181 lines
20 KiB
XML
181 lines
20 KiB
XML
<document id="C1E62558A9E73EB9D5162D4D16F7AE82" ID-DOI="10.11646/zootaxa.5574.1.1" ID-ISSN="1175-5334" ID-Zenodo-Dep="14746103" ID-ZooBank="DA62BE83-26E8-487F-985A-1D48C3853025" IM.bibliography_approvedBy="felipe" IM.illustrations_approvedBy="felipe" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" IM.treatmentCitations_approvedBy="felipe" IM.treatments_approvedBy="felipe" checkinTime="1737969833979" checkinUser="plazi" docAuthor="Kc, Sajan & Pollock, Darren A." docDate="2025" docId="E14D87AEFF846E3F2A8DFF006A462841" docLanguage="en" docName="zootaxa.5574.1.1.pdf" docOrigin="Zootaxa 5574 (1)" docSource="https://doi.org/10.11646/zootaxa.5574.1.1" docStyle="DocumentStyle:5EBBA59367AD13919D70D935FA04F6A3.19:Zootaxa.2013-.monograph" docStyleId="5EBBA59367AD13919D70D935FA04F6A3" docStyleName="Zootaxa.2013-.monograph" docStyleVersion="19" docTitle="Hemipeplus madagascariensis Grouvelle & NOT 1906" docType="treatment" docVersion="2" lastPageNumber="64" masterDocId="1D74FFD6FFBA6E002A1AFF97694E2A20" masterDocTitle="Review of the Hemipeplinae (Coleoptera: Mycteridae) fauna of the world with descriptions of twenty-nine new species" masterLastPageNumber="140" masterPageNumber="1" pageNumber="63" updateTime="1737970979626" updateUser="ExternalLinkService">
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<mods:title id="93ABCB9DEFD0E9B29424F2F8799CD294">Review of the Hemipeplinae (Coleoptera: Mycteridae) fauna of the world with descriptions of twenty-nine new species</mods:title>
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<mods:namePart id="31A6DDB137EC93BA60B5F741ECF14275">Kc, Sajan</mods:namePart>
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<mods:affiliation id="3462A4DB698447C242DE0BBF6A81F848">Department of Biology, Eastern New Mexico University, Portales, NM 88130 USA & Corresponding author: Sajankc 143 @ gmail. com; https: // orcid. org / 0000 - 0002 - 2749 - 0738</mods:affiliation>
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<mods:nameIdentifier id="903D587E6C3E31D7D17C7E7D849EC4C8" type="email">Sajankc143@gmail.com</mods:nameIdentifier>
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<mods:namePart id="D1F9C16AA3BF9FD042E444B52360B648">Pollock, Darren A.</mods:namePart>
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<mods:affiliation id="BEEA1F240EF871278F59FBB9BE89FD79">Department of Biology, Eastern New Mexico University, Portales, NM 88130 USA & Darren. pollock @ enmu. edu; https: // orcid. org / 0000 - 0002 - 8709 - 0269 & Department of Biology, Eastern New Mexico University, Portales, NM 88130 USA & Corresponding author: Sajankc 143 @ gmail. com; https: // orcid. org / 0000 - 0002 - 2749 - 0738</mods:affiliation>
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<mods:nameIdentifier id="36731F0CAF953CA7885F317D509FA9FB" type="email">Darren.pollock@enmu.edu</mods:nameIdentifier>
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<mods:title id="D13B195F0DF1E9A2D033F716B8AFBED0">Zootaxa</mods:title>
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<mods:date id="C518A8E5C21BF3E61B9879384E37B81B">2025</mods:date>
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<mods:number id="567FDC747AF552D0BF936F5BD5EBC673">5574</mods:number>
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<treatment id="E14D87AEFF846E3F2A8DFF006A462841" ID-DOI="http://doi.org/10.5281/zenodo.14746032" ID-Zenodo-Dep="14746032" LSID="urn:lsid:plazi:treatment:E14D87AEFF846E3F2A8DFF006A462841" httpUri="http://treatment.plazi.org/id/E14D87AEFF846E3F2A8DFF006A462841" lastPageId="63" lastPageNumber="64" pageId="62" pageNumber="63">
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<subSubSection id="21FE6533FF846E3E2A8DFF0068692AF5" pageId="62" pageNumber="63" type="nomenclature">
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<paragraph id="695B36B8FF846E3E2A8DFF006B9D2A91" blockId="62.[151,723,151,213]" box="[151,723,151,178]" pageId="62" pageNumber="63">
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<heading id="321381D4FF846E3E2A8DFF006B9D2A91" bold="true" box="[151,723,151,178]" fontSize="11" level="2" pageId="62" pageNumber="63" reason="4">
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<taxonomicName id="AEE44D3BFF846E3E2A8DFF006B9D2A91" authority="Grouvelle, 1906" authorityName="Grouvelle & NOT" authorityYear="1906" box="[151,723,151,178]" class="Insecta" family="Mycteridae" genus="Hemipeplus" kingdom="Animalia" order="Coleoptera" pageId="62" pageNumber="63" phylum="Arthropoda" rank="species" species="madagascariensis">
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<emphasis id="5B90EAAAFF846E3E2A8DFF006B9D2A91" bold="true" box="[151,723,151,178]" pageId="62" pageNumber="63">
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<emphasis id="5B90EAAAFF846E3E2A8DFF006B492A92" bold="true" box="[151,519,151,178]" italics="true" pageId="62" pageNumber="63">Hemipeplus madagascariensis</emphasis>
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<bibRefCitation id="0D754B49FF846E3E2814FF006B9D2A91" author="Grouvelle, A." box="[526,723,151,177]" pageId="62" pageNumber="63" pagination="67 - 168" refId="ref83137" refString="Grouvelle, A. (1906) Contribution a l'etude des coleopteres de Madagascar. Nitidulidae, Colydiidae, Cucujidae, Monotomidae, Cryptophagidae, Mycetophagidae, Dryopidae, Heteroceridae. Annales de la Societe Entomologique de France, 75 (1), 67 - 168. https: // doi. org / 10.1080 / 21686351.1906.12279361" type="journal article" year="1906">Grouvelle, 1906</bibRefCitation>
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</emphasis>
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||
</taxonomicName>
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||
</heading>
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</paragraph>
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<paragraph id="695B36B8FF846E3E2A8DFF2C68692AF5" blockId="62.[151,723,151,213]" box="[151,295,187,213]" pageId="62" pageNumber="63">(Figs 29, 63)</paragraph>
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</subSubSection>
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<subSubSection id="21FE6533FF846E3E2A8DFF686DEA2DF9" pageId="62" pageNumber="63" type="reference_group">
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<paragraph id="695B36B8FF846E3E2A8DFF6868162B16" blockId="62.[151,1437,255,310]" pageId="62" pageNumber="63">
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<treatmentCitationGroup id="49F41196FF846E3E2A8DFF6868162B16" pageId="62" pageNumber="63">
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<treatmentCitation id="E84510A9FF846E3E2A8DFF686BFC2B37" author="Grouvelle, A." box="[151,690,255,279]" page="116" pageId="62" pageNumber="63" year="1906">
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<taxonomicName id="AEE44D3BFF846E3E2A8DFF686BFC2B37" authority="Grouvelle, 1906: 116" authorityName="Grouvelle & NOT" authorityPageNumber="116" authorityYear="1906" box="[151,690,255,279]" class="Insecta" family="Mycteridae" genus="Hemipeplus" kingdom="Animalia" order="Coleoptera" pageId="62" pageNumber="63" phylum="Arthropoda" rank="species" species="madagascariensis">
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<emphasis id="5B90EAAAFF846E3E2A8DFF68689B2B37" bold="true" box="[151,469,255,279]" italics="true" pageId="62" pageNumber="63">Hemipeplus madagascariensis</emphasis>
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<bibRefCitation id="0D754B49FF846E3E2BC6FF686BFC2B37" author="Grouvelle, A." box="[476,690,255,279]" pageId="62" pageNumber="63" pagination="67 - 168" refId="ref83137" refString="Grouvelle, A. (1906) Contribution a l'etude des coleopteres de Madagascar. Nitidulidae, Colydiidae, Cucujidae, Monotomidae, Cryptophagidae, Mycetophagidae, Dryopidae, Heteroceridae. Annales de la Societe Entomologique de France, 75 (1), 67 - 168. https: // doi. org / 10.1080 / 21686351.1906.12279361" type="journal article" year="1906">Grouvelle, 1906: 116</bibRefCitation>
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</taxonomicName>
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||
</treatmentCitation>
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.—
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<treatmentCitation id="E84510A9FF846E3E28CAFF686AD82B38" author="Hetschko, A." box="[720,918,255,280]" page="90" pageId="62" pageNumber="63" year="1930">
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<bibRefCitation id="0D754B49FF846E3E28CAFF686AD82B38" author="Hetschko, A." box="[720,918,255,280]" pageId="62" pageNumber="63" pagination="1 - 22" refId="ref83335" refString="Hetschko, A. (1930) Cucujidae. In: Schenkling, S. (Ed.), Coleopterorum Catalogus. Vol. 15. Pt. 109. W. Junk, Berlin, pp. 1 - 22." type="book chapter" year="1930">Hetschko 1930: 90</bibRefCitation>
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</treatmentCitation>
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;
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<treatmentCitation id="E84510A9FF846E3E29BBFE976D0B2B37" author="Scott, H." box="[929,1093,255,280]" page="607" pageId="62" pageNumber="63" year="1933">
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<bibRefCitation id="0D754B49FF846E3E29BBFE976D0B2B37" author="Scott, H." box="[929,1093,255,280]" pageId="62" pageNumber="63" pagination="595 - 611" refId="ref84778" refString="Scott, H. (1933) The systematic position of the family Hemipeplidae (Coleoptera), and descriptions of new species. Annals and Magazine of Natural History, 12 (72), 595 - 611. https: // doi. org / 10.1080 / 00222933308673726" type="journal article" year="1933">Scott 1933: 607</bibRefCitation>
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</treatmentCitation>
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||
;
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||
<treatmentCitation id="E84510A9FF846E3E2E4AFF686C4D2B38" author="Kamiya, H." box="[1104,1283,255,280]" page="14" pageId="62" pageNumber="63" year="1963">
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<bibRefCitation id="0D754B49FF846E3E2E4AFF686C4D2B38" author="Kamiya, H." box="[1104,1283,255,280]" pageId="62" pageNumber="63" pagination="11 - 16" refId="ref83597" refString="Kamiya, H. (1963) On the family Hemipeplidae (Coleoptera). Life Study, Fukui, 7, 11 - 16." type="journal article" year="1963">Kamiya 1963: 14</bibRefCitation>
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||
</treatmentCitation>
|
||
;
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||
<treatmentCitation id="E84510A9FF846E3E2F14FF6869AC2B16" author="Pollock, D. A." page="86" pageId="62" pageNumber="63" year="1997">
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||
<bibRefCitation id="0D754B49FF846E3E2F14FF6869AC2B16" author="Pollock, D. A." pageId="62" pageNumber="63" pagination="79 - 92" refId="ref84398" refString="Pollock, D. A. (1997) Review of the Afrotropical species of Hemipeplus Latreille (Coleoptera: Mycteridae: Hemipeplinae). Annales Zoologici, 47 (1 / 2), 79 - 92." type="journal article" year="1997">Pollock 1997: 86</bibRefCitation>
|
||
</treatmentCitation>
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||
, Figs 2, 13.
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||
</treatmentCitationGroup>
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||
</paragraph>
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||
<paragraph id="695B36B8FF846E3E2A82F80D6DEA2DF9" blockId="62.[151,1436,1946,2009]" pageId="62" pageNumber="63">
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||
<emphasis id="5B90EAAAFF846E3E2A82F80D68632D95" bold="true" box="[152,301,1946,1973]" pageId="62" pageNumber="63">FIGURE 29</emphasis>
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||
.
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<emphasis id="5B90EAAAFF846E3E2B21F80D6AAB2D94" bold="true" box="[315,997,1946,1973]" pageId="62" pageNumber="63">
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||
<taxonomicName id="AEE44D3BFF846E3E2B21F80D6A132D95" authority="Grouvelle, 1906" authorityName="Grouvelle & NOT" authorityYear="1906" box="[315,861,1946,1973]" class="Insecta" family="Mycteridae" genus="Hemipeplus" kingdom="Animalia" order="Coleoptera" pageId="62" pageNumber="63" phylum="Arthropoda" rank="species" species="madagascariensis">
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||
<emphasis id="5B90EAAAFF846E3E2B21F80D6BD42D95" bold="true" box="[315,666,1946,1973]" italics="true" pageId="62" pageNumber="63">Hemipeplus madagascariensis</emphasis>
|
||
<bibRefCitation id="0D754B49FF846E3E28B8F80D6A132D95" author="Grouvelle, A." box="[674,861,1946,1973]" pageId="62" pageNumber="63" pagination="67 - 168" refId="ref83137" refString="Grouvelle, A. (1906) Contribution a l'etude des coleopteres de Madagascar. Nitidulidae, Colydiidae, Cucujidae, Monotomidae, Cryptophagidae, Mycetophagidae, Dryopidae, Heteroceridae. Annales de la Societe Entomologique de France, 75 (1), 67 - 168. https: // doi. org / 10.1080 / 21686351.1906.12279361" type="journal article" year="1906">Grouvelle, 1906</bibRefCitation>
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</taxonomicName>
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||
(
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<typeStatus id="B65F881AFF846E3E2976F80D6A922D94" box="[876,988,1946,1972]" pageId="62" pageNumber="63" type="holotype">Holotype</typeStatus>
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||
)
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</emphasis>
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||
. A. Dorsal habitus; B. Dorsal head and prothorax; C. Lateral view of head and prothorax.
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<emphasis id="5B90EAAAFF846E3E28D6F8286A242DF8" bold="true" box="[716,874,1982,2009]" pageId="62" pageNumber="63">Image Credit</emphasis>
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||
.
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||
<collectingRegion id="AB20F85AFF846E3E2963F8286A802DF9" box="[889,974,1983,2009]" country="Ireland" name="Monaghan" pageId="62" pageNumber="63">MNHN</collectingRegion>
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||
/Christophe
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<collectingRegion id="AB20F85AFF846E3E2E42F8286DEE2DF9" box="[1112,1184,1983,2009]" country="Nigeria" name="Rivers" pageId="62" pageNumber="63">Rivier</collectingRegion>
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||
.
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||
</paragraph>
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||
</subSubSection>
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||
<subSubSection id="21FE6533FF856E3F2A8DFF006C322B3D" pageId="63" pageNumber="64" type="materials_examined">
|
||
<paragraph id="695B36B8FF856E3F2A8DFF006C322B3D" blockId="63.[151,1437,150,609]" pageId="63" pageNumber="64">
|
||
<emphasis id="5B90EAAAFF856E3F2A8DFF0068702A90" bold="true" box="[151,318,150,177]" pageId="63" pageNumber="64">Type Material</emphasis>
|
||
.
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||
<materialsCitation id="D98C3CE5FF856E3F2B50FF006BE62AD9" collectionCode="TYPE, MUSEUM" location="Grouv" pageId="63" pageNumber="64" specimenCode="PARIS 1917" specimenCount="1" stateProvince="Monaghan" typeStatus="holotype">
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||
<typeStatus id="B65F881AFF856E3F2B50FF0068FB2A91" box="[330,437,151,177]" pageId="63" pageNumber="64" type="holotype">Holotype</typeStatus>
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||
, sex unknown, only images sent from
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||
<collectingRegion id="AB20F85AFF856E3F2949FF006AE52A91" box="[851,939,151,177]" country="Ireland" name="Monaghan" pageId="63" pageNumber="64">MNHN</collectingRegion>
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||
, labeled: ‘[light green blue label] Madagasc. / [in Grouvelle’s handwriting]
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||
<taxonomicName id="AEE44D3BFF856E3F2BC2FF2C6A662AF5" authorityName="Grouvelle & NOT" authorityYear="1906" box="[472,808,187,213]" class="Insecta" family="Mycteridae" genus="Hemipeplus" kingdom="Animalia" order="Coleoptera" pageId="63" pageNumber="64" phylum="Arthropoda" rank="species" species="madagascariensis">Hemipeplus madagascariensis</taxonomicName>
|
||
5
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||
<location id="6C3B6063FF856E3F295AFF2C6AC62AF5" LSID="urn:lsid:plazi:treatment:E14D87AEFF846E3F2A8DFF006A462841:6C3B6063FF856E3F295AFF2C6AC62AF5" box="[832,904,187,213]" name="Grouv" pageId="63" pageNumber="64" stateProvince="Monaghan">Grouv</location>
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||
/ [red label]
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||
<collectionCode id="0FF5AE7DFF856E3F2E15FF2C6D1B2AF5" box="[1039,1109,187,213]" pageId="63" pageNumber="64">TYPE</collectionCode>
|
||
/ [light blue label]
|
||
<collectionCode id="0FF5AE7DFF856E3F2F3BFF2C6CD22AF5" box="[1313,1436,187,213]" pageId="63" pageNumber="64">MUSEUM</collectionCode>
|
||
<specimenCode id="39429EC3FF856E3F2A8DFF48686B2AD9" box="[151,293,223,249]" collectionCode="PARIS" pageId="63" pageNumber="64">PARIS 1917</specimenCode>
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||
Coll. GROUVELLE / [red label]
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||
</materialsCitation>
|
||
<materialsCitation id="D98C3CE5FF856E3F28A8FF486C362B3D" collectorName="Paris EC" location="MNHN" pageId="63" pageNumber="64" specimenCount="1" stateProvince="Monaghan" typeStatus="holotype">
|
||
<typeStatus id="B65F881AFF856E3F28A8FF486A082AD9" box="[690,838,223,249]" pageId="63" pageNumber="64" type="holotype">HOLOTYPE</typeStatus>
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||
[handwritten]
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||
<taxonomicName id="AEE44D3BFF856E3F29EBFF486CD32AD9" authority="Grouv." authorityName="Grouvelle & NOT" authorityYear="1906" box="[1009,1437,223,249]" class="Insecta" family="Mycteridae" genus="Hemipeplus" kingdom="Animalia" order="Coleoptera" pageId="63" pageNumber="64" phylum="Arthropoda" rank="species" species="madagascariensis">Hemipeplus madagascariensis Grouv.</taxonomicName>
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||
Examined by D.A.
|
||
<bibRefCitation id="0D754B49FF856E3F2B6AFE946B4B2B3D" author="Pollock, D. A." box="[368,517,259,285]" pageId="63" pageNumber="64" pagination="79 - 92" refId="ref84398" refString="Pollock, D. A. (1997) Review of the Afrotropical species of Hemipeplus Latreille (Coleoptera: Mycteridae: Hemipeplinae). Annales Zoologici, 47 (1 / 2), 79 - 92." type="journal article" year="1997">Pollock 1997</bibRefCitation>
|
||
/ [with a barcode]
|
||
<collectingRegion id="AB20F85AFF856E3F28C3FE946A7F2B3D" box="[729,817,259,285]" country="Ireland" name="Monaghan" pageId="63" pageNumber="64">MNHN</collectingRegion>
|
||
,
|
||
<collectorName id="C411536EFF856E3F2927FE946AEC2B3D" box="[829,930,259,285]" pageId="63" pageNumber="64">Paris EC</collectorName>
|
||
26369’, in
|
||
<collectingRegion id="AB20F85AFF856E3F2E03FE946D212B3D" box="[1049,1135,259,285]" country="Ireland" name="Monaghan" pageId="63" pageNumber="64">MNHN</collectingRegion>
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||
(Grouvelle Collection)
|
||
</materialsCitation>
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||
.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="21FE6533FF856E3F2ADDFEB16A462841" pageId="63" pageNumber="64" type="diagnosis">
|
||
<paragraph id="695B36B8FF856E3F2ADDFEB16D25281D" blockId="63.[151,1437,150,609]" pageId="63" pageNumber="64">
|
||
<emphasis id="5B90EAAAFF856E3F2ADDFEB168742B60" bold="true" box="[199,314,294,320]" pageId="63" pageNumber="64">Diagnosis</emphasis>
|
||
.
|
||
<taxonomicName id="AEE44D3BFF856E3F2B50FEB06BEC2B61" authorityName="Grouvelle & NOT" authorityYear="1906" box="[330,674,295,321]" class="Insecta" family="Mycteridae" genus="Hemipeplus" kingdom="Animalia" order="Coleoptera" pageId="63" pageNumber="64" phylum="Arthropoda" rank="species" species="madagascariensis">
|
||
<emphasis id="5B90EAAAFF856E3F2B50FEB06BEC2B61" box="[330,674,295,321]" italics="true" pageId="63" pageNumber="64">Hemipeplus madagascariensis</emphasis>
|
||
</taxonomicName>
|
||
can be distinguished by its large eyes, large, rounded temples, and distinct pronotal lobe.
|
||
<taxonomicName id="AEE44D3BFF856E3F2BBAFEDC6BB52B45" authorityName="Grouvelle & NOT" authorityYear="1906" box="[416,763,331,357]" class="Insecta" family="Mycteridae" genus="Hemipeplus" kingdom="Animalia" order="Coleoptera" pageId="63" pageNumber="64" phylum="Arthropoda" rank="species" species="madagascariensis">
|
||
<emphasis id="5B90EAAAFF856E3F2BBAFEDC6BB52B45" box="[416,763,331,357]" italics="true" pageId="63" pageNumber="64">Hemipeplus madagascariensis</emphasis>
|
||
</taxonomicName>
|
||
has the following diagnostic features: temples very large (about 1/3 eye length) and rounded; scape moniliform externally produced on one side; eyes very large (subequal to scape + pedicel + antennomere III), very wide and convex; pronotum subcordiform, wide and flat without distinct anterior emargination; pronotum strongly sinuate anterolaterally, anterior half strongly produced and wider than posterior half; anterolateral angles narrowly rounded; pronotal pad present; posterolateral angles rounded; pronotal lobe broad and rounded without median notch; scutellar shield slightly wider than long, shield-shaped; elytra slightly convex apically; color uniformly light brown; distribution:
|
||
<collectingCountry id="11F37628FF856E3F2935FDB46AF8281D" box="[815,950,547,573]" name="Madagascar" pageId="63" pageNumber="64">Madagascar</collectingCountry>
|
||
(
|
||
<bibRefCitation id="0D754B49FF856E3F29DEFDB46D11281D" author="Pollock, D. A." box="[964,1119,547,573]" pageId="63" pageNumber="64" pagination="79 - 92" refId="ref84398" refString="Pollock, D. A. (1997) Review of the Afrotropical species of Hemipeplus Latreille (Coleoptera: Mycteridae: Hemipeplinae). Annales Zoologici, 47 (1 / 2), 79 - 92." type="journal article" year="1997">Pollock 1997</bibRefCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="695B36B8FF856E3F2ADDFDD16A462841" blockId="63.[151,1437,150,609]" box="[199,776,582,609]" pageId="63" pageNumber="64">
|
||
<emphasis id="5B90EAAAFF856E3F2ADDFDD16B742840" bold="true" box="[199,570,582,608]" pageId="63" pageNumber="64">Geographical Distribution (Fig</emphasis>
|
||
.
|
||
<emphasis id="5B90EAAAFF856E3F285DFDD16B232840" bold="true" box="[583,621,582,608]" pageId="63" pageNumber="64">63)</emphasis>
|
||
.
|
||
<collectingCountry id="11F37628FF856E3F2861FDD06A4A2841" box="[635,772,583,609]" name="Madagascar" pageId="63" pageNumber="64">Madagascar</collectingCountry>
|
||
.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |