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<document id="66FAA9A4D00284048DF47B5792D3A740" ID-CLB-Dataset="24943" ID-DOI="10.11646/zootaxa.4695.2.5" ID-GBIF-Dataset="7f34a626-f4fc-4484-83cf-ea8a2fdc963c" ID-ISSN="1175-5326" ID-Zenodo-Dep="3531280" ID-ZooBank="CC111070-3A28-4D52-A26F-B808CC200975" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1573110041946" checkinUser="plazi" docAuthor="Sobral, Rafael, De Morais, José W. &amp; Grossi, Paschoal C." docDate="2019" docId="03F74F327A49162167D8FDDEFE3AFE30" docLanguage="en" docName="zootaxa.4695.2.5.pdf" docOrigin="Zootaxa 4695 (2)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Colacus morio Ohaus 1910" docType="treatment" docVersion="8" lastPageNumber="167" masterDocId="FFCE374A7A4F1629674FFFDFFF8EFF9F" masterDocTitle="A new species of Colacus Ohaus, 1910 (Coleoptera: Scarabaeidae: Dynastinae) from the Mata Seca biotope of Brazil, and notes on Colacus morio Ohaus, 1910" masterLastPageNumber="167" masterPageNumber="159" pageNumber="165" updateTime="1698763786448" updateUser="ExternalLinkService">
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<mods:title id="635E684018DED6E290B480F8C7E40F1A">A new species of Colacus Ohaus, 1910 (Coleoptera: Scarabaeidae: Dynastinae) from the Mata Seca biotope of Brazil, and notes on Colacus morio Ohaus, 1910</mods:title>
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<mods:namePart id="D1043284250F0FC94737BA8F527E69F9">Sobral, Rafael</mods:namePart>
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<mods:namePart id="ACB84750FB4DB6175469D7FDF21061C8">De Morais, José W.</mods:namePart>
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<treatment id="03F74F327A49162167D8FDDEFE3AFE30" ID-DOI="http://doi.org/10.5281/zenodo.5922986" ID-GBIF-Taxon="160164835" ID-Zenodo-Dep="5922986" LSID="urn:lsid:plazi:treatment:03F74F327A49162167D8FDDEFE3AFE30" httpUri="http://treatment.plazi.org/id/03F74F327A49162167D8FDDEFE3AFE30" lastPageId="8" lastPageNumber="167" pageId="6" pageNumber="165">
<subSubSection id="C344ADAF7A49162F67D8FDDEFE65FD84" box="[151,491,513,540]" pageId="6" pageNumber="165" type="nomenclature">
<paragraph id="8BE1FE247A49162F67D8FDDEFE65FD84" blockId="6.[151,491,513,575]" box="[151,491,513,540]" pageId="6" pageNumber="165">
<heading id="D0A949487A49162F67D8FDDEFE65FD84" bold="true" box="[151,491,513,540]" fontSize="11" level="1" pageId="6" pageNumber="165" reason="1">
<taxonomicName id="4C5E85A77A49162F67D8FDDEFE65FD84" authority="Ohaus, 1910" authorityName="Ohaus" authorityYear="1910" box="[151,491,513,540]" class="Insecta" family="Dynastidae" genus="Colacus" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="165" phylum="Arthropoda" rank="species" species="morio">
<emphasis id="B92A22367A49162F67D8FDDEFE65FD84" bold="true" box="[151,491,513,540]" pageId="6" pageNumber="165">
<emphasis id="B92A22367A49162F67D8FDDEFEC6FD84" bold="true" box="[151,328,513,540]" italics="true" pageId="6" pageNumber="165">Colacus morio</emphasis>
Ohaus, 1910
</emphasis>
</taxonomicName>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C344ADAF7A49162E67D8FDFAFB8FF833" lastPageId="7" lastPageNumber="166" pageId="6" pageNumber="165" type="description">
<paragraph id="8BE1FE247A49162F67D8FDFAFE9CFDA0" blockId="6.[151,491,513,575]" box="[151,274,548,575]" pageId="6" pageNumber="165">
(
<figureCitation id="1365E2A17A49162F67D0FDFAFE84FDA0" box="[159,266,548,575]" captionStart-0="FIGURE 5" captionStart-1="FIGURE 6" captionStart-2="FIGURE 7" captionStartId-0="5.[151,250,1868,1893]" captionStartId-1="6.[151,250,1987,2012]" captionStartId-2="7.[151,250,1759,1784]" captionTargetBox-0="[151,1435,171,1846]" captionTargetBox-1="[157,1424,812,1955]" captionTargetBox-2="[151,1435,526,1739]" captionTargetId-0="figure@5.[151,1435,171,1846]" captionTargetId-1="figure@6.[151,1435,803,1966]" captionTargetId-2="figure@7.[151,1435,517,1739]" captionTargetPageId-0="5" captionTargetPageId-1="6" captionTargetPageId-2="7" captionText-0="FIGURE 5. A, Known distribution map of Colacus species; B, Mata Seca biotope." captionText-1="FIGURE 6. Female of Colacus morio. A, Habitus in dorsal view; B, habitus in ventral view. Scale bars: 3 mm." captionText-2="FIGURE 7. Female of Colacus morio. A, Head and anterior portion of pronotum in dorsal view; B, head in ventral view, black arrow indicating outer side of right mandible, white arrows indicating apical projections of mentum; C, canthus in dorsal view; D, head and pronotum in lateral view, black arrows indicating cephalic tubercle and pronotal tubercle. Scale bars: AC, 1 mm; D, 3 mm." figureDoi-0="http://doi.org/10.5281/zenodo.3531290" figureDoi-1="http://doi.org/10.5281/zenodo.3531292" figureDoi-2="http://doi.org/10.5281/zenodo.3531294" httpUri-0="https://zenodo.org/record/3531290/files/figure.png" httpUri-1="https://zenodo.org/record/3531292/files/figure.png" httpUri-2="https://zenodo.org/record/3531294/files/figure.png" pageId="6" pageNumber="165">Figs. 57</figureCitation>
)
</paragraph>
<paragraph id="8BE1FE247A49162F67D8FDB2FA1FFC88" blockId="6.[151,1437,620,791]" pageId="6" pageNumber="165">
<emphasis id="B92A22367A49162F67D8FDB2FE0FFD18" bold="true" box="[151,385,620,647]" pageId="6" pageNumber="165">Female description.</emphasis>
Equal to males in general aspect (
<figureCitation id="1365E2A17A49162F65B8FDB2FCCDFD17" box="[759,835,621,648]" captionStart="FIGURE 6" captionStartId="6.[151,250,1987,2012]" captionTargetBox="[157,1424,812,1955]" captionTargetId="figure@6.[151,1435,803,1966]" captionTargetPageId="6" captionText="FIGURE 6. Female of Colacus morio. A, Habitus in dorsal view; B, habitus in ventral view. Scale bars: 3 mm." figureDoi="http://doi.org/10.5281/zenodo.3531292" httpUri="https://zenodo.org/record/3531292/files/figure.png" pageId="6" pageNumber="165">Figs. 6</figureCitation>
AB).
<emphasis id="B92A22367A49162F64C1FDB2FC59FD18" bold="true" box="[910,983,621,647]" pageId="6" pageNumber="165">Head:</emphasis>
Cephalic tubercle short (
<figureCitation id="1365E2A17A49162F63A4FDB2FACAFD18" box="[1259,1348,621,647]" captionStart="FIGURE 7" captionStartId="7.[151,250,1759,1784]" captionTargetBox="[151,1435,526,1739]" captionTargetId="figure@7.[151,1435,517,1739]" captionTargetPageId="7" captionText="FIGURE 7. Female of Colacus morio. A, Head and anterior portion of pronotum in dorsal view; B, head in ventral view, black arrow indicating outer side of right mandible, white arrows indicating apical projections of mentum; C, canthus in dorsal view; D, head and pronotum in lateral view, black arrows indicating cephalic tubercle and pronotal tubercle. Scale bars: AC, 1 mm; D, 3 mm." figureDoi="http://doi.org/10.5281/zenodo.3531294" httpUri="https://zenodo.org/record/3531294/files/figure.png" pageId="6" pageNumber="165">Fig. 7D</figureCitation>
), lateral carina of tubercle inconspicuous. Punctures coalescent and dense on vertex. Clypeus slightly shorter and broader than in male; wrinkles dense on surface of clypeus (
<figureCitation id="1365E2A17A49162F659DFD6AFCA5FD50" box="[722,811,693,719]" captionStart="FIGURE 7" captionStartId="7.[151,250,1759,1784]" captionTargetBox="[151,1435,526,1739]" captionTargetId="figure@7.[151,1435,517,1739]" captionTargetPageId="7" captionText="FIGURE 7. Female of Colacus morio. A, Head and anterior portion of pronotum in dorsal view; B, head in ventral view, black arrow indicating outer side of right mandible, white arrows indicating apical projections of mentum; C, canthus in dorsal view; D, head and pronotum in lateral view, black arrows indicating cephalic tubercle and pronotal tubercle. Scale bars: AC, 1 mm; D, 3 mm." figureDoi="http://doi.org/10.5281/zenodo.3531294" httpUri="https://zenodo.org/record/3531294/files/figure.png" pageId="6" pageNumber="165">Fig. 7A</figureCitation>
). Ocular canthus slightly convex on outer side (
<figureCitation id="1365E2A17A49162F6277FD6AFA01FD50" box="[1336,1423,693,719]" captionStart="FIGURE 7" captionStartId="7.[151,250,1759,1784]" captionTargetBox="[151,1435,526,1739]" captionTargetId="figure@7.[151,1435,517,1739]" captionTargetPageId="7" captionText="FIGURE 7. Female of Colacus morio. A, Head and anterior portion of pronotum in dorsal view; B, head in ventral view, black arrow indicating outer side of right mandible, white arrows indicating apical projections of mentum; C, canthus in dorsal view; D, head and pronotum in lateral view, black arrows indicating cephalic tubercle and pronotal tubercle. Scale bars: AC, 1 mm; D, 3 mm." figureDoi="http://doi.org/10.5281/zenodo.3531294" httpUri="https://zenodo.org/record/3531294/files/figure.png" pageId="6" pageNumber="165">Fig. 7C</figureCitation>
). Antennae shorter than in male, lamellae slender than lamellae of male and shorter than the other antennomeres combined.
<emphasis id="B92A22367A49162F67ACFD22FEF2FC88" bold="true" box="[227,380,765,791]" pageId="6" pageNumber="165">Mouthparts:</emphasis>
Mandibles slightly more slender, apically with a slight concavity on outer side (
<figureCitation id="1365E2A17A49162F63B1FD22FAC5FC88" box="[1278,1355,765,791]" captionStart="FIGURE 7" captionStartId="7.[151,250,1759,1784]" captionTargetBox="[151,1435,526,1739]" captionTargetId="figure@7.[151,1435,517,1739]" captionTargetPageId="7" captionText="FIGURE 7. Female of Colacus morio. A, Head and anterior portion of pronotum in dorsal view; B, head in ventral view, black arrow indicating outer side of right mandible, white arrows indicating apical projections of mentum; C, canthus in dorsal view; D, head and pronotum in lateral view, black arrows indicating cephalic tubercle and pronotal tubercle. Scale bars: AC, 1 mm; D, 3 mm." figureDoi="http://doi.org/10.5281/zenodo.3531294" httpUri="https://zenodo.org/record/3531294/files/figure.png" pageId="6" pageNumber="165">Figs. 7</figureCitation>
AB).
</paragraph>
<caption id="DF21AEAC7A49162F67D8F81CFA8EF843" ID-DOI="http://doi.org/10.5281/zenodo.3531292" ID-Zenodo-Dep="3531292" box="[151,1280,1987,2012]" httpUri="https://zenodo.org/record/3531292/files/figure.png" pageId="6" pageNumber="165" startId="6.[151,250,1987,2012]" targetBox="[157,1424,812,1955]" targetPageId="6">
<paragraph id="8BE1FE247A49162F67D8F81CFA8EF843" blockId="6.[151,1280,1987,2012]" box="[151,1280,1987,2012]" pageId="6" pageNumber="165">
<emphasis id="B92A22367A49162F67D8F81CFE9DF843" bold="true" box="[151,275,1987,2012]" pageId="6" pageNumber="165">FIGURE 6.</emphasis>
Female of
<taxonomicName id="4C5E85A77A49162F66C9F81BFD92F843" authorityName="Ohaus" authorityYear="1910" box="[390,540,1988,2012]" class="Insecta" family="Dynastidae" genus="Colacus" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="165" phylum="Arthropoda" rank="species" species="morio">
<emphasis id="B92A22367A49162F66C9F81BFD92F843" box="[390,540,1988,2012]" italics="true" pageId="6" pageNumber="165">Colacus morio</emphasis>
</taxonomicName>
.
<emphasis id="B92A22367A49162F6567F81CFDB4F843" bold="true" box="[552,570,1987,2012]" pageId="6" pageNumber="165">A</emphasis>
, Habitus in dorsal view;
<emphasis id="B92A22367A49162F6476F81BFCC4F843" bold="true" box="[825,842,1988,2012]" pageId="6" pageNumber="165">B</emphasis>
, habitus in ventral view. Scale bars: 3 mm.
</paragraph>
</caption>
<paragraph id="8BE1FE247A48162E67D8FF46FC0AFE30" blockId="7.[151,1437,153,503]" pageId="7" pageNumber="166">
Mentum with 2 apical projections in females (
<figureCitation id="1365E2A17A48162E65DBFF46FD64FF2C" box="[660,746,153,179]" captionStart="FIGURE 7" captionStartId="7.[151,250,1759,1784]" captionTargetBox="[151,1435,526,1739]" captionTargetId="figure@7.[151,1435,517,1739]" captionTargetPageId="7" captionText="FIGURE 7. Female of Colacus morio. A, Head and anterior portion of pronotum in dorsal view; B, head in ventral view, black arrow indicating outer side of right mandible, white arrows indicating apical projections of mentum; C, canthus in dorsal view; D, head and pronotum in lateral view, black arrows indicating cephalic tubercle and pronotal tubercle. Scale bars: AC, 1 mm; D, 3 mm." figureDoi="http://doi.org/10.5281/zenodo.3531294" httpUri="https://zenodo.org/record/3531294/files/figure.png" pageId="7" pageNumber="166">Fig. 7B</figureCitation>
), absent in males.
<emphasis id="B92A22367A48162E64F9FF46FB96FF2C" bold="true" box="[950,1048,153,179]" pageId="7" pageNumber="166">Thorax:</emphasis>
Pronotum more convex than male, with a slight anterior protrusion between pronotal tubercles; tubercles nearly obsolete (
<figureCitation id="1365E2A17A48162E6312FF62FB39FF48" box="[1117,1207,189,215]" captionStart="FIGURE 7" captionStartId="7.[151,250,1759,1784]" captionTargetBox="[151,1435,526,1739]" captionTargetId="figure@7.[151,1435,517,1739]" captionTargetPageId="7" captionText="FIGURE 7. Female of Colacus morio. A, Head and anterior portion of pronotum in dorsal view; B, head in ventral view, black arrow indicating outer side of right mandible, white arrows indicating apical projections of mentum; C, canthus in dorsal view; D, head and pronotum in lateral view, black arrows indicating cephalic tubercle and pronotal tubercle. Scale bars: AC, 1 mm; D, 3 mm." figureDoi="http://doi.org/10.5281/zenodo.3531294" httpUri="https://zenodo.org/record/3531294/files/figure.png" pageId="7" pageNumber="166">Fig. 7D</figureCitation>
). Pronotal punctures more coalescent anteriorly and medially, wrinkled near anterior angles. Scutellar surface more punctate basally than medially, punctures on base smaller than on disc, discal punctures setose.
<emphasis id="B92A22367A48162E6498FEDAFBA3FE80" bold="true" box="[983,1069,261,287]" pageId="7" pageNumber="166">Elytra:</emphasis>
Shape more round than in male. Disc with deep, annular punctures mixed with coalescent punctures. Surface more rugose, and punctures more coalescent near scutellum than on disc.
<emphasis id="B92A22367A48162E657DFE92FDFDFEF8" bold="true" box="[562,627,333,359]" pageId="7" pageNumber="166">Legs:</emphasis>
Protibiae tridentate with trace of a fourth basal tooth as in males.
<emphasis id="B92A22367A48162E6207FE93FF5BFE14" bold="true" pageId="7" pageNumber="166">Tergite VIII:</emphasis>
Disc with small punctures more scattered than in males, posterior angles with punctures smaller than on disc, wrinkles present. Long bristles more dense on sides, disc glabrous.
</paragraph>
<paragraph id="8BE1FE247A48162E6788FE66FD7EFE68" blockId="7.[151,1437,153,503]" pageId="7" pageNumber="166">
<emphasis id="B92A22367A48162E6788FE66FEF0FE4C" bold="true" box="[199,382,441,467]" pageId="7" pageNumber="166">Measurements.</emphasis>
Body length:
<quantity id="4CA653C17A48162E656DFE66FD05FE4C" box="[546,651,441,467]" metricMagnitude="-2" metricUnit="m" metricValue="2.69" pageId="7" pageNumber="166" unit="mm" value="26.9">26.9 mm</quantity>
. Head width:
<quantity id="4CA653C17A48162E6463FE67FC09FE4C" box="[812,903,440,467]" metricMagnitude="-3" metricUnit="m" metricValue="5.2" pageId="7" pageNumber="166" unit="mm" value="5.2">5.2 mm</quantity>
. Pronotal width:
<quantity id="4CA653C17A48162E6304FE66FB3DFE4C" box="[1099,1203,441,467]" metricMagnitude="-2" metricUnit="m" metricValue="1.1099999999999999" pageId="7" pageNumber="166" unit="mm" value="11.1">11.1 mm</quantity>
. Elytral length:
<quantity id="4CA653C17A48162E6224FE66FF48FE68" metricMagnitude="-2" metricUnit="m" metricValue="1.72" pageId="7" pageNumber="166" unit="mm" value="17.2">17.2 mm</quantity>
. Elytral width:
<quantity id="4CA653C17A48162E663CFE02FE42FE68" box="[371,460,477,503]" metricMagnitude="-3" metricUnit="m" metricValue="9.4" pageId="7" pageNumber="166" unit="mm" value="9.4">9.4 mm</quantity>
. Protibial length:
<quantity id="4CA653C17A48162E65DDFE03FD65FE68" box="[658,747,476,503]" metricMagnitude="-3" metricUnit="m" metricValue="5.2" pageId="7" pageNumber="166" unit="mm" value="5.2">5.2 mm</quantity>
.
</paragraph>
<caption id="DF21AEAC7A48162E67D8F900FF76F8FC" ID-DOI="http://doi.org/10.5281/zenodo.3531294" ID-Zenodo-Dep="3531294" httpUri="https://zenodo.org/record/3531294/files/figure.png" pageId="7" pageNumber="166" startId="7.[151,250,1759,1784]" targetBox="[151,1435,526,1739]" targetPageId="7">
<paragraph id="8BE1FE247A48162E67D8F900FF76F8FC" blockId="7.[151,1436,1759,1892]" pageId="7" pageNumber="166">
<emphasis id="B92A22367A48162E67D8F900FE9DF967" bold="true" box="[151,275,1759,1784]" pageId="7" pageNumber="166">FIGURE 7.</emphasis>
Female of
<taxonomicName id="4C5E85A77A48162E66CAF93FFD95F968" authorityName="Ohaus" authorityYear="1910" box="[389,539,1760,1784]" class="Insecta" family="Dynastidae" genus="Colacus" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="166" phylum="Arthropoda" rank="species" species="morio">
<emphasis id="B92A22367A48162E66CAF93FFD95F968" box="[389,539,1760,1784]" italics="true" pageId="7" pageNumber="166">Colacus morio</emphasis>
</taxonomicName>
.
<emphasis id="B92A22367A48162E6568F900FDB7F967" bold="true" box="[551,569,1759,1784]" pageId="7" pageNumber="166">A</emphasis>
, Head and anterior portion of pronotum in dorsal view;
<emphasis id="B92A22367A48162E633FF93FFB0FF967" bold="true" box="[1136,1153,1760,1784]" pageId="7" pageNumber="166">B</emphasis>
, head in ventral view, black arrow indicating outer side of right mandible, white arrows indicating apical projections of mentum;
<emphasis id="B92A22367A48162E63DBF8DCFB28F883" bold="true" box="[1172,1190,1795,1820]" pageId="7" pageNumber="166">C</emphasis>
, canthus in dorsal view;
<emphasis id="B92A22367A48162E67D8F8F7FF27F8DF" bold="true" box="[151,169,1832,1856]" pageId="7" pageNumber="166">D</emphasis>
, head and pronotum in lateral view, black arrows indicating cephalic tubercle and pronotal tubercle. Scale bars:
<emphasis id="B92A22367A48162E6252F8F8FAC0F8DF" bold="true" box="[1309,1358,1831,1856]" pageId="7" pageNumber="166">AC</emphasis>
, 1 mm;
<emphasis id="B92A22367A48162E67D8F893FF27F8FB" bold="true" box="[151,169,1868,1892]" pageId="7" pageNumber="166">D</emphasis>
, 3 mm.
</paragraph>
</caption>
<paragraph id="8BE1FE247A48162E6788F84EFB8FF833" blockId="7.[151,1436,1937,2037]" box="[199,1025,1937,1964]" pageId="7" pageNumber="166">
<emphasis id="B92A22367A48162E6788F84EFE61F833" bold="true" box="[199,495,1937,1964]" pageId="7" pageNumber="166">Geographic distribution.</emphasis>
<materialsCitation id="3B36F4797A48162E66B9F84DFC73F833" ID-GBIF-Occurrence="2447668904" box="[502,1021,1937,1964]" country="Argentina" location="Catamarca" pageId="7" pageNumber="166" specimenCount="1" stateProvince="Catamarca">
<collectingCountry id="F349BEB47A48162E66B9F84DFDE4F833" box="[502,618,1938,1964]" name="Argentina" pageId="7" pageNumber="166">Argentina</collectingCountry>
:
<collectingRegion id="499A30C67A48162E653BF84DFD62F833" box="[628,748,1938,1964]" country="Argentina" name="Catamarca" pageId="7" pageNumber="166">Catamarca</collectingRegion>
and
<collectingRegion id="499A30C67A48162E646DF84DFC05F833" box="[802,907,1938,1964]" country="Argentina" name="Tucuman" pageId="7" pageNumber="166">Tucumán</collectingRegion>
(
<figureCitation id="1365E2A17A48162E64D5F84DFC7AF833" box="[922,1012,1937,1964]" captionStart="FIGURE 5" captionStartId="5.[151,250,1868,1893]" captionTargetBox="[151,1435,171,1846]" captionTargetId="figure@5.[151,1435,171,1846]" captionTargetPageId="5" captionText="FIGURE 5. A, Known distribution map of Colacus species; B, Mata Seca biotope." figureDoi="http://doi.org/10.5281/zenodo.3531290" httpUri="https://zenodo.org/record/3531290/files/figure.png" pageId="7" pageNumber="166">Fig. 5A</figureCitation>
)
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="C344ADAF7A4816216788F86AFC25FF64" lastPageId="8" lastPageNumber="167" pageId="7" pageNumber="166" type="materials_examined">
<paragraph id="8BE1FE247A4816216788F86AFC25FF64" blockId="7.[151,1436,1937,2037]" lastBlockId="8.[151,1436,153,431]" lastPageId="8" lastPageNumber="167" pageId="7" pageNumber="166">
<emphasis id="B92A22367A48162E6788F86AFE20F84F" bold="true" box="[199,430,1973,2000]" pageId="7" pageNumber="166">Material examined.</emphasis>
One female and one male at
<collectionCode id="ED4F66E17A48162E65AAF869FCB2F84F" box="[741,828,1974,2000]" pageId="7" pageNumber="166">CERPE</collectionCode>
with the following data: a) “
<collectingCountry id="F349BEB47A48162E6321F869FA99F84F" box="[1134,1303,1974,2000]" name="Argentina" pageId="7" pageNumber="166">ARGENTINA</collectingCountry>
, /
<collectingRegion id="499A30C67A48162E6263F869FA16F84F" box="[1324,1432,1974,2000]" country="Argentina" name="Tucuman" pageId="7" pageNumber="166">Tucumán</collectingRegion>
, Infiernillo, / 3048 m—
<date id="FFE0D8E47A48162E66D5F805FE70F86B" box="[410,510,2010,2036]" pageId="7" pageNumber="166" value="2008-01-10">10.i.2008</date>
/ E. Abadie Leg.”; one male at
<collectionCode id="ED4F66E17A48162E6413F804FC10F86A" box="[860,926,2011,2037]" pageId="7" pageNumber="166">IFML</collectionCode>
with the following data: a) “Cumbre Infierniello /
<date id="FFE0D8E47A4716216798FF46FEA6FF2C" box="[215,296,153,179]" pageId="8" pageNumber="167" value="2018-11-03">3.xi.18</date>
”; two males at
<collectionCode id="ED4F66E17A471621669AFF46FD99FF2C" box="[469,535,153,179]" pageId="8" pageNumber="167">IFML</collectionCode>
with the following data: a) “Valle S. Maria /
<collectingRegion id="499A30C67A471621635DFF46FB06FF2C" box="[1042,1160,153,179]" country="Argentina" name="Entre Rios" pageId="8" pageNumber="167">Entre Rios</collectingRegion>
/
<date id="FFE0D8E47A47162163D0FF46FB71FF2C" box="[1183,1279,153,179]" pageId="8" pageNumber="167" value="1933-12">xii.1933</date>
”, b) “
<emphasis id="B92A22367A471621620FFF46FF74FF48" italics="true" pageId="8" pageNumber="167">
<taxonomicName id="4C5E85A77A471621620FFF46FA12FF2C" authorityName="Ohaus" authorityYear="1910" box="[1344,1436,153,179]" class="Insecta" family="Dynastidae" genus="Colacus" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="167" phylum="Arthropoda" rank="genus">Colacus</taxonomicName>
/
<taxonomicName id="4C5E85A77A47162167E8FF61FF64FF48" authorityName="Ohaus" authorityYear="1910" box="[167,234,190,215]" class="Insecta" family="Dynastidae" genus="Colacus" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="167" phylum="Arthropoda" rank="species" species="morio">morio</taxonomicName>
/
</emphasis>
Ohaus / J. M. Bosq det. 1946”; two males at
<collectionCode id="ED4F66E17A471621644DFF62FCCAFF48" box="[770,836,189,215]" pageId="8" pageNumber="167">IFML</collectionCode>
with the following data: a) “Qda. Carapunco / (Tafi)
<quantity id="4CA653C17A47162167D8FF3FFF62FF64" box="[151,236,224,251]" metricMagnitude="3" metricUnit="m" metricValue="2.5" pageId="8" pageNumber="167" unit="m" value="2500.0">2500 m</quantity>
/
<date id="FFE0D8E47A471621664DFF3EFED4FF64" box="[258,346,225,251]" pageId="8" pageNumber="167" value="1932-11">xi.1932</date>
”, b) “
<emphasis id="B92A22367A47162166D3FF3EFDDFFF64" box="[412,593,225,251]" italics="true" pageId="8" pageNumber="167">
<taxonomicName id="4C5E85A77A47162166D3FF3EFE7DFF64" authorityName="Ohaus" authorityYear="1910" box="[412,499,225,251]" class="Insecta" family="Dynastidae" genus="Colacus" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="167" phylum="Arthropoda" rank="genus">Colacus</taxonomicName>
/
<taxonomicName id="4C5E85A77A4716216549FF3EFDCAFF64" authorityName="Ohaus" authorityYear="1910" box="[518,580,225,251]" class="Insecta" family="Dynastidae" genus="Colacus" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="167" phylum="Arthropoda" rank="species" species="morio">morio</taxonomicName>
/
</emphasis>
Ohaus/ J. M. Bosq det. 1946”.
</paragraph>
</subSubSection>
<subSubSection id="C344ADAF7A4716216788FEDAFE3AFE30" pageId="8" pageNumber="167" type="discussion">
<paragraph id="8BE1FE247A4716216788FEDAFE3AFE30" blockId="8.[151,1436,153,431]" pageId="8" pageNumber="167">
<emphasis id="B92A22367A4716216788FEDAFEB5FE80" bold="true" box="[199,315,261,287]" pageId="8" pageNumber="167">Remarks.</emphasis>
In his review of
<taxonomicName id="4C5E85A77A47162166B2FEDAFCD7FE80" authority=", Neita-Moreno (2015)" authorityName="Neita-Moreno" authorityYear="2015" box="[509,857,260,287]" class="Insecta" family="Dynastidae" genus="Colacus" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="167" phylum="Arthropoda" rank="genus">
<emphasis id="B92A22367A47162166B2FEDAFDD7FE80" box="[509,601,261,287]" italics="true" pageId="8" pageNumber="167">Colacus</emphasis>
,
<bibRefCitation id="EFCF83D57A4716216527FEDAFCD7FE80" author="Neita-Moreno, J. C." box="[616,857,260,287]" pageId="8" pageNumber="167" pagination="187 - 199" refId="ref3531" refString="Neita-Moreno, J. C. (2015) Revision del genero Colacus Ohaus (Coleoptera: Scarabaeidae: Dynastinae: Agaocephalini). Dugesiana, 22, 187 - 199." type="journal article" year="2015">Neita-Moreno (2015)</bibRefCitation>
</taxonomicName>
synonymized
<taxonomicName id="4C5E85A77A47162164B1FEDAFB78FE80" authority="Martinez" authorityName="Martinez" authorityYear="1988" box="[1022,1270,261,287]" class="Insecta" family="Dynastidae" genus="Colacus" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="167" phylum="Arthropoda" rank="species" species="endroedii">
<emphasis id="B92A22367A47162164B1FEDAFB07FE80" box="[1022,1161,261,287]" italics="true" pageId="8" pageNumber="167">C. endroedii</emphasis>
Martínez
</taxonomicName>
with
<taxonomicName id="4C5E85A77A4716216277FEDAFF6FFEDC" authority="Ohaus" authorityName="Ohaus" authorityYear="1910" class="Insecta" family="Dynastidae" genus="Colacus" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="167" phylum="Arthropoda" rank="species" species="morio">
<emphasis id="B92A22367A4716216277FEDAFA12FE80" box="[1336,1436,261,287]" italics="true" pageId="8" pageNumber="167">C. morio</emphasis>
Ohaus
</taxonomicName>
, redescribing the male but apparently not considering the description of the female made by
<bibRefCitation id="EFCF83D57A4716216246FEF6FF6BFEF8" author="Abadie, E. I. &amp; Grossi, P. C. &amp; Wagner, P. S." pageId="8" pageNumber="167" refId="ref3426" refString="Abadie, E. I., Grossi, P. C. &amp; Wagner, P. S. (2008) A field guide of the Dynastidae family of the south of South America. Design &amp; Digital Edition Javier Canete, Buenos Aires, 120 pp." type="book" year="2008">
Abadie
<emphasis id="B92A22367A471621622DFEF5FA1BFEDC" box="[1378,1429,297,323]" italics="true" pageId="8" pageNumber="167">et al</emphasis>
. (2008)
</bibRefCitation>
. In that work,
<bibRefCitation id="EFCF83D57A47162166C6FE92FDE0FEF8" author="Abadie, E. I. &amp; Grossi, P. C. &amp; Wagner, P. S." box="[393,622,333,359]" pageId="8" pageNumber="167" refId="ref3426" refString="Abadie, E. I., Grossi, P. C. &amp; Wagner, P. S. (2008) A field guide of the Dynastidae family of the south of South America. Design &amp; Digital Edition Javier Canete, Buenos Aires, 120 pp." type="book" year="2008">
Abadie
<emphasis id="B92A22367A47162166ADFE91FD9BFEF8" box="[482,533,333,359]" italics="true" pageId="8" pageNumber="167">et al</emphasis>
. (2008)
</bibRefCitation>
opted to avoid large and repetitive descriptions and emphasized mainly six traits that separate males from females. Here we expand on the basic characters used by
<bibRefCitation id="EFCF83D57A47162163D2FEAEFAF1FE14" author="Abadie, E. I. &amp; Grossi, P. C. &amp; Wagner, P. S." box="[1181,1407,369,395]" pageId="8" pageNumber="167" refId="ref3426" refString="Abadie, E. I., Grossi, P. C. &amp; Wagner, P. S. (2008) A field guide of the Dynastidae family of the south of South America. Design &amp; Digital Edition Javier Canete, Buenos Aires, 120 pp." type="book" year="2008">
Abadie
<emphasis id="B92A22367A47162163BAFEADFAA9FE14" box="[1269,1319,369,395]" italics="true" pageId="8" pageNumber="167">et al</emphasis>
. (2008)
</bibRefCitation>
to fully describe the female.
</paragraph>
</subSubSection>
</treatment>
</document>