377 lines
46 KiB
XML
377 lines
46 KiB
XML
<document id="A0E72E5017014715CD67DAD0734445BA" ID-CLB-Dataset="20422" ID-DOI="10.5852/ejt.2022.798.1669" ID-GBIF-Dataset="31fec661-5dac-4626-856d-4ba4680efee7" ID-ISSN="2118-9773" ID-Zenodo-Dep="6323002" ID-ZooBank="50692D26-A41C-4F85-B207-A6747FD07470" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" IM.treatmentCitations_approvedBy="felipe" checkinTime="1646225219873" checkinUser="felipe" docAuthor="Vagalinski, Boyan, Borissov, Simeon, Bobeva, Aneliya, Canciani, Giacomo & Antić, Dragan Ž." docDate="2022" docId="4A307579CF490D05FD95FB2DFB16F9E2" docLanguage="en" docName="ejt-798_30-69.pdf" docOrigin="European Journal of Taxonomy 798" docSource="http://dx.doi.org/10.5852/ejt.2022.798.1669" docStyle="DocumentStyle:EF2B578F1D15862ADE45B0C07C620911.14:EJT.2018-.journal_article.type1" docStyleId="EF2B578F1D15862ADE45B0C07C620911" docStyleName="EJT.2018-.journal_article.type1" docStyleVersion="14" docTitle="Stygiiulus ausugi Vagalinski & Borissov & Bobeva & Canciani & Antić 2022, comb. nov." docType="treatment" docVersion="11" lastPageNumber="35" masterDocId="B6090D01CF4D0D00FF95FFEAFFB8FF83" masterDocTitle="The mostly cavernicolous millipede genus Stygiiulus Verhoeff, 1929, stat. nov.: taxonomy, distribution and phylogenetic relationships (Diplopoda, Julida, Julidae)" masterLastPageNumber="69" masterPageNumber="30" pageNumber="34" updateTime="1699300459248" updateUser="plazi" zenodo-license-document="CC-BY-4.0" zenodo-license-figures="CC-BY-4.0" zenodo-license-treatments="UNSPECIFIED">
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<mods:title id="8BE11F5360B337D062CD200B5EA24AD4">The mostly cavernicolous millipede genus Stygiiulus Verhoeff, 1929, stat. nov.: taxonomy, distribution and phylogenetic relationships (Diplopoda, Julida, Julidae)</mods:title>
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<mods:namePart id="5AF68C9131875B9A8B78A5392014448C">Vagalinski, Boyan</mods:namePart>
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<mods:affiliation id="D95F860EC5D1E77E9A87196AB9F36EF1">Institute of Biodiversity and Ecosystem Research, Bulgarian Academy of Sciences, 2 Gagarin Street, 1113, Sofia, Bulgaria.</mods:affiliation>
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<mods:nameIdentifier id="436B45FA3FDA62BF2B9A7C48644C450C" type="email">boyanv84@gmail.com</mods:nameIdentifier>
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<mods:namePart id="9251F42461A9158AB9BEC88994A1C461">Borissov, Simeon</mods:namePart>
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<mods:affiliation id="E94211378FF00379B0A202B0ABE3466E">Institute of Biodiversity and Ecosystem Research, Bulgarian Academy of Sciences, 2 Gagarin Street, 1113, Sofia, Bulgaria.</mods:affiliation>
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<mods:nameIdentifier id="836B8EE5185A7CEA5773D04162EDDF34" type="email">borissovsb@gmail.com</mods:nameIdentifier>
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<mods:namePart id="7C3CEE7CC9034C31D303361D04384CAF">Bobeva, Aneliya</mods:namePart>
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<mods:affiliation id="FF45243D1BFEA673F6F92A46859B05F0">Institute of Biodiversity and Ecosystem Research, Bulgarian Academy of Sciences, 2 Gagarin Street, 1113, Sofia, Bulgaria.</mods:affiliation>
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<mods:nameIdentifier id="90A08C0E5548D20814B5D1F53821FD99" type="email">aneliabobeva@gmail.com</mods:nameIdentifier>
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<mods:namePart id="F5168B29D1C94743F766DDF545C8437F">Canciani, Giacomo</mods:namePart>
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<mods:affiliation id="51CDAA2F2609350731D134D8E70193A7">Jamarski Klub Kraški Krti / G. S. Talpe del Carso, loc. San Michele del Carso / Vrh, I- 34070 Savogna d’Isonzo (GO), Italy. & Associazione Speleologica Forum Julii Speleo, Via Carraria 99, I- 33043 Cividale del Friuli (UD), Italy.</mods:affiliation>
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<mods:nameIdentifier id="65A7B32DF072D8D14DA5635B3F05826C" type="email">giacomo.canciani90@gmail.com</mods:nameIdentifier>
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<mods:namePart id="D6557A31B4F8B55ECC0F2879327B90DA">Antić, Dragan Ž.</mods:namePart>
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<mods:affiliation id="2270556680F4AA3CE61DAE0AC217496A">University of Belgrade - Faculty of Biology, Institute of Zoology, Studentski Trg 16, 11000 Belgrade, Serbia. & Serbian Biospeleological Society, Trg Dositeja Obradovića 2, 21000 Novi Sad, Serbia.</mods:affiliation>
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<mods:nameIdentifier id="6C6D6A105E00CF089B356B339716A96F" type="email">dragan.antic@bio.bg.ac.rs</mods:nameIdentifier>
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<mods:title id="6584F36251D122DF587CA19A6BBE3B72">European Journal of Taxonomy</mods:title>
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<mods:date id="23150F22E1489E26A8A39FC7414A1686">2022</mods:date>
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<treatment id="4A307579CF490D05FD95FB2DFB16F9E2" ID-DOI="http://doi.org/10.5281/zenodo.6323848" ID-GBIF-Taxon="193854116" ID-Zenodo-Dep="6323848" LSID="urn:lsid:plazi:treatment:4A307579CF490D05FD95FB2DFB16F9E2" httpUri="http://treatment.plazi.org/id/4A307579CF490D05FD95FB2DFB16F9E2" lastPageId="5" lastPageNumber="35" pageId="4" pageNumber="34">
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<subSubSection id="8A8397E4CF490D04FD95FB2DFB8CFB61" box="[512,1076,1223,1250]" pageId="4" pageNumber="34" type="nomenclature">
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<paragraph id="C226C46FCF490D04FD95FB2DFB8CFB61" blockId="4.[512,1076,1223,1284]" box="[512,1076,1223,1250]" pageId="4" pageNumber="34">
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<heading id="996E7303CF490D04FD95FB2DFB8CFB61" box="[512,1076,1223,1250]" centered="true" fontSize="11" level="2" pageId="4" pageNumber="34" reason="2">
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<taxonomicName id="0599BFECCF490D04FD95FB2DFC11FB61" ID-CoL="8YKQL" authority="(Manfredi, 1953)" authorityName="Vagalinski & Borissov & Bobeva & Canciani & Antić" authorityYear="2022" baseAuthorityName="Manfredi" baseAuthorityYear="1953" box="[512,937,1223,1250]" class="Diplopoda" family="Julidae" genus="Stygiiulus" kingdom="Animalia" order="Julida" pageId="4" pageNumber="34" phylum="Arthropoda" rank="species" species="ausugi" status="comb. nov.">
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<emphasis id="F0ED187DCF490D04FD95FB2DFD6CFB62" bold="true" box="[512,724,1223,1250]" italics="true" pageId="4" pageNumber="34">Stygiiulus ausugi</emphasis>
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(Manfredi, 1953)
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<taxonomicNameLabel id="EBDEA506CF490D04FC25FB2DFB8CFB61" box="[944,1076,1223,1250]" pageId="4" pageNumber="34" rank="species">comb. nov.</taxonomicNameLabel>
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</paragraph>
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<subSubSection id="8A8397E4CF490D04FD39FB00FC30FA87" box="[684,904,1258,1284]" pageId="4" pageNumber="34" type="description">
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<paragraph id="C226C46FCF490D04FD39FB00FC30FA87" blockId="4.[512,1076,1223,1284]" box="[684,904,1258,1284]" pageId="4" pageNumber="34">
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<figureCitation id="5AA2D8EACF490D04FD39FB00FCA5FA87" box="[684,797,1258,1284]" captionStart="Fig" captionStartId="26.[189,232,1822,1848]" captionTargetBox="[197,1384,282,1779]" captionTargetId="figure-12@26.[189,1398,263,1787]" captionTargetPageId="26" captionText="Fig. 10. Left gonopods of species of Stygiiulus stat. nov., mesal views. A. S. ausugi (Manfredi, 1953) comb.nov.,topotype♂(NHMD).B.S.illyricus(Verhoeff,1929)comb.nov.,redrawnfromVerhoeff (1929). C. S. insularis (Strasser, 1938) comb. nov., redrawn from Antić et al. (2018). D. S. maximus (Verhoeff, 1929) comb. nov., redrawn from Verhoeff (1930). E. S. seewaldi (Strasser, 1967) comb. nov., redrawn and modified from Strasser (1967). F. S. tobias (Berlese, 1886) comb. nov., ♂ from Grotta [Cave] del Subiolo (NHMD). Abbreviations: do = distal outgrowth of velum. Drawn not to scale." figureDoi="http://doi.org/10.5281/zenodo.6323028" httpUri="https://zenodo.org/record/6323028/files/figure.png" pageId="4" pageNumber="34">Figs 10A</figureCitation>
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,
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<figureCitation id="5AA2D8EACF490D04FCBCFB00FCE6FA87" box="[809,862,1258,1284]" captionStart="Fig" captionStartId="31.[189,231,1681,1707]" captionTargetBox="[345,1243,265,1635]" captionTargetId="figure-15@31.[325,1262,262,1645]" captionTargetPageId="31" captionText="Fig. 11. Left vulvae of species of Stygiiulus stat. nov. (A–D) and representatives of (other) Julini/ Leptoiulini (E) and Typhloiulini/blind Julini/Leptoiulini in caudal, somewhat mesal views. A. S. ausugi (Manfredi, 1953) comb. nov., ♀ from Grotta [Cave] del Calgeron (NHMD). B. S. illyricus (Verhoef, 1929) comb. nov., ♀ from Rabakova Cave (IZB). C. S. insularis (Strasser, 1938) comb. nov., topotype ♀ (CBSS). D. S. tobias (Berlese, 1886) comb. nov., ♀ from Grotta [Cave] del Subiolo (NHMD). E. Leptoiulus cf. trilineatus(C.L. Koch, 1847), ♀ from Belasitsa Mtn, Bulgaria. F. Serboiulus spelaeophilus Gulička, 1967, ♀ from Vodni Pech Cave (NMNHS). G. Typhloiulus (Inversotyphlus) lobifer Attems, 1951, ♀ from Minjera-Bauxite Mine (IBER). H. Typhloiulus (s. str.) strictus (Latzel, 1882), ♀ from Kunino (IBER). Abbreviations: at = anterior tube; la = lateral ampulla; lt = lateral tube; ma = median ampulla; mt = median tube; op = operculum; pa = posterior ampulla; pt = posterior tube. Scale bar = 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.6323030" httpUri="https://zenodo.org/record/6323030/files/figure.png" pageId="4" pageNumber="34">11A</figureCitation>
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,
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<figureCitation id="5AA2D8EACF490D04FCFFFB00FC30FA87" box="[874,904,1258,1284]" captionStart="Fig" captionStartId="33.[189,232,1998,2024]" captionTargetBox="[189,1398,1099,1960]" captionTargetId="figure-332@33.[189,1398,1099,1960]" captionTargetPageId="33" captionText="Fig. 13. General distribution of the species of the genus Stygiiulus stat. nov." figureDoi="http://doi.org/10.5281/zenodo.6323034" httpUri="https://zenodo.org/record/6323034/files/figure.png" pageId="4" pageNumber="34">13</figureCitation>
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</paragraph>
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<subSubSection id="8A8397E4CF490D04FF28FAC6FC87FA72" pageId="4" pageNumber="34" type="reference_group">
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<paragraph id="C226C46FCF490D04FF28FAC6FCE2FAC5" blockId="4.[189,858,1324,1351]" box="[189,858,1324,1351]" pageId="4" pageNumber="34">
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<treatmentCitationGroup id="E289E341CF490D04FF28FAC6FCE2FAC5" box="[189,858,1324,1351]" pageId="4" pageNumber="34">
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<treatmentCitation id="4338E27ECF490D04FF28FAC6FD50FAC5" author="Manfredi P." box="[189,744,1324,1351]" page="138" pageId="4" pageNumber="34" year="1953">
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<taxonomicName id="0599BFECCF490D04FF28FAC6FD50FAC5" ID-CoL="59WSP" authority="Manfredi, 1953 b: 136 - 138" authorityName="Manfredi" authorityPageNumber="136 - 138" authorityYear="1953" box="[189,744,1324,1351]" class="Diplopoda" family="Julidae" genus="Typhloiulus" kingdom="Animalia" order="Julida" pageId="4" pageNumber="34" phylum="Arthropoda" rank="species" species="ausugi">
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<emphasis id="F0ED187DCF490D04FF28FAC6FE26FAC5" box="[189,414,1324,1350]" italics="true" pageId="4" pageNumber="34">Typhloiulus ausugi</emphasis>
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<bibRefCitation id="A608B99ECF490D04FE33FAC6FD50FAC5" author="Manfredi P." box="[422,744,1324,1351]" pageId="4" pageNumber="34" pagination="117 - 122" refId="ref21673" refString="Manfredi P. 1953 b. Miriapodi italiani - VIII. contributo. Atti della Societa italiana di Scienze naturali, e del Museo civile di Storia naturale 92: 117 - 122." type="journal article" year="1953">Manfredi, 1953b: 136–138</bibRefCitation>
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</taxonomicName>
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</treatmentCitation>
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, figs 1–2.
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</treatmentCitationGroup>
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</paragraph>
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<paragraph id="C226C46FCF490D04FF28FA87FA99FA04" blockId="4.[189,1313,1389,1522]" box="[189,1313,1389,1416]" pageId="4" pageNumber="34">
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<treatmentCitationGroup id="E289E341CF490D04FF28FA87FA99FA04" box="[189,1313,1389,1416]" pageId="4" pageNumber="34">
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<taxonomicName id="0599BFECCF490D04FF28FA87FD97FA04" ID-CoL="59WSP" authorityName="Manfredi" authorityYear="1953" box="[189,559,1389,1416]" class="Diplopoda" family="Julidae" genus="Typhloiulus" kingdom="Animalia" order="Julida" pageId="4" pageNumber="34" phylum="Arthropoda" rank="species" species="ausugi" subGenus="Stygiiulus">
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<emphasis id="F0ED187DCF490D04FF28FA87FEFFFA04" box="[189,327,1389,1415]" italics="true" pageId="4" pageNumber="34">Typhloiulus</emphasis>
|
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(
|
||
<emphasis id="F0ED187DCF490D04FECCFA87FE76FA04" box="[345,462,1389,1415]" italics="true" pageId="4" pageNumber="34">Stygiiulus</emphasis>
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)
|
||
<emphasis id="F0ED187DCF490D04FE75FA84FD97FA04" box="[480,559,1390,1415]" italics="true" pageId="4" pageNumber="34">ausugi</emphasis>
|
||
</taxonomicName>
|
||
–
|
||
<treatmentCitation id="4338E27ECF490D04FDD8FA87FCA5FA04" author="Strasser K." box="[589,797,1389,1416]" page="11" pageId="4" pageNumber="34" year="1962">
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<bibRefCitation id="A608B99ECF490D04FDD8FA87FCA5FA04" author="Strasser K." box="[589,797,1389,1416]" pageId="4" pageNumber="34" pagination="1 - 77" refId="ref22282" refString="Strasser K. 1962. Die Typhloiulini (Diplopoda Symphyognatha). Atti del Museo civico di Storia naturale di Trieste 23 (1): 1 - 77." type="journal article" year="1962">Strasser 1962: 11</bibRefCitation>
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||
</treatmentCitation>
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||
, 12, 18, 37, 38, figs 1–2, 8, 11h, 15, 41–44.
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||
</treatmentCitationGroup>
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</paragraph>
|
||
<paragraph id="C226C46FCF490D04FF28FA7AFC3EFA28" blockId="4.[189,1313,1389,1522]" box="[189,902,1424,1451]" pageId="4" pageNumber="34">
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||
<treatmentCitationGroup id="E289E341CF490D04FF28FA7AFC3EFA28" box="[189,902,1424,1451]" pageId="4" pageNumber="34">
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||
<taxonomicName id="0599BFECCF490D04FF28FA7AFD3DFA29" ID-CoL="59WSP" authority="ausugi" box="[189,645,1424,1451]" class="Diplopoda" family="Julidae" genus="Typhloiulus" kingdom="Animalia" order="Julida" pageId="4" pageNumber="34" phylum="Arthropoda" rank="subSpecies" species="ausugi" subGenus="Stygiiulus" subSpecies="ausugi">
|
||
<emphasis id="F0ED187DCF490D04FF28FA7AFEFFFA29" box="[189,327,1424,1450]" italics="true" pageId="4" pageNumber="34">Typhloiulus</emphasis>
|
||
(
|
||
<emphasis id="F0ED187DCF490D04FECCFA7AFE76FA29" box="[345,462,1424,1450]" italics="true" pageId="4" pageNumber="34">Stygiiulus</emphasis>
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||
)
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<emphasis id="F0ED187DCF490D04FE75FA7BFD3DFA29" box="[480,645,1425,1450]" italics="true" pageId="4" pageNumber="34">ausugi ausugi</emphasis>
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</taxonomicName>
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||
–
|
||
<treatmentCitation id="4338E27ECF490D04FD31FA7BFC3BFA28" author="Strasser K." box="[676,899,1425,1451]" page="13" pageId="4" pageNumber="34" year="1971">
|
||
<bibRefCitation id="A608B99ECF490D04FD31FA7BFC3BFA28" author="Strasser K." box="[676,899,1425,1451]" pageId="4" pageNumber="34" pagination="1 - 21" refId="ref22340" refString="Strasser K. 1971 a. Uber italienische, besonders kavernikole Diplopoden. Memorie del Museo civico di Storia naturale di Verona 19: 1 - 21." type="journal article" year="1971">Strasser 1971a: 13</bibRefCitation>
|
||
</treatmentCitation>
|
||
.
|
||
</treatmentCitationGroup>
|
||
</paragraph>
|
||
<paragraph id="C226C46FCF490D04FF28FA5EFD6CFA4D" blockId="4.[189,1313,1389,1522]" box="[189,724,1459,1486]" pageId="4" pageNumber="34">
|
||
<treatmentCitationGroup id="E289E341CF490D04FF28FA5EFD6CFA4D" box="[189,724,1459,1486]" pageId="4" pageNumber="34">
|
||
<taxonomicName id="0599BFECCF490D04FF28FA5EFE4DFA4E" ID-CoL="59WSP" authority="ausugi" box="[189,501,1460,1486]" class="Diplopoda" family="Julidae" genus="Typhloiulus" kingdom="Animalia" order="Julida" pageId="4" pageNumber="34" phylum="Arthropoda" rank="subSpecies" species="ausugi" subSpecies="ausugi">
|
||
<emphasis id="F0ED187DCF490D04FF28FA5EFE4DFA4E" box="[189,501,1460,1486]" italics="true" pageId="4" pageNumber="34">Typhloiulus ausugi ausugi</emphasis>
|
||
</taxonomicName>
|
||
–
|
||
<treatmentCitation id="4338E27ECF490D04FD86FA5EFD77FA4D" author="Minelli A." box="[531,719,1459,1486]" page="9" pageId="4" pageNumber="34" year="1985">
|
||
<bibRefCitation id="A608B99ECF490D04FD86FA5EFD77FA4D" author="Minelli A." box="[531,719,1459,1486]" pageId="4" pageNumber="34" pagination="1 - 50" refId="ref21769" refString="Minelli A. 1985. Catalogo dei Diplopodi e dei Chilopodi cavernicoli italiani. Memorie del Museo civico di Storia naturale di Verona, Serie 2 (Sezione scienze della Vita) 4: 1 - 50." type="journal article" year="1985">Minelli 1985: 9</bibRefCitation>
|
||
</treatmentCitation>
|
||
.
|
||
</treatmentCitationGroup>
|
||
</paragraph>
|
||
<paragraph id="C226C46FCF490D04FF28FA3DFC87FA72" blockId="4.[189,1313,1389,1522]" box="[189,831,1495,1522]" pageId="4" pageNumber="34">
|
||
<treatmentCitationGroup id="E289E341CF490D04FF28FA3DFC87FA72" box="[189,831,1495,1522]" pageId="4" pageNumber="34">
|
||
<taxonomicName id="0599BFECCF490D04FF28FA3DFE26FA72" ID-CoL="59WSP" authorityName="Manfredi" authorityYear="1953" box="[189,414,1495,1521]" class="Diplopoda" family="Julidae" genus="Typhloiulus" kingdom="Animalia" order="Julida" pageId="4" pageNumber="34" phylum="Arthropoda" rank="species" species="ausugi">
|
||
<emphasis id="F0ED187DCF490D04FF28FA3DFE26FA72" box="[189,414,1495,1521]" italics="true" pageId="4" pageNumber="34">Typhloiulus ausugi</emphasis>
|
||
</taxonomicName>
|
||
–
|
||
<treatmentCitation id="4338E27ECF490D04FE29FA3DFC82FA72" ID-DOI="http://doi.org/10.5281/zenodo.5664796" ID-Zenodo-Dep="5664796" author="Vagalinski B. & Stoev P. & Enghoff H." box="[444,826,1495,1522]" httpUri="http://treatment.plazi.org/id/990B87FC34673236FF437815F8607E97" page="336" pageId="4" pageNumber="34" year="2015">
|
||
<bibRefCitation id="A608B99ECF490D04FE29FA3DFC82FA72" author="Vagalinski B. & Stoev P. & Enghoff H." box="[444,826,1495,1522]" pageId="4" pageNumber="34" pagination="334 - 362" refId="ref22493" refString="Vagalinski B., Stoev P. & Enghoff H. 2015. A review of the millipede genus Typhloiulus Latzel, 1884 (Diplopoda: Julida: Julidae), with a description of three new species from Bulgaria and Greece. Zootaxa 3999 (3): 334 - 362. https: // doi. org / 10.11646 / zootaxa. 3999.3.2" type="journal article" year="2015">
|
||
Vagalinski
|
||
<emphasis id="F0ED187DCF490D04FDD4FA32FDC4FA72" box="[577,636,1495,1521]" italics="true" pageId="4" pageNumber="34">et al.</emphasis>
|
||
2015: 336–337
|
||
</bibRefCitation>
|
||
</treatmentCitation>
|
||
.
|
||
</treatmentCitationGroup>
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="8A8397E4CF490D04FF28F9F2FEC0F96E" pageId="4" pageNumber="34" type="diagnosis">
|
||
<paragraph id="C226C46FCF490D04FF28F9F2FE8FF9B1" blockId="4.[189,311,1560,1586]" box="[189,311,1560,1586]" pageId="4" pageNumber="34">
|
||
<heading id="996E7303CF490D04FF28F9F2FE8FF9B1" bold="true" box="[189,311,1560,1586]" fontSize="11" level="3" pageId="4" pageNumber="34" reason="3">
|
||
<emphasis id="F0ED187DCF490D04FF28F9F2FE8FF9B1" bold="true" box="[189,311,1560,1586]" pageId="4" pageNumber="34">Diagnosis</emphasis>
|
||
</heading>
|
||
</paragraph>
|
||
<paragraph id="C226C46FCF490D04FF28F9ADFEC0F96E" blockId="4.[188,1399,1607,1773]" pageId="4" pageNumber="34">
|
||
One of the three species of
|
||
<taxonomicName id="0599BFECCF490D04FDBDF9ADFD25F9E2" authorityName="Vagalinski & Borissov & Bobeva & Canciani & Antić" authorityYear="2022" box="[552,669,1607,1633]" class="Diplopoda" family="Julidae" genus="Stygiiulus" kingdom="Animalia" order="Julida" pageId="4" pageNumber="34" phylum="Arthropoda" rank="genus" status="stat. nov.">
|
||
<emphasis id="F0ED187DCF490D04FDBDF9ADFD25F9E2" box="[552,669,1607,1633]" italics="true" pageId="4" pageNumber="34">Stygiiulus</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="EBDEA506CF490D04FD3EF9ADFCA4F9E2" box="[683,796,1607,1633]" pageId="4" pageNumber="34">stat. nov.</taxonomicNameLabel>
|
||
with modified mouthparts, the other two being
|
||
<taxonomicName id="0599BFECCF490D04FF28F980FEECF907" authorityName="Vagalinski & Borissov & Bobeva & Canciani & Antić" authorityYear="2022" baseAuthorityName="Strasser" baseAuthorityYear="1971" box="[189,340,1642,1668]" class="Diplopoda" family="Julidae" genus="Stygiiulus" kingdom="Animalia" order="Julida" pageId="4" pageNumber="34" phylum="Arthropoda" rank="species" species="fimbriatus" status="comb. et stat. nov.">
|
||
<emphasis id="F0ED187DCF490D04FF28F980FEECF907" box="[189,340,1642,1668]" italics="true" pageId="4" pageNumber="34">S. fimbriatus</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="EBDEA506CF490D04FEC8F980FD81F906" box="[349,569,1642,1669]" pageId="4" pageNumber="34">comb. et stat. nov.</taxonomicNameLabel>
|
||
and
|
||
<taxonomicName id="0599BFECCF490D04FDE3F980FCB1F907" authorityName="Vagalinski & Borissov & Bobeva & Canciani & Antić" authorityYear="2022" baseAuthorityName="Strasser" baseAuthorityYear="1971" box="[630,777,1642,1668]" class="Diplopoda" family="Julidae" genus="Stygiiulus" kingdom="Animalia" order="Julida" pageId="4" pageNumber="34" phylum="Arthropoda" rank="species" species="gentianae" status="comb. et stat. nov.">
|
||
<emphasis id="F0ED187DCF490D04FDE3F980FCB1F907" box="[630,777,1642,1668]" italics="true" pageId="4" pageNumber="34">S. gentianae</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="EBDEA506CF490D04FC84F980FC55F906" box="[785,1005,1642,1669]" pageId="4" pageNumber="34">comb. et stat. nov.</taxonomicNameLabel>
|
||
Differs from both mainly by the complete absence of posterior hump on opisthomere, the very large velum with minute fringes on posterodistal margin, and the anterior and posterior solenomeral branches both being very short, hardly distinguishable.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="8A8397E4CF490D04FF28F8FFFB64F84B" pageId="4" pageNumber="34" type="materials_examined">
|
||
<paragraph id="C226C46FCF490D04FF28F8FFFE15F8AC" blockId="4.[189,429,1813,1839]" box="[189,429,1813,1839]" pageId="4" pageNumber="34">
|
||
<heading id="996E7303CF490D04FF28F8FFFE15F8AC" bold="true" box="[189,429,1813,1839]" fontSize="11" level="3" pageId="4" pageNumber="34" reason="3">
|
||
<emphasis id="F0ED187DCF490D04FF28F8FFFE15F8AC" bold="true" box="[189,429,1813,1839]" pageId="4" pageNumber="34">Material examined</emphasis>
|
||
</heading>
|
||
</paragraph>
|
||
<paragraph id="C226C46FCF490D04FF28F8AEFB64F84B" blockId="4.[187,1398,1859,1992]" pageId="4" pageNumber="34">
|
||
<materialsCitation id="72F1CE32CF490D04FF28F8AEFDA5F827" ID-GBIF-Occurrence="3504034332" collectingDate="1996-03-10" collectionCode="NHMD" collectorName="della Bigonda & G. Peretto & E. Piva" country="Italy" county="Altopiano" determinerName="H. Enghoff" elevation="450" location="Trentino" municipality="Sette Comuni" pageId="4" pageNumber="34" specimenCount="1" specimenCount-male="1" stateProvince="Autonomous Province of Trento">
|
||
<collectingCountry id="BA8E84FFCF490D04FF28F8AEFEB7F8DD" box="[189,271,1860,1886]" name="Italy" pageId="4" pageNumber="34">ITALY</collectingCountry>
|
||
–
|
||
<emphasis id="F0ED187DCF490D04FEA7F8A9FCE8F8DE" bold="true" box="[306,848,1859,1885]" pageId="4" pageNumber="34">
|
||
<location id="C74692B4CF490D04FEA7F8A9FE26F8DE" LSID="urn:lsid:plazi:treatment:4A307579CF490D05FD95FB2DFB16F9E2:C74692B4CF490D04FEA7F8A9FE26F8DE" box="[306,414,1859,1885]" country="Italy" county="Altopiano" municipality="Sette Comuni" name="Trentino" pageId="4" pageNumber="34" stateProvince="Autonomous Province of Trento">Trentino</location>
|
||
(
|
||
<collectingRegion id="005D0A8DCF490D04FE3AF8A9FCF1F8DE" box="[431,841,1859,1885]" pageId="4" pageNumber="34">Autonomous Province of Trento</collectingRegion>
|
||
)
|
||
</emphasis>
|
||
•
|
||
<specimenCount id="D49F0FE6CF490D04FCFEF8AEFC24F8DD" box="[875,924,1860,1886]" pageId="4" pageNumber="34" type="male">1 ♂</specimenCount>
|
||
; topotype;
|
||
<location id="C74692B4CF490D04FBB4F8A9FACBF8DE" LSID="urn:lsid:plazi:treatment:4A307579CF490D05FD95FB2DFB16F9E2:C74692B4CF490D04FBB4F8A9FACBF8DE" box="[1057,1395,1859,1886]" country="Italy" county="Altopiano" municipality="Sette Comuni" name="Altopiano dei Sette Comuni" pageId="4" pageNumber="34" stateProvince="Autonomous Province of Trento">Altopiano dei Sette Comuni</location>
|
||
,
|
||
<location id="C74692B4CF490D04FF28F88DFEABF802" LSID="urn:lsid:plazi:treatment:4A307579CF490D05FD95FB2DFB16F9E2:C74692B4CF490D04FF28F88DFEABF802" box="[189,275,1895,1921]" country="Italy" county="Altopiano" municipality="Sette Comuni" name="Grigno" pageId="4" pageNumber="34" stateProvince="Autonomous Province of Trento">Grigno</location>
|
||
,
|
||
<location id="C74692B4CF490D04FE89F88DFEDFF802" LSID="urn:lsid:plazi:treatment:4A307579CF490D05FD95FB2DFB16F9E2:C74692B4CF490D04FE89F88DFEDFF802" box="[284,359,1895,1921]" country="Italy" county="Altopiano" municipality="Sette Comuni" name="Grotta" pageId="4" pageNumber="34" stateProvince="Autonomous Province of Trento">Grotta</location>
|
||
[Cave]
|
||
<collectorName id="6F6CA1B9CF490D04FE54F88DFDDAF802" box="[449,610,1895,1921]" pageId="4" pageNumber="34">della Bigonda</collectorName>
|
||
(243 VT/TN);
|
||
<elevation id="49B4235CCF490D04FC9BF88DFC2BF802" box="[782,915,1895,1922]" metricMagnitude="2" metricUnit="m" metricValue="4.5" pageId="4" pageNumber="34" unit="m" value="450.0">
|
||
<quantity id="0561698ACF490D04FC9BF88DFCEEF802" box="[782,854,1895,1922]" metricMagnitude="2" metricUnit="m" metricValue="4.5" pageId="4" pageNumber="34" unit="m" value="450.0">450 m</quantity>
|
||
a.s.l.
|
||
</elevation>
|
||
;
|
||
<date id="B627E2AFCF490D04FC0BF88DFB82F802" box="[926,1082,1895,1921]" pageId="4" pageNumber="34" value="1996-03-10">
|
||
<collectingDate id="A6631B47CF490D04FC0BF88DFB82F802" box="[926,1082,1895,1921]" pageId="4" pageNumber="34" value="1996-03-10">10 Mar. 1996</collectingDate>
|
||
</date>
|
||
;
|
||
<collectorName id="6F6CA1B9CF490D04FBD6F88DFB00F802" box="[1091,1208,1895,1921]" pageId="4" pageNumber="34">G. Peretto</collectorName>
|
||
and
|
||
<collectorName id="6F6CA1B9CF490D04FB79F88DFA86F802" box="[1260,1342,1895,1921]" pageId="4" pageNumber="34">E. Piva</collectorName>
|
||
leg.;
|
||
<determinerName id="37FA28ABCF490D04FF2EF860FEF9F827" box="[187,321,1930,1956]" pageId="4" pageNumber="34">H. Enghoff</determinerName>
|
||
det. 2013;
|
||
<collectionCode id="A4885CAACF490D04FE56F860FDA5F827" box="[451,541,1930,1956]" pageId="4" pageNumber="34">NHMD</collectionCode>
|
||
</materialsCitation>
|
||
•
|
||
<materialsCitation id="72F1CE32CF490D04FDA0F860FB6FF84B" ID-GBIF-Occurrence="3504034313" collectingDate="1973-12" collectionCode="NHMD" collectorName="Ischia & A. Minelli" country="Italy" county="Grigno" determinerName="H. Enghoff" location="del Calgeron" municipality="Grotta" pageId="4" pageNumber="34" specimenCount="1" specimenCount-female="1" stateProvince="Autonomous Province of Trento">
|
||
<specimenCount id="D49F0FE6CF490D04FDA0F860FDDAF827" box="[565,610,1930,1956]" pageId="4" pageNumber="34" type="female">1 ♀</specimenCount>
|
||
;
|
||
<collectingCounty id="2B47BCE3CF490D04FDE5F860FD7EF827" box="[624,710,1930,1956]" pageId="4" pageNumber="34">Grigno</collectingCounty>
|
||
,
|
||
<collectingMunicipality id="22425E15CF490D04FD47F860FCA5F827" box="[722,797,1930,1956]" pageId="4" pageNumber="34">Grotta</collectingMunicipality>
|
||
[Cave]
|
||
<location id="C74692B4CF490D04FCEEF860FBA9F827" LSID="urn:lsid:plazi:treatment:4A307579CF490D05FD95FB2DFB16F9E2:C74692B4CF490D04FCEEF860FBA9F827" box="[891,1041,1930,1956]" country="Italy" county="Grigno" municipality="Grotta" name="del Calgeron" pageId="4" pageNumber="34" stateProvince="Autonomous Province of Trento">del Calgeron</location>
|
||
(new record), a
|
||
<locationDeviation id="D394DD4DCF490D04FB44F860FEECF844" pageId="4" pageNumber="34">side branch of the waterfall</locationDeviation>
|
||
;
|
||
<date id="B627E2AFCF490D04FEF5F844FE64F844" box="[352,476,1965,1992]" pageId="4" pageNumber="34" value="1973-12">
|
||
<collectingDate id="A6631B47CF490D04FEF5F844FE64F844" box="[352,476,1965,1992]" pageId="4" pageNumber="34" value="1973-12">Dec. 1973</collectingDate>
|
||
</date>
|
||
;
|
||
<collectorName id="6F6CA1B9CF490D04FE7DF847FD96F844" box="[488,558,1965,1991]" pageId="4" pageNumber="34">Ischia</collectorName>
|
||
leg.;
|
||
<determinerName id="37FA28ABCF490D04FDE5F844FD4CF844" box="[624,756,1965,1992]" pageId="4" pageNumber="34">H. Enghoff</determinerName>
|
||
det. 1984;
|
||
<collectorName id="6F6CA1B9CF490D04FCE3F847FC48F844" box="[886,1008,1965,1991]" pageId="4" pageNumber="34">A. Minelli</collectorName>
|
||
ded. 1985;
|
||
<collectionCode id="A4885CAACF490D04FBEFF844FB6FF84B" box="[1146,1239,1966,1992]" pageId="4" pageNumber="34">NHMD</collectionCode>
|
||
</materialsCitation>
|
||
.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="8A8397E4CF480D05FF28FEE2FE8FFBA6" pageId="5" pageNumber="35" type="description">
|
||
<paragraph id="C226C46FCF480D05FF28FEE2FE29FEA0" blockId="5.[189,401,264,291]" box="[189,401,264,291]" pageId="5" pageNumber="35">
|
||
<heading id="996E7303CF480D05FF28FEE2FE29FEA0" bold="true" box="[189,401,264,291]" fontSize="11" level="3" pageId="5" pageNumber="35" reason="3">
|
||
<emphasis id="F0ED187DCF480D05FF28FEE2FE29FEA0" bold="true" box="[189,401,264,291]" pageId="5" pageNumber="35">Descriptive notes</emphasis>
|
||
</heading>
|
||
</paragraph>
|
||
<paragraph id="C226C46FCF480D05FF28FEDDFCA5FE38" blockId="5.[189,1399,311,443]" pageId="5" pageNumber="35">ANTENNAE. 2–2.1 times as long as head and 1.7–1.75 times as long as H in males, and 1.7–1.8 times and ca 1.4 times, respectively, in females; antennomere 5 ca twice as long as broad; antennomeres 2, 3 and 4 subequal in length, ca 1.2 times as long as 5, and 1.7–1.8 times as long as 6; 6 visibly broader than 5, giving a clavate appearance of the antenna.</paragraph>
|
||
<paragraph id="C226C46FCF480D05FF28FE09FD6AFD2F" blockId="5.[188,1401,482,685]" pageId="5" pageNumber="35">
|
||
MOUTHPARTS. With strong hydrophilous modifications (sensu
|
||
<bibRefCitation id="A608B99ECF480D05FC2BFE08FBD1FE7E" author="Enghoff H." box="[958,1129,482,509]" pageId="5" pageNumber="35" pagination="67 - 77" refId="ref20760" refString="Enghoff H. 1985. Modified mouthparts in hydrophilous cave millipedes (Diplopoda). In: Ellis W. N., Jeekel C. A. W. & Pieters F. F. J. M. (eds) Proceedings of the 6 th International Congress of Myriapodology. Amsterdam, 12 - 17 April 1984. Bijdragen tot de Dierkunde 55 (1): 67 - 77." type="journal article" year="1985">Enghoff 1985</bibRefCitation>
|
||
): labrum edentate or with three minute, vestigial teeth. Gnathochilarium short and distally markedly broad, stipites with conspicuously large palps. Gnathal lobes of mandibles with the external and the internal tooth strongly reduced, both being distinct but very small and pointed, deeply hidden in the buccal cavity; molar plate much smaller than the normal julid condition; pectinate lamellae five instead of the usual four, consisting of very fine and densely set teeth.
|
||
</paragraph>
|
||
<paragraph id="C226C46FCF480D05FF28FD3EFC5BFC92" blockId="5.[188,1398,723,785]" pageId="5" pageNumber="35">TARSUS OF MID- BODY LEGS. Ca 2.5 times as long as tibia and ca 5 times as long as apical claw. Mid-body legs ca 1.7 times as long as H in males and ca 1.4 times in females.</paragraph>
|
||
<paragraph id="C226C46FCF480D05FF28FCD3FE8FFBA6" blockId="5.[188,1399,824,1062]" pageId="5" pageNumber="35">
|
||
FEMALE SEXUAL CHARACTERS. Legs 1 and 2 slightly shorter but not thicker than following legs. Vulva (
|
||
<figureCitation id="5AA2D8EACF480D05FF52FCB1FE8FFCF6" box="[199,311,859,885]" captionStart="Fig" captionStartId="31.[189,231,1681,1707]" captionTargetBox="[345,1243,265,1635]" captionTargetId="figure-15@31.[325,1262,262,1645]" captionTargetPageId="31" captionText="Fig. 11. Left vulvae of species of Stygiiulus stat. nov. (A–D) and representatives of (other) Julini/ Leptoiulini (E) and Typhloiulini/blind Julini/Leptoiulini in caudal, somewhat mesal views. A. S. ausugi (Manfredi, 1953) comb. nov., ♀ from Grotta [Cave] del Calgeron (NHMD). B. S. illyricus (Verhoef, 1929) comb. nov., ♀ from Rabakova Cave (IZB). C. S. insularis (Strasser, 1938) comb. nov., topotype ♀ (CBSS). D. S. tobias (Berlese, 1886) comb. nov., ♀ from Grotta [Cave] del Subiolo (NHMD). E. Leptoiulus cf. trilineatus(C.L. Koch, 1847), ♀ from Belasitsa Mtn, Bulgaria. F. Serboiulus spelaeophilus Gulička, 1967, ♀ from Vodni Pech Cave (NMNHS). G. Typhloiulus (Inversotyphlus) lobifer Attems, 1951, ♀ from Minjera-Bauxite Mine (IBER). H. Typhloiulus (s. str.) strictus (Latzel, 1882), ♀ from Kunino (IBER). Abbreviations: at = anterior tube; la = lateral ampulla; lt = lateral tube; ma = median ampulla; mt = median tube; op = operculum; pa = posterior ampulla; pt = posterior tube. Scale bar = 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.6323030" httpUri="https://zenodo.org/record/6323030/files/figure.png" pageId="5" pageNumber="35">Fig. 11A</figureCitation>
|
||
) symmetric; bursa very broad, strongly compressed in the sagittal plane; each valve of bursa with one vertical row of setae; operculum (op) distally bulging, with a distinct apical concavity, exceeding bursa by nearly ⅓ of total height of vulva, with just several setae each side. Receptaculum seminis consisting of two small tubes: a very fine, somewhat bent, mesal one (mt) ending in a small ovoid ampulla (ma), and a significantly broader, mostly straight, lateral one (lt), not forming ampulla at bottom.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="8A8397E4CF480D05FF28FBA6FACFFB5E" pageId="5" pageNumber="35" type="distribution">
|
||
<paragraph id="C226C46FCF480D05FF28FBA6FEE1FBE5" blockId="5.[189,345,1100,1126]" box="[189,345,1100,1126]" pageId="5" pageNumber="35">
|
||
<heading id="996E7303CF480D05FF28FBA6FEE1FBE5" bold="true" box="[189,345,1100,1126]" fontSize="11" level="3" pageId="5" pageNumber="35" reason="3">
|
||
<emphasis id="F0ED187DCF480D05FF28FBA6FEE1FBE5" bold="true" box="[189,345,1100,1126]" pageId="5" pageNumber="35">Distribution</emphasis>
|
||
</heading>
|
||
</paragraph>
|
||
<paragraph id="C226C46FCF480D05FF28FB91FACFFB5E" blockId="5.[188,1399,1147,1245]" pageId="5" pageNumber="35">
|
||
Prior to this study, the species was known only from its
|
||
<typeStatus id="1D227ACDCF480D05FCD3FB96FCC1FB15" box="[838,889,1148,1174]" pageId="5" pageNumber="35">type</typeStatus>
|
||
locality – the Grotta della Bigonda – on the northern border of the central part of the Venetian Prealps. The new locality of this species – the Grotta del Calgeron – is located in the same area, some
|
||
<quantity id="0561698ACF480D05FD6DFB28FCFAFB5F" box="[760,834,1218,1244]" metricMagnitude="4" metricUnit="m" metricValue="2.0" pageId="5" pageNumber="35" unit="km" value="20.0">20 km</quantity>
|
||
south of the
|
||
<typeStatus id="1D227ACDCF480D05FC4EFB28FBB6FB5F" box="[987,1038,1218,1244]" pageId="5" pageNumber="35">type</typeStatus>
|
||
locality (
|
||
<figureCitation id="5AA2D8EACF480D05FBE8FB28FB6AFB5F" box="[1149,1234,1218,1244]" captionStart="Fig" captionStartId="33.[189,232,1998,2024]" captionTargetBox="[189,1398,1099,1960]" captionTargetId="figure-332@33.[189,1398,1099,1960]" captionTargetPageId="33" captionText="Fig. 13. General distribution of the species of the genus Stygiiulus stat. nov." figureDoi="http://doi.org/10.5281/zenodo.6323034" httpUri="https://zenodo.org/record/6323034/files/figure.png" pageId="5" pageNumber="35">Fig. 13</figureCitation>
|
||
, blue circles).
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="8A8397E4CF480D05FF28FAE9FB16F9E2" pageId="5" pageNumber="35" type="discussion">
|
||
<paragraph id="C226C46FCF480D05FF28FAE9FE89FA9E" blockId="5.[189,305,1283,1309]" box="[189,305,1283,1309]" pageId="5" pageNumber="35">
|
||
<heading id="996E7303CF480D05FF28FAE9FE89FA9E" bold="true" box="[189,305,1283,1309]" fontSize="11" level="3" pageId="5" pageNumber="35" reason="3">
|
||
<emphasis id="F0ED187DCF480D05FF28FAE9FE89FA9E" bold="true" box="[189,305,1283,1309]" pageId="5" pageNumber="35">Remarks</emphasis>
|
||
</heading>
|
||
</paragraph>
|
||
<paragraph id="C226C46FCF480D05FF28FAD8FAD3F99C" blockId="5.[189,1399,1330,1567]" pageId="5" pageNumber="35">
|
||
<bibRefCitation id="A608B99ECF480D05FF28FAD8FE3DFACF" author="Strasser K." box="[189,389,1330,1356]" pageId="5" pageNumber="35" pagination="1 - 21" refId="ref22340" refString="Strasser K. 1971 a. Uber italienische, besonders kavernikole Diplopoden. Memorie del Museo civico di Storia naturale di Verona 19: 1 - 21." type="journal article" year="1971">Strasser (1971a)</bibRefCitation>
|
||
described two subspecies of
|
||
<taxonomicName id="0599BFECCF480D05FD6FFAD8FCF2FAC8" authorityName="Manfredi" authorityYear="1953" box="[762,842,1330,1355]" class="Diplopoda" family="Julidae" genus="Typhloiulus" kingdom="Animalia" order="Julida" pageId="5" pageNumber="35" phylum="Arthropoda" rank="species" species="ausugi">
|
||
<emphasis id="F0ED187DCF480D05FD6FFAD8FCF2FAC8" box="[762,842,1330,1355]" italics="true" pageId="5" pageNumber="35">ausugi</emphasis>
|
||
</taxonomicName>
|
||
, viz.,
|
||
<taxonomicName id="0599BFECCF480D05FC0BFAD8FBAFFACF" authorityName="Vagalinski & Borissov & Bobeva & Canciani & Antić" authorityYear="2022" baseAuthorityName="Strasser" baseAuthorityYear="1971" box="[926,1047,1330,1356]" class="Diplopoda" family="Julidae" genus="Stygiiulus" kingdom="Animalia" order="Julida" pageId="5" pageNumber="35" phylum="Arthropoda" rank="species" species="fimbriatus">
|
||
<emphasis id="F0ED187DCF480D05FC0BFAD8FBAFFACF" box="[926,1047,1330,1356]" italics="true" pageId="5" pageNumber="35">fimbriatus</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<emphasis id="F0ED187DCF480D05FBCFFAD8FB60FAC8" box="[1114,1240,1330,1355]" italics="true" pageId="5" pageNumber="35">
|
||
<taxonomicName id="0599BFECCF480D05FBCFFAD8FB6CFAC8" authorityName="Vagalinski & Borissov & Bobeva & Canciani & Antić" authorityYear="2022" baseAuthorityName="Strasser" baseAuthorityYear="1971" box="[1114,1236,1330,1355]" class="Diplopoda" family="Julidae" genus="Stygiiulus" kingdom="Animalia" order="Julida" pageId="5" pageNumber="35" phylum="Arthropoda" rank="species" species="gentianae">gentianae</taxonomicName>
|
||
.
|
||
</emphasis>
|
||
Considering the gonopod conformations of the two latter forms, both of which differ significantly from
|
||
<taxonomicName id="0599BFECCF480D05FA9DFABFFACFFAEC" authorityName="Vagalinski & Borissov & Bobeva & Canciani & Antić" authorityYear="2022" baseAuthorityName="Manfredi" baseAuthorityYear="1953" box="[1288,1399,1365,1391]" class="Diplopoda" family="Julidae" genus="Typhloiulus" kingdom="Animalia" order="Julida" pageId="5" pageNumber="35" phylum="Arthropoda" rank="species" species="ausugi" status="comb. nov.">
|
||
<emphasis id="F0ED187DCF480D05FA9DFABFFACFFAEC" box="[1288,1399,1365,1391]" italics="true" pageId="5" pageNumber="35">S. ausugi</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="EBDEA506CF480D05FF28FA92FEFCFA10" box="[189,324,1400,1427]" pageId="5" pageNumber="35" rank="species">comb. nov.</taxonomicNameLabel>
|
||
and are instead much more similar to
|
||
<taxonomicName id="0599BFECCF480D05FCBCFA92FC16FA11" authorityName="Vagalinski & Borissov & Bobeva & Canciani & Antić" authorityYear="2022" baseAuthorityName="Verhoeff" baseAuthorityYear="1929" box="[809,942,1400,1426]" class="Diplopoda" family="Julidae" genus="Typhloiulus" kingdom="Animalia" order="Julida" pageId="5" pageNumber="35" phylum="Arthropoda" rank="species" species="illyricus" status="comb. nov.">
|
||
<emphasis id="F0ED187DCF480D05FCBCFA92FC16FA11" box="[809,942,1400,1426]" italics="true" pageId="5" pageNumber="35">S. illyricus</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="EBDEA506CF480D05FC2DFA92FBFAFA10" box="[952,1090,1400,1427]" pageId="5" pageNumber="35" rank="species">comb. nov.</taxonomicNameLabel>
|
||
,
|
||
<taxonomicName id="0599BFECCF480D05FBC7FA92FB65FA11" authorityName="Vagalinski & Borissov & Bobeva & Canciani & Antić" authorityYear="2022" baseAuthorityName="Verhoeff" baseAuthorityYear="1929" box="[1106,1245,1400,1426]" class="Diplopoda" family="Julidae" genus="Stygiiulus" kingdom="Animalia" order="Julida" pageId="5" pageNumber="35" phylum="Arthropoda" rank="species" species="maximus" status="comb. nov.">
|
||
<emphasis id="F0ED187DCF480D05FBC7FA92FB65FA11" box="[1106,1245,1400,1426]" italics="true" pageId="5" pageNumber="35">S. maximus</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="EBDEA506CF480D05FB7DFA92FACAFA10" box="[1256,1394,1400,1427]" pageId="5" pageNumber="35" rank="species">comb. nov.</taxonomicNameLabel>
|
||
,
|
||
<taxonomicName id="0599BFECCF480D05FF28FA76FEC3FA36" authorityName="Vagalinski & Borissov & Bobeva & Canciani & Antić" authorityYear="2022" baseAuthorityName="Verhoeff" baseAuthorityYear="1930" box="[189,379,1435,1462]" class="Diplopoda" family="Julidae" genus="Stygiiulus" kingdom="Animalia" order="Julida" pageId="5" pageNumber="35" phylum="Arthropoda" rank="species" species="montellensis" status="comb. nov.">
|
||
<emphasis id="F0ED187DCF480D05FF28FA76FEC3FA36" box="[189,379,1435,1462]" italics="true" pageId="5" pageNumber="35">S. montellensis</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="EBDEA506CF480D05FE18FA71FDA5FA35" box="[397,541,1435,1462]" pageId="5" pageNumber="35" rank="species">comb. nov.</taxonomicNameLabel>
|
||
, and
|
||
<taxonomicName id="0599BFECCF480D05FDE4FA76FCA2FA36" authorityName="Vagalinski & Borissov & Bobeva & Canciani & Antić" authorityYear="2022" baseAuthorityName="Strasser" baseAuthorityYear="1962" box="[625,794,1435,1462]" class="Diplopoda" family="Julidae" genus="Stygiiulus" kingdom="Animalia" order="Julida" pageId="5" pageNumber="35" phylum="Arthropoda" rank="species" species="rotundatus" status="comb. et stat. nov.">
|
||
<emphasis id="F0ED187DCF480D05FDE4FA76FCA2FA36" box="[625,794,1435,1462]" italics="true" pageId="5" pageNumber="35">S. rotundatus</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="EBDEA506CF480D05FCBEFA71FB9EFA35" box="[811,1062,1435,1462]" pageId="5" pageNumber="35">comb. et stat. nov.</taxonomicNameLabel>
|
||
, it becomes obvious that
|
||
<bibRefCitation id="A608B99ECF480D05FF28FA55FE38FA5A" author="Strasser K." box="[189,384,1471,1497]" pageId="5" pageNumber="35" pagination="1 - 21" refId="ref22340" refString="Strasser K. 1971 a. Uber italienische, besonders kavernikole Diplopoden. Memorie del Museo civico di Storia naturale di Verona 19: 1 - 21." type="journal article" year="1971">Strasser (1971a)</bibRefCitation>
|
||
treated the modified mouthparts as a taxonomic feature of primary importance, being unaware of the adaptive nature of such modifications, as revealed later by
|
||
<bibRefCitation id="A608B99ECF480D05FBD3FA08FABAFA7F" author="Enghoff H." box="[1094,1282,1505,1532]" pageId="5" pageNumber="35" pagination="67 - 77" refId="ref20760" refString="Enghoff H. 1985. Modified mouthparts in hydrophilous cave millipedes (Diplopoda). In: Ellis W. N., Jeekel C. A. W. & Pieters F. F. J. M. (eds) Proceedings of the 6 th International Congress of Myriapodology. Amsterdam, 12 - 17 April 1984. Bijdragen tot de Dierkunde 55 (1): 67 - 77." type="journal article" year="1985">Enghoff (1985)</bibRefCitation>
|
||
. Thus we here elevate
|
||
<taxonomicName id="0599BFECCF480D05FEC0F9EFFE76F99C" authorityName="Vagalinski & Borissov & Bobeva & Canciani & Antić" authorityYear="2022" baseAuthorityName="Strasser" baseAuthorityYear="1971" box="[341,462,1541,1567]" class="Diplopoda" family="Julidae" genus="Stygiiulus" kingdom="Animalia" order="Julida" pageId="5" pageNumber="35" phylum="Arthropoda" rank="species" species="fimbriatus">
|
||
<emphasis id="F0ED187DCF480D05FEC0F9EFFE76F99C" box="[341,462,1541,1567]" italics="true" pageId="5" pageNumber="35">fimbriatus</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="0599BFECCF480D05FD9CF9ECFDC7F99C" authorityName="Vagalinski & Borissov & Bobeva & Canciani & Antić" authorityYear="2022" baseAuthorityName="Strasser" baseAuthorityYear="1971" box="[521,639,1542,1567]" class="Diplopoda" family="Julidae" genus="Stygiiulus" kingdom="Animalia" order="Julida" pageId="5" pageNumber="35" phylum="Arthropoda" rank="species" species="gentianae">
|
||
<emphasis id="F0ED187DCF480D05FD9CF9ECFDC7F99C" box="[521,639,1542,1567]" italics="true" pageId="5" pageNumber="35">gentianae</emphasis>
|
||
</taxonomicName>
|
||
to the species level and describe both of them in detail below.
|
||
</paragraph>
|
||
<paragraph id="C226C46FCF480D05FF28F9ADFB16F9E2" blockId="5.[189,1198,1606,1633]" box="[189,1198,1606,1633]" pageId="5" pageNumber="35">
|
||
In the Grotta della Bigonda, this species lives in sympatry with
|
||
<taxonomicName id="0599BFECCF480D05FC2EF9ADFB9AF9E3" authorityName="Vagalinski & Borissov & Bobeva & Canciani & Antić" authorityYear="2022" baseAuthorityName="Berlese" baseAuthorityYear="1886" box="[955,1058,1606,1633]" class="Diplopoda" family="Julidae" genus="Stygiiulus" kingdom="Animalia" order="Julida" pageId="5" pageNumber="35" phylum="Arthropoda" rank="species" species="tobias" status="comb. nov.">
|
||
<emphasis id="F0ED187DCF480D05FC2EF9ADFB9AF9E3" box="[955,1058,1606,1633]" italics="true" pageId="5" pageNumber="35">S. tobias</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="EBDEA506CF480D05FBBFF9ACFB16F9E2" box="[1066,1198,1606,1633]" pageId="5" pageNumber="35" rank="species">comb. nov.</taxonomicNameLabel>
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |