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<document id="7096D93F7E8BD528160D70C2C6238093" ID-CLB-Dataset="4089" ID-DOI="10.5852/ejt.2021.773.1507" ID-GBIF-Dataset="737e33ca-4d43-4320-9c3e-e5e942ceb079" ID-ISSN="2118-9773" ID-Zenodo-Dep="5536285" ID-ZooBank="51990779-969E-412C-B6F5-5E01A8103410" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1632918963903" checkinUser="felipe" docAuthor="Taylor, Paul D., Harmelin, Jean-Georges, Waeschenbach, Andrea &amp; Bouchon, Claude" docDate="2021" docId="0451D341FFD4772DD96E7154FBF0FB24" docLanguage="en" docName="ejt-773_1-18.pdf" docOrigin="European Journal of Taxonomy 773" docSource="http://dx.doi.org/10.5852/ejt.2021.773.1507" docStyle="DocumentStyle:EF2B578F1D15862ADE45B0C07C620911.14:EJT.2018-.journal_article.type1" docStyleId="EF2B578F1D15862ADE45B0C07C620911" docStyleName="EJT.2018-.journal_article.type1" docStyleVersion="14" docTitle="Disporella Gray 1848" docType="treatment" docVersion="9" lastPageNumber="4" masterDocId="F868AB39FFD6772EDB037650FFEFFF93" masterDocTitle="Disporella guada sp. nov., an erect-ramose rectangulate cyclostome (Bryozoa, Stenolaemata) from the Caribbean Sea: convergent evolution in bryozoan colony morphology" masterLastPageNumber="18" masterPageNumber="1" pageNumber="3" updateTime="1699458169252" updateUser="ExternalLinkService" zenodo-license-document="CC-BY-4.0" zenodo-license-figures="CC-BY-4.0" zenodo-license-treatments="UNSPECIFIED">
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<mods:title id="C77476EB50208A4BD494CEC6775F8957">Disporella guada sp. nov., an erect-ramose rectangulate cyclostome (Bryozoa, Stenolaemata) from the Caribbean Sea: convergent evolution in bryozoan colony morphology</mods:title>
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<mods:affiliation id="892CB1C4C10289EA62281B55F182E8B3">Department of Earth Sciences, Natural History Museum, Cromwell Road, London SW 7 5 BD, UK.</mods:affiliation>
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<mods:affiliation id="24CB318166C6339AFE0F74D9D7AC9C86">Aix-Marseille University, Mediterranean Institute of Oceanography (MIO) &amp; GIS Posidonie, OSU Pytheas, Station Marine dEndoume, F- 13007 Marseille, France.</mods:affiliation>
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<mods:affiliation id="F172C924FEA8A4C4A05B535E7E45195C">Department of Life Sciences, Natural History Museum, Cromwell Road, London SW 7 5 BD, UK.</mods:affiliation>
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Genus
<taxonomicName id="4BF819D4FFD4772CD9BC7154FC28F88D" ID-CoL="84N99" authority="Gray, 1848" authorityName="Gray" authorityYear="1848" box="[703,967,1796,1823]" class="Stenolaemata" family="Lichenoporidae" genus="Disporella" kingdom="Animalia" order="Cyclostomatida" pageId="2" pageNumber="3" phylum="Bryozoa" rank="genus">
<emphasis id="BE8CBE45FFD4772CD9BC7154FCD3F88C" bold="true" box="[703,828,1796,1823]" italics="true" pageId="2" pageNumber="3">Disporella</emphasis>
<bibRefCitation id="E8691FA6FFD4772CD8477154FC28F88D" author="Gray J. E." box="[836,967,1796,1822]" pageId="2" pageNumber="3" refId="ref7709" refString="Gray J. E. 1848. List of the Specimens of British Animals in the Collections of the British Museum. Part 1. Centrionae or Radiated Animals. Trustees of the British Museum, London. https: // doi. org / 10.5962 / bhl. title. 1582" type="book" year="1848">Gray, 1848</bibRefCitation>
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<treatmentCitation id="0D594446FFD4772CDBBE7116FDEEF8F3" author="Gray J. E." box="[189,513,1862,1888]" page="138" pageId="2" pageNumber="3" year="1848">
<taxonomicName id="4BF819D4FFD4772CDBBE7116FDEEF8F3" ID-CoL="84N99" authority="Gray, 1848: 138" authorityName="Gray" authorityPageNumber="138" authorityYear="1848" box="[189,513,1862,1888]" class="Stenolaemata" family="Lichenoporidae" genus="Disporella" kingdom="Animalia" order="Cyclostomatida" pageId="2" pageNumber="3" phylum="Bryozoa" rank="genus">
<emphasis id="BE8CBE45FFD4772CDBBE7116FED6F8F3" box="[189,313,1862,1888]" italics="true" pageId="2" pageNumber="3">Disporella</emphasis>
<bibRefCitation id="E8691FA6FFD4772CDA427116FDEEF8F3" author="Gray J. E." box="[321,513,1862,1888]" pageId="2" pageNumber="3" refId="ref7709" refString="Gray J. E. 1848. List of the Specimens of British Animals in the Collections of the British Museum. Part 1. Centrionae or Radiated Animals. Trustees of the British Museum, London. https: // doi. org / 10.5962 / bhl. title. 1582" type="book" year="1848">Gray, 1848: 138</bibRefCitation>
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.
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<emphasis id="BE8CBE45FFD4772CDBBE71D7FEB7F832" bold="true" box="[189,344,1927,1953]" pageId="2" pageNumber="3">Type species</emphasis>
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<taxonomicName id="4BF819D4FFD4772CDBBE71E7FCA8F842" authority="Fleming, 1828" authorityName="Fleming" authorityYear="1828" box="[189,839,1974,2001]" class="Gymnolaemata" family="Umbonulidae" genus="Discopora" higherTaxonomySource="GBIF" kingdom="Animalia" order="Cheilostomatida" pageId="2" pageNumber="3" phylum="Bryozoa" rank="species" species="hispida">
<emphasis id="BE8CBE45FFD4772CDBBE71E7FE79F843" box="[189,406,1974,2000]" italics="true" pageId="2" pageNumber="3">Discopora hispida</emphasis>
<bibRefCitation id="E8691FA6FFD4772CDA9D71E6FDA6F843" author="Fleming J." box="[414,585,1974,2000]" pageId="2" pageNumber="3" refId="ref7633" refString="Fleming J. 1828. A History of British Animals. Bell &amp; Bradfute, Edinburgh. https: // doi. org / 10.5962 / bhl. title. 12859" type="book" year="1828">Fleming, 1828</bibRefCitation>
(
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<bibRefCitation id="E8691FA6FFD4772CD95B71E6FCD1F842" author="Fleming J." box="[600,830,1974,2001]" pageId="2" pageNumber="3" refId="ref7633" refString="Fleming J. 1828. A History of British Animals. Bell &amp; Bradfute, Edinburgh. https: // doi. org / 10.5962 / bhl. title. 12859" type="book" year="1828">Fleming 1828: 530</bibRefCitation>
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.
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<emphasis id="BE8CBE45FFD5772DDBBE7759FEC0FEB0" bold="true" box="[189,303,265,291]" pageId="3" pageNumber="4">Remarks</emphasis>
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<taxonomicName id="4BF819D4FFD5772DDBBE7767FED7FEC2" authorityName="Gray" authorityYear="1848" box="[189,312,311,337]" class="Stenolaemata" family="Lichenoporidae" genus="Disporella" kingdom="Animalia" order="Cyclostomatida" pageId="3" pageNumber="4" phylum="Bryozoa" rank="genus">
<emphasis id="BE8CBE45FFD5772DDBBE7767FED7FEC2" box="[189,312,311,337]" italics="true" pageId="3" pageNumber="4">Disporella</emphasis>
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was introduced by
<bibRefCitation id="E8691FA6FFD5772DD91D7767FD42FEC2" author="Gray J. E." box="[542,685,311,337]" pageId="3" pageNumber="4" refId="ref7709" refString="Gray J. E. 1848. List of the Specimens of British Animals in the Collections of the British Museum. Part 1. Centrionae or Radiated Animals. Trustees of the British Museum, London. https: // doi. org / 10.5962 / bhl. title. 1582" type="book" year="1848">Gray (1848)</bibRefCitation>
in a list of animal specimens in the collections of the British Museum. He included only one species in the genus, albeit with several synonyms, listed by him as
<taxonomicName id="4BF819D4FFD5772DDBBE772DFE76FE04" box="[189,409,381,407]" class="Stenolaemata" family="Tubuliporidae" genus="Tubulipora" kingdom="Animalia" order="Cyclostomatida" pageId="3" pageNumber="4" phylum="Bryozoa" rank="species" species="hispida">
<emphasis id="BE8CBE45FFD5772DDBBE772DFE76FE04" box="[189,409,381,407]" italics="true" pageId="3" pageNumber="4">Tubulipora hispida</emphasis>
</taxonomicName>
.
<bibRefCitation id="E8691FA6FFD5772DDAA4772EFD80FE0B" author="Gray J. E." box="[423,623,382,408]" pageId="3" pageNumber="4" refId="ref7709" refString="Gray J. E. 1848. List of the Specimens of British Animals in the Collections of the British Museum. Part 1. Centrionae or Radiated Animals. Trustees of the British Museum, London. https: // doi. org / 10.5962 / bhl. title. 1582" type="book" year="1848">Gray (1848: 138)</bibRefCitation>
diagnosed
<taxonomicName id="4BF819D4FFD5772DD9EC772DFC85FE04" authorityName="Gray" authorityYear="1848" box="[751,874,381,407]" class="Stenolaemata" family="Lichenoporidae" genus="Disporella" kingdom="Animalia" order="Cyclostomatida" pageId="3" pageNumber="4" phylum="Bryozoa" rank="genus">
<emphasis id="BE8CBE45FFD5772DD9EC772DFC85FE04" box="[751,874,381,407]" italics="true" pageId="3" pageNumber="4">Disporella</emphasis>
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thus: “Orbicular, edge thin, tubes in radiating ridges”. This diagnosis by itself is totally inadequate to recognise the genus. However,
<bibRefCitation id="E8691FA6FFD5772DDE0277F1FEAAFE4C" author="Fleming J." pageId="3" pageNumber="4" refId="ref7633" refString="Fleming J. 1828. A History of British Animals. Bell &amp; Bradfute, Edinburgh. https: // doi. org / 10.5962 / bhl. title. 12859" type="book" year="1828">Flemings (1828: 530)</bibRefCitation>
description of the type species (as
<taxonomicName id="4BF819D4FFD5772DD9D67795FC43FE4D" authorityName="Fleming" authorityYear="1828" box="[725,940,452,478]" class="Gymnolaemata" family="Umbonulidae" genus="Discopora" higherTaxonomySource="GBIF" kingdom="Animalia" order="Cheilostomatida" pageId="3" pageNumber="4" phylum="Bryozoa" rank="species" species="hispida">
<emphasis id="BE8CBE45FFD5772DD9D67795FC43FE4D" box="[725,940,452,478]" italics="true" pageId="3" pageNumber="4">Discopora hispida</emphasis>
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) provides some additional information: “Margin thin and waved, the cells distributed or radiated, with denticulated orifices. Coral resembling the cups and foliage of flowers…. Breadth nearly an inch; hispid, the cells seem distributed over the whole surface, and more vertical than the preceding [
<taxonomicName id="4BF819D4FFD5772DD9DA747EFC38FDD4" box="[729,983,558,583]" class="Gymnolaemata" family="Umbonulidae" genus="Discopora" higherTaxonomySource="GBIF" kingdom="Animalia" order="Cheilostomatida" pageId="3" pageNumber="4" phylum="Bryozoa" rank="species" species="verrucaria">
<emphasis id="BE8CBE45FFD5772DD9DA747EFC38FDD4" box="[729,983,558,583]" italics="true" pageId="3" pageNumber="4">Discopora verrucaria</emphasis>
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]; there are, however, waved porous grooves, and the cells seem disposed on each side of these in irregular transverse rows, united or free, short, with expanding orifices, dividing into irregular spinous processes. This species is very common in Zetland [=
<collectingCountry id="F4EF22C7FFD5772DDA3774C6FE4DFD22" box="[308,418,662,689]" name="United Kingdom" pageId="3" pageNumber="4">Shetland</collectingCountry>
], adhering to
<taxonomicName id="4BF819D4FFD5772DD93D74C7FCD2FD22" box="[574,829,663,689]" class="Gymnolaemata" family="Celleporidae" genus="Cellepora" kingdom="Animalia" order="Cheilostomatida" pageId="3" pageNumber="4" phylum="Bryozoa" rank="species" species="cervicornis">
<emphasis id="BE8CBE45FFD5772DD93D74C7FCD2FD22" box="[574,829,663,689]" italics="true" pageId="3" pageNumber="4">Cellepora cervicornis</emphasis>
</taxonomicName>
[=
<taxonomicName id="4BF819D4FFD5772DD86774C7FB13FD22" authority="(Pallas, 1766)" baseAuthorityName="Pallas" baseAuthorityYear="1766" box="[868,1276,662,689]" class="Gymnolaemata" family="Smittinidae" genus="Smittina" kingdom="Animalia" order="Cheilostomatida" pageId="3" pageNumber="4" phylum="Bryozoa" rank="species" species="cervicornis">
<emphasis id="BE8CBE45FFD5772DD86774C7FBA0FD22" box="[868,1103,663,689]" italics="true" pageId="3" pageNumber="4">Smittina cervicornis</emphasis>
(Pallas, 1766)
</taxonomicName>
]...”
</paragraph>
<paragraph id="8C476257FFD5772DDBBE7489FB46FB9F" blockId="3.[188,1401,728,1037]" pageId="3" pageNumber="4">
As remarked by
<bibRefCitation id="E8691FA6FFD5772DDA877488FDD3FD60" author="Alvarez J. A." box="[388,572,728,755]" pageId="3" pageNumber="4" pagination="201 - 243" refId="ref7249" refString="Alvarez J. A. 1992. Sobre algunas especies de la familia Lichenoporidae Smitt, 1866 (Bryozoa, Cyclostomida) en la region Atlantico-Mediterranea. Parte I: genero Disporella Gray, 1848. Cahiers de Biologie marine 33: 201 - 243." type="journal article" year="1992">Alvarez (1992)</bibRefCitation>
, there is no known holotype of
<taxonomicName id="4BF819D4FFD5772DD8BC7488FB76FD61" baseAuthorityName="Fleming" baseAuthorityYear="1828" box="[959,1177,728,754]" class="Stenolaemata" family="Lichenoporidae" genus="Disporella" kingdom="Animalia" order="Cyclostomatida" pageId="3" pageNumber="4" phylum="Bryozoa" rank="species" species="hispida">
<emphasis id="BE8CBE45FFD5772DD8BC7488FB76FD61" box="[959,1177,728,754]" italics="true" pageId="3" pageNumber="4">Disporella hispida</emphasis>
</taxonomicName>
, nor are there any Fleming specimens from which a lectotype could be chosen. Consequently,
<bibRefCitation id="E8691FA6FFD5772DDF7274ACFAC7FC85" author="Alvarez J. A." box="[1137,1320,764,790]" pageId="3" pageNumber="4" pagination="201 - 243" refId="ref7249" refString="Alvarez J. A. 1992. Sobre algunas especies de la familia Lichenoporidae Smitt, 1866 (Bryozoa, Cyclostomida) en la region Atlantico-Mediterranea. Parte I: genero Disporella Gray, 1848. Cahiers de Biologie marine 33: 201 - 243." type="journal article" year="1992">Alvarez (1992)</bibRefCitation>
chose NHMUK 99.7.1.4187 as the neotype of
<taxonomicName id="4BF819D4FFD5772DD9AC7570FCC1FCAA" baseAuthorityName="Fleming" baseAuthorityYear="1828" box="[687,814,799,825]" class="Stenolaemata" family="Lichenoporidae" genus="Disporella" kingdom="Animalia" order="Cyclostomatida" pageId="3" pageNumber="4" phylum="Bryozoa" rank="species" species="hispida">
<emphasis id="BE8CBE45FFD5772DD9AC7570FCC1FCAA" box="[687,814,799,825]" italics="true" pageId="3" pageNumber="4">D. hispida</emphasis>
</taxonomicName>
. This specimen is the holotype of
<taxonomicName id="4BF819D4FFD5772DDFE2754FFE08FCCE" authority="Lagaaij, 1952" authorityName="Lagaaij" authorityYear="1952" class="Stenolaemata" family="Lichenoporidae" genus="Lichenopora" kingdom="Animalia" order="Cyclostomatida" pageId="3" pageNumber="4" phylum="Bryozoa" rank="species" species="mamillata">
<emphasis id="BE8CBE45FFD5772DDFE2754FFEDBFCCF" italics="true" pageId="3" pageNumber="4">Lichenopora mamillata</emphasis>
<bibRefCitation id="E8691FA6FFD5772DDA437512FE08FCCE" author="Lagaaij R." box="[320,487,834,861]" pageId="3" pageNumber="4" pagination="1 - 233" refId="ref8023" refString="Lagaaij R. 1952. The Pliocene Bryozoa of the Low Countries and their bearing on the marine stratigraphy of the North Sea region. Mededelingen van de Geologische Stichting, Serie C 5: 1 - 233." type="journal article" year="1952">Lagaaij, 1952</bibRefCitation>
</taxonomicName>
, which Alvarez considered to be a junior synonym of
<taxonomicName id="4BF819D4FFD5772DDF8F7513FAE5FCCF" baseAuthorityName="Fleming" baseAuthorityYear="1828" box="[1164,1290,834,860]" class="Stenolaemata" family="Lichenoporidae" genus="Disporella" kingdom="Animalia" order="Cyclostomatida" pageId="3" pageNumber="4" phylum="Bryozoa" rank="species" species="hispida">
<emphasis id="BE8CBE45FFD5772DDF8F7513FAE5FCCF" box="[1164,1290,834,860]" italics="true" pageId="3" pageNumber="4">D. hispida</emphasis>
</taxonomicName>
;
<bibRefCitation id="E8691FA6FFD5772DDE1D7512FEA6FCEC" author="Lagaaij R." pageId="3" pageNumber="4" pagination="1 - 233" refId="ref8023" refString="Lagaaij R. 1952. The Pliocene Bryozoa of the Low Countries and their bearing on the marine stratigraphy of the North Sea region. Mededelingen van de Geologische Stichting, Serie C 5: 1 - 233." type="journal article" year="1952">Lagaaij (1952: 181)</bibRefCitation>
had introduced his new species for a form referred to by
<bibRefCitation id="E8691FA6FFD5772DDF0A7535FB5BFCEC" author="Hincks T." box="[1033,1204,869,895]" pageId="3" pageNumber="4" refId="ref7894" refString="Hincks T. 1880. A History of the British Marine Polyzoa. J. Van Voorst, London. https: // doi. org / 10.5962 / bhl. title. 3978" type="book" year="1880">Hincks (1880)</bibRefCitation>
as
<taxonomicName id="4BF819D4FFD5772DDFE27535FE7CFC30" authority="(Fleming)" baseAuthorityName="Fleming" baseAuthorityYear="1828" class="Stenolaemata" family="Lichenoporidae" genus="Lichenopora" kingdom="Animalia" order="Cyclostomatida" pageId="3" pageNumber="4" phylum="Bryozoa" rank="species" species="hispida">
<emphasis id="BE8CBE45FFD5772DDFE27535FEFCFC31" italics="true" pageId="3" pageNumber="4">Lichenopora hispida</emphasis>
(Fleming)
</taxonomicName>
var. b. It was noted by
<bibRefCitation id="E8691FA6FFD5772DD9BD75D8FBC9FC30" author="Gordon D. P. &amp; Taylor P. D." box="[702,1062,904,931]" pageId="3" pageNumber="4" pagination="243 - 290" refId="ref7664" refString="Gordon D. P. &amp; Taylor P. D. 2001. New Zealand Recent Densiporidae and Lichenoporidae (Bryozoa: Cyclostomata). Species Diversity 6: 243 - 290. https: // doi. org / 10.12782 / specdiv. 6.243" type="journal article" year="2001">Gordon &amp; Taylor (2001: 259)</bibRefCitation>
that the neotype chosen by
<bibRefCitation id="E8691FA6FFD5772DDBBE75FCFE81FC55" author="Alvarez J. A." box="[189,366,940,966]" pageId="3" pageNumber="4" pagination="201 - 243" refId="ref7249" refString="Alvarez J. A. 1992. Sobre algunas especies de la familia Lichenoporidae Smitt, 1866 (Bryozoa, Cyclostomida) en la region Atlantico-Mediterranea. Parte I: genero Disporella Gray, 1848. Cahiers de Biologie marine 33: 201 - 243." type="journal article" year="1992">Alvarez (1992)</bibRefCitation>
is neither topotypic it was collected at Tenby in Wales, not the Shetland Islands almost
<quantity id="4B00CFB2FFD5772DDBBF759FFECCFC7A" box="[188,291,975,1001]" metricMagnitude="6" metricUnit="m" metricValue="1.0" pageId="3" pageNumber="4" unit="km" value="1000.0">1000 km</quantity>
to the north nor does it conform to the original description given by
<bibRefCitation id="E8691FA6FFD5772DDF58759FFAFDFC7A" author="Fleming J." box="[1115,1298,975,1001]" pageId="3" pageNumber="4" refId="ref7633" refString="Fleming J. 1828. A History of British Animals. Bell &amp; Bradfute, Edinburgh. https: // doi. org / 10.5962 / bhl. title. 12859" type="book" year="1828">Fleming (1828)</bibRefCitation>
because it is not hispid (i.e., covered in spines) and the apertures (orifices) are not expanded.
</paragraph>
<paragraph id="8C476257FFD5772DDBBE7263FBF0FB24" blockId="3.[189,1398,1075,1208]" pageId="3" pageNumber="4">
Stabilization of
<taxonomicName id="4BF819D4FFD5772DDA817263FE11FBDE" authorityName="Gray" authorityYear="1848" box="[386,510,1075,1101]" class="Stenolaemata" family="Lichenoporidae" genus="Disporella" kingdom="Animalia" order="Cyclostomatida" pageId="3" pageNumber="4" phylum="Bryozoa" rank="genus">
<emphasis id="BE8CBE45FFD5772DDA817263FE11FBDE" box="[386,510,1075,1101]" italics="true" pageId="3" pageNumber="4">Disporella</emphasis>
</taxonomicName>
awaits comprehensive morphological description of topotype specimens matching
<bibRefCitation id="E8691FA6FFD5772DDA3D7207FDF9FBE2" author="Fleming J." box="[318,534,1111,1137]" pageId="3" pageNumber="4" refId="ref7633" refString="Fleming J. 1828. A History of British Animals. Bell &amp; Bradfute, Edinburgh. https: // doi. org / 10.5962 / bhl. title. 12859" type="book" year="1828">Flemings (1828)</bibRefCitation>
original description of the type species, coupled with molecular characterization. Until this is achieved, the genus name is here used as applied, for instance, by
<bibRefCitation id="E8691FA6FFD5772DDBBE72CDFE15FB2B" author="Hayward P. J. &amp; Ryland J. S." box="[189,506,1181,1208]" pageId="3" pageNumber="4" pagination="1 - 147" refId="ref7854" refString="Hayward P. J. &amp; Ryland J. S. 1985. Cyclostome bryozoans. Keys and notes for the identification of the species. Synopses of the British Fauna. New Series 34: 1 - 147." type="book chapter" year="1985">Hayward &amp; Ryland (1985)</bibRefCitation>
,
<bibRefCitation id="E8691FA6FFD5772DD90772CDFD56FB24" author="Alvarez J. A." box="[516,697,1181,1207]" pageId="3" pageNumber="4" pagination="201 - 243" refId="ref7249" refString="Alvarez J. A. 1992. Sobre algunas especies de la familia Lichenoporidae Smitt, 1866 (Bryozoa, Cyclostomida) en la region Atlantico-Mediterranea. Parte I: genero Disporella Gray, 1848. Cahiers de Biologie marine 33: 201 - 243." type="journal article" year="1992">Alvarez (1992)</bibRefCitation>
, and
<bibRefCitation id="E8691FA6FFD5772DD9F472CDFBF4FB24" author="Gordon D. P. &amp; Taylor P. D." box="[759,1051,1181,1207]" pageId="3" pageNumber="4" pagination="243 - 290" refId="ref7664" refString="Gordon D. P. &amp; Taylor P. D. 2001. New Zealand Recent Densiporidae and Lichenoporidae (Bryozoa: Cyclostomata). Species Diversity 6: 243 - 290. https: // doi. org / 10.12782 / specdiv. 6.243" type="journal article" year="2001">Gordon &amp; Taylor (2001)</bibRefCitation>
.
</paragraph>
</subSubSection>
</treatment>
</document>