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<document id="32F888D9EEFBDE5E27A55DE99FE39F0F" ID-DOI="10.26107/RBZ-2020-0001" ID-ISSN="2345-7600" ID-Zenodo-Dep="4577165" ID-ZooBank="4A76F694-DD52-48CC-A0D4-C1AD9D33B0A1" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1614784698889" checkinUser="carolina" docAuthor="Páll-Gergely, Barna, Reischütz, Alexander, Maassen, Wim J. M., Grego, Jozef &amp; Hunyadi, András" docDate="2020" docId="03EE1927C5051408FC9AFF7DFBEE18DD" docLanguage="en" docName="RafflesBZool.68.1-13.pdf" docOrigin="Raffles Bulletin of Zoology 68" docStyle="DocumentStyle:AF09A0581917EACD12E7F0991C3E7C26.5:RafflesBZool.2019-.journal_article" docStyleId="AF09A0581917EACD12E7F0991C3E7C26" docStyleName="RafflesBZool.2019-.journal_article" docStyleVersion="5" docTitle="Laoennea carychioides Pall-Gergely, A. Reischutz &amp; Maassen 2020, new species" docType="treatment" docVersion="3" lastPageNumber="2" masterDocId="FFD7615FC5041409FFDAFFC0FF961E28" masterDocTitle="New taxa of Diapheridae Panha &amp; Naggs in Sutcharit et al., 2010 from Laos and Thailand (Gastropoda: Eupulmonata: Stylommatophora)" masterLastPageNumber="13" masterPageNumber="1" pageNumber="2" updateTime="1698878575407" updateUser="plazi" zenodo-license-document="CC-BY-4.0">
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<mods:title id="1271DA56E960B6F6FB3517F59CF3EB22">New taxa of Diapheridae Panha &amp; Naggs in Sutcharit et al., 2010 from Laos and Thailand (Gastropoda: Eupulmonata: Stylommatophora)</mods:title>
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<mods:name id="5055E0496F70E9C93648649CE5422418" type="personal">
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<mods:namePart id="F0E5264623519A27CA216B94AECCC329">Páll-Gergely, Barna</mods:namePart>
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<mods:namePart id="44C0CC91E53E9272004807756964E83E">Reischütz, Alexander</mods:namePart>
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<mods:namePart id="3A3A652E36DC48D318CD6FD743EAE08B">Maassen, Wim J. M.</mods:namePart>
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<mods:namePart id="2DE785F4E8B0D59E87C0D9098343AEB3">Grego, Jozef</mods:namePart>
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<mods:namePart id="25FCE320376A29D43E058E1479033E3E">Hunyadi, András</mods:namePart>
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<mods:title id="B899D8D2243C89736A435055C765F112">Raffles Bulletin of Zoology</mods:title>
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<mods:date id="11F39967D52DD3FBE661C6F8D1403E7B">2020</mods:date>
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<treatment id="03EE1927C5051408FC9AFF7DFBEE18DD" ID-DOI="http://doi.org/10.5281/zenodo.4577211" ID-Zenodo-Dep="4577211" LSID="urn:lsid:plazi:treatment:03EE1927C5051408FC9AFF7DFBEE18DD" httpUri="http://treatment.plazi.org/id/03EE1927C5051408FC9AFF7DFBEE18DD" lastPageNumber="2" pageId="1" pageNumber="2">
<subSubSection id="C35DFBBAC5051408FC9AFF7DFB7C1EDD" pageId="1" pageNumber="2" type="nomenclature">
<paragraph id="8BF8A831C5051408FC9AFF7DFB7C1EDD" blockId="1.[832,1432,188,277]" pageId="1" pageNumber="2">
<heading id="D0B01F5DC5051408FC9AFF7DFB7C1EDD" bold="true" centered="true" fontSize="10" level="2" pageId="1" pageNumber="2" reason="2">
<emphasis id="B9337423C5051408FC9AFF7DFB7C1EDD" bold="true" pageId="1" pageNumber="2">
<taxonomicName id="4C47D3B2C5051408FC9AFF7DFBC11EDD" ID-CoL="7RTK7" authority="Pall-Gergely, A. Reischutz &amp; Maassen" authorityName="Pall-Gergely, A. Reischutz &amp; Maassen" authorityYear="2020" class="Gastropoda" family="Diapheridae" genus="Laoennea" higherTaxonomySource="GBIF" kingdom="Animalia" order="Stylommatophora" pageId="1" pageNumber="2" phylum="Mollusca" rank="species" species="carychioides" status="sp. nov.">
<emphasis id="B9337423C5051408FC9AFF7DFBD61EFC" bold="true" box="[832,1088,188,212]" italics="true" pageId="1" pageNumber="2">Laoennea carychioides</emphasis>
Páll-Gergely, A. Reischütz &amp; Maassen
</taxonomicName>
,
<taxonomicNameLabel id="A200C958C5051408FBBFFF1DFB7C1EDD" box="[1125,1258,221,245]" pageId="1" pageNumber="2" rank="species">new species</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
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<subSubSection id="C35DFBBAC5051408FBD6FF3DFB5B1F3D" box="[1036,1229,253,277]" pageId="1" pageNumber="2" type="description">
<paragraph id="8BF8A831C5051408FBD6FF3DFB5B1F3D" blockId="1.[832,1432,188,277]" box="[1036,1229,253,277]" pageId="1" pageNumber="2">
(
<figureCitation id="137CB4B4C5051408FBCEFF3DFB001F3C" box="[1044,1174,253,277]" captionStart="Fig" captionStartId="2.[135,170,1935,1957]" captionTargetBox="[135,1473,317,1919]" captionTargetId="figure-7@2.[135,1473,317,1919]" captionTargetPageId="2" captionText="Fig. 1. New species of Diapheridae from Laos. A, Laoennea carychioides Páll-Gergely, A. Reischütz &amp; Maassen, new species (holotype); B, L. carychioides (paratype); C, Sinoennea infantilis Páll-Gergely &amp; Grego, new species (holotype). Scale bar represents 1 mm." figureDoi="http://doi.org/10.5281/zenodo.4577167" httpUri="https://zenodo.org/record/4577167/files/figure.png" pageId="1" pageNumber="2">Figs. 1A, B</figureCitation>
,
<figureCitation id="137CB4B4C5051408FB79FF3DFB531F3D" box="[1187,1221,253,277]" captionStart="Fig" captionStartId="10.[135,170,1193,1215]" captionTargetBox="[135,1473,190,1177]" captionTargetId="figure-112@10.[135,1473,190,1177]" captionTargetPageId="10" captionText="Fig. 6. Synoptic view of Sinoennea and Laoennea, new genus, species of Laos (all species shown) and northern Thailand. A, Laoennea carychioides Páll-Gergely, A. Reischütz &amp; Maassen, new species; B, Sinoennea infantilis Páll-Gergely &amp; Grego, new species; CE, Sinoennea variabilis Páll-Gergely &amp; Grego, new species; F, Sinoennea sutchariti Páll-Gergely &amp; Hunyadi, new species; G, Sinoennea panhai Páll-Gergely &amp; Hunyadi, new species; H, Sinoennea euryomphala Inkhavilay &amp; Panha, 2016; I, Sinoennea ljudmilena Páll-Gergely, new species; J, Sinoennea lizae Maassen, 2008; K, Sinoennea otostoma Páll-Gergely, A. Reischütz &amp; Maassen, new species; L, Sinoennea angustistoma Páll-Gergely, A. Reischütz &amp; Maassen, new species. Scale bar represents 1 mm." figureDoi="http://doi.org/10.5281/zenodo.4577179" httpUri="https://zenodo.org/record/4577179/files/figure.png" pageId="1" pageNumber="2">6A</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="C35DFBBAC5051408FCF7FEFDFC731C3D" pageId="1" pageNumber="2" type="materials_examined">
<paragraph id="8BF8A831C5051408FCF7FEFDFC731C3D" blockId="1.[813,1452,317,533]" pageId="1" pageNumber="2">
<emphasis id="B9337423C5051408FCF7FEFDFC4D1F7D" bold="true" box="[813,987,317,341]" pageId="1" pageNumber="2">Type material.</emphasis>
<materialsCitation id="3B2FA26CC5051408FC3CFEFDFC731C3D" collectingDate="2010-03" collectionCode="H, D" collectorName="A. Reischutz" country="Laos" elevation="265" latitude="18.926683" location="Tham Pou Kham Cave" longLatPrecision="1" longitude="102.39518" pageId="1" pageNumber="2" specimenCode="NHMW 113107" specimenCount="1" stateProvince="Viangchan" typeStatus="holotype">
<typeStatus id="54FC1693C5051408FC3CFEFDFBDD1F7D" box="[998,1099,317,341]" pageId="1" pageNumber="2" type="holotype">Holotype</typeStatus>
(1 shell,
<collectionCode id="ED5630F4C5051408FB66FEFDFB451F7C" box="[1212,1235,317,340]" pageId="1" pageNumber="2">H</collectionCode>
:
<quantity id="4CBF05D4C5051408FB3BFEFDFADE1F7C" box="[1249,1352,317,341]" metricMagnitude="-3" metricUnit="m" metricValue="2.37" pageId="1" pageNumber="2" unit="mm" value="2.37">2.37 mm</quantity>
,
<collectionCode id="ED5630F4C5051408FA8DFEFDFAF81F7C" box="[1367,1390,317,340]" pageId="1" pageNumber="2">D</collectionCode>
:
<quantity id="4CBF05D4C5051408FAA6FEFDFCCC1F5C" metricMagnitude="-3" metricUnit="m" metricValue="1.13" pageId="1" pageNumber="2" unit="mm" value="1.13">1.13 mm</quantity>
;
<figureCitation id="137CB4B4C5051408FCBFFE9DFC211F5D" box="[869,951,349,373]" captionStart="Fig" captionStartId="2.[135,170,1935,1957]" captionTargetBox="[135,1473,317,1919]" captionTargetId="figure-7@2.[135,1473,317,1919]" captionTargetPageId="2" captionText="Fig. 1. New species of Diapheridae from Laos. A, Laoennea carychioides Páll-Gergely, A. Reischütz &amp; Maassen, new species (holotype); B, L. carychioides (paratype); C, Sinoennea infantilis Páll-Gergely &amp; Grego, new species (holotype). Scale bar represents 1 mm." figureDoi="http://doi.org/10.5281/zenodo.4577167" httpUri="https://zenodo.org/record/4577167/files/figure.png" pageId="1" pageNumber="2">Fig. 1A</figureCitation>
) (
<specimenCode id="DBE1004AC5051408FC14FE9DFB141F5D" box="[974,1154,349,373]" collectionCode="NHMW" country="Austria" httpUri="http://grbio.org/cool/91g3-0mnw" name="Naturhistorisches Museum, Wien" pageId="1" pageNumber="2" type="Museum">NHMW 113107</specimenCode>
),
<collectingCountry id="F350E8A1C5051408FB4FFE9DFB5D1F5C" box="[1173,1227,349,372]" name="Laos" pageId="1" pageNumber="2">Laos</collectingCountry>
,
<collectingRegion id="498366D3C5051408FB0DFE9DFA3E1F5D" box="[1239,1448,349,373]" country="Laos" name="Viangchan" pageId="1" pageNumber="2">Vientiane Province</collectingRegion>
,
<location id="8E98FEEAC5051408FCF7FEBDFBA41FBD" LSID="urn:lsid:plazi:treatment:03EE1927C5051408FC9AFF7DFBEE18DD:8E98FEEAC5051408FCF7FEBDFBA41FBD" box="[813,1074,381,405]" country="Laos" latitude="18.926683" longLatPrecision="1" longitude="102.39518" name="Tham Pou Kham Cave" pageId="1" pageNumber="2" stateProvince="Viangchan">Tham Pou Kham Cave</location>
,
<locationDeviation id="9A4AB113C5051408FB9BFEBDFA861FBD" box="[1089,1296,381,405]" location="Vang Vieng" pageId="1" pageNumber="2">W of Vang Vieng</locationDeviation>
,
<elevation id="006A4F02C5051408FAC5FEBDFA301FBD" box="[1311,1446,381,405]" metricMagnitude="2" metricUnit="m" metricValue="2.65" pageId="1" pageNumber="2" unit="m" value="265.0">
<quantity id="4CBF05D4C5051408FAC5FEBDFAF11FBC" box="[1311,1383,381,405]" metricMagnitude="2" metricUnit="m" metricValue="2.65" pageId="1" pageNumber="2" unit="m" value="265.0">265 m</quantity>
a.s.l.
</elevation>
,
<geoCoordinate id="EE73CEF6C5051408FCF7FE5BFC211F9C" box="[813,951,411,436]" degrees="18" direction="north" minutes="55.601" orientation="latitude" pageId="1" pageNumber="2" precision="1" value="18.926683">18°55.601N</geoCoordinate>
,
<geoCoordinate id="EE73CEF6C5051408FC1EFE5BFBCE1F9C" box="[964,1112,411,436]" degrees="102" direction="east" minutes="23.711" orientation="longitude" pageId="1" pageNumber="2" precision="1" value="102.39518">102°23.711E</geoCoordinate>
, at the base of limestone rocks near the cave entrance, coll.
<collectorName id="26B2CDE7C5051408FBB6FE7DFB6D1FFD" box="[1132,1275,445,469]" pageId="1" pageNumber="2">A. Reischütz</collectorName>
,
<date id="FFF98EF1C5051408FAD3FE7DFA071FFD" box="[1289,1425,445,469]" pageId="1" pageNumber="2" value="2010-03">
<collectingDate id="EFBD7719C5051408FAD3FE7DFA071FFD" box="[1289,1425,445,469]" pageId="1" pageNumber="2" value="2010-03">March 2010</collectingDate>
</date>
;
<specimenCount id="9D4163B8C5051408FA44FE7DFC1F1FDD" pageId="1" pageNumber="2" type="generic" typeStatus="paratype">1 paratype</specimenCount>
(RE/1,
<collectionCode id="ED5630F4C5051408FC3BFE1DFC6E1FDC" box="[993,1016,477,500]" pageId="1" pageNumber="2">H</collectionCode>
:
<quantity id="4CBF05D4C5051408FBDEFE1DFBFE1FDC" box="[1028,1128,477,501]" metricMagnitude="-3" metricUnit="m" metricValue="2.27" pageId="1" pageNumber="2" unit="mm" value="2.27">2.27 mm</quantity>
,
<collectionCode id="ED5630F4C5051408FBAFFE1DFB1A1FDC" box="[1141,1164,477,500]" pageId="1" pageNumber="2">D</collectionCode>
:
<quantity id="4CBF05D4C5051408FB42FE1DFB6B1FDC" box="[1176,1277,477,501]" metricMagnitude="-3" metricUnit="m" metricValue="1.19" pageId="1" pageNumber="2" unit="mm" value="1.19">1.19 mm</quantity>
;
<figureCitation id="137CB4B4C5051408FAD0FE1DFACB1FDD" box="[1290,1373,477,501]" captionStart="Fig" captionStartId="2.[135,170,1935,1957]" captionTargetBox="[135,1473,317,1919]" captionTargetId="figure-7@2.[135,1473,317,1919]" captionTargetPageId="2" captionText="Fig. 1. New species of Diapheridae from Laos. A, Laoennea carychioides Páll-Gergely, A. Reischütz &amp; Maassen, new species (holotype); B, L. carychioides (paratype); C, Sinoennea infantilis Páll-Gergely &amp; Grego, new species (holotype). Scale bar represents 1 mm." figureDoi="http://doi.org/10.5281/zenodo.4577167" httpUri="https://zenodo.org/record/4577167/files/figure.png" pageId="1" pageNumber="2">Fig. 1B</figureCitation>
), same data as holotype.
</materialsCitation>
</paragraph>
</subSubSection>
<subSubSection id="C35DFBBAC5051408FCF7FDFDFB2D1C9D" pageId="1" pageNumber="2" type="diagnosis">
<paragraph id="8BF8A831C5051408FCF7FDFDFB2D1C9D" blockId="1.[811,1451,573,693]" pageId="1" pageNumber="2">
<emphasis id="B9337423C5051408FCF7FDFDFC371C7D" bold="true" box="[813,929,573,597]" pageId="1" pageNumber="2">Diagnosis.</emphasis>
Shell minute, elongate with wide penultimate and conspicuously narrow last whorl. Swelling behind peristome absent, parietal lamella nearly continuous, long. Palatal wall with a single pointed lablar denticle.
</paragraph>
</subSubSection>
<subSubSection id="C35DFBBAC5051408FCF7FD1DFAB81B1D" pageId="1" pageNumber="2" type="description">
<paragraph id="8BF8A831C5051408FCF7FD1DFAB81B1D" blockId="1.[811,1452,733,1333]" pageId="1" pageNumber="2">
<emphasis id="B9337423C5051408FCF7FD1DFC2E1CDD" bold="true" box="[813,952,733,757]" pageId="1" pageNumber="2">Description.</emphasis>
Shell minute, whitish, transparent when fresh. Shell elongate with wide penultimate and conspicuously narrow last whorl, entire shell with 6.75 whorls. It is nearly impossible to determine the boundary between protoconch and teleoconch. Teleoconch and protoconch glossy, extremely finely pitted, with some indication of weak radial growth lines that develop to thin ribs behind peristome. Aperture suboval, parietal callus rounded, extends to more than a half of preceding whorl, strongly thinned, “smeared” onto the preceding whorl. Apertural dentition three-fold, weak. Parietal lamella nearly straight from apertural view, continuous without changing direction or bending, although the “turning point” is identifiable, situated close to the outer end of the lamella. Sinulus relatively wide, rounded, opening shifted slightly laterally. There is a single, palatal (lablar) denticle, pointing in the direction of parietal lamella. Columellar lamella low, situated close to peristome. Peristome expanded and slightly reflected. Umbilicus slit-like, situated behind the columellar peristome, showing only last whorl.
</paragraph>
</subSubSection>
<subSubSection id="C35DFBBAC5051408FCF7FA9DFA9E185D" pageId="1" pageNumber="2" type="diagnosis">
<paragraph id="8BF8A831C5051408FCF7FA9DFA9E185D" blockId="1.[811,1452,1373,1653]" pageId="1" pageNumber="2">
<emphasis id="B9337423C5051408FCF7FA9DFBBB1B5D" bold="true" box="[813,1069,1373,1397]" pageId="1" pageNumber="2">Differential diagnosis.</emphasis>
This species can be distinguished from all tiny
<taxonomicName id="4C47D3B2C5051408FC1EFABDFBA21BBC" authorityName="Kobelt" authorityYear="1904" box="[964,1076,1405,1428]" class="Gastropoda" family="Streptaxidae" genus="Sinoennea" higherTaxonomySource="GBIF" kingdom="Animalia" order="Stylommatophora" pageId="1" pageNumber="2" phylum="Mollusca" rank="genus">
<emphasis id="B9337423C5051408FC1EFABDFBA21BBC" box="[964,1076,1405,1428]" italics="true" pageId="1" pageNumber="2">Sinoennea</emphasis>
</taxonomicName>
species at first sight by the wide penultimate and narrow last whorl.
<taxonomicName id="4C47D3B2C5051408FB77FA5EFA321B9D" authority="Páll-Gergely &amp; Reischütz &amp; Maassen &amp; Grego &amp; Hunyadi, 2020" authorityName="Páll-Gergely &amp; Reischütz &amp; Maassen &amp; Grego &amp; Hunyadi" authorityYear="2020" box="[1197,1444,1437,1461]" class="Gastropoda" family="Diapheridae" genus="Laoennea" higherTaxonomySource="GBIF" kingdom="Animalia" order="Stylommatophora" pageId="1" pageNumber="2" phylum="Mollusca" rank="species" species="carychioides" status="sp. nov.">
<emphasis id="B9337423C5051408FB77FA5EFA321B9D" box="[1197,1444,1437,1461]" italics="true" pageId="1" pageNumber="2">Laoennea carychioides</emphasis>
</taxonomicName>
,
<taxonomicNameLabel id="A200C958C5051408FCF7FA7DFC391BFD" box="[813,943,1469,1493]" pageId="1" pageNumber="2" rank="species">new species</taxonomicNameLabel>
, is superficially similar to members of the genus
<taxonomicName id="4C47D3B2C5051408FCF7FA1DFBFF1BDD" authority="O. F. Muller, 1773" authorityName="O. F. Muller" authorityYear="1773" box="[813,1129,1501,1525]" class="Gastropoda" family="Ellobiidae" genus="Carychium" higherTaxonomySource="GBIF" kingdom="Animalia" order="Pulmonata" pageId="1" pageNumber="2" phylum="Mollusca" rank="genus">
<emphasis id="B9337423C5051408FCF7FA1DFC351BDD" box="[813,931,1501,1525]" italics="true" pageId="1" pageNumber="2">Carychium</emphasis>
O.F.
<bibRefCitation id="EFD6D5C0C5051408FC07FA1DFBFF1BDD" author="Muller OF" box="[989,1129,1501,1525]" pageId="1" pageNumber="2" refId="ref8298" refString="Muller OF (1773) Vermium terrestrium et fluviatilium, seu animalium infusoriorum, helminthicorum, et testaecorum, non marinorum, succincta historia. Heineck &amp; Faber, Havniae [(= Copenhagen] &amp; Lipsiae [= Leipzig], 136 pp." type="book" year="1773">Müller, 1773</bibRefCitation>
</taxonomicName>
, due to the tiny, elongate shell. However,
<taxonomicName id="4C47D3B2C5051408FC46FA3DFB85183D" authorityName="O. F. Muller" authorityYear="1773" box="[924,1043,1533,1557]" class="Gastropoda" family="Ellobiidae" genus="Carychium" higherTaxonomySource="GBIF" kingdom="Animalia" order="Pulmonata" pageId="1" pageNumber="2" phylum="Mollusca" rank="genus">
<emphasis id="B9337423C5051408FC46FA3DFB85183D" box="[924,1043,1533,1557]" italics="true" pageId="1" pageNumber="2">Carychium</emphasis>
</taxonomicName>
shells have a more pointed apex, their parietal callus does not climb on the preceding whorl that much, and their parietal lamella is oblique, and situated closer to the columella on the parietal wall.
</paragraph>
</subSubSection>
<subSubSection id="C35DFBBAC5051408FCF7F95DFBEE18DD" pageId="1" pageNumber="2" type="etymology">
<paragraph id="8BF8A831C5051408FCF7F95DFBEE18DD" blockId="1.[813,1452,1693,1781]" pageId="1" pageNumber="2">
<emphasis id="B9337423C5051408FCF7F95DFC25189D" bold="true" box="[813,947,1693,1717]" pageId="1" pageNumber="2">Etymology.</emphasis>
The specific epithet indicates that this new species is similar in size, shape, and colour to the genus
<taxonomicName id="4C47D3B2C5051408FCF7F91DFBE318DD" authority="O. F. Muller, 1773" authorityName="O. F. Muller" authorityYear="1773" box="[813,1141,1757,1781]" class="Gastropoda" family="Ellobiidae" genus="Carychium" higherTaxonomySource="GBIF" kingdom="Animalia" order="Pulmonata" pageId="1" pageNumber="2" phylum="Mollusca" rank="genus">
<emphasis id="B9337423C5051408FCF7F91DFC3218DD" box="[813,932,1757,1781]" italics="true" pageId="1" pageNumber="2">Carychium</emphasis>
O.F.
<bibRefCitation id="EFD6D5C0C5051408FC3EF91DFBE318DD" author="Muller OF" box="[996,1141,1757,1781]" pageId="1" pageNumber="2" refId="ref8298" refString="Muller OF (1773) Vermium terrestrium et fluviatilium, seu animalium infusoriorum, helminthicorum, et testaecorum, non marinorum, succincta historia. Heineck &amp; Faber, Havniae [(= Copenhagen] &amp; Lipsiae [= Leipzig], 136 pp." type="book" year="1773">Müller, 1773</bibRefCitation>
</taxonomicName>
.
</paragraph>
</subSubSection>
</treatment>
</document>