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<document id="4B9B6B448F1FCE757AAADE6F171D5244" ID-DOI="10.11646/zootaxa.5374.1.1" ID-ISSN="1175-5326" ID-Zenodo-Dep="10148159" ID-ZooBank="072341AB-7693-4DF3-9286-D69D0BBD8FAD" IM.bibliography_approvedBy="carolina" IM.illustrations_approvedBy="carolina" IM.materialsCitations_approvedBy="carolina" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="GgImagineBatch" IM.taxonomicNames_approvedBy="carolina" IM.treatments_approvedBy="carolina" checkinTime="1700206341287" checkinUser="plazi" docAuthor="MACFARLAN, DONALD ALEXANDER BANKIER" docDate="2023" docId="073F87ADFFA35A6DFF4BFBD6FC86FE93" docLanguage="en" docName="zootaxa.5374.1.1.pdf" docOrigin="Zootaxa 5374 (1)" docSource="https://www.mapress.com/zt/article/download/zootaxa.5374.1.1/52274" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Rhaetina rainei MACFARLAN 2023, n. sp." docType="treatment" docVersion="6" lastPageNumber="17" masterDocId="FB06FFD5FFAF5A7DFFDCFFF9FFCCFFF6" masterDocTitle="Otapirian (Rhaetian) Terebratulida (Brachiopoda) of Zealandia" masterLastPageNumber="34" masterPageNumber="1" pageNumber="13" updateTime="1706309636962" updateUser="ExternalLinkService" zenodo-license-document="CLOSED">
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<mods:title id="9604B4EB2CF3194F946E0061549072C4">Otapirian (Rhaetian) Terebratulida (Brachiopoda) of Zealandia</mods:title>
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<taxonomicName id="48964D38FFA35A71FF4BFBD6FE99FBBF" authority="MACFARLAN, 2023" authorityName="MACFARLAN" authorityYear="2023" box="[151,341,1071,1098]" class="Rhynchonellata" family="Dielasmatidae" genus="Rhaetina" higherTaxonomySource="GBIF" kingdom="Animalia" order="Terebratulida" pageId="12" pageNumber="13" phylum="Brachiopoda" rank="species" species="rainei" status="sp. nov.">
<emphasis id="BDE2EAA9FFA35A71FF4BFBD6FE99FBBF" bold="true" box="[151,341,1071,1098]" italics="true" pageId="12" pageNumber="13">Rhaetina rainei</emphasis>
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<paragraph id="8F2936BBFFA35A71FF4BFBAAFD9EFB9B" blockId="12.[151,594,1071,1133]" box="[151,594,1107,1133]" pageId="12" pageNumber="13">
<figureCitation id="17AD2A3EFFA35A71FF4BFBAAFF25FB9B" box="[151,233,1107,1133]" captionStart="FIGURE 10" captionStartId="13.[151,250,1866,1890]" captionTargetBox="[170,1418,717,1839]" captionTargetId="figure-275@13.[158,1430,705,1843]" captionTargetPageId="13" captionText="FIGURE 10. Rhaetina rainei n. sp. (New Caledonia). 1 Holotype B876 (NC/f0519) shelly specimen (a) dorsal (b) ventral (c) lateral (d) posterior (e) anterior. 2 B875 (NC/f0519) shelly specimen (a) dorsal (b) lateral. 3 B871 (NC/f0503) shelly specimen, ventral. 4 B844 (NC/f0007A) shelly specimen, ventral. 5 B881 (NC/f0005A) partly shelly specimen (a) dorsal (b) ventral (c) lateral. 6 BR 3414 (NC/f0020) shelly specimen (a) dorsal (b) ventral (c) lateral. 7 B880 (NC/f0005A) dorsal view of beak. 8 B820 (NC/f0087A) shelly specimen, anterior." figureDoi="http://doi.org/10.5281/zenodo.10148179" httpUri="https://zenodo.org/record/10148179/files/figure.png" pageId="12" pageNumber="13">Fig. 10</figureCitation>
,
<figureCitation id="17AD2A3EFFA35A71FF29FBAAFEEDFB9B" box="[245,289,1107,1133]" captionStart-0="FIGURE 1" captionStart-1="FIGURE 2" captionStart-2="FIGURE 3" captionStart-3="FIGURE 4" captionStart-4="FIGURE 5" captionStart-5="FIGURE 6" captionStart-6="FIGURE 7" captionStart-7="FIGURE 8" captionStartId-0="1.[151,260,1422,1448]" captionStartId-1="3.[151,250,907,931]" captionStartId-2="3.[151,250,1855,1879]" captionStartId-3="4.[151,250,1826,1850]" captionStartId-4="7.[151,250,1823,1847]" captionStartId-5="8.[151,250,974,998]" captionStartId-6="9.[151,250,1544,1568]" captionStartId-7="10.[151,250,1811,1835]" captionTargetBox-0="[151,1435,882,1398]" captionTargetBox-1="[151,1435,179,882]" captionTargetBox-2="[151,1435,1028,1830]" captionTargetBox-3="[151,1436,181,1802]" captionTargetBox-4="[233,1361,193,1800]" captionTargetBox-5="[151,1435,184,950]" captionTargetBox-6="[164,1432,184,1510]" captionTargetBox-7="[151,1436,1003,1788]" captionTargetId-0="figure-285@1.[151,1435,882,1398]" captionTargetId-1="figure-20@3.[151,1435,179,882]" captionTargetId-2="figure-55@3.[151,1435,1028,1831]" captionTargetId-3="figure-34@4.[151,1436,181,1802]" captionTargetId-4="figure-20@7.[222,1365,181,1800]" captionTargetId-5="figure-34@8.[151,1435,181,950]" captionTargetId-6="figure-20@9.[151,1436,179,1518]" captionTargetId-7="figure-337@10.[151,1436,1003,1788]" captionTargetPageId-0="1" captionTargetPageId-1="3" captionTargetPageId-2="3" captionTargetPageId-3="4" captionTargetPageId-4="7" captionTargetPageId-5="8" captionTargetPageId-6="9" captionTargetPageId-7="10" captionText-0="FIGURE 1. Measured dimensions." captionText-1="FIGURE 2. Late Triassic - Jurassic timescale and range chart showing correlations between New Zealand and international stages (adapted from Raine et al. 2015) and ranges of Zealandian terebratulide species." captionText-2="FIGURE 3. A Zealandia, showing the area of submerged continental crust surrounding New Zealand and extending to New Caledonia. Base map: NIWA (National Institute of Water and Atmospheric Research). B New Caledonia, showing location of Moindou-Téremba area and Baie de St Vincent (Fig. 4A). C New Zealand, showing Kawhia, (Fig. 4B) Nelson and Southland (Fig. 4C) synclines, Alpine Fault and plate boundary." captionText-3="FIGURE 4. Main areas of TriassicJurassic strata in this study with main terebratulide localities shown (red stars): A Baie de St Vincent, New Caledonia, with areas of Late Triassic and Jurassic rocks indicated. Base and geology from Gouvernement de la Nouvelle-Calédonie data. B Southwest Auckland, showing southern part of Kawhia Syncline, with areas of Late Triassic and Jurassic rocks indicated. C Western Southland, showing northwestern part of Southland Syncline. Base for B and C: LINZ (Land Information New Zealand) topographic data with geology from GNS Science (Institute of Geological and Nuclear Sciences) 1: 250,000 geological map (Q-Map; Edbrooke et al. 2014, Heron 2014)." captionText-4="FIGURE 5. Tibetothyris johnstoni n. sp. 1 Holotype BR 3420 (N28/f7454) (a) dorsal (b) ventral (c) lateral (d) posterior (e) anterior. 2 BR 3419 (N28/f7454) (a) dorsal (b) ventral (c) lateral (d) posterior. 3 B890 (NC/f0109B) shelly specimen, ventral. 4 C1963 (E45/f6635) partly shelly specimen (a) dorsal (b) ventral (c) lateral (d) anterior. 5 C1965 (E45/f6635) internal mould, distorted, dorsal. 6 C1967 (E45/f6635) internal mould (a) dorsal (b) ventral (c) lateral (d) posterior (e) anterior. 7 C1970 (E45/f6635) internal mould (a) dorsal (b) ventral (c) lateral (d) posterior (e) anterior. 8 C1971 (E45/f6635) internal mould (a) dorsal (b) ventral (c) lateral." captionText-5="FIGURE 6. Detailed view of beak area, Tibetothyris johnstoni n. sp. A Holotype BR 3420 (N28/f7454), dorsal view of internal mould showing pedicle collar, ridged septalium, broad median septum and short, widely divergent dental plates. B C1970 (E45/f6635) dorsal view of internal mould, showing small cardinal process." captionText-6="FIGURE 7. Tibetothyris hamishi n. sp. 1 Holotype, OU 47309 (F45/f0329) internal mould (a) dorsal (b) ventral (c) lateral (d) anterior. 2 BR 3502 (F45/f8523) internal mould (a) dorsal (b) ventral. 3 BR3417 (NC/f0050) internal mould (a) dorsal (b) ventral (c) lateral. 4 B664 (R15/f8837) shelly specimen (a) dorsal (b) ventral (c) lateral (d) posterior (e) anterior. 5 OU 46810 (E45/f0279) internal mould (a) dorsal (b) ventral. 6 B822 (R16/f0377) internal mould, dorsal. 7 B665 (R15/f8838) shelly specimen (a) dorsal (b) ventral (c) lateral (d) posterior (e) anterior. 8 OU 46809 (E45/f9622) internal mould (a) dorsal (b) ventral. 9 B905 (NC/f0087) (a) dorsal (b) ventral (c) lateral (d) anterior." captionText-7="FIGURE 8. Detailed view of beak area, Tibetothyris hamishi n. sp. A B822 (R16/f0377) internal mould of dorsal valve, showing median septum and septalium. B BR 3417 (NC/f0050) dorsal view of internal mould, slightly distorted, showing dental plates, foramen with pedicle collar and cardinal process." figureDoi-0="http://doi.org/10.5281/zenodo.10148161" figureDoi-1="http://doi.org/10.5281/zenodo.10148163" figureDoi-2="http://doi.org/10.5281/zenodo.10148165" figureDoi-3="http://doi.org/10.5281/zenodo.10148167" figureDoi-4="http://doi.org/10.5281/zenodo.10148169" figureDoi-5="http://doi.org/10.5281/zenodo.10148171" figureDoi-6="http://doi.org/10.5281/zenodo.10148173" figureDoi-7="http://doi.org/10.5281/zenodo.10148175" httpUri-0="https://zenodo.org/record/10148161/files/figure.png" httpUri-1="https://zenodo.org/record/10148163/files/figure.png" httpUri-2="https://zenodo.org/record/10148165/files/figure.png" httpUri-3="https://zenodo.org/record/10148167/files/figure.png" httpUri-4="https://zenodo.org/record/10148169/files/figure.png" httpUri-5="https://zenodo.org/record/10148171/files/figure.png" httpUri-6="https://zenodo.org/record/10148173/files/figure.png" httpUri-7="https://zenodo.org/record/10148175/files/figure.png" pageId="12" pageNumber="13">18</figureCitation>
,
<figureCitation id="17AD2A3EFFA35A71FEF1FBAAFE0FFB9B" box="[301,451,1107,1133]" captionStart-0="FIGURE 11" captionStart-1="FIGURE 1" captionStart-10="FIGURE 10" captionStart-2="FIGURE 2" captionStart-3="FIGURE 3" captionStart-4="FIGURE 4" captionStart-5="FIGURE 5" captionStart-6="FIGURE 6" captionStart-7="FIGURE 7" captionStart-8="FIGURE 8" captionStart-9="FIGURE 9" captionStartId-0="14.[151,250,1758,1782]" captionStartId-1="1.[151,260,1422,1448]" captionStartId-10="13.[151,250,1866,1890]" captionStartId-2="3.[151,250,907,931]" captionStartId-3="3.[151,250,1855,1879]" captionStartId-4="4.[151,250,1826,1850]" captionStartId-5="7.[151,250,1823,1847]" captionStartId-6="8.[151,250,974,998]" captionStartId-7="9.[151,250,1544,1568]" captionStartId-8="10.[151,250,1811,1835]" captionStartId-9="11.[151,250,1489,1513]" captionTargetBox-0="[164,1428,194,1697]" captionTargetBox-1="[151,1435,882,1398]" captionTargetBox-10="[170,1418,717,1839]" captionTargetBox-2="[151,1435,179,882]" captionTargetBox-3="[151,1435,1028,1830]" captionTargetBox-4="[151,1436,181,1802]" captionTargetBox-5="[233,1361,193,1800]" captionTargetBox-6="[151,1435,184,950]" captionTargetBox-7="[164,1432,184,1510]" captionTargetBox-8="[151,1436,1003,1788]" captionTargetBox-9="[153,1434,181,1465]" captionTargetId-0="figure-34@14.[151,1436,181,1698]" captionTargetId-1="figure-285@1.[151,1435,882,1398]" captionTargetId-10="figure-275@13.[158,1430,705,1843]" captionTargetId-2="figure-20@3.[151,1435,179,882]" captionTargetId-3="figure-55@3.[151,1435,1028,1831]" captionTargetId-4="figure-34@4.[151,1436,181,1802]" captionTargetId-5="figure-20@7.[222,1365,181,1800]" captionTargetId-6="figure-34@8.[151,1435,181,950]" captionTargetId-7="figure-20@9.[151,1436,179,1518]" captionTargetId-8="figure-337@10.[151,1436,1003,1788]" captionTargetId-9="figure-20@11.[153,1434,181,1465]" captionTargetPageId-0="14" captionTargetPageId-1="1" captionTargetPageId-10="13" captionTargetPageId-2="3" captionTargetPageId-3="3" captionTargetPageId-4="4" captionTargetPageId-5="7" captionTargetPageId-6="8" captionTargetPageId-7="9" captionTargetPageId-8="10" captionTargetPageId-9="11" captionText-0="FIGURE 11. Rhaetina rainei n. sp. (Nelson). 1 BR 3430 (N28/f7454) internal mould (a) dorsal (b) ventral. 2 BR 3461 (N28/f7454) internal mould (a) dorsal (b) ventral. 3 BR 3490 (N28/f7454) latex of dorsal valve interior. 4 BR 3452 (N28/f7454 internal mould (a) dorsal (b) ventral (c) lateral (d) anterior. 5 BR 1014 (N28/f7522) ventral view (Specimen figured by Marwick (1953) as Coenothyris sp.; photo Marianna Terezow). 6 BR 3456 (N28/f7454) internal mould (a) dorsal (b) ventral (c) lateral (d) posterior. 7 BR 3424 (N28/f7454) internal mould with remnant shell (a) dorsal (b) ventral (c) lateral (d) posterior (e) anterior. 8 BR3500 (N28/f7455) internal mould, ventral. 9 BR 3498 (N28/f7454) latex of ventral valve interior and beak. 10 BR 3439 (N28/f7454) ventral view. 11 BR 3422 (N28/f7454) internal mould (a) dorsal (b) ventral." captionText-1="FIGURE 1. Measured dimensions." captionText-10="FIGURE 10. Rhaetina rainei n. sp. (New Caledonia). 1 Holotype B876 (NC/f0519) shelly specimen (a) dorsal (b) ventral (c) lateral (d) posterior (e) anterior. 2 B875 (NC/f0519) shelly specimen (a) dorsal (b) lateral. 3 B871 (NC/f0503) shelly specimen, ventral. 4 B844 (NC/f0007A) shelly specimen, ventral. 5 B881 (NC/f0005A) partly shelly specimen (a) dorsal (b) ventral (c) lateral. 6 BR 3414 (NC/f0020) shelly specimen (a) dorsal (b) ventral (c) lateral. 7 B880 (NC/f0005A) dorsal view of beak. 8 B820 (NC/f0087A) shelly specimen, anterior." captionText-2="FIGURE 2. Late Triassic - Jurassic timescale and range chart showing correlations between New Zealand and international stages (adapted from Raine et al. 2015) and ranges of Zealandian terebratulide species." captionText-3="FIGURE 3. A Zealandia, showing the area of submerged continental crust surrounding New Zealand and extending to New Caledonia. Base map: NIWA (National Institute of Water and Atmospheric Research). B New Caledonia, showing location of Moindou-Téremba area and Baie de St Vincent (Fig. 4A). C New Zealand, showing Kawhia, (Fig. 4B) Nelson and Southland (Fig. 4C) synclines, Alpine Fault and plate boundary." captionText-4="FIGURE 4. Main areas of TriassicJurassic strata in this study with main terebratulide localities shown (red stars): A Baie de St Vincent, New Caledonia, with areas of Late Triassic and Jurassic rocks indicated. Base and geology from Gouvernement de la Nouvelle-Calédonie data. B Southwest Auckland, showing southern part of Kawhia Syncline, with areas of Late Triassic and Jurassic rocks indicated. C Western Southland, showing northwestern part of Southland Syncline. Base for B and C: LINZ (Land Information New Zealand) topographic data with geology from GNS Science (Institute of Geological and Nuclear Sciences) 1: 250,000 geological map (Q-Map; Edbrooke et al. 2014, Heron 2014)." captionText-5="FIGURE 5. Tibetothyris johnstoni n. sp. 1 Holotype BR 3420 (N28/f7454) (a) dorsal (b) ventral (c) lateral (d) posterior (e) anterior. 2 BR 3419 (N28/f7454) (a) dorsal (b) ventral (c) lateral (d) posterior. 3 B890 (NC/f0109B) shelly specimen, ventral. 4 C1963 (E45/f6635) partly shelly specimen (a) dorsal (b) ventral (c) lateral (d) anterior. 5 C1965 (E45/f6635) internal mould, distorted, dorsal. 6 C1967 (E45/f6635) internal mould (a) dorsal (b) ventral (c) lateral (d) posterior (e) anterior. 7 C1970 (E45/f6635) internal mould (a) dorsal (b) ventral (c) lateral (d) posterior (e) anterior. 8 C1971 (E45/f6635) internal mould (a) dorsal (b) ventral (c) lateral." captionText-6="FIGURE 6. Detailed view of beak area, Tibetothyris johnstoni n. sp. A Holotype BR 3420 (N28/f7454), dorsal view of internal mould showing pedicle collar, ridged septalium, broad median septum and short, widely divergent dental plates. B C1970 (E45/f6635) dorsal view of internal mould, showing small cardinal process." captionText-7="FIGURE 7. Tibetothyris hamishi n. sp. 1 Holotype, OU 47309 (F45/f0329) internal mould (a) dorsal (b) ventral (c) lateral (d) anterior. 2 BR 3502 (F45/f8523) internal mould (a) dorsal (b) ventral. 3 BR3417 (NC/f0050) internal mould (a) dorsal (b) ventral (c) lateral. 4 B664 (R15/f8837) shelly specimen (a) dorsal (b) ventral (c) lateral (d) posterior (e) anterior. 5 OU 46810 (E45/f0279) internal mould (a) dorsal (b) ventral. 6 B822 (R16/f0377) internal mould, dorsal. 7 B665 (R15/f8838) shelly specimen (a) dorsal (b) ventral (c) lateral (d) posterior (e) anterior. 8 OU 46809 (E45/f9622) internal mould (a) dorsal (b) ventral. 9 B905 (NC/f0087) (a) dorsal (b) ventral (c) lateral (d) anterior." captionText-8="FIGURE 8. Detailed view of beak area, Tibetothyris hamishi n. sp. A B822 (R16/f0377) internal mould of dorsal valve, showing median septum and septalium. B BR 3417 (NC/f0050) dorsal view of internal mould, slightly distorted, showing dental plates, foramen with pedicle collar and cardinal process." captionText-9="FIGURE 9. Length vs width graphs for ventral valve. A Tibetothyris johnstoni and T. hamishi. B Rhaetina rainei, New Caledonia and Nelson. C Lobothyris richardsi, (E44/f6635 South Taringatura Hills and other localities). D Zeilleria minima (E44/ f6635 and other localities), Zeilleria spiculata and Zeilleria n. sp. Z. spiculata data from MacFarlan (2021)." figureDoi-0="http://doi.org/10.5281/zenodo.10148181" figureDoi-1="http://doi.org/10.5281/zenodo.10148161" figureDoi-10="http://doi.org/10.5281/zenodo.10148179" figureDoi-2="http://doi.org/10.5281/zenodo.10148163" figureDoi-3="http://doi.org/10.5281/zenodo.10148165" figureDoi-4="http://doi.org/10.5281/zenodo.10148167" figureDoi-5="http://doi.org/10.5281/zenodo.10148169" figureDoi-6="http://doi.org/10.5281/zenodo.10148171" figureDoi-7="http://doi.org/10.5281/zenodo.10148173" figureDoi-8="http://doi.org/10.5281/zenodo.10148175" figureDoi-9="http://doi.org/10.5281/zenodo.10148177" httpUri-0="https://zenodo.org/record/10148181/files/figure.png" httpUri-1="https://zenodo.org/record/10148161/files/figure.png" httpUri-10="https://zenodo.org/record/10148179/files/figure.png" httpUri-2="https://zenodo.org/record/10148163/files/figure.png" httpUri-3="https://zenodo.org/record/10148165/files/figure.png" httpUri-4="https://zenodo.org/record/10148167/files/figure.png" httpUri-5="https://zenodo.org/record/10148169/files/figure.png" httpUri-6="https://zenodo.org/record/10148171/files/figure.png" httpUri-7="https://zenodo.org/record/10148173/files/figure.png" httpUri-8="https://zenodo.org/record/10148175/files/figure.png" httpUri-9="https://zenodo.org/record/10148177/files/figure.png" pageId="12" pageNumber="13">Fig. 11, 111</figureCitation>
.
<figureCitation id="17AD2A3EFFA35A71FE12FBAAFD9EFB9B" box="[462,594,1107,1133]" captionStart="FIGURE 12" captionStartId="15.[151,250,1683,1707]" captionTargetBox="[159,1431,214,1659]" captionTargetId="figure-20@15.[156,1431,181,1659]" captionTargetPageId="15" captionText="FIGURE 12. Detailed views of beak area, Rhaetina rainei n.sp. A BR3458 (N28/f7454) dorsal view of internal mould, showing foramen with pedicle collar, and dorsal valve with low, ridgelike median septum. B BR3456 (N28/f7454) dorsal view of internal mould showing cardinal process. C B877 (NC/f0519) dorsal view of shelly specimen, slightly abraded, showing disjunct deltidial plates and foramen surrounded by pedicle collar." figureDoi="http://doi.org/10.5281/zenodo.10148183" httpUri="https://zenodo.org/record/10148183/files/figure.png" pageId="12" pageNumber="13">Fig. 12AC</figureCitation>
</paragraph>
</subSubSection>
<subSubSection id="C78C6530FFA35A71FF4BFB6EFC56FB38" pageId="12" pageNumber="13" type="reference_group">
<paragraph id="8F2936BBFFA35A71FF4BFB6EFCA3FB46" blockId="12.[151,922,1175,1230]" box="[151,879,1175,1200]" pageId="12" pageNumber="13">
1951
<taxonomicName id="48964D38FFA35A71FF0CFB61FEBCFB59" box="[208,368,1176,1200]" class="Rhynchonellata" family="Dielasmatidae" genus="Coenothyris" kingdom="Animalia" order="Terebratulida" pageId="12" pageNumber="13" phylum="Brachiopoda" rank="species" species="undetermined">
<emphasis id="BDE2EAA9FFA35A71FF0CFB61FE80FB46" box="[208,332,1176,1200]" italics="true" pageId="12" pageNumber="13">Coenothyris</emphasis>
sp.
</taxonomicName>
<bibRefCitation id="EB074B4AFFA35A71FEA9FB61FDE8FB46" author="Trechmann, C. T." box="[373,548,1176,1200]" pageId="12" pageNumber="13" pagination="3" refId="ref17196" refString="Trechmann, C. T. (1918) TheTriasofNewZealand. QuarterlyJournaloftheGeologicalSociety, London, 73 (3), 165 - 246,9 pl, map, sect. https: // doi. org / 10.1144 / GSL. JGS. 1917.073.01 - 04.10" type="journal article" year="1918">Trechmann 1918</bibRefCitation>
;
<bibRefCitation id="EB074B4AFFA35A71FDF2FB61FD32FB46" author="Marwick, J." box="[558,766,1175,1200]" pageId="12" pageNumber="13" refId="ref15397" refString="Marwick, J. (1953) Divisions and faunas of the Hokonui System (Triassic and Jurassic). New Zealand Geological Survey Paleontological Bulletin 21. New Zealand Geological Survey, Wellington, 141 pp." type="book" year="1953">Marwick 1953 p. 34</bibRefCitation>
, pl. 1: 8, 9.
</paragraph>
<paragraph id="8F2936BBFFA35A71FF4BFB4FFC56FB38" blockId="12.[151,922,1175,1230]" box="[151,922,1206,1230]" pageId="12" pageNumber="13">
NON
<taxonomicName id="48964D38FFA35A71FF08FB4FFEB8FB38" box="[212,372,1206,1230]" class="Rhynchonellata" family="Dielasmatidae" genus="Coenothyris" kingdom="Animalia" order="Terebratulida" pageId="12" pageNumber="13" phylum="Brachiopoda" rank="species" species="undetermined">
<emphasis id="BDE2EAA9FFA35A71FF08FB4FFE9CFB38" box="[212,336,1206,1230]" italics="true" pageId="12" pageNumber="13">Coenothyris</emphasis>
sp.
</taxonomicName>
Trechmann, p. 218, pl. 23:7 (fide J.D. Campbell, MS)
</paragraph>
</subSubSection>
<subSubSection id="C78C6530FFA35A71FF4BFB04FC88F989" pageId="12" pageNumber="13" type="materials_examined">
<paragraph id="8F2936BBFFA35A71FF4BFB04FD9CFACC" blockId="12.[151,1437,1276,2023]" pageId="12" pageNumber="13">
<emphasis id="BDE2EAA9FFA35A71FF4BFB04FEC8FAE1" bold="true" box="[151,260,1277,1303]" pageId="12" pageNumber="13">
<typeStatus id="502D8819FFA35A71FF4BFB04FEC8FAE1" box="[151,260,1277,1303]" pageId="12" pageNumber="13" type="holotype">Holotype</typeStatus>
</emphasis>
. B876 from NC/f0519 AU 7774, île Hugon. Double-valved shelly specimen with some damage to dorsal valve. Collected by J.A. Grant-Mackie.
</paragraph>
<paragraph id="8F2936BBFFA35A71FF1BFABDFBFEFAA8" blockId="12.[151,1437,1276,2023]" box="[199,1074,1348,1375]" pageId="12" pageNumber="13">
<emphasis id="BDE2EAA9FFA35A71FF1BFABDFE61FAA9" bold="true" box="[199,429,1348,1375]" pageId="12" pageNumber="13">Derivation of name</emphasis>
. This species is named for Dr Ian Raine of GNS Science.
</paragraph>
<paragraph id="8F2936BBFFA35A71FF1BFA91FC7EFA51" blockId="12.[151,1437,1276,2023]" pageId="12" pageNumber="13">
<emphasis id="BDE2EAA9FFA35A71FF1BFA91FEE3FA74" bold="true" box="[199,303,1384,1410]" pageId="12" pageNumber="13">Material</emphasis>
. The data series consists of
<specimenCount id="9990FD32FFA35A71FDABFA90FCEEFA74" box="[631,802,1384,1411]" count="129" pageId="12" pageNumber="13" type="generic">129 specimens</specimenCount>
, 38 from
<collectingCountry id="F781762BFFA35A71FC4FFA90FB8FFA74" box="[915,1091,1384,1411]" name="New Caledonia" pageId="12" pageNumber="13">New Caledonia</collectingCountry>
and 91 from Nelson; 23
<collectingCountry id="F781762BFFA35A71FABBFA90FEC5FA50" name="New Caledonia" pageId="12" pageNumber="13">New Caledonia</collectingCountry>
specimens and 68 from Nelson yielded valid measurements.
</paragraph>
<paragraph id="8F2936BBFFA35A71FF1BFA48FA8EF9E4" blockId="12.[151,1437,1276,2023]" pageId="12" pageNumber="13">
<emphasis id="BDE2EAA9FFA35A71FF1BFA48FE4EFA3C" bold="true" box="[199,386,1456,1483]" pageId="12" pageNumber="13">
<collectingCountry id="F781762BFFA35A71FF1BFA48FEB1FA3C" box="[199,381,1456,1483]" name="New Caledonia" pageId="12" pageNumber="13">New Caledonia</collectingCountry>
.
</emphasis>
île Hugon: NC/f0004A (AU 7148), NC/f0005 (AU 7146), NC/f0007A (AU 7149), NC/f0008A (AU 7795), NC/f0010A (AU 4155), NC/f503 (AU 7174), NC/f518 (AU 7773), NC/f519 (AU 7774). île Ducos: NC/f0020 (
<collectionCode id="E987AE7EFFA35A71FECAFA00FEF7F9E5" box="[278,315,1529,1555]" pageId="12" pageNumber="13">GS</collectionCode>
12733), NC/f0023 (?
<collectionCode id="E987AE7EFFA35A71FDF6FA00FD9CF9E5" box="[554,592,1529,1555]" pageId="12" pageNumber="13">GS</collectionCode>
12376), NC/f60A (?AU 7214), NC/f0087A (?AU 6073, AU 7168).
</paragraph>
<paragraph id="8F2936BBFFA35A71FF1BF9E4FD25F9AD" blockId="12.[151,1437,1276,2023]" pageId="12" pageNumber="13">
<emphasis id="BDE2EAA9FFA35A71FF1BF9E4FED3F9C1" bold="true" box="[199,287,1565,1591]" pageId="12" pageNumber="13">
<collectingRegion id="4D52F859FFA35A71FF1BF9E4FED6F9C1" box="[199,282,1565,1591]" country="New Zealand" name="Nelson" pageId="12" pageNumber="13">Nelson</collectingRegion>
.
</emphasis>
Eighty-Eight Valley: N28/f7454 (
<collectionCode id="E987AE7EFFA35A71FD4AF9E4FD77F9C1" box="[662,699,1565,1591]" pageId="12" pageNumber="13">GS</collectionCode>
196), N28/f7455 (
<collectionCode id="E987AE7EFFA35A71FC54F9E4FC61F9C1" box="[904,941,1565,1591]" pageId="12" pageNumber="13">GS</collectionCode>
197). Ram Creek: N28/f0045 (
<collectionCode id="E987AE7EFFA35A71FAD8F9E4FAF5F9C1" box="[1284,1337,1565,1591]" pageId="12" pageNumber="13">PGS</collectionCode>
40, McF D21), N28/f7860 (AU 7885), N28/f7864 (AU 6521).
</paragraph>
<paragraph id="8F2936BBFFA35A71FF1BF99DFC88F989" blockId="12.[151,1437,1276,2023]" box="[199,836,1636,1663]" pageId="12" pageNumber="13">
<emphasis id="BDE2EAA9FFA35A71FF1BF99DFE79F988" bold="true" box="[199,437,1636,1662]" pageId="12" pageNumber="13">
<collectingRegion id="4D52F859FFA35A71FF1BF99DFE8EF988" box="[199,322,1636,1662]" country="New Zealand" name="Southland" pageId="12" pageNumber="13">Southland</collectingRegion>
Syncline.
</emphasis>
Oreti Valley: E45/f7484 (?
<collectionCode id="E987AE7EFFA35A71FD3FF99CFCC5F989" box="[739,777,1637,1663]" pageId="12" pageNumber="13">GS</collectionCode>
372).
</paragraph>
</subSubSection>
<subSubSection id="C78C6530FFA35A72FF1BF971FC1FF851" lastPageId="15" lastPageNumber="16" pageId="12" pageNumber="13" type="description">
<paragraph id="8F2936BBFFA35A71FF1BF971FE39F91C" blockId="12.[151,1437,1276,2023]" pageId="12" pageNumber="13">
<emphasis id="BDE2EAA9FFA35A71FF1BF971FE9DF954" bold="true" box="[199,337,1672,1698]" pageId="12" pageNumber="13">Description</emphasis>
. Medium sized to large terebratulide with an elongate-elliptical to rounded-triangular outline. The dorsal valve is moderately and evenly convex posteriorly, and generally has two rounded folds anteriorly. Some large specimens are not folded.
</paragraph>
<paragraph id="8F2936BBFFA35A71FF1BF90DFEF7F88C" blockId="12.[151,1437,1276,2023]" pageId="12" pageNumber="13">The ventral valve is strongly convex posteriorly, and in some specimens is strongly folded anteriorly. The initiation of folds is variable, from 1/5 of valve length in some specimens to near anterior in others. The folds are often stronger on the dorsal valve. The strength can be accentuated by distortion. The anterior margin is uniplicate or parasulcate.</paragraph>
<paragraph id="8F2936BBFFA35A71FF1BF87DFAD1F810" blockId="12.[151,1437,1276,2023]" pageId="12" pageNumber="13">
The shell surface is smooth or has fine closely spaced growth lines. Some specimens have stronger, rounded concentrics, which may be visible on the valve interior. Punctae appear to be fine but are poorly shown on shelly
<collectingCountry id="F781762BFFA35A71FF4BF834FE89F810" box="[151,325,1996,2023]" name="New Caledonia" pageId="12" pageNumber="13">New Caledonia</collectingCountry>
specimens and the remnant shell material on Nelson specimens due to recrystallisation.
</paragraph>
<paragraph id="8F2936BBFFA25A70FF1BFF6EFC04FF0F" blockId="13.[151,1437,151,681]" pageId="13" pageNumber="14">
The beak is triangular and suberect with a bluntly rounded tip. The foramen is large, subcircular (
<figureCitation id="17AD2A3EFFA25A70FB28FF6EFAAAFF47" box="[1268,1382,151,177]" captionStart="FIGURE 10" captionStartId="13.[151,250,1866,1890]" captionTargetBox="[170,1418,717,1839]" captionTargetId="figure-275@13.[158,1430,705,1843]" captionTargetPageId="13" captionText="FIGURE 10. Rhaetina rainei n. sp. (New Caledonia). 1 Holotype B876 (NC/f0519) shelly specimen (a) dorsal (b) ventral (c) lateral (d) posterior (e) anterior. 2 B875 (NC/f0519) shelly specimen (a) dorsal (b) lateral. 3 B871 (NC/f0503) shelly specimen, ventral. 4 B844 (NC/f0007A) shelly specimen, ventral. 5 B881 (NC/f0005A) partly shelly specimen (a) dorsal (b) ventral (c) lateral. 6 BR 3414 (NC/f0020) shelly specimen (a) dorsal (b) ventral (c) lateral. 7 B880 (NC/f0005A) dorsal view of beak. 8 B820 (NC/f0087A) shelly specimen, anterior." figureDoi="http://doi.org/10.5281/zenodo.10148179" httpUri="https://zenodo.org/record/10148179/files/figure.png" pageId="13" pageNumber="14">Fig. 10.1a</figureCitation>
) and permesothyrid (
<figureCitation id="17AD2A3EFFA25A70FE94FF42FE77FF23" box="[328,443,187,213]" captionStart="FIGURE 11" captionStartId="14.[151,250,1758,1782]" captionTargetBox="[164,1428,194,1697]" captionTargetId="figure-34@14.[151,1436,181,1698]" captionTargetPageId="14" captionText="FIGURE 11. Rhaetina rainei n. sp. (Nelson). 1 BR 3430 (N28/f7454) internal mould (a) dorsal (b) ventral. 2 BR 3461 (N28/f7454) internal mould (a) dorsal (b) ventral. 3 BR 3490 (N28/f7454) latex of dorsal valve interior. 4 BR 3452 (N28/f7454 internal mould (a) dorsal (b) ventral (c) lateral (d) anterior. 5 BR 1014 (N28/f7522) ventral view (Specimen figured by Marwick (1953) as Coenothyris sp.; photo Marianna Terezow). 6 BR 3456 (N28/f7454) internal mould (a) dorsal (b) ventral (c) lateral (d) posterior. 7 BR 3424 (N28/f7454) internal mould with remnant shell (a) dorsal (b) ventral (c) lateral (d) posterior (e) anterior. 8 BR3500 (N28/f7455) internal mould, ventral. 9 BR 3498 (N28/f7454) latex of ventral valve interior and beak. 10 BR 3439 (N28/f7454) ventral view. 11 BR 3422 (N28/f7454) internal mould (a) dorsal (b) ventral." figureDoi="http://doi.org/10.5281/zenodo.10148181" httpUri="https://zenodo.org/record/10148181/files/figure.png" pageId="13" pageNumber="14">Fig. 11.6a</figureCitation>
,
<figureCitation id="17AD2A3EFFA25A70FE1AFF42FE29FF23" box="[454,485,187,213]" captionStart="FIGURE 6" captionStartId="8.[151,250,974,998]" captionTargetBox="[151,1435,184,950]" captionTargetId="figure-34@8.[151,1435,181,950]" captionTargetPageId="8" captionText="FIGURE 6. Detailed view of beak area, Tibetothyris johnstoni n. sp. A Holotype BR 3420 (N28/f7454), dorsal view of internal mould showing pedicle collar, ridged septalium, broad median septum and short, widely divergent dental plates. B C1970 (E45/f6635) dorsal view of internal mould, showing small cardinal process." figureDoi="http://doi.org/10.5281/zenodo.10148171" httpUri="https://zenodo.org/record/10148171/files/figure.png" pageId="13" pageNumber="14">6d</figureCitation>
,
<figureCitation id="17AD2A3EFFA25A70FE2CFF42FD88FF23" box="[496,580,187,213]" captionStart="FIGURE 12" captionStartId="15.[151,250,1683,1707]" captionTargetBox="[159,1431,214,1659]" captionTargetId="figure-20@15.[156,1431,181,1659]" captionTargetPageId="15" captionText="FIGURE 12. Detailed views of beak area, Rhaetina rainei n.sp. A BR3458 (N28/f7454) dorsal view of internal mould, showing foramen with pedicle collar, and dorsal valve with low, ridgelike median septum. B BR3456 (N28/f7454) dorsal view of internal mould showing cardinal process. C B877 (NC/f0519) dorsal view of shelly specimen, slightly abraded, showing disjunct deltidial plates and foramen surrounded by pedicle collar." figureDoi="http://doi.org/10.5281/zenodo.10148183" httpUri="https://zenodo.org/record/10148183/files/figure.png" pageId="13" pageNumber="14">12AC</figureCitation>
), and is surrounded by a large, thick pedicle collar. The beak ridges are blunt and poorly defined. The deltidial plates are large and disjunct (
<figureCitation id="17AD2A3EFFA25A70FC89FF26FC70FF0F" box="[853,956,223,249]" captionStart="FIGURE 12" captionStartId="15.[151,250,1683,1707]" captionTargetBox="[159,1431,214,1659]" captionTargetId="figure-20@15.[156,1431,181,1659]" captionTargetPageId="15" captionText="FIGURE 12. Detailed views of beak area, Rhaetina rainei n.sp. A BR3458 (N28/f7454) dorsal view of internal mould, showing foramen with pedicle collar, and dorsal valve with low, ridgelike median septum. B BR3456 (N28/f7454) dorsal view of internal mould showing cardinal process. C B877 (NC/f0519) dorsal view of shelly specimen, slightly abraded, showing disjunct deltidial plates and foramen surrounded by pedicle collar." figureDoi="http://doi.org/10.5281/zenodo.10148183" httpUri="https://zenodo.org/record/10148183/files/figure.png" pageId="13" pageNumber="14">Fig. 12C</figureCitation>
).
</paragraph>
<paragraph id="8F2936BBFFA25A70FF1BFEFAFB6DFE93" blockId="13.[151,1437,151,681]" pageId="13" pageNumber="14">
In the dorsal valve the hingeplate has broad inner socket ridges and narrow outer ridges, with a distinct cardinal process (
<figureCitation id="17AD2A3EFFA25A70FF26FEDEFEA1FEB7" box="[250,365,295,321]" captionStart="FIGURE 11" captionStartId="14.[151,250,1758,1782]" captionTargetBox="[164,1428,194,1697]" captionTargetId="figure-34@14.[151,1436,181,1698]" captionTargetPageId="14" captionText="FIGURE 11. Rhaetina rainei n. sp. (Nelson). 1 BR 3430 (N28/f7454) internal mould (a) dorsal (b) ventral. 2 BR 3461 (N28/f7454) internal mould (a) dorsal (b) ventral. 3 BR 3490 (N28/f7454) latex of dorsal valve interior. 4 BR 3452 (N28/f7454 internal mould (a) dorsal (b) ventral (c) lateral (d) anterior. 5 BR 1014 (N28/f7522) ventral view (Specimen figured by Marwick (1953) as Coenothyris sp.; photo Marianna Terezow). 6 BR 3456 (N28/f7454) internal mould (a) dorsal (b) ventral (c) lateral (d) posterior. 7 BR 3424 (N28/f7454) internal mould with remnant shell (a) dorsal (b) ventral (c) lateral (d) posterior (e) anterior. 8 BR3500 (N28/f7455) internal mould, ventral. 9 BR 3498 (N28/f7454) latex of ventral valve interior and beak. 10 BR 3439 (N28/f7454) ventral view. 11 BR 3422 (N28/f7454) internal mould (a) dorsal (b) ventral." figureDoi="http://doi.org/10.5281/zenodo.10148181" httpUri="https://zenodo.org/record/10148181/files/figure.png" pageId="13" pageNumber="14">Fig. 11.4a</figureCitation>
,
<figureCitation id="17AD2A3EFFA25A70FEABFEDEFE58FEB7" box="[375,404,295,321]" captionStart="FIGURE 6" captionStartId="8.[151,250,974,998]" captionTargetBox="[151,1435,184,950]" captionTargetId="figure-34@8.[151,1435,181,950]" captionTargetPageId="8" captionText="FIGURE 6. Detailed view of beak area, Tibetothyris johnstoni n. sp. A Holotype BR 3420 (N28/f7454), dorsal view of internal mould showing pedicle collar, ridged septalium, broad median septum and short, widely divergent dental plates. B C1970 (E45/f6635) dorsal view of internal mould, showing small cardinal process." figureDoi="http://doi.org/10.5281/zenodo.10148171" httpUri="https://zenodo.org/record/10148171/files/figure.png" pageId="13" pageNumber="14">6a</figureCitation>
,
<figureCitation id="17AD2A3EFFA25A70FE42FEDEFE62FEB7" box="[414,430,295,321]" captionStart="FIGURE 9" captionStartId="11.[151,250,1489,1513]" captionTargetBox="[153,1434,181,1465]" captionTargetId="figure-20@11.[153,1434,181,1465]" captionTargetPageId="11" captionText="FIGURE 9. Length vs width graphs for ventral valve. A Tibetothyris johnstoni and T. hamishi. B Rhaetina rainei, New Caledonia and Nelson. C Lobothyris richardsi, (E44/f6635 South Taringatura Hills and other localities). D Zeilleria minima (E44/ f6635 and other localities), Zeilleria spiculata and Zeilleria n. sp. Z. spiculata data from MacFarlan (2021)." figureDoi="http://doi.org/10.5281/zenodo.10148177" httpUri="https://zenodo.org/record/10148177/files/figure.png" pageId="13" pageNumber="14">9</figureCitation>
,
<figureCitation id="17AD2A3EFFA25A70FE66FEDEFDF3FEB7" box="[442,575,295,321]" captionStart="FIGURE 12" captionStartId="15.[151,250,1683,1707]" captionTargetBox="[159,1431,214,1659]" captionTargetId="figure-20@15.[156,1431,181,1659]" captionTargetPageId="15" captionText="FIGURE 12. Detailed views of beak area, Rhaetina rainei n.sp. A BR3458 (N28/f7454) dorsal view of internal mould, showing foramen with pedicle collar, and dorsal valve with low, ridgelike median septum. B BR3456 (N28/f7454) dorsal view of internal mould showing cardinal process. C B877 (NC/f0519) dorsal view of shelly specimen, slightly abraded, showing disjunct deltidial plates and foramen surrounded by pedicle collar." figureDoi="http://doi.org/10.5281/zenodo.10148183" httpUri="https://zenodo.org/record/10148183/files/figure.png" pageId="13" pageNumber="14">Fig. 12A, B</figureCitation>
). There is a low, ridge-like median septum which is about 1/3 of valve length, with a shallow muscle field on either side of the septum (
<figureCitation id="17AD2A3EFFA25A70FCC4FEB2FC41FE93" box="[792,909,331,357]" captionStart="FIGURE 11" captionStartId="14.[151,250,1758,1782]" captionTargetBox="[164,1428,194,1697]" captionTargetId="figure-34@14.[151,1436,181,1698]" captionTargetPageId="14" captionText="FIGURE 11. Rhaetina rainei n. sp. (Nelson). 1 BR 3430 (N28/f7454) internal mould (a) dorsal (b) ventral. 2 BR 3461 (N28/f7454) internal mould (a) dorsal (b) ventral. 3 BR 3490 (N28/f7454) latex of dorsal valve interior. 4 BR 3452 (N28/f7454 internal mould (a) dorsal (b) ventral (c) lateral (d) anterior. 5 BR 1014 (N28/f7522) ventral view (Specimen figured by Marwick (1953) as Coenothyris sp.; photo Marianna Terezow). 6 BR 3456 (N28/f7454) internal mould (a) dorsal (b) ventral (c) lateral (d) posterior. 7 BR 3424 (N28/f7454) internal mould with remnant shell (a) dorsal (b) ventral (c) lateral (d) posterior (e) anterior. 8 BR3500 (N28/f7455) internal mould, ventral. 9 BR 3498 (N28/f7454) latex of ventral valve interior and beak. 10 BR 3439 (N28/f7454) ventral view. 11 BR 3422 (N28/f7454) internal mould (a) dorsal (b) ventral." figureDoi="http://doi.org/10.5281/zenodo.10148181" httpUri="https://zenodo.org/record/10148181/files/figure.png" pageId="13" pageNumber="14">Fig. 11.6a</figureCitation>
). The loop was not seen.
</paragraph>
<paragraph id="8F2936BBFFA25A70FF1BFE97FB98FE5B" blockId="13.[151,1437,151,681]" pageId="13" pageNumber="14">
<emphasis id="BDE2EAA9FFA25A70FF1BFE97FE95FE7E" bold="true" box="[199,345,366,392]" pageId="13" pageNumber="14">Dimensions.</emphasis>
<materialsCitation id="3FFE3CE6FFA25A70FEBDFE96FB83FE5B" country="New Caledonia" location="Nelson" pageId="13" pageNumber="14" specimenCount="1" typeStatus="holotype">
Dimensions of the
<typeStatus id="502D8819FFA25A70FDE4FE96FD56FE7F" box="[568,666,367,393]" pageId="13" pageNumber="14" type="holotype">holotype</typeStatus>
and representative specimens, and statistics for
<collectingCountry id="F781762BFFA25A70FB61FE96FAA0FE7F" box="[1213,1388,367,393]" name="New Caledonia" pageId="13" pageNumber="14">New Caledonia</collectingCountry>
and
<location id="8A496060FFA25A70FF4BFE6AFF2AFE5B" LSID="urn:lsid:plazi:treatment:073F87ADFFA35A6DFF4BFBD6FC86FE93:8A496060FFA25A70FF4BFE6AFF2AFE5B" box="[151,230,403,429]" country="New Caledonia" name="Nelson" pageId="13" pageNumber="14">Nelson</location>
specimens are shown in
<tableCitation id="C2140300FFA25A70FE23FE6AFDBDFE5B" box="[511,625,403,429]" captionStart="TABLE 3" captionStartId="16.[151,239,399,423]" captionTargetPageId="16" captionText="TABLE 3. Dimensions, Rhaetina rainei n. sp." httpUri="http://table.plazi.org/id/DBE96633FFBF5A6DFF4BFE76FD48FE51" pageId="13" pageNumber="14" tableUuid="DBE96633FFBF5A6DFF4BFE76FD48FE51">Table 3. A</tableCitation>
length vs width graph is shown in
<figureCitation id="17AD2A3EFFA25A70FC25FE6AFB83FE5B" box="[1017,1103,403,429]" captionStart="FIGURE 9" captionStartId="11.[151,250,1489,1513]" captionTargetBox="[153,1434,181,1465]" captionTargetId="figure-20@11.[153,1434,181,1465]" captionTargetPageId="11" captionText="FIGURE 9. Length vs width graphs for ventral valve. A Tibetothyris johnstoni and T. hamishi. B Rhaetina rainei, New Caledonia and Nelson. C Lobothyris richardsi, (E44/f6635 South Taringatura Hills and other localities). D Zeilleria minima (E44/ f6635 and other localities), Zeilleria spiculata and Zeilleria n. sp. Z. spiculata data from MacFarlan (2021)." figureDoi="http://doi.org/10.5281/zenodo.10148177" httpUri="https://zenodo.org/record/10148177/files/figure.png" pageId="13" pageNumber="14">Fig. 9B</figureCitation>
</materialsCitation>
.
</paragraph>
<paragraph id="8F2936BBFFA25A70FF1BFE4EFC2DFE03" blockId="13.[151,1437,151,681]" pageId="13" pageNumber="14">
<emphasis id="BDE2EAA9FFA25A70FF1BFE4EFE2CFE26" bold="true" box="[199,480,438,465]" pageId="13" pageNumber="14">Range and Distribution</emphasis>
. Otapirian of Nelson and
<collectingCountry id="F781762BFFA25A70FCDEFE4EFC63FE27" box="[770,943,439,465]" name="New Caledonia" pageId="13" pageNumber="14">New Caledonia</collectingCountry>
(Hugon and Ducos). The Nelson material is late Otapirian, the
<collectingCountry id="F781762BFFA25A70FEB4FE22FDD9FE03" box="[360,533,475,501]" name="New Caledonia" pageId="13" pageNumber="14">New Caledonia</collectingCountry>
material is both early and late Otapirian.
</paragraph>
<paragraph id="8F2936BBFFA25A70FF1BFE06FCDCFD5F" blockId="13.[151,1437,151,681]" pageId="13" pageNumber="14">
<bibRefCitation id="EB074B4AFFA25A70FF1BFE06FEB0FDEF" author="Marwick, J." box="[199,380,510,537]" pageId="13" pageNumber="14" refId="ref15397" refString="Marwick, J. (1953) Divisions and faunas of the Hokonui System (Triassic and Jurassic). New Zealand Geological Survey Paleontological Bulletin 21. New Zealand Geological Survey, Wellington, 141 pp." type="book" year="1953">Marwick (1953)</bibRefCitation>
figured
<taxonomicName id="48964D38FFA25A70FE04FE06FD44FDEF" box="[472,648,511,537]" class="Rhynchonellata" family="Dielasmatidae" genus="Coenothyris" kingdom="Animalia" order="Terebratulida" pageId="13" pageNumber="14" phylum="Brachiopoda" rank="species" species="undetermined">
<emphasis id="BDE2EAA9FFA25A70FE04FE06FDADFDEF" box="[472,609,511,537]" italics="true" pageId="13" pageNumber="14">Coenothyris</emphasis>
sp.
</taxonomicName>
, which was originally reported by
<bibRefCitation id="EB074B4AFFA25A70FBDFFE06FB1DFDEF" author="Trechmann, C. T." box="[1027,1233,511,537]" pageId="13" pageNumber="14" pagination="3" refId="ref17196" refString="Trechmann, C. T. (1918) TheTriasofNewZealand. QuarterlyJournaloftheGeologicalSociety, London, 73 (3), 165 - 246,9 pl, map, sect. https: // doi. org / 10.1144 / GSL. JGS. 1917.073.01 - 04.10" type="journal article" year="1918">Trechmann (1918)</bibRefCitation>
from the Warepan of Ram Hill,
<collectingRegion id="4D52F859FFA25A70FEF7FDDAFEB6FDCB" box="[299,378,547,573]" country="New Zealand" name="Nelson" pageId="13" pageNumber="14">Nelson</collectingRegion>
(N28/f7522, GS 4559). Re-examination of Marwicks figured specimens (BR 1013 and 1014) show them to belong to the new species (
<figureCitation id="17AD2A3EFFA25A70FDA5FDBEFD28FD97" box="[633,740,582,609]" captionStart="FIGURE 11" captionStartId="14.[151,250,1758,1782]" captionTargetBox="[164,1428,194,1697]" captionTargetId="figure-34@14.[151,1436,181,1698]" captionTargetPageId="14" captionText="FIGURE 11. Rhaetina rainei n. sp. (Nelson). 1 BR 3430 (N28/f7454) internal mould (a) dorsal (b) ventral. 2 BR 3461 (N28/f7454) internal mould (a) dorsal (b) ventral. 3 BR 3490 (N28/f7454) latex of dorsal valve interior. 4 BR 3452 (N28/f7454 internal mould (a) dorsal (b) ventral (c) lateral (d) anterior. 5 BR 1014 (N28/f7522) ventral view (Specimen figured by Marwick (1953) as Coenothyris sp.; photo Marianna Terezow). 6 BR 3456 (N28/f7454) internal mould (a) dorsal (b) ventral (c) lateral (d) posterior. 7 BR 3424 (N28/f7454) internal mould with remnant shell (a) dorsal (b) ventral (c) lateral (d) posterior (e) anterior. 8 BR3500 (N28/f7455) internal mould, ventral. 9 BR 3498 (N28/f7454) latex of ventral valve interior and beak. 10 BR 3439 (N28/f7454) ventral view. 11 BR 3422 (N28/f7454) internal mould (a) dorsal (b) ventral." figureDoi="http://doi.org/10.5281/zenodo.10148181" httpUri="https://zenodo.org/record/10148181/files/figure.png" pageId="13" pageNumber="14">Fig. 11.5</figureCitation>
). Trechmanns original
<taxonomicName id="48964D38FFA25A70FC29FDBEFB6BFD97" box="[1013,1191,583,609]" class="Rhynchonellata" family="Dielasmatidae" genus="Coenothyris" kingdom="Animalia" order="Terebratulida" pageId="13" pageNumber="14" phylum="Brachiopoda" rank="species" species="undetermined">
<emphasis id="BDE2EAA9FFA25A70FC29FDBEFBB2FD97" box="[1013,1150,583,609]" italics="true" pageId="13" pageNumber="14">Coenothyris</emphasis>
sp.
</taxonomicName>
was examined at the Natural History Museum, London, by Dr J.D. Campbell. He stated in an unpublished MS on Late Triassic athyrids of
<collectingCountry id="F781762BFFA25A70FF6AFD76FE87FD5F" box="[182,331,655,681]" name="New Zealand" pageId="13" pageNumber="14">New Zealand</collectingCountry>
and
<collectingCountry id="F781762BFFA25A70FE5EFD76FDFCFD5F" box="[386,560,655,681]" name="New Caledonia" pageId="13" pageNumber="14">New Caledonia</collectingCountry>
that it is an athyrid.
</paragraph>
<caption id="DBE96633FFA25A70FF4BF8B3FA9CF804" ID-DOI="http://doi.org/10.5281/zenodo.10148179" ID-Zenodo-Dep="10148179" httpUri="https://zenodo.org/record/10148179/files/figure.png" pageId="13" pageNumber="14" startId="13.[151,250,1866,1890]" targetBox="[170,1418,717,1839]" targetPageId="13" targetType="figure">
<paragraph id="8F2936BBFFA25A70FF4BF8B3FD73F894" blockId="13.[151,1436,1866,2035]" box="[151,703,1866,1891]" pageId="13" pageNumber="14">
<emphasis id="BDE2EAA9FFA25A70FF4BF8B3FEECF894" bold="true" box="[151,288,1866,1890]" pageId="13" pageNumber="14">FIGURE 10.</emphasis>
<taxonomicName id="48964D38FFA25A70FEFAF8B3FE09F894" authority="MACFARLAN, 2023" authorityName="MACFARLAN" authorityYear="2023" box="[294,453,1866,1890]" class="Rhynchonellata" family="Dielasmatidae" genus="Rhaetina" higherTaxonomySource="GBIF" kingdom="Animalia" order="Terebratulida" pageId="13" pageNumber="14" phylum="Brachiopoda" rank="species" species="rainei" status="sp. nov.">
<emphasis id="BDE2EAA9FFA25A70FEFAF8B3FE09F894" box="[294,453,1866,1890]" italics="true" pageId="13" pageNumber="14">Rhaetina rainei</emphasis>
</taxonomicName>
<emphasis id="BDE2EAA9FFA25A70FE17F8B2FDC8F895" bold="true" box="[459,516,1867,1891]" pageId="13" pageNumber="14">
<taxonomicNameLabel id="A6D157D2FFA25A70FE17F8B2FDC8F895" box="[459,516,1867,1891]" pageId="13" pageNumber="14" rank="species">n. sp.</taxonomicNameLabel>
</emphasis>
(New Caledonia).
</paragraph>
<paragraph id="8F2936BBFFA25A70FF4BF897FA9CF804" blockId="13.[151,1436,1866,2035]" pageId="13" pageNumber="14">
<emphasis id="BDE2EAA9FFA25A70FF4BF897FF68F870" bold="true" box="[151,164,1902,1926]" pageId="13" pageNumber="14">1</emphasis>
Holotype B876 (NC/f0519) shelly specimen (a) dorsal (b) ventral (c) lateral (d) posterior (e) anterior.
<emphasis id="BDE2EAA9FFA25A70FB0EF897FB13F870" bold="true" box="[1234,1247,1902,1926]" pageId="13" pageNumber="14">2</emphasis>
B875 (NC/f0519) shelly specimen (a) dorsal (b) lateral.
<emphasis id="BDE2EAA9FFA25A70FDC9F86BFDEEF85C" bold="true" box="[533,546,1938,1962]" pageId="13" pageNumber="14">3</emphasis>
B871 (NC/f0503) shelly specimen, ventral.
<emphasis id="BDE2EAA9FFA25A70FC3EF86BFC23F85C" bold="true" box="[994,1007,1938,1962]" pageId="13" pageNumber="14">4</emphasis>
B844 (NC/f0007A) shelly specimen, ventral.
<emphasis id="BDE2EAA9FFA25A70FF1BF84EFF18F839" bold="true" box="[199,212,1975,1999]" pageId="13" pageNumber="14">5</emphasis>
B881 (NC/f0005A) partly shelly specimen (a) dorsal (b) ventral (c) lateral.
<emphasis id="BDE2EAA9FFA25A70FC3DF84FFC22F838" bold="true" box="[993,1006,1974,1998]" pageId="13" pageNumber="14">6</emphasis>
BR 3414 (NC/f0020) shelly specimen (a) dorsal (b) ventral (c) lateral.
<emphasis id="BDE2EAA9FFA25A70FE66F822FE0BF805" bold="true" box="[442,455,2011,2035]" pageId="13" pageNumber="14">7</emphasis>
B880 (NC/f0005A) dorsal view of beak.
<emphasis id="BDE2EAA9FFA25A70FCB1F823FCB6F804" bold="true" box="[877,890,2010,2034]" pageId="13" pageNumber="14">8</emphasis>
B820 (NC/f0087A) shelly specimen, anterior.
</paragraph>
</caption>
<caption id="DBE96633FFA15A73FF4BF927FDC6F804" ID-DOI="http://doi.org/10.5281/zenodo.10148181" ID-Zenodo-Dep="10148181" httpUri="https://zenodo.org/record/10148181/files/figure.png" pageId="14" pageNumber="15" startId="14.[151,250,1758,1782]" targetBox="[164,1428,194,1697]" targetPageId="14" targetType="figure">
<paragraph id="8F2936BBFFA15A73FF4BF927FDA5F900" blockId="14.[151,1437,1758,2034]" box="[151,617,1758,1782]" pageId="14" pageNumber="15">
<emphasis id="BDE2EAA9FFA15A73FF4BF927FED2F900" bold="true" box="[151,286,1758,1782]" pageId="14" pageNumber="15">FIGURE 11.</emphasis>
<taxonomicName id="48964D38FFA15A73FEF9F927FE0FF900" authority="MACFARLAN, 2023" authorityName="MACFARLAN" authorityYear="2023" box="[293,451,1758,1782]" class="Rhynchonellata" family="Dielasmatidae" genus="Rhaetina" higherTaxonomySource="GBIF" kingdom="Animalia" order="Terebratulida" pageId="14" pageNumber="15" phylum="Brachiopoda" rank="species" species="rainei" status="sp. nov.">
<emphasis id="BDE2EAA9FFA15A73FEF9F927FE0FF900" box="[293,451,1758,1782]" italics="true" pageId="14" pageNumber="15">Rhaetina rainei</emphasis>
</taxonomicName>
<emphasis id="BDE2EAA9FFA15A73FE15F927FDCEF900" bold="true" box="[457,514,1758,1782]" pageId="14" pageNumber="15">
<taxonomicNameLabel id="A6D157D2FFA15A73FE15F927FDCEF900" box="[457,514,1758,1782]" pageId="14" pageNumber="15" rank="species">n. sp.</taxonomicNameLabel>
</emphasis>
(Nelson).
</paragraph>
<paragraph id="8F2936BBFFA15A73FF4BF8FBFDC6F804" blockId="14.[151,1437,1758,2034]" pageId="14" pageNumber="15">
<emphasis id="BDE2EAA9FFA15A73FF4BF8FBFF68F8EC" bold="true" box="[151,164,1794,1818]" pageId="14" pageNumber="15">1</emphasis>
BR 3430 (N28/f7454) internal mould (a) dorsal (b) ventral.
<emphasis id="BDE2EAA9FFA15A73FCDCF8FBFCC1F8EC" bold="true" box="[768,781,1794,1818]" pageId="14" pageNumber="15">2</emphasis>
BR 3461 (N28/f7454) internal mould (a) dorsal (b) ventral.
<emphasis id="BDE2EAA9FFA15A73FAB4F8FBFAB9F8EC" bold="true" box="[1384,1397,1794,1818]" pageId="14" pageNumber="15">3</emphasis>
BR 3490 (N28/f7454) latex of dorsal valve interior.
<emphasis id="BDE2EAA9FFA15A73FDA2F8DFFD47F8C8" bold="true" box="[638,651,1830,1854]" pageId="14" pageNumber="15">4</emphasis>
BR 3452 (N28/f7454 internal mould (a) dorsal (b) ventral (c) lateral (d) anterior.
<emphasis id="BDE2EAA9FFA15A73FF1AF8B3FF1FF894" bold="true" box="[198,211,1866,1890]" pageId="14" pageNumber="15">5</emphasis>
BR 1014 (N28/f7522) ventral view (Specimen figured by
<bibRefCitation id="EB074B4AFFA15A73FCFCF8B3FC09F894" author="Marwick, J." box="[800,965,1866,1890]" pageId="14" pageNumber="15" refId="ref15397" refString="Marwick, J. (1953) Divisions and faunas of the Hokonui System (Triassic and Jurassic). New Zealand Geological Survey Paleontological Bulletin 21. New Zealand Geological Survey, Wellington, 141 pp." type="book" year="1953">Marwick (1953)</bibRefCitation>
as
<taxonomicName id="48964D38FFA15A73FC3AF8B3FB4BF897" box="[998,1159,1866,1890]" class="Rhynchonellata" family="Dielasmatidae" genus="Coenothyris" kingdom="Animalia" order="Terebratulida" pageId="14" pageNumber="15" phylum="Brachiopoda" rank="species" species="undetermined">
<emphasis id="BDE2EAA9FFA15A73FC3AF8B3FBAEF894" box="[998,1122,1866,1890]" italics="true" pageId="14" pageNumber="15">Coenothyris</emphasis>
sp.
</taxonomicName>
; photo Marianna Terezow).
<emphasis id="BDE2EAA9FFA15A73FF4BF897FF68F870" bold="true" box="[151,164,1902,1926]" pageId="14" pageNumber="15">6</emphasis>
BR 3456 (N28/f7454) internal mould (a) dorsal (b) ventral (c) lateral (d) posterior.
<emphasis id="BDE2EAA9FFA15A73FBD4F897FBD9F870" bold="true" box="[1032,1045,1902,1926]" pageId="14" pageNumber="15">7</emphasis>
BR 3424 (N28/f7454) internal mould with remnant shell (a) dorsal (b) ventral (c) lateral (d) posterior (e) anterior.
<emphasis id="BDE2EAA9FFA15A73FC78F86BFC7DF85C" bold="true" box="[932,945,1938,1962]" pageId="14" pageNumber="15">8</emphasis>
BR3500 (N28/f7455) internal mould, ventral.
<emphasis id="BDE2EAA9FFA15A73FA4CF86BFA51F85C" bold="true" box="[1424,1437,1938,1962]" pageId="14" pageNumber="15">9</emphasis>
BR 3498 (N28/f7454) latex of ventral valve interior and beak.
<emphasis id="BDE2EAA9FFA15A73FCCDF84FFCE6F838" bold="true" box="[785,810,1974,1998]" pageId="14" pageNumber="15">10</emphasis>
BR 3439 (N28/f7454) ventral view.
<emphasis id="BDE2EAA9FFA15A73FB43F84FFB7BF838" bold="true" box="[1183,1207,1974,1998]" pageId="14" pageNumber="15">11</emphasis>
BR 3422 (N28/f7454) internal mould (a) dorsal (b) ventral.
</paragraph>
</caption>
<caption id="DBE96633FFA05A72FF4BF96AFD13F8E1" ID-DOI="http://doi.org/10.5281/zenodo.10148183" ID-Zenodo-Dep="10148183" httpUri="https://zenodo.org/record/10148183/files/figure.png" pageId="15" pageNumber="16" startId="15.[151,250,1683,1707]" targetBox="[159,1431,214,1659]" targetPageId="15" targetType="figure">
<paragraph id="8F2936BBFFA05A72FF4BF96AFD13F8E1" blockId="15.[151,1436,1683,1815]" pageId="15" pageNumber="16">
<emphasis id="BDE2EAA9FFA05A72FF4BF96AFED3F95D" bold="true" box="[151,287,1683,1707]" pageId="15" pageNumber="16">FIGURE 12.</emphasis>
Detailed views of beak area,
<taxonomicName id="48964D38FFA05A72FD98F96AFD2DF95C" authority="MACFARLAN, 2023" authorityName="MACFARLAN" authorityYear="2023" box="[580,737,1683,1707]" class="Rhynchonellata" family="Dielasmatidae" genus="Rhaetina" higherTaxonomySource="GBIF" kingdom="Animalia" order="Terebratulida" pageId="15" pageNumber="16" phylum="Brachiopoda" rank="species" species="rainei" status="sp. nov.">
<emphasis id="BDE2EAA9FFA05A72FD98F96AFD2DF95C" box="[580,737,1683,1707]" italics="true" pageId="15" pageNumber="16">Rhaetina rainei</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A6D157D2FFA05A72FD3AF96AFCDAF95D" box="[742,790,1683,1707]" pageId="15" pageNumber="16" rank="species">n.sp.</taxonomicNameLabel>
<emphasis id="BDE2EAA9FFA05A72FCFCF96AFCFDF95D" bold="true" box="[800,817,1683,1707]" pageId="15" pageNumber="16">A</emphasis>
BR3458 (N28/f7454) dorsal view of internal mould, showing foramen with pedicle collar, and dorsal valve with low, ridgelike median septum.
<emphasis id="BDE2EAA9FFA05A72FC04F94EFC25F939" bold="true" box="[984,1001,1719,1743]" pageId="15" pageNumber="16">B</emphasis>
BR3456 (N28/f7454) dorsal view of internal mould showing cardinal process.
<emphasis id="BDE2EAA9FFA05A72FDC1F922FDE3F905" bold="true" box="[541,559,1755,1779]" pageId="15" pageNumber="16">C</emphasis>
B877 (NC/f0519) dorsal view of shelly specimen, slightly abraded, showing disjunct deltidial plates and foramen surrounded by pedicle collar.
</paragraph>
</caption>
<paragraph id="8F2936BBFFA05A72FF1BF8BCFC1FF851" blockId="15.[151,1436,1861,1960]" pageId="15" pageNumber="16">
The largest collection is from N28/f7454, GS 196 (Eighty-Eight Valley, Nelson), and the species is common in several collections from shellbeds in the Bouraké Formation in île Hugon,
<collectingCountry id="F781762BFFA05A72FC26F893FB67F875" box="[1018,1195,1897,1924]" name="New Caledonia" pageId="15" pageNumber="16">New Caledonia</collectingCountry>
. Fragments of a large terebratulide from one collection in Southland may belong to this species.
</paragraph>
</subSubSection>
<subSubSection id="C78C6530FFBF5A6DFF1BFF6EFC86FE93" pageId="16" pageNumber="17" type="discussion">
<paragraph id="8F2936BBFFBF5A6DFF1BFF6EFE37FF0F" blockId="16.[151,1437,151,357]" pageId="16" pageNumber="17">
<emphasis id="BDE2EAA9FFBF5A6DFF1BFF6EFEF8FF47" bold="true" box="[199,308,151,177]" pageId="16" pageNumber="17">Remarks</emphasis>
. This species is present in large numbers in the Otapirian of île Hugon and of the Eighty-Eight Valley of
<collectingRegion id="4D52F859FFBF5A6DFF6AFF42FEC5FF23" box="[182,265,187,213]" country="New Zealand" name="Nelson" pageId="16" pageNumber="17">Nelson</collectingRegion>
. It is also present in île Ducos, and a few specimens are known elsewhere. The distribution suggests that it may be a shallow water form.
</paragraph>
<paragraph id="8F2936BBFFBF5A6DFF1BFEFAFC86FE93" blockId="16.[151,1437,151,357]" pageId="16" pageNumber="17">
The variation in shape in this species is considerable, with the larger collections from île Hugon and
<collectingRegion id="4D52F859FFBF5A6DFA91FEFAFA50FEEB" box="[1357,1436,259,285]" country="New Zealand" name="Nelson" pageId="16" pageNumber="17">Nelson</collectingRegion>
showing a range from medium-sized forms with strong folding anteriorly to large, elongate-elliptical specimens with blunt, rounded folds developed only on the anterior part.
</paragraph>
</subSubSection>
</treatment>
</document>