treatments-xml/data/1E/8B/E7/1E8BE7195A7D5507812A131B5161B2AA.xml
2024-06-21 12:30:59 +02:00

287 lines
36 KiB
XML

<document ID-DOI="http://dx.doi.org/10.3897/zookeys.1103.84015" ID-Pensoft-Pub="1313-2970-1103-123" ID-Pensoft-UUID="A820088C785659F18D1C8E0F14FC642D" ID-ZooBank="58199DE7FAD743E1A029DC969C6E9BFC" ModsDocID="1313-2970-1103-123" checkinTime="1654168745574" checkinUser="pensoft" docAuthor="Qiu, Lu &amp; Kundrata, Robin" docDate="2022" docId="1E8BE7195A7D5507812A131B5161B2AA" docLanguage="en" docName="ZooKeys 1103: 123-138" docOrigin="ZooKeys 1103" docPubDate="2022-06-01" docSource="http://dx.doi.org/10.3897/zookeys.1103.84015" docTitle="Pseudocsikia choui Qiu &amp; Kundrata 2022, sp. nov." docType="treatment" docUuid="9AD84855-59B9-4C47-9734-FC6A2AAC723E" docUuidSource="ZooBank" docVersion="1" id="A820088C785659F18D1C8E0F14FC642D" lastPageNumber="123" masterDocId="A820088C785659F18D1C8E0F14FC642D" masterDocTitle="Descriptions of two new flightless species of Pseudocsikia Schimmel &amp; Platia (Coleoptera, Elateridae, Dimini) from Taiwan Island, China, with a definition of the formosana species-group" masterLastPageNumber="138" masterPageNumber="123" pageNumber="123" updateTime="1654168745574" updateUser="pensoft">
<mods:mods xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo>
<mods:title>Descriptions of two new flightless species of Pseudocsikia Schimmel &amp; Platia (Coleoptera, Elateridae, Dimini) from Taiwan Island, China, with a definition of the formosana species-group</mods:title>
</mods:titleInfo>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Qiu, Lu</mods:namePart>
<mods:nameIdentifier type="ORCID">https://orcid.org/0000-0002-0946-1634</mods:nameIdentifier>
<mods:affiliation>Engineering Research Center for Forest and Grassland Disaster Prevention and Reduction, Mianyang Normal University, Mianxing West Road, 621000, Mianyang, China</mods:affiliation>
<mods:nameIdentifier type="email">123church@163.com</mods:nameIdentifier>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Kundrata, Robin</mods:namePart>
<mods:nameIdentifier type="ORCID">https://orcid.org/0000-0001-9397-1030</mods:nameIdentifier>
<mods:affiliation>Department of Zoology, Faculty of Science, Palacky University, 17. listopadu 50, 77146, Olomouc, Czech Republic</mods:affiliation>
</mods:name>
<mods:typeOfResource>text</mods:typeOfResource>
<mods:relatedItem type="host">
<mods:titleInfo>
<mods:title>ZooKeys</mods:title>
</mods:titleInfo>
<mods:part>
<mods:date>2022</mods:date>
<mods:detail type="pubDate">
<mods:number>2022-06-01</mods:number>
</mods:detail>
<mods:detail type="volume">
<mods:number>1103</mods:number>
</mods:detail>
<mods:extent unit="page">
<mods:start>123</mods:start>
<mods:end>138</mods:end>
</mods:extent>
</mods:part>
</mods:relatedItem>
<mods:location>
<mods:url>http://dx.doi.org/10.3897/zookeys.1103.84015</mods:url>
</mods:location>
<mods:classification>journal article</mods:classification>
<mods:identifier type="DOI">http://dx.doi.org/10.3897/zookeys.1103.84015</mods:identifier>
<mods:identifier type="Pensoft-Pub">1313-2970-1103-123</mods:identifier>
<mods:identifier type="ZooBank">58199DE7FAD743E1A029DC969C6E9BFC</mods:identifier>
<mods:identifier type="Pensoft-UUID">A820088C785659F18D1C8E0F14FC642D</mods:identifier>
</mods:mods>
<treatment LSID="urn:lsid:zoobank.org:act:9AD84855-59B9-4C47-9734-FC6A2AAC723E" httpUri="http://treatment.plazi.org/id/1E8BE7195A7D5507812A131B5161B2AA" lastPageNumber="123" pageId="0" pageNumber="123">
<subSubSection pageId="0" pageNumber="123" type="nomenclature">
<paragraph pageId="0" pageNumber="123">
<taxonomicName LSID="http://zoobank.org/9AD84855-59B9-4C47-9734-FC6A2AAC723E" authority="Qiu &amp; Kundrata, 2022" authorityName="Qiu &amp; Kundrata" authorityYear="2022" class="Insecta" family="Elateridae" genus="Pseudocsikia" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Pseudocsikia choui" order="Coleoptera" pageId="0" pageNumber="123" phylum="Arthropoda" rank="species" species="choui" status="sp. nov.">Pseudocsikia choui</taxonomicName>
<taxonomicNameLabel pageId="0" pageNumber="123">sp. nov.</taxonomicNameLabel>
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="123" type="description">
<paragraph pageId="0" pageNumber="123">
<figureCitation captionStart="Figure 1" captionStartId="F1" captionText="Figure 1. Habitus of Pseudocsikia spp. A Pseudocsikia choui sp. nov., male holotype, dorsal view B Pseudocsikia chanjuan sp. nov., male holotype, dorsal view. Scale bar: 1.0 mm." figureDoi="10.3897/zookeys.1103.84015.figure1" httpUri="https://binary.pensoft.net/fig/691080" pageId="0" pageNumber="123">Figs 1A</figureCitation>
<figureCitation captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. Characters of Pseudocsikia choui sp. nov., male, holotype A pronorum, dorsal view B pro- and mesothorax, ventral view C anterior protrusion of pronotum, dorsal view D anterior protrusion of hypomeron, ventral view E antenna F posterior angle of pronotum, dorsal view G metacoxal plate, ventral view H prosternal process, lateral view. Scale bars: 0.5 mm (A, B, E, F); 0.2 mm (C, D, G, H)." figureDoi="10.3897/zookeys.1103.84015.figure3" httpUri="https://binary.pensoft.net/fig/691082" pageId="0" pageNumber="123">, 3A-H</figureCitation>
<figureCitation captionStart="Figure 5" captionStartId="F5" captionText="Figure 5. Characters of Pseudocsikia spp. A-F Pseudocsikia choui sp. nov., male holotype G-L Pseudocsikia chanjuan sp. nov., male, holotype A, G abdominal sternite IX, dorsal view B, H abdominal tergites IX-X, dorsal view C, I abdominal sternite VIII and tergite VIII, ventral view D, J aedeagus, ventral view E, K aedeagus, lateral view F, L aedeagus, dorsal view. Scale bar: 0.2 mm." figureDoi="10.3897/zookeys.1103.84015.figure5" httpUri="https://binary.pensoft.net/fig/691084" pageId="0" pageNumber="123">, 5A-F</figureCitation>
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="123" type="type material">
<paragraph pageId="0" pageNumber="123">Type material.</paragraph>
<paragraph pageId="0" pageNumber="123">
<emphasis bold="true" italics="true" pageId="0" pageNumber="123">Holotype</emphasis>
, male, &quot;2017.IX.13,台湾嘉义县阿里山二万坪, 2000m,
<normalizedToken originalValue="周文一”">周文-&quot;</normalizedToken>
[Erwanping, Mount Alishan, Chiayi County, Taiwan, 2000 m, 13.IX.2017, Wen-I Chou leg.], &quot;
<taxonomicName authorityName="Qiu &amp; Kundrata" authorityYear="2022" class="Insecta" family="Elateridae" genus="Pseudocsikia" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Pseudocsikia choui" order="Coleoptera" pageId="0" pageNumber="123" phylum="Arthropoda" rank="species" species="choui">
<emphasis italics="true" pageId="0" pageNumber="123">Pseudocsikia choui</emphasis>
</taxonomicName>
sp. nov.
<normalizedToken originalValue="周氏伪斯叩甲">周氏伪斯叩甲</normalizedToken>
HOLOTYPE des. Qiu et Kundrata 2022&quot; (MYNU).
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="123" type="diagnosis">
<paragraph pageId="0" pageNumber="123">Diagnosis.</paragraph>
<paragraph pageId="0" pageNumber="123">
Head, pronotum, and elytra dark brown, with paler lateral portions of pronotum and elytra, legs yellow (Fig.
<figureCitation captionStart="Figure 1" captionStartId="F1" captionText="Figure 1. Habitus of Pseudocsikia spp. A Pseudocsikia choui sp. nov., male holotype, dorsal view B Pseudocsikia chanjuan sp. nov., male holotype, dorsal view. Scale bar: 1.0 mm." figureDoi="10.3897/zookeys.1103.84015.figure1" httpUri="https://binary.pensoft.net/fig/691080" pageId="0" pageNumber="123">1A</figureCitation>
). Antennomere II subequal in length to antennomere III. Pronotum (Fig.
<figureCitation captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. Characters of Pseudocsikia choui sp. nov., male, holotype A pronorum, dorsal view B pro- and mesothorax, ventral view C anterior protrusion of pronotum, dorsal view D anterior protrusion of hypomeron, ventral view E antenna F posterior angle of pronotum, dorsal view G metacoxal plate, ventral view H prosternal process, lateral view. Scale bars: 0.5 mm (A, B, E, F); 0.2 mm (C, D, G, H)." figureDoi="10.3897/zookeys.1103.84015.figure3" httpUri="https://binary.pensoft.net/fig/691082" pageId="0" pageNumber="123">3A</figureCitation>
) smooth, with sparse punctures (intervals usually equal to 4-6 puncture diameters). Anterior angle of pronotum with apex of protrusion closer to inner angle. Posterior angles divergent. Metaventrite sparsely punctate, intervals between punctures on average subequal to 2-5 puncture diameters. Metacoxal plate (Fig.
<figureCitation captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. Characters of Pseudocsikia choui sp. nov., male, holotype A pronorum, dorsal view B pro- and mesothorax, ventral view C anterior protrusion of pronotum, dorsal view D anterior protrusion of hypomeron, ventral view E antenna F posterior angle of pronotum, dorsal view G metacoxal plate, ventral view H prosternal process, lateral view. Scale bars: 0.5 mm (A, B, E, F); 0.2 mm (C, D, G, H)." figureDoi="10.3897/zookeys.1103.84015.figure3" httpUri="https://binary.pensoft.net/fig/691082" pageId="0" pageNumber="123">3G</figureCitation>
) enlarged mesally, sparsely punctate. Tergite IX (Fig.
<figureCitation captionStart="Figure 5" captionStartId="F5" captionText="Figure 5. Characters of Pseudocsikia spp. A-F Pseudocsikia choui sp. nov., male holotype G-L Pseudocsikia chanjuan sp. nov., male, holotype A, G abdominal sternite IX, dorsal view B, H abdominal tergites IX-X, dorsal view C, I abdominal sternite VIII and tergite VIII, ventral view D, J aedeagus, ventral view E, K aedeagus, lateral view F, L aedeagus, dorsal view. Scale bar: 0.2 mm." figureDoi="10.3897/zookeys.1103.84015.figure5" httpUri="https://binary.pensoft.net/fig/691084" pageId="0" pageNumber="123">5B</figureCitation>
) subtriangular, with two narrow lobes. Aedeagus with median lobe with acute lateral projetions near midlength, narrowed to apex, apex blunt with small acute lateral projections. Paramere with apex pointed and projecting laterad. Phallobase with basal angles rounded (Fig.
<figureCitation captionStart="Figure 5" captionStartId="F5" captionText="Figure 5. Characters of Pseudocsikia spp. A-F Pseudocsikia choui sp. nov., male holotype G-L Pseudocsikia chanjuan sp. nov., male, holotype A, G abdominal sternite IX, dorsal view B, H abdominal tergites IX-X, dorsal view C, I abdominal sternite VIII and tergite VIII, ventral view D, J aedeagus, ventral view E, K aedeagus, lateral view F, L aedeagus, dorsal view. Scale bar: 0.2 mm." figureDoi="10.3897/zookeys.1103.84015.figure5" httpUri="https://binary.pensoft.net/fig/691084" pageId="0" pageNumber="123">5D</figureCitation>
).
</paragraph>
<caption doi="10.3897/zookeys.1103.84015.figure2" httpUri="https://binary.pensoft.net/fig/691081" pageId="0" pageNumber="123" start="Figure 2" startId="F2">
<paragraph pageId="0" pageNumber="123">
<emphasis bold="true" pageId="0" pageNumber="123">Figure 2.</emphasis>
Habitus and characters of
<taxonomicName baseAuthorityName="Ohira" baseAuthorityYear="1972" class="Insecta" family="Elateridae" genus="Pseudocsikia" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Pseudocsikia formosana" order="Coleoptera" pageId="0" pageNumber="123" phylum="Arthropoda" rank="species" species="formosana">
<emphasis italics="true" pageId="0" pageNumber="123">Pseudocsikia formosana</emphasis>
</taxonomicName>
(
<normalizedToken originalValue="Ôhira">Ohira</normalizedToken>
, 1972)
<emphasis bold="true" pageId="0" pageNumber="123">A-H</emphasis>
male holotype
<emphasis bold="true" pageId="0" pageNumber="123">A</emphasis>
habitus, dorsal view
<emphasis bold="true" pageId="0" pageNumber="123">B</emphasis>
habitus, ventral view
<emphasis bold="true" pageId="0" pageNumber="123">C</emphasis>
habitus, lateral view
<emphasis bold="true" pageId="0" pageNumber="123">D</emphasis>
labels
<emphasis bold="true" pageId="0" pageNumber="123">E</emphasis>
anterior protrusion of pronotum (indicated by an arrow), dorsal view
<emphasis bold="true" pageId="0" pageNumber="123">F</emphasis>
anterior protrusion of hypomeron (indicated by an arrow), ventral view
<emphasis bold="true" pageId="0" pageNumber="123">G</emphasis>
posterior angle of pronotum, dorsal view
<emphasis bold="true" pageId="0" pageNumber="123">H</emphasis>
abdominal tip, ventral view (arrow indicates the apex of median lobe)
<emphasis bold="true" pageId="0" pageNumber="123">I, J</emphasis>
paratype of an unknown sex
<emphasis bold="true" pageId="0" pageNumber="123">I</emphasis>
habitus, dorsal view
<emphasis bold="true" pageId="0" pageNumber="123">J</emphasis>
labels. Scale bars: 1.0 mm (
<emphasis bold="true" pageId="0" pageNumber="123">A-C, I</emphasis>
);
<emphasis bold="true" pageId="0" pageNumber="123">E-H</emphasis>
not to scale. All photos provided by Jeremy Frank (BPBM).
</paragraph>
</caption>
</subSubSection>
<subSubSection pageId="0" pageNumber="123" type="comparisons">
<paragraph pageId="0" pageNumber="123">Comparison.</paragraph>
<paragraph pageId="0" pageNumber="123">
This species superficially resembles
<taxonomicName class="Insecta" genus="P." kingdom="Animalia" lsidName="P. fomosana" order="Coleoptera" pageId="0" pageNumber="123" phylum="Arthropoda" rank="species" species="fomosana">
<emphasis italics="true" pageId="0" pageNumber="123">P. fomosana</emphasis>
</taxonomicName>
by the pale coloration of its pronotum and elytral sides and by the sparse punctures of pronotum, but it can be easily distinguished from the latter by the larger body length (5.9 mm, while 4.0 mm in
<taxonomicName class="Insecta" genus="P." kingdom="Animalia" lsidName="P. formosana" order="Coleoptera" pageId="0" pageNumber="123" phylum="Arthropoda" rank="species" species="formosana">
<emphasis italics="true" pageId="0" pageNumber="123">P. formosana</emphasis>
</taxonomicName>
), darker coloration of pronotum and elytra medially, more forwardly protruded anterior angles of pronotum (pointing more outward in the pronotum of
<taxonomicName class="Insecta" genus="P." kingdom="Animalia" lsidName="P. formosana" order="Coleoptera" pageId="0" pageNumber="123" phylum="Arthropoda" rank="species" species="formosana">
<emphasis italics="true" pageId="0" pageNumber="123">P. formosana</emphasis>
</taxonomicName>
), and shorter and more divergent posterior angle of pronotum (longer, more robust and nearly straight in
<taxonomicName class="Insecta" genus="P." kingdom="Animalia" lsidName="P. formosana" order="Coleoptera" pageId="0" pageNumber="123" phylum="Arthropoda" rank="species" species="formosana">
<emphasis italics="true" pageId="0" pageNumber="123">P. formosana</emphasis>
</taxonomicName>
). The shape of aedeagus also readily differentiates these two species. Based on the illustration of
<bibRefCitation author="Ohira, H" journalOrPublisher="Pacific Insects" pageId="0" pageNumber="123" pagination="1 - 14" refId="B8" refString="Ohira, H, 1972. Elaterid-beetles from Taiwan in Bishop Museum (Coleoptera). Pacific Insects 14: 1 - 14" title="Elaterid-beetles from Taiwan in Bishop Museum (Coleoptera)." volume="14" year="1972">
<normalizedToken originalValue="Ôhira">Ohira</normalizedToken>
(1972
</bibRefCitation>
: fig. 3), the distal half of the median lobe of
<taxonomicName class="Insecta" genus="P." kingdom="Animalia" lsidName="P. formosana" order="Coleoptera" pageId="0" pageNumber="123" phylum="Arthropoda" rank="species" species="formosana">
<emphasis italics="true" pageId="0" pageNumber="123">P. formosana</emphasis>
</taxonomicName>
has four large acute processes laterally, and its apex is somewhat rectangular; and the paramere of
<taxonomicName class="Insecta" genus="P." kingdom="Animalia" lsidName="P. formosana" order="Coleoptera" pageId="0" pageNumber="123" phylum="Arthropoda" rank="species" species="formosana">
<emphasis italics="true" pageId="0" pageNumber="123">P. formosana</emphasis>
</taxonomicName>
has the apex rounded and slightly outward.
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="123" type="description">
<paragraph pageId="0" pageNumber="123">Description</paragraph>
<paragraph pageId="0" pageNumber="123">
<emphasis bold="true" pageId="0" pageNumber="123">(male holotype).</emphasis>
Body smooth, surface covered with curved, semi-erect, and moderately long pubescence. Body length 5.9 mm; width 2.2 mm; antenna length 3.0 mm; pronotum length 1.4 mm, pronotum width 1.9 mm (measured at posterior angles), elytra length 3.5 mm.
</paragraph>
<paragraph pageId="0" pageNumber="123">
Body brown, pubescence yellow (Fig.
<figureCitation captionStart="Figure 1" captionStartId="F1" captionText="Figure 1. Habitus of Pseudocsikia spp. A Pseudocsikia choui sp. nov., male holotype, dorsal view B Pseudocsikia chanjuan sp. nov., male holotype, dorsal view. Scale bar: 1.0 mm." figureDoi="10.3897/zookeys.1103.84015.figure1" httpUri="https://binary.pensoft.net/fig/691080" pageId="0" pageNumber="123">1A</figureCitation>
). Head dark brown, antennae yellowish brown, labrum and mandibles brown, remaining mouthparts yellowish brown. Pronotum dark brown centrally; lateral, anterior and hind portions, and hind angles yellowish brown, with darker outlined margins. Scutellar shield brown, with dark outlined margins, especially anteriorly. Elytra dark brown centrally, yellowish brown laterally; yellowish-brown portions gradually lightened toward apices, basal margins of elytra dark outlined. Underside reddish brown, prosternum darker than hypomeron, sternites VI-VII and legs yellow.
</paragraph>
<paragraph pageId="0" pageNumber="123">
Head including eyes 0.5 times as wide as pronotum. Supra-antennal carinae short, directed mesad and fading medially so that median portion of frontoclypeus is not formed by sharp carina; frontoclypeus overhanging base of labrum in lateral view. Head surface sparsely punctate; punctures small, intervals between punctures mostly equal 2-3 puncture diameters. Maxillary palpus with palpomere III longer than wide. Antenna (Fig.
<figureCitation captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. Characters of Pseudocsikia choui sp. nov., male, holotype A pronorum, dorsal view B pro- and mesothorax, ventral view C anterior protrusion of pronotum, dorsal view D anterior protrusion of hypomeron, ventral view E antenna F posterior angle of pronotum, dorsal view G metacoxal plate, ventral view H prosternal process, lateral view. Scale bars: 0.5 mm (A, B, E, F); 0.2 mm (C, D, G, H)." figureDoi="10.3897/zookeys.1103.84015.figure3" httpUri="https://binary.pensoft.net/fig/691082" pageId="0" pageNumber="123">3E</figureCitation>
) surpassing hind angle of pronotum by about one antennomere; scape robust and longest, remaining antennomeres subequal in length; ultimate antennomere obliquely truncate, with apex rounded.
</paragraph>
<paragraph pageId="0" pageNumber="123">
Pronotum (Fig.
<figureCitation captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. Characters of Pseudocsikia choui sp. nov., male, holotype A pronorum, dorsal view B pro- and mesothorax, ventral view C anterior protrusion of pronotum, dorsal view D anterior protrusion of hypomeron, ventral view E antenna F posterior angle of pronotum, dorsal view G metacoxal plate, ventral view H prosternal process, lateral view. Scale bars: 0.5 mm (A, B, E, F); 0.2 mm (C, D, G, H)." figureDoi="10.3897/zookeys.1103.84015.figure3" httpUri="https://binary.pensoft.net/fig/691082" pageId="0" pageNumber="123">3A</figureCitation>
) wider than long (measured at midline), widest near middle. In lateral view, pronotum convex. Anterior angles of pronotum protruding (Fig.
<figureCitation captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. Characters of Pseudocsikia choui sp. nov., male, holotype A pronorum, dorsal view B pro- and mesothorax, ventral view C anterior protrusion of pronotum, dorsal view D anterior protrusion of hypomeron, ventral view E antenna F posterior angle of pronotum, dorsal view G metacoxal plate, ventral view H prosternal process, lateral view. Scale bars: 0.5 mm (A, B, E, F); 0.2 mm (C, D, G, H)." figureDoi="10.3897/zookeys.1103.84015.figure3" httpUri="https://binary.pensoft.net/fig/691082" pageId="0" pageNumber="123">3C</figureCitation>
); protrusion of anterior angle subquadrate, inner angle more protruded than outer angle, posterior part of protrusion with deep, crescent-shape pit. Lateral margins of pronotum arched medially, sides near middle narrowing anteriad and posteriad, anteriorly narrowing more sharply than posteriorly; posterior angles (Fig.
<figureCitation captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. Characters of Pseudocsikia choui sp. nov., male, holotype A pronorum, dorsal view B pro- and mesothorax, ventral view C anterior protrusion of pronotum, dorsal view D anterior protrusion of hypomeron, ventral view E antenna F posterior angle of pronotum, dorsal view G metacoxal plate, ventral view H prosternal process, lateral view. Scale bars: 0.5 mm (A, B, E, F); 0.2 mm (C, D, G, H)." figureDoi="10.3897/zookeys.1103.84015.figure3" httpUri="https://binary.pensoft.net/fig/691082" pageId="0" pageNumber="123">3F</figureCitation>
) long, slightly divergent, apical portion of posterior angle slightly enlarged, then narrowed, apex blunt. Disc of pronotum sparsely covered with small, shallow punctures; intervals between punctures on average subequal to four to six puncture diameters; interstices smooth. Pubescence mostly directed outwards; basal portion directed anteriorly.
</paragraph>
<paragraph pageId="0" pageNumber="123">
Hypomeron (Fig.
<figureCitation captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. Characters of Pseudocsikia choui sp. nov., male, holotype A pronorum, dorsal view B pro- and mesothorax, ventral view C anterior protrusion of pronotum, dorsal view D anterior protrusion of hypomeron, ventral view E antenna F posterior angle of pronotum, dorsal view G metacoxal plate, ventral view H prosternal process, lateral view. Scale bars: 0.5 mm (A, B, E, F); 0.2 mm (C, D, G, H)." figureDoi="10.3897/zookeys.1103.84015.figure3" httpUri="https://binary.pensoft.net/fig/691082" pageId="0" pageNumber="123">3B</figureCitation>
) more densely punctate than pronotum, punctures small and shallow, intervals between punctures on average subequal to 3-4 puncture diameters; apex of hypomeron strongly protruded, margin wrinkled. Pronotosternal sutures nearly straight, anterior excavation wide, long carination paralleled with suture from base of hypomeron and reaching anterior protrusion of hypomeron, forming hook-shaped carination anteriorly, end of the carination slightly extending backwards, with a small pit partly enclosed by curving hook of carination (Fig.
<figureCitation captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. Characters of Pseudocsikia choui sp. nov., male, holotype A pronorum, dorsal view B pro- and mesothorax, ventral view C anterior protrusion of pronotum, dorsal view D anterior protrusion of hypomeron, ventral view E antenna F posterior angle of pronotum, dorsal view G metacoxal plate, ventral view H prosternal process, lateral view. Scale bars: 0.5 mm (A, B, E, F); 0.2 mm (C, D, G, H)." figureDoi="10.3897/zookeys.1103.84015.figure3" httpUri="https://binary.pensoft.net/fig/691082" pageId="0" pageNumber="123">3D</figureCitation>
). Prosternum (Fig.
<figureCitation captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. Characters of Pseudocsikia choui sp. nov., male, holotype A pronorum, dorsal view B pro- and mesothorax, ventral view C anterior protrusion of pronotum, dorsal view D anterior protrusion of hypomeron, ventral view E antenna F posterior angle of pronotum, dorsal view G metacoxal plate, ventral view H prosternal process, lateral view. Scale bars: 0.5 mm (A, B, E, F); 0.2 mm (C, D, G, H)." figureDoi="10.3897/zookeys.1103.84015.figure3" httpUri="https://binary.pensoft.net/fig/691082" pageId="0" pageNumber="123">3B</figureCitation>
) including prosternal process about 2.00 times as long as wide; chin piece with large, dense punctures, intervals between punctures approximately one puncture diameter; punctures in remaining area sparser and smaller, intervals between punctures 3-6 puncture diameters, punctures on prosternal process sparse, small. Prosternal process (Fig.
<figureCitation captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. Characters of Pseudocsikia choui sp. nov., male, holotype A pronorum, dorsal view B pro- and mesothorax, ventral view C anterior protrusion of pronotum, dorsal view D anterior protrusion of hypomeron, ventral view E antenna F posterior angle of pronotum, dorsal view G metacoxal plate, ventral view H prosternal process, lateral view. Scale bars: 0.5 mm (A, B, E, F); 0.2 mm (C, D, G, H)." figureDoi="10.3897/zookeys.1103.84015.figure3" httpUri="https://binary.pensoft.net/fig/691082" pageId="0" pageNumber="123">3H</figureCitation>
) with ventral surface horizontal in lateral view, with elongate notch ventroapically, roundly enlarged dorsoapically.
</paragraph>
<caption doi="10.3897/zookeys.1103.84015.figure3" httpUri="https://binary.pensoft.net/fig/691082" pageId="0" pageNumber="123" start="Figure 3" startId="F3">
<paragraph pageId="0" pageNumber="123">
<emphasis bold="true" pageId="0" pageNumber="123">Figure 3.</emphasis>
Characters of
<taxonomicName authorityName="Qiu &amp; Kundrata" authorityYear="2022" class="Insecta" family="Elateridae" genus="Pseudocsikia" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Pseudocsikia choui" order="Coleoptera" pageId="0" pageNumber="123" phylum="Arthropoda" rank="species" species="choui">
<emphasis italics="true" pageId="0" pageNumber="123">Pseudocsikia choui</emphasis>
</taxonomicName>
sp. nov., male, holotype
<emphasis bold="true" pageId="0" pageNumber="123">A</emphasis>
pronorum, dorsal view
<emphasis bold="true" pageId="0" pageNumber="123">B</emphasis>
pro- and mesothorax, ventral view
<emphasis bold="true" pageId="0" pageNumber="123">C</emphasis>
anterior protrusion of pronotum, dorsal view
<emphasis bold="true" pageId="0" pageNumber="123">D</emphasis>
anterior protrusion of hypomeron, ventral view
<emphasis bold="true" pageId="0" pageNumber="123">E</emphasis>
antenna
<emphasis bold="true" pageId="0" pageNumber="123">F</emphasis>
posterior angle of pronotum, dorsal view
<emphasis bold="true" pageId="0" pageNumber="123">G</emphasis>
metacoxal plate, ventral view
<emphasis bold="true" pageId="0" pageNumber="123">H</emphasis>
prosternal process, lateral view. Scale bars: 0.5 mm (
<emphasis bold="true" pageId="0" pageNumber="123">A, B, E, F</emphasis>
); 0.2 mm (
<emphasis bold="true" pageId="0" pageNumber="123">C, D, G, H</emphasis>
).
</paragraph>
</caption>
<paragraph pageId="0" pageNumber="123">
Scutellar shield (Fig.
<figureCitation captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. Characters of Pseudocsikia choui sp. nov., male, holotype A pronorum, dorsal view B pro- and mesothorax, ventral view C anterior protrusion of pronotum, dorsal view D anterior protrusion of hypomeron, ventral view E antenna F posterior angle of pronotum, dorsal view G metacoxal plate, ventral view H prosternal process, lateral view. Scale bars: 0.5 mm (A, B, E, F); 0.2 mm (C, D, G, H)." figureDoi="10.3897/zookeys.1103.84015.figure3" httpUri="https://binary.pensoft.net/fig/691082" pageId="0" pageNumber="123">3F</figureCitation>
) suboval, about 1.2 times as wide as long; anterior margin rounded, posterior margin slightly pointed. Mesoventrite (Fig.
<figureCitation captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. Characters of Pseudocsikia choui sp. nov., male, holotype A pronorum, dorsal view B pro- and mesothorax, ventral view C anterior protrusion of pronotum, dorsal view D anterior protrusion of hypomeron, ventral view E antenna F posterior angle of pronotum, dorsal view G metacoxal plate, ventral view H prosternal process, lateral view. Scale bars: 0.5 mm (A, B, E, F); 0.2 mm (C, D, G, H)." figureDoi="10.3897/zookeys.1103.84015.figure3" httpUri="https://binary.pensoft.net/fig/691082" pageId="0" pageNumber="123">3B</figureCitation>
) with deep procoxal rests. Mesoventral process elevated, hind margin wide. Mesanepisternum with large, curved lateral extensions of procoxal rests. Metaventrite medially with sparse punctures, intervals between punctures on average subequal to 3-5 puncture diameters. Anterior portion of discrimen with sharp groove, occupying half-length of metaventrite. Metacoxal plate enlarged inward, narrowed laterad (Fig.
<figureCitation captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. Characters of Pseudocsikia choui sp. nov., male, holotype A pronorum, dorsal view B pro- and mesothorax, ventral view C anterior protrusion of pronotum, dorsal view D anterior protrusion of hypomeron, ventral view E antenna F posterior angle of pronotum, dorsal view G metacoxal plate, ventral view H prosternal process, lateral view. Scale bars: 0.5 mm (A, B, E, F); 0.2 mm (C, D, G, H)." figureDoi="10.3897/zookeys.1103.84015.figure3" httpUri="https://binary.pensoft.net/fig/691082" pageId="0" pageNumber="123">3G</figureCitation>
), surface with very sparse punctures.
</paragraph>
<paragraph pageId="0" pageNumber="123">
Elytra (Fig.
<figureCitation captionStart="Figure 1" captionStartId="F1" captionText="Figure 1. Habitus of Pseudocsikia spp. A Pseudocsikia choui sp. nov., male holotype, dorsal view B Pseudocsikia chanjuan sp. nov., male holotype, dorsal view. Scale bar: 1.0 mm." figureDoi="10.3897/zookeys.1103.84015.figure1" httpUri="https://binary.pensoft.net/fig/691080" pageId="0" pageNumber="123">1A</figureCitation>
) elongate, together 1.6 times as long as wide, widest at 1/3 of their length from base. Humeri (Fig.
<figureCitation captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. Characters of Pseudocsikia choui sp. nov., male, holotype A pronorum, dorsal view B pro- and mesothorax, ventral view C anterior protrusion of pronotum, dorsal view D anterior protrusion of hypomeron, ventral view E antenna F posterior angle of pronotum, dorsal view G metacoxal plate, ventral view H prosternal process, lateral view. Scale bars: 0.5 mm (A, B, E, F); 0.2 mm (C, D, G, H)." figureDoi="10.3897/zookeys.1103.84015.figure3" httpUri="https://binary.pensoft.net/fig/691082" pageId="0" pageNumber="123">3A, F</figureCitation>
) elevated. Sides from humeri roundly widened to 1/3 of elytral length in dorsal view, then gradually narrowed towards apices; apices slightly independently rounded. Elytral striae shallow, formed by lines of small punctures, intervals between punctures in stria on average subequal to 2-3 puncture diameters. Interstriae flat, smooth, with some micropunctures. Hind wings absent. Abdomen with ventrites more densely punctate than pronotum, intervals between punctures on average subequal to 2-3 puncture diameters; pubescence directed backwards. Apical ventrite with blunt apex. Tergite VIII (Fig.
<figureCitation captionStart="Figure 5" captionStartId="F5" captionText="Figure 5. Characters of Pseudocsikia spp. A-F Pseudocsikia choui sp. nov., male holotype G-L Pseudocsikia chanjuan sp. nov., male, holotype A, G abdominal sternite IX, dorsal view B, H abdominal tergites IX-X, dorsal view C, I abdominal sternite VIII and tergite VIII, ventral view D, J aedeagus, ventral view E, K aedeagus, lateral view F, L aedeagus, dorsal view. Scale bar: 0.2 mm." figureDoi="10.3897/zookeys.1103.84015.figure5" httpUri="https://binary.pensoft.net/fig/691084" pageId="0" pageNumber="123">5C</figureCitation>
) subtriangular, 1.6 times as long as wide, distal margin pointed medially, apically covered with sparse pubescence. Sternite VIII with two dark colored lobes, shape as Fig.
<figureCitation captionStart="Figure 5" captionStartId="F5" captionText="Figure 5. Characters of Pseudocsikia spp. A-F Pseudocsikia choui sp. nov., male holotype G-L Pseudocsikia chanjuan sp. nov., male, holotype A, G abdominal sternite IX, dorsal view B, H abdominal tergites IX-X, dorsal view C, I abdominal sternite VIII and tergite VIII, ventral view D, J aedeagus, ventral view E, K aedeagus, lateral view F, L aedeagus, dorsal view. Scale bar: 0.2 mm." figureDoi="10.3897/zookeys.1103.84015.figure5" httpUri="https://binary.pensoft.net/fig/691084" pageId="0" pageNumber="123">5C</figureCitation>
, with long setae, remaining portion membranous. Tergite IX (Fig.
<figureCitation captionStart="Figure 5" captionStartId="F5" captionText="Figure 5. Characters of Pseudocsikia spp. A-F Pseudocsikia choui sp. nov., male holotype G-L Pseudocsikia chanjuan sp. nov., male, holotype A, G abdominal sternite IX, dorsal view B, H abdominal tergites IX-X, dorsal view C, I abdominal sternite VIII and tergite VIII, ventral view D, J aedeagus, ventral view E, K aedeagus, lateral view F, L aedeagus, dorsal view. Scale bar: 0.2 mm." figureDoi="10.3897/zookeys.1103.84015.figure5" httpUri="https://binary.pensoft.net/fig/691084" pageId="0" pageNumber="123">5B</figureCitation>
) subtriangular, 1.3 times as long as wide, medially deeply emarginate; two lobes elongate, lateral sides with long setae; tergite X (Fig.
<figureCitation captionStart="Figure 5" captionStartId="F5" captionText="Figure 5. Characters of Pseudocsikia spp. A-F Pseudocsikia choui sp. nov., male holotype G-L Pseudocsikia chanjuan sp. nov., male, holotype A, G abdominal sternite IX, dorsal view B, H abdominal tergites IX-X, dorsal view C, I abdominal sternite VIII and tergite VIII, ventral view D, J aedeagus, ventral view E, K aedeagus, lateral view F, L aedeagus, dorsal view. Scale bar: 0.2 mm." figureDoi="10.3897/zookeys.1103.84015.figure5" httpUri="https://binary.pensoft.net/fig/691084" pageId="0" pageNumber="123">5B</figureCitation>
) membranous, exceeding apices of lobes of tergite IX. Sternite IX (Fig.
<figureCitation captionStart="Figure 5" captionStartId="F5" captionText="Figure 5. Characters of Pseudocsikia spp. A-F Pseudocsikia choui sp. nov., male holotype G-L Pseudocsikia chanjuan sp. nov., male, holotype A, G abdominal sternite IX, dorsal view B, H abdominal tergites IX-X, dorsal view C, I abdominal sternite VIII and tergite VIII, ventral view D, J aedeagus, ventral view E, K aedeagus, lateral view F, L aedeagus, dorsal view. Scale bar: 0.2 mm." figureDoi="10.3897/zookeys.1103.84015.figure5" httpUri="https://binary.pensoft.net/fig/691084" pageId="0" pageNumber="123">5A</figureCitation>
) slightly stout, 2.7 times as long as wide, apically widely rounded and setose.
</paragraph>
<paragraph pageId="0" pageNumber="123">
<emphasis bold="true" italics="true" pageId="0" pageNumber="123">Aedeagus</emphasis>
(Fig.
<figureCitation captionStart="Figure 5" captionStartId="F5" captionText="Figure 5. Characters of Pseudocsikia spp. A-F Pseudocsikia choui sp. nov., male holotype G-L Pseudocsikia chanjuan sp. nov., male, holotype A, G abdominal sternite IX, dorsal view B, H abdominal tergites IX-X, dorsal view C, I abdominal sternite VIII and tergite VIII, ventral view D, J aedeagus, ventral view E, K aedeagus, lateral view F, L aedeagus, dorsal view. Scale bar: 0.2 mm." figureDoi="10.3897/zookeys.1103.84015.figure5" httpUri="https://binary.pensoft.net/fig/691084" pageId="0" pageNumber="123">5D-F</figureCitation>
) with robust median lobe, two times as long as one paramere; distal half of median lobe arrow-shaped, apex with small protrusion, apex blunt, laterally with small acute projections; long, needle-like sclerite present on ventral side of median lobe. Paramere stout, reaching half of median lobe; apex pointed outward. Phallobase subquadrate, margins thickened, medially with longitudinal thickened line, basal angles rounded.
</paragraph>
<paragraph pageId="0" pageNumber="123">
<emphasis bold="true" pageId="0" pageNumber="123">Female.</emphasis>
Unknown.
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="123" type="immature stages">
<paragraph pageId="0" pageNumber="123">Immature stages.</paragraph>
<paragraph pageId="0" pageNumber="123">Unknown.</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="123" type="distribution">
<paragraph pageId="0" pageNumber="123">Distribution.</paragraph>
<paragraph pageId="0" pageNumber="123">China: Taiwan (Chiayi).</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="123" type="etymology">
<paragraph pageId="0" pageNumber="123">Etymology.</paragraph>
<paragraph pageId="0" pageNumber="123">The specific patronymic epithet is dedicated to Dr Wen-I Chou (Taiwan, China), the collector of the holotype.</paragraph>
</subSubSection>
</treatment>
</document>