662361AEBD6FF07C1FC" pageId="0" pageNumber="47">sp. nov.</taxonomicNameLabel>
</paragraph>
</subSubSection>
<subSubSection id="BB24680F8A326CF111C5444681CB7B54" pageId="0" pageNumber="47" type="materials_examined">
<paragraph id="3DB1CE612197D10AEF80409B58F80CD5" pageId="0" pageNumber="47">Material examined.</paragraph>
<paragraph id="076F1D75343CEAF0636B4FFFA9B68DD1" pageId="0" pageNumber="47">
<materialsCitation id="85878D71CC8ECE4BD6634739775460B3" collectingDate="1987-06-01" collectingDateMax="1987-06-02" collectingDateMin="1987-06-01" collectorName="S. Golovatch, K. Eskov" country="Azerbaijan" elevation="825" location="Drmbon" specimenCount="♂" typeStatus="Holotype">
<emphasis id="B91AE9CF9F4D0DC4C42CF83B8B2E466F" bold="true" italics="true" pageId="0" pageNumber="47">
<typeStatus id="F5806DD1B07D98967761B94317A133B0">Holotype</typeStatus>
</emphasis>
:
<specimenCount id="6276413BB1DBB6732AFF1AF512C529F8">♂</specimenCount>
(unbroken) (ZMUM):
<collectingCountry id="531D61BAE8319CC0FA71A3F9CC2355E4" name="Azerbaijan">Azerbaijan</collectingCountry>
,
<location id="C1BEAA9F92F5271F5D42EC8DA4E9211B" LSID="urn:lsid:plazi:treatment:0D955BFA370F5C2DB82F8D8EDE4F6E00:C1BEAA9F92F5271F5D42EC8DA4E9211B" country="Azerbaijan" name="Drmbon">Drmbon</location>
[
<normalizedToken id="FB457E834D0A8772986B377CD06B6F94" originalValue="Heyvalı">Heyvali</normalizedToken>
] ca
<locationDeviation id="8C3AC4657AC33554A5C6960390C05E98" location="Mardakert">
<quantity id="706E29065ADA5B06643F69D8D6FF3CEA" metricMagnitude="4" metricUnit="m" metricValue="3.0" unit="km" value="30.0">30 km</quantity>
WSW of Mardakert
</locationDeviation>
[
<normalizedToken id="C350573A5DAF4753032F1A31810AE293" originalValue="Ağdərə">Agdərə</normalizedToken>
],
<elevation id="E7F80B95DA0ACA34A1FEB497BA21D162" metricMagnitude="2" metricUnit="m" metricValue="8.25" metricValueMax="8.5" metricValueMin="8.0" unit="m" value="825.0" valueMax="850.0" valueMin="800.0">
<quantity id="BA5D53287E4C32ACA610832ABD39C7DE" metricMagnitude="2" metricUnit="m" metricValue="8.25" metricValueMax="8.5" metricValueMin="8.0" unit="m" value="825.0" valueMax="850.0" valueMin="800.0">800-850 m</quantity>
a.s.l.
</elevation>
,
<taxonomicName id="597059137AFC4F1847F4FF653D974396" class="Magnoliopsida" family="Fagaceae" genus="Quercus" kingdom="Plantae" lsidName="Quercus" order="Fagales" pageId="0" pageNumber="47" phylum="Tracheophyta" rank="genus">
<emphasis id="BB029DB9C1C6E3C94B2CC8C584EE7BB2" italics="true" pageId="0" pageNumber="47">Quercus</emphasis>
</taxonomicName>
,
<taxonomicName id="63257E275221A3E33108EDEA0632BBDC" class="Magnoliopsida" family="Betulaceae" genus="Carpinus" kingdom="Plantae" lsidName="Carpinus" order="Fagales" pageId="0" pageNumber="47" phylum="Tracheophyta" rank="genus">
<emphasis id="5EE5C613607BF394BE88365C699099CC" italics="true" pageId="0" pageNumber="47">Carpinus</emphasis>
</taxonomicName>
,
<taxonomicName id="19483BE05DDBCCBC98D775F965397068" class="Magnoliopsida" family="Sapindaceae" genus="Acer" kingdom="Plantae" lsidName="Acer" order="Sapindales" pageId="0" pageNumber="47" phylum="Tracheophyta" rank="genus">
<emphasis id="D0394F283757753E9A7F98232F293BBA" italics="true" pageId="0" pageNumber="47">Acer</emphasis>
</taxonomicName>
, etc. forest, litter,
<collectingDate id="95F1E90ADF72DF0966DD8BC5E7B7BD49" value="1987-06-01" valueMax="1987-06-02" valueMin="1987-06-01">1-2.VI.1987</collectingDate>
,
<collectorName id="24AD79A2DDD65FE6054508BD1A29591F">S. Golovatch</collectorName>
&amp;
<collectorName id="24224C13B5464D69C6AFB5232217AA61">K. Eskov</collectorName>
leg.
</materialsCitation>
<emphasis id="B0EC5A345591E7649E9C0E1D40D0ADC6" bold="true" italics="true" pageId="0" pageNumber="47">
<typeStatus id="60BC1CB913FC836D507FF986CC55210B">Paratypes</typeStatus>
</emphasis>
:
<specimenCount id="FF78FF20EC01A800EE2767D66770C315" type="male">1 ♂</specimenCount>
(in 3 pieces, with dissected gonopods, leg-pair 2, a mid-body and an end-body leg),
<specimenCount id="0BB51D63508C624F8761652956B22101" type="female">1 ♀</specimenCount>
(in 2 pieces, right vulva dissected) (NMNHS),
<specimenCount id="2D5491F75D53B9A2C38449AC8D53E9F2" type="male">1 ♂</specimenCount>
(in 2 pieces, with dissected leg-pair 2 and a mid-body leg),
<specimenCount id="B9F76DE3C66FBDA03DDB826AFAA7135A" type="female">1 ♀</specimenCount>
(unbroken) (NHMD),
<specimenCount id="9CA5DE41F525183CB3840380AEF920D3" type="male">6 ♂♂</specimenCount>
(4 unbroken,
<quantity id="2CE67CEBC16E2FFF76BA589279DC2A2A" metricMagnitude="-2" metricUnit="m" metricValue="5.08" unit="in" value="2.0">2 in</quantity>
2 pieces, with dissected gonopods, head, antenna, leg-pairs 1, 2, 3, 8 and a mid-body leg),
<specimenCount id="64ABFBFC7CF34384C8E163DAA7119875" type="female">42 ♀♀</specimenCount>
(30 unbroken, 11 broken in two or more pieces, 1 with dissected vulvae and leg-pair 2),
<materialsCitation id="0720CE58E78A158B7933F507B2157F54" collectingDate="2022-01-01" collectingDateMax="2022-12-31" collectingDateMin="2022-01-01" collectorName="S. Golovatch &amp; K. Eskov" country="Azerbaijan" elevation="825" location="Drmbon" specimenCount="21" specimenCount-juvenile="21" typeStatus="paratype">
<specimenCount id="F575FCE2E1891D25AE8CDDBD65B72843" type="generic">21 juv.</specimenCount>
(ZMUM), same collecting data as for holotype
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="80B9646616D9695C035D0CC844A17E7B" pageId="0" pageNumber="47" type="type material">
<paragraph id="638A29D493BCEC7BC468DA95746C2651" pageId="0" pageNumber="47">Non-type material:</paragraph>
<paragraph id="32B56AFF7B8545AB74506ED9C6E2FE64" pageId="0" pageNumber="47">
7 ♂♂, 12 ♀♀, 2 juv. (ZMUM), Azerbaijan, Nadirkhanly [
<normalizedToken id="0E1237330D5A07337CFFAA1C77630082" originalValue="Nadirxanlı">Nadirxanli</normalizedToken>
] ca 12 km NE of Kelbajar [Kəlbəcər], 1200 m a.s.l.,
<taxonomicName id="117E81581ADD673B86850C62495D5061" class="Magnoliopsida" family="Oleaceae" genus="Fraxinus" kingdom="Plantae" lsidName="Fraxinus" order="Lamiales" pageId="0" pageNumber="47" phylum="Tracheophyta" rank="genus">
<emphasis id="234A911B3CAEE2913740FF72ACF8F9C7" italics="true" pageId="0" pageNumber="47">Fraxinus</emphasis>
</taxonomicName>
&amp;
<taxonomicName id="A60C1AED10F4C93BA777228FD6F44C7A" class="Magnoliopsida" family="Juglandaceae" genus="Juglans" kingdom="Plantae" lsidName="Juglans" order="Fagales" pageId="0" pageNumber="47" phylum="Tracheophyta" rank="genus">
<emphasis id="3D5FE3EBD4DA831A57686A60565ED8B2" italics="true" pageId="0" pageNumber="47">Juglans</emphasis>
</taxonomicName>
stand, litter, 1.VI.1987, S. Golovatch &amp; K. Eskov leg.; 2 ♂♂ (1 ♂ in 2 pieces, with dissected gonopods), 10 ♀♀, 8 juv. (ZMUM), Azerbaijan, ca 15 km WSW of Mardakert [
<normalizedToken id="8DF04C003B24B7A6B72FDEAE05FE2689" originalValue="Ağdərə">Agdərə</normalizedToken>
], 1100 m a.s.l.,
<taxonomicName id="6D65215651971FBDE192FA834E309F4D" class="Magnoliopsida" family="Fagaceae" genus="Quercus" kingdom="Plantae" lsidName="Quercus" order="Fagales" pageId="0" pageNumber="47" phylum="Tracheophyta" rank="genus">
<emphasis id="70E11C8DAAA0D65F47EC0E8A4FC01AA1" italics="true" pageId="0" pageNumber="47">Quercus</emphasis>
</taxonomicName>
,
<taxonomicName id="04B4EE067898790E7A8EB1CE059537DB" class="Magnoliopsida" family="Fagaceae" genus="Fagus" kingdom="Plantae" lsidName="Fagus" order="Fagales" pageId="0" pageNumber="47" phylum="Tracheophyta" rank="genus">
<emphasis id="3A3EDBD1CCE4E89E6BA402788A2EF847" italics="true" pageId="0" pageNumber="47">Fagus</emphasis>
</taxonomicName>
,
<taxonomicName id="7D0B78E1DBDEE30EFEC7D5429133811E" class="Magnoliopsida" family="Sapindaceae" genus="Acer" kingdom="Plantae" lsidName="Acer" order="Sapindales" pageId="0" pageNumber="47" phylum="Tracheophyta" rank="genus">
<emphasis id="A692AD722F7371BB7225122194C8571D" italics="true" pageId="0" pageNumber="47">Acer</emphasis>
</taxonomicName>
, etc. forest, litter, 2.VI.1987, S. Golovatch &amp; K. Eskov leg.
</paragraph>
</subSubSection>
<subSubSection id="064F8F7EB1386EE09C2B745C01CB542D" pageId="0" pageNumber="47" type="cybertypes">
<paragraph id="0077EAE260DB81B069183F006C1E5551" pageId="0" pageNumber="47">Cybertypes:</paragraph>
<paragraph id="654760D045622C96A695F3136641AFB5" pageId="0" pageNumber="47">http://morphobank.org/permalink/?P4180. Created from the physical ♂ holotype and a ♀ paratype.</paragraph>
</subSubSection>
<subSubSection id="37198E396B3C0B3A3F073508E980B5F6" pageId="0" pageNumber="47" type="name">
<paragraph id="EDBDA172615B128DB5D8659D33EEA367" pageId="0" pageNumber="47">Name.</paragraph>
<paragraph id="56451B155411DB28BC9BC048942C2D08" pageId="0" pageNumber="47">The new species honours our friend and colleague, Prof. Sergei Golovatch (Moscow, Russia), an outstanding myriapodologist and one of the collectors of the material used for the present species description.</paragraph>
</subSubSection>
<subSubSection id="94DC77A2C3BB9E6AD8EEA38ECB311F52" pageId="0" pageNumber="47" type="description">
<paragraph id="3C6DA437128CA0E2507F96886A13A951" pageId="0" pageNumber="47">Description.</paragraph>
<paragraph id="6D0C74B583FAD2C921C4F737A4F0A81A" pageId="0" pageNumber="47">Measurements: holotype with BRF 45+2+T, L = 12 mm, H = 0.6 mm; paratype and non-type ♂♂ with BRF 40-46+1-3+T, L = 10-12 mm, H = 0.5-0.6 mm; paratype and non-type ♀♀ with BRF 37-51+1-4+T, L = 9.5-14 mm, H = 0.5-0.7 mm.</paragraph>
<paragraph id="4F59EE084F9A058849535D684F5AFF95" pageId="0" pageNumber="47">
Colouration (Fig.
<figureCitation id="10631B971872338AF4D5A64B1A0AD01C" captionStart="Figure 1" captionStartId="F1" captionText="Figure 1. Habitus of Bellatoiulus golovatchi gen. et sp. nov. A male paratype B female paratype (both ZMUM). Scale bars: 1.0 mm." figureDoi="10.3897/zookeys.1097.81792.figure1" httpUri="https://binary.pensoft.net/fig/674257" pageId="0" pageNumber="47">1</figureCitation>
): completely pallid, gut and defence glands partly visible by transparency (alcohol-fixed material).
</paragraph>
<caption id="C6A73D4A1409B1C1B99A2CCC29BA766B" doi="10.3897/zookeys.1097.81792.figure1" httpUri="https://binary.pensoft.net/fig/674257" pageId="0" pageNumber="47" start="Figure 1" startId="F1">
<paragraph id="B6B952F37116FDEF8E144D05B49F2282" pageId="0" pageNumber="47">
<emphasis id="B0DBBA4A1F38DCCC7DD50969A6BC790A" bold="true" pageId="0" pageNumber="47">Figure 1.</emphasis>
Habitus of
<taxonomicName id="AE664AAA393075F473CEEBB28CFAB415" authorityName="gen. et" authorityYear="2022" class="Diplopoda" family="Julidae" genus="Bellatoiulus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Bellatoiulus golovatchi" order="Julida" pageId="0" pageNumber="47" phylum="Arthropoda" rank="species" species="golovatchi">
<emphasis id="73E8B465F099B84DC8CE432FE3F20CBE" italics="true" pageId="0" pageNumber="47">Bellatoiulus golovatchi</emphasis>
</taxonomicName>
gen. et sp. nov.
<emphasis id="0C9CC363B61562D74E37EBB94D13C908" bold="true" pageId="0" pageNumber="47">A</emphasis>
male paratype
<emphasis id="323B9473EE5BBFE08AAEDD563982D34B" bold="true" pageId="0" pageNumber="47">B</emphasis>
female paratype (both ZMUM). Scale bars: 1.0 mm.
</paragraph>
</caption>
<paragraph id="ED077E4848D40BCCF239266EC7EC5924" pageId="0" pageNumber="47">
Head (Figs
<figureCitation id="B68A525AED34DF8373DE0BC1F0AB5A21" captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. Bellatoiulus golovatchi gen. et sp. nov., male paratype (ZMUM). A head, ventral view B right antenna, lateral view C leg pair 1, frontal view D right leg 2, frontal view E left leg 3, caudal view F ozopores in midbody rings, lateral view G posterior part of body, lateral view. Abbreviation: ap adhesive pads. Scale bars: 0.1 mm (A, B, G); 0.01 mm (C); 0.02 mm (D, E, F)." figureDoi="10.3897/zookeys.1097.81792.figure2" httpUri="https://binary.pensoft.net/fig/674258" pageId="0" pageNumber="47">2A</figureCitation>
,
<figureCitation id="5795F4BECDE6D4A719A0ED56A588F994" captionStart="Figure 1" captionStartId="F1" captionText="Figure 1. Habitus of Bellatoiulus golovatchi gen. et sp. nov. A male paratype B female paratype (both ZMUM). Scale bars: 1.0 mm." figureDoi="10.3897/zookeys.1097.81792.figure1" httpUri="https://binary.pensoft.net/fig/674257" pageId="0" pageNumber="47">1</figureCitation>
,
<figureCitation id="335AD7AEE2D2E5BDCA454EE68F48C1FE" captionStart="Figure 5" captionStartId="F5" captionText="Figure 5. Bellatoiulus golovatchi gen. et sp. nov., male cybertype. A habitus, lateral view B anterior half of body, lateral view C anterior half of body with highlighted gonopods, lateral view D anterior body part with highlighted gonopods, ventral view. Abbreviation: g gonopods. Images not to scale." figureDoi="10.3897/zookeys.1097.81792.figure5" httpUri="https://binary.pensoft.net/fig/674261" pageId="0" pageNumber="47">5</figureCitation>
,
<figureCitation id="B2E321999864A2782B7DA4298D0248E5" captionStart="Figure 8" captionStartId="F8" captionText="Figure 8. Bellatoiulus golovatchi gen. et sp. nov., female cybertype. A habitus, lateral view B anterior and posterior body parts, ventrolateral and dorso-lateral views, respectively C anterior and posterior body parts with highlighted vulvae, lateral views D anterior body part with highlighted vulvae, ventral view. Abbreviation: v vulvae. Images not to scale." figureDoi="10.3897/zookeys.1097.81792.figure8" httpUri="https://binary.pensoft.net/fig/674264" pageId="0" pageNumber="47">8</figureCitation>
): ommatidia absent. 2 vertigial, 4 (usually) or 5 (rarely) supralabral and 8-14 labral setae. Antennae (Fig.
<figureCitation id="BB365486C8A63B3DB83030E39DA5242E" captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. Bellatoiulus golovatchi gen. et sp. nov., male paratype (ZMUM). A head, ventral view B right antenna, lateral view C leg pair 1, frontal view D right leg 2, frontal view E left leg 3, caudal view F ozopores in midbody rings, lateral view G posterior part of body, lateral view. Abbreviation: ap adhesive pads. Scale bars: 0.1 mm (A, B, G); 0.01 mm (C); 0.02 mm (D, E, F)." figureDoi="10.3897/zookeys.1097.81792.figure2" httpUri="https://binary.pensoft.net/fig/674258" pageId="0" pageNumber="47">2B</figureCitation>
) 1.4-1.7 times as long as head in males, and 1.3-1.4 times in females; antennomeres 2 and 5 subequal in length, somewhat longer than 3 and 4, and much longer than 6; 5 ca 1.6 times as long as broad and ca 1.5 times as broad as 2; the four apical cones relatively small; distal margins of antennomeres 5 and 6 dorsolaterally with several bacilliform sensilla basiconica, those on antennomere 5 of similar size to the apical cones, those on antennomere 6 somewhat finer. Mandibular stipites in males not expanded. Labrum tridentate. Gnathochilarium without peculiarities; proximal part of stipites non-setose, distally with three setae usual for the family, stipital palps normally developed, each bearing a group of extremely small apical sensilla; promentum small, rhomboid, separating lamellae linguales in roughly their proximal third, the latter each bearing three or four setae in a longitudinal row.
</paragraph>
<caption id="E6344079C2103D79CF51CB135B391180" doi="10.3897/zookeys.1097.81792.figure2" httpUri="https://binary.pensoft.net/fig/674258" pageId="0" pageNumber="47" start="Figure 2" startId="F2">
<paragraph id="6585B1ADC37405110028B4B5BF456462" pageId="0" pageNumber="47">
<emphasis id="ACB05470A1A08418AC4EDFE509147A39" bold="true" pageId="0" pageNumber="47">Figure 2.</emphasis>
<taxonomicName id="A371B39DE0D794C171D5221381026A81" authorityName="gen. et" authorityYear="2022" class="Diplopoda" family="Julidae" genus="Bellatoiulus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Bellatoiulus golovatchi" order="Julida" pageId="0" pageNumber="47" phylum="Arthropoda" rank="species" species="golovatchi">
<emphasis id="4B43A9A1E76DB5F3B71559A4CDFE7FBF" italics="true" pageId="0" pageNumber="47">Bellatoiulus golovatchi</emphasis>
</taxonomicName>
gen. et sp. nov., male paratype (ZMUM).
<emphasis id="27E6F8287A18D87CAC5E67308A0C9476" bold="true" pageId="0" pageNumber="47">A</emphasis>
head, ventral view
<emphasis id="8873186BB41F0A4008EF6E9636244AAA" bold="true" pageId="0" pageNumber="47">B</emphasis>
right antenna, lateral view
<emphasis id="6C66988DC7476929F1395F1E629AE5F2" bold="true" pageId="0" pageNumber="47">C</emphasis>
leg pair 1, frontal view
<emphasis id="AF0D9FD0F70A6D96CE1563248AAFC1FD" bold="true" pageId="0" pageNumber="47">D</emphasis>
right leg 2, frontal view
<emphasis id="55F653D155AEB4D58D686BBFF7CB8606" bold="true" pageId="0" pageNumber="47">E</emphasis>
left leg 3, caudal view
<emphasis id="A86C778E01A67776CE5922971BA75CEA" bold="true" pageId="0" pageNumber="47">F</emphasis>
ozopores in midbody rings, lateral view
<emphasis id="A80B5FBE9754A12712D8872ED7AA12DE" bold="true" pageId="0" pageNumber="47">G</emphasis>
posterior part of body, lateral view. Abbreviation:
<emphasis id="03507AD8D9B89AE36528AB3012AD33AA" bold="true" pageId="0" pageNumber="47">ap</emphasis>
adhesive pads. Scale bars: 0.1 mm (
<emphasis id="006A477E9BBD6E61593921A33986FE91" bold="true" pageId="0" pageNumber="47">A, B, G</emphasis>
); 0.01 mm (
<emphasis id="ABC3784FA34E0826B2EC9CF8AA0CC73C" bold="true" pageId="0" pageNumber="47">C</emphasis>
); 0.02 mm (
<emphasis id="CAFF30379663039B43A4E860937D2555" bold="true" pageId="0" pageNumber="47">D, E, F</emphasis>
).
</paragraph>
</caption>
<paragraph id="B25EB6C401A69AFFC2B7E0EE8A6326AB" pageId="0" pageNumber="47">
Trunk and legs: collum mostly smooth, with only two or three faint longitudinal grooves just next to posterolateral corner. Body rings (Figs
<figureCitation id="1FCFC02E2F600B2C7D96535AE97F920D" captionStart="Figure 1" captionStartId="F1" captionText="Figure 1. Habitus of Bellatoiulus golovatchi gen. et sp. nov. A male paratype B female paratype (both ZMUM). Scale bars: 1.0 mm." figureDoi="10.3897/zookeys.1097.81792.figure1" httpUri="https://binary.pensoft.net/fig/674257" pageId="0" pageNumber="47">1</figureCitation>
,
<figureCitation id="AC1D68F9D4D6F979BDD0AB731C3D2B31" captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. Bellatoiulus golovatchi gen. et sp. nov., male paratype (ZMUM). A head, ventral view B right antenna, lateral view C leg pair 1, frontal view D right leg 2, frontal view E left leg 3, caudal view F ozopores in midbody rings, lateral view G posterior part of body, lateral view. Abbreviation: ap adhesive pads. Scale bars: 0.1 mm (A, B, G); 0.01 mm (C); 0.02 mm (D, E, F)." figureDoi="10.3897/zookeys.1097.81792.figure2" httpUri="https://binary.pensoft.net/fig/674258" pageId="0" pageNumber="47">2F, G</figureCitation>
,
<figureCitation id="37F8516F9349302FAC3F32A31AC0D24B" captionStart="Figure 5" captionStartId="F5" captionText="Figure 5. Bellatoiulus golovatchi gen. et sp. nov., male cybertype. A habitus, lateral view B anterior half of body, lateral view C anterior half of body with highlighted gonopods, lateral view D anterior body part with highlighted gonopods, ventral view. Abbreviation: g gonopods. Images not to scale." figureDoi="10.3897/zookeys.1097.81792.figure5" httpUri="https://binary.pensoft.net/fig/674261" pageId="0" pageNumber="47">5</figureCitation>
,
<figureCitation id="F7D0D965E92854654A0432F8B29A4C46" captionStart="Figure 8" captionStartId="F8" captionText="Figure 8. Bellatoiulus golovatchi gen. et sp. nov., female cybertype. A habitus, lateral view B anterior and posterior body parts, ventrolateral and dorso-lateral views, respectively C anterior and posterior body parts with highlighted vulvae, lateral views D anterior body part with highlighted vulvae, ventral view. Abbreviation: v vulvae. Images not to scale." figureDoi="10.3897/zookeys.1097.81792.figure8" httpUri="https://binary.pensoft.net/fig/674264" pageId="0" pageNumber="47">8</figureCitation>
) slightly to moderately vaulted. Prozonae generally smooth, with sparse, very short and fine, irregular striae. Metazonae with sparse, shallow longitudinal striations, most striae not crossing entire length of metazona; hind margins with a rather dense whorl of erect to somewhat slanting setae, these being 0.12-0.2 times as long as H in males, and 0.1-0.17 times in females. Ozopores (Fig.
<figureCitation id="66C558F077470E2FD122B4A3B29E7AA6" captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. Bellatoiulus golovatchi gen. et sp. nov., male paratype (ZMUM). A head, ventral view B right antenna, lateral view C leg pair 1, frontal view D right leg 2, frontal view E left leg 3, caudal view F ozopores in midbody rings, lateral view G posterior part of body, lateral view. Abbreviation: ap adhesive pads. Scale bars: 0.1 mm (A, B, G); 0.01 mm (C); 0.02 mm (D, E, F)." figureDoi="10.3897/zookeys.1097.81792.figure2" httpUri="https://binary.pensoft.net/fig/674258" pageId="0" pageNumber="47">2F</figureCitation>
) small, placed behind pro-metazonal suture at ca 1/3-2/5 of metazonal length measured from front to back. Walking legs (Figs
<figureCitation id="761757086D1E7A786FD7B3F2B379CC8E" captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. Bellatoiulus golovatchi gen. et sp. nov., male paratype (ZMUM). A head, ventral view B right antenna, lateral view C leg pair 1, frontal view D right leg 2, frontal view E left leg 3, caudal view F ozopores in midbody rings, lateral view G posterior part of body, lateral view. Abbreviation: ap adhesive pads. Scale bars: 0.1 mm (A, B, G); 0.01 mm (C); 0.02 mm (D, E, F)." figureDoi="10.3897/zookeys.1097.81792.figure2" httpUri="https://binary.pensoft.net/fig/674258" pageId="0" pageNumber="47">2D, E</figureCitation>
,
<figureCitation id="C3DC2B468657A3F8DB109E0B01FDBC7D" captionStart="Figure 7" captionStartId="F7" captionText="Figure 7. Bellatoiulus golovatchi gen. et sp. nov., female paratypes (ZMUM) A-C and NMNHS D). A right leg 2, caudal view B left vulva, caudal view C right vulva, fronto-mesal, slightly apical view D right vulva, fronto-mesal view. Abbreviations: at anterior tube bu bursa hp hyaline protrusions mc median cleft mf median field op operculum pt posterior tube. Scale bars: 0.03 mm (A); 0.02 mm (B, C); 0.1 mm (D)." figureDoi="10.3897/zookeys.1097.81792.figure7" httpUri="https://binary.pensoft.net/fig/674263" pageId="0" pageNumber="47">7A</figureCitation>
) of moderate length: mid-body legs (Fig.
<figureCitation id="80F62C34433C2F10A5869B17234440A9" captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. Bellatoiulus golovatchi gen. et sp. nov., male paratype (ZMUM). A head, ventral view B right antenna, lateral view C leg pair 1, frontal view D right leg 2, frontal view E left leg 3, caudal view F ozopores in midbody rings, lateral view G posterior part of body, lateral view. Abbreviation: ap adhesive pads. Scale bars: 0.1 mm (A, B, G); 0.01 mm (C); 0.02 mm (D, E, F)." figureDoi="10.3897/zookeys.1097.81792.figure2" httpUri="https://binary.pensoft.net/fig/674258" pageId="0" pageNumber="47">2E</figureCitation>
) 0.8-0.9 times as long as H in males and 0.7-0.8 times in females. Tarsus of mid-body legs 1.5-1.7 times as long as tibia and ca 3.5 times as long as apical claw. Legs 2 in males with accessory claw, legs 3 and following pairs without; female legs altogether without accessory claw.
</paragraph>
<paragraph id="F37887D3ECC85A8DA271AC9086172E1D" pageId="0" pageNumber="47">
Telson (Fig.
<figureCitation id="300028618473F594DBD8EED93841B0F0" captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. Bellatoiulus golovatchi gen. et sp. nov., male paratype (ZMUM). A head, ventral view B right antenna, lateral view C leg pair 1, frontal view D right leg 2, frontal view E left leg 3, caudal view F ozopores in midbody rings, lateral view G posterior part of body, lateral view. Abbreviation: ap adhesive pads. Scale bars: 0.1 mm (A, B, G); 0.01 mm (C); 0.02 mm (D, E, F)." figureDoi="10.3897/zookeys.1097.81792.figure2" httpUri="https://binary.pensoft.net/fig/674258" pageId="0" pageNumber="47">2G</figureCitation>
): Pre-anal ring sparsely covered with long setae. Epiproct short and stout (considerably surpassed by the longest paraproctal setae), straight to slightly bent ventrad, roof-like, ending bluntly without distinct hyaline tip. Hypoproct small, rounded, not protruding behind hind margins of paraprocts in both sexes; ventral surface with a pair of median setae. Paraprocts sparsely to moderately setose, with a row of shorter setae along each
<normalizedToken id="5E006469B64B1E71F3B5C059CD7C8198" originalValue="paraproct’s">paraproct's</normalizedToken>
caudal margin.
</paragraph>
<paragraph id="B9BB9379F6E8F13ACDE35D69E1BE264B" pageId="0" pageNumber="47">
Male sexual characters: leg-pair 1 (Fig.
<figureCitation id="360E860AC850C284A9C85122DAE9D70A" captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. Bellatoiulus golovatchi gen. et sp. nov., male paratype (ZMUM). A head, ventral view B right antenna, lateral view C leg pair 1, frontal view D right leg 2, frontal view E left leg 3, caudal view F ozopores in midbody rings, lateral view G posterior part of body, lateral view. Abbreviation: ap adhesive pads. Scale bars: 0.1 mm (A, B, G); 0.01 mm (C); 0.02 mm (D, E, F)." figureDoi="10.3897/zookeys.1097.81792.figure2" httpUri="https://binary.pensoft.net/fig/674258" pageId="0" pageNumber="47">2C</figureCitation>
) 3-segmented, typical hooks oriented fronto-mesad; tibial outgrowth slender, tarsal remnant absent. Leg-pair 2 (Fig.
<figureCitation id="46182BD9F7E4DFD95C2B9094EE154C2E" captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. Bellatoiulus golovatchi gen. et sp. nov., male paratype (ZMUM). A head, ventral view B right antenna, lateral view C leg pair 1, frontal view D right leg 2, frontal view E left leg 3, caudal view F ozopores in midbody rings, lateral view G posterior part of body, lateral view. Abbreviation: ap adhesive pads. Scale bars: 0.1 mm (A, B, G); 0.01 mm (C); 0.02 mm (D, E, F)." figureDoi="10.3897/zookeys.1097.81792.figure2" httpUri="https://binary.pensoft.net/fig/674258" pageId="0" pageNumber="47">2D</figureCitation>
) slightly thicker than following pairs, ventrally with crested adhesive pads (
<emphasis id="E594358007D64323F405325E383E386F" bold="true" pageId="0" pageNumber="47">ap</emphasis>
), one each on postfemur and tibia; next several pairs (Fig.
<figureCitation id="63C7BC0EEDE40028D07EC1DAA7BA3CA2" captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. Bellatoiulus golovatchi gen. et sp. nov., male paratype (ZMUM). A head, ventral view B right antenna, lateral view C leg pair 1, frontal view D right leg 2, frontal view E left leg 3, caudal view F ozopores in midbody rings, lateral view G posterior part of body, lateral view. Abbreviation: ap adhesive pads. Scale bars: 0.1 mm (A, B, G); 0.01 mm (C); 0.02 mm (D, E, F)." figureDoi="10.3897/zookeys.1097.81792.figure2" httpUri="https://binary.pensoft.net/fig/674258" pageId="0" pageNumber="47">2E</figureCitation>
) with a small femoral pad in addition; the pads gradually diminishing towards end-body, completely disappearing in legs of the last third. Pleurotergum 7 (Fig.
<figureCitation id="2880D5D1300EACCDAD3D0EC2CB0F1EC9" captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. Bellatoiulus golovatchi gen. et sp. nov., male paratype (NMNHS). A right ventral edge of pleurotergum 7, ventro-lateral view B penis, caudal view C left promere, caudal view D left opisthomere, lateral, slightly caudal view E left opisthomere, mesal view. Abbreviations: bl basofrontal lobe fo fovea lp lateral process mg median grove mp mesal process mr mesal ridge ms mesomeral process ic (supposed) inner canal s solenomere, sc sperm canal. Scale bars: 0.1 mm." figureDoi="10.3897/zookeys.1097.81792.figure3" httpUri="https://binary.pensoft.net/fig/674259" pageId="0" pageNumber="47">3A</figureCitation>
) ventrally forming well-pronounced, shovel-like lobes originating from the border zone between pro- and metazona, protruding ventrad behind gonopods, not concealing them from lateral view. Penis (Fig.
<figureCitation id="FC57FBEC98508F573E1CD099FD6565EF" captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. Bellatoiulus golovatchi gen. et sp. nov., male paratype (NMNHS). A right ventral edge of pleurotergum 7, ventro-lateral view B penis, caudal view C left promere, caudal view D left opisthomere, lateral, slightly caudal view E left opisthomere, mesal view. Abbreviations: bl basofrontal lobe fo fovea lp lateral process mg median grove mp mesal process mr mesal ridge ms mesomeral process ic (supposed) inner canal s solenomere, sc sperm canal. Scale bars: 0.1 mm." figureDoi="10.3897/zookeys.1097.81792.figure3" httpUri="https://binary.pensoft.net/fig/674259" pageId="0" pageNumber="47">3B</figureCitation>
) very small, non-sclerotised, set deeply above coxae 2, with a short basal part and rather slender, flattened, completely transparent apical lobes.
</paragraph>
<caption id="994E42CAF551D8A8B03D8F57C4FD7ADC" doi="10.3897/zookeys.1097.81792.figure3" httpUri="https://binary.pensoft.net/fig/674259" pageId="0" pageNumber="47" start="Figure 3" startId="F3">
<paragraph id="78B5E4A9B94C44E78C48CED9812EAF3D" pageId="0" pageNumber="47">
<emphasis id="2616EE37D8261280D51DA452E1A4A193" bold="true" pageId="0" pageNumber="47">Figure 3.</emphasis>
<taxonomicName id="7726C61E1D236A8A919FFC1658CC584A" authorityName="gen. et" authorityYear="2022" class="Diplopoda" family="Julidae" genus="Bellatoiulus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Bellatoiulus golovatchi" order="Julida" pageId="0" pageNumber="47" phylum="Arthropoda" rank="species" species="golovatchi">
<emphasis id="D1E056CA5B831A3FAB56940F5C1E1E34" italics="true" pageId="0" pageNumber="47">Bellatoiulus golovatchi</emphasis>
</taxonomicName>
gen. et sp. nov., male paratype (NMNHS).
<emphasis id="65D1F3FB1FBCF8FB8D494802F14D22F0" bold="true" pageId="0" pageNumber="47">A</emphasis>
right ventral edge of pleurotergum 7, ventro-lateral view
<emphasis id="5543B14DD43C9F27F67157A6F1BB70E6" bold="true" pageId="0" pageNumber="47">B</emphasis>
penis, caudal view
<emphasis id="3C7640333B551DF44D27062A7A38EAC3" bold="true" pageId="0" pageNumber="47">C</emphasis>
left promere, caudal view
<emphasis id="1E2AC38C41F8223DB0F3BDDBCB63E649" bold="true" pageId="0" pageNumber="47">D</emphasis>
left opisthomere, lateral, slightly caudal view
<emphasis id="4537F5C524113B6F4BD99E28A99F7F96" bold="true" pageId="0" pageNumber="47">E</emphasis>
left opisthomere, mesal view. Abbreviations:
<emphasis id="8DBE6304E3581FC6D3ECF4349D6FAD63" bold="true" pageId="0" pageNumber="47">bl</emphasis>
basofrontal lobe
<emphasis id="5E0E6624C557CED8A3641CF03A737C31" bold="true" pageId="0" pageNumber="47">fo</emphasis>
fovea
<emphasis id="95394FE45045743AD2DAFD3E93306C41" bold="true" pageId="0" pageNumber="47">lp</emphasis>
lateral process
<emphasis id="BA87DC18E573A237DA7E7CF3DB09C87E" bold="true" pageId="0" pageNumber="47">mg</emphasis>
median grove
<emphasis id="26561CA56761BAE0405C79D23B18AEB6" bold="true" pageId="0" pageNumber="47">mp</emphasis>
mesal process
<emphasis id="B2054EA850B423FCC31D77C909CF6BC5" bold="true" pageId="0" pageNumber="47">mr</emphasis>
mesal ridge
<emphasis id="BA01A21240A7308841C4EC03E887C09F" bold="true" pageId="0" pageNumber="47">ms</emphasis>
mesomeral process
<emphasis id="E217D88448BD80E85A5860A7C5729F44" bold="true" pageId="0" pageNumber="47">ic</emphasis>
(supposed) inner canal
<emphasis id="1993C45336B2F8E55A88D751638CF119" bold="true" pageId="0" pageNumber="47">s</emphasis>
solenomere,
<emphasis id="D1E01CECAD0D511EA6A3C096D57D0234" bold="true" pageId="0" pageNumber="47">sc</emphasis>
sperm canal. Scale bars: 0.1 mm.
</paragraph>
</caption>
<paragraph id="37A6D1FE1BC0B7912C8497274B9A3F8E" pageId="0" pageNumber="47">
Gonopods (Figs
<figureCitation id="8398032B91B9E956D150AD69B062102C" captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. Bellatoiulus golovatchi gen. et sp. nov., male paratype (NMNHS). A right ventral edge of pleurotergum 7, ventro-lateral view B penis, caudal view C left promere, caudal view D left opisthomere, lateral, slightly caudal view E left opisthomere, mesal view. Abbreviations: bl basofrontal lobe fo fovea lp lateral process mg median grove mp mesal process mr mesal ridge ms mesomeral process ic (supposed) inner canal s solenomere, sc sperm canal. Scale bars: 0.1 mm." figureDoi="10.3897/zookeys.1097.81792.figure3" httpUri="https://binary.pensoft.net/fig/674259" pageId="0" pageNumber="47">3C-E</figureCitation>
,
<figureCitation id="257E0629EF63ADB60A0B048FD231B3B0" captionStart="Figure 4" captionStartId="F4" captionText="Figure 4. Bellatoiulus golovatchi gen. et sp. nov., male paratype (ZMUM). A left gonopods, mesal view (promere strongly turned mesad) B right gonopods, fronto-lateral view C left gonopods, frontal view D mesal process of right promere, fronto-lateral view E end of left opisthomere, meso-frontal view F end of right opisthomere, fronto-lateral view. Abbreviations: bl basofrontal lobe fo apical fovea lp lateral process mg median groove mp mesal process mr mesal ridge ms mesomeral process o opisthomere p promere s solenomere sc sperm canal. Scale bars: 0.03 mm (A, B, F); 0.02 mm (C, D, E)." figureDoi="10.3897/zookeys.1097.81792.figure4" httpUri="https://binary.pensoft.net/fig/674260" pageId="0" pageNumber="47">4</figureCitation>
,
<figureCitation id="A11C7185395BA138F879148344033AEA" captionStart="Figure 6" captionStartId="F6" captionText="Figure 6. Bellatoiulus golovatchi gen. et sp. nov., male cybertype. A gonopods, caudal view B gonopods, frontal view C gonopods, lateral view D gonopods, ventral view E left gonopod, mesal view. Abbreviations: bl basofrontal lobe lp lateral process mp mesal process ms mesomeral process o opisthomere p promere s solenomere. Images not to scale." figureDoi="10.3897/zookeys.1097.81792.figure6" httpUri="https://binary.pensoft.net/fig/674262" pageId="0" pageNumber="47">6</figureCitation>
, and
<emphasis id="7207F680A9F0CD5376D2443DB27DE151" bold="true" pageId="0" pageNumber="47">g</emphasis>
in Fig.
<figureCitation id="3094F0B3A7319FD3762B725125A54881" captionStart="Figure 5" captionStartId="F5" captionText="Figure 5. Bellatoiulus golovatchi gen. et sp. nov., male cybertype. A habitus, lateral view B anterior half of body, lateral view C anterior half of body with highlighted gonopods, lateral view D anterior body part with highlighted gonopods, ventral view. Abbreviation: g gonopods. Images not to scale." figureDoi="10.3897/zookeys.1097.81792.figure5" httpUri="https://binary.pensoft.net/fig/674261" pageId="0" pageNumber="47">5C, D</figureCitation>
):
<emphasis id="7F5FF666F293732EF4AD55CA8AE65925" italics="true" pageId="0" pageNumber="47">in situ</emphasis>
protruding from gonopodal sinus with their apical parts. Promere (Fig.
<figureCitation id="FE2B60E23E78FF4499F0AF99EE5FD6EB" captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. Bellatoiulus golovatchi gen. et sp. nov., male paratype (NMNHS). A right ventral edge of pleurotergum 7, ventro-lateral view B penis, caudal view C left promere, caudal view D left opisthomere, lateral, slightly caudal view E left opisthomere, mesal view. Abbreviations: bl basofrontal lobe fo fovea lp lateral process mg median grove mp mesal process mr mesal ridge ms mesomeral process ic (supposed) inner canal s solenomere, sc sperm canal. Scale bars: 0.1 mm." figureDoi="10.3897/zookeys.1097.81792.figure3" httpUri="https://binary.pensoft.net/fig/674259" pageId="0" pageNumber="47">3C</figureCitation>
, and
<emphasis id="9C00B810D526B864B91D3D42FD950765" bold="true" pageId="0" pageNumber="47">p</emphasis>
in Figs
<figureCitation id="27E0293423F3587311A81D7DC8F925FC" captionStart="Figure 4" captionStartId="F4" captionText="Figure 4. Bellatoiulus golovatchi gen. et sp. nov., male paratype (ZMUM). A left gonopods, mesal view (promere strongly turned mesad) B right gonopods, fronto-lateral view C left gonopods, frontal view D mesal process of right promere, fronto-lateral view E end of left opisthomere, meso-frontal view F end of right opisthomere, fronto-lateral view. Abbreviations: bl basofrontal lobe fo apical fovea lp lateral process mg median groove mp mesal process mr mesal ridge ms mesomeral process o opisthomere p promere s solenomere sc sperm canal. Scale bars: 0.03 mm (A, B, F); 0.02 mm (C, D, E)." figureDoi="10.3897/zookeys.1097.81792.figure4" httpUri="https://binary.pensoft.net/fig/674260" pageId="0" pageNumber="47">4A-C</figureCitation>
,
<figureCitation id="EC72B4A8CCB18CA028BBDFF03CEEC152" captionStart="Figure 6" captionStartId="F6" captionText="Figure 6. Bellatoiulus golovatchi gen. et sp. nov., male cybertype. A gonopods, caudal view B gonopods, frontal view C gonopods, lateral view D gonopods, ventral view E left gonopod, mesal view. Abbreviations: bl basofrontal lobe lp lateral process mp mesal process ms mesomeral process o opisthomere p promere s solenomere. Images not to scale." figureDoi="10.3897/zookeys.1097.81792.figure6" httpUri="https://binary.pensoft.net/fig/674262" pageId="0" pageNumber="47">6B-E</figureCitation>
) rather stout, roughly quadrangular, insignificantly widening distally, with slightly concave mesal margin and gently sigmoid lateral margin, connected in a flat to broadly rounded apical margin; caudal face with a broad median groove (
<emphasis id="78688AC556A908007EBBC94F763A88BB" bold="true" pageId="0" pageNumber="47">mg</emphasis>
), and a broad, not too strongly pronounced mesal ridge (
<emphasis id="729D2479B7A409F5356BED6EE8FE82A5" bold="true" pageId="0" pageNumber="47">mr</emphasis>
) bearing one or two short setae at mid-height, distally extending in a long and straight mesal process (Fig.
<figureCitation id="7E200032B4DBBD4BF499A1214CB71F20" captionStart="Figure 4" captionStartId="F4" captionText="Figure 4. Bellatoiulus golovatchi gen. et sp. nov., male paratype (ZMUM). A left gonopods, mesal view (promere strongly turned mesad) B right gonopods, fronto-lateral view C left gonopods, frontal view D mesal process of right promere, fronto-lateral view E end of left opisthomere, meso-frontal view F end of right opisthomere, fronto-lateral view. Abbreviations: bl basofrontal lobe fo apical fovea lp lateral process mg median groove mp mesal process mr mesal ridge ms mesomeral process o opisthomere p promere s solenomere sc sperm canal. Scale bars: 0.03 mm (A, B, F); 0.02 mm (C, D, E)." figureDoi="10.3897/zookeys.1097.81792.figure4" httpUri="https://binary.pensoft.net/fig/674260" pageId="0" pageNumber="47">4D</figureCitation>
, and
<emphasis id="BFA490158AC5B35BFC696DF762DA1BDB" bold="true" pageId="0" pageNumber="47">mp</emphasis>
in Figs
<figureCitation id="8B1AD2AC622724AA5D91221DF908B938" captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. Bellatoiulus golovatchi gen. et sp. nov., male paratype (NMNHS). A right ventral edge of pleurotergum 7, ventro-lateral view B penis, caudal view C left promere, caudal view D left opisthomere, lateral, slightly caudal view E left opisthomere, mesal view. Abbreviations: bl basofrontal lobe fo fovea lp lateral process mg median grove mp mesal process mr mesal ridge ms mesomeral process ic (supposed) inner canal s solenomere, sc sperm canal. Scale bars: 0.1 mm." figureDoi="10.3897/zookeys.1097.81792.figure3" httpUri="https://binary.pensoft.net/fig/674259" pageId="0" pageNumber="47">3C</figureCitation>
,
<figureCitation id="1D5B438AD7CE8ED4412ECF05FAF75269" captionStart="Figure 4" captionStartId="F4" captionText="Figure 4. Bellatoiulus golovatchi gen. et sp. nov., male paratype (ZMUM). A left gonopods, mesal view (promere strongly turned mesad) B right gonopods, fronto-lateral view C left gonopods, frontal view D mesal process of right promere, fronto-lateral view E end of left opisthomere, meso-frontal view F end of right opisthomere, fronto-lateral view. Abbreviations: bl basofrontal lobe fo apical fovea lp lateral process mg median groove mp mesal process mr mesal ridge ms mesomeral process o opisthomere p promere s solenomere sc sperm canal. Scale bars: 0.03 mm (A, B, F); 0.02 mm (C, D, E)." figureDoi="10.3897/zookeys.1097.81792.figure4" httpUri="https://binary.pensoft.net/fig/674260" pageId="0" pageNumber="47">4A-C</figureCitation>
,
<figureCitation id="0B384D8643E2BDB702FE899D037D04A3" captionStart="Figure 6" captionStartId="F6" captionText="Figure 6. Bellatoiulus golovatchi gen. et sp. nov., male cybertype. A gonopods, caudal view B gonopods, frontal view C gonopods, lateral view D gonopods, ventral view E left gonopod, mesal view. Abbreviations: bl basofrontal lobe lp lateral process mp mesal process ms mesomeral process o opisthomere p promere s solenomere. Images not to scale." figureDoi="10.3897/zookeys.1097.81792.figure6" httpUri="https://binary.pensoft.net/fig/674262" pageId="0" pageNumber="47">6</figureCitation>
), with a somewhat clavate apex bearing several blunt teeth on its frontal side. Opisthomere (Figs
<figureCitation id="3C4D2C927CBF4C2DA22287289F72B3D5" captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. Bellatoiulus golovatchi gen. et sp. nov., male paratype (NMNHS). A right ventral edge of pleurotergum 7, ventro-lateral view B penis, caudal view C left promere, caudal view D left opisthomere, lateral, slightly caudal view E left opisthomere, mesal view. Abbreviations: bl basofrontal lobe fo fovea lp lateral process mg median grove mp mesal process mr mesal ridge ms mesomeral process ic (supposed) inner canal s solenomere, sc sperm canal. Scale bars: 0.1 mm." figureDoi="10.3897/zookeys.1097.81792.figure3" httpUri="https://binary.pensoft.net/fig/674259" pageId="0" pageNumber="47">3D, E</figureCitation>
,
<figureCitation id="DD64394B88AA2ECAB5C1C20C603C6584" captionStart="Figure 4" captionStartId="F4" captionText="Figure 4. Bellatoiulus golovatchi gen. et sp. nov., male paratype (ZMUM). A left gonopods, mesal view (promere strongly turned mesad) B right gonopods, fronto-lateral view C left gonopods, frontal view D mesal process of right promere, fronto-lateral view E end of left opisthomere, meso-frontal view F end of right opisthomere, fronto-lateral view. Abbreviations: bl basofrontal lobe fo apical fovea lp lateral process mg median groove mp mesal process mr mesal ridge ms mesomeral process o opisthomere p promere s solenomere sc sperm canal. Scale bars: 0.03 mm (A, B, F); 0.02 mm (C, D, E)." figureDoi="10.3897/zookeys.1097.81792.figure4" httpUri="https://binary.pensoft.net/fig/674260" pageId="0" pageNumber="47">4E, F</figureCitation>
, and
<emphasis id="B35BECCB10EEE96DA423AA2AEF4C9CAC" bold="true" pageId="0" pageNumber="47">o</emphasis>
in Figs
<figureCitation id="9CCC87577024A698541C848A8EDB1BC9" captionStart="Figure 4" captionStartId="F4" captionText="Figure 4. Bellatoiulus golovatchi gen. et sp. nov., male paratype (ZMUM). A left gonopods, mesal view (promere strongly turned mesad) B right gonopods, fronto-lateral view C left gonopods, frontal view D mesal process of right promere, fronto-lateral view E end of left opisthomere, meso-frontal view F end of right opisthomere, fronto-lateral view. Abbreviations: bl basofrontal lobe fo apical fovea lp lateral process mg median groove mp mesal process mr mesal ridge ms mesomeral process o opisthomere p promere s solenomere sc sperm canal. Scale bars: 0.03 mm (A, B, F); 0.02 mm (C, D, E)." figureDoi="10.3897/zookeys.1097.81792.figure4" httpUri="https://binary.pensoft.net/fig/674260" pageId="0" pageNumber="47">4A, B</figureCitation>
,
<figureCitation id="DDE828F3E4BA85E0F1E80675D7A03467" captionStart="Figure 6" captionStartId="F6" captionText="Figure 6. Bellatoiulus golovatchi gen. et sp. nov., male cybertype. A gonopods, caudal view B gonopods, frontal view C gonopods, lateral view D gonopods, ventral view E left gonopod, mesal view. Abbreviations: bl basofrontal lobe lp lateral process mp mesal process ms mesomeral process o opisthomere p promere s solenomere. Images not to scale." figureDoi="10.3897/zookeys.1097.81792.figure6" httpUri="https://binary.pensoft.net/fig/674262" pageId="0" pageNumber="47">6A, C-E</figureCitation>
) relatively slender, reaching nearly level to mesal process of promere; frontal face with a broad, massive, microsquamous, basofrontal lobe (
<emphasis id="074656F7C7F51CEDA9CF684D0E6A95E1" bold="true" pageId="0" pageNumber="47">bl</emphasis>
); mesomeral process (
<emphasis id="E5A15E2AA90454CC6C29A4B41FC15F2A" bold="true" pageId="0" pageNumber="47">ms</emphasis>
) originating just distally to the lobe, being strongly flattened, somewhat twisted around its axis, distally widening, apically bearing a group of minute seti- or spiniform filaments; a well-developed club or ping-pong paddle-like lateral process (
<emphasis id="EDB7BD08D2EB8398E79BD8D02AF611CB" bold="true" pageId="0" pageNumber="47">lp</emphasis>
) with smooth surface, almost reaching level to solenomere; solenomere (
<emphasis id="04CFC630134BE0C2E7ACC2C9895AE66C" bold="true" pageId="0" pageNumber="47">s</emphasis>
) tubular, reaching higher than mesomeral process, distally somewhat bent latero-frontad, with a deep and narrow apical fovea (
<emphasis id="5B96F71E8F5B386B2F0C2BC36976ED95" bold="true" pageId="0" pageNumber="47">fo</emphasis>
), and an (presumably) inner canal (
<emphasis id="4E4B13B2138D109FB61AA2F869785983" bold="true" pageId="0" pageNumber="47">ic</emphasis>
) running vertically right basally to fovea, mesal side with a sperm canal (
<emphasis id="E285F32C91B7496EEF9D4FB717394835" bold="true" pageId="0" pageNumber="47">sc</emphasis>
), distally somewhat deviating frontad, caudal lamella absent.
</paragraph>
<caption id="36F754B3E84F5C291231CC9DC1BBD3CC" doi="10.3897/zookeys.1097.81792.figure4" httpUri="https://binary.pensoft.net/fig/674260" pageId="0" pageNumber="47" start="Figure 4" startId="F4">
<paragraph id="87F50F9270CA9F2456ABC85DDF8B6712" pageId="0" pageNumber="47">
<emphasis id="E4CF6039E14DF63BEBA979A50F5FB05E" bold="true" pageId="0" pageNumber="47">Figure 4.</emphasis>
<taxonomicName id="AB9F03E9F39A9EDC51C3A30703005782" authorityName="gen. et" authorityYear="2022" class="Diplopoda" family="Julidae" genus="Bellatoiulus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Bellatoiulus golovatchi" order="Julida" pageId="0" pageNumber="47" phylum="Arthropoda" rank="species" species="golovatchi">
<emphasis id="8AEF6076BD4E491C2FDB0EF89BFAFA4C" italics="true" pageId="0" pageNumber="47">Bellatoiulus golovatchi</emphasis>
</taxonomicName>
gen. et sp. nov., male paratype (ZMUM).
<emphasis id="68540BADFDBE6C33F1BAE3CE2D29BB6E" bold="true" pageId="0" pageNumber="47">A</emphasis>
left gonopods, mesal view (promere strongly turned mesad)
<emphasis id="FD1D3FBD83751BC6C8D1398D284C857E" bold="true" pageId="0" pageNumber="47">B</emphasis>
right gonopods, fronto-lateral view
<emphasis id="A35E33554570B73648C3B60935B3E045" bold="true" pageId="0" pageNumber="47">C</emphasis>
left gonopods, frontal view
<emphasis id="083A16CD55A44157E6B4967DB1EAE129" bold="true" pageId="0" pageNumber="47">D</emphasis>
mesal process of right promere, fronto-lateral view
<emphasis id="79324A9E0B6CBA150D6A7F0B29F6CE73" bold="true" pageId="0" pageNumber="47">E</emphasis>
end of left opisthomere, meso-frontal view
<emphasis id="9387A880C9B5DB4ADB08B9F6DAB80D01" bold="true" pageId="0" pageNumber="47">F</emphasis>
end of right opisthomere, fronto-lateral view. Abbreviations:
<emphasis id="2B8145F681EB2EDB3A145983BCF0764F" bold="true" pageId="0" pageNumber="47">bl</emphasis>
basofrontal lobe
<emphasis id="72BE44C99443D4C7926E660DFD18D7BB" bold="true" pageId="0" pageNumber="47">fo</emphasis>
apical fovea
<emphasis id="89A2A20C7520A63887FE2229B97BD612" bold="true" pageId="0" pageNumber="47">lp</emphasis>
lateral process
<emphasis id="357498BEEEA140DB5E4E16DF042BF2E6" bold="true" pageId="0" pageNumber="47">mg</emphasis>
median groove
<emphasis id="69DF1915ECA2EBEDA14F0A5571CEFB36" bold="true" pageId="0" pageNumber="47">mp</emphasis>
mesal process
<emphasis id="B1CA0A0780BC36DEA4F63B72F163B958" bold="true" pageId="0" pageNumber="47">mr</emphasis>
mesal ridge
<emphasis id="48CF86A53307A76646863D5A44685E3F" bold="true" pageId="0" pageNumber="47">ms</emphasis>
mesomeral process
<emphasis id="E3B70D53C3B1CB761B9495CA3212B888" bold="true" pageId="0" pageNumber="47">o</emphasis>
opisthomere
<emphasis id="BBCBA2FE38AE896AE439D1A57B77F9E2" bold="true" pageId="0" pageNumber="47">p</emphasis>
promere
<emphasis id="6DE340CC72109744A6F61B4FA9E37DC7" bold="true" pageId="0" pageNumber="47">s</emphasis>
solenomere
<emphasis id="A3737EF086C697FF56287E9D0BADBE91" bold="true" pageId="0" pageNumber="47">sc</emphasis>
sperm canal. Scale bars: 0.03 mm (
<emphasis id="49165399DADD7C2D4031EE70974E6862" bold="true" pageId="0" pageNumber="47">A, B, F</emphasis>
); 0.02 mm (
<emphasis id="4B45E93137806965D4A1D04EF78B436F" bold="true" pageId="0" pageNumber="47">C, D, E</emphasis>
).
</paragraph>
</caption>
<caption id="E3502643A87D5F3E493DD1CE5C23D5A1" doi="10.3897/zookeys.1097.81792.figure5" httpUri="https://binary.pensoft.net/fig/674261" pageId="0" pageNumber="47" start="Figure 5" startId="F5">
<paragraph id="6F3F03562C409A507A094A8AB6F981A1" pageId="0" pageNumber="47">
<emphasis id="8F06B89D0AE6A59E843B7B164B199BB3" bold="true" pageId="0" pageNumber="47">Figure 5.</emphasis>
<taxonomicName id="14FF8B8C0B1E0D0DC995371AEBB63875" authorityName="gen. et" authorityYear="2022" class="Diplopoda" family="Julidae" genus="Bellatoiulus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Bellatoiulus golovatchi" order="Julida" pageId="0" pageNumber="47" phylum="Arthropoda" rank="species" species="golovatchi">
<emphasis id="00F0A3D5DC6B32CF87F067716C47470C" italics="true" pageId="0" pageNumber="47">Bellatoiulus golovatchi</emphasis>
</taxonomicName>
gen. et sp. nov., male cybertype.
<emphasis id="FB0E822ACD31583203C8120897AE02DF" bold="true" pageId="0" pageNumber="47">A</emphasis>
habitus, lateral view
<emphasis id="F16AD3B45A10C238F0AB31DB284697E6" bold="true" pageId="0" pageNumber="47">B</emphasis>
anterior half of body, lateral view
<emphasis id="2ABE9004B00D455B7EB12A27888B98BB" bold="true" pageId="0" pageNumber="47">C</emphasis>
anterior half of body with highlighted gonopods, lateral view
<emphasis id="9B2E33B77D370115D9CA5F0F9593AC59" bold="true" pageId="0" pageNumber="47">D</emphasis>
anterior body part with highlighted gonopods, ventral view. Abbreviation:
<emphasis id="097A8FF8FF8A27773F9E6F781BB90DE6" bold="true" pageId="0" pageNumber="47">g</emphasis>
gonopods. Images not to scale.
</paragraph>
</caption>
<caption id="784459A221F8643BEB9E6AFDA0B7197A" doi="10.3897/zookeys.1097.81792.figure6" httpUri="https://binary.pensoft.net/fig/674262" pageId="0" pageNumber="47" start="Figure 6" startId="F6">
<paragraph id="0B27782158E11846D4D87962A159BBD3" pageId="0" pageNumber="47">
<emphasis id="2E411920618379C45063387E5CBD56B0" bold="true" pageId="0" pageNumber="47">Figure 6.</emphasis>
<taxonomicName id="437659E86D86A863360615CC248E4372" authorityName="gen. et" authorityYear="2022" class="Diplopoda" family="Julidae" genus="Bellatoiulus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Bellatoiulus golovatchi" order="Julida" pageId="0" pageNumber="47" phylum="Arthropoda" rank="species" species="golovatchi">
<emphasis id="AEA13C05E3AFE2A7FB53FF21877C3AD9" italics="true" pageId="0" pageNumber="47">Bellatoiulus golovatchi</emphasis>
</taxonomicName>
gen. et sp. nov., male cybertype.
<emphasis id="257F7F869BE361ACEACD0CF23D4F540D" bold="true" pageId="0" pageNumber="47">A</emphasis>
gonopods, caudal view
<emphasis id="6169829002E79E78F3540591D9DEE77A" bold="true" pageId="0" pageNumber="47">B</emphasis>
gonopods, frontal view
<emphasis id="2EA09E42CA9127D9D69D632AF34F941E" bold="true" pageId="0" pageNumber="47">C</emphasis>
gonopods, lateral view
<emphasis id="F159FEC4218E656F258FC9B407DD5533" bold="true" pageId="0" pageNumber="47">D</emphasis>
gonopods, ventral view
<emphasis id="EC94A68DC100BD14B6924EFDB5FD7054" bold="true" pageId="0" pageNumber="47">E</emphasis>
left gonopod, mesal view. Abbreviations:
<emphasis id="4220C8C594B03A28C4EE3B540202C93C" bold="true" pageId="0" pageNumber="47">bl</emphasis>
basofrontal lobe
<emphasis id="9E7899C93D53E8A4C12148465D5A9521" bold="true" pageId="0" pageNumber="47">lp</emphasis>
lateral process
<emphasis id="B6EE93C3F8D28032A631087A72224C0E" bold="true" pageId="0" pageNumber="47">mp</emphasis>
mesal process
<emphasis id="55E4398233FD81D45164B403A0EA60DF" bold="true" pageId="0" pageNumber="47">ms</emphasis>
mesomeral process
<emphasis id="AE03B6096813F8D5EAEAECBB31E2FACD" bold="true" pageId="0" pageNumber="47">o</emphasis>
opisthomere
<emphasis id="12B75389EE05A60AE7672E7575CBA593" bold="true" pageId="0" pageNumber="47">p</emphasis>
promere
<emphasis id="015DA8B1206A7AA28B59871C7666AFAA" bold="true" pageId="0" pageNumber="47">s</emphasis>
solenomere. Images not to scale.
</paragraph>
</caption>
<paragraph id="9BEB878F7C7BEE0922D6E30B613EB824" pageId="0" pageNumber="47">
Female sexual characters: leg-pairs 1 (Fig.
<figureCitation id="52FB2D697197E8A2511FB2398AFBC023" captionStart="Figure 7" captionStartId="F7" captionText="Figure 7. Bellatoiulus golovatchi gen. et sp. nov., female paratypes (ZMUM) A-C and NMNHS D). A right leg 2, caudal view B left vulva, caudal view C right vulva, fronto-mesal, slightly apical view D right vulva, fronto-mesal view. Abbreviations: at anterior tube bu bursa hp hyaline protrusions mc median cleft mf median field op operculum pt posterior tube. Scale bars: 0.03 mm (A); 0.02 mm (B, C); 0.1 mm (D)." figureDoi="10.3897/zookeys.1097.81792.figure7" httpUri="https://binary.pensoft.net/fig/674263" pageId="0" pageNumber="47">7A</figureCitation>
) and 2 only slightly thicker than following legs. Vulva (Figs
<figureCitation id="8286EF4A3622B354F4F5EB07C4ACEDEF" captionStart="Figure 7" captionStartId="F7" captionText="Figure 7. Bellatoiulus golovatchi gen. et sp. nov., female paratypes (ZMUM) A-C and NMNHS D). A right leg 2, caudal view B left vulva, caudal view C right vulva, fronto-mesal, slightly apical view D right vulva, fronto-mesal view. Abbreviations: at anterior tube bu bursa hp hyaline protrusions mc median cleft mf median field op operculum pt posterior tube. Scale bars: 0.03 mm (A); 0.02 mm (B, C); 0.1 mm (D)." figureDoi="10.3897/zookeys.1097.81792.figure7" httpUri="https://binary.pensoft.net/fig/674263" pageId="0" pageNumber="47">7B-D</figureCitation>
,
<figureCitation id="31B6A33147C39A311FC177B6BFAB122C" captionStart="Figure 9" captionStartId="F9" captionText="Figure 9. Bellatoiulus golovatchi gen. et sp. nov., female cybertype. A vulvae, caudal view B vulvae, frontal view C vulvae, ventral view D vulvae, dorsal view E right vulva, lateral view D right vulva, mesal view. Abbreviations: bu bursa mc median cleft mf median field op operculum. Images not to scale." figureDoi="10.3897/zookeys.1097.81792.figure9" httpUri="https://binary.pensoft.net/fig/674265" pageId="0" pageNumber="47">9</figureCitation>
, and
<emphasis id="FB9B5216344EB6B886EDEBF26811E9BC" bold="true" pageId="0" pageNumber="47">v</emphasis>
in Fig.
<figureCitation id="63AD532AE7DFA474A802F7E870CD0939" captionStart="Figure 8" captionStartId="F8" captionText="Figure 8. Bellatoiulus golovatchi gen. et sp. nov., female cybertype. A habitus, lateral view B anterior and posterior body parts, ventrolateral and dorso-lateral views, respectively C anterior and posterior body parts with highlighted vulvae, lateral views D anterior body part with highlighted vulvae, ventral view. Abbreviation: v vulvae. Images not to scale." figureDoi="10.3897/zookeys.1097.81792.figure8" httpUri="https://binary.pensoft.net/fig/674264" pageId="0" pageNumber="47">8C, D</figureCitation>
) stout, roughly cylindrical, slightly compressed on sides; bursa (
<emphasis id="7E60CD2203F9B8F691716ECBC7EA5923" bold="true" pageId="0" pageNumber="47">bu</emphasis>
) mostly symmetric, median cleft (
<emphasis id="E341BE5AB2ABB677E30FDED14C870C2E" bold="true" pageId="0" pageNumber="47">mc</emphasis>
) positioned on top within a rather deep and narrow median field (
<emphasis id="5355191E0007D69422B3A8FFEB8F0346" bold="true" pageId="0" pageNumber="47">mf</emphasis>
); operculum (
<emphasis id="5BB13580037878F3F27E3F543B41A531" bold="true" pageId="0" pageNumber="47">op</emphasis>
) higher than bursa, ending with two very long, tapering, hyaline protrusions (
<emphasis id="8189D66A09BEB7D00FB2B023C90149F8" bold="true" pageId="0" pageNumber="47">hp</emphasis>
); a row of four or five setae on each bursal valve and a group of several setae distally on each side of the operculum; bursal side sclerites non-setose. Receptaculum seminis composed of two nearly straight tubes: a shorter and somewhat thinner posterior one (
<emphasis id="323843E77F9CA2B04229EDB390117C62" bold="true" pageId="0" pageNumber="47">pt</emphasis>
), and an anterior one (
<emphasis id="C6E388CD947CE5BC8DD551727B2C7E48" bold="true" pageId="0" pageNumber="47">at</emphasis>
), slightly widening at bottom.
</paragraph>
<caption id="888E2BAD3990D573B1E10612CFA1A96C" doi="10.3897/zookeys.1097.81792.figure7" httpUri="https://binary.pensoft.net/fig/674263" pageId="0" pageNumber="47" start="Figure 7" startId="F7">
<paragraph id="68D0B63D6B9D329008CFB2506615AE76" pageId="0" pageNumber="47">
<emphasis id="E780466DDB09803F8496EDB24FC94034" bold="true" pageId="0" pageNumber="47">Figure 7.</emphasis>
<taxonomicName id="F93D3582D0091482905B86CABD46AFDC" authorityName="gen. et" authorityYear="2022" class="Diplopoda" family="Julidae" genus="Bellatoiulus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Bellatoiulus golovatchi" order="Julida" pageId="0" pageNumber="47" phylum="Arthropoda" rank="species" species="golovatchi">
<emphasis id="E87187D149577A80D13CBD18F0C3AB13" italics="true" pageId="0" pageNumber="47">Bellatoiulus golovatchi</emphasis>
</taxonomicName>
gen. et sp. nov., female paratypes (ZMUM)
<emphasis id="E68866D9B89C13213AA8D8DD3EC834A6" bold="true" pageId="0" pageNumber="47">A-C</emphasis>
and NMNHS
<emphasis id="EECE7D772FB778CDE88CB39A067902C5" bold="true" pageId="0" pageNumber="47">D</emphasis>
).
<emphasis id="84950AFE22AD0345C2CD68693E58BC9C" bold="true" pageId="0" pageNumber="47">A</emphasis>
right leg 2, caudal view
<emphasis id="5159DD15E3A769C019AFD99C544FD4CA" bold="true" pageId="0" pageNumber="47">B</emphasis>
left vulva, caudal view
<emphasis id="3F09B102EB3208970547CC5D9B6E3D8B" bold="true" pageId="0" pageNumber="47">C</emphasis>
right vulva, fronto-mesal, slightly apical view
<emphasis id="F7356EF34CB7993F8CAC7655A766E642" bold="true" pageId="0" pageNumber="47">D</emphasis>
right vulva, fronto-mesal view. Abbreviations:
<emphasis id="095555A5922C4250ED5067AC8ADF40E8" bold="true" pageId="0" pageNumber="47">at</emphasis>
anterior tube
<emphasis id="069E1C58F324E3D07EA40465E36875B7" bold="true" pageId="0" pageNumber="47">bu</emphasis>
bursa
<emphasis id="77241D4E92710A4B5753507688D495BE" bold="true" pageId="0" pageNumber="47">hp</emphasis>
hyaline protrusions
<emphasis id="893CEF3ACCED01756ABC2A1FC942CF33" bold="true" pageId="0" pageNumber="47">mc</emphasis>
median cleft
<emphasis id="C162705DAB32155F4E7F2E65562760B6" bold="true" pageId="0" pageNumber="47">mf</emphasis>
median field
<emphasis id="BE122A39374B2F7839336BAD6A18CAD8" bold="true" pageId="0" pageNumber="47">op</emphasis>
operculum
<emphasis id="58552110CEAAA2E114AAB6DB3A2C898C" bold="true" pageId="0" pageNumber="47">pt</emphasis>
posterior tube. Scale bars: 0.03 mm (
<emphasis id="070096A387E529DDAF53EF7C7929716B" bold="true" pageId="0" pageNumber="47">A</emphasis>
); 0.02 mm (
<emphasis id="6FCC03E06B10CC493192BFD78DEDA6E2" bold="true" pageId="0" pageNumber="47">B, C</emphasis>
); 0.1 mm (
<emphasis id="7EDE91718AB3A57CFD3A6DE721A707B6" bold="true" pageId="0" pageNumber="47">D</emphasis>
).
</paragraph>
</caption>
<caption id="8526A0548197BBB78670A526458701D4" doi="10.3897/zookeys.1097.81792.figure8" httpUri="https://binary.pensoft.net/fig/674264" pageId="0" pageNumber="47" start="Figure 8" startId="F8">
<paragraph id="325DA36502B045F7066722A6D5E4C979" pageId="0" pageNumber="47">
<emphasis id="88C2F8C6483F2222A2AFE755EF1EC7CE" bold="true" pageId="0" pageNumber="47">Figure 8.</emphasis>
<taxonomicName id="A23B519F346366E6D77D71E914D7EB1B" authorityName="gen. et" authorityYear="2022" class="Diplopoda" family="Julidae" genus="Bellatoiulus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Bellatoiulus golovatchi" order="Julida" pageId="0" pageNumber="47" phylum="Arthropoda" rank="species" species="golovatchi">
<emphasis id="CC3497BD7AA7F8D94253EA844A409F61" italics="true" pageId="0" pageNumber="47">Bellatoiulus golovatchi</emphasis>
</taxonomicName>
gen. et sp. nov., female cybertype.
<emphasis id="140A7F0EF56CE83CA42A244133E04C10" bold="true" pageId="0" pageNumber="47">A</emphasis>
habitus, lateral view
<emphasis id="5E2814BD7A3A188566535841BE6E0303" bold="true" pageId="0" pageNumber="47">B</emphasis>
anterior and posterior body parts, ventrolateral and dorso-lateral views, respectively
<emphasis id="804A5A41BAE3EBE9D17C1F270F69620B" bold="true" pageId="0" pageNumber="47">C</emphasis>
anterior and posterior body parts with highlighted vulvae, lateral views
<emphasis id="5E9E472969753B9EC3C82B81BC7D8449" bold="true" pageId="0" pageNumber="47">D</emphasis>
anterior body part with highlighted vulvae, ventral view. Abbreviation:
<emphasis id="02CF47BFE58FD88D77DB347E043D8FC2" bold="true" pageId="0" pageNumber="47">v</emphasis>
vulvae. Images not to scale.
</paragraph>
</caption>
<caption id="AA59A9FD27AA72F0C83785500F139242" doi="10.3897/zookeys.1097.81792.figure9" httpUri="https://binary.pensoft.net/fig/674265" pageId="0" pageNumber="47" start="Figure 9" startId="F9">
<paragraph id="868E27D7F5CEB5EAEB5A2A6781D0D79C" pageId="0" pageNumber="47">
<emphasis id="97D812B648909FDFC98F7EC56A83FDA6" bold="true" pageId="0" pageNumber="47">Figure 9.</emphasis>
<taxonomicName id="FC054F96DB61142E74A9DAC2AC0A2D0B" authorityName="gen. et" authorityYear="2022" class="Diplopoda" family="Julidae" genus="Bellatoiulus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Bellatoiulus golovatchi" order="Julida" pageId="0" pageNumber="47" phylum="Arthropoda" rank="species" species="golovatchi">
<emphasis id="3C2C73C43A2FD86C16468F8BB9AF2DE6" italics="true" pageId="0" pageNumber="47">Bellatoiulus golovatchi</emphasis>
</taxonomicName>
gen. et sp. nov., female cybertype.
<emphasis id="F8B84ADDDC448CB45B7019FBEB1F1DF3" bold="true" pageId="0" pageNumber="47">A</emphasis>
vulvae, caudal view
<emphasis id="3A4CB2A3066DBD2F369AA6FB3B3AAA15" bold="true" pageId="0" pageNumber="47">B</emphasis>
vulvae, frontal view
<emphasis id="0F227804191D5AEC1104A45FC7629ED5" bold="true" pageId="0" pageNumber="47">C</emphasis>
vulvae, ventral view
<emphasis id="03CA5D2C403A038656D231189CC42B71" bold="true" pageId="0" pageNumber="47">D</emphasis>
vulvae, dorsal view
<emphasis id="08B51FC610E985FFDAD64EFAC4EA7714" bold="true" pageId="0" pageNumber="47">E</emphasis>
right vulva, lateral view
<emphasis id="3843D3179B27B34F315DF68DCEB6D5FF" bold="true" pageId="0" pageNumber="47">D</emphasis>
right vulva, mesal view. Abbreviations:
<emphasis id="CA503B5751D3CAB5EA6B0F6DBD3F00AC" bold="true" pageId="0" pageNumber="47">bu</emphasis>
bursa
<emphasis id="A38034E742DBD8E28358110068F8D21F" bold="true" pageId="0" pageNumber="47">mc</emphasis>
median cleft
<emphasis id="BBC9166DC8A97F2448CFF79AC2D2D4C4" bold="true" pageId="0" pageNumber="47">mf</emphasis>
median field
<emphasis id="CCE428F335103E37AB99EA56F43BAF70" bold="true" pageId="0" pageNumber="47">op</emphasis>
operculum. Images not to scale.
</paragraph>
</caption>
</subSubSection>
</treatment>
</subSection>
</document>
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