404 lines
56 KiB
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404 lines
56 KiB
XML
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<mods:title id="B07D8E19AB7A734A570B02EA5B5B82C5">Historical perspective, new contributions and an enlightening dispersal mechanism for the endogean genus Typhlocharis Dieck 1869 (Coleoptera: Carabidae: Trechinae)</mods:title>
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<mods:namePart id="F8A18B4A9B505AC8AB34A7744BAD0991">Gilgado, José D.</mods:namePart>
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<mods:title id="4A48EFC2AF1522EC399C3C08333DC3F1">Journal of Natural History</mods:title>
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<mods:date id="E513F31E1B4D842D657AEF81CA00176A">2011</mods:date>
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<emphasis id="649CAE1877534542FE574E52DB8B3A47" bold="true" box="[405,654,823,847]" italics="true" pageId="8" pageNumber="1239">Typhlocharis josabelae</emphasis>
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Ortuño and Gilgado (
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<figureCitation id="CED36E8F77534542FDA54E32DBE93A67" box="[615,748,855,879]" captionStart-0="Figure 2" captionStart-1="Figure 3" captionStart-2="Figure 4" captionStartId-0="9.[144,211,1654,1676]" captionStartId-1="10.[213,280,1238,1260]" captionStartId-2="11.[144,211,1393,1415]" captionTargetBox-0="[145,1099,164,1619]" captionTargetBox-1="[211,1171,160,1206]" captionTargetBox-2="[142,1102,159,1360]" captionTargetId-0="figure-6@9.[142,1102,161,1622]" captionTargetId-1="figure-5@10.[211,1171,160,1206]" captionTargetId-2="figure-6@11.[142,1102,159,1360]" captionTargetPageId-0="9" captionTargetPageId-1="10" captionTargetPageId-2="11" captionText-0="Figure 2. Habitus of Typhlocharis josabelae sp. nov. Scale bar: 0.3 mm." captionText-1="Figure 3. Morphological details of Typhlocharis josabelae sp. nov. (A) Labium and prebasilar; (B) posterior angle of pronotum; (C) left protibia and protarsum in ventral view; (D) last abdominal sternite of male; (E) last abdominal sternite of female. Scale bars: A–C, 0.05 mm; D,E, 0.1 mm." captionText-2="Figure 4. Details of the genitalia of Typhlocharis josabelae sp. nov. (A) Median lobe of aedeagus in left lateral view; (B) median lobe and right paramere in dorsal view; (C) left paramere in left lateral view; (D) right paramere in right lateral view; (E) apodemal ring; (F) right gonocoxite in ventral view of paratype 1; (G) right gonocoxite in ventral view of paratype 2; (H) spermatheca, glandule and conduct. Scale bars: A–D, F–H, 0.05 mm; E, 0.1 mm." figureDoi-0="http://doi.org/10.5281/zenodo.5203657" figureDoi-1="http://doi.org/10.5281/zenodo.5203659" figureDoi-2="http://doi.org/10.5281/zenodo.5203661" httpUri-0="https://zenodo.org/record/5203657/files/figure.png" httpUri-1="https://zenodo.org/record/5203659/files/figure.png" httpUri-2="https://zenodo.org/record/5203661/files/figure.png" pageId="8" pageNumber="1239">Figures 2–4</figureCitation>
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)
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(
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and
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leg.
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One
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, Idem,
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and
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leg.
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(
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;
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.M.
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and
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leg. The
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series is deposited in the collection of
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|
||
<collectionCode id="30F9EACF77534542FCDF49A4DA363DD0" box="[797,819,1217,1240]" country="Canada" lsid="urn:lsid:biocol.org:col:13946" name="Royal British Columbia Museum - Herbarium" pageId="8" pageNumber="1239" type="Museum">V</collectionCode>
|
||
.
|
||
<collectorName id="FB1D17DC77534542FCF449A4DAB53DD0" box="[822,944,1216,1240]" pageId="8" pageNumber="1239">M. Ortuño</collectorName>
|
||
in the
|
||
<location id="533724D177534542FC3D49A4D8573DF0" LSID="urn:lsid:plazi:treatment:DE41C31C77534547FE574E52D8593A0C:533724D177534542FC3D49A4D8573DF0" country="Spain" county="Alcala University" municipality="Physical Anthropology" name="Deptartment of Zoology" pageId="8" pageNumber="1239">Deptartment of Zoology</location>
|
||
and
|
||
<collectingMunicipality id="B633E87077534542FE484985DB8B3DF0" box="[394,654,1248,1272]" pageId="8" pageNumber="1239">Physical Anthropology</collectingMunicipality>
|
||
,
|
||
<collectingCounty id="BF360A8677534542FD5A4985DA5A3DF0" box="[664,863,1248,1272]" pageId="8" pageNumber="1239">Alcalá University</collectingCounty>
|
||
,
|
||
<collectingCountry id="2EFF329A77534542FCAB4985DAA83DF0" box="[873,941,1248,1272]" name="Spain" pageId="8" pageNumber="1239">Spain</collectingCountry>
|
||
</materialsCitation>
|
||
.
|
||
</paragraph>
|
||
<paragraph id="5657720A77534542FF17485ED8443C5A" blockId="8.[213,321,1339,1362]" box="[213,321,1339,1362]" pageId="8" pageNumber="1239">
|
||
<emphasis id="649CAE1877534542FF17485ED8443C5A" box="[213,321,1339,1362]" italics="true" pageId="8" pageNumber="1239">Diagnosis</emphasis>
|
||
</paragraph>
|
||
<paragraph id="5657720A77534542FF174804D8623F90" blockId="8.[213,1170,1377,1688]" pageId="8" pageNumber="1239">
|
||
Body long, parallel and flattened. Length:
|
||
<quantity id="9110DFEF77534542FD734804DA4A3C71" box="[689,847,1377,1401]" metricMagnitude="-3" metricUnit="m" metricValue="1.3" metricValueMax="1.33" metricValueMin="1.27" pageId="8" pageNumber="1239" unit="mm" value="1.3" valueMax="1.33" valueMin="1.27">1.27–1.33 mm</quantity>
|
||
for males and
|
||
<quantity id="9110DFEF77534542FC364804DD973C71" box="[1012,1170,1377,1401]" metricMagnitude="-3" metricUnit="m" metricValue="1.285" metricValueMax="1.3" metricValueMin="1.27" pageId="8" pageNumber="1239" unit="mm" value="1.285" valueMax="1.3" valueMin="1.27">1.27–1.30 mm</quantity>
|
||
for females. Anophthalmous. Integument brownish-yellow, microreticulated, with scattered pubescence. Elytron with 7 (4+3) marginal umbilicate setae. Apical edge slightly sinuate without conspicuous teeth (males and females). Trochanters, femora and tibiae inermous in both sexes. Abdominal sternum II without special characters. Aedeagus with median lobe sickle-shaped, and basal lamina arcuate; internal sac with one small sclerite; parameres with two setae, left paramere subtriangular and right paramere narrow, unusually long and slightly sclerotized. Female genitalia with gonocoxite unguiform: a trait present in the
|
||
<taxonomicName id="91E8098977534542FD224B07DA753F71" authorityName="Zaballos and Wrase" authorityYear="1998" box="[736,880,1634,1657]" class="Insecta" family="Carabidae" genus="Typhlocharis" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="1239" phylum="Arthropoda" rank="species" species="monastica">
|
||
<emphasis id="649CAE1877534542FD224B07DA753F71" box="[736,880,1634,1657]" italics="true" pageId="8" pageNumber="1239">T. monastica</emphasis>
|
||
</taxonomicName>
|
||
group (see
|
||
<bibRefCitation id="32790FFB77534542FC3E4B05D8593F90" author="Zaballos JP & Wrase DW" pageId="8" pageNumber="1239" pagination="43 - 52" refId="ref10972" refString="Zaballos JP, Wrase DW. 1998. Tres nuevos Typhlocharis Dieck, 1869 (Coleoptera, Caraboidea, Trechidae) de Navarra (Espana). Graellsia 54: 43 - 52." type="journal article" year="1998">Zaballos and Wrase 1998</bibRefCitation>
|
||
).
|
||
</paragraph>
|
||
<caption id="0297228277524543FF524B13DA583F84" ID-DOI="http://doi.org/10.5281/zenodo.5203657" ID-Zenodo-Dep="5203657" box="[144,861,1654,1677]" httpUri="https://zenodo.org/record/5203657/files/figure.png" pageId="9" pageNumber="1240" startId="9.[144,211,1654,1676]" targetBox="[145,1099,164,1619]" targetPageId="9">
|
||
<paragraph id="5657720A77524543FF524B13DA583F84" blockId="9.[144,861,1654,1677]" box="[144,861,1654,1677]" pageId="9" pageNumber="1240">
|
||
Figure 2. Habitus of
|
||
<taxonomicName id="91E8098977524543FEAB4B12DB4C3F85" authority="Ortuño & Gilgado, 2011" authorityName="Ortuño & Gilgado" authorityYear="2011" box="[361,585,1655,1677]" class="Insecta" family="Carabidae" genus="Typhlocharis" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="1240" phylum="Arthropoda" rank="species" species="josabelae" status="sp. nov.">
|
||
<emphasis id="649CAE1877524543FEAB4B12DB4C3F85" box="[361,585,1655,1677]" italics="true" pageId="9" pageNumber="1240">Typhlocharis josabelae</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="7FAF136377524543FD8D4B12DB993F84" box="[591,668,1655,1676]" pageId="9" pageNumber="1240" rank="species">sp. nov.</taxonomicNameLabel>
|
||
Scale bar: 0.3 mm.
|
||
</paragraph>
|
||
</caption>
|
||
<caption id="0297228277514540FF1749B3D85B3C4C" ID-DOI="http://doi.org/10.5281/zenodo.5203659" ID-Zenodo-Dep="5203659" httpUri="https://zenodo.org/record/5203659/files/figure.png" pageId="10" pageNumber="1241" startId="10.[213,280,1238,1260]" targetBox="[211,1171,160,1206]" targetPageId="10">
|
||
<paragraph id="5657720A77514540FF1749B3D85B3C4C" blockId="10.[213,1170,1238,1348]" pageId="10" pageNumber="1241">
|
||
Figure 3. Morphological details of
|
||
<taxonomicName id="91E8098977514540FD8849B2DA283DE5" authority="Ortuño & Gilgado, 2011" authorityName="Ortuño & Gilgado" authorityYear="2011" box="[586,813,1239,1261]" class="Insecta" family="Carabidae" genus="Typhlocharis" kingdom="Animalia" order="Coleoptera" pageId="10" pageNumber="1241" phylum="Arthropoda" rank="species" species="josabelae" status="sp. nov.">
|
||
<emphasis id="649CAE1877514540FD8849B2DA283DE5" box="[586,813,1239,1261]" italics="true" pageId="10" pageNumber="1241">Typhlocharis josabelae</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="7FAF136377514540FCF449B2DA803DE4" box="[822,901,1239,1260]" pageId="10" pageNumber="1241" rank="species">sp. nov.</taxonomicNameLabel>
|
||
(A) Labium and prebasilar; (B) posterior angle of pronotum; (C) left protibia and protarsum in ventral view; (D) last abdominal sternite of male; (E) last abdominal sternite of female. Scale bars: A–C, 0.05 mm; D,E, 0.1 mm.
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="5657720A77514540FF174809D8513C8B" blockId="10.[213,340,1388,1411]" box="[213,340,1388,1411]" pageId="10" pageNumber="1241">
|
||
<emphasis id="649CAE1877514540FF174809D8513C8B" box="[213,340,1388,1411]" italics="true" pageId="10" pageNumber="1241">Description</emphasis>
|
||
</paragraph>
|
||
<paragraph id="5657720A77514540FF1748F6D83C3FA2" blockId="10.[213,1170,1427,1706]" pageId="10" pageNumber="1241">
|
||
Head lenticular, slightly narrower (
|
||
<quantity id="9110DFEF77514540FD9E48F6DBF83CA3" box="[604,765,1427,1451]" metricMagnitude="-4" metricUnit="m" metricValue="2.65" metricValueMax="2.7" metricValueMin="2.6" pageId="10" pageNumber="1241" unit="mm" value="0.265" valueMax="0.27" valueMin="0.26">0.26–0.27 mm</quantity>
|
||
) than pronotum (
|
||
<figureCitation id="CED36E8F77514540FC0448F6DD2C3CA3" box="[966,1065,1427,1451]" captionStart="Figure 2" captionStartId="9.[144,211,1654,1676]" captionTargetBox="[145,1099,164,1619]" captionTargetId="figure-6@9.[142,1102,161,1622]" captionTargetPageId="9" captionText="Figure 2. Habitus of Typhlocharis josabelae sp. nov. Scale bar: 0.3 mm." figureDoi="http://doi.org/10.5281/zenodo.5203657" httpUri="https://zenodo.org/record/5203657/files/figure.png" pageId="10" pageNumber="1241">Figure 2</figureCitation>
|
||
). Frontal sulci divergent. Microsculpture of cephalic disc polygonal, mostly isodiametric except in a small central area where microsculpture is lessened (no striated surface is observed). Antennae moniliform (
|
||
<figureCitation id="CED36E8F77514540FD944897DBBF3F02" box="[598,698,1522,1546]" captionStart="Figure 2" captionStartId="9.[144,211,1654,1676]" captionTargetBox="[145,1099,164,1619]" captionTargetId="figure-6@9.[142,1102,161,1622]" captionTargetPageId="9" captionText="Figure 2. Habitus of Typhlocharis josabelae sp. nov. Scale bar: 0.3 mm." figureDoi="http://doi.org/10.5281/zenodo.5203657" httpUri="https://zenodo.org/record/5203657/files/figure.png" pageId="10" pageNumber="1241">Figure 2</figureCitation>
|
||
), with noticeably setulated antennomeres. Labrum rectangular. Mandibles sharpened. Labium transverse, with a notch that is armed with a conspicuous tooth (blunt) that does not exceed the anterior end of the epilobe (
|
||
<figureCitation id="CED36E8F77514540FE534B37DB0A3F62" box="[401,527,1618,1642]" captionStart="Figure 3" captionStartId="10.[213,280,1238,1260]" captionTargetBox="[211,1171,160,1206]" captionTargetId="figure-5@10.[211,1171,160,1206]" captionTargetPageId="10" captionText="Figure 3. Morphological details of Typhlocharis josabelae sp. nov. (A) Labium and prebasilar; (B) posterior angle of pronotum; (C) left protibia and protarsum in ventral view; (D) last abdominal sternite of male; (E) last abdominal sternite of female. Scale bars: A–C, 0.05 mm; D,E, 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.5203659" httpUri="https://zenodo.org/record/5203659/files/figure.png" pageId="10" pageNumber="1241">Figure 3A</figureCitation>
|
||
). Prebasilar is separated from labium by a transverse suture (
|
||
<figureCitation id="CED36E8F77514540FEEC4B17D8A33F82" box="[302,422,1650,1674]" captionStart="Figure 3" captionStartId="10.[213,280,1238,1260]" captionTargetBox="[211,1171,160,1206]" captionTargetId="figure-5@10.[211,1171,160,1206]" captionTargetPageId="10" captionText="Figure 3. Morphological details of Typhlocharis josabelae sp. nov. (A) Labium and prebasilar; (B) posterior angle of pronotum; (C) left protibia and protarsum in ventral view; (D) last abdominal sternite of male; (E) last abdominal sternite of female. Scale bars: A–C, 0.05 mm; D,E, 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.5203659" httpUri="https://zenodo.org/record/5203659/files/figure.png" pageId="10" pageNumber="1241">Figure 3B</figureCitation>
|
||
). Labial and maxillary palps have typical shape and chaetotaxy of genus.
|
||
</paragraph>
|
||
<caption id="0297228277504541FF524814DA323CF4" ID-DOI="http://doi.org/10.5281/zenodo.5203661" ID-Zenodo-Dep="5203661" httpUri="https://zenodo.org/record/5203661/files/figure.png" pageId="11" pageNumber="1242" startId="11.[144,211,1393,1415]" targetBox="[142,1102,159,1360]" targetPageId="11">
|
||
<paragraph id="5657720A77504541FF524814DA323CF4" blockId="11.[144,1101,1393,1532]" pageId="11" pageNumber="1242">
|
||
Figure 4. Details of the genitalia of
|
||
<taxonomicName id="91E8098977504541FE3A4814DBD23C8F" authority="Ortuño & Gilgado, 2011" authorityName="Ortuño & Gilgado" authorityYear="2011" box="[504,727,1393,1415]" class="Insecta" family="Carabidae" genus="Typhlocharis" kingdom="Animalia" order="Coleoptera" pageId="11" pageNumber="1242" phylum="Arthropoda" rank="species" species="josabelae" status="sp. nov.">
|
||
<emphasis id="649CAE1877504541FE3A4814DBD23C8F" box="[504,727,1393,1415]" italics="true" pageId="11" pageNumber="1242">Typhlocharis josabelae</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="7FAF136377504541FD1E4817DA223C8F" box="[732,807,1394,1415]" pageId="11" pageNumber="1242" rank="species">sp. nov.</taxonomicNameLabel>
|
||
(A) Median lobe of aedeagus in left lateral view; (B) median lobe and right paramere in dorsal view; (C) left paramere in left lateral view; (D) right paramere in right lateral view; (E) apodemal ring; (F) right gonocoxite in ventral view of paratype 1; (G) right gonocoxite in ventral view of paratype 2; (H) spermatheca, glandule and conduct. Scale bars: A–D, F–H, 0.05 mm; E, 0.1 mm.
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="5657720A77504546FF794B44DA1439F6" blockId="11.[144,1101,1569,1688]" lastBlockId="12.[213,1170,166,1497]" lastPageId="12" lastPageNumber="1243" pageId="11" pageNumber="1242">
|
||
Cephalic chaetotaxy (
|
||
<figureCitation id="CED36E8F77504541FE734B44DB1A3F31" box="[433,543,1569,1593]" captionStart="Figure 2" captionStartId="9.[144,211,1654,1676]" captionTargetBox="[145,1099,164,1619]" captionTargetId="figure-6@9.[142,1102,161,1622]" captionTargetPageId="9" captionText="Figure 2. Habitus of Typhlocharis josabelae sp. nov. Scale bar: 0.3 mm." figureDoi="http://doi.org/10.5281/zenodo.5203657" httpUri="https://zenodo.org/record/5203657/files/figure.png" pageId="11" pageNumber="1242">Figures 2</figureCitation>
|
||
,
|
||
<figureCitation id="CED36E8F77504541FDE94B44DB553F31" box="[555,592,1569,1593]" captionStart="Figure 3" captionStartId="10.[213,280,1238,1260]" captionTargetBox="[211,1171,160,1206]" captionTargetId="figure-5@10.[211,1171,160,1206]" captionTargetPageId="10" captionText="Figure 3. Morphological details of Typhlocharis josabelae sp. nov. (A) Labium and prebasilar; (B) posterior angle of pronotum; (C) left protibia and protarsum in ventral view; (D) last abdominal sternite of male; (E) last abdominal sternite of female. Scale bars: A–C, 0.05 mm; D,E, 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.5203659" httpUri="https://zenodo.org/record/5203659/files/figure.png" pageId="11" pageNumber="1242">3A</figureCitation>
|
||
): on dorsal surface there are a series of large setae on each side of symmetry plane (a couple on the supraocular region, a seta near lateral carina, a seta on frontal sulcus, and two setae on clypeus); six setae on labrum, the longest lateral, a seta in the mandibular scrobe, two large setae inserted beside basal part of tooth in labium, and there is a pair of setae (one long and one very short) in middle region of each epilobe. The basilar has a pair of setae on both sides of plane of symmetry (the basal one is considerably longer).
|
||
</paragraph>
|
||
<paragraph id="5657720A77574546FEC24C63DA9D3875" blockId="12.[213,1170,166,1497]" pageId="12" pageNumber="1243">
|
||
Pronotum trapezoidal (
|
||
<figureCitation id="CED36E8F77574546FDED4C63DBA03816" box="[559,677,262,286]" captionStart="Figure 2" captionStartId="9.[144,211,1654,1676]" captionTargetBox="[145,1099,164,1619]" captionTargetId="figure-6@9.[142,1102,161,1622]" captionTargetPageId="9" captionText="Figure 2. Habitus of Typhlocharis josabelae sp. nov. Scale bar: 0.3 mm." figureDoi="http://doi.org/10.5281/zenodo.5203657" httpUri="https://zenodo.org/record/5203657/files/figure.png" pageId="12" pageNumber="1243">Figure 2</figureCitation>
|
||
), longer (
|
||
<quantity id="9110DFEF77574546FCF54C63DAE93816" box="[823,1004,262,286]" metricMagnitude="-4" metricUnit="m" metricValue="3.4499999999999997" metricValueMax="3.5" metricValueMin="3.4" pageId="12" pageNumber="1243" unit="mm" value="0.345" valueMax="0.35" valueMin="0.34">0.34–0.35 mm</quantity>
|
||
) than wide (
|
||
<quantity id="9110DFEF77574546FF1F4C40D8853835" box="[221,384,293,317]" metricMagnitude="-4" metricUnit="m" metricValue="3.25" metricValueMax="3.3" metricValueMin="3.2" pageId="12" pageNumber="1243" unit="mm" value="0.325" valueMax="0.33" valueMin="0.32">0.32–0.33 mm</quantity>
|
||
). Proximal margin nearly as wide as distal margin. Posterior angles indicated by a small tooth preceded by marginal edge, which is slightly wavy (
|
||
<figureCitation id="CED36E8F77574546FF1F4C00D8543875" box="[221,337,357,381]" captionStart="Figure 3" captionStartId="10.[213,280,1238,1260]" captionTargetBox="[211,1171,160,1206]" captionTargetId="figure-5@10.[211,1171,160,1206]" captionTargetPageId="10" captionText="Figure 3. Morphological details of Typhlocharis josabelae sp. nov. (A) Labium and prebasilar; (B) posterior angle of pronotum; (C) left protibia and protarsum in ventral view; (D) last abdominal sternite of male; (E) last abdominal sternite of female. Scale bars: A–C, 0.05 mm; D,E, 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.5203659" httpUri="https://zenodo.org/record/5203659/files/figure.png" pageId="12" pageNumber="1243">Figure 3B</figureCitation>
|
||
). The microsculpture is polygonal and isodiametric.
|
||
</paragraph>
|
||
<paragraph id="5657720A77574546FEC24CE0DA5C38D5" blockId="12.[213,1170,166,1497]" pageId="12" pageNumber="1243">
|
||
Pronotal chaetotaxy (
|
||
<figureCitation id="CED36E8F77574546FE324CE0DB593895" box="[496,604,389,413]" captionStart="Figure 2" captionStartId="9.[144,211,1654,1676]" captionTargetBox="[145,1099,164,1619]" captionTargetId="figure-6@9.[142,1102,161,1622]" captionTargetPageId="9" captionText="Figure 2. Habitus of Typhlocharis josabelae sp. nov. Scale bar: 0.3 mm." figureDoi="http://doi.org/10.5281/zenodo.5203657" httpUri="https://zenodo.org/record/5203657/files/figure.png" pageId="12" pageNumber="1243">Figures 2</figureCitation>
|
||
,
|
||
<figureCitation id="CED36E8F77574546FDA44CE0DB823895" box="[614,647,389,413]" captionStart="Figure 3" captionStartId="10.[213,280,1238,1260]" captionTargetBox="[211,1171,160,1206]" captionTargetId="figure-5@10.[211,1171,160,1206]" captionTargetPageId="10" captionText="Figure 3. Morphological details of Typhlocharis josabelae sp. nov. (A) Labium and prebasilar; (B) posterior angle of pronotum; (C) left protibia and protarsum in ventral view; (D) last abdominal sternite of male; (E) last abdominal sternite of female. Scale bars: A–C, 0.05 mm; D,E, 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.5203659" httpUri="https://zenodo.org/record/5203659/files/figure.png" pageId="12" pageNumber="1243">3B</figureCitation>
|
||
): disc with numerous setae, distal and proximal margins with rows of short setae, lateral margin with pair of long setae, one inserted near first quarter and the other next to the angular tooth.
|
||
</paragraph>
|
||
<paragraph id="5657720A77574546FEC24C80DBAC3B74" blockId="12.[213,1170,166,1497]" pageId="12" pageNumber="1243">
|
||
Elytra (from shoulder to apical edge) almost twice as long as wide (length:
|
||
<quantity id="9110DFEF77574546FB974C80D83B3B15" metricMagnitude="-4" metricUnit="m" metricValue="6.3" metricValueMax="6.4" metricValueMin="6.2" pageId="12" pageNumber="1243" unit="mm" value="0.63" valueMax="0.64" valueMin="0.62">0.62– 0.64 mm</quantity>
|
||
, width:
|
||
<quantity id="9110DFEF77574546FE5F4F60DB463B15" box="[413,579,517,541]" metricMagnitude="-4" metricUnit="m" metricValue="3.25" metricValueMax="3.3" metricValueMin="3.2" pageId="12" pageNumber="1243" unit="mm" value="0.325" valueMax="0.33" valueMin="0.32">0.32–0.33 mm</quantity>
|
||
)..Longitudinal carina parallel to lateral margin of elytra, which runs from humeral region to subapical region. Elytral margin finely serrulated, diminishing towards apex. Rounded apex, showing no conspicuous teeth. Isodiametric polygonal microreticulation.
|
||
</paragraph>
|
||
<paragraph id="5657720A77574546FEC24FE1DA373BF4" blockId="12.[213,1170,166,1497]" pageId="12" pageNumber="1243">
|
||
Elytral chaetotaxy (
|
||
<figureCitation id="CED36E8F77574546FE234FE1DB403B94" box="[481,581,644,668]" captionStart="Figure 2" captionStartId="9.[144,211,1654,1676]" captionTargetBox="[145,1099,164,1619]" captionTargetId="figure-6@9.[142,1102,161,1622]" captionTargetPageId="9" captionText="Figure 2. Habitus of Typhlocharis josabelae sp. nov. Scale bar: 0.3 mm." figureDoi="http://doi.org/10.5281/zenodo.5203657" httpUri="https://zenodo.org/record/5203657/files/figure.png" pageId="12" pageNumber="1243">Figure 2</figureCitation>
|
||
): numerous setae on disc, although some are somewhat longer than others, such as scutellar pore seta, and three or four discal setae, longitudinally aligned parallel to longitudinal carina. Setae of umbilicate series, whose formula is (4+3), stand out because of their large size.
|
||
</paragraph>
|
||
<paragraph id="5657720A77574546FEC24E60DBAA3AB3" blockId="12.[213,1170,166,1497]" pageId="12" pageNumber="1243">
|
||
Legs of males and females do not exhibit sexual dimorphism, so there are no special structures. However, it is interesting to note a relevant structure that has only been cited in
|
||
<bibRefCitation id="32790FFB77574546FE644E26DB4C3A53" author="Ortuno VM" box="[422,585,835,859]" pageId="12" pageNumber="1243" pagination="639 - 646" refId="ref10125" refString="Ortuno VM. 2005. A new endogaeic ground beetle from eastern Spain: Typhlocharis gonzaloi sp. n. (Coleoptera: Carabidae: Anillini). Rev suisse Zool. 112 (3): 639 - 646." type="journal article" year="2005">Ortuño (2005)</bibRefCitation>
|
||
and
|
||
<bibRefCitation id="32790FFB77574546FD414E26DA753A53" author="Andujar C & Lencina JL & Serrano J" box="[643,880,835,859]" pageId="12" pageNumber="1243" pagination="35 - 44" refId="ref9142" refString="Andujar C, Lencina JL, Serrano J. 2008. Typhlocharis Dieck, 1869 (Coleoptera; Carabidae, Anillini): a new species from the Iberian Peninsula, with notes about its relationships and the evolution of the diecki species group. Zootaxa 1842: 35 - 44." type="journal article" year="2008">Andújar et al. (2008)</bibRefCitation>
|
||
for
|
||
<taxonomicName id="91E8098977574546FC634E21DD943A54" authorityName="Ortuno" authorityYear="2005" box="[929,1169,836,860]" class="Insecta" family="Carabidae" genus="Typhlocharis" kingdom="Animalia" order="Coleoptera" pageId="12" pageNumber="1243" phylum="Arthropoda" rank="species" species="gonzaloi">
|
||
<emphasis id="649CAE1877574546FC634E21DD943A54" box="[929,1169,836,860]" italics="true" pageId="12" pageNumber="1243">Typhlocharis gonzaloi</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="91E8098977574546FECA4E01D8E83A73" authorityName="Andujar; Lencina & A. Serrano" authorityYear="2008" box="[264,493,868,892]" class="Insecta" family="Carabidae" genus="Typhlocharis" kingdom="Animalia" order="Coleoptera" pageId="12" pageNumber="1243" phylum="Arthropoda" rank="species" species="martini">
|
||
<emphasis id="649CAE1877574546FECA4E01D8E83A73" box="[264,493,868,892]" italics="true" pageId="12" pageNumber="1243">Typhlocharis martini</emphasis>
|
||
</taxonomicName>
|
||
, respectively: the clip seta of the protibial antennal cleaner. As in other species of the genus, it has a spatuliform shape (
|
||
<figureCitation id="CED36E8F77574546FC634EE6DD1F3A93" box="[929,1050,899,923]" captionStart="Figure 3" captionStartId="10.[213,280,1238,1260]" captionTargetBox="[211,1171,160,1206]" captionTargetId="figure-5@10.[211,1171,160,1206]" captionTargetPageId="10" captionText="Figure 3. Morphological details of Typhlocharis josabelae sp. nov. (A) Labium and prebasilar; (B) posterior angle of pronotum; (C) left protibia and protarsum in ventral view; (D) last abdominal sternite of male; (E) last abdominal sternite of female. Scale bars: A–C, 0.05 mm; D,E, 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.5203659" httpUri="https://zenodo.org/record/5203659/files/figure.png" pageId="12" pageNumber="1243">Figure 3C</figureCitation>
|
||
), which is substantially different from other
|
||
<taxonomicName id="91E8098977574546FD934EC6DBAE3AB3" baseAuthorityName="Vigna Taglianti" baseAuthorityYear="1972" box="[593,683,931,955]" class="Insecta" family="Carabidae" kingdom="Animalia" order="Coleoptera" pageId="12" pageNumber="1243" phylum="Arthropoda" rank="tribe" tribe="Anillini">Anillini</taxonomicName>
|
||
.
|
||
</paragraph>
|
||
<paragraph id="5657720A77574546FEC24EA6DA553D72" blockId="12.[213,1170,166,1497]" pageId="12" pageNumber="1243">
|
||
Abdomen with two long setae in males and four in females in last sternite,. Apodemal ring elliptical (
|
||
<figureCitation id="CED36E8F77574546FDC44E86DB853AF3" box="[518,640,995,1019]" captionStart="Figure 4" captionStartId="11.[144,211,1393,1415]" captionTargetBox="[142,1102,159,1360]" captionTargetId="figure-6@11.[142,1102,159,1360]" captionTargetPageId="11" captionText="Figure 4. Details of the genitalia of Typhlocharis josabelae sp. nov. (A) Median lobe of aedeagus in left lateral view; (B) median lobe and right paramere in dorsal view; (C) left paramere in left lateral view; (D) right paramere in right lateral view; (E) apodemal ring; (F) right gonocoxite in ventral view of paratype 1; (G) right gonocoxite in ventral view of paratype 2; (H) spermatheca, glandule and conduct. Scale bars: A–D, F–H, 0.05 mm; E, 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.5203661" httpUri="https://zenodo.org/record/5203661/files/figure.png" pageId="12" pageNumber="1243">Figure 4E</figureCitation>
|
||
). Median lobe of aedeagus (
|
||
<figureCitation id="CED36E8F77574546FC1A4E86DD6E3AF3" box="[984,1131,995,1019]" captionStart="Figure 4" captionStartId="11.[144,211,1393,1415]" captionTargetBox="[142,1102,159,1360]" captionTargetId="figure-6@11.[142,1102,159,1360]" captionTargetPageId="11" captionText="Figure 4. Details of the genitalia of Typhlocharis josabelae sp. nov. (A) Median lobe of aedeagus in left lateral view; (B) median lobe and right paramere in dorsal view; (C) left paramere in left lateral view; (D) right paramere in right lateral view; (E) apodemal ring; (F) right gonocoxite in ventral view of paratype 1; (G) right gonocoxite in ventral view of paratype 2; (H) spermatheca, glandule and conduct. Scale bars: A–D, F–H, 0.05 mm; E, 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.5203661" httpUri="https://zenodo.org/record/5203661/files/figure.png" pageId="12" pageNumber="1243">Figure 4A,B</figureCitation>
|
||
) is 174 µm in length. It is slightly curved and the apex points downward slightly compared with basal lamina. Internal sac is small and resembles the Greek letter ‘gamma’. Parameres with two setae, left one with a normal, subtriangular appearance, and right one narrow, abnormally long and hyaline (
|
||
<figureCitation id="CED36E8F77574546FD734907DA463D72" box="[689,835,1122,1146]" captionStart="Figure 4" captionStartId="11.[144,211,1393,1415]" captionTargetBox="[142,1102,159,1360]" captionTargetId="figure-6@11.[142,1102,159,1360]" captionTargetPageId="11" captionText="Figure 4. Details of the genitalia of Typhlocharis josabelae sp. nov. (A) Median lobe of aedeagus in left lateral view; (B) median lobe and right paramere in dorsal view; (C) left paramere in left lateral view; (D) right paramere in right lateral view; (E) apodemal ring; (F) right gonocoxite in ventral view of paratype 1; (G) right gonocoxite in ventral view of paratype 2; (H) spermatheca, glandule and conduct. Scale bars: A–D, F–H, 0.05 mm; E, 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.5203661" httpUri="https://zenodo.org/record/5203661/files/figure.png" pageId="12" pageNumber="1243">Figure 4C,D</figureCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="5657720A77574546FEC249E7DA253C12" blockId="12.[213,1170,166,1497]" pageId="12" pageNumber="1243">
|
||
Female genitalia (
|
||
<figureCitation id="CED36E8F77574546FE0A49E7DB513D92" box="[456,596,1154,1178]" captionStart="Figure 4" captionStartId="11.[144,211,1393,1415]" captionTargetBox="[142,1102,159,1360]" captionTargetId="figure-6@11.[142,1102,159,1360]" captionTargetPageId="11" captionText="Figure 4. Details of the genitalia of Typhlocharis josabelae sp. nov. (A) Median lobe of aedeagus in left lateral view; (B) median lobe and right paramere in dorsal view; (C) left paramere in left lateral view; (D) right paramere in right lateral view; (E) apodemal ring; (F) right gonocoxite in ventral view of paratype 1; (G) right gonocoxite in ventral view of paratype 2; (H) spermatheca, glandule and conduct. Scale bars: A–D, F–H, 0.05 mm; E, 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.5203661" httpUri="https://zenodo.org/record/5203661/files/figure.png" pageId="12" pageNumber="1243">Figure 4F,G</figureCitation>
|
||
) trimmer and slightly sclerotized. It has unguiform gonocoxites, with a seta inserted along external edge and a long seta in a fovea located on ventral surface. Sub-oval gonosubcoxite. Joint line with the gonocoxite almost imperceptible. Rhombus-shaped lateroterguite IX, generally smaller, in relation to gonocoxites and gonosubcoxites, than in
|
||
<taxonomicName id="91E8098977574546FD664867DA193C12" authorityName="Latreille" authorityYear="1802" box="[676,796,1282,1306]" class="Insecta" family="Carabidae" kingdom="Animalia" order="Coleoptera" pageId="12" pageNumber="1243" phylum="Arthropoda" rank="family">Carabidae</taxonomicName>
|
||
.
|
||
</paragraph>
|
||
<paragraph id="5657720A77574546FEC24847D8553CD1" blockId="12.[213,1170,166,1497]" pageId="12" pageNumber="1243">
|
||
Spermathecal complex (
|
||
<figureCitation id="CED36E8F77574546FDD54847DB973C32" box="[535,658,1314,1338]" captionStart="Figure 4" captionStartId="11.[144,211,1393,1415]" captionTargetBox="[142,1102,159,1360]" captionTargetId="figure-6@11.[142,1102,159,1360]" captionTargetPageId="11" captionText="Figure 4. Details of the genitalia of Typhlocharis josabelae sp. nov. (A) Median lobe of aedeagus in left lateral view; (B) median lobe and right paramere in dorsal view; (C) left paramere in left lateral view; (D) right paramere in right lateral view; (E) apodemal ring; (F) right gonocoxite in ventral view of paratype 1; (G) right gonocoxite in ventral view of paratype 2; (H) spermatheca, glandule and conduct. Scale bars: A–D, F–H, 0.05 mm; E, 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.5203661" httpUri="https://zenodo.org/record/5203661/files/figure.png" pageId="12" pageNumber="1243">Figure 4H</figureCitation>
|
||
) with membranous canal that is 86 µm in length and 3.5 µm in diameter, expanding (6.8 µm) at distal end. Spermatheca is pyriform and very sclerotized, with a diameter between 5 and 15 µm. There is a short membranous tube on this structure that expands gradually towards the end in a sclerotic and fusiform section that matches the spermathecal gland of other
|
||
<taxonomicName id="91E8098977574546FF1748A4D84E3CD1" authorityName="Latreille" authorityYear="1802" box="[213,331,1473,1497]" class="Insecta" family="Carabidae" kingdom="Animalia" order="Coleoptera" pageId="12" pageNumber="1243" phylum="Arthropoda" rank="family">Carabidae</taxonomicName>
|
||
.
|
||
</paragraph>
|
||
<paragraph id="5657720A77574546FF174B7BD8483F3E" blockId="12.[213,333,1566,1590]" box="[213,333,1566,1590]" pageId="12" pageNumber="1243">
|
||
<emphasis id="649CAE1877574546FF174B7BD8483F3E" box="[213,333,1566,1590]" italics="true" pageId="12" pageNumber="1243">Etymology</emphasis>
|
||
</paragraph>
|
||
<paragraph id="5657720A77574546FF174B20DBC43F96" blockId="12.[213,1170,1605,1694]" pageId="12" pageNumber="1243">
|
||
This new species is named after Dr Josabel Belliure from the Department of Ecology at Alcalá University (
|
||
<collectingCountry id="2EFF329A77574546FE124B00DB103F75" box="[464,533,1637,1661]" name="Spain" pageId="12" pageNumber="1243">Spain</collectingCountry>
|
||
), who worked on the first exploration of the endogean environment of the Ponoig mountain range.
|
||
</paragraph>
|
||
<paragraph id="5657720A77564547FF524DC3D86939B6" blockId="13.[144,364,166,190]" box="[144,364,166,190]" pageId="13" pageNumber="1244">
|
||
<emphasis id="649CAE1877564547FF524DC3D86939B6" bold="true" box="[144,364,166,190]" italics="true" pageId="13" pageNumber="1244">Habitat and biology</emphasis>
|
||
</paragraph>
|
||
<paragraph id="5657720A77564547FF524DABDADC3B4C" blockId="13.[144,1101,206,772]" pageId="13" pageNumber="1244">
|
||
<taxonomicName id="91E8098977564547FF524DABD88C39EE" authority="Ortuño & Gilgado, 2011" authorityName="Ortuño & Gilgado" authorityYear="2011" box="[144,393,206,230]" class="Insecta" family="Carabidae" genus="Typhlocharis" kingdom="Animalia" order="Coleoptera" pageId="13" pageNumber="1244" phylum="Arthropoda" rank="species" species="josabelae" status="sp. nov.">
|
||
<emphasis id="649CAE1877564547FF524DABD88C39EE" box="[144,393,206,230]" italics="true" pageId="13" pageNumber="1244">Typhlocharis josabelae</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="7FAF136377564547FE4D4DAAD8E639EE" box="[399,483,207,230]" pageId="13" pageNumber="1244" rank="species">sp. nov.</taxonomicNameLabel>
|
||
was found in the “Collado del Llamp” (Llamp Pass), a small mountain pass located
|
||
<quantity id="9110DFEF77564547FE184D8BDB25380E" box="[474,544,238,262]" metricMagnitude="2" metricUnit="m" metricValue="9.3" pageId="13" pageNumber="1244" unit="m" value="930.0">930 m</quantity>
|
||
above sea level on the northwest slope of Mount Ponoig. The lithology of this mountain is mainly limestone. The characteristics of this soil
|
||
<typeStatus id="8953CCA877564547FF024C4BD9F7384E" box="[192,242,302,326]" pageId="13" pageNumber="1244">type</typeStatus>
|
||
, limestone marl, allow the infiltration of water in the form of precipitation. The vegetation is thermophilic and shows adaptations to the characteristic xeric conditions of the Mediterranean forest. In addition to abundant grasses, the following plant species are dominant:
|
||
<taxonomicName id="91E8098977564547FE4F4CEBDB8138AD" authority="Mill." authorityName="Mill." box="[397,644,397,421]" class="Pinopsida" family="Pinaceae" genus="Pinus" kingdom="Plantae" order="Pinales" pageId="13" pageNumber="1244" phylum="Tracheophyta" rank="species" species="halepensis">
|
||
<emphasis id="649CAE1877564547FE4F4CEBDB4738AD" box="[397,578,397,421]" italics="true" pageId="13" pageNumber="1244">Pinus halepensis</emphasis>
|
||
Mill.
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="91E8098977564547FD534CE8DA5638AD" box="[657,851,397,421]" class="Magnoliopsida" family="Fagaceae" genus="Quercus" kingdom="Plantae" order="Fagales" pageId="13" pageNumber="1244" phylum="Tracheophyta" rank="species" species="coccifera">
|
||
<emphasis id="649CAE1877564547FD534CE8DA5638AD" box="[657,851,397,421]" italics="true" pageId="13" pageNumber="1244">Quercus coccifera</emphasis>
|
||
</taxonomicName>
|
||
L.,
|
||
<taxonomicName id="91E8098977564547FC464CE8DD2038AD" box="[900,1061,397,421]" class="Insecta" family="Carabidae" genus="Cystus" kingdom="Animalia" order="Coleoptera" pageId="13" pageNumber="1244" phylum="Arthropoda" rank="species" species="albidus">
|
||
<emphasis id="649CAE1877564547FC464CE8DD2038AD" box="[900,1061,397,421]" italics="true" pageId="13" pageNumber="1244">Cystus albidus</emphasis>
|
||
</taxonomicName>
|
||
L.,
|
||
<taxonomicName id="91E8098977564547FF524CCBD88D38CD" box="[144,392,429,453]" class="Magnoliopsida" family="Lamiaceae" genus="Rosmarinus" higherTaxonomySource="GBIF" kingdom="Plantae" order="Lamiales" pageId="13" pageNumber="1244" phylum="Tracheophyta" rank="species" species="officinalis">
|
||
<emphasis id="649CAE1877564547FF524CCBD88D38CD" box="[144,392,429,453]" italics="true" pageId="13" pageNumber="1244">Rosmarinus officinalis</emphasis>
|
||
</taxonomicName>
|
||
L.,
|
||
<taxonomicName id="91E8098977564547FE7F4CC8DB8438CD" box="[445,641,429,453]" class="Magnoliopsida" family="Fabaceae" genus="Coronilla" kingdom="Plantae" order="Fabales" pageId="13" pageNumber="1244" phylum="Tracheophyta" rank="species" species="minima">
|
||
<emphasis id="649CAE1877564547FE7F4CC8DB8438CD" box="[445,641,429,453]" italics="true" pageId="13" pageNumber="1244">Coronilla minima</emphasis>
|
||
</taxonomicName>
|
||
L. and
|
||
<taxonomicName id="91E8098977564547FD264CC8DD2B38CD" authority="L. However" authorityName="L. However" box="[740,1070,429,453]" class="Magnoliopsida" family="Lamiaceae" genus="Thymus" kingdom="Plantae" order="Lamiales" pageId="13" pageNumber="1244" phylum="Tracheophyta" rank="species" species="vulgaris">
|
||
<emphasis id="649CAE1877564547FD264CC8DA9E38CD" box="[740,923,429,453]" italics="true" pageId="13" pageNumber="1244">Thymus vulgaris</emphasis>
|
||
L. However
|
||
</taxonomicName>
|
||
, it should be noted that the
|
||
<taxonomicName id="91E8098977564547FE674CABDB2238ED" authority="Ortuño & Gilgado, 2011" authorityName="Ortuño & Gilgado" authorityYear="2011" box="[421,551,461,485]" class="Insecta" family="Carabidae" genus="Typhlocharis" kingdom="Animalia" order="Coleoptera" pageId="13" pageNumber="1244" phylum="Arthropoda" rank="species" species="josabelae" status="sp. nov.">
|
||
<emphasis id="649CAE1877564547FE674CABDB2238ED" box="[421,551,461,485]" italics="true" pageId="13" pageNumber="1244">T. josabelae</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="7FAF136377564547FDEF4CABDB8438ED" box="[557,641,462,485]" pageId="13" pageNumber="1244" rank="species">sp. nov.</taxonomicNameLabel>
|
||
specimens were collected at the head of a seasonal watercourse, so the substrate they inhabit maintains relatively high humidity and a lower temperature than other regions. The temperature of the soil was 14.4
|
||
<superScript id="A19DDF4277564547FBD14F6CDD193B12" attach="right" box="[1043,1052,521,538]" fontSize="7" pageId="13" pageNumber="1244">◦</superScript>
|
||
C at the time of collection at a depth of
|
||
<quantity id="9110DFEF77564547FDD84F49DB5A3B4C" box="[538,607,556,580]" metricMagnitude="-1" metricUnit="m" metricValue="1.5" pageId="13" pageNumber="1244" unit="cm" value="15.0">15 cm</quantity>
|
||
, compared with 22–24
|
||
<superScript id="A19DDF4277564547FC9E4F4CDA603B32" attach="right" box="[860,869,553,570]" fontSize="7" pageId="13" pageNumber="1244">◦</superScript>
|
||
C outside.
|
||
</paragraph>
|
||
<paragraph id="5657720A77564547FF794F29D8593A0C" blockId="13.[144,1101,206,772]" pageId="13" pageNumber="1244">
|
||
Other endogean beetles have been found in the same locality where
|
||
<taxonomicName id="91E8098977564547FC064F2BDD483B6D" authority="Ortuño & Gilgado, 2011" authorityName="Ortuño & Gilgado" authorityYear="2011" box="[964,1101,589,613]" class="Insecta" family="Carabidae" genus="Typhlocharis" kingdom="Animalia" order="Coleoptera" pageId="13" pageNumber="1244" phylum="Arthropoda" rank="species" species="josabelae" status="sp. nov.">
|
||
<emphasis id="649CAE1877564547FC064F2BDD483B6D" box="[964,1101,589,613]" italics="true" pageId="13" pageNumber="1244">T. josabelae</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="7FAF136377564547FF524F08D9EA3B8C" box="[144,239,621,644]" pageId="13" pageNumber="1244" rank="species">sp. nov.</taxonomicNameLabel>
|
||
was collected, including
|
||
<taxonomicName id="91E8098977564547FDED4F08DA8E3B8C" box="[559,907,620,645]" class="Insecta" family="Carabidae" genus="Microtyphlus" kingdom="Animalia" order="Coleoptera" pageId="13" pageNumber="1244" phylum="Arthropoda" rank="species" species="undetermined">
|
||
<emphasis id="649CAE1877564547FDED4F08DBC73B8D" box="[559,706,621,645]" italics="true" pageId="13" pageNumber="1244">Microtyphlus</emphasis>
|
||
sp. (Carabidae)
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="91E8098977564547FC624F09D83E3BAC" baseAuthorityName="Coiffait" baseAuthorityYear="1972" class="Insecta" family="Staphylinidae" kingdom="Animalia" order="Coleoptera" pageId="13" pageNumber="1244" phylum="Arthropoda" rank="subFamily" subFamily="Leptotyphlinae">Leptotyphlinae (Staphylinidae)</taxonomicName>
|
||
and
|
||
<taxonomicName id="91E8098977564547FE4A4FE9DB293BAC" authorityName="Leach" authorityYear="1815" box="[392,556,652,676]" class="Insecta" family="Scydmaenidae" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="13" pageNumber="1244" phylum="Arthropoda" rank="family">Scydmaenidae</taxonomicName>
|
||
. They are also accompanied by other soilendogean arthropods such as Protura,
|
||
<taxonomicName id="91E8098977564547FD424FC9DA183BCC" box="[640,797,684,708]" class="Entognatha" family="Campodeidae" kingdom="Animalia" order="Diplura" pageId="13" pageNumber="1244" phylum="Arthropoda" rank="family">Campodeidae</taxonomicName>
|
||
and
|
||
<taxonomicName id="91E8098977564547FCAE4FC9DADF3BCC" box="[876,986,684,708]" class="Entognatha" family="Japygidae" kingdom="Animalia" order="Diplura" pageId="13" pageNumber="1244" phylum="Arthropoda" rank="family">Japygidae</taxonomicName>
|
||
<taxonomicName id="91E8098977564547FC2F4FC9D8983BEC" authority=", Symphyla, Pauropoda" authorityName="Symphyla, Pauropoda" class="Entognatha" kingdom="Animalia" order="Diplura" pageId="13" pageNumber="1244" phylum="Arthropoda" rank="order">Diplura, Symphyla, Pauropoda</taxonomicName>
|
||
, and numerous small oribatid mites and Chilopoda Geophilomorpha.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |