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<document id="C86F1C7632EFF55A142CFA9626851143" ID-DOI="10.11646/zootaxa.5574.1.1" ID-ISSN="1175-5334" ID-Zenodo-Dep="14746103" ID-ZooBank="DA62BE83-26E8-487F-985A-1D48C3853025" IM.bibliography_approvedBy="felipe" IM.illustrations_approvedBy="felipe" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" IM.treatmentCitations_approvedBy="felipe" IM.treatments_approvedBy="felipe" checkinTime="1737969833979" checkinUser="plazi" docAuthor="Kc, Sajan &amp; Pollock, Darren A." docDate="2025" docId="E14D87AEFFC26E7A2A8DFA156AC22E31" docLanguage="en" docName="zootaxa.5574.1.1.pdf" docOrigin="Zootaxa 5574 (1)" docSource="https://doi.org/10.11646/zootaxa.5574.1.1" docStyle="DocumentStyle:5EBBA59367AD13919D70D935FA04F6A3.19:Zootaxa.2013-.monograph" docStyleId="5EBBA59367AD13919D70D935FA04F6A3" docStyleName="Zootaxa.2013-.monograph" docStyleVersion="19" docTitle="Hemipeplus nigerrimus KC &amp; Pollock 2025, sp. nov." docType="treatment" docVersion="2" lastPageNumber="123" masterDocId="1D74FFD6FFBA6E002A1AFF97694E2A20" masterDocTitle="Review of the Hemipeplinae (Coleoptera: Mycteridae) fauna of the world with descriptions of twenty-nine new species" masterLastPageNumber="140" masterPageNumber="1" pageNumber="121" updateTime="1737970979626" updateUser="ExternalLinkService">
<mods:mods id="B0E974099A0DFFDB4938E3FC2AE8B83F" xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo id="EB9610EC8F19467A3D69442C27B1FC2C">
<mods:title id="EABDB225C49DB6CC83236CF2215FDD33">Review of the Hemipeplinae (Coleoptera: Mycteridae) fauna of the world with descriptions of twenty-nine new species</mods:title>
</mods:titleInfo>
<mods:name id="498621BEBC576994D871AB30FE7F86FE" type="personal">
<mods:role id="7E7ECB3071B84687438E53E69E879A07">
<mods:roleTerm id="07628DEA595BCDAAC97697DFF1BDF06A">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="18AADCF125211A4DDBE413A399D2F7C0">Kc, Sajan</mods:namePart>
<mods:nameIdentifier id="1345AFB44EBCF8E518697872115092E4" type="ORCID">0000-0002-2749-0738</mods:nameIdentifier>
<mods:affiliation id="036E638704E5162BB69EBA43560D37C8">Department of Biology, Eastern New Mexico University, Portales, NM 88130 USA &amp; Corresponding author: Sajankc 143 @ gmail. com; https: // orcid. org / 0000 - 0002 - 2749 - 0738</mods:affiliation>
<mods:nameIdentifier id="33A2706CB98FE34A0F92E6319928A69D" type="email">Sajankc143@gmail.com</mods:nameIdentifier>
</mods:name>
<mods:name id="6F06BF1039BCC818FCF623B7855028F9" type="personal">
<mods:role id="E6E18130E9DFA17D6AA86AB1425FC0D6">
<mods:roleTerm id="CEDEA8C17BDA57A659A327037B821076">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="AC487D9936118C99BF715A4E53750D8E">Pollock, Darren A.</mods:namePart>
<mods:nameIdentifier id="047A5C6EAD200144219DCB2F6093C35E" type="ORCID">0000-0002-8709-0269</mods:nameIdentifier>
<mods:affiliation id="8F205D64A424542388DF78D380BA8074">Department of Biology, Eastern New Mexico University, Portales, NM 88130 USA &amp; Darren. pollock @ enmu. edu; https: // orcid. org / 0000 - 0002 - 8709 - 0269 &amp; Department of Biology, Eastern New Mexico University, Portales, NM 88130 USA &amp; Corresponding author: Sajankc 143 @ gmail. com; https: // orcid. org / 0000 - 0002 - 2749 - 0738</mods:affiliation>
<mods:nameIdentifier id="511F7B8DA1B02A176BDAD8ED934DCE52" type="email">Darren.pollock@enmu.edu</mods:nameIdentifier>
</mods:name>
<mods:typeOfResource id="0D23DA07A9D6A7087DBC5D57EDCF7A3A">text</mods:typeOfResource>
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<mods:titleInfo id="B12CAD4384B360B21BCECA54E15A9F38">
<mods:title id="34C14A3B20B1A4964CB3F6EADCBD2A48">Zootaxa</mods:title>
</mods:titleInfo>
<mods:part id="FB2DCDA1EB04673690876EE7FBCEDC62">
<mods:date id="BECA7C28F9645AC6375BF57279D35351">2025</mods:date>
<mods:detail id="6E640DEA169DA78F6837C6520266D793" type="pubDate">
<mods:number id="A8A7C66A97C5FCED1502B6511B818448">2025-01-21</mods:number>
</mods:detail>
<mods:detail id="DEDCC48F6A9ADCEEACBF0AA28F37482B" type="volume">
<mods:number id="FE314C6A29A4CF9DCEA8BDBD89D50215">5574</mods:number>
</mods:detail>
<mods:detail id="572C600934F810245CCF6F973772F503" type="issue">
<mods:number id="1B18B7AF08C69350B1366E304A34B0A0">1</mods:number>
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<mods:extent id="2F7A0D264D12FC1B649D1ABBF8C054B5" unit="page">
<mods:start id="47E6B90D6433304C0A606EFEB2A7C724">1</mods:start>
<mods:end id="2EC4D4519007C45E812F8DA5FEC60D33">140</mods:end>
</mods:extent>
</mods:part>
</mods:relatedItem>
<mods:location id="E5CE6B401FEF631CFC861793B5D77A53">
<mods:url id="CF6ED2CD7C00AAC5B718778111CFCFB8">https://doi.org/10.11646/zootaxa.5574.1.1</mods:url>
</mods:location>
<mods:classification id="6236DC064C0DB69A52A07470D8A2B950">journal article</mods:classification>
<mods:identifier id="001474DC6B5AB12145C922BB3F984E0C" type="DOI">10.11646/zootaxa.5574.1.1</mods:identifier>
<mods:identifier id="6B0D40A0D434588E5D25BAD3025A5A79" type="ISSN">1175-5334</mods:identifier>
<mods:identifier id="F998D5E930B7540C4059F3E98F404619" type="Zenodo-Dep">14746103</mods:identifier>
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</mods:mods>
<treatment id="E14D87AEFFC26E7A2A8DFA156AC22E31" ID-DOI="http://doi.org/10.5281/zenodo.14746163" ID-Zenodo-Dep="14746163" LSID="urn:lsid:plazi:treatment:E14D87AEFFC26E7A2A8DFA156AC22E31" httpUri="http://treatment.plazi.org/id/E14D87AEFFC26E7A2A8DFA156AC22E31" lastPageId="122" lastPageNumber="123" pageId="120" pageNumber="121">
<subSubSection id="21FE6533FFC26E782A8DFA156DC62FE1" pageId="120" pageNumber="121" type="nomenclature">
<paragraph id="695B36B8FFC26E782A8DFA156B972FBC" blockId="120.[151,1160,1410,1509]" box="[151,729,1410,1437]" pageId="120" pageNumber="121">
<heading id="321381D4FFC26E782A8DFA156B972FBC" bold="true" box="[151,729,1410,1437]" fontSize="11" level="2" pageId="120" pageNumber="121" reason="4">
<emphasis id="5B90EAAAFFC26E782A8DFA156B972FBC" bold="true" box="[151,729,1410,1437]" pageId="120" pageNumber="121">
<taxonomicName id="AEE44D3BFFC26E782A8DFA156B212FBC" authority="KC &amp; Pollock" authorityName="KC &amp; Pollock" authorityYear="2025" box="[151,623,1410,1437]" class="Insecta" family="Mycteridae" genus="Hemipeplus" kingdom="Animalia" order="Coleoptera" pageId="120" pageNumber="121" phylum="Arthropoda" rank="species" species="nigerrimus" status="sp. nov.">
<emphasis id="5B90EAAAFFC26E782A8DFA1568F82FBC" bold="true" box="[151,438,1410,1437]" italics="true" pageId="120" pageNumber="121">Hemipeplus nigerrimus</emphasis>
KC &amp; Pollock
</taxonomicName>
,
<taxonomicNameLabel id="40A357D1FFC26E782861FA156B972FBC" box="[635,729,1410,1436]" pageId="120" pageNumber="121" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
<paragraph id="695B36B8FFC26E782A8DFA316DC62FE1" blockId="120.[151,1160,1410,1509]" box="[151,1160,1446,1473]" pageId="120" pageNumber="121">
http://zoobank.org/
<uri id="1D753ABAFFC26E782B69FA316DC62FE1" box="[371,1160,1446,1473]" pageId="120" pageNumber="121">
urn:lsid:zoobank.org:act:
<uuid id="1D420C6DFFC26E78289EFA316DC62FE1" box="[644,1160,1446,1473]" pageId="120" pageNumber="121">9149255B-69C8-4043-89A9-01C4E4158FB0</uuid>
</uri>
</paragraph>
</subSubSection>
<subSubSection id="21FE6533FFC26E782A8DFA5D68692FC5" box="[151,295,1482,1509]" pageId="120" pageNumber="121" type="description">
<paragraph id="695B36B8FFC26E782A8DFA5D68692FC5" blockId="120.[151,1160,1410,1509]" box="[151,295,1482,1509]" pageId="120" pageNumber="121">(Figs 56, 65)</paragraph>
</subSubSection>
<subSubSection id="21FE6533FFC26E782A8DF9856A0F2D08" pageId="120" pageNumber="121" type="materials_examined">
<paragraph id="695B36B8FFC26E782A8DF9856D212C9D" blockId="120.[151,1437,1554,2013]" pageId="120" pageNumber="121">
<emphasis id="5B90EAAAFFC26E782A8DF985680A2C0C" bold="true" box="[151,324,1554,1580]" pageId="120" pageNumber="121">Type Material</emphasis>
.
<materialsCitation id="D98C3CE5FFC26E782B4FF9856BB32C70" collectingDate="1916-08-30" collectorName="Surigao &amp; Bottcher" country="Philippines" location="Philippinen" pageId="120" pageNumber="121" specimenCount="1" specimenCount-male="1" typeStatus="holotype">
<typeStatus id="B65F881AFFC26E782B4FF985688E2C0C" box="[341,448,1554,1580]" pageId="120" pageNumber="121" type="holotype">Holotype</typeStatus>
, male, labeled: ‘
<collectingCountry id="11F37628FFC26E782866F9856A4C2C0C" box="[636,770,1554,1580]" name="Philippines" pageId="120" pageNumber="121">Philippinen</collectingCountry>
Mindanao,
<collectorName id="C411536EFFC26E78299EF9856A952C0C" box="[900,987,1554,1580]" pageId="120" pageNumber="121">Surigao</collectorName>
leg.
<collectorName id="C411536EFFC26E782E11F9856D212C0C" box="[1035,1135,1554,1580]" pageId="120" pageNumber="121">Böttcher</collectorName>
[underside of label]
<date id="1D5A1078FFC26E782F7EF98569812C70" pageId="120" pageNumber="121" value="1916-08-30">
<collectingDate id="0D1EE990FFC26E782F7EF98569812C70" pageId="120" pageNumber="121" value="1916-08-30">30.8. 1916</collectingDate>
</date>
/ Zool. Mus. Berlin / [green label]
<specimenCount id="7FE2FD31FFC26E782847F9A06B3C2C71" box="[605,626,1591,1617]" count="1" pageId="120" pageNumber="121" type="male">♂</specimenCount>
/ [red label]
</materialsCitation>
<materialsCitation id="D98C3CE5FFC26E78291FF9A16D252C9D" collectingDate="1916-08-28" collectingDateMax="1916-08-30" collectingDateMin="1916-08-28" collectionCode="ZMHB" collectorName="Surigao &amp; Bottcher" country="Philippines" location="Philippinen" pageId="120" pageNumber="121" specimenCount="5" specimenCount-male="1" typeStatus="holotype">
<typeStatus id="B65F881AFFC26E78291FF9A16AD72C70" box="[773,921,1590,1616]" pageId="120" pageNumber="121" type="holotype">HOLOTYPE</typeStatus>
<specimenCount id="7FE2FD31FFC26E7829BBF9A06AF82C71" box="[929,950,1591,1617]" count="1" pageId="120" pageNumber="121" type="male">♂</specimenCount>
<taxonomicName id="AEE44D3BFFC26E7829A7F9A16DBA2C70" authority="KC" authorityName="KC &amp; Pollock" authorityYear="2025" box="[957,1268,1590,1616]" class="Insecta" family="Mycteridae" genus="Hemipeplus" kingdom="Animalia" order="Coleoptera" pageId="120" pageNumber="121" phylum="Arthropoda" rank="species" species="nigerrimus">
<emphasis id="5B90EAAAFFC26E7829A7F9A16D882C70" box="[957,1222,1590,1616]" italics="true" pageId="120" pageNumber="121">Hemipeplus nigerrimus</emphasis>
KC
</taxonomicName>
&amp; Pollock’, in
<collectionCode id="0FF5AE7DFFC26E782A8DF9CC69A22C55" box="[151,236,1627,1653]" pageId="120" pageNumber="121">ZMHB</collectionCode>
.
<specimenCount id="7FE2FD31FFC26E782AEFF9CC68D22C55" box="[245,412,1627,1653]" count="2" pageId="120" pageNumber="121" type="generic" typeStatus="paratype">Two paratypes</specimenCount>
:
<specimenCount id="7FE2FD31FFC26E782BBFF9CD6B692C54" box="[421,551,1626,1652]" count="1" pageId="120" pageNumber="121" type="generic">1 specimen</specimenCount>
, sex unknown, labeled: ‘
<collectingCountry id="11F37628FFC26E78292EF9CD6AF42C54" box="[820,954,1626,1652]" name="Philippines" pageId="120" pageNumber="121">Philippinen</collectingCountry>
Mindanao,
<collectorName id="C411536EFFC26E782E23F9CD6DDE2C54" box="[1081,1168,1626,1652]" pageId="120" pageNumber="121">Surigao</collectorName>
leg.
<collectorName id="C411536EFFC26E782EA1F9CD6C512C54" box="[1211,1311,1626,1652]" pageId="120" pageNumber="121">Böttcher</collectorName>
[underside of label]
<date id="1D5A1078FFC26E782AE6F9E968222CB8" box="[252,364,1662,1688]" pageId="120" pageNumber="121" value="1916-08-30">
<collectingDate id="0D1EE990FFC26E782AE6F9E968222CB8" box="[252,364,1662,1688]" pageId="120" pageNumber="121" value="1916-08-30">30.8.1916</collectingDate>
</date>
/
<taxonomicName id="AEE44D3BFFC26E782B98F9E96B262CB8" authority="Gestro" authorityName="Gestro" box="[386,616,1662,1688]" class="Insecta" family="Mycteridae" genus="Hemipeplus" kingdom="Animalia" order="Coleoptera" pageId="120" pageNumber="121" phylum="Arthropoda" rank="species" species="klematanicus">klematanicus Gestro</taxonomicName>
[handwritten in red ink]’, in
<collectionCode id="0FF5AE7DFFC26E7829AFF9E86D472CB9" box="[949,1033,1663,1689]" pageId="120" pageNumber="121">ZMHB</collectionCode>
;
<specimenCount id="7FE2FD31FFC26E782E0EF9E96DD82CB8" box="[1044,1174,1662,1688]" count="1" pageId="120" pageNumber="121" type="generic">1 specimen</specimenCount>
, sex unknown, labeled: ‘
<collectingCountry id="11F37628FFC26E782A81F935686F2C9C" box="[155,289,1698,1724]" name="Philippines" pageId="120" pageNumber="121">Philippinen</collectingCountry>
Mindanao,
<collectorName id="C411536EFFC26E782BBBF93568B62C9C" box="[417,504,1698,1724]" pageId="120" pageNumber="121">Surigao</collectorName>
leg.
<collectorName id="C411536EFFC26E78283FF9356BC72C9C" box="[549,649,1698,1724]" pageId="120" pageNumber="121">Böttcher</collectorName>
[underside of label]
<date id="1D5A1078FFC26E782968F9356AA42C9C" box="[882,1002,1698,1724]" pageId="120" pageNumber="121" value="1916-08-28">
<collectingDate id="0D1EE990FFC26E782968F9356AA42C9C" box="[882,1002,1698,1724]" pageId="120" pageNumber="121" value="1916-08-28">28.8.1916</collectingDate>
</date>
’, in
<collectionCode id="0FF5AE7DFFC26E782E0CF9346D252C9D" box="[1046,1131,1699,1725]" pageId="120" pageNumber="121">ZMHB</collectionCode>
</materialsCitation>
.
</paragraph>
<paragraph id="695B36B8FFC26E782ADDF9516A0F2D08" blockId="120.[151,1437,1554,2013]" pageId="120" pageNumber="121">
<emphasis id="5B90EAAAFFC26E782ADDF9516B692CC0" bold="true" box="[199,551,1734,1760]" pageId="120" pageNumber="121">Derivation of Specific Epithet</emphasis>
. The specific name (adjective in the nominative case, in gender agreement with substantive) is given in allusion to the holotype’s darker body color, distinguishing it from
<taxonomicName id="AEE44D3BFFC26E782EA8F97C6C252D24" box="[1202,1387,1770,1797]" class="Insecta" family="Mycteridae" genus="Hemipeplus" kingdom="Animalia" order="Coleoptera" pageId="120" pageNumber="121" phylum="Arthropoda" rank="species" species="klematanicus">
<emphasis id="5B90EAAAFFC26E782EA8F97C6D882D24" box="[1202,1222,1771,1796]" italics="true" pageId="120" pageNumber="121">H</emphasis>
.
<emphasis id="5B90EAAAFFC26E782ECDF97D6C252D24" box="[1239,1387,1770,1796]" italics="true" pageId="120" pageNumber="121">klematanicus</emphasis>
</taxonomicName>
and prompting a detailed examination of these darker specimens.
</paragraph>
</subSubSection>
<subSubSection id="21FE6533FFC26E792ADDF8A56D6A2DCA" lastPageId="121" lastPageNumber="122" pageId="120" pageNumber="121" type="diagnosis">
<paragraph id="695B36B8FFC26E792ADDF8A5686D2B3D" blockId="120.[151,1437,1554,2013]" lastBlockId="121.[151,1436,151,285]" lastPageId="121" lastPageNumber="122" pageId="120" pageNumber="121">
<emphasis id="5B90EAAAFFC26E782ADDF8A568742D6C" bold="true" box="[199,314,1842,1868]" pageId="120" pageNumber="121">Diagnosis</emphasis>
.
<taxonomicName id="AEE44D3BFFC26E782B50F8A56B1D2D6C" authorityName="KC &amp; Pollock" authorityYear="2025" box="[330,595,1842,1868]" class="Insecta" family="Mycteridae" genus="Hemipeplus" kingdom="Animalia" order="Coleoptera" pageId="120" pageNumber="121" phylum="Arthropoda" rank="species" species="nigerrimus" status="sp. nov.">
<emphasis id="5B90EAAAFFC26E782B50F8A56B1D2D6C" box="[330,595,1842,1868]" italics="true" pageId="120" pageNumber="121">Hemipeplus nigerrimus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="40A357D1FFC26E782846F8A46BFA2D6D" box="[604,692,1843,1869]" pageId="120" pageNumber="121" rank="species">sp. nov.</taxonomicNameLabel>
can be distinguished from the similar-appearing
<taxonomicName id="AEE44D3BFFC26E782EFFF8A46CD22D6C" box="[1253,1436,1842,1869]" class="Insecta" family="Mycteridae" genus="Hemipeplus" kingdom="Animalia" order="Coleoptera" pageId="120" pageNumber="121" phylum="Arthropoda" rank="species" species="klematanicus">
<emphasis id="5B90EAAAFFC26E782EFFF8A46DB72D6C" box="[1253,1273,1843,1868]" italics="true" pageId="120" pageNumber="121">H</emphasis>
.
<emphasis id="5B90EAAAFFC26E782F12F8A56CD22D6C" box="[1288,1436,1842,1868]" italics="true" pageId="120" pageNumber="121">klematanicus</emphasis>
</taxonomicName>
(Fig. 46) by its relatively smaller eyes, less acutely pointed posterior pronotal angles, and darker color.
<taxonomicName id="AEE44D3BFFC26E782F0CF8C1685C2DB4" authorityName="KC &amp; Pollock" authorityYear="2025" class="Insecta" family="Mycteridae" genus="Hemipeplus" kingdom="Animalia" order="Coleoptera" pageId="120" pageNumber="121" phylum="Arthropoda" rank="species" species="nigerrimus" status="sp. nov.">
<emphasis id="5B90EAAAFFC26E782F0CF8C1685C2DB4" italics="true" pageId="120" pageNumber="121">Hemipeplus nigerrimus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="40A357D1FFC26E782B01F8EC683D2DB5" box="[283,371,1915,1941]" pageId="120" pageNumber="121" rank="species">sp. nov.</taxonomicNameLabel>
has the following diagnostic features: temples short and angulate; scape moniliform, apically dilated; eyes moderately large (as long as scape + pedicel), moderately wide and moderately convex; pronotum subcordiform; anterior emargination distinct; anterior half slightly wider than posterior half; pronotal pad narrow, moderately distinct; anterior pronotal angles rounded, posterior angles slightly obtuse to square, more angled than rounded (never pointed acutely as in
<taxonomicName id="AEE44D3BFFC36E792854FF2C6A462AF5" box="[590,776,187,213]" class="Insecta" family="Mycteridae" genus="Hemipeplus" kingdom="Animalia" order="Coleoptera" pageId="121" pageNumber="122" phylum="Arthropoda" rank="species" species="klematanicus">
<emphasis id="5B90EAAAFFC36E792854FF2C6B2C2AF4" box="[590,610,187,212]" italics="true" pageId="121" pageNumber="122">H</emphasis>
.
<emphasis id="5B90EAAAFFC36E79286EFF2C6A462AF5" box="[628,776,187,213]" italics="true" pageId="121" pageNumber="122">klematanicus</emphasis>
</taxonomicName>
); pronotal lobe indistinct with faint median notch; body long; elytra convex; color dull, uniformly rufopiceous to darker head and pronotum; distribution:
<collectingCountry id="11F37628FFC36E792F04FF486CD22AD9" box="[1310,1436,223,249]" name="Philippines" pageId="121" pageNumber="122">Philippines</collectingCountry>
(Mindanao).
</paragraph>
<paragraph id="695B36B8FFC36E792A82F81F6D6A2DCA" blockId="121.[151,1436,1928,2027]" pageId="121" pageNumber="122">
<emphasis id="5B90EAAAFFC36E792A82F81F68632D82" bold="true" box="[152,301,1928,1954]" pageId="121" pageNumber="122">FIGURE 56</emphasis>
.
<taxonomicName id="AEE44D3BFFC36E792B20F81F6B022D82" authorityName="KC &amp; Pollock" authorityYear="2025" box="[314,588,1928,1955]" class="Insecta" family="Mycteridae" genus="Hemipeplus" kingdom="Animalia" order="Coleoptera" pageId="121" pageNumber="122" phylum="Arthropoda" rank="species" species="nigerrimus" status="sp. nov.">Hemipeplus nigerrimus</taxonomicName>
<taxonomicNameLabel id="40A357D1FFC36E792848F81F6BE32D83" box="[594,685,1928,1955]" pageId="121" pageNumber="122" rank="species">
<emphasis id="5B90EAAAFFC36E792848F81F6B222D82" bold="true" box="[594,620,1928,1954]" pageId="121" pageNumber="122">sp</emphasis>
.
<emphasis id="5B90EAAAFFC36E792860F81F6BE82D82" bold="true" box="[634,678,1928,1954]" pageId="121" pageNumber="122">nov</emphasis>
.
</taxonomicNameLabel>
<emphasis id="5B90EAAAFFC36E7928AEF81F6A7D2D82" bold="true" box="[692,819,1928,1954]" pageId="121" pageNumber="122">
(
<typeStatus id="B65F881AFFC36E7928A1F81F6A652D82" box="[699,811,1928,1954]" pageId="121" pageNumber="122" type="holotype">Holotype</typeStatus>
)
</emphasis>
. A. Dorsal habitus (with scale bar); B. Dorsal head and prothorax; C. Lateral view of head and prothorax; D. Male genitalia dorsal view with spiculum gastrale; E. Tegmen dorsal view (with scale bar); F. Tegmen lateral view with one paramere removed.
</paragraph>
</subSubSection>
<subSubSection id="21FE6533FFC06E7A2ADDFF016AB42985" pageId="122" pageNumber="123" type="description">
<paragraph id="695B36B8FFC06E7A2ADDFF016A742AF5" blockId="122.[151,1437,150,1041]" pageId="122" pageNumber="123">
<emphasis id="5B90EAAAFFC06E7A2ADDFF01681F2A90" bold="true" box="[199,337,150,176]" pageId="122" pageNumber="123">Description</emphasis>
. Measurements (in mm)—GHW: 0.50–0.56; HL: 0.45–0.52; AL: 0.85–1.01; GPW: 0.55–0.63; PL: 0.47–0.52; GEW: 0.60–0.66; EL: 2.63–2.96; TL: 3.55–4.00.
</paragraph>
<paragraph id="695B36B8FFC06E7A2ADDFF4868D628ED" blockId="122.[151,1437,150,1041]" pageId="122" pageNumber="123">
<emphasis id="5B90EAAAFFC06E7A2ADDFF4868492AD9" bold="true" box="[199,263,223,249]" pageId="122" pageNumber="123">Head</emphasis>
(Fig. 56B) wide (GHW/HL: 1.08–1.11); eyes moderately large (as long as scape + pedicel), moderately wide and moderately convex; temples short and angulate; occiput with strongly raised areas; antennae about subequal to head and pronotum length combined; antennal scape moniliform, apically dilated; pedicel small moniliform; antennomere III more elongate and apically dilated; IV shorter, moniliform; V–X subtriangular or bowl shaped, each wider than the next; XI subpyriform with apex narrowed after the notch; lateral mandibular tooth absent.
<emphasis id="5B90EAAAFFC06E7A2A8DFE04685E2B8D" bold="true" box="[151,272,403,429]" pageId="122" pageNumber="123">Pronotum</emphasis>
(Fig. 56B) wide (GPW/PL: 1.17–1.21); anterior margin with distinct emargination; anterolateral angles rounded, produced; lateral margins sinuate; posterior half much more contracted than anterior half; posterolateral angles slightly obtuse to square, more angled than rounded but not pointed (Fig. 56B); pronotal lobe indistinct with faint median notch; pronotal pad (Fig. 56C) narrow, moderately distinct; pronotal pits moderately deep with moderately developed grooves.
<emphasis id="5B90EAAAFFC06E7A2BE7FDB56BFB281C" bold="true" box="[509,693,546,572]" pageId="122" pageNumber="123">Scutellar shield</emphasis>
elliptical, transverse, less than twice as wide as long.
<emphasis id="5B90EAAAFFC06E7A2F09FDB46C11281D" bold="true" box="[1299,1375,547,573]" pageId="122" pageNumber="123">Elytra</emphasis>
(Fig. 56A) long (EL/GEW: 4.38–4.48), convex; vestiture short and dense; apical patch broad and slightly darker than elytra.
<emphasis id="5B90EAAAFFC06E7A2AFCFDFC68D028A4" bold="true" box="[230,414,618,645]" pageId="122" pageNumber="123">Ventral surface</emphasis>
. Prosternal process short, not extending posterior of procoxae; hypomeral edges narrowly separated; mesosternal process narrow and linear.
<emphasis id="5B90EAAAFFC06E7A28D7FD196A5F2888" bold="true" box="[717,785,654,680]" pageId="122" pageNumber="123">Color</emphasis>
dull, uniformly rufopiceous to head and pronotum darker than other dorsal parts.
</paragraph>
<paragraph id="695B36B8FFC06E7A2ADDFD40682929A1" blockId="122.[151,1437,150,1041]" pageId="122" pageNumber="123">
<emphasis id="5B90EAAAFFC06E7A2ADDFD406B6A28D0" bold="true" box="[199,548,726,753]" pageId="122" pageNumber="123">Male Genitalia (Figs 56D–F)</emphasis>
. (Tegmen length =
<quantity id="AE1C9B5DFFC06E7A2916FD416A3D28D1" box="[780,883,726,753]" metricMagnitude="-4" metricUnit="m" metricValue="3.5" pageId="122" pageNumber="123" unit="mm" value="0.35">0.35 mm</quantity>
(n = 1)) This is the only species in the Austro-Oriental region in which the apices of parameres are not arcuate or bulged, and are straight; lobe of basale long, slender with blunt apex; shoulders obtusely angled with moderate slant; apicale and basale with distinct suture; parameres with bases thicker than apices; apices straight without distinct curves; penis much longer than tegmen, medially widened.
</paragraph>
<paragraph id="695B36B8FFC06E7A2ADDFC1D6AB42985" blockId="122.[151,1437,150,1041]" box="[199,1018,906,933]" pageId="122" pageNumber="123">
<emphasis id="5B90EAAAFFC06E7A2ADDFC1D68762984" bold="true" box="[199,312,906,932]" pageId="122" pageNumber="123">Variation</emphasis>
. Specimens vary in the intensity of darkness of the body color.
</paragraph>
</subSubSection>
<subSubSection id="21FE6533FFC06E7A2ADDFC386AC22E31" pageId="122" pageNumber="123" type="discussion">
<paragraph id="695B36B8FFC06E7A2ADDFC386C1229CD" blockId="122.[151,1437,150,1041]" pageId="122" pageNumber="123">
<emphasis id="5B90EAAAFFC06E7A2ADDFC38687A29E9" bold="true" box="[199,308,943,969]" pageId="122" pageNumber="123">Remarks</emphasis>
.
<taxonomicName id="AEE44D3BFFC06E7A2B58FC386B0429E8" authorityName="KC &amp; Pollock" authorityYear="2025" box="[322,586,943,969]" class="Insecta" family="Mycteridae" genus="Hemipeplus" kingdom="Animalia" order="Coleoptera" pageId="122" pageNumber="123" phylum="Arthropoda" rank="species" species="nigerrimus" status="sp. nov.">
<emphasis id="5B90EAAAFFC06E7A2B58FC386B0429E8" box="[322,586,943,969]" italics="true" pageId="122" pageNumber="123">Hemipeplus nigerrimus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="40A357D1FFC06E7A284AFC386BE929E9" box="[592,679,943,969]" pageId="122" pageNumber="123" rank="species">sp. nov.</taxonomicNameLabel>
is similar to
<taxonomicName id="AEE44D3BFFC06E7A2922FC386AA029E9" box="[824,1006,943,969]" class="Insecta" family="Mycteridae" genus="Hemipeplus" kingdom="Animalia" order="Coleoptera" pageId="122" pageNumber="123" phylum="Arthropoda" rank="species" species="klematanicus">
<emphasis id="5B90EAAAFFC06E7A2922FC386A0229E8" box="[824,844,943,968]" italics="true" pageId="122" pageNumber="123">H</emphasis>
.
<emphasis id="5B90EAAAFFC06E7A2940FC386AA029E9" box="[858,1006,943,969]" italics="true" pageId="122" pageNumber="123">klematanicus</emphasis>
</taxonomicName>
(Fig. 46) (particularly the rufopiceous forms of
<taxonomicName id="AEE44D3BFFC06E7A2AE4FC4468FA29CD" box="[254,436,979,1005]" class="Insecta" family="Mycteridae" genus="Hemipeplus" kingdom="Animalia" order="Coleoptera" pageId="122" pageNumber="123" phylum="Arthropoda" rank="species" species="klematanicus">
<emphasis id="5B90EAAAFFC06E7A2AE4FC44685C29CC" box="[254,274,979,1004]" italics="true" pageId="122" pageNumber="123">H</emphasis>
.
<emphasis id="5B90EAAAFFC06E7A2B3AFC4468FA29CD" box="[288,436,979,1005]" italics="true" pageId="122" pageNumber="123">klematanicus</emphasis>
</taxonomicName>
); however, the genitalia are distinct between the two species (Figs 46D–F, 56D–F).
</paragraph>
<paragraph id="695B36B8FFC06E7A2ADDFC616AC22E31" blockId="122.[151,1437,150,1041]" box="[199,908,1014,1041]" pageId="122" pageNumber="123">
<emphasis id="5B90EAAAFFC06E7A2ADDFC616B742E30" bold="true" box="[199,570,1014,1040]" pageId="122" pageNumber="123">Geographical Distribution (Fig</emphasis>
.
<emphasis id="5B90EAAAFFC06E7A285DFC616B232E30" bold="true" box="[583,621,1014,1040]" pageId="122" pageNumber="123">65)</emphasis>
.
<collectingCountry id="11F37628FFC06E7A2861FC606BB72E31" box="[635,761,1015,1041]" name="Philippines" pageId="122" pageNumber="123">Philippines</collectingCountry>
(Mindanao).
</paragraph>
</subSubSection>
</treatment>
</document>
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