338 lines
51 KiB
XML
338 lines
51 KiB
XML
<document ID-CLB-Dataset="198853" ID-DOI="http://dx.doi.org/10.3897/zookeys.1133.94530" ID-GBIF-Dataset="c5acfacc-834a-4abc-ad38-d6d2964f72f5" ID-PMC="PMC9836506" ID-Pensoft-Pub="1313-2970-1133-1" ID-Pensoft-UUID="4C6A7FCDB0B55088AED4D861AF5BE503" ID-PubMed="36760608" ID-ZooBank="D7A37FE0DC2A4ECCA6A10E873C7C7A5A" ModsDocID="1313-2970-1133-1" checkinTime="1669899240784" checkinUser="pensoft" docAuthor="Kato, Makoto, Yamamori, Luna & Imada, Yume" docDate="2022" docId="FC660752E10E56E9A0134CA0D9B34DC2" docLanguage="en" docName="ZooKeys 1133: 1-164" docOrigin="ZooKeys 1133" docPubDate="2022-11-30" docSource="http://dx.doi.org/10.3897/zookeys.1133.94530" docTitle="Phytoliriomyza crepusculum Kato 2022, sp. nov." docType="treatment" docUuid="421AB9E5-4908-450E-8960-79A097FA8EBA" docUuidSource="ZooBank" docVersion="6" id="4C6A7FCDB0B55088AED4D861AF5BE503" lastPageNumber="1" masterDocId="4C6A7FCDB0B55088AED4D861AF5BE503" masterDocTitle="Diversity underfoot of agromyzids (Agromyzidae, Diptera) mining thalli of liverworts and hornworts" masterLastPageNumber="164" masterPageNumber="1" pageNumber="1" updateTime="1720471835971" updateUser="ExternalLinkService">
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<mods:mods id="C17DABF5E825F04C54793EAADF43EC5D" xmlns:mods="http://www.loc.gov/mods/v3">
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<mods:titleInfo id="A346454E44396D41420888C29E9CCFE0">
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<mods:title id="A16A4E16FDB88D4558A80952A25D14F8">Diversity underfoot of agromyzids (Agromyzidae, Diptera) mining thalli of liverworts and hornworts</mods:title>
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</mods:titleInfo>
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<mods:name id="313769A5FF67327640E0820ED2D7C8AE" type="personal">
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<mods:roleTerm id="8A9F1038205E3BCD53E8B405E91525ED">Author</mods:roleTerm>
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<mods:namePart id="8CCA16F6CBC50F2DAE0E8B5DC733E9FB">Kato, Makoto</mods:namePart>
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<mods:nameIdentifier id="C0613CF91CF6ABA62178421E0DE3E554" type="email">kato@zoo.zool.kyoto-u.ac.jp</mods:nameIdentifier>
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<mods:name id="2DFAC8281DF4BDBE4C9E665BBC6C3B9F" type="personal">
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<mods:roleTerm id="0D8C1F5391C4EB4DCB3179CF72DF97F8">Author</mods:roleTerm>
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</mods:role>
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<mods:namePart id="7BCACA6CBFD456D4A3DDF0B04662B176">Yamamori, Luna</mods:namePart>
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<mods:nameIdentifier id="EAF5E915562DC433305017EE92DA4B7F" type="ORCID">https://orcid.org/0000-0002-5342-1277</mods:nameIdentifier>
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</mods:role>
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<mods:namePart id="3B088E8BCB42C9FAD86B06AF118800BA">Imada, Yume</mods:namePart>
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<mods:nameIdentifier id="A94A742B4EB4D3FE138732C69B1BBC17" type="ORCID">https://orcid.org/0000-0003-2173-7389</mods:nameIdentifier>
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<mods:typeOfResource id="E9A63095212A27CE9347741D8166A793">text</mods:typeOfResource>
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<mods:title id="DDC4D36B43B4F564389A1F3DA50096C3">ZooKeys</mods:title>
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<mods:part id="2BB894A8FF814BF70434277BE1DAC518">
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<mods:date id="54C2A83666C57CEEA7D9EEE7FBE1CDA0">2022</mods:date>
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<mods:detail id="416715A5A2F4E9CDB3886E8DAF8D6C1D" type="pubDate">
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<mods:number id="00C8F9CAAC21DEC3C680331C61C299F1">2022-11-30</mods:number>
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<mods:detail id="ABD3FBB8EFFA1DADA342D1A8E5EC776A" type="volume">
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<mods:number id="8EA918EB8676D86C7D4DE130E559A55D">1133</mods:number>
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<mods:start id="13B69104C523C11BC73A2B5C6128E8EE">1</mods:start>
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<mods:end id="68CCB0D193B26E08BA60292D62FB34D4">164</mods:end>
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<mods:location id="6089F8CF370FDE07F9C0259BB09D46DF">
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<mods:url id="E67DF4251290150487165F466106D71A">http://dx.doi.org/10.3897/zookeys.1133.94530</mods:url>
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</mods:location>
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<mods:classification id="CE99874AF677DEC1E12BAF18A9616681">journal article</mods:classification>
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<mods:identifier id="81A2D54EEC358FE422C151D746576A14" type="DOI">http://dx.doi.org/10.3897/zookeys.1133.94530</mods:identifier>
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<mods:identifier id="E5A705E982AD425BDE05C91A4DE3A353" type="Pensoft-Pub">1313-2970-1133-1</mods:identifier>
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<mods:identifier id="E6CC828A476368BBC4773D01BB4F59DC" type="ZooBank">D7A37FE0DC2A4ECCA6A10E873C7C7A5A</mods:identifier>
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<treatment id="FC660752E10E56E9A0134CA0D9B34DC2" ID-GBIF-Taxon="205344540" LSID="urn:lsid:zoobank.org:act:421AB9E5-4908-450E-8960-79A097FA8EBA" httpUri="http://treatment.plazi.org/id/FC660752E10E56E9A0134CA0D9B34DC2" lastPageNumber="1" pageId="0" pageNumber="1">
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<subSubSection id="2BBD1D6FAA51786C7B0E2FD2E118BCAA" pageId="0" pageNumber="1" type="nomenclature">
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<paragraph id="65E6FFBAC1E9B6D6385B562CED584CF5" pageId="0" pageNumber="1">
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7.
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<taxonomicName id="148B6E1FB31795A452BA42F10DE1ED52" ID-CoL="BDJLF" LSID="https://zoobank.org/421AB9E5-4908-450E-8960-79A097FA8EBA" authority="Kato" authorityName="Kato" authorityYear="2022" class="Insecta" family="Agromyzidae" genus="Phytoliriomyza" higherTaxonomySource="treatment-meta" kingdom="Animalia" lsidName="Phytoliriomyza crepusculum" order="Diptera" pageId="0" pageNumber="1" phylum="Arthropoda" rank="species" species="crepusculum" status="sp. nov.">Phytoliriomyza crepusculum Kato</taxonomicName>
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<taxonomicNameLabel id="2F15409F9F282D3A0B76E6FBF46CD21C" pageId="0" pageNumber="1">sp. nov.</taxonomicNameLabel>
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</paragraph>
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</subSubSection>
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<subSubSection id="0371DDBD420D2384D02503A7EA51BF6B" pageId="0" pageNumber="1" type="description">
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<paragraph id="D5694C03FD6290C8F43C0543AB665942" pageId="0" pageNumber="1">
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<figureCitation id="CE7410D5F2401D3310E39264F0647287" captionStart="Figure 13" captionStartId="F13" captionText="Figure 13. Phytoliriomyza crepusculum sp. nov. A-C holotype male A habitus B lateral C frontal D paratype male (MK-AG-a 309), dorsal E paratype female (MK-AG- 274), dorsal F-I male genitalia F ejaculatory apodeme, dorsal G whole genitalia, ventral H phallic complex, lateral I epandrium, ventral J puparium K-L female postabdomen K oviscape and spermatheca L tergite 10 M habitat of type locality N-P mined thalli of Dumortiera hirsuta (N type locality O Kiyosumi P Mt. Nabejiri). An arrow in N indicate an puparium." figureDoi="10.3897/zookeys.1133.94530.figure13" httpUri="https://binary.pensoft.net/fig/775386" pageId="0" pageNumber="1">Fig. 13.</figureCitation>
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</paragraph>
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</subSubSection>
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<subSubSection id="FCBCF53D1229B27B69C42F61A386D31E" pageId="0" pageNumber="1" type="material examined">
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<paragraph id="1827CAD014B08F0B2EA1BEE545EED777" pageId="0" pageNumber="1">Material examined.</paragraph>
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<paragraph id="A53EBC9EBDDA91AB2BD0B42421635F51" pageId="0" pageNumber="1">
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<emphasis id="786C467DCA3D8D7D868631611F27B944" bold="true" italics="true" pageId="0" pageNumber="1">Holotype</emphasis>
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<emphasis id="AA5EEF821E6CFB8F69643898CD4F9CEB" bold="true" pageId="0" pageNumber="1">:</emphasis>
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Japan: 1♂ (MK-AG-a29), Naiku, Oe, Fukuchiyama, Kyoto Pref. (
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<geoCoordinate id="5C7F402124A65D3529CE1B90D8D95D2E" degrees="35.433016" direction="north" orientation="latitude" precision="1" value="35.433018">35.433016°N</geoCoordinate>
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,
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<geoCoordinate id="0220DDB5A8AB8E4EFBC89F86889E6C84" degrees="135.150397" direction="east" orientation="longitude" precision="1" value="135.15039">135.150397°E</geoCoordinate>
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, 75 m asl), 17-III-2017 (as larva), emerged on 17-V-2017, NSMT-I-Dip 31929.
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<emphasis id="30E85EE9CF82452932E6ECEB626E9EE5" bold="true" italics="true" pageId="0" pageNumber="1">Paratypes</emphasis>
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<emphasis id="4419F48AB7F99F0F0AF728253CB0DFA4" bold="true" pageId="0" pageNumber="1">:</emphasis>
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Japan: 1♂ (MK-AG-a457), type locality, 19-V-2021 (as larva), on 1-VII-2021, NSMT-I-Dip 31930; 1♂1 ♀ (MK-AG-a399, 274), Inago, Shibakawa, Fujinomiya, Shizuoka Pref., 17-II-2002 (as larva), emerged on 17-23-IV-2002, NSMT-I-Dip 31931, 31932; 1♂ (MK-AG-a309), Mitake, Kamitsushima, Nagasaki Pref., 19-IV-2009 (as larva), emerged on 6-VI-2009, NSMT-I-Dip 31933.
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</paragraph>
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</subSubSection>
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<subSubSection id="ACC7782874678B6DBB1CF8FCE97E49D0" pageId="0" pageNumber="1" type="other material">
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<paragraph id="EFC5E941BB65773A19C13E46586942AC" pageId="0" pageNumber="1">Other material.</paragraph>
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<paragraph id="BE1D6293BDC6C3559609CC17704CB792" pageId="0" pageNumber="1">Japan: 1♀, Sanekawa-keikoku, Iide, Aga, Higashikanbara, Niigata Pref., 3-V-2015 (as larva), emerged on 12-VI -2015; 1♀, Mt. Kiyosumi, Kamogawa, Chiba Pref., 24-I-2012 (as larva), emerged on 25-V-5-VI-2012; 17♂32♀, Inago, Shibakawa, Fujinomiya, Shizuoka Pref., 17-II-2002 (as larva), emerged on 23-IV-5-VI-2021; 1♂, Shogawa-kyo, Tonami, Toyama Pref., 3-V-2016 (as larva), emerged on 9-VI -2016; 1♀, Takeda-gawa, Maruoka, Sakai, Fukui Pref., 6-IV-2002 (as larva), emerged on 13-VI-2019; 2♂2♀, Mt. Nabejiri, Taga, Shiga Pref., 23-V-2015 (as larva), emerged on 26-15-VI-2015; 1♂1♀, Boumura, Kazuragawa, Otsu, Shiga Pref., 7-IV-2002 (as larva), emerged on 24-V-2002; 1♂1♀, Mitake, Kamitsushima, Nagasaki Pref., 9-IV-2009 (as larva), emerged on 6-VI-2009.</paragraph>
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</subSubSection>
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<subSubSection id="607D92E3FBCFF69CFB46C04F7DDE7157" pageId="0" pageNumber="1" type="diagnosis">
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<paragraph id="96392181443CBFA81944BB8280A95C71" pageId="0" pageNumber="1">Diagnosis.</paragraph>
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<paragraph id="9BD8D2F95C87B5467F83D21B9FB2523E" pageId="0" pageNumber="1">
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A medium-sized species (wing length 1.4-1.8 mm) having subshiny brown scutum with brownish yellow pattern extending from mid-posterior margin to scutellum, black 1st flagellomere, brown maxillary palpus, brown halteres, and brownish yellow legs. Male epandrium inner-laterally with a comb comprising five long fused tubercle-like setae. Larva mines the thallus of
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<taxonomicName id="2114E942976501EF042DDC33FF14663D" class="Marchantiopsida" family="Dumortieraceae" genus="Dumortiera" kingdom="Plantae" lsidName="Dumortiera hirsuta" order="Marchantiales" pageId="0" pageNumber="1" phylum="Marchantiophyta" rank="species" species="hirsuta">
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<emphasis id="614A3D0FA91C78187B15A3DA9D0E8A00" italics="true" pageId="0" pageNumber="1">Dumortiera hirsuta</emphasis>
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</taxonomicName>
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.
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</paragraph>
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</subSubSection>
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<subSubSection id="F2BA4A5E976EB6190AC46CA07BF8D184" pageId="0" pageNumber="1" type="description">
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<paragraph id="AC621FAA9C610E4B3FB40A032CCA6C16" pageId="0" pageNumber="1">Description.</paragraph>
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<paragraph id="4676338A697C5671D3E0EDDF2E0F72E4" pageId="0" pageNumber="1">
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<emphasis id="EF59A56BF090E3F5C84913047F6FCB60" bold="true" pageId="0" pageNumber="1">Adult male</emphasis>
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(Fig.
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<figureCitation id="96571E0011A8D020AE4153B4171BAFF9" captionStart="Figure 13" captionStartId="F13" captionText="Figure 13. Phytoliriomyza crepusculum sp. nov. A-C holotype male A habitus B lateral C frontal D paratype male (MK-AG-a 309), dorsal E paratype female (MK-AG- 274), dorsal F-I male genitalia F ejaculatory apodeme, dorsal G whole genitalia, ventral H phallic complex, lateral I epandrium, ventral J puparium K-L female postabdomen K oviscape and spermatheca L tergite 10 M habitat of type locality N-P mined thalli of Dumortiera hirsuta (N type locality O Kiyosumi P Mt. Nabejiri). An arrow in N indicate an puparium." figureDoi="10.3897/zookeys.1133.94530.figure13" httpUri="https://binary.pensoft.net/fig/775386" pageId="0" pageNumber="1">13A-D</figureCitation>
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).
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</paragraph>
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<paragraph id="E1823BFC2EBC9A5D2B42D6B1987ECA30" pageId="0" pageNumber="1">
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<emphasis id="08EA0C8BF4B7BF388D6CF269B191AD1E" bold="true" italics="true" pageId="0" pageNumber="1">Head</emphasis>
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<emphasis id="601BBDE447B312B3B5D0DD91A5306CAF" bold="true" pageId="0" pageNumber="1">:</emphasis>
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Head largely light yellow; ocellar triangle yellow but ocellar tubercle dark brown; back of head dark brown above foramen excluding margin (Fig.
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<figureCitation id="E0BEA7898FF2D324DFDEDAA54C7D8A62" captionStart="Figure 13" captionStartId="F13" captionText="Figure 13. Phytoliriomyza crepusculum sp. nov. A-C holotype male A habitus B lateral C frontal D paratype male (MK-AG-a 309), dorsal E paratype female (MK-AG- 274), dorsal F-I male genitalia F ejaculatory apodeme, dorsal G whole genitalia, ventral H phallic complex, lateral I epandrium, ventral J puparium K-L female postabdomen K oviscape and spermatheca L tergite 10 M habitat of type locality N-P mined thalli of Dumortiera hirsuta (N type locality O Kiyosumi P Mt. Nabejiri). An arrow in N indicate an puparium." figureDoi="10.3897/zookeys.1133.94530.figure13" httpUri="https://binary.pensoft.net/fig/775386" pageId="0" pageNumber="1">13C</figureCitation>
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). Antenna porrect, first flagellomere black, pedicel and scape yellow. Arista subbasal, pubescent. Face, gena, parafacial and postgena light yellow. Proboscis normal, yellow; palpus yellow, cylindrical.
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<emphasis id="7AD2190382FEF25FC5A1B7571418071E" bold="true" italics="true" pageId="0" pageNumber="1">Chaetotaxy</emphasis>
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<emphasis id="C06BA0C3055F47B9F0C73756A31D502A" bold="true" pageId="0" pageNumber="1">:</emphasis>
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Front orbitals three pairs; one ori directed inward; two ors directed upward (Fig.
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<figureCitation id="E92CE5A7072929CCEF66076CCCAD4B13" captionStart="Figure 13" captionStartId="F13" captionText="Figure 13. Phytoliriomyza crepusculum sp. nov. A-C holotype male A habitus B lateral C frontal D paratype male (MK-AG-a 309), dorsal E paratype female (MK-AG- 274), dorsal F-I male genitalia F ejaculatory apodeme, dorsal G whole genitalia, ventral H phallic complex, lateral I epandrium, ventral J puparium K-L female postabdomen K oviscape and spermatheca L tergite 10 M habitat of type locality N-P mined thalli of Dumortiera hirsuta (N type locality O Kiyosumi P Mt. Nabejiri). An arrow in N indicate an puparium." figureDoi="10.3897/zookeys.1133.94530.figure13" httpUri="https://binary.pensoft.net/fig/775386" pageId="0" pageNumber="1">13B</figureCitation>
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). Orbital setulae minute and erect, in a single row.
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</paragraph>
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<caption id="6D3D629E67DAECAEE11C7C15D6D7EE58" doi="10.3897/zookeys.1133.94530.figure13" httpUri="https://binary.pensoft.net/fig/775386" pageId="0" pageNumber="1" start="Figure 13" startId="F13">
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<paragraph id="92FC47BCCBFD6AA85CE951F272467A3C" pageId="0" pageNumber="1">
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<emphasis id="25743D6D6450B31ACA8329347A8388E7" bold="true" pageId="0" pageNumber="1">Figure 13.</emphasis>
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<taxonomicName id="52898DA07D07F088F8173031059BA38B" authorityName="Kato" authorityYear="2022" class="Insecta" family="Dendrocerotaceae" genus="Phytoliriomyza" kingdom="Animalia" lsidName="Phytoliriomyza crepusculum" order="Diptera" pageId="0" pageNumber="1" phylum="Arthropoda" rank="species" species="crepusculum">
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<emphasis id="0B44FB3D265A829466C9B66F44573A3A" italics="true" pageId="0" pageNumber="1">Phytoliriomyza crepusculum</emphasis>
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</taxonomicName>
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sp. nov.
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<emphasis id="307E075AEC7C7CB07540F8A060F2B962" bold="true" pageId="0" pageNumber="1">A-C</emphasis>
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holotype male
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<emphasis id="65C43E8CB6803C4894DFECAACE7E2617" bold="true" pageId="0" pageNumber="1">A</emphasis>
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habitus
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<emphasis id="11ED77E062B5E8011EF707BB6057F797" bold="true" pageId="0" pageNumber="1">B</emphasis>
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lateral
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<emphasis id="E653D2C3481BD95A64DE131ABE11CD80" bold="true" pageId="0" pageNumber="1">C</emphasis>
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frontal
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<emphasis id="F7E88CC644AF532A388DA72C59EE2AA7" bold="true" pageId="0" pageNumber="1">D</emphasis>
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paratype male (MK-AG-a309), dorsal
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<emphasis id="3A0137248646512AE35103860B4B5629" bold="true" pageId="0" pageNumber="1">E</emphasis>
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paratype female (MK-AG-274), dorsal
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<emphasis id="0FF20AB59DF23E84D2B2A16945FE29EA" bold="true" pageId="0" pageNumber="1">F-I</emphasis>
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male genitalia
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<emphasis id="A587B1691C8EFDC4533AF6EC2DF03CFE" bold="true" pageId="0" pageNumber="1">F</emphasis>
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ejaculatory apodeme, dorsal
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<emphasis id="E7C3EC946265D6C22E2DB532A57A66BE" bold="true" pageId="0" pageNumber="1">G</emphasis>
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whole genitalia, ventral
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<emphasis id="087457A1DF1D1185E454E86FFC57AD88" bold="true" pageId="0" pageNumber="1">H</emphasis>
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phallic complex, lateral
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<emphasis id="70792704BA453F1A8D68C4A714235278" bold="true" pageId="0" pageNumber="1">I</emphasis>
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epandrium, ventral
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<emphasis id="E5E12B72C04873DE18CE9654A6C8D189" bold="true" pageId="0" pageNumber="1">J</emphasis>
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puparium
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<emphasis id="741E91B17A8E00D6C999426FDA525BF4" bold="true" pageId="0" pageNumber="1">K-L</emphasis>
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female postabdomen
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<emphasis id="560ACFA58D168BEF4D1CFD8674FE88C0" bold="true" pageId="0" pageNumber="1">K</emphasis>
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oviscape and spermatheca
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<emphasis id="BA81EF51CEDBC14EB87565D5D383B5CC" bold="true" pageId="0" pageNumber="1">L</emphasis>
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tergite 10
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<emphasis id="BDAEF0F215B93A160ED5F8973AECD9D3" bold="true" pageId="0" pageNumber="1">M</emphasis>
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habitat of type locality
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<emphasis id="5A6B4D7B59BC9F3CAF7DEEF53E875630" bold="true" pageId="0" pageNumber="1">N-P</emphasis>
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mined thalli of
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<taxonomicName id="F4193536507BD87CB1E283E24C75E169" class="Marchantiopsida" family="Dumortieraceae" genus="Dumortiera" kingdom="Plantae" lsidName="Dumortiera hirsuta" order="Marchantiales" pageId="0" pageNumber="1" phylum="Marchantiophyta" rank="species" species="hirsuta">
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<emphasis id="6848BB0EA6FC81562FAAEB51B9ACAD5B" italics="true" pageId="0" pageNumber="1">Dumortiera hirsuta</emphasis>
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</taxonomicName>
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<emphasis id="2F1E8FC7FCBDA42B6840D9B32AE6192A" bold="true" pageId="0" pageNumber="1">(N</emphasis>
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type locality
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<emphasis id="2D7F0F782330152FAF26CD7649356153" bold="true" pageId="0" pageNumber="1">O</emphasis>
|
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Kiyosumi
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<emphasis id="3B99EA161FF79A3EAFA4F5FA78B7A1C2" bold="true" pageId="0" pageNumber="1">P</emphasis>
|
|
Mt. Nabejiri). An arrow in
|
|
<emphasis id="1242D12025B84D1AC9B864E82DB1AC66" bold="true" pageId="0" pageNumber="1">N</emphasis>
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|
indicate an puparium.
|
|
</paragraph>
|
|
</caption>
|
|
<paragraph id="23C7DC9902D1107C4FE009D63D579A85" pageId="0" pageNumber="1">
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|
<emphasis id="A136E118C8F81F8084D9C72EC52E5E8B" bold="true" italics="true" pageId="0" pageNumber="1">Thorax</emphasis>
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<emphasis id="AC1D2AFFFF6887A058716F582C8E3A45" bold="true" pageId="0" pageNumber="1">:</emphasis>
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Thorax subshiny. Scutum brown with a large posterior trapezoid yellow patch adjacent to scutellum (Fig.
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|
<figureCitation id="9C9ED8FEBAAB7D1C33C451253ED6B18E" captionStart="Figure 13" captionStartId="F13" captionText="Figure 13. Phytoliriomyza crepusculum sp. nov. A-C holotype male A habitus B lateral C frontal D paratype male (MK-AG-a 309), dorsal E paratype female (MK-AG- 274), dorsal F-I male genitalia F ejaculatory apodeme, dorsal G whole genitalia, ventral H phallic complex, lateral I epandrium, ventral J puparium K-L female postabdomen K oviscape and spermatheca L tergite 10 M habitat of type locality N-P mined thalli of Dumortiera hirsuta (N type locality O Kiyosumi P Mt. Nabejiri). An arrow in N indicate an puparium." figureDoi="10.3897/zookeys.1133.94530.figure13" httpUri="https://binary.pensoft.net/fig/775386" pageId="0" pageNumber="1">13D</figureCitation>
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). Scutellum yellow with lateral corner narrowly brown. Subscutellum yellow. Mediotergite, katatergite and anatergite brown. Pleuron yellow, but propleuron, meron, anepimeron centrally pale brown (Fig.
|
|
<figureCitation id="E92551632D672A7ED375F2C68943A695" captionStart="Figure 13" captionStartId="F13" captionText="Figure 13. Phytoliriomyza crepusculum sp. nov. A-C holotype male A habitus B lateral C frontal D paratype male (MK-AG-a 309), dorsal E paratype female (MK-AG- 274), dorsal F-I male genitalia F ejaculatory apodeme, dorsal G whole genitalia, ventral H phallic complex, lateral I epandrium, ventral J puparium K-L female postabdomen K oviscape and spermatheca L tergite 10 M habitat of type locality N-P mined thalli of Dumortiera hirsuta (N type locality O Kiyosumi P Mt. Nabejiri). An arrow in N indicate an puparium." figureDoi="10.3897/zookeys.1133.94530.figure13" httpUri="https://binary.pensoft.net/fig/775386" pageId="0" pageNumber="1">13B</figureCitation>
|
|
). Haltere brown. Calypter margin and hairs gray. Leg segments yellow; tibia and tarsus dorsally brown (Fig.
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|
<figureCitation id="9CA81A9A1055AA1F9C204149D3BD4351" captionStart="Figure 13" captionStartId="F13" captionText="Figure 13. Phytoliriomyza crepusculum sp. nov. A-C holotype male A habitus B lateral C frontal D paratype male (MK-AG-a 309), dorsal E paratype female (MK-AG- 274), dorsal F-I male genitalia F ejaculatory apodeme, dorsal G whole genitalia, ventral H phallic complex, lateral I epandrium, ventral J puparium K-L female postabdomen K oviscape and spermatheca L tergite 10 M habitat of type locality N-P mined thalli of Dumortiera hirsuta (N type locality O Kiyosumi P Mt. Nabejiri). An arrow in N indicate an puparium." figureDoi="10.3897/zookeys.1133.94530.figure13" httpUri="https://binary.pensoft.net/fig/775386" pageId="0" pageNumber="1">13D</figureCitation>
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).
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|
<emphasis id="3ABBCFB9B136935ADFD989771B2A5FF5" bold="true" italics="true" pageId="0" pageNumber="1">Chaetotaxy</emphasis>
|
|
<emphasis id="C8ABB5078E56C08D190A774EB5189BF7" bold="true" pageId="0" pageNumber="1">:</emphasis>
|
|
Scutum with 1+3 dorsocentrals, shortened anteriorly (Fig.
|
|
<figureCitation id="9263507B909CFFA05C9C22C897923A56" captionStart="Figure 13" captionStartId="F13" captionText="Figure 13. Phytoliriomyza crepusculum sp. nov. A-C holotype male A habitus B lateral C frontal D paratype male (MK-AG-a 309), dorsal E paratype female (MK-AG- 274), dorsal F-I male genitalia F ejaculatory apodeme, dorsal G whole genitalia, ventral H phallic complex, lateral I epandrium, ventral J puparium K-L female postabdomen K oviscape and spermatheca L tergite 10 M habitat of type locality N-P mined thalli of Dumortiera hirsuta (N type locality O Kiyosumi P Mt. Nabejiri). An arrow in N indicate an puparium." figureDoi="10.3897/zookeys.1133.94530.figure13" httpUri="https://binary.pensoft.net/fig/775386" pageId="0" pageNumber="1">13D</figureCitation>
|
|
). Acrostichal setulae five pairs in two rows.
|
|
<emphasis id="E796B061016BE1CB02049F64B1179081" bold="true" italics="true" pageId="0" pageNumber="1">Wing</emphasis>
|
|
<emphasis id="0BBE6822018868C727FACA5CB55F8FBD" bold="true" pageId="0" pageNumber="1">:</emphasis>
|
|
Wing length 1.4 mm, costa reaching M1 (Fig.
|
|
<figureCitation id="B531F7E1249DC8D6800D1620A99911A1" captionStart="Figure 13" captionStartId="F13" captionText="Figure 13. Phytoliriomyza crepusculum sp. nov. A-C holotype male A habitus B lateral C frontal D paratype male (MK-AG-a 309), dorsal E paratype female (MK-AG- 274), dorsal F-I male genitalia F ejaculatory apodeme, dorsal G whole genitalia, ventral H phallic complex, lateral I epandrium, ventral J puparium K-L female postabdomen K oviscape and spermatheca L tergite 10 M habitat of type locality N-P mined thalli of Dumortiera hirsuta (N type locality O Kiyosumi P Mt. Nabejiri). An arrow in N indicate an puparium." figureDoi="10.3897/zookeys.1133.94530.figure13" httpUri="https://binary.pensoft.net/fig/775386" pageId="0" pageNumber="1">13A</figureCitation>
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|
). Length of ultimate section of vein M4 divided by penultimate section 1.1.
|
|
</paragraph>
|
|
<paragraph id="442654E7F3F3CD3D7E9268B9DEF92998" pageId="0" pageNumber="1">
|
|
<emphasis id="21C7A10B1A643E3C758269A137995B2A" bold="true" italics="true" pageId="0" pageNumber="1">Abdomen</emphasis>
|
|
<emphasis id="5DEBDFB8FD1E871832807331462ABE67" bold="true" pageId="0" pageNumber="1">:</emphasis>
|
|
Abdomen dorsally subshiny brown (Fig.
|
|
<figureCitation id="F8828FA46B87F3A5F0EF834418505A92" captionStart="Figure 13" captionStartId="F13" captionText="Figure 13. Phytoliriomyza crepusculum sp. nov. A-C holotype male A habitus B lateral C frontal D paratype male (MK-AG-a 309), dorsal E paratype female (MK-AG- 274), dorsal F-I male genitalia F ejaculatory apodeme, dorsal G whole genitalia, ventral H phallic complex, lateral I epandrium, ventral J puparium K-L female postabdomen K oviscape and spermatheca L tergite 10 M habitat of type locality N-P mined thalli of Dumortiera hirsuta (N type locality O Kiyosumi P Mt. Nabejiri). An arrow in N indicate an puparium." figureDoi="10.3897/zookeys.1133.94530.figure13" httpUri="https://binary.pensoft.net/fig/775386" pageId="0" pageNumber="1">13D</figureCitation>
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|
).
|
|
<emphasis id="607B1616C97D1728D0B7FF98BFB371E4" bold="true" italics="true" pageId="0" pageNumber="1">Genitalia</emphasis>
|
|
<emphasis id="6BC5292E50948CF97BBE37BDAA576CB3" bold="true" pageId="0" pageNumber="1">:</emphasis>
|
|
(Fig.
|
|
<figureCitation id="C6F661F1A7D4AA853389CEB24A3FB470" captionStart="Figure 13" captionStartId="F13" captionText="Figure 13. Phytoliriomyza crepusculum sp. nov. A-C holotype male A habitus B lateral C frontal D paratype male (MK-AG-a 309), dorsal E paratype female (MK-AG- 274), dorsal F-I male genitalia F ejaculatory apodeme, dorsal G whole genitalia, ventral H phallic complex, lateral I epandrium, ventral J puparium K-L female postabdomen K oviscape and spermatheca L tergite 10 M habitat of type locality N-P mined thalli of Dumortiera hirsuta (N type locality O Kiyosumi P Mt. Nabejiri). An arrow in N indicate an puparium." figureDoi="10.3897/zookeys.1133.94530.figure13" httpUri="https://binary.pensoft.net/fig/775386" pageId="0" pageNumber="1">13F-I</figureCitation>
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|
) Epandrium dark brown, rounded apically; posterior end of inner margin with a stout tubercle-like seta; inner-ventral surface of epandrium with a comb comprising five fused long tubercle-like setae; inner-ventral margin with a row of several short tubercle-like setae (Fig.
|
|
<figureCitation id="3D02022F6881E18C2D3FDC767422CBBA" captionStart="Figure 13" captionStartId="F13" captionText="Figure 13. Phytoliriomyza crepusculum sp. nov. A-C holotype male A habitus B lateral C frontal D paratype male (MK-AG-a 309), dorsal E paratype female (MK-AG- 274), dorsal F-I male genitalia F ejaculatory apodeme, dorsal G whole genitalia, ventral H phallic complex, lateral I epandrium, ventral J puparium K-L female postabdomen K oviscape and spermatheca L tergite 10 M habitat of type locality N-P mined thalli of Dumortiera hirsuta (N type locality O Kiyosumi P Mt. Nabejiri). An arrow in N indicate an puparium." figureDoi="10.3897/zookeys.1133.94530.figure13" httpUri="https://binary.pensoft.net/fig/775386" pageId="0" pageNumber="1">13I</figureCitation>
|
|
). Surstylus rounded, directed inwards, setose apically but without tubercle-like seta. Cercus narrow, setose. Subepandrial sclerite with one pair of posteriorly directed plate-like arms (Fig.
|
|
<figureCitation id="3BBA26926C493143FEDEC97493A1595B" captionStart="Figure 13" captionStartId="F13" captionText="Figure 13. Phytoliriomyza crepusculum sp. nov. A-C holotype male A habitus B lateral C frontal D paratype male (MK-AG-a 309), dorsal E paratype female (MK-AG- 274), dorsal F-I male genitalia F ejaculatory apodeme, dorsal G whole genitalia, ventral H phallic complex, lateral I epandrium, ventral J puparium K-L female postabdomen K oviscape and spermatheca L tergite 10 M habitat of type locality N-P mined thalli of Dumortiera hirsuta (N type locality O Kiyosumi P Mt. Nabejiri). An arrow in N indicate an puparium." figureDoi="10.3897/zookeys.1133.94530.figure13" httpUri="https://binary.pensoft.net/fig/775386" pageId="0" pageNumber="1">13I</figureCitation>
|
|
). Hypandrium slightly sclerotized along outer margin (Fig.
|
|
<figureCitation id="4BA24505E31318E9224947D2F0766222" captionStart="Figure 13" captionStartId="F13" captionText="Figure 13. Phytoliriomyza crepusculum sp. nov. A-C holotype male A habitus B lateral C frontal D paratype male (MK-AG-a 309), dorsal E paratype female (MK-AG- 274), dorsal F-I male genitalia F ejaculatory apodeme, dorsal G whole genitalia, ventral H phallic complex, lateral I epandrium, ventral J puparium K-L female postabdomen K oviscape and spermatheca L tergite 10 M habitat of type locality N-P mined thalli of Dumortiera hirsuta (N type locality O Kiyosumi P Mt. Nabejiri). An arrow in N indicate an puparium." figureDoi="10.3897/zookeys.1133.94530.figure13" httpUri="https://binary.pensoft.net/fig/775386" pageId="0" pageNumber="1">13G</figureCitation>
|
|
). Postgonite bare, goose barnacle-shaped, narrowly rounded, and cleft apically (Fig.
|
|
<figureCitation id="649D91BA20A90BCCE80B8949FF5AE2E2" captionStart="Figure 13" captionStartId="F13" captionText="Figure 13. Phytoliriomyza crepusculum sp. nov. A-C holotype male A habitus B lateral C frontal D paratype male (MK-AG-a 309), dorsal E paratype female (MK-AG- 274), dorsal F-I male genitalia F ejaculatory apodeme, dorsal G whole genitalia, ventral H phallic complex, lateral I epandrium, ventral J puparium K-L female postabdomen K oviscape and spermatheca L tergite 10 M habitat of type locality N-P mined thalli of Dumortiera hirsuta (N type locality O Kiyosumi P Mt. Nabejiri). An arrow in N indicate an puparium." figureDoi="10.3897/zookeys.1133.94530.figure13" httpUri="https://binary.pensoft.net/fig/775386" pageId="0" pageNumber="1">13H</figureCitation>
|
|
). Phallophorus with deep incision below (Fig.
|
|
<figureCitation id="B56B1531AF5B749376A69CEDBBA285EC" captionStart="Figure 13" captionStartId="F13" captionText="Figure 13. Phytoliriomyza crepusculum sp. nov. A-C holotype male A habitus B lateral C frontal D paratype male (MK-AG-a 309), dorsal E paratype female (MK-AG- 274), dorsal F-I male genitalia F ejaculatory apodeme, dorsal G whole genitalia, ventral H phallic complex, lateral I epandrium, ventral J puparium K-L female postabdomen K oviscape and spermatheca L tergite 10 M habitat of type locality N-P mined thalli of Dumortiera hirsuta (N type locality O Kiyosumi P Mt. Nabejiri). An arrow in N indicate an puparium." figureDoi="10.3897/zookeys.1133.94530.figure13" httpUri="https://binary.pensoft.net/fig/775386" pageId="0" pageNumber="1">13G</figureCitation>
|
|
), and articulated with phallapodeme, fused to epiphallus (Fig.
|
|
<figureCitation id="C8CC4E117F093BE654F36ABFADB9485B" captionStart="Figure 13" captionStartId="F13" captionText="Figure 13. Phytoliriomyza crepusculum sp. nov. A-C holotype male A habitus B lateral C frontal D paratype male (MK-AG-a 309), dorsal E paratype female (MK-AG- 274), dorsal F-I male genitalia F ejaculatory apodeme, dorsal G whole genitalia, ventral H phallic complex, lateral I epandrium, ventral J puparium K-L female postabdomen K oviscape and spermatheca L tergite 10 M habitat of type locality N-P mined thalli of Dumortiera hirsuta (N type locality O Kiyosumi P Mt. Nabejiri). An arrow in N indicate an puparium." figureDoi="10.3897/zookeys.1133.94530.figure13" httpUri="https://binary.pensoft.net/fig/775386" pageId="0" pageNumber="1">13H</figureCitation>
|
|
). Basiphallus composing of a left anterodorsal sclerite and a right L-shaped lateral sclerite; former narrow, protruding rightward, the latter protruding ventrally (Fig.
|
|
<figureCitation id="B47001E5CB3868DA9352CF43F9AF69A7" captionStart="Figure 13" captionStartId="F13" captionText="Figure 13. Phytoliriomyza crepusculum sp. nov. A-C holotype male A habitus B lateral C frontal D paratype male (MK-AG-a 309), dorsal E paratype female (MK-AG- 274), dorsal F-I male genitalia F ejaculatory apodeme, dorsal G whole genitalia, ventral H phallic complex, lateral I epandrium, ventral J puparium K-L female postabdomen K oviscape and spermatheca L tergite 10 M habitat of type locality N-P mined thalli of Dumortiera hirsuta (N type locality O Kiyosumi P Mt. Nabejiri). An arrow in N indicate an puparium." figureDoi="10.3897/zookeys.1133.94530.figure13" httpUri="https://binary.pensoft.net/fig/775386" pageId="0" pageNumber="1">13G-H</figureCitation>
|
|
). Hypophallus broad and membranous, supported by one pair of fused sclerites ventrally (Fig.
|
|
<figureCitation id="A6FE70093E1D1BB9A499AD9230700E44" captionStart="Figure 13" captionStartId="F13" captionText="Figure 13. Phytoliriomyza crepusculum sp. nov. A-C holotype male A habitus B lateral C frontal D paratype male (MK-AG-a 309), dorsal E paratype female (MK-AG- 274), dorsal F-I male genitalia F ejaculatory apodeme, dorsal G whole genitalia, ventral H phallic complex, lateral I epandrium, ventral J puparium K-L female postabdomen K oviscape and spermatheca L tergite 10 M habitat of type locality N-P mined thalli of Dumortiera hirsuta (N type locality O Kiyosumi P Mt. Nabejiri). An arrow in N indicate an puparium." figureDoi="10.3897/zookeys.1133.94530.figure13" httpUri="https://binary.pensoft.net/fig/775386" pageId="0" pageNumber="1">13G, H</figureCitation>
|
|
). Mesophallus dark, cylindrical, widest subbasally, slightly shorter than distiphallus (Fig.
|
|
<figureCitation id="BBDB7266E9D2D61A5D016A8CA5669450" captionStart="Figure 13" captionStartId="F13" captionText="Figure 13. Phytoliriomyza crepusculum sp. nov. A-C holotype male A habitus B lateral C frontal D paratype male (MK-AG-a 309), dorsal E paratype female (MK-AG- 274), dorsal F-I male genitalia F ejaculatory apodeme, dorsal G whole genitalia, ventral H phallic complex, lateral I epandrium, ventral J puparium K-L female postabdomen K oviscape and spermatheca L tergite 10 M habitat of type locality N-P mined thalli of Dumortiera hirsuta (N type locality O Kiyosumi P Mt. Nabejiri). An arrow in N indicate an puparium." figureDoi="10.3897/zookeys.1133.94530.figure13" httpUri="https://binary.pensoft.net/fig/775386" pageId="0" pageNumber="1">13H</figureCitation>
|
|
). Distiphallus consists of a pair of moderately long tubules slightly diverging apically; basal half composed of dark narrow lobate sclerite and weaker medial region; distal half cylindrical, dorsally pigmented, with truncated, unpigmented apex (Fig.
|
|
<figureCitation id="748137BEEC0DBD5A5A922829B995B793" captionStart="Figure 13" captionStartId="F13" captionText="Figure 13. Phytoliriomyza crepusculum sp. nov. A-C holotype male A habitus B lateral C frontal D paratype male (MK-AG-a 309), dorsal E paratype female (MK-AG- 274), dorsal F-I male genitalia F ejaculatory apodeme, dorsal G whole genitalia, ventral H phallic complex, lateral I epandrium, ventral J puparium K-L female postabdomen K oviscape and spermatheca L tergite 10 M habitat of type locality N-P mined thalli of Dumortiera hirsuta (N type locality O Kiyosumi P Mt. Nabejiri). An arrow in N indicate an puparium." figureDoi="10.3897/zookeys.1133.94530.figure13" httpUri="https://binary.pensoft.net/fig/775386" pageId="0" pageNumber="1">13H</figureCitation>
|
|
). Ejaculatory apodeme fan-shaped with short broad stalk, broad base, and clear sperm pump (Fig.
|
|
<figureCitation id="98852BAEF41046DB4FB64417E481484E" captionStart="Figure 13" captionStartId="F13" captionText="Figure 13. Phytoliriomyza crepusculum sp. nov. A-C holotype male A habitus B lateral C frontal D paratype male (MK-AG-a 309), dorsal E paratype female (MK-AG- 274), dorsal F-I male genitalia F ejaculatory apodeme, dorsal G whole genitalia, ventral H phallic complex, lateral I epandrium, ventral J puparium K-L female postabdomen K oviscape and spermatheca L tergite 10 M habitat of type locality N-P mined thalli of Dumortiera hirsuta (N type locality O Kiyosumi P Mt. Nabejiri). An arrow in N indicate an puparium." figureDoi="10.3897/zookeys.1133.94530.figure13" httpUri="https://binary.pensoft.net/fig/775386" pageId="0" pageNumber="1">13F</figureCitation>
|
|
).
|
|
</paragraph>
|
|
<paragraph id="86A350A06DBF87BD46061111425166A5" pageId="0" pageNumber="1">
|
|
<emphasis id="326CC858C7F633D1F4727EAE161BEA97" bold="true" pageId="0" pageNumber="1">Female</emphasis>
|
|
(Fig.
|
|
<figureCitation id="236E87DB279A320DB14D6786FE44DED8" captionStart="Figure 13" captionStartId="F13" captionText="Figure 13. Phytoliriomyza crepusculum sp. nov. A-C holotype male A habitus B lateral C frontal D paratype male (MK-AG-a 309), dorsal E paratype female (MK-AG- 274), dorsal F-I male genitalia F ejaculatory apodeme, dorsal G whole genitalia, ventral H phallic complex, lateral I epandrium, ventral J puparium K-L female postabdomen K oviscape and spermatheca L tergite 10 M habitat of type locality N-P mined thalli of Dumortiera hirsuta (N type locality O Kiyosumi P Mt. Nabejiri). An arrow in N indicate an puparium." figureDoi="10.3897/zookeys.1133.94530.figure13" httpUri="https://binary.pensoft.net/fig/775386" pageId="0" pageNumber="1">13E</figureCitation>
|
|
). Similar to male (Fig.
|
|
<figureCitation id="CBF53BCADC058EC45DF689FA904DBAFB" captionStart="Figure 13" captionStartId="F13" captionText="Figure 13. Phytoliriomyza crepusculum sp. nov. A-C holotype male A habitus B lateral C frontal D paratype male (MK-AG-a 309), dorsal E paratype female (MK-AG- 274), dorsal F-I male genitalia F ejaculatory apodeme, dorsal G whole genitalia, ventral H phallic complex, lateral I epandrium, ventral J puparium K-L female postabdomen K oviscape and spermatheca L tergite 10 M habitat of type locality N-P mined thalli of Dumortiera hirsuta (N type locality O Kiyosumi P Mt. Nabejiri). An arrow in N indicate an puparium." figureDoi="10.3897/zookeys.1133.94530.figure13" httpUri="https://binary.pensoft.net/fig/775386" pageId="0" pageNumber="1">13E</figureCitation>
|
|
). Wing length 1.4-1.8 mm.
|
|
<emphasis id="76B5328614B87E5FE713C3F5B09C57C3" bold="true" italics="true" pageId="0" pageNumber="1">Postabdomen</emphasis>
|
|
<emphasis id="2875BC98518B54D49A20998CB7E5D314" bold="true" pageId="0" pageNumber="1">:</emphasis>
|
|
(Fig.
|
|
<figureCitation id="DFED9A8010C52CCF4132B6B1ADE27D2F" captionStart="Figure 13" captionStartId="F13" captionText="Figure 13. Phytoliriomyza crepusculum sp. nov. A-C holotype male A habitus B lateral C frontal D paratype male (MK-AG-a 309), dorsal E paratype female (MK-AG- 274), dorsal F-I male genitalia F ejaculatory apodeme, dorsal G whole genitalia, ventral H phallic complex, lateral I epandrium, ventral J puparium K-L female postabdomen K oviscape and spermatheca L tergite 10 M habitat of type locality N-P mined thalli of Dumortiera hirsuta (N type locality O Kiyosumi P Mt. Nabejiri). An arrow in N indicate an puparium." figureDoi="10.3897/zookeys.1133.94530.figure13" httpUri="https://binary.pensoft.net/fig/775386" pageId="0" pageNumber="1">13K, L</figureCitation>
|
|
) Oviscape brown sparsely setigerous on lateral sides (Fig.
|
|
<figureCitation id="9193199CDEFC2019FCA431C8E586E612" captionStart="Figure 13" captionStartId="F13" captionText="Figure 13. Phytoliriomyza crepusculum sp. nov. A-C holotype male A habitus B lateral C frontal D paratype male (MK-AG-a 309), dorsal E paratype female (MK-AG- 274), dorsal F-I male genitalia F ejaculatory apodeme, dorsal G whole genitalia, ventral H phallic complex, lateral I epandrium, ventral J puparium K-L female postabdomen K oviscape and spermatheca L tergite 10 M habitat of type locality N-P mined thalli of Dumortiera hirsuta (N type locality O Kiyosumi P Mt. Nabejiri). An arrow in N indicate an puparium." figureDoi="10.3897/zookeys.1133.94530.figure13" httpUri="https://binary.pensoft.net/fig/775386" pageId="0" pageNumber="1">13K</figureCitation>
|
|
). Tergite 10 trifurcate, laterally uniting narrow pleural sclerites (Fig.
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|
<figureCitation id="C23016D3F25747759B15E797F3CD3354" captionStart="Figure 13" captionStartId="F13" captionText="Figure 13. Phytoliriomyza crepusculum sp. nov. A-C holotype male A habitus B lateral C frontal D paratype male (MK-AG-a 309), dorsal E paratype female (MK-AG- 274), dorsal F-I male genitalia F ejaculatory apodeme, dorsal G whole genitalia, ventral H phallic complex, lateral I epandrium, ventral J puparium K-L female postabdomen K oviscape and spermatheca L tergite 10 M habitat of type locality N-P mined thalli of Dumortiera hirsuta (N type locality O Kiyosumi P Mt. Nabejiri). An arrow in N indicate an puparium." figureDoi="10.3897/zookeys.1133.94530.figure13" httpUri="https://binary.pensoft.net/fig/775386" pageId="0" pageNumber="1">13L</figureCitation>
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). Cercus with two stout, apical, trichoid sensilla, ca. 1/2 length of cercus (Fig.
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<figureCitation id="CBDC102AA2F68C4B9427EB0775F627C6" captionStart="Figure 13" captionStartId="F13" captionText="Figure 13. Phytoliriomyza crepusculum sp. nov. A-C holotype male A habitus B lateral C frontal D paratype male (MK-AG-a 309), dorsal E paratype female (MK-AG- 274), dorsal F-I male genitalia F ejaculatory apodeme, dorsal G whole genitalia, ventral H phallic complex, lateral I epandrium, ventral J puparium K-L female postabdomen K oviscape and spermatheca L tergite 10 M habitat of type locality N-P mined thalli of Dumortiera hirsuta (N type locality O Kiyosumi P Mt. Nabejiri). An arrow in N indicate an puparium." figureDoi="10.3897/zookeys.1133.94530.figure13" httpUri="https://binary.pensoft.net/fig/775386" pageId="0" pageNumber="1">13L</figureCitation>
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). Spermathecae semi-orbicular, with truncate proximal ends (Fig.
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<figureCitation id="B310F142E831153EF524F78CDD07D8C9" captionStart="Figure 13" captionStartId="F13" captionText="Figure 13. Phytoliriomyza crepusculum sp. nov. A-C holotype male A habitus B lateral C frontal D paratype male (MK-AG-a 309), dorsal E paratype female (MK-AG- 274), dorsal F-I male genitalia F ejaculatory apodeme, dorsal G whole genitalia, ventral H phallic complex, lateral I epandrium, ventral J puparium K-L female postabdomen K oviscape and spermatheca L tergite 10 M habitat of type locality N-P mined thalli of Dumortiera hirsuta (N type locality O Kiyosumi P Mt. Nabejiri). An arrow in N indicate an puparium." figureDoi="10.3897/zookeys.1133.94530.figure13" httpUri="https://binary.pensoft.net/fig/775386" pageId="0" pageNumber="1">13K</figureCitation>
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).
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</paragraph>
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<paragraph id="1A70992AEB70094016FE8323BEA6B4BF" pageId="0" pageNumber="1">
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|
<emphasis id="8FFC6917BDCD000F27F8E69920F87DB8" bold="true" pageId="0" pageNumber="1">Immatures.</emphasis>
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|
(Fig.
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|
<figureCitation id="6769CBD5F8DBB137E05E770B669B7985" captionStart="Figure 13" captionStartId="F13" captionText="Figure 13. Phytoliriomyza crepusculum sp. nov. A-C holotype male A habitus B lateral C frontal D paratype male (MK-AG-a 309), dorsal E paratype female (MK-AG- 274), dorsal F-I male genitalia F ejaculatory apodeme, dorsal G whole genitalia, ventral H phallic complex, lateral I epandrium, ventral J puparium K-L female postabdomen K oviscape and spermatheca L tergite 10 M habitat of type locality N-P mined thalli of Dumortiera hirsuta (N type locality O Kiyosumi P Mt. Nabejiri). An arrow in N indicate an puparium." figureDoi="10.3897/zookeys.1133.94530.figure13" httpUri="https://binary.pensoft.net/fig/775386" pageId="0" pageNumber="1">13J</figureCitation>
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) Puparium internal, slender, and pale brown.
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</paragraph>
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</subSubSection>
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<subSubSection id="36CB0E06F63FB4FE6F1717D757023EC7" pageId="0" pageNumber="1" type="etymology">
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<paragraph id="D3EA0F79551574E3C7FEC0623EC034AB" pageId="0" pageNumber="1">Etymology.</paragraph>
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<paragraph id="61CF4E37B734BB790345AD784A87115E" pageId="0" pageNumber="1">
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|
The specific name (
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<emphasis id="97788C07FBC7ADE76A537A260EE96181" italics="true" pageId="0" pageNumber="1">crepusculum</emphasis>
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= twilight) refers to the brownish yellow patch on the scutum.
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</paragraph>
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</subSubSection>
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<subSubSection id="E9B7D791BFED5B4823E3284BA626690B" pageId="0" pageNumber="1" type="japanese name">
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|
<paragraph id="ABA7125E993478AA2E553552EC8C7C73" pageId="0" pageNumber="1">Japanese name.</paragraph>
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<paragraph id="8A47254F5130527D7C6933C12B334711" pageId="0" pageNumber="1">Yuuzuki-kezenigoke-hamoguribae.</paragraph>
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</subSubSection>
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<subSubSection id="011CF5626486769895E60690F02D5205" pageId="0" pageNumber="1" type="host plant">
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<paragraph id="86EAD2CE2B241C4012E9C782EA598519" pageId="0" pageNumber="1">Host plant.</paragraph>
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<paragraph id="3AEA8CCE5A85D9BFA5C195E45E814972" pageId="0" pageNumber="1">
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|
<taxonomicName id="2D345AD3DDC8DAD77204B9629A3CDBF9" class="Marchantiopsida" family="Dumortieraceae" genus="Dumortiera" kingdom="Plantae" lsidName="Dumortiera hirsuta" order="Marchantiales" pageId="0" pageNumber="1" phylum="Marchantiophyta" rank="species" species="hirsuta">
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|
<emphasis id="CC0E8CAB062BC9832CB4077832746C01" italics="true" pageId="0" pageNumber="1">Dumortiera hirsuta</emphasis>
|
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</taxonomicName>
|
|
(
|
|
<taxonomicName id="88B7309413BA58B7F9D6F70D4C81EA02" authorityName="C.G.D.Nees" authorityYear="1824" class="Marchantiopsida" family="Dumortieraceae" kingdom="Plantae" lsidName="" order="Marchantiales" pageId="0" pageNumber="1" phylum="Marchantiophyta" rank="family">Dumortieraceae</taxonomicName>
|
|
).
|
|
</paragraph>
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|
</subSubSection>
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|
<subSubSection id="2A2E9FA52CF8B642369C0BB033ADCA62" pageId="0" pageNumber="1" type="mine">
|
|
<paragraph id="15D4C638B8C662358C71159DB93ECF35" pageId="0" pageNumber="1">Mine.</paragraph>
|
|
<paragraph id="64986B2633092D24DD549C4E7762B8D5" pageId="0" pageNumber="1">
|
|
(Fig.
|
|
<figureCitation id="18D44889FB07D406C2150317395518F6" captionStart="Figure 13" captionStartId="F13" captionText="Figure 13. Phytoliriomyza crepusculum sp. nov. A-C holotype male A habitus B lateral C frontal D paratype male (MK-AG-a 309), dorsal E paratype female (MK-AG- 274), dorsal F-I male genitalia F ejaculatory apodeme, dorsal G whole genitalia, ventral H phallic complex, lateral I epandrium, ventral J puparium K-L female postabdomen K oviscape and spermatheca L tergite 10 M habitat of type locality N-P mined thalli of Dumortiera hirsuta (N type locality O Kiyosumi P Mt. Nabejiri). An arrow in N indicate an puparium." figureDoi="10.3897/zookeys.1133.94530.figure13" httpUri="https://binary.pensoft.net/fig/775386" pageId="0" pageNumber="1">13N-P</figureCitation>
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|
) Larvae mine the midrib of the thallus in early instars, later construct radiate mines by coming out from the midrib, and pupate in the mines. Puparium is smooth and pale brown, posterior spiracles projecting (Fig.
|
|
<figureCitation id="E5212FA9B94625EEC7740D3E835F8FAD" captionStart="Figure 13" captionStartId="F13" captionText="Figure 13. Phytoliriomyza crepusculum sp. nov. A-C holotype male A habitus B lateral C frontal D paratype male (MK-AG-a 309), dorsal E paratype female (MK-AG- 274), dorsal F-I male genitalia F ejaculatory apodeme, dorsal G whole genitalia, ventral H phallic complex, lateral I epandrium, ventral J puparium K-L female postabdomen K oviscape and spermatheca L tergite 10 M habitat of type locality N-P mined thalli of Dumortiera hirsuta (N type locality O Kiyosumi P Mt. Nabejiri). An arrow in N indicate an puparium." figureDoi="10.3897/zookeys.1133.94530.figure13" httpUri="https://binary.pensoft.net/fig/775386" pageId="0" pageNumber="1">13J</figureCitation>
|
|
).
|
|
</paragraph>
|
|
</subSubSection>
|
|
<subSubSection id="198EBCE248D25B501C7E7DA17DCD8B6C" pageId="0" pageNumber="1" type="biological notes">
|
|
<paragraph id="DFA3351FCE4462DCF1064858A4F69862" pageId="0" pageNumber="1">Biological notes.</paragraph>
|
|
<paragraph id="3705E2B832D98B0CF99CF7413137B674" pageId="0" pageNumber="1">
|
|
The main habitats of this species are along streams in warm temperate forests and cool temperate deciduous forests (Fig.
|
|
<figureCitation id="C5D46FD7DD56F1CF03D04EA3C1541362" captionStart="Figure 13" captionStartId="F13" captionText="Figure 13. Phytoliriomyza crepusculum sp. nov. A-C holotype male A habitus B lateral C frontal D paratype male (MK-AG-a 309), dorsal E paratype female (MK-AG- 274), dorsal F-I male genitalia F ejaculatory apodeme, dorsal G whole genitalia, ventral H phallic complex, lateral I epandrium, ventral J puparium K-L female postabdomen K oviscape and spermatheca L tergite 10 M habitat of type locality N-P mined thalli of Dumortiera hirsuta (N type locality O Kiyosumi P Mt. Nabejiri). An arrow in N indicate an puparium." figureDoi="10.3897/zookeys.1133.94530.figure13" httpUri="https://binary.pensoft.net/fig/775386" pageId="0" pageNumber="1">13M</figureCitation>
|
|
), and this species was sympatric with
|
|
<taxonomicName id="6342E7B13303B8164091D08969FD9401" class="Marchantiopsida" family="Aytoniaceae" kingdom="Plantae" lsidName="P. dumortierae" order="Marchantiales" pageId="0" pageNumber="1" phylum="Marchantiophyta" rank="species" species="dumortierae">
|
|
<emphasis id="192BAC9D4D992DEF62A7F36710A7F4C5" italics="true" pageId="0" pageNumber="1">P. dumortierae</emphasis>
|
|
</taxonomicName>
|
|
in some localities. Our rearing records suggest that this species is univoltine, and that adults emerge from overwintered pupae in spring.
|
|
</paragraph>
|
|
</subSubSection>
|
|
<subSubSection id="B014188E4B7A70EF4E52BD08AED6B465" pageId="0" pageNumber="1" type="distribution">
|
|
<paragraph id="452F4F5641B27953DFDEBC72FA244EB7" pageId="0" pageNumber="1">Distribution.</paragraph>
|
|
<paragraph id="7FB388B68A155567B719EAEDBCDC122F" pageId="0" pageNumber="1">
|
|
Japan: Honshu, Shikoku, Kyushu, Tsushima Island (Fig.
|
|
<figureCitation id="65B29D722F399D0A53139D2325743792" captionStart="Figure 12" captionStartId="F12" captionText="Figure 12. Locality records of three Phytoliriomyza species associated with Dumortiera spp.: P. dumortierae and P. crepusculum." figureDoi="10.3897/zookeys.1133.94530.figure12" httpUri="https://binary.pensoft.net/fig/775385" pageId="0" pageNumber="1">12</figureCitation>
|
|
).
|
|
</paragraph>
|
|
</subSubSection>
|
|
<subSubSection id="BB38FF83A6B49D627D62BF65C5859EAB" pageId="0" pageNumber="1" type="remarks">
|
|
<paragraph id="3F4C8FA119FC267DC69754C9F10F987B" pageId="0" pageNumber="1">Remarks.</paragraph>
|
|
<paragraph id="CC66BD0EEEA10902BD7EF22E615E5C48" pageId="0" pageNumber="1">
|
|
Although this species used the same host plant as
|
|
<taxonomicName id="2E422001514DC85D8C134E7CED932BFC" class="Marchantiopsida" family="Aytoniaceae" kingdom="Plantae" lsidName="P. dumortierae" order="Marchantiales" pageId="0" pageNumber="1" phylum="Marchantiophyta" rank="species" species="dumortierae">
|
|
<emphasis id="39DC6D3ADCBC31C3436B05A75E662D8C" italics="true" pageId="0" pageNumber="1">P. dumortierae</emphasis>
|
|
</taxonomicName>
|
|
, these two species were not sympatric in any localities. This species resembles
|
|
<taxonomicName id="38F0FED6B6C2D3FEB3EE90DC75C81031" class="Marchantiopsida" family="Aytoniaceae" kingdom="Plantae" lsidName="P. arcus" order="Marchantiales" pageId="0" pageNumber="1" phylum="Marchantiophyta" rank="species" species="arcus">
|
|
<emphasis id="D7783B85143569B37B1F64BA7740696E" italics="true" pageId="0" pageNumber="1">P. arcus</emphasis>
|
|
</taxonomicName>
|
|
,
|
|
<taxonomicName id="C7F45CAD03AB912F8248B6916D35DE2D" class="Marchantiopsida" family="Aytoniaceae" kingdom="Plantae" lsidName="P. plagiochasmatos" order="Marchantiales" pageId="0" pageNumber="1" phylum="Marchantiophyta" rank="species" species="plagiochasmatos">
|
|
<emphasis id="960080EB210B98D4FF3C56E03F68FA9C" italics="true" pageId="0" pageNumber="1">P. plagiochasmatos</emphasis>
|
|
</taxonomicName>
|
|
and
|
|
<taxonomicName id="55EA6E1FAB275892DDDBC33B3C32CA11" class="Marchantiopsida" family="Aytoniaceae" kingdom="Plantae" lsidName="P. falcata" order="Marchantiales" pageId="0" pageNumber="1" phylum="Marchantiophyta" rank="species" species="falcata">
|
|
<emphasis id="45EFB05CA9B8B161601D71F7143CFEAE" italics="true" pageId="0" pageNumber="1">P. falcata</emphasis>
|
|
</taxonomicName>
|
|
in having a pair of brown lateral bands on the scutum; it is distinguished from them by the brownish yellow color of the medial mark on scutum (light yellow in the other species).
|
|
</paragraph>
|
|
</subSubSection>
|
|
<subSubSection id="2726F7733EB0597B7B60789D349E2F38" pageId="0" pageNumber="1" type="species associated with plagiochasma">
|
|
<paragraph id="79ABA42CA296CA0006EC7844F6844BD5" pageId="0" pageNumber="1">
|
|
Species associated with
|
|
<taxonomicName id="EB51B3CA2C05DBAB5425957950C9D5E0" authorityName="Pomel" authorityYear="1883" class="Marchantiopsida" family="Aytoniaceae" genus="Plagiochasma" kingdom="Plantae" lsidName="Plagiochasma" order="Marchantiales" pageId="0" pageNumber="1" phylum="Marchantiophyta" rank="genus">
|
|
<emphasis id="51EAC4212926F0463A3617419A827C29" italics="true" pageId="0" pageNumber="1">Plagiochasma</emphasis>
|
|
</taxonomicName>
|
|
</paragraph>
|
|
</subSubSection>
|
|
</treatment>
|
|
</document> |