treatments-xml/data/03/F8/C1/03F8C140FF4F94C5FF4EF944FE68FC82.xml
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<document id="1A12A2C55ED84751E9A3740C949B14E6" ID-DOI="10.5281/zenodo.5171097" ID-ISSN="1942-1354" ID-Zenodo-Dep="5171097" ID-ZooBank="0011FDFF-35F5-4B7E-B952-7FD2B29D538B" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1628293865072" checkinUser="felipe" docAuthor="Canepari, Claudio, Gordon, Robert D. &amp; Hanley, Guy A." docDate="2016" docId="03F8C140FF4F94C5FF4EF944FE68FC82" docLanguage="en" docName="InsectaMundi.0486.1-180.pdf" docOrigin="Insecta Mundi 2016 (486)" docStyle="DocumentStyle:6D85C66E4B52897F25DA165649BC06DC.3:InsectaMundi.2009-2020.journal_article" docStyleId="6D85C66E4B52897F25DA165649BC06DC" docStyleName="InsectaMundi.2009-2020.journal_article" docStyleVersion="3" docTitle="Tiphysa Mulsant 1850" docType="treatment" docVersion="8" lastPageNumber="133" masterDocId="FFC1B938FFCA9443FF8EFFC7FFC2FFF9" masterDocTitle="South American Coccinellidae (Coleoptera), Part XVII: Systematic revision of the genera Cyrea Gordon and Canepari and Tiphysa Mulsant (Hyperaspidinae: Brachiacanthini)" masterLastPageNumber="180" masterPageNumber="1" pageNumber="132" updateTime="1699041177162" updateUser="plazi" zenodo-license-document="CC-BY-NC-3.0">
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<mods:title id="18B61D9804BC949D06CC52202F7E0F92">South American Coccinellidae (Coleoptera), Part XVII: Systematic revision of the genera Cyrea Gordon and Canepari and Tiphysa Mulsant (Hyperaspidinae: Brachiacanthini)</mods:title>
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Genus
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<emphasis id="B925AC44FF4F94C6FE95F944FE4AF962" bold="true" box="[283,392,1667,1691]" italics="true" pageId="133" pageNumber="132">Tiphysa</emphasis>
Mulsant
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<taxonomicName id="4C510BD5FF4F94C6FF4EF904FDD3F922" authority="Mulsant, 1850: 517" authorityName="Mulsant" authorityPageNumber="517" authorityYear="1850" box="[192,529,1731,1755]" class="Insecta" family="Coccinellidae" genus="Tiphysa" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="133" pageNumber="132" phylum="Arthropoda" rank="genus">
<emphasis id="B925AC44FF4F94C6FF4EF904FEE1F922" box="[192,291,1731,1755]" italics="true" pageId="133" pageNumber="132">Tiphysa</emphasis>
<bibRefCitation id="EFC00DA7FF4F94C6FEA4F904FDD3F922" author="Mulsant, M. E." box="[298,529,1731,1755]" pageId="133" pageNumber="132" pagination="1 - 1104" refId="ref106003" refString="Mulsant, M. E. 1850. Species de Coleopteres trimeres securipalpes. Annales des Sciences Physiques et Naturelles, Lyon 2: 1 - 1104." type="journal article" year="1850">Mulsant, 1850:517</bibRefCitation>
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;
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<bibRefCitation id="EFC00DA7FF4F94C6FD93F904FD33F922" author="Crotch, G. R." box="[541,753,1731,1755]" pageId="133" pageNumber="132" refId="ref105276" refString="Crotch, G. R. 1874. A revision of the coleopterous family Coccinellidae. E. W. Janson. London, 311 p." type="book" year="1874">Crotch, 1874:210</bibRefCitation>
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;
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;
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<bibRefCitation id="EFC00DA7FF4F94C6FAB3F904FE91F902" author="Gordon, R. D." pageId="133" pageNumber="132" pagination="1 - 46" refId="ref105422" refString="Gordon, R. D. 1987. A catalogue of the Crotch collection of Coccinellidae (Coleoptera). Occasional Papers on Systematic Entomology 3: 1 - 46." type="journal article" year="1987">Gordon, 1987:26</bibRefCitation>
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;
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;
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<bibRefCitation id="EFC00DA7FF4F94C6FDD9F924FC03F902" author="Milleo, J. M. &amp; M. Almeida" box="[599,961,1763,1787]" pageId="133" pageNumber="132" pagination="274 - 280" refId="ref105924" refString="Milleo, J. M., and M. Almeida L. 2003. Revision of the Genus Tiphysa Mulsant (Coleoptera, Coccinellidae, Hyperaspinae). Coleopterists Bulletin 57: 274 - 280." type="journal article" year="2003">Millèo and Almeida, 2003:274</bibRefCitation>
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.
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<typeStatus id="54EACEF4FF4F94C6FF4EF8C3FF3FF8E5" box="[192,253,1796,1820]" pageId="133" pageNumber="132">Type</typeStatus>
species:
<taxonomicName id="4C510BD5FF4F94C6FEEAF8C4FD2AF8E2" authority="Mulsant, 1850" authorityName="Mulsant" authorityYear="1850" box="[356,744,1795,1819]" class="Insecta" family="Coccinellidae" genus="Tiphysa" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="133" pageNumber="132" phylum="Arthropoda" rank="species" species="plumbea">
<emphasis id="B925AC44FF4F94C6FEEAF8C4FDF1F8E2" box="[356,563,1795,1819]" italics="true" pageId="133" pageNumber="132">Tiphysa plumbea</emphasis>
<bibRefCitation id="EFC00DA7FF4F94C6FDB7F8C4FD2AF8E2" author="Mulsant, M. E." box="[569,744,1795,1819]" pageId="133" pageNumber="132" pagination="1 - 1104" refId="ref106003" refString="Mulsant, M. E. 1850. Species de Coleopteres trimeres securipalpes. Annales des Sciences Physiques et Naturelles, Lyon 2: 1 - 1104." type="journal article" year="1850">Mulsant, 1850</bibRefCitation>
</taxonomicName>
, by monotypy.
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<subSubSection id="C34B23DDFF4F94C5FF4EF884FDE2FE25" lastPageId="134" lastPageNumber="133" pageId="133" pageNumber="132" type="description">
<paragraph id="8BEE7056FF4F94C5FF4EF884FDE2FE25" blockId="133.[192,1441,1859,1916]" lastBlockId="134.[192,1441,195,476]" lastPageId="134" lastPageNumber="133" pageId="133" pageNumber="132">
<emphasis id="B925AC44FF4F94C6FF4EF884FEA1F8A2" bold="true" box="[192,355,1859,1883]" pageId="133" pageNumber="132">Description</emphasis>
:
<taxonomicName id="4C510BD5FF4F94C6FEFCF884FDFAF8A2" authorityName="Mulsant" authorityYear="1850" box="[370,568,1859,1883]" class="Insecta" family="Coccinellidae" kingdom="Animalia" order="Coleoptera" pageId="133" pageNumber="132" phylum="Arthropoda" rank="tribe" tribe="Brachiacanthini">Brachiacanthini</taxonomicName>
with form oval, convex. Dorsal surface of pronotum and elytra entirely black with metallic blue or green tint (
<figureCitation id="136A6CD3FF4F94C6FD18F8A4FD3CF882" box="[662,766,1891,1915]" captionStart="Figures 655-662" captionStartId="181.[192,285,1129,1151]" captionTargetBox="[227,1405,215,1107]" captionTargetId="figure-16@181.[215,1415,197,1117]" captionTargetPageId="181" captionText="Figures 655-662. Tiphysa spp. 655-661) Tiphysa plumbea 655) Habitus. 656. Protibia.657) Maxilla.658) Phallobase ventral. 659) Phallobase lateral. 660) Sipho. 661). Female genitalia. 662) Tiphysa egae 662) Female genitalia." figureDoi="http://doi.org/10.5281/zenodo.5171224" httpUri="https://zenodo.org/record/5171224/files/figure.png" pageId="133" pageNumber="132">Fig. 655</figureCitation>
). Head partly yellow in male, brown or black in female. Antenna with 11 articles, basal article twice as long as wide, antennal insertion exposed. Clypeus with apical border emarginate. Labrum rectangular. Apical maxillary palpomere securiform with sides diverging (
<figureCitation id="136A6CD3FF4C94C5FF76FEC4FEA3FEE2" box="[248,353,259,283]" captionStart="Figures 655-662" captionStartId="181.[192,285,1129,1151]" captionTargetBox="[227,1405,215,1107]" captionTargetId="figure-16@181.[215,1415,197,1117]" captionTargetPageId="181" captionText="Figures 655-662. Tiphysa spp. 655-661) Tiphysa plumbea 655) Habitus. 656. Protibia.657) Maxilla.658) Phallobase ventral. 659) Phallobase lateral. 660) Sipho. 661). Female genitalia. 662) Tiphysa egae 662) Female genitalia." figureDoi="http://doi.org/10.5281/zenodo.5171224" httpUri="https://zenodo.org/record/5171224/files/figure.png" pageId="134" pageNumber="133">Fig. 657</figureCitation>
). Mandible large, robust, with apical tooth and subapical tooth. Scutellum small, longer than wide. Elytral epipleuron narrow, deeply excavated for reception of tibiae, strongly descending externally. Prosternal process slightly convex, with two carinae. Protibia short, with large, arcuate flange smooth on outer margin (
<figureCitation id="136A6CD3FF4C94C5FD8FFEA4FDA7FE82" box="[513,613,355,379]" captionStart="Figures 655-662" captionStartId="181.[192,285,1129,1151]" captionTargetBox="[227,1405,215,1107]" captionTargetId="figure-16@181.[215,1415,197,1117]" captionTargetPageId="181" captionText="Figures 655-662. Tiphysa spp. 655-661) Tiphysa plumbea 655) Habitus. 656. Protibia.657) Maxilla.658) Phallobase ventral. 659) Phallobase lateral. 660) Sipho. 661). Female genitalia. 662) Tiphysa egae 662) Female genitalia." figureDoi="http://doi.org/10.5281/zenodo.5171224" httpUri="https://zenodo.org/record/5171224/files/figure.png" pageId="134" pageNumber="133">Fig 656</figureCitation>
). Abdomen without visible gland pores between ventrites 4 and 5. Tarsal claw with subquadrate basal tooth. Postcoxal line on basal abdominal sternite incomplete, of
<taxonomicName id="4C510BD5FF4C94C5FF4EFE64FE77FE42" authority="(Scymnus)" box="[192,437,419,443]" class="Insecta" family="Coccinellidae" genus="Scymnus" kingdom="Animalia" order="Coleoptera" pageId="134" pageNumber="133" phylum="Arthropoda" rank="subGenus" subGenus="Scymnus">
<emphasis id="B925AC44FF4C94C5FF4EFE64FEECFE43" box="[192,302,419,442]" italics="true" pageId="134" pageNumber="133">Scymnus</emphasis>
(
<emphasis id="B925AC44FF4C94C5FEB0FE64FE6EFE43" box="[318,428,419,442]" italics="true" pageId="134" pageNumber="133">Scymnus</emphasis>
)
</taxonomicName>
<typeStatus id="54EACEF4FF4C94C5FE32FE63FE36FE45" box="[444,500,420,444]" pageId="134" pageNumber="133">type</typeStatus>
. Male genitalia with basal lobe asymmetrical (
<figureCitation id="136A6CD3FF4C94C5FBADFE64FB06FE42" box="[1059,1220,419,443]" captionStart="Figures 655-662" captionStartId="181.[192,285,1129,1151]" captionTargetBox="[227,1405,215,1107]" captionTargetId="figure-16@181.[215,1415,197,1117]" captionTargetPageId="181" captionText="Figures 655-662. Tiphysa spp. 655-661) Tiphysa plumbea 655) Habitus. 656. Protibia.657) Maxilla.658) Phallobase ventral. 659) Phallobase lateral. 660) Sipho. 661). Female genitalia. 662) Tiphysa egae 662) Female genitalia." figureDoi="http://doi.org/10.5281/zenodo.5171224" httpUri="https://zenodo.org/record/5171224/files/figure.png" pageId="134" pageNumber="133">Fig. 658, 659</figureCitation>
). Female genitalia typical
<taxonomicName id="4C510BD5FF4C94C5FE96FE04FE1CFE22" authorityName="Mulsant" authorityYear="1850" box="[280,478,451,475]" class="Insecta" family="Coccinellidae" kingdom="Animalia" order="Coleoptera" pageId="134" pageNumber="133" phylum="Arthropoda" rank="tribe" tribe="Brachiacanthini">Brachiacanthini</taxonomicName>
<typeStatus id="54EACEF4FF4C94C5FE6AFE03FDDEFE25" box="[484,540,452,476]" pageId="134" pageNumber="133">type</typeStatus>
.
</paragraph>
</subSubSection>
<subSubSection id="C34B23DDFF4C94C5FF4EFDC4FE68FC82" pageId="134" pageNumber="133" type="discussion">
<paragraph id="8BEE7056FF4C94C5FF4EFDC4FADEFDC2" blockId="134.[192,1440,515,891]" pageId="134" pageNumber="133">
<emphasis id="B925AC44FF4C94C5FF4EFDC4FEF9FDE2" bold="true" box="[192,315,515,539]" pageId="134" pageNumber="133">Remarks</emphasis>
:
<taxonomicName id="4C510BD5FF4C94C5FEC7FDC4FE69FDE2" authorityName="Mulsant" authorityYear="1850" box="[329,427,515,539]" class="Insecta" family="Coccinellidae" genus="Tiphysa" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="134" pageNumber="133" phylum="Arthropoda" rank="genus">
<emphasis id="B925AC44FF4C94C5FEC7FDC4FE69FDE2" box="[329,427,515,539]" italics="true" pageId="134" pageNumber="133">Tiphysa</emphasis>
</taxonomicName>
was reviewed and correctly assigned to
<taxonomicName id="4C510BD5FF4C94C5FC09FDC4FB8EFDE2" authorityName="Mulsant" authorityYear="1850" box="[903,1100,515,539]" class="Insecta" family="Coccinellidae" kingdom="Animalia" order="Coleoptera" pageId="134" pageNumber="133" phylum="Arthropoda" rank="tribe" tribe="Brachiacanthini">Brachiacanthini</taxonomicName>
by
<bibRefCitation id="EFC00DA7FF4C94C5FBFAFDC4FA5FFDE2" author="Milleo, J. M. &amp; M. Almeida" box="[1140,1437,515,539]" pageId="134" pageNumber="133" pagination="274 - 280" refId="ref105924" refString="Milleo, J. M., and M. Almeida L. 2003. Revision of the Genus Tiphysa Mulsant (Coleoptera, Coccinellidae, Hyperaspinae). Coleopterists Bulletin 57: 274 - 280." type="journal article" year="2003">Milléo &amp; Almeida (2003)</bibRefCitation>
. That publication should be referred to for detailed illustrations of morphological characters.
</paragraph>
<paragraph id="8BEE7056FF4C94C5FF7EFD84FD5AFD22" blockId="134.[192,1440,515,891]" pageId="134" pageNumber="133">
<taxonomicName id="4C510BD5FF4C94C5FF7EFD84FE90FDA2" authorityName="Mulsant" authorityYear="1850" box="[240,338,579,603]" class="Insecta" family="Coccinellidae" genus="Tiphysa" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="134" pageNumber="133" phylum="Arthropoda" rank="genus">
<emphasis id="B925AC44FF4C94C5FF7EFD84FE90FDA2" box="[240,338,579,603]" italics="true" pageId="134" pageNumber="133">Tiphysa</emphasis>
</taxonomicName>
is not strongly distinguished from
<taxonomicName id="4C510BD5FF4C94C5FD7BFD83FCF9FDA2" box="[757,827,580,603]" class="Globothalamea" genus="Cyrea" kingdom="Chromista" order="Textulariida" pageId="134" pageNumber="133" phylum="Foraminifera" rank="genus">
<emphasis id="B925AC44FF4C94C5FD7BFD83FCF9FDA2" box="[757,827,580,603]" italics="true" pageId="134" pageNumber="133">Cyrea</emphasis>
</taxonomicName>
, they both lack abdominal gland openings between abdominal ventrites 4-5, but
<taxonomicName id="4C510BD5FF4C94C5FD90FDA4FD43FD82" authorityName="Mulsant" authorityYear="1850" box="[542,641,611,635]" class="Insecta" family="Coccinellidae" genus="Tiphysa" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="134" pageNumber="133" phylum="Arthropoda" rank="genus">
<emphasis id="B925AC44FF4C94C5FD90FDA4FD43FD82" box="[542,641,611,635]" italics="true" pageId="134" pageNumber="133">Tiphysa</emphasis>
</taxonomicName>
has epipleurae strongly descending externally, a black, dorsal surface with a metallic luster, and strongly flanged protibia. Nearly all members of
<taxonomicName id="4C510BD5FF4C94C5FBF8FD43FB7EFD62" box="[1142,1212,644,667]" class="Globothalamea" genus="Cyrea" kingdom="Chromista" order="Textulariida" pageId="134" pageNumber="133" phylum="Foraminifera" rank="genus">
<emphasis id="B925AC44FF4C94C5FBF8FD43FB7EFD62" box="[1142,1212,644,667]" italics="true" pageId="134" pageNumber="133">Cyrea</emphasis>
</taxonomicName>
lack a well-defined protibial flange, the epipleurae do not have a strongly descending external margin, and no known species has an entirely black, metallic dorsum.
</paragraph>
<paragraph id="8BEE7056FF4C94C5FF7EFD23FE68FC82" blockId="134.[192,1440,515,891]" pageId="134" pageNumber="133">
<typeStatus id="54EACEF4FF4C94C5FF7EFD23FEEFFD05" box="[240,301,740,764]" pageId="134" pageNumber="133">Type</typeStatus>
specimens of
<taxonomicName id="4C510BD5FF4C94C5FE5DFD24FDA6FD02" box="[467,612,739,763]" class="Insecta" family="Coccinellidae" genus="Tiphysa" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="134" pageNumber="133" phylum="Arthropoda" rank="species" species="ega">
<emphasis id="B925AC44FF4C94C5FE5DFD24FDA6FD02" box="[467,612,739,763]" italics="true" pageId="134" pageNumber="133">Tiphysa ega</emphasis>
</taxonomicName>
e and two unlabeled specimens of
<taxonomicName id="4C510BD5FF4C94C5FC77FD23FB43FD02" authorityName="Mulsant" authorityYear="1850" box="[1017,1153,739,763]" class="Insecta" family="Coccinellidae" genus="Tiphysa" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="134" pageNumber="133" phylum="Arthropoda" rank="species" species="plumbea">
<emphasis id="B925AC44FF4C94C5FC77FD23FBCEFD02" box="[1017,1036,740,763]" italics="true" pageId="134" pageNumber="133">T</emphasis>
.
<emphasis id="B925AC44FF4C94C5FB97FD24FB43FD02" box="[1049,1153,739,763]" italics="true" pageId="134" pageNumber="133">plumbea</emphasis>
</taxonomicName>
in the Crotch collection are the only
<taxonomicName id="4C510BD5FF4C94C5FED1FCC4FE00FCE2" authorityName="Mulsant" authorityYear="1850" box="[351,450,771,795]" class="Insecta" family="Coccinellidae" genus="Tiphysa" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="134" pageNumber="133" phylum="Arthropoda" rank="genus">
<emphasis id="B925AC44FF4C94C5FED1FCC4FE00FCE2" box="[351,450,771,795]" italics="true" pageId="134" pageNumber="133">Tiphysa</emphasis>
</taxonomicName>
found among many
<taxonomicName id="4C510BD5FF4C94C5FD4DFCC4FC4BFCE2" authorityName="Mulsant" authorityYear="1850" box="[707,905,771,795]" class="Insecta" family="Coccinellidae" kingdom="Animalia" order="Coleoptera" pageId="134" pageNumber="133" phylum="Arthropoda" rank="tribe" tribe="Brachiacanthini">Brachiacanthini</taxonomicName>
taxa examined.
<taxonomicName id="4C510BD5FF4C94C5FBD8FCC4FAE8FCE2" authorityName="Mulsant" authorityYear="1850" box="[1110,1322,771,795]" class="Insecta" family="Coccinellidae" genus="Tiphysa" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="134" pageNumber="133" phylum="Arthropoda" rank="species" species="plumbea">
<emphasis id="B925AC44FF4C94C5FBD8FCC4FAE8FCE2" box="[1110,1322,771,795]" italics="true" pageId="134" pageNumber="133">Tiphysa plumbea</emphasis>
</taxonomicName>
, the
<typeStatus id="54EACEF4FF4C94C5FAE5FCC3FA62FCE5" box="[1387,1440,772,796]" pageId="134" pageNumber="133">type</typeStatus>
species, was described from Cayenne (
<collectingCountry id="F34630C6FF4C94C5FD19FCE4FC94FCC2" box="[663,854,803,827]" name="French Guiana" pageId="134" pageNumber="133">French Guiana</collectingCountry>
), and
<taxonomicName id="4C510BD5FF4C94C5FC2CFCE3FC38FCC2" box="[930,1018,804,827]" class="Insecta" family="Coccinellidae" genus="Tiphysa" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="134" pageNumber="133" phylum="Arthropoda" rank="species" species="egae">
<emphasis id="B925AC44FF4C94C5FC2CFCE3FC77FCC2" box="[930,949,804,827]" italics="true" pageId="134" pageNumber="133">T</emphasis>
.
<emphasis id="B925AC44FF4C94C5FC4BFCE3FC38FCC2" box="[965,1018,804,827]" italics="true" pageId="134" pageNumber="133">egae</emphasis>
</taxonomicName>
from
<collectingCountry id="F34630C6FF4C94C5FBC8FCE4FB52FCC2" box="[1094,1168,803,827]" name="Brazil" pageId="134" pageNumber="133">Brazil</collectingCountry>
(Amazonas).
<bibRefCitation id="EFC00DA7FF4C94C5FAB4FCE4FECCFCA2" author="Mulsant, M. E." pageId="134" pageNumber="133" pagination="1 - 1104" refId="ref106003" refString="Mulsant, M. E. 1850. Species de Coleopteres trimeres securipalpes. Annales des Sciences Physiques et Naturelles, Lyon 2: 1 - 1104." type="journal article" year="1850">Mulsant (1850)</bibRefCitation>
stated that his
<typeStatus id="54EACEF4FF4C94C5FE40FC84FDC4FCA2" box="[462,518,835,859]" pageId="134" pageNumber="133">type</typeStatus>
(s) were in the Dejean collection, now deposited in the MNHL, but no specimens remain there.
</paragraph>
</subSubSection>
</treatment>
</document>