treatments-xml/data/CF/D4/60/CFD460610D325A180374B6538C0CC74B.xml

<document ID-DOI="http://dx.doi.org/10.3897/zookeys.699.12031" ID-GBIF-Dataset="dbf6bb9f-7ea4-4fa3-83ee-f58263427e39" ID-PMC="PMC5673864" ID-Pensoft-Pub="1313-2970-699-1" ID-PubMed="29200927" ID-ZBK="0F92401F3F7C4896AD9D72BC84348C7D" ModsDocAuthor="" ModsDocDate="2017" ModsDocID="1313-2970-699-1" ModsDocOrigin="ZooKeys 699" ModsDocTitle="A taxonomic revision of Liogenys occurring in Brazil with an interactive key and remarks on New World Diplotaxini (Coleoptera, Melolonthidae)" checkinTime="1505836577638" checkinUser="pensoft" docAuthor="Cherman, Mariana Alejandra, Mise, Kleber Makoto, Moron †, Miguel Angel, Vaz-de-Mello, Fernando Z. &amp; Almeida, Lucia Massutti de" docDate="2017" docId="CFD460610D325A180374B6538C0CC74B" docLanguage="en" docName="ZooKeys 699: 1-120" docOrigin="ZooKeys 699" docSource="http://dx.doi.org/10.3897/zookeys.699.12031" docTitle="Liogenys acutidens Moser 1919" docType="treatment" docVersion="4" lastPageNumber="12" masterDocId="FFD9636FB945C41DFF8AB133FFFA9D76" masterDocTitle="A taxonomic revision of Liogenys occurring in Brazil with an interactive key and remarks on New World Diplotaxini (Coleoptera, Melolonthidae)" masterLastPageNumber="120" masterPageNumber="1" pageNumber="11" updateTime="1668164813467" updateUser="ExternalLinkService">
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<mods:title>A taxonomic revision of Liogenys occurring in Brazil with an interactive key and remarks on New World Diplotaxini (Coleoptera, Melolonthidae)</mods:title>
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<mods:name type="personal">
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<mods:roleTerm>Author</mods:roleTerm>
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<mods:namePart>Cherman, Mariana Alejandra</mods:namePart>
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<mods:name type="personal">
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<mods:roleTerm>Author</mods:roleTerm>
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<mods:namePart>Mise, Kleber Makoto</mods:namePart>
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<mods:namePart>Moron †, Miguel Angel</mods:namePart>
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<mods:roleTerm>Author</mods:roleTerm>
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<mods:namePart>Vaz-de-Mello, Fernando Z.</mods:namePart>
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<mods:name type="personal">
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<mods:roleTerm>Author</mods:roleTerm>
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<mods:namePart>Almeida, Lucia Massutti de</mods:namePart>
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<mods:typeOfResource>text</mods:typeOfResource>
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<mods:title>ZooKeys</mods:title>
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<mods:date>2017</mods:date>
<mods:detail type="volume">
<mods:number>699</mods:number>
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<mods:url>http://dx.doi.org/10.3897/zookeys.699.12031</mods:url>
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<mods:classification>journal article</mods:classification>
<mods:identifier type="DOI">http://dx.doi.org/10.3897/zookeys.699.12031</mods:identifier>
<mods:identifier type="Pensoft-Pub">1313-2970-699-1</mods:identifier>
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<treatment ID-GBIF-Taxon="134311168" LSID="urn:lsid:plazi:treatment:CFD460610D325A180374B6538C0CC74B" httpUri="http://treatment.plazi.org/id/CFD460610D325A180374B6538C0CC74B" lastPageId="11" lastPageNumber="12" pageId="10" pageNumber="11">
<subSubSection pageId="10" pageNumber="11" type="nomenclature">
<paragraph pageId="10" pageNumber="11">
<taxonomicName authority="Moser, 1919" authorityName="Moser" authorityYear="1919" class="Insecta" family="Melolonthidae" genus="Liogenys" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Liogenys acutidens" order="Coleoptera" pageId="10" pageNumber="11" phylum="Arthropoda" rank="species" species="acutidens">Liogenys acutidens Moser, 1919</taxonomicName>
Figs 56, 93
</paragraph>
</subSubSection>
<subSubSection pageId="10" pageNumber="11" type="reference_group">
<paragraph pageId="10" pageNumber="11">
<taxonomicName class="Insecta" family="Melolonthidae" genus="Liogenys" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Liogenys acutidens" order="Coleoptera" pageId="10" pageNumber="11" phylum="Arthropoda" rank="species" species="acutidens">Liogenys acutidens</taxonomicName>
Moser, 1919: 14 (orig. desc.); 
<bibRefCitation author="Blackwelder, RE" journalOrPublisher="United States National Museum Bulletin" pageId="103" pageNumber="104" pagination="189 - 341" title="Checklist of the coleopterous insects of Mexico, Central America, the West Indies, and South America." volume="185" year="1944">Blackwelder 1944</bibRefCitation>
: 228 (check.); 
<bibRefCitation author="Frey, G" journalOrPublisher="Frey" pageId="105" pageNumber="106" pagination="36 - 64" title="Betimmumgstabelle und Revision der Gattung Liogenys Guerrez. (Coleoptera-Melolonthinae-Macrodactylini). Entomologischen Arbeiten aus dem Museum G." volume="20" year="1969">Frey 1969</bibRefCitation>
: 47 (key); 
<bibRefCitation author="Evans, AV" journalOrPublisher="Zootaxa" pageId="104" pageNumber="105" pagination="1 - 458" title="A checklist of the New World chafers (Coleoptera: Scarabaeidae: Melolonthinae)." url="https://doi.org/10.11646/zootaxa.211.1.1" volume="211" year="2003">Evans 2003</bibRefCitation>
: 207 (check.); 
<bibRefCitation author="Evans, AV" journalOrPublisher="Zootaxa" pageId="104" pageNumber="105" url="http://digitalcommons.unl.edu/entomologypapers/2" year="2005">Evans and Smith 2005</bibRefCitation>
: 171 (check.); 
<bibRefCitation author="Evans, AV" journalOrPublisher="Zootaxa" pageId="104" pageNumber="105" url="http://museum.unl.edu/research/entomology/SSSA/nwmelos.htm" year="2009">Evans and Smith 2009</bibRefCitation>
: 175 (check.)
</paragraph>
</subSubSection>
<subSubSection pageId="10" pageNumber="11" type="type material">
<paragraph pageId="10" pageNumber="11">Type material.</paragraph>
<paragraph pageId="10" pageNumber="11">
<taxonomicName class="Insecta" family="Melolonthidae" genus="Liogenys" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Liogenys acutidens" order="Coleoptera" pageId="10" pageNumber="11" phylum="Arthropoda" rank="species" species="acutidens">Liogenys acutidens</taxonomicName>
male holotype (ZMHB): [white printed] 
<normalizedToken originalValue="“Brasília">&quot;Brasilia</normalizedToken>
/ [handwritten] Cuyaba&quot;, [white handwritten] &quot; 
<taxonomicName class="Insecta" family="Melolonthidae" genus="Liogenys" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Liogenys" order="Coleoptera" pageId="10" pageNumber="11" phylum="Arthropoda" rank="genus">Liogenys</taxonomicName>
/ 
<taxonomicName lsidName="acutidens" pageId="10" pageNumber="11" rank="species" species="acutidens">acutidens</taxonomicName>
/Mos./Typen&quot;, [light red printed] 
<normalizedToken originalValue="“Typus”">&quot;Typus&quot;</normalizedToken>
, &quot; 
<taxonomicName class="Insecta" family="Melolonthidae" genus="Liogenys" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Liogenys" order="Coleoptera" pageId="10" pageNumber="11" phylum="Arthropoda" rank="genus">Liogenys</taxonomicName>
/ 
<taxonomicName lsidName="acutidens" pageId="10" pageNumber="11" rank="species" species="acutidens">acutidens</taxonomicName>
/Mos&quot;, [red printed] &quot;HOLOTYPUS/ 
<taxonomicName class="Insecta" family="Melolonthidae" genus="Liogenys" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Liogenys" order="Coleoptera" pageId="10" pageNumber="11" phylum="Arthropoda" rank="genus">Liogenys</taxonomicName>
/ 
<taxonomicName lsidName="acutidens" pageId="10" pageNumber="11" rank="species" species="acutidens">acutidens</taxonomicName>
Moser, 1919/ labelled by MNHUB 2011&quot;. Genitalia mounted.
</paragraph>
</subSubSection>
<subSubSection pageId="10" pageNumber="11" type="type material">
<paragraph pageId="10" pageNumber="11">Non-type material.</paragraph>
<paragraph pageId="10" pageNumber="11">
BRAZIL. MT: 
<normalizedToken originalValue="Cuiabá">Cuiaba</normalizedToken>
, without date and collector, 1 ex. (ZMHB); MG: 
<normalizedToken originalValue="Unaí">Unai</normalizedToken>
(Faz. 
<normalizedToken originalValue="Bolívia">Bolivia</normalizedToken>
), 22-24/X/1964, Exp. Dep. Zool. col., 1 ex. (MZSP).
</paragraph>
</subSubSection>
<subSubSection pageId="10" pageNumber="11" type="diagnosis">
<paragraph pageId="10" pageNumber="11">Diagnosis.</paragraph>
<paragraph pageId="10" pageNumber="11">Body yellowish brown; elongate; elytra testaceous, lighter in color than pronotum; clypeal emargination wide and rounded; outer sides of anterior teeth parallel; lateral margin convex and slightly produced; distance between clypeal lateral projection and anterior tooth equal to basal width of anterior tooth, distance between clypeal lateral projection and anterior margin of eye shorter than one eye, obtuse angle between outer side of anterior teeth and clypeal lateral projection; mesotibia quadrate in cross section; pygidium convex, bristled throughout, reticulated punctures; in males, inner margins of parameres convergent; subapical projections on outer margins; parameres strongly narrowed between subapical projections and apex; apex spatula-like, sharp edges, curved outwards, not reaching the level of the parameres outer margin (Fig. 56F).</paragraph>
<caption pageId="10" pageNumber="11">
<paragraph pageId="10" pageNumber="11">
Figure 56. 
<taxonomicName class="Insecta" family="Melolonthidae" genus="Liogenys" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Liogenys acutidens" order="Coleoptera" pageId="10" pageNumber="11" phylum="Arthropoda" rank="species" species="acutidens">Liogenys acutidens</taxonomicName>
Moser. A Dorsal view B Lateral view C Frontal view D Clypeus and pronotum E Pygidium F Parameres, dorsal view G Parameres, lateral view.
</paragraph>
</caption>
</subSubSection>
<subSubSection lastPageId="11" lastPageNumber="12" pageId="10" pageNumber="11" type="redescription">
<paragraph pageId="10" pageNumber="11">Redescription.</paragraph>
<paragraph lastPageId="11" lastPageNumber="12" pageId="10" pageNumber="11">
Length: 8.0-9.0 mm; width 3.7-4.2 mm. Yellowish to testaceous. Head: distance between eyes nearly twice the width of one eye; frons equal in length to clypeus; clypeus bristled anteriorly, emargination shallow, wide and rounded; outer sides of anterior teeth parallel; outer margin of anterior teeth shorter than the eye; lateral margin convex and slightly produced; distance between clypeal lateral projection and anterior tooth equal to the basal width of anterior tooth, distance between clypeal lateral projection and anterior margin of eye shorter than one eye, obtuse angle between outer side of anterior teeth and clypeal lateral projection; canthus not exceeding the outer margin of the eye; distal maxillary palpomere, maximum width less than twice width of apex; fovea deep, extending up to the transverse midline of the palpomere; labium transversely carinated, as wide as it is long; antenna 10-articulated, lamellae lighter in color than flagellum, and equal in length. Thorax: anterior margin of pronotum straight; maximum length of pronotum exceeding the length of tarsomeres I, II and III together; disc glabrous, punctures sparse and fine; pronotal posterior corners sub-angled, obtuse; proepisternum with short bristles; mesepisternum scaly, as are the sides of metasternum, also with few long bristles on the anterior margin; distance between meso- and metacoxae up to twice the metacoxa length; scutellum triangular, smooth or basally punctured at the sides. Elytra: shiny, glabrous, uniform yellowish to testaceous, barely lighter in color than the pronotum; elytra more than three times longer than the pronotum; elytral suture slightly darker than elytron and not elevated; all four elytral ridges barely noticeable. Legs: procoxa, sparse scales on infra-carinal surface, punctures visible at 12 
<normalizedToken originalValue="×">x</normalizedToken>
magnification; three protibial teeth, middle and apical equal in size, distance between basal and middle teeth slightly longer than between middle and apical; protibial inner apical spur present; mesofemural disc setose, mesotibia quadrate in cross section, disc finely sculptured, two mesotibial transverse carinae, the apical one incomplete; basal apophysis of metacoxa produced beyond the outer margin of trochanter; inner margin of metatibia carinated towards apex, apical inner surface setose; metatibial disc finely sculptured; a metatibial transverse carina present posteriorly and posterior discontinuous longitudinal carina; basal metatarsomere and tarsomere II equal in size, protarsomere II short and wide; pro- and mesotarsomeres I 
<pageBreakToken pageId="11" pageNumber="12" start="start">to</pageBreakToken>
IV enlarged, protarsomeres slightly wider than the mesotarsomeres, less than twice as wide as the metatarsi; claw bifid, symmetrical, superior tooth narrower than the inferior and equal in length; distance between teeth shorter than the inferior tooth. Abdomen: ventrites bristled and on sides also scaly; propygidium visible, bristled and scaly; pygidium convex, sub-trapezoidal, wide; pygidial width not exceeding distance between spiracles of propygidium; pygidial disc bristled throughout, reticulated punctures; pygidial apex sub-quadrate. Parameres: width of basal region equal to the parameres together at its maximum width; parameral split at 2/3; inner margins of parameres convergent; outer margins with subapical projections; strongly narrowed between subapical projections and apex; apex spatula-like, edges sharp, curved outwards, not reaching the level of the parameral outer margin (Fig. 56F). In lateral view parameres concave; apex slightly curved downwards (Fig. 56G).
</paragraph>
</subSubSection>
<subSubSection pageId="11" pageNumber="12" type="type-locality">
<paragraph pageId="11" pageNumber="12">Type-locality.</paragraph>
<paragraph pageId="11" pageNumber="12">
BRAZIL, 
<normalizedToken originalValue="Cuiabá">Cuiaba</normalizedToken>
, Mato Grosso.
</paragraph>
</subSubSection>
<subSubSection pageId="11" pageNumber="12" type="distribution">
<paragraph pageId="11" pageNumber="12">Geographical distribution.</paragraph>
<paragraph pageId="11" pageNumber="12">BRAZIL (MT, MG).</paragraph>
</subSubSection>
<subSubSection pageId="11" pageNumber="12" type="remarks">
<paragraph pageId="11" pageNumber="12">Remarks.</paragraph>
<paragraph pageId="11" pageNumber="12">
<taxonomicName class="Insecta" family="Melolonthidae" genus="Liogenys" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Liogenys acutidens" order="Coleoptera" pageId="11" pageNumber="12" phylum="Arthropoda" rank="species" species="acutidens">Liogenys acutidens</taxonomicName>
resembles 
<taxonomicName lsidName="L. bidenticeps" pageId="11" pageNumber="12" rank="species" species="bidenticeps">L. bidenticeps</taxonomicName>
(Fig. 58) in size and color, however, 
<taxonomicName lsidName="L. acutidens" pageId="11" pageNumber="12" rank="species" species="acutidens">L. acutidens</taxonomicName>
elytra are somewhat lighter in color; the clypeus is bristled anteriorly; the clypeal teeth are sharper and slightly longer-, the lateral margin of clypeus is slightly projected; the meso- and metatibia are quadrate in cross section with discs both finely sculptured; the mesotibial apical carina is incomplete; the pygidium is convex with reticulated punctures on disc alike 
<taxonomicName lsidName="L. santaecrucis" pageId="11" pageNumber="12" rank="species" species="santaecrucis">L. santaecrucis</taxonomicName>
(Fig. 71) and protarsomeres wider. Female of 
<taxonomicName lsidName="L. acutidens" pageId="11" pageNumber="12" rank="species" species="acutidens">L. acutidens</taxonomicName>
remains unknown.
</paragraph>
</subSubSection>
</treatment>
</document>