160 lines
16 KiB
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160 lines
16 KiB
XML
<document ID-CLB-Dataset="7157" ID-DOI="http://dx.doi.org/10.3897/zookeys.1035.65767" ID-GBIF-Dataset="cbd813c3-228e-4bce-b30a-1a150d778c5b" ID-PMC="PMC8096803" ID-Pensoft-Pub="1313-2970-1035-145" ID-Pensoft-UUID="55CAE096B4A451D88EA433175454DA3F" ID-PubMed="33958933" ID-ZooBank="ADD73A162DB34873890B1E490C2C8841" ModsDocID="1313-2970-1035-145" checkinTime="1619617199447" checkinUser="pensoft" docAuthor="Zamani, Alireza & Marusik, Yuri M." docDate="2021" docId="02249EDF73C059D1843C3DE96E998ED5" docLanguage="en" docName="ZooKeys 1035: 145-193" docOrigin="ZooKeys 1035" docPubDate="2021-04-27" docSource="http://dx.doi.org/10.3897/zookeys.1035.65767" docTitle="Zodarion Walckenaer 1826" docType="treatment" docVersion="6" id="55CAE096B4A451D88EA433175454DA3F" lastPageNumber="145" masterDocId="55CAE096B4A451D88EA433175454DA3F" masterDocTitle="Revision of the spider family Zodariidae (Arachnida, Araneae) in Iran and Turkmenistan, with seventeen new species" masterLastPageNumber="193" masterPageNumber="145" pageNumber="145" updateTime="1732670661048" updateUser="ExternalLinkService">
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<mods:mods id="F03E758FF8C1912DD47346DF1875EA49" xmlns:mods="http://www.loc.gov/mods/v3">
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<mods:titleInfo id="2395D094FD08DB16E675C667B8CFE280">
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<mods:title id="34459F93891F9C0B18430BC8A08E140E">Revision of the spider family Zodariidae (Arachnida, Araneae) in Iran and Turkmenistan, with seventeen new species</mods:title>
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</mods:titleInfo>
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<mods:name id="CFB5185B47D3FD81D0AB2DCB78559074" type="personal">
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<mods:roleTerm id="C65DEF1AD017900DCDA6B8E9FD8F46E2">Author</mods:roleTerm>
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</mods:role>
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<mods:namePart id="5DCAEDDD51F55C0230567504371D58B6">Zamani, Alireza</mods:namePart>
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<mods:nameIdentifier id="E507CB4B0F400B42B5D01B141D474A60" type="ORCID">https://orcid.org/0000-0002-8084-9666</mods:nameIdentifier>
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<mods:affiliation id="BD17B663E76AA73150AA05A44B39CB55">Zoological Museum, Biodiversity Unit, University of Turku, FI- 20014, Finland</mods:affiliation>
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<mods:nameIdentifier id="A5ED81BA533D5FE193B5F4F8F12AC294" type="email">zamani.alireza5@gmail.com</mods:nameIdentifier>
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</mods:name>
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<mods:name id="98A87F847332F7262BCEB9567B23C541" type="personal">
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<mods:roleTerm id="06EAC65B0AF40E04A6F828521D5C490D">Author</mods:roleTerm>
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</mods:role>
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<mods:namePart id="29C03CB1D9B6A22C936F7E24453615C9">Marusik, Yuri M.</mods:namePart>
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<mods:nameIdentifier id="6D2F4B97F2BCB1A34FA6D806E3A5E3D6" type="ORCID">https://orcid.org/0000-0002-4499-5148</mods:nameIdentifier>
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<mods:affiliation id="B5650E5E3C067A3AF4CA6EA206615630">Institute for Biological Problems of the North RAS, Portovaya Str. 18, Magadan, Russia & Department of Zoology & Entomology, University of the Free State, Bloemfontein 9300, South Africa</mods:affiliation>
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</mods:name>
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<mods:typeOfResource id="BED4492627427761D195F40DE1572C04">text</mods:typeOfResource>
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<mods:titleInfo id="1A7FAE6646616414180F71ED1F36B273">
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<mods:title id="A9A27B4414D9F7B427DCF1A7B06637D0">ZooKeys</mods:title>
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</mods:titleInfo>
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<mods:part id="72750BDFCAC6B1219785D88E01B1C6F9">
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<mods:date id="0E2643F48FDDCF790642102F667B776C">2021</mods:date>
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<mods:detail id="2CFB1775CFD547BE7C882487069FB152" type="pubDate">
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<mods:number id="4D571986FAB23B44859B27E2960E07DD">2021-04-27</mods:number>
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</mods:detail>
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<mods:detail id="E9D3F22DD2BEC34BBD0A821A9F623F74" type="volume">
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<mods:number id="F064BE232BB2EF9287AD33BFB39F3789">1035</mods:number>
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</mods:detail>
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<mods:start id="A5A811B8BED9FEB0E6DB2884B2441655">145</mods:start>
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<mods:end id="554E25B80AC8D9D9D73A4E135FF211AB">193</mods:end>
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<mods:location id="3BB052D783B78357441AF27336DEB5CD">
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<mods:url id="9F40D50AA9E2B3D60BD472658A339119">http://dx.doi.org/10.3897/zookeys.1035.65767</mods:url>
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</mods:location>
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<mods:classification id="B6DBA8AC9A2365E784C2C87031330A36">journal article</mods:classification>
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<mods:identifier id="009FE69559633778F49D708F98B02D61" type="DOI">http://dx.doi.org/10.3897/zookeys.1035.65767</mods:identifier>
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<mods:identifier id="4E933EE352932D4E9D2BAD5640086D7F" type="Pensoft-Pub">1313-2970-1035-145</mods:identifier>
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<mods:identifier id="AA77D41AEBDFBE8F191B9975C8FF5D02" type="ZooBank">ADD73A162DB34873890B1E490C2C8841</mods:identifier>
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</mods:mods>
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<treatment id="02249EDF73C059D1843C3DE96E998ED5" ID-GBIF-Taxon="181753573" LSID="urn:lsid:plazi:treatment:02249EDF73C059D1843C3DE96E998ED5" httpUri="http://treatment.plazi.org/id/02249EDF73C059D1843C3DE96E998ED5" lastPageNumber="145" pageId="0" pageNumber="145" scope_class="Arachnida" scope_family="Zodariidae" scope_order="Araneae" scope_subFamily="Zodariinae">
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<subSubSection id="20827C5608BC70AFE03991121EA42FBB" pageId="0" pageNumber="145" type="nomenclature">
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<paragraph id="F57323D5F4BF387FAEB4E4A1F52DCF9B" pageId="0" pageNumber="145">
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Genus
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<taxonomicName id="47A03107E1710609DCE974C709322D57" ID-CoL="8B2B" LSID="02249EDF-73C0-59D1-843C-3DE96E998ED5" authority="Walckenaer, 1826" authorityName="Walckenaer" authorityYear="1826" class="Arachnida" family="Zodariidae" genus="Zodarion" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Zodarion" order="Araneae" pageId="0" pageNumber="145" phylum="Arthropoda" rank="genus">Zodarion Walckenaer, 1826</taxonomicName>
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</paragraph>
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</subSubSection>
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<subSubSection id="0C756A4F82E62D458828E26411AEA910" pageId="0" pageNumber="145" type="type species">
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<paragraph id="AA3D977C13206B9747DA3EFED6760182" pageId="0" pageNumber="145">Type species.</paragraph>
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<paragraph id="7F8D3C2D3C5A9A833515AD0FAD217570" pageId="0" pageNumber="145">
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<taxonomicName id="115C472EE2D87AE4E48D6A38CFE76810" class="Arachnida" family="Zodariidae" genus="Enyo" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Enyo nitida" order="Araneae" pageId="0" pageNumber="145" phylum="Arthropoda" rank="species" species="nitida">
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<emphasis id="117FDD52C239BE7B18D49F660B263B6A" italics="true" pageId="0" pageNumber="145">Enyo nitida</emphasis>
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</taxonomicName>
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Audouin, 1826 from Egypt.
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</paragraph>
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</subSubSection>
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<subSubSection id="FECE839F907964B1787D21132432E37C" pageId="0" pageNumber="145" type="comments">
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<paragraph id="C64B8E8ED26644473A6B18383A27FA40" pageId="0" pageNumber="145">Comments.</paragraph>
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<paragraph id="770B3FA50E3FE105D288BF2C86B81558" pageId="0" pageNumber="145">
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With 177 named species, this is the largest genus within
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<taxonomicName id="DFBEA76ADD13A99871546F92434385E4" authorityName="Thorell" authorityYear="1881" lsidName="" pageId="0" pageNumber="145" rank="subFamily" subFamily="Zodariinae">Zodariinae</taxonomicName>
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(
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<bibRefCitation id="C05AB38B7C6D7AF1D0F69EE51548D4D8" author="WSC" journalOrPublisher="Trudy Soveta po Izuceniju Proizvoditeljnych Sil, Serija Turkmenskaja" pageId="0" pageNumber="145" publicationUrl="http://wsc.nmbe.ch" refId="B26" refString="WSC, 2021. World Spider Catalog. Version 21.5. Natural History Museum Bern, http://wsc.nmbe.ch" title="World Spider Catalog. Version 21.5. Natural History Museum Bern," url="http://wsc.nmbe.ch" year="2021">WSC 2021</bibRefCitation>
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). Based on the copulatory organs of the species currently considered in this genus,
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<taxonomicName id="B3FA8F757BAE53D3B79F767ECACF0022" authorityName="Walckenaer" authorityYear="1826" class="Arachnida" family="Zodariidae" genus="Zodarion" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Zodarion" order="Araneae" pageId="0" pageNumber="145" phylum="Arthropoda" rank="genus">
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<emphasis id="515E6BA3F5881C9037F4ADCF4569B76B" italics="true" pageId="0" pageNumber="145">Zodarion</emphasis>
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</taxonomicName>
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does not appear to be monophyletic. There are only two species that are morphologically similar to the generotype:
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<taxonomicName id="DC0AE2A849AC5F3D8C255AB65F217583" lsidName="Z. luctuosum" pageId="0" pageNumber="145" rank="species" species="luctuosum">
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<emphasis id="F48A6AB60F66F3BBE2A2DEB115D5537E" italics="true" pageId="0" pageNumber="145">Z. luctuosum</emphasis>
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</taxonomicName>
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(O. Pickard-Cambridge, 1872) and
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<taxonomicName id="0A42897BDB14512B2D3DDC403E6EEB0D" lsidName="Z. lutipes" pageId="0" pageNumber="145" rank="species" species="lutipes">
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<emphasis id="49C06D8660EA049914C72E95F23C302F" italics="true" pageId="0" pageNumber="145">Z. lutipes</emphasis>
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</taxonomicName>
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(O. Pickard-Cambridge, 1872), from the eastern Mediterranean (east of Tunisia to Iran).
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</paragraph>
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</subSubSection>
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<subSubSection id="AFC0DECB1E432B5AC42F55A6D1EF941C" pageId="0" pageNumber="145" type="materials_examined">
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<paragraph id="FDA07F83F6FE446D984509F47C3CDDC8" pageId="0" pageNumber="145">Comparative material.</paragraph>
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<paragraph id="9E9F4963AA4FB08881BFC4A3E8A72F4E" pageId="0" pageNumber="145">
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<taxonomicName id="3B29D01F09ED3D6BA6BBB59E99944192" class="Arachnida" family="Zodariidae" genus="Zodarion" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Zodarion expers" order="Araneae" pageId="0" pageNumber="145" phylum="Arthropoda" rank="species" species="expers">
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<emphasis id="D1292117F21BA97FB2AF0697ABDC4782" italics="true" pageId="0" pageNumber="145">Zodarion expers</emphasis>
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</taxonomicName>
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(O. Pickard-Cambridge, 1876),
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<materialsCitation id="4D435058BD88808C86C253D33C016488" ID-GBIF-Occurrence="3093634307" collectingDate="2021-01-01" collectingDateMax="2021-12-31" collectingDateMin="2021-01-01" country="Israel" location="Israel" specimenCount="2" specimenCount-male="2">
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<specimenCount id="5071CE3BD50261CB86F0EF22CA7BDAD9" type="male">1♂</specimenCount>
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from
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<collectingCountry id="0DD4215C26A66F76E65A44991DB195A6" name="Israel">Israel</collectingCountry>
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(Figs
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<figureCitation id="C0FD670EA72F7D67F5A8E954F92A0F02" captionStart="Figure 19" captionStartId="F19" captionText="Figure 19. Male habitus and palps of Acanthinozodium parmida sp. nov. (A-C) and Zodarion expers (D-F) A, D habitus, dorsal B, E palp, ventral C, F palp, retrolateral. Scale bar: 0.2 mm (A)." figureDoi="10.3897/zookeys.1035.65767.figure19" httpUri="https://binary.pensoft.net/fig/536896" pageId="0" pageNumber="145">19D-F</figureCitation>
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,
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<figureCitation id="C66B6858C629A09A41F52678942D56D4" captionStart="Figure 25" captionStartId="F25" captionText="Figure 25. SEM images of the male palps of Acanthinozodium parmida sp. nov. (A-C) and Zodarion expers (D-F) A, B, D ventral C, E retrolateral F dorsoretrolateral. Abbreviations: Cg - cymbial groove, Co - conductor, Em - embolus, Er - embolar ridge, Ma - median apophysis, Ra - retrolateral tibial apophysis. Scale bars: 0.1 mm." figureDoi="10.3897/zookeys.1035.65767.figure25" httpUri="https://binary.pensoft.net/fig/536902" pageId="0" pageNumber="145">25D-F</figureCitation>
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) and
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<taxonomicName id="7146295478D7D1F6E9244C53097055C6" baseAuthorityName="Audouin" baseAuthorityYear="1826" class="Arachnida" family="Zodariidae" genus="Zodarion" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Zodarion nitidum" order="Araneae" pageId="0" pageNumber="145" phylum="Arthropoda" rank="species" species="nitidum">
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<emphasis id="0E8CC7027D40D607B0E85F92C8447757" italics="true" pageId="0" pageNumber="145">Zodarion nitidum</emphasis>
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</taxonomicName>
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(Audouin, 1826):
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<specimenCount id="155930A6EE02471FB027C119F53F5F27" type="male">1♂</specimenCount>
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from
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<collectingCountry id="8B71DF92A1B97C49683FFE5DF75A1EF7" name="Israel">Israel</collectingCountry>
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(Fig.
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<figureCitation id="7B50D0951E658D5C0DBE12A183D27F8F" captionStart="Figure 28" captionStartId="F28" captionText="Figure 28. SEM images of the male palps of Zodarion nitidum (A-C) and Z. bactrianum (D-F) A, D ventral B, E retrolateral C dorsal F retrodorsal. Abbreviations: Co - conductor, Eb - embolar base, Em - embolus, Ma - median apophysis, Mc - claw of median apophysis, Ra - retrolateral tibial apophysis, Tp - tegular process, Tu - tutaculum, Va - ventral apophysis. Scale bars: 0.1 mm." figureDoi="10.3897/zookeys.1035.65767.figure28" httpUri="https://binary.pensoft.net/fig/536905" pageId="0" pageNumber="145">28A-C</figureCitation>
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)
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</materialsCitation>
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</paragraph>
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</subSubSection>
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<subSubSection id="0FADA4FBE19B27EE4A69AFF650F35BD3" pageId="0" pageNumber="145" type="comments">
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<paragraph id="164E39919DF6E328EF23608B29B61DAF" pageId="0" pageNumber="145">Comments.</paragraph>
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<paragraph id="8C34E0AB325456F06198E651D00EBDFC" pageId="0" pageNumber="145">
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Until recently,
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<taxonomicName id="BB865E18A2720E571E982147199500BE" lsidName="Z. expers" pageId="0" pageNumber="145" rank="species" species="expers">
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<emphasis id="C71D61FAE2379104D1B2F2C9B85CED09" italics="true" pageId="0" pageNumber="145">Z. expers</emphasis>
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</taxonomicName>
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was placed in
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<taxonomicName id="A6858C0602702035FA45B5A15E759F5D" authorityName="Jocque" authorityYear="1991" class="Arachnida" family="Zodariidae" genus="Ranops" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Ranops" order="Araneae" pageId="0" pageNumber="145" phylum="Arthropoda" rank="genus">
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<emphasis id="91EE6D153201F2F4EB13F397B9C8CB33" italics="true" pageId="0" pageNumber="145">Ranops</emphasis>
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</taxonomicName>
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<normalizedToken id="C848D4BC653BD01B5E7FFAE7BE307045" originalValue="Jocqué">Jocque</normalizedToken>
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, 1991, but now it is placed in
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<taxonomicName id="1C7D7790739EBA24233509A649D2BFA2" authorityName="Walckenaer" authorityYear="1826" class="Arachnida" family="Zodariidae" genus="Zodarion" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Zodarion" order="Araneae" pageId="0" pageNumber="145" phylum="Arthropoda" rank="genus">
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<emphasis id="E9327B15B34E680F8EBDB863DC117FD1" italics="true" pageId="0" pageNumber="145">Zodarion</emphasis>
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</taxonomicName>
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. It differs from the generotype of
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<taxonomicName id="C67344BB633707FF4126777440A68D12" authorityName="Jocque" authorityYear="1991" class="Arachnida" family="Zodariidae" genus="Ranops" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Ranops" order="Araneae" pageId="0" pageNumber="145" phylum="Arthropoda" rank="genus">
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<emphasis id="78F8CB800925D0FC93C169CE540C2C0A" italics="true" pageId="0" pageNumber="145">Ranops</emphasis>
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</taxonomicName>
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, but the copulatory organs are also very different from those of
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<taxonomicName id="24BFF4C6E6F2596F0726A0F138E158B5" lsidName="Z. nitidum" pageId="0" pageNumber="145" rank="species" species="nitidum">
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<emphasis id="DDB00CC34670707472BD2151C274A103" italics="true" pageId="0" pageNumber="145">Z. nitidum</emphasis>
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</taxonomicName>
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, and most likely it represents a separate genus. Although it is absent in Iran or Turkmenistan, we have provided figures of this species to illustrate the differences with the generotype of
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<taxonomicName id="90E2A520B9CCACA16748ADC34DDD3C31" authorityName="Walckenaer" authorityYear="1826" class="Arachnida" family="Zodariidae" genus="Zodarion" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Zodarion" order="Araneae" pageId="0" pageNumber="145" phylum="Arthropoda" rank="genus">
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<emphasis id="B4A13C221605FAB2545AE40877D8F3D9" italics="true" pageId="0" pageNumber="145">Zodarion</emphasis>
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</taxonomicName>
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. Furthermore, although
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<taxonomicName id="0836D45646F457A4FA4B25B0DE5823C6" lsidName="Z. nitidum" pageId="0" pageNumber="145" rank="species" species="nitidum">
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<emphasis id="E009DD5F9048FB07135881C4BF4DD618" italics="true" pageId="0" pageNumber="145">Z. nitidum</emphasis>
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</taxonomicName>
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has not been recorded in the current study area, because it is the type species of the genus, we have provided illustartions for this species as well, to demonstrate its differences with other
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<taxonomicName id="ADED0C1D71FB722A1EB3020E9E7265B0" authorityName="Thorell" authorityYear="1881" lsidName="" pageId="0" pageNumber="145" rank="subFamily" subFamily="Zodariinae">Zodariinae</taxonomicName>
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genera in Iran and Turkmenistan, as well as with other species currently placed in
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<taxonomicName id="E1B59239019B906425258E8D7E3FB10B" authorityName="Walckenaer" authorityYear="1826" class="Arachnida" family="Zodariidae" genus="Zodarion" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Zodarion" order="Araneae" pageId="0" pageNumber="145" phylum="Arthropoda" rank="genus">
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<emphasis id="CF42DDAA6AC98587437FB890BF576FD4" italics="true" pageId="0" pageNumber="145">Zodarion</emphasis>
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</taxonomicName>
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.
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</paragraph>
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</subSubSection>
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</treatment>
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</document> |