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<mods:title id="72CD12F0783E2F6A379AF3E09BFFFBE7">Review of the Hemipeplinae (Coleoptera: Mycteridae) fauna of the world with descriptions of twenty-nine new species</mods:title>
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<mods:namePart id="D79B5E1BB886216C8D29A3F57493EB3A">Kc, Sajan</mods:namePart>
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<mods:affiliation id="A5E71AF60B3A9FD892BF5DB1830A81F8">Department of Biology, Eastern New Mexico University, Portales, NM 88130 USA &amp; Corresponding author: Sajankc 143 @ gmail. com; https: // orcid. org / 0000 - 0002 - 2749 - 0738</mods:affiliation>
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<mods:namePart id="DA7D219DFA436F9FE21C9F8F04D01947">Pollock, Darren A.</mods:namePart>
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<mods:affiliation id="9579343035CCE1B921C88D42102AB5FF">Department of Biology, Eastern New Mexico University, Portales, NM 88130 USA &amp; Darren. pollock @ enmu. edu; https: // orcid. org / 0000 - 0002 - 8709 - 0269 &amp; Department of Biology, Eastern New Mexico University, Portales, NM 88130 USA &amp; Corresponding author: Sajankc 143 @ gmail. com; https: // orcid. org / 0000 - 0002 - 2749 - 0738</mods:affiliation>
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<mods:title id="066539420A950DEBB26C02389AEDA96E">Zootaxa</mods:title>
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<mods:date id="70424211A37CF608E7DB316AD2B060B2">2025</mods:date>
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<subSubSection id="21FE6533FFA26E182A8DFD19685428ED" pageId="24" pageNumber="25" type="nomenclature">
<paragraph id="695B36B8FFA26E182A8DFD196B322888" blockId="24.[151,636,654,717]" box="[151,636,654,681]" pageId="24" pageNumber="25">
<heading id="321381D4FFA26E182A8DFD196B322888" bold="true" box="[151,636,654,681]" fontSize="11" level="2" pageId="24" pageNumber="25" reason="3">
<taxonomicName id="AEE44D3BFFA26E182A8DFD196B322888" authority="Pollock, 1999" authorityName="Pollock" authorityYear="1999" box="[151,636,654,681]" class="Insecta" family="Mycteridae" genus="Hemipeplus" kingdom="Animalia" order="Coleoptera" pageId="24" pageNumber="25" phylum="Arthropoda" rank="species" species="dominicensis">
<emphasis id="5B90EAAAFFA26E182A8DFD196B322888" bold="true" box="[151,636,654,681]" pageId="24" pageNumber="25">
<emphasis id="5B90EAAAFFA26E182A8DFD1968802889" bold="true" box="[151,462,654,681]" italics="true" pageId="24" pageNumber="25">Hemipeplus dominicensis</emphasis>
<bibRefCitation id="0D754B49FFA26E182BCFFD186B322888" author="Pollock, D. A." box="[469,636,654,681]" pageId="24" pageNumber="25" pagination="47 - 73" refId="ref84437" refString="Pollock, D. A. (1999) Review of the New World Hemipeplinae (Coleoptera: Mycteridae) with descriptions of ten new species. Entomologica Scandinavica, 30 (1), 47 - 73. https: // doi. org / 10.1163 / 187631200 X 00200" type="journal article" year="1999">Pollock, 1999</bibRefCitation>
</emphasis>
</taxonomicName>
</heading>
</paragraph>
<paragraph id="695B36B8FFA26E182A8DFD24685428ED" blockId="24.[151,636,654,717]" box="[151,282,691,717]" pageId="24" pageNumber="25">(Figs 7, 61)</paragraph>
</subSubSection>
<subSubSection id="21FE6533FFA26E182A8DFD6068852980" pageId="24" pageNumber="25" type="reference_group">
<paragraph id="695B36B8FFA26E182A8DFD606A41292F" blockId="24.[151,783,759,783]" box="[151,783,759,783]" pageId="24" pageNumber="25">
<treatmentCitationGroup id="49F41196FFA26E182A8DFD606A41292F" box="[151,783,759,783]" pageId="24" pageNumber="25">
<treatmentCitation id="E84510A9FFA26E182A8DFD606B11292F" author="Pollock, D. A." box="[151,607,759,783]" page="65" pageId="24" pageNumber="25" year="1999">
<taxonomicName id="AEE44D3BFFA26E182A8DFD606B11292F" authority="Pollock, 1999: 65" authorityName="Pollock" authorityPageNumber="65" authorityYear="1999" box="[151,607,759,783]" class="Insecta" family="Mycteridae" genus="Hemipeplus" kingdom="Animalia" order="Coleoptera" pageId="24" pageNumber="25" phylum="Arthropoda" rank="species" species="dominicensis">
<emphasis id="5B90EAAAFFA26E182A8DFD6068EA292F" bold="true" box="[151,420,759,783]" italics="true" pageId="24" pageNumber="25">Hemipeplus dominicensis</emphasis>
<bibRefCitation id="0D754B49FFA26E182BB0FD606B11292F" author="Pollock, D. A." box="[426,607,759,783]" pageId="24" pageNumber="25" pagination="47 - 73" refId="ref84437" refString="Pollock, D. A. (1999) Review of the New World Hemipeplinae (Coleoptera: Mycteridae) with descriptions of ten new species. Entomologica Scandinavica, 30 (1), 47 - 73. https: // doi. org / 10.1163 / 187631200 X 00200" type="journal article" year="1999">Pollock, 1999: 65</bibRefCitation>
</taxonomicName>
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, Figs 2C, 5C, 11.
</treatmentCitationGroup>
</paragraph>
<paragraph id="695B36B8FFA26E182A8DFCAA68852980" blockId="24.[151,1437,829,1576]" pageId="24" pageNumber="25">
<emphasis id="5B90EAAAFFA26E182A8DFCAA68132977" bold="true" box="[151,349,829,855]" pageId="24" pageNumber="25">Taxonomic Note</emphasis>
. Although
<bibRefCitation id="0D754B49FFA26E182BF9FCAA6BC02977" author="Pollock, D. A." box="[483,654,829,855]" pageId="24" pageNumber="25" pagination="47 - 73" refId="ref84437" refString="Pollock, D. A. (1999) Review of the New World Hemipeplinae (Coleoptera: Mycteridae) with descriptions of ten new species. Entomologica Scandinavica, 30 (1), 47 - 73. https: // doi. org / 10.1163 / 187631200 X 00200" type="journal article" year="1999">Pollock (1999)</bibRefCitation>
stated that the
<typeStatus id="B65F881AFFA26E18295DFCAA6AE72977" box="[839,937,829,855]" pageId="24" pageNumber="25" type="holotype">holotype</typeStatus>
is the middle specimen of three, the label indicates that the top specimen is the
<typeStatus id="B65F881AFFA26E182856FCF66BFC295B" box="[588,690,865,891]" pageId="24" pageNumber="25" type="holotype">holotype</typeStatus>
. Therefore, we have given precedence to the
<typeStatus id="B65F881AFFA26E182EA5FCF66C6F295B" box="[1215,1313,865,891]" pageId="24" pageNumber="25" type="holotype">holotype</typeStatus>
label over
<bibRefCitation id="0D754B49FFA26E182A8DFC12681F29BF" author="Pollock, D. A." box="[151,337,901,927]" pageId="24" pageNumber="25" pagination="47 - 73" refId="ref84437" refString="Pollock, D. A. (1999) Review of the New World Hemipeplinae (Coleoptera: Mycteridae) with descriptions of ten new species. Entomologica Scandinavica, 30 (1), 47 - 73. https: // doi. org / 10.1163 / 187631200 X 00200" type="journal article" year="1999">Pollocks (1999)</bibRefCitation>
statement.
</paragraph>
</subSubSection>
<subSubSection id="21FE6533FFA26E182ADDFC3E6BAE2E0F" pageId="24" pageNumber="25" type="materials_examined">
<paragraph id="695B36B8FFA26E182ADDFC3E6BAE2E0F" blockId="24.[151,1437,829,1576]" pageId="24" pageNumber="25">
<emphasis id="5B90EAAAFFA26E182ADDFC3E683F29E3" bold="true" box="[199,369,937,963]" pageId="24" pageNumber="25">Type Material</emphasis>
.
<materialsCitation id="D98C3CE5FFA26E182B9AFC3E6CD229C7" collectingDate="1973-04-11" collectingDateMax="1976-01-19" collectingDateMin="1973-04-11" collectionCode="SMNS" collectorName="Republic Dominican Bani &amp; J. S. Klapperich" pageId="24" pageNumber="25" specimenCount="4" specimenCount-male="1" typeStatus="holotype">
<typeStatus id="B65F881AFFA26E182B9AFC3E68A529E3" box="[384,491,937,963]" pageId="24" pageNumber="25" type="holotype">Holotype</typeStatus>
, male, top specimen of
<specimenCount id="7FE2FD31FFA26E1828EDFC3E6AE229E3" box="[759,940,937,963]" count="3" pageId="24" pageNumber="25" type="generic">three specimens</specimenCount>
on same pin, labeled:
<collectorName id="C411536EFFA26E182EB4FC3E698229C7" pageId="24" pageNumber="25">Republic Dominican Bani</collectorName>
<date id="1D5A1078FFA26E182ACDFC5A680929C7" box="[215,327,973,999]" pageId="24" pageNumber="25" value="1976-01-19">
<collectingDate id="0D1EE990FFA26E182ACDFC5A680929C7" box="[215,327,973,999]" pageId="24" pageNumber="25" value="1976-01-19">19.1.1976</collectingDate>
</date>
leg.
<collectorName id="C411536EFFA26E182B9CFC5A6B7829C7" box="[390,566,973,999]" pageId="24" pageNumber="25">J.S. Klapperich</collectorName>
[underside of label]
<date id="1D5A1078FFA26E182935FC5A6AD029C7" box="[815,926,973,999]" pageId="24" pageNumber="25" value="1973-04-11">
<collectingDate id="0D1EE990FFA26E182935FC5A6AD029C7" box="[815,926,973,999]" pageId="24" pageNumber="25" value="1973-04-11">4.11.1973</collectingDate>
</date>
/
<collectionCode id="0FF5AE7DFFA26E1829A6FC5A6D4629C7" box="[956,1032,973,999]" country="Germany" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34840" lsid="urn:lsid:biocol.org:col:34840" name="Staatliches Museum fuer Naturkund Stuttgart" pageId="24" pageNumber="25">SMNS</collectionCode>
/ [green label]
<specimenCount id="7FE2FD31FFA26E182ED9FC596D9629C8" box="[1219,1240,974,1000]" count="1" pageId="24" pageNumber="25" type="male"></specimenCount>
top / [red label]
</materialsCitation>
<materialsCitation id="D98C3CE5FFA26E182A8DFC666B922E0F" collectionCode="SMNS" collectorName="D. A. Pollock'" pageId="24" pageNumber="25" specimenCount="2" typeStatus="holotype">
<typeStatus id="B65F881AFFA26E182A8DFC6668652E2B" box="[151,299,1009,1035]" pageId="24" pageNumber="25" type="holotype">HOLOTYPE</typeStatus>
(top spec.) &amp;
<typeStatus id="B65F881AFFA26E182BCFFC656B0E2E2C" box="[469,576,1010,1036]" pageId="24" pageNumber="25" type="paratype">paratypes</typeStatus>
<taxonomicName id="AEE44D3BFFA26E182853FC666A3F2E2B" authorityName="Kc &amp; Pollock" authorityYear="2025" box="[585,881,1009,1035]" class="Insecta" family="Mycteridae" genus="Hemipeplus" kingdom="Animalia" order="Coleoptera" pageId="24" pageNumber="25" phylum="Arthropoda" rank="species" species="dominicensis" status="sp. nov.">Hemipeplus dominicensis</taxonomicName>
,
<taxonomicNameLabel id="40A357D1FFA26E182964FC656AF22E2C" box="[894,956,1010,1036]" pageId="24" pageNumber="25" rank="species">n. sp.</taxonomicNameLabel>
<collectorName id="C411536EFFA26E1829DFFC656D292E2B" box="[965,1127,1009,1036]" pageId="24" pageNumber="25">D.A. Pollock</collectorName>
, in
<collectionCode id="0FF5AE7DFFA26E182E89FC666DAD2E2B" box="[1171,1251,1009,1035]" country="Germany" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34840" lsid="urn:lsid:biocol.org:col:34840" name="Staatliches Museum fuer Naturkund Stuttgart" pageId="24" pageNumber="25">SMNS</collectionCode>
.
<specimenCount id="7FE2FD31FFA26E182EF5FC656CD62E2C" box="[1263,1432,1010,1036]" count="2" pageId="24" pageNumber="25" type="generic" typeStatus="paratype">Two paratypes</specimenCount>
, mounted on the same pin as the
<typeStatus id="B65F881AFFA26E182BE5FB826B2A2E0F" box="[511,612,1045,1071]" pageId="24" pageNumber="25" type="holotype">holotype</typeStatus>
, in
<collectionCode id="0FF5AE7DFFA26E182896FB826B922E0F" box="[652,732,1045,1071]" country="Germany" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34840" lsid="urn:lsid:biocol.org:col:34840" name="Staatliches Museum fuer Naturkund Stuttgart" pageId="24" pageNumber="25">SMNS</collectionCode>
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="21FE6533FFA26E182ADDFBAE6D182F9B" pageId="24" pageNumber="25" type="diagnosis">
<paragraph id="695B36B8FFA26E182ADDFBAE6A362F0B" blockId="24.[151,1437,829,1576]" pageId="24" pageNumber="25">
<emphasis id="5B90EAAAFFA26E182ADDFBAE68742E73" bold="true" box="[199,314,1081,1107]" pageId="24" pageNumber="25">Diagnosis</emphasis>
.
<taxonomicName id="AEE44D3BFFA26E182B5DFBAE6B2B2E73" authorityName="Kc &amp; Pollock" authorityYear="2025" box="[327,613,1081,1107]" class="Insecta" family="Mycteridae" genus="Hemipeplus" kingdom="Animalia" order="Coleoptera" pageId="24" pageNumber="25" phylum="Arthropoda" rank="species" species="dominicensis">
<emphasis id="5B90EAAAFFA26E182B5DFBAE6B2B2E73" box="[327,613,1081,1107]" italics="true" pageId="24" pageNumber="25">Hemipeplus dominicensis</emphasis>
</taxonomicName>
can be distinguished by its elongate scape, lack of temples, and uniformly rufotestaceous color.
<taxonomicName id="AEE44D3BFFA26E182B98FBCA6BD02E57" authorityName="Kc &amp; Pollock" authorityYear="2025" box="[386,670,1117,1143]" class="Insecta" family="Mycteridae" genus="Hemipeplus" kingdom="Animalia" order="Coleoptera" pageId="24" pageNumber="25" phylum="Arthropoda" rank="species" species="dominicensis">
<emphasis id="5B90EAAAFFA26E182B98FBCA6BD02E57" box="[386,670,1117,1143]" italics="true" pageId="24" pageNumber="25">Hemipeplus dominicensis</emphasis>
</taxonomicName>
has the following diagnostic features: temples absent; scape elongate, widened apically; eyes large (as long as scape + pedicel), moderately wide and moderately convex; pronotum subcordiform with an indistinct but broad anterior emargination; anterolateral angles slightly produced, broadly rounded; pronotal pad present but less distinct; posterolateral angles obtusely rounded; pronotal lobe brief with indistinct median notch; scutellar shield subhexagonal, transverse, about twice as long as wide; elytra flat; color uniformly rufotestaceous, dull; distribution:
<collectingCountry id="11F37628FFA26E182892FA866A3B2F0B" box="[648,885,1297,1323]" name="Dominican Republic" pageId="24" pageNumber="25">Dominican Republic</collectingCountry>
.
</paragraph>
<paragraph id="695B36B8FFA26E182ADDFAA26D182F9B" blockId="24.[151,1437,829,1576]" pageId="24" pageNumber="25">
<emphasis id="5B90EAAAFFA26E182ADDFAA26B542F6F" bold="true" box="[199,538,1333,1359]" pageId="24" pageNumber="25">Male Genitalia (Figs 7DF)</emphasis>
. (Tegmen length =
<quantity id="AE1C9B5DFFA26E18291DFAA26A212F70" box="[775,879,1333,1360]" metricMagnitude="-4" metricUnit="m" metricValue="4.2" pageId="24" pageNumber="25" unit="mm" value="0.42">0.42 mm</quantity>
(n = 1)) Lobe of basale short, sharp, and thin; shoulders acutely pointed with an upward slant; apicale and basale without distinct suture; parameres sinuate with pointed apices; inner margin of parameres with a notch near halfway; penis subequal in length to tegmen; parameres laterally surrounded by sheath-like structures (hereafter referred to as lateral sheaths).
</paragraph>
</subSubSection>
<subSubSection id="21FE6533FFA26E182ADDFA526A992C07" pageId="24" pageNumber="25" type="discussion">
<paragraph id="695B36B8FFA26E182ADDFA526D4E2C24" blockId="24.[151,1437,829,1576]" pageId="24" pageNumber="25">
<emphasis id="5B90EAAAFFA26E182ADDFA52687A2FFF" bold="true" box="[199,308,1477,1503]" pageId="24" pageNumber="25">Remarks</emphasis>
.
<taxonomicName id="AEE44D3BFFA26E182B5BFA526B112FFF" authorityName="Kc &amp; Pollock" authorityYear="2025" box="[321,607,1477,1503]" class="Insecta" family="Mycteridae" genus="Hemipeplus" kingdom="Animalia" order="Coleoptera" pageId="24" pageNumber="25" phylum="Arthropoda" rank="species" species="dominicensis">
<emphasis id="5B90EAAAFFA26E182B5BFA526B112FFF" box="[321,607,1477,1503]" italics="true" pageId="24" pageNumber="25">Hemipeplus dominicensis</emphasis>
</taxonomicName>
is closely related to
<taxonomicName id="AEE44D3BFFA26E182958FA516AEA2FFF" authorityName="Thomas" authorityYear="1985" box="[834,932,1477,1504]" class="Insecta" family="Mycteridae" genus="Hemipeplus" kingdom="Animalia" order="Coleoptera" pageId="24" pageNumber="25" phylum="Arthropoda" rank="species" species="chaos">
<emphasis id="5B90EAAAFFA26E182958FA516A182FFF" box="[834,854,1478,1503]" italics="true" pageId="24" pageNumber="25">H</emphasis>
.
<emphasis id="5B90EAAAFFA26E182979FA526AEA2FFF" box="[867,932,1477,1503]" italics="true" pageId="24" pageNumber="25">chaos</emphasis>
</taxonomicName>
(
<collectingCountry id="11F37628FFA26E1829A9FA526AA12FFF" box="[947,1007,1477,1503]" name="United States of America" pageId="24" pageNumber="25">USA</collectingCountry>
,
<collectingCountry id="11F37628FFA26E1829E3FA526D1E2FFF" box="[1017,1104,1477,1503]" name="Mexico" pageId="24" pageNumber="25">Mexico</collectingCountry>
) and
<taxonomicName id="AEE44D3BFFA26E182E97FA516C5E2FFF" authorityName="Grouvelle" authorityYear="1896" box="[1165,1296,1477,1504]" class="Insecta" family="Mycteridae" genus="Hemipeplus" kingdom="Animalia" order="Coleoptera" pageId="24" pageNumber="25" phylum="Arthropoda" rank="species" species="insularis">
<emphasis id="5B90EAAAFFA26E182E97FA516DEF2FFF" box="[1165,1185,1478,1503]" italics="true" pageId="24" pageNumber="25">H</emphasis>
.
<emphasis id="5B90EAAAFFA26E182EB4FA526C5E2FFF" box="[1198,1296,1477,1503]" italics="true" pageId="24" pageNumber="25">insularis</emphasis>
</taxonomicName>
(
<collectingCountry id="11F37628FFA26E182F04FA5269B02C23" name="Dominican Republic" pageId="24" pageNumber="25">Dominican Republic</collectingCountry>
) owing to the presence of lateral sheaths surrounding the parameres.
</paragraph>
<paragraph id="695B36B8FFA26E182ADDF99A6A992C07" blockId="24.[151,1437,829,1576]" box="[199,983,1549,1576]" pageId="24" pageNumber="25">
<emphasis id="5B90EAAAFFA26E182ADDF99A6B742C07" bold="true" box="[199,570,1549,1575]" pageId="24" pageNumber="25">Geographical Distribution (Fig</emphasis>
.
<emphasis id="5B90EAAAFFA26E18285DF99A6B222C07" bold="true" box="[583,620,1549,1575]" pageId="24" pageNumber="25">61)</emphasis>
. Southern
<collectingCountry id="11F37628FFA26E1828FDF99A6A9D2C07" box="[743,979,1549,1575]" name="Dominican Republic" pageId="24" pageNumber="25">Dominican Republic</collectingCountry>
.
</paragraph>
</subSubSection>
</treatment>
</document>