treatments-xml/data/03/E5/E9/03E5E958FFDB917462CFFF18FE5EFEAB.xml

<document id="7269E21A11533116BAC62C7DE766CE75" ID-CLB-Dataset="27548" ID-DOI="10.11646/zootaxa.4555.2.3" ID-GBIF-Dataset="2373752d-6333-4f9d-9476-6f2e163a67da" ID-ISSN="1175-5326" ID-Zenodo-Dep="2624293" ID-ZooBank="A7CF86E1-C763-4082-B1C2-9B8B66428142" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1554272273913" checkinUser="plazi" docAuthor="Magalhães, Wagner F., Rizzo, Alexandra E. &amp; Bailey-Brock, Julie H." docDate="2019" docId="03E5E958FFDB917462CFFF18FE5EFEAB" docLanguage="en" docName="zootaxa.4555.2.3.pdf" docOrigin="Zootaxa 4555 (2)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Polyophthalmus mauliola Magalhães &amp; Rizzo &amp; Bailey-Brock 2019, sp. nov." docType="treatment" docVersion="13" lastPageNumber="225" masterDocId="FFDC9120FFD791646258FF8FFFD1FFCE" masterDocTitle="Opheliidae (Annelida: Polychaeta) from the western Pacific islands, including five new species" masterLastPageNumber="235" masterPageNumber="209" pageNumber="221" updateTime="1701997935202" updateUser="plazi">
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<mods:title id="4F825F309143783725B0070B5609C156">Opheliidae (Annelida: Polychaeta) from the western Pacific islands, including five new species</mods:title>
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<mods:namePart id="59D1C9C7599A6761FE2718E70A6D1083">Magalhães, Wagner F.</mods:namePart>
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<mods:namePart id="AA7A09F1099A9F3D6FC81EF158F93995">Rizzo, Alexandra E.</mods:namePart>
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<mods:namePart id="8045B03099143C2C92EE5D8AB799AE3D">Bailey-Brock, Julie H.</mods:namePart>
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<treatment id="03E5E958FFDB917462CFFF18FE5EFEAB" ID-DOI="http://doi.org/10.5281/zenodo.5282927" ID-GBIF-Taxon="154533566" ID-Zenodo-Dep="5282927" LSID="urn:lsid:plazi:treatment:03E5E958FFDB917462CFFF18FE5EFEAB" httpUri="http://treatment.plazi.org/id/03E5E958FFDB917462CFFF18FE5EFEAB" lastPageId="16" lastPageNumber="225" pageId="12" pageNumber="221">
<subSubSection id="C3560BC5FFDB916862CFFF18FDE5FF7C" box="[151,564,151,178]" pageId="12" pageNumber="221" type="nomenclature">
<paragraph id="8BF3584EFFDB916862CFFF18FDE5FF7C" blockId="12.[151,564,151,211]" box="[151,564,151,178]" pageId="12" pageNumber="221">
<heading id="D0BBEF22FFDB916862CFFF18FDE5FF7C" bold="true" box="[151,564,151,178]" fontSize="11" level="1" pageId="12" pageNumber="221" reason="1">
<emphasis id="B938845CFFDB916862CFFF18FDE5FF7C" bold="true" box="[151,564,151,178]" pageId="12" pageNumber="221">
<taxonomicName id="4C4C23CDFFDB916862CFFF18FE1EFF7F" ID-CoL="4LCTB" authority="Magalhães &amp; Rizzo &amp; Bailey-Brock, 2019" authorityName="Magalhães &amp; Rizzo &amp; Bailey-Brock" authorityYear="2019" box="[151,463,151,177]" class="Polychaeta" family="Opheliidae" genus="Polyophthalmus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Opheliida" pageId="12" pageNumber="221" phylum="Annelida" rank="species" species="mauliola" status="sp. nov.">
<emphasis id="B938845CFFDB916862CFFF18FE1EFF7F" bold="true" box="[151,463,151,177]" italics="true" pageId="12" pageNumber="221">Polyophthalmus mauliola</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A20B3927FFDB9168638EFF17FDE5FF7C" box="[470,564,152,178]" pageId="12" pageNumber="221" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C3560BC5FFDB916862CFFF35FE06FF1D" box="[151,471,186,211]" pageId="12" pageNumber="221" type="description">
<paragraph id="8BF3584EFFDB916862CFFF35FE06FF1D" blockId="12.[151,564,151,211]" box="[151,471,186,211]" pageId="12" pageNumber="221">
<figureCitation id="137744CBFFDB916862CFFF35FF2FFF1D" box="[151,254,186,211]" captionStart="FIGURE 8" captionStartId="15.[151,250,1887,1909]" captionTargetBox="[253,1294,443,1847]" captionTargetId="figure@15.[253,1333,419,1859]" captionTargetPageId="15" captionText="FIGURE 8. Polyophthalmus mauliola sp. nov. from Mamala Bay, Hawaii: A, complete specimen in dorsal view; B, anterior end in lateral view; C, anterior end in dorsal view; D, parapodium of the posterior end; E, posterior end with pygidium in lateral view; F, posterior end with pygidium in dorsal view." httpUri="https://zenodo.org/record/2624309/files/figure.png" pageId="12" pageNumber="221">Figures 8</figureCitation>
(A–F) and 9 (A–E)
</paragraph>
</subSubSection>
<subSubSection id="C3560BC5FFDB916862CFFF72FEDAFDDF" pageId="12" pageNumber="221" type="materials_examined">
<paragraph id="8BF3584EFFDB916862CFFF72FEDAFDDF" blockId="12.[151,1437,253,2033]" pageId="12" pageNumber="221">
<emphasis id="B938845CFFDB916862CFFF72FE57FED8" bold="true" box="[151,390,253,278]" pageId="12" pageNumber="221">Material examined.</emphasis>
<materialsCitation id="3B245213FFDB916863CAFF72FC8FFEF7" ID-GBIF-Occurrence="2234219933" collectionCode="BPBM-R" collectorName="Sta. D" elevation="50" latitude="21.282055" location="Mamala Bay" longLatPrecision="1" longitude="-157.897" municipality="Sand Island" pageId="12" pageNumber="221" specimenCode="BPBM-R3887" specimenCount="1" typeStatus="holotype">
<typeStatus id="54F7E6ECFFDB916863CAFF72FE2BFED8" box="[402,506,253,278]" pageId="12" pageNumber="221" type="holotype">Holotype</typeStatus>
: 
<location id="8E930E95FFDB91686057FF72FD74FED8" LSID="urn:lsid:plazi:treatment:03E5E958FFDB917462CFFF18FE5EFEAB:8E930E95FFDB91686057FF72FD74FED8" box="[527,677,253,278]" latitude="21.282055" longLatPrecision="1" longitude="-157.897" municipality="Sand Island" name="Mamala Bay" pageId="12" pageNumber="221">Mamala Bay</location>
, Oahu, Hawaii: vicinities of 
<collectingMunicipality id="6B97C234FFDB9168665BFF72FB5DFED8" box="[1027,1164,253,278]" pageId="12" pageNumber="221">Sand Island</collectingMunicipality>
outfall, Oct/2015, 
<collectorName id="26B93D98FFDB9168672AFF71FF7DFEF7" pageId="12" pageNumber="221">Sta. D</collectorName>
3AR3, 
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, 
<geoCoordinate id="EE783E89FFDB916863F0FEAEFD8DFEF7" box="[424,604,289,314]" direction="west" orientation="longitude" pageId="12" pageNumber="221" precision="1" value="-157.897">157°53'49.2&quot; W</geoCoordinate>
, 
<quantity id="4CB4F5ABFFDB9168603FFEAEFD4EFEF7" box="[615,671,289,314]" metricMagnitude="1" metricUnit="m" metricValue="5.0" pageId="12" pageNumber="221" unit="m" value="50.0">
<elevation id="0061BF7DFFDB9168603FFEAEFD4EFEF7" box="[615,671,289,314]" metricMagnitude="1" metricUnit="m" metricValue="5.0" pageId="12" pageNumber="221" unit="m" value="50.0">50 m</elevation>
</quantity>
(
<specimenCode id="DBEAF035FFDB916860F5FEAEFC84FEF7" box="[685,853,289,313]" collectionCode="BPBM-R" pageId="12" pageNumber="221">BPBM-R3887</specimenCode>
)
</materialsCitation>
; 
<materialsCitation id="3B245213FFDB91686130FEADFB6EFE93" ID-GBIF-Occurrence="2234219890" collectionCode="BPBM-R" collectorName="Sta. D" elevation="50" latitude="21.282" location="Mamala Bay" longLatPrecision="1" longitude="-157.91016" municipality="Sand Island" pageId="12" pageNumber="221" specimenCode="BPBM-R3888" specimenCount="2" typeStatus="paratype">
<typeStatus id="54F7E6ECFFDB91686130FEADFC04FEF4" box="[872,981,290,314]" pageId="12" pageNumber="221" type="paratype">Paratypes</typeStatus>
: same locality, date as 
<typeStatus id="54F7E6ECFFDB9168668DFEAEFAE6FEF4" box="[1237,1335,289,314]" pageId="12" pageNumber="221" type="holotype">holotype</typeStatus>
(4 spms, 
<specimenCode id="DBEAF035FFDB916862CFFEC9FEEFFE90" box="[151,318,326,350]" collectionCode="BPBM-R" pageId="12" pageNumber="221">BPBM-R3888</specimenCode>
). Additional material examined: 
<collectorName id="26B93D98FFDB91686099FEC9FCDEFE90" box="[705,783,326,350]" pageId="12" pageNumber="221">Sta. D</collectorName>
2R1, 
<geoCoordinate id="EE783E89FFDB91686115FECAFC24FE93" box="[845,1013,325,350]" direction="north" orientation="latitude" pageId="12" pageNumber="221" precision="1" value="21.282">21°16'55.2&quot; N</geoCoordinate>
, 
<geoCoordinate id="EE783E89FFDB9168665DFECAFB6EFE93" box="[1029,1215,325,350]" direction="west" orientation="longitude" pageId="12" pageNumber="221" precision="1" value="-157.91016">157°54'36.6&quot; W</geoCoordinate>
</materialsCitation>
, 
<quantity id="4CB4F5ABFFDB91686697FECAFAF3FE93" box="[1231,1314,325,350]" metricMagnitude="1" metricUnit="m" metricValue="5.64" pageId="12" pageNumber="221" unit="m" value="56.4">56.4 m</quantity>
(2 spms); Waianae outfall, May/2015, Sta. W2R6, 
<geoCoordinate id="EE783E89FFDB9168603AFEE6FCD7FE4F" box="[610,774,361,386]" direction="north" orientation="latitude" pageId="12" pageNumber="221" precision="1" value="21.412916">21°24'46.5&quot; N</geoCoordinate>
, 
<geoCoordinate id="EE783E89FFDB9168614AFEE6FC16FE4F" box="[786,967,361,386]" direction="west" orientation="longitude" pageId="12" pageNumber="221" precision="1" value="-158.196">158°11'45.6&quot; W</geoCoordinate>
, 
<quantity id="4CB4F5ABFFDB9168618AFEE6FBF0FE4F" box="[978,1057,361,385]" metricMagnitude="1" metricUnit="m" metricValue="2.77" pageId="12" pageNumber="221" unit="m" value="27.7">27.7 m</quantity>
(17); Sta. ZWR4, 
<geoCoordinate id="EE783E89FFDB916866ACFEE6FA49FE4F" box="[1268,1432,361,386]" direction="north" orientation="latitude" pageId="12" pageNumber="221" precision="1" value="21.423637">21°25'25.1&quot; N</geoCoordinate>
, 
<geoCoordinate id="EE783E89FFDB916862CFFE02FE9DFE6B" box="[151,332,397,422]" direction="west" orientation="longitude" pageId="12" pageNumber="221" precision="1" value="-158.19873">158°11'55.4&quot; W</geoCoordinate>
, 
<quantity id="4CB4F5ABFFDB91686301FE01FE78FE6B" box="[345,425,398,422]" metricMagnitude="1" metricUnit="m" metricValue="3.41" pageId="12" pageNumber="221" unit="m" value="34.1">34.1 m</quantity>
(2); Sta. W1R1, 
<geoCoordinate id="EE783E89FFDB91686037FE02FCC5FE6B" box="[623,788,397,422]" direction="north" orientation="latitude" pageId="12" pageNumber="221" precision="1" value="21.39236">21°23'32.5&quot; N</geoCoordinate>
, 
<geoCoordinate id="EE783E89FFDB91686179FE02FC06FE6B" box="[801,983,397,422]" direction="west" orientation="longitude" pageId="12" pageNumber="221" precision="1" value="-158.19102">158°11'27.7&quot; W</geoCoordinate>
, 
<materialsCitation id="3B245213FFDB916861BBFE01FB68FE07" ID-GBIF-Occurrence="2234220027" collectorName="Sta. &amp; R. Brock" elevation="32" latitude="21.423138" location="Kaneohe Bay" longLatPrecision="1" longitude="-158.19653" municipality="Oct" pageId="12" pageNumber="221" specimenCount="1" typeStatus="paratype">
<quantity id="4CB4F5ABFFDB916861BBFE01FBE2FE6B" box="[995,1075,398,422]" metricMagnitude="1" metricUnit="m" metricValue="3.17" pageId="12" pageNumber="221" unit="m" value="31.7">
<elevation id="0061BF7DFFDB916861BBFE01FBE2FE6B" box="[995,1075,398,422]" metricMagnitude="1" metricUnit="m" metricValue="3.17" pageId="12" pageNumber="221" unit="m" value="31.7">31.7 m</elevation>
</quantity>
(2); 
<collectorName id="26B93D98FFDB91686635FE01FB49FE68" box="[1133,1176,398,422]" pageId="12" pageNumber="221">Sta.</collectorName>
ZER1, 
<geoCoordinate id="EE783E89FFDB916866ABFE02FA49FE6B" box="[1267,1432,397,422]" direction="north" orientation="latitude" pageId="12" pageNumber="221" precision="1" value="21.423138">21°25'23.3&quot; N</geoCoordinate>
, 
<geoCoordinate id="EE783E89FFDB916862CFFE3EFE9BFE07" box="[151,330,433,458]" direction="west" orientation="longitude" pageId="12" pageNumber="221" precision="1" value="-158.19653">158°11'47.5&quot; W</geoCoordinate>
, 
<quantity id="4CB4F5ABFFDB9168630CFE3EFE70FE07" box="[340,417,433,458]" metricMagnitude="1" metricUnit="m" metricValue="3.05" pageId="12" pageNumber="221" unit="m" value="30.5">
<elevation id="0061BF7DFFDB9168630CFE3EFE70FE07" box="[340,417,433,458]" metricMagnitude="1" metricUnit="m" metricValue="3.05" pageId="12" pageNumber="221" unit="m" value="30.5">30.5 m</elevation>
</quantity>
(7). 
<location id="8E930E95FFDB9168638DFE3EFDBCFE04" LSID="urn:lsid:plazi:treatment:03E5E958FFDB917462CFFF18FE5EFEAB:8E930E95FFDB9168638DFE3EFDBCFE04" box="[469,621,433,458]" latitude="21.423138" longLatPrecision="1" longitude="-158.19653" municipality="Oct" name="Kaneohe Bay" pageId="12" pageNumber="221">Kaneohe Bay</location>
, 
<quantity id="4CB4F5ABFFDB9168602FFE3EFD18FE04" box="[631,713,433,458]" metricMagnitude="4" metricUnit="m" metricValue="1.2874752" pageId="12" pageNumber="221" unit="mi" value="8.0">8 miles</quantity>
offshore, 
<quantity id="4CB4F5ABFFDB91686161FE3EFCA2FE07" box="[825,883,433,457]" metricMagnitude="1" metricUnit="m" metricValue="3.0" pageId="12" pageNumber="221" unit="m" value="30.0">
<elevation id="0061BF7DFFDB91686161FE3EFCA2FE07" box="[825,883,433,457]" metricMagnitude="1" metricUnit="m" metricValue="3.0" pageId="12" pageNumber="221" unit="m" value="30.0">30 m</elevation>
</quantity>
, 
<collectingMunicipality id="6B97C234FFDB91686127FE3EFC78FE07" box="[895,937,433,457]" pageId="12" pageNumber="221">Oct</collectingMunicipality>
/1982, coll. 
<collectorName id="26B93D98FFDB91686673FE3DFB5EFE04" box="[1067,1167,433,458]" pageId="12" pageNumber="221">R. Brock</collectorName>
(3)
</materialsCitation>
; 
<materialsCitation id="3B245213FFDB9168669DFE3EFF0CFDDC" ID-GBIF-Occurrence="2234220001" collectorName="Waikiki &amp; Sta. SW &amp; Feb &amp; M. Hixon" county="About" location="Halape" municipality="Park" pageId="12" pageNumber="221" specimenCount="1" typeStatus="paratype">
<location id="8E930E95FFDB9168669DFE3EFAC6FE04" LSID="urn:lsid:plazi:treatment:03E5E958FFDB917462CFFF18FE5EFEAB:8E930E95FFDB9168669DFE3EFAC6FE04" box="[1221,1303,433,458]" county="About" municipality="Park" name="Halape" pageId="12" pageNumber="221">Halape</location>
, 
<location id="8E930E95FFDB9168677AFE3EFA49FE04" LSID="urn:lsid:plazi:treatment:03E5E958FFDB917462CFFF18FE5EFEAB:8E930E95FFDB9168677AFE3EFA49FE04" box="[1314,1432,433,458]" county="About" municipality="Park" name="Big Island" pageId="12" pageNumber="221">Big Island</location>
, 
<location id="8E930E95FFDB916862CFFE5AFED3FE20" LSID="urn:lsid:plazi:treatment:03E5E958FFDB917462CFFF18FE5EFEAB:8E930E95FFDB916862CFFE5AFED3FE20" box="[151,258,469,494]" county="About" municipality="Park" name="Hawaiian" pageId="12" pageNumber="221">Hawaiian</location>
volcano national 
<collectingMunicipality id="6B97C234FFDB91686397FE5AFDD4FE20" box="[463,517,469,494]" pageId="12" pageNumber="221">Park</collectingMunicipality>
, shallow subtidal (2). 
<collectingCounty id="629220C2FFDB9168615AFE5AFC99FE20" box="[770,840,469,494]" pageId="12" pageNumber="221">About</collectingCounty>
<quantity id="4CB4F5ABFFDB91686109FE59FC5AFE20" box="[849,907,469,494]" metricMagnitude="3" metricUnit="m" metricValue="1.0" pageId="12" pageNumber="221" unit="km" value="1.0">1 km</quantity>
offshore 
<collectorName id="26B93D98FFDB916861A1FE5AFB85FE20" box="[1017,1108,469,494]" pageId="12" pageNumber="221">Waikiki</collectorName>
, 
<collectorName id="26B93D98FFDB91686638FE59FB6CFE20" box="[1120,1213,470,494]" pageId="12" pageNumber="221">Sta. SW</collectorName>
, 
<collectorName id="26B93D98FFDB91686692FE5AFB24FE20" box="[1226,1269,469,494]" pageId="12" pageNumber="221">Feb</collectorName>
/2016, coll. 
<collectorName id="26B93D98FFDB91686724FE59FF0CFDDC" pageId="12" pageNumber="221">M. Hixon</collectorName>
</materialsCitation>
(9).
</paragraph>
</subSubSection>
<subSubSection id="C3560BC5FFDB9168629FFD92FF26FD6C" pageId="12" pageNumber="221" type="diagnosis">
<paragraph id="8BF3584EFFDB9168629FFD92FF26FD6C" blockId="12.[151,1437,253,2033]" pageId="12" pageNumber="221">
<emphasis id="B938845CFFDB9168629FFD92FEEEFDF8" bold="true" box="[199,319,541,566]" pageId="12" pageNumber="221">Diagnosis.</emphasis>
Parapodia biramous, chaetae emerging from body wall throughout; minute digitate ventral cirrus; interparapodial ciliated sensorial cilia present anteriorly and forming distinct lobes on last 5–7 chaetigers. Anal tube very short; posterior border with a basal pair of conical papillae, and up to four pairs of short, digitate marginal papillae.
</paragraph>
</subSubSection>
<subSubSection id="C3560BC5FFDB9168629FFD22FE6AFABC" pageId="12" pageNumber="221" type="description">
<paragraph id="8BF3584EFFDB9168629FFD22FE75FC0C" blockId="12.[151,1437,253,2033]" pageId="12" pageNumber="221">
<emphasis id="B938845CFFDB9168629FFD22FE87FD08" bold="true" box="[199,342,685,710]" pageId="12" pageNumber="221">Description.</emphasis>
<typeStatus id="54F7E6ECFFDB91686304FD22FE13FD08" box="[348,450,685,710]" pageId="12" pageNumber="221" type="holotype">Holotype</typeStatus>
<quantity id="4CB4F5ABFFDB91686392FD22FDCFFD0B" box="[458,542,685,710]" metricMagnitude="-3" metricUnit="m" metricValue="7.5" pageId="12" pageNumber="221" unit="mm" value="7.5">7.5 mm</quantity>
long, 
<quantity id="4CB4F5ABFFDB9168603CFD21FD69FD0B" box="[612,696,686,710]" metricMagnitude="-4" metricUnit="m" metricValue="3.0" pageId="12" pageNumber="221" unit="mm" value="0.3">0.3 mm</quantity>
wide for 26 chaetigers. 
<typeStatus id="54F7E6ECFFDB9168619AFD21FBFCFD08" box="[962,1069,686,710]" pageId="12" pageNumber="221" type="paratype">Paratypes</typeStatus>
ranging from 
<quantity id="4CB4F5ABFFDB91686693FD22FAF7FD0B" box="[1227,1318,685,710]" metricMagnitude="-3" metricUnit="m" metricValue="6.5" metricValueMax="8.0" metricValueMin="5.0" pageId="12" pageNumber="221" unit="mm" value="6.5" valueMax="8.0" valueMin="5.0">5–8 mm</quantity>
long, 
<quantity id="4CB4F5ABFFDB91686734FD21FF3DFD27" metricMagnitude="-4" metricUnit="m" metricValue="4.0" metricValueMax="5.0" metricValueMin="3.0" pageId="12" pageNumber="221" unit="mm" value="0.4" valueMax="0.5" valueMin="0.3">0.3– 0.5 mm</quantity>
wide for up to 28 chaetigers. Body slender, tubular tapering from last five chaetigers with deep ventral and lateral grooves (
<figureCitation id="137744CBFFDB91686312FD7AFE18FCC3" box="[330,457,757,782]" captionStart="FIGURE 8" captionStartId="15.[151,250,1887,1909]" captionTargetBox="[253,1294,443,1847]" captionTargetId="figure@15.[253,1333,419,1859]" captionTargetPageId="15" captionText="FIGURE 8. Polyophthalmus mauliola sp. nov. from Mamala Bay, Hawaii: A, complete specimen in dorsal view; B, anterior end in lateral view; C, anterior end in dorsal view; D, parapodium of the posterior end; E, posterior end with pygidium in lateral view; F, posterior end with pygidium in dorsal view." httpUri="https://zenodo.org/record/2624309/files/figure.png" pageId="12" pageNumber="221">Figs 8A, B</figureCitation>
; 
<figureCitation id="137744CBFFDB9168638EFD79FE2DFCC0" box="[470,508,758,782]" captionStart="FIGURE 9" captionStartId="16.[151,249,1540,1562]" captionTargetBox="[253,1333,411,1512]" captionTargetId="figure@16.[253,1333,411,1518]" captionTargetPageId="16" captionText="FIGURE 9. SEM of Polyophthalmus mauliola sp. nov. from Mamala Bay, Hawaii: A, complete specimen in lateral view; B, anterior end, dorsal view; C, parapodia of mid-body; D, parapodium of mid-body; E, pygidium and last chaetigers in lateral view." httpUri="https://zenodo.org/record/2624311/files/figure.png" pageId="12" pageNumber="221">9A</figureCitation>
); lateral groove beginning anterior to chaetiger 1 (
<figureCitation id="137744CBFFDB91686676FD7AFB7CFCC3" box="[1070,1197,757,782]" captionStart="FIGURE 8" captionStartId="15.[151,250,1887,1909]" captionTargetBox="[253,1294,443,1847]" captionTargetId="figure@15.[253,1333,419,1859]" captionTargetPageId="15" captionText="FIGURE 8. Polyophthalmus mauliola sp. nov. from Mamala Bay, Hawaii: A, complete specimen in dorsal view; B, anterior end in lateral view; C, anterior end in dorsal view; D, parapodium of the posterior end; E, posterior end with pygidium in lateral view; F, posterior end with pygidium in dorsal view." httpUri="https://zenodo.org/record/2624309/files/figure.png" pageId="12" pageNumber="221">Figs 8A, B</figureCitation>
; 
<figureCitation id="137744CBFFDB916866E2FD79FAD1FCC0" box="[1210,1280,758,782]" captionStart="FIGURE 9" captionStartId="16.[151,249,1540,1562]" captionTargetBox="[253,1333,411,1512]" captionTargetId="figure@16.[253,1333,411,1518]" captionTargetPageId="16" captionText="FIGURE 9. SEM of Polyophthalmus mauliola sp. nov. from Mamala Bay, Hawaii: A, complete specimen in lateral view; B, anterior end, dorsal view; C, parapodia of mid-body; D, parapodium of mid-body; E, pygidium and last chaetigers in lateral view." httpUri="https://zenodo.org/record/2624311/files/figure.png" pageId="12" pageNumber="221">9A, B</figureCitation>
). Prostomium broadly rounded, palpode absent (
<figureCitation id="137744CBFFDB91686053FC96FD9FFCFC" box="[523,590,793,818]" captionStart="FIGURE 8" captionStartId="15.[151,250,1887,1909]" captionTargetBox="[253,1294,443,1847]" captionTargetId="figure@15.[253,1333,419,1859]" captionTargetPageId="15" captionText="FIGURE 8. Polyophthalmus mauliola sp. nov. from Mamala Bay, Hawaii: A, complete specimen in dorsal view; B, anterior end in lateral view; C, anterior end in dorsal view; D, parapodium of the posterior end; E, posterior end with pygidium in lateral view; F, posterior end with pygidium in dorsal view." httpUri="https://zenodo.org/record/2624309/files/figure.png" pageId="12" pageNumber="221">Figs 8</figureCitation>
A–C; 9A, B). Nuchal organs large, deep oval lateral depressions (
<figureCitation id="137744CBFFDB91686745FC96FA46FCFC" box="[1309,1431,793,818]" captionStart="FIGURE 8" captionStartId="15.[151,250,1887,1909]" captionTargetBox="[253,1294,443,1847]" captionTargetId="figure@15.[253,1333,419,1859]" captionTargetPageId="15" captionText="FIGURE 8. Polyophthalmus mauliola sp. nov. from Mamala Bay, Hawaii: A, complete specimen in dorsal view; B, anterior end in lateral view; C, anterior end in dorsal view; D, parapodium of the posterior end; E, posterior end with pygidium in lateral view; F, posterior end with pygidium in dorsal view." httpUri="https://zenodo.org/record/2624309/files/figure.png" pageId="12" pageNumber="221">Figs 8B, C</figureCitation>
; 
<figureCitation id="137744CBFFDB916862CFFCB1FF6BFC98" box="[151,186,830,854]" captionStart="FIGURE 9" captionStartId="16.[151,249,1540,1562]" captionTargetBox="[253,1333,411,1512]" captionTargetId="figure@16.[253,1333,411,1518]" captionTargetPageId="16" captionText="FIGURE 9. SEM of Polyophthalmus mauliola sp. nov. from Mamala Bay, Hawaii: A, complete specimen in lateral view; B, anterior end, dorsal view; C, parapodia of mid-body; D, parapodium of mid-body; E, pygidium and last chaetigers in lateral view." httpUri="https://zenodo.org/record/2624311/files/figure.png" pageId="12" pageNumber="221">9B</figureCitation>
). A pair of small reddish eyespots deeply and ventrally embedded in prostomium seen in all specimens and a third and smaller eyespot on dorsal side observed in some specimens (
<figureCitation id="137744CBFFDB916861D4FCEEFBDAFCB4" box="[908,1035,865,890]" captionStart="FIGURE 8" captionStartId="15.[151,250,1887,1909]" captionTargetBox="[253,1294,443,1847]" captionTargetId="figure@15.[253,1333,419,1859]" captionTargetPageId="15" captionText="FIGURE 8. Polyophthalmus mauliola sp. nov. from Mamala Bay, Hawaii: A, complete specimen in dorsal view; B, anterior end in lateral view; C, anterior end in dorsal view; D, parapodium of the posterior end; E, posterior end with pygidium in lateral view; F, posterior end with pygidium in dorsal view." httpUri="https://zenodo.org/record/2624309/files/figure.png" pageId="12" pageNumber="221">Fig. 8B, C</figureCitation>
). Pharynx not observed. 
<typeStatus id="54F7E6ECFFDB9168676CFCEEFA4DFCB4" box="[1332,1436,865,890]" pageId="12" pageNumber="221" type="holotype">Holotype</typeStatus>
preserved pale yellow in color and some 
<typeStatus id="54F7E6ECFFDB91686035FC09FD09FC50" box="[621,728,902,926]" pageId="12" pageNumber="221" type="paratype">paratypes</typeStatus>
with lightly colored brown rings present on dorsal and lateral region, absent ventrally.
</paragraph>
<paragraph id="8BF3584EFFDB9168629FFC42FD4FFAC8" blockId="12.[151,1437,253,2033]" pageId="12" pageNumber="221">
Parapodia biramous, chaetae emerging from body wall throughout; minute digitate ventral cirrus also present throughout; interparapodial ciliated sensorial cilia present throughout half way between notopodia and neuropodia chaetae and forming a distinct lobe on last 5–7 chaetigers (
<figureCitation id="137744CBFFDB9168611DFB9AFC5DFBE0" box="[837,908,1045,1070]" captionStart="FIGURE 8" captionStartId="15.[151,250,1887,1909]" captionTargetBox="[253,1294,443,1847]" captionTargetId="figure@15.[253,1333,419,1859]" captionTargetPageId="15" captionText="FIGURE 8. Polyophthalmus mauliola sp. nov. from Mamala Bay, Hawaii: A, complete specimen in dorsal view; B, anterior end in lateral view; C, anterior end in dorsal view; D, parapodium of the posterior end; E, posterior end with pygidium in lateral view; F, posterior end with pygidium in dorsal view." httpUri="https://zenodo.org/record/2624309/files/figure.png" pageId="12" pageNumber="221">Figs 8</figureCitation>
D–F; 9D, E). Simple capillary chaetae in two bundles; 2–3 capillaries per bundle anteriorly reducing to 1–2 posteriorly; notochaetae slightly longer than neurochaetae; additional 3–4 short capillaries per bundle seen only with SEM. Chaetae of last 5–7 posterior chaetigers longer and surpassing pygidium (
<figureCitation id="137744CBFFDB916860CAFB0EFCE7FB57" box="[658,822,1153,1178]" captionStart="FIGURE 8" captionStartId="15.[151,250,1887,1909]" captionTargetBox="[253,1294,443,1847]" captionTargetId="figure@15.[253,1333,419,1859]" captionTargetPageId="15" captionText="FIGURE 8. Polyophthalmus mauliola sp. nov. from Mamala Bay, Hawaii: A, complete specimen in dorsal view; B, anterior end in lateral view; C, anterior end in dorsal view; D, parapodium of the posterior end; E, posterior end with pygidium in lateral view; F, posterior end with pygidium in dorsal view." httpUri="https://zenodo.org/record/2624309/files/figure.png" pageId="12" pageNumber="221">Figs 8A, E, F</figureCitation>
; 
<figureCitation id="137744CBFFDB9168611CFB0DFCB6FB54" box="[836,871,1154,1178]" captionStart="FIGURE 9" captionStartId="16.[151,249,1540,1562]" captionTargetBox="[253,1333,411,1512]" captionTargetId="figure@16.[253,1333,411,1518]" captionTargetPageId="16" captionText="FIGURE 9. SEM of Polyophthalmus mauliola sp. nov. from Mamala Bay, Hawaii: A, complete specimen in lateral view; B, anterior end, dorsal view; C, parapodia of mid-body; D, parapodium of mid-body; E, pygidium and last chaetigers in lateral view." httpUri="https://zenodo.org/record/2624311/files/figure.png" pageId="12" pageNumber="221">9E</figureCitation>
). Smallest individual (
<quantity id="4CB4F5ABFFDB91686636FB0DFB61FB57" box="[1134,1200,1154,1178]" metricMagnitude="-3" metricUnit="m" metricValue="1.0" pageId="12" pageNumber="221" unit="mm" value="1.0">1 mm</quantity>
long, 19 chaetigers) with capillaries of last segment very long, almost 1/6 of total body length. Lateral eyespots anterior to parapodia on 11 chaetigers (chaetigers 7–17), reddish brown, rounded; eyespots of chaetiger 7 and chaetiger 17 smaller than others; eyespots on posterior edge of segment.
</paragraph>
<paragraph id="8BF3584EFFDB9168629FFA9EFE6AFABC" blockId="12.[151,1437,253,2033]" pageId="12" pageNumber="221">
Anal tube very short, preserved specimens with tapering posterior end (
<figureCitation id="137744CBFFDB916861A8FA9EFBB7FAE4" box="[1008,1126,1297,1322]" captionStart="FIGURE 8" captionStartId="15.[151,250,1887,1909]" captionTargetBox="[253,1294,443,1847]" captionTargetId="figure@15.[253,1333,419,1859]" captionTargetPageId="15" captionText="FIGURE 8. Polyophthalmus mauliola sp. nov. from Mamala Bay, Hawaii: A, complete specimen in dorsal view; B, anterior end in lateral view; C, anterior end in dorsal view; D, parapodium of the posterior end; E, posterior end with pygidium in lateral view; F, posterior end with pygidium in dorsal view." httpUri="https://zenodo.org/record/2624309/files/figure.png" pageId="12" pageNumber="221">Fig. 8E, F</figureCitation>
). Posterior border provided with a basal pair of conical papillae, and up to four pairs of short, digitate marginal papillae can be completely retracted (
<figureCitation id="137744CBFFDB91686350FAD6FE51FABF" box="[264,384,1369,1394]" captionStart="FIGURE 8" captionStartId="15.[151,250,1887,1909]" captionTargetBox="[253,1294,443,1847]" captionTargetId="figure@15.[253,1333,419,1859]" captionTargetPageId="15" captionText="FIGURE 8. Polyophthalmus mauliola sp. nov. from Mamala Bay, Hawaii: A, complete specimen in dorsal view; B, anterior end in lateral view; C, anterior end in dorsal view; D, parapodium of the posterior end; E, posterior end with pygidium in lateral view; F, posterior end with pygidium in dorsal view." httpUri="https://zenodo.org/record/2624309/files/figure.png" pageId="12" pageNumber="221">Figs 8E, F</figureCitation>
; 
<figureCitation id="137744CBFFDB916863D4FAD5FE7EFABC" box="[396,431,1370,1394]" captionStart="FIGURE 9" captionStartId="16.[151,249,1540,1562]" captionTargetBox="[253,1333,411,1512]" captionTargetId="figure@16.[253,1333,411,1518]" captionTargetPageId="16" captionText="FIGURE 9. SEM of Polyophthalmus mauliola sp. nov. from Mamala Bay, Hawaii: A, complete specimen in lateral view; B, anterior end, dorsal view; C, parapodia of mid-body; D, parapodium of mid-body; E, pygidium and last chaetigers in lateral view." httpUri="https://zenodo.org/record/2624311/files/figure.png" pageId="12" pageNumber="221">9E</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="C3560BC5FFDB916B629FFAF2FA57FEAA" lastPageId="15" lastPageNumber="224" pageId="12" pageNumber="221" type="discussion">
<paragraph id="8BF3584EFFDB9168629FFAF2FF0EF940" blockId="12.[151,1437,253,2033]" pageId="12" pageNumber="221">
<emphasis id="B938845CFFDB9168629FFAF2FEEAFA58" bold="true" box="[199,315,1405,1430]" pageId="12" pageNumber="221">Remarks.</emphasis>
<tableCitation id="C6CE6DF5FFDB9168631CFAF2FE4BFA58" box="[324,410,1405,1430]" captionStart="TABLE 1" captionStartId="13.[147,228,678,701]" captionText="TABLE 1. Selected morphological characters of species of Polyophthalmus from the Indian and Pacific Oceans. Note that P. australis, P. ceylonensis, P. collaris, P. longisetosus and P. striatus are currently in synonymy with P. pictus sensu Hartman (1959). Polyophthalmus translucens was not included as it may belong to Ophelina abranchiata as suggested by Sene Silva (2007)." pageId="12" pageNumber="221">Table 1</tableCitation>
shows morphological features of eight 
<taxonomicName id="4C4C23CDFFDB9168613CFAF2FBCBFA58" authorityName="Quatrefages" authorityYear="1850" box="[868,1050,1405,1430]" class="Polychaeta" family="Opheliidae" genus="Polyophthalmus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Opheliida" pageId="12" pageNumber="221" phylum="Annelida" rank="genus">
<emphasis id="B938845CFFDB9168613CFAF2FBCBFA58" box="[868,1050,1405,1430]" italics="true" pageId="12" pageNumber="221">Polyophthalmus</emphasis>
</taxonomicName>
species present in the Indian and Pacific oceans, including 
<taxonomicName id="4C4C23CDFFDB916863E4FA2DFDCCFA77" baseAuthorityName="Dujardin" baseAuthorityYear="1839" box="[444,541,1442,1465]" class="Polychaeta" family="Opheliidae" genus="Polyophthalmus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Opheliida" pageId="12" pageNumber="221" phylum="Annelida" rank="species" species="pictus">
<emphasis id="B938845CFFDB916863E4FA2DFDCCFA77" box="[444,541,1442,1465]" italics="true" pageId="12" pageNumber="221">P. pictus</emphasis>
</taxonomicName>
from the Mediterranean for comparison. 
<taxonomicName id="4C4C23CDFFDB916861A1FA2FFACDFA77" authority="Magalhães &amp; Rizzo &amp; Bailey-Brock, 2019" authorityName="Magalhães &amp; Rizzo &amp; Bailey-Brock" authorityYear="2019" box="[1017,1308,1440,1465]" class="Polychaeta" family="Opheliidae" genus="Polyophthalmus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Opheliida" pageId="12" pageNumber="221" phylum="Annelida" rank="species" species="mauliola" status="sp. nov.">
<emphasis id="B938845CFFDB916861A1FA2FFACDFA77" box="[1017,1308,1440,1465]" italics="true" pageId="12" pageNumber="221">Polyophthalmus mauliola</emphasis>
</taxonomicName>
<emphasis id="B938845CFFDB9168677DFA2FFA51FA77" bold="true" box="[1317,1408,1440,1465]" pageId="12" pageNumber="221">
<taxonomicNameLabel id="A20B3927FFDB9168677DFA2FFA51FA77" box="[1317,1408,1440,1465]" pageId="12" pageNumber="221" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
is unique among all congeners by the presence of a minute, digitate ventral cirri in all segments and enlarged interparapodial ciliated sensory lobes on the very posterior segments. These lobes can be interpreted as interramal papillae because they emerge in between notopodial and neuropodial chaetae that are clearly separated and covered with sensory cilia. The distribution of the segmental eyes from chaetigers 7–17 is similar to what have been reported for 
<taxonomicName id="4C4C23CDFFDB9168637BF9DCFE71F9A5" authorityName="Grube" authorityYear="1869" box="[291,416,1618,1643]" class="Polychaeta" family="Opheliidae" genus="Polyophthalmus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Opheliida" pageId="12" pageNumber="221" phylum="Annelida" rank="species" species="australis">
<emphasis id="B938845CFFDB9168637BF9DCFE71F9A5" box="[291,416,1618,1643]" italics="true" pageId="12" pageNumber="221">P. australis</emphasis>
</taxonomicName>
and distinct from 
<taxonomicName id="4C4C23CDFFDB91686036F9DCFCDAF9A5" authorityName="Kukenthal" authorityYear="1887" box="[622,779,1618,1643]" class="Polychaeta" family="Opheliidae" genus="Polyophthalmus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Opheliida" pageId="12" pageNumber="221" phylum="Annelida" rank="species" species="ceylonensis">
<emphasis id="B938845CFFDB91686036F9DCFCDAF9A5" box="[622,779,1618,1643]" italics="true" pageId="12" pageNumber="221">P. ceylonensis</emphasis>
</taxonomicName>
(ch. 7–15), 
<taxonomicName id="4C4C23CDFFDB916861CBF9DCFB92F9A5" authorityName="Purschke, Ding &amp; Muller" authorityYear="1995" box="[915,1091,1618,1643]" class="Polychaeta" family="Opheliidae" genus="Polyophthalmus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Opheliida" pageId="12" pageNumber="221" phylum="Annelida" rank="species" species="qingdaoensis">
<emphasis id="B938845CFFDB916861CBF9DCFB92F9A5" box="[915,1091,1618,1643]" italics="true" pageId="12" pageNumber="221">P. qingdaoensis</emphasis>
</taxonomicName>
(ch. 7–18), and 
<taxonomicName id="4C4C23CDFFDB916866A2F9DCFABBF9A4" authorityName="Kukenthal" authorityYear="1887" box="[1274,1386,1619,1642]" class="Polychaeta" family="Opheliidae" genus="Polyophthalmus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Opheliida" pageId="12" pageNumber="221" phylum="Annelida" rank="species" species="striatus">
<emphasis id="B938845CFFDB916866A2F9DCFABBF9A4" box="[1274,1386,1619,1642]" italics="true" pageId="12" pageNumber="221">P. striatus</emphasis>
</taxonomicName>
(ch. 5–16).
</paragraph>
<paragraph id="8BF3584EFFDB9168629FF916FEC2F83F" blockId="12.[151,1437,253,2033]" pageId="12" pageNumber="221">
The body pigmentation has extensively been used to separate 
<taxonomicName id="4C4C23CDFFDB9168612EF916FBFDF97C" authorityName="Quatrefages" authorityYear="1850" box="[886,1068,1689,1714]" class="Polychaeta" family="Opheliidae" genus="Polyophthalmus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Opheliida" pageId="12" pageNumber="221" phylum="Annelida" rank="genus">
<emphasis id="B938845CFFDB9168612EF916FBFDF97C" box="[886,1068,1689,1714]" italics="true" pageId="12" pageNumber="221">Polyophthalmus</emphasis>
</taxonomicName>
species but preserved specimens of 
<taxonomicName id="4C4C23CDFFDB916862EFF931FEE9F91B" authority="Magalhães &amp; Rizzo &amp; Bailey-Brock, 2019" authorityName="Magalhães &amp; Rizzo &amp; Bailey-Brock" authorityYear="2019" box="[183,312,1724,1749]" class="Polychaeta" family="Opheliidae" genus="Polyophthalmus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Opheliida" pageId="12" pageNumber="221" phylum="Annelida" rank="species" species="mauliola" status="sp. nov.">
<emphasis id="B938845CFFDB916862EFF931FEE9F91B" box="[183,312,1724,1749]" italics="true" pageId="12" pageNumber="221">P. mauliola</emphasis>
</taxonomicName>
<emphasis id="B938845CFFDB91686319F933FE4DF91B" bold="true" box="[321,412,1724,1749]" pageId="12" pageNumber="221">
<taxonomicNameLabel id="A20B3927FFDB91686319F933FE4DF91B" box="[321,412,1724,1749]" pageId="12" pageNumber="221" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
have shown some variability from completely unpigmented and translucent to presenting transverse bands across dorsum. This is mostly due to fixation and preservation techniques, so the use of pigmentation to separate species should be done with caution and based on live material, if possible. 
<bibRefCitation id="EFDD25BFFFDB91686770F88CFF30F8F1" author="Kukenthal, W." pageId="12" pageNumber="221" pagination="361 - 373" refId="ref13264" refString="Kukenthal, W. (1887) Die Opheliaceen der expedition der Vettore Pisani. Jenaische Zeitschrift fur Naturwissenschaft, 21, 361 - 373." type="journal article" year="1887">Kükenthal (1887)</bibRefCitation>
presented illustrations of two species with distinct pigmentation, 
<taxonomicName id="4C4C23CDFFDB91686183F8A7FBABF8F1" authorityName="Kukenthal" authorityYear="1887" box="[987,1146,1830,1855]" class="Polychaeta" family="Opheliidae" genus="Polyophthalmus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Opheliida" pageId="12" pageNumber="221" phylum="Annelida" rank="species" species="ceylonensis">
<emphasis id="B938845CFFDB91686183F8A7FBABF8F1" box="[987,1146,1830,1855]" italics="true" pageId="12" pageNumber="221">P. ceylonensis</emphasis>
</taxonomicName>
and 
<taxonomicName id="4C4C23CDFFDB916866EEF8A7FAF9F8F1" authorityName="Kukenthal" authorityYear="1887" box="[1206,1320,1832,1855]" class="Polychaeta" family="Opheliidae" genus="Polyophthalmus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Opheliida" pageId="12" pageNumber="221" phylum="Annelida" rank="species" species="striatus">
<emphasis id="B938845CFFDB916866EEF8A7FAF9F8F1" box="[1206,1320,1832,1855]" italics="true" pageId="12" pageNumber="221">P. striatus</emphasis>
</taxonomicName>
. While 
<taxonomicName id="4C4C23CDFFDB916867DFF8A7FF3AF8AC" authorityName="Kukenthal" authorityYear="1887" class="Polychaeta" family="Opheliidae" genus="Polyophthalmus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Opheliida" pageId="12" pageNumber="221" phylum="Annelida" rank="species" species="striatus">
<emphasis id="B938845CFFDB916867DFF8A7FF3AF8AC" italics="true" pageId="12" pageNumber="221">P. striatus</emphasis>
</taxonomicName>
had complete transverse bands of dark brown color across the dorsum along the entire body, 
<taxonomicName id="4C4C23CDFFDB916866A7F8C4FA4DF8AD" authorityName="Kukenthal" authorityYear="1887" box="[1279,1436,1866,1891]" class="Polychaeta" family="Opheliidae" genus="Polyophthalmus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Opheliida" pageId="12" pageNumber="221" phylum="Annelida" rank="species" species="ceylonensis">
<emphasis id="B938845CFFDB916866A7F8C4FA4DF8AD" box="[1279,1436,1866,1891]" italics="true" pageId="12" pageNumber="221">P. ceylonensis</emphasis>
</taxonomicName>
only showed transverse bands on anterior five segments and patches of dark brown pigments on the dorsum from mid- to posterior segments. 
<bibRefCitation id="EFDD25BFFFDB91686381F81EFD07F864" author="Purschke, G. &amp; Ding, Z. &amp; Muller, M. C." box="[473,726,1937,1962]" pageId="12" pageNumber="221" pagination="229 - 241" refId="ref13839" refString="Purschke, G., Ding, Z. &amp; Muller, M. C. (1995) Ultrastructural differences as a taxonomic marker: the segmental ocelli of Polyophthalmus pictus and Polyophthalmus qingdaoensis sp. n. (Polychaeta, Opheliidae). Zoomorphology, 115 (4), 229 - 241. https: // doi. org / 10.1007 / BF 00393803" type="journal article" year="1995">
Purschke 
<emphasis id="B938845CFFDB91686011F81DFD52F864" box="[585,643,1937,1962]" italics="true" pageId="12" pageNumber="221">et al.</emphasis>
(1995)
</bibRefCitation>
showed that 
<taxonomicName id="4C4C23CDFFDB91686129F81DFBC4F867" box="[881,1045,1938,1961]" class="Polychaeta" family="Opheliidae" genus="Polyophthalmus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Opheliida" pageId="12" pageNumber="221" phylum="Annelida" rank="species" species="qingaoensis">
<emphasis id="B938845CFFDB91686129F81DFBC4F867" box="[881,1045,1938,1961]" italics="true" pageId="12" pageNumber="221">P. qingaoensis</emphasis>
</taxonomicName>
presented pigmented spots on the brain region of the prostomium in addition to dorsal ribbon-like spot of red pigments in most segments of live specimens.
</paragraph>
<caption id="DF3308C6FFDA916962CBFD29FE17FD36" pageId="13" pageNumber="222">
<paragraph id="8BF3584EFFDA916962CBFD29FE17FD36" blockId="13.[147,1849,677,760]" pageId="13" pageNumber="222">
<emphasis id="B938845CFFDA916962CBFD29FF2DFD73" bold="true" box="[147,252,678,701]" pageId="13" pageNumber="222">TABLE 1.</emphasis>
Selected morphological characters of species of 
<taxonomicName id="4C4C23CDFFDA91696096FD2AFCBBFD73" authorityName="Quatrefages" authorityYear="1850" box="[718,874,677,701]" class="Polychaeta" family="Opheliidae" genus="Polyophthalmus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Opheliida" pageId="13" pageNumber="222" phylum="Annelida" rank="genus">
<emphasis id="B938845CFFDA91696096FD2AFCBBFD73" box="[718,874,677,701]" italics="true" pageId="13" pageNumber="222">Polyophthalmus</emphasis>
</taxonomicName>
from the Indian and Pacific Oceans. Note that 
<taxonomicName id="4C4C23CDFFDA91696772FD2AFA49FD73" authorityName="Grube" authorityYear="1869" box="[1322,1432,677,701]" class="Polychaeta" family="Opheliidae" genus="Polyophthalmus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Opheliida" pageId="13" pageNumber="222" phylum="Annelida" rank="species" species="australis">
<emphasis id="B938845CFFDA91696772FD2AFA49FD73" box="[1322,1432,677,701]" italics="true" pageId="13" pageNumber="222">P. australis</emphasis>
</taxonomicName>
, 
<emphasis id="B938845CFFDA916967FCFD2AF94DFD73" box="[1444,1692,677,701]" italics="true" pageId="13" pageNumber="222">
<taxonomicName id="4C4C23CDFFDA916967FCFD2AF9FFFD73" authorityName="Kukenthal" authorityYear="1887" box="[1444,1582,677,701]" class="Polychaeta" family="Opheliidae" genus="Polyophthalmus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Opheliida" pageId="13" pageNumber="222" phylum="Annelida" rank="species" species="ceylonensis">P. ceylonensis</taxonomicName>
, 
<taxonomicName id="4C4C23CDFFDA91696460FD2AF94DFD73" authorityName="Michaelsen" authorityYear="1892" box="[1592,1692,677,701]" class="Polychaeta" family="Opheliidae" genus="Polyophthalmus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Opheliida" pageId="13" pageNumber="222" phylum="Annelida" rank="species" species="collaris">P. collaris</taxonomicName>
</emphasis>
, 
<taxonomicName id="4C4C23CDFFDA916964FFFD2AF8E8FD73" authorityName="Michaelsen" authorityYear="1892" box="[1703,1849,677,701]" class="Polychaeta" family="Opheliidae" genus="Polyophthalmus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Opheliida" pageId="13" pageNumber="222" phylum="Annelida" rank="species" species="longisetosus">
<emphasis id="B938845CFFDA916964FFFD2AF8E8FD73" box="[1703,1849,677,701]" italics="true" pageId="13" pageNumber="222">P. longisetosus</emphasis>
</taxonomicName>
and 
<taxonomicName id="4C4C23CDFFDA916962E3FD4CFECCFD15" authorityName="Kukenthal" authorityYear="1887" box="[187,285,707,731]" class="Polychaeta" family="Opheliidae" genus="Polyophthalmus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Opheliida" pageId="13" pageNumber="222" phylum="Annelida" rank="species" species="striatus">
<emphasis id="B938845CFFDA916962E3FD4CFECCFD15" box="[187,285,707,731]" italics="true" pageId="13" pageNumber="222">P. striatus</emphasis>
</taxonomicName>
are currently in synonymy with 
<taxonomicName id="4C4C23CDFFDA9169600CFD4CFC54FD15" authority="sensu Hartman (1959)" authorityName="Hartman" authorityYear="1959" box="[596,901,707,731]" class="Polychaeta" family="Opheliidae" genus="Polyophthalmus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Opheliida" pageId="13" pageNumber="222" phylum="Annelida" rank="species" species="pictus">
<emphasis id="B938845CFFDA9169600CFD4CFD32FD15" box="[596,739,707,731]" italics="true" pageId="13" pageNumber="222">P. pictus sensu</emphasis>
Hartman (1959)
</taxonomicName>
. 
<taxonomicName id="4C4C23CDFFDA916961D6FD4CFB4CFD15" baseAuthorityName="Blake &amp; Maciolek" baseAuthorityYear="2016" box="[910,1181,707,731]" class="Polychaeta" family="Opheliidae" genus="Polyophthalmus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Opheliida" pageId="13" pageNumber="222" phylum="Annelida" rank="species" species="translucens">
<emphasis id="B938845CFFDA916961D6FD4CFB4CFD15" box="[910,1181,707,731]" italics="true" pageId="13" pageNumber="222">Polyophthalmus translucens</emphasis>
</taxonomicName>
was not included as it may belong to 
<taxonomicName id="4C4C23CDFFDA9169645EFD4CF90BFD15" authorityName="Stop-Bowitz" authorityYear="1948" box="[1542,1754,707,731]" class="Polychaeta" family="Opheliidae" genus="Ophelina" higherTaxonomySource="GBIF" kingdom="Animalia" order="Opheliida" pageId="13" pageNumber="222" phylum="Annelida" rank="species" species="abranchiata">
<emphasis id="B938845CFFDA9169645EFD4CF90BFD15" box="[1542,1754,707,731]" italics="true" pageId="13" pageNumber="222">Ophelina abranchiata</emphasis>
</taxonomicName>
as suggested by Sene Silva (2007).
</paragraph>
</caption>
<paragraph id="8BF3584EFFDA916962CBFC91F98AFA7F" pageId="13" pageNumber="222">
<table id="F94CAAEEFFDA6E9B62CBFC91F8EFFA7F" box="[147,1854,798,1457]" gridcols="4" gridrows="9" pageId="13" pageNumber="222" rowsContinueIn="14.[147,1864,569,1692]">
<tr id="357C5A0CFFDA6E9B62CBFC91F8EFFCFB" box="[147,1854,798,821]" gridrow="0" pageId="13" pageNumber="222">
<th id="76AD3370FFDA6E9B62CBFC91FD88FCFB" box="[147,601,798,821]" gridcol="0" gridrow="0" pageId="13" pageNumber="222">
<emphasis id="B938845CFFDA916962CBFC91FF0CFCFB" bold="true" box="[147,221,798,821]" pageId="13" pageNumber="222">Species</emphasis>
</th>
<th id="76AD3370FFDA6E9B602EFC91FD2CFCFB" box="[630,765,798,821]" gridcol="1" gridrow="0" pageId="13" pageNumber="222">
<emphasis id="B938845CFFDA9169602FFC91FD2CFCFB" bold="true" box="[631,765,798,821]" pageId="13" pageNumber="222">Length (mm)</emphasis>
</th>
<th id="76AD3370FFDA6E9B6147FC91FC5CFCFB" box="[799,909,798,821]" gridcol="2" gridrow="0" pageId="13" pageNumber="222">
<emphasis id="B938845CFFDA91696147FC91FC5CFCFB" bold="true" box="[799,909,798,821]" pageId="13" pageNumber="222">Chaetigers</emphasis>
</th>
<th id="76AD3370FFDA6E9B6190FC91F8EFFCFB" box="[968,1854,798,821]" gridcol="3" gridrow="0" pageId="13" pageNumber="222">
<emphasis id="B938845CFFDA91696191FC91FB83FCFB" bold="true" box="[969,1106,798,821]" pageId="13" pageNumber="222">Pigmentation</emphasis>
</th>
</tr>
<tr id="357C5A0CFFDA6E9B62CBFCC6F8EFFCB1" box="[147,1854,841,895]" gridrow="1" pageId="13" pageNumber="222">
<th id="76AD3370FFDA6E9B62CBFCC6FD88FCB1" box="[147,601,841,895]" gridcol="0" gridrow="1" pageId="13" pageNumber="222">
<taxonomicName id="4C4C23CDFFDA916962CBFCC6FE59FCAF" authority="Magalhães &amp; Rizzo &amp; Bailey-Brock, 2019" authorityName="Magalhães &amp; Rizzo &amp; Bailey-Brock" authorityYear="2019" box="[147,392,841,865]" class="Polychaeta" family="Opheliidae" genus="Polyophthalmus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Opheliida" pageId="13" pageNumber="222" phylum="Annelida" rank="species" species="mauliola" status="sp. nov.">
<emphasis id="B938845CFFDA916962CBFCC6FE59FCAF" box="[147,392,841,865]" italics="true" pageId="13" pageNumber="222">Polyophthalmus mauliola</emphasis>
</taxonomicName>
<emphasis id="B938845CFFDA916963D6FCC5FE0AFCAF" bold="true" box="[398,475,842,865]" pageId="13" pageNumber="222">
<taxonomicNameLabel id="A20B3927FFDA916963D6FCC5FE0AFCAF" box="[398,475,842,865]" pageId="13" pageNumber="222" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
</th>
<td id="76AD3370FFDA6E9B602EFCC6FD2CFCB1" box="[630,765,841,895]" gridcol="1" gridrow="1" pageId="13" pageNumber="222">5–8</td>
<td id="76AD3370FFDA6E9B6147FCC6FC5CFCB1" box="[799,909,841,895]" gridcol="2" gridrow="1" pageId="13" pageNumber="222">27–28</td>
<td id="76AD3370FFDA6E9B6190FCC6F8EFFCB1" box="[968,1854,841,895]" gridcol="3" gridrow="1" pageId="13" pageNumber="222">Pale yellow in color and some paratypes with colored brown rings present on dorsal and lateral region, absent ventrally.</td>
</tr>
<tr id="357C5A0CFFDA6E9B62CBFC2DF8EFFC16" box="[147,1854,930,984]" gridrow="2" pageId="13" pageNumber="222">
<th id="76AD3370FFDA6E9B62CBFC2DFD88FC16" box="[147,601,930,984]" gridcol="0" gridrow="2" pageId="13" pageNumber="222">
<taxonomicName id="4C4C23CDFFDA916962CBFC2DFDD4FC74" authority="Grube, 1869" authorityName="Grube" authorityYear="1869" box="[147,517,930,954]" class="Polychaeta" family="Opheliidae" genus="Polyophthalmus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Opheliida" pageId="13" pageNumber="222" phylum="Annelida" rank="species" species="australis">
<emphasis id="B938845CFFDA916962CBFC2DFE59FC74" box="[147,392,930,954]" italics="true" pageId="13" pageNumber="222">Polyophthalmus australis</emphasis>
Grube, 1869
</taxonomicName>
</th>
<td id="76AD3370FFDA6E9B602EFC2DFD2CFC16" box="[630,765,930,984]" gridcol="1" gridrow="2" pageId="13" pageNumber="222">9–15</td>
<td id="76AD3370FFDA6E9B6147FC2DFC5CFC16" box="[799,909,930,984]" gridcol="2" gridrow="2" pageId="13" pageNumber="222">28</td>
<td id="76AD3370FFDA6E9B6190FC2DF8EFFC16" box="[968,1854,930,984]" gridcol="3" gridrow="2" pageId="13" pageNumber="222">Brown pigmentation in transverse bands reaching the parapodial region above segmental eyes.</td>
</tr>
<tr id="357C5A0CFFDA6E9B62CBFC75F8EFFBFE" box="[147,1854,1018,1072]" gridrow="3" pageId="13" pageNumber="222">
<th id="76AD3370FFDA6E9B62CBFC75FD88FBFE" box="[147,601,1018,1072]" gridcol="0" gridrow="3" pageId="13" pageNumber="222">
<taxonomicName id="4C4C23CDFFDA916962CBFC75FD96FBDC" authority="Kukenthal, 1887" authorityName="Kukenthal" authorityYear="1887" box="[147,583,1018,1042]" class="Polychaeta" family="Opheliidae" genus="Polyophthalmus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Opheliida" pageId="13" pageNumber="222" phylum="Annelida" rank="species" species="ceylonensis">
<emphasis id="B938845CFFDA916962CBFC75FE73FBDC" box="[147,418,1018,1042]" italics="true" pageId="13" pageNumber="222">Polyophthalmus ceylonensis</emphasis>
Kükenthal, 1887
</taxonomicName>
</th>
<td id="76AD3370FFDA6E9B602EFC75FD2CFBFE" box="[630,765,1018,1072]" gridcol="1" gridrow="3" pageId="13" pageNumber="222">11</td>
<td id="76AD3370FFDA6E9B6147FC75FC5CFBFE" box="[799,909,1018,1072]" gridcol="2" gridrow="3" pageId="13" pageNumber="222">27</td>
<td id="76AD3370FFDA6E9B6190FC75F8EFFBFE" box="[968,1854,1018,1072]" gridcol="3" gridrow="3" pageId="13" pageNumber="222">First five segments with two light brown bands; the following 15–18 segments with dorsal dark brown patches.</td>
</tr>
<tr id="357C5A0CFFDA6E9B62CBFBDCF8EFFBA5" box="[147,1854,1107,1131]" gridrow="4" pageId="13" pageNumber="222">
<th id="76AD3370FFDA6E9B62CBFBDCFD88FBA5" box="[147,601,1107,1131]" gridcol="0" gridrow="4" pageId="13" pageNumber="222">
<taxonomicName id="4C4C23CDFFDA916962CBFBDCFDFDFBA5" authority="Michaelsen, 1892" authorityName="Michaelsen" authorityYear="1892" box="[147,556,1107,1131]" class="Polychaeta" family="Opheliidae" genus="Polyophthalmus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Opheliida" pageId="13" pageNumber="222" phylum="Annelida" rank="species" species="collaris">
<emphasis id="B938845CFFDA916962CBFBDCFEACFBA5" box="[147,381,1107,1131]" italics="true" pageId="13" pageNumber="222">Polyophthalmus collaris</emphasis>
Michaelsen, 1892
</taxonomicName>
</th>
<td id="76AD3370FFDA6E9B602EFBDCFD2CFBA5" box="[630,765,1107,1131]" gridcol="1" gridrow="4" pageId="13" pageNumber="222">20</td>
<td id="76AD3370FFDA6E9B6147FBDCFC5CFBA5" box="[799,909,1107,1131]" gridcol="2" gridrow="4" pageId="13" pageNumber="222">~31</td>
<td id="76AD3370FFDA6E9B6190FBDCF8EFFBA5" box="[968,1854,1107,1131]" gridcol="3" gridrow="4" pageId="13" pageNumber="222">Pigmentation present on dorsal region of prostomium and peristomium.</td>
</tr>
<tr id="357C5A0CFFDA6E9B62CBFB01F8EFFB0A" box="[147,1854,1166,1220]" gridrow="5" pageId="13" pageNumber="222">
<th id="76AD3370FFDA6E9B62CBFB01FD88FB0A" box="[147,601,1166,1220]" gridcol="0" gridrow="5" pageId="13" pageNumber="222">
<taxonomicName id="4C4C23CDFFDA916962CBFB01FD88FB68" authority="Michaelsen, 1892" authorityName="Michaelsen" authorityYear="1892" box="[147,601,1166,1190]" class="Polychaeta" family="Opheliidae" genus="Polyophthalmus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Opheliida" pageId="13" pageNumber="222" phylum="Annelida" rank="species" species="longisetosus">
<emphasis id="B938845CFFDA916962CBFB01FE7BFB68" box="[147,426,1166,1190]" italics="true" pageId="13" pageNumber="222">Polyophthalmus longisetosus</emphasis>
Michaelsen, 1892
</taxonomicName>
</th>
<td id="76AD3370FFDA6E9B602EFB01FD2CFB0A" box="[630,765,1166,1220]" gridcol="1" gridrow="5" pageId="13" pageNumber="222">6.5</td>
<td id="76AD3370FFDA6E9B6147FB01FC5CFB0A" box="[799,909,1166,1220]" gridcol="2" gridrow="5" pageId="13" pageNumber="222">~26</td>
<td id="76AD3370FFDA6E9B6190FB01F8EFFB0A" box="[968,1854,1166,1220]" gridcol="3" gridrow="5" pageId="13" pageNumber="222">Translucent, dark patches present along the body, forming transverse bands (Michaelsen 1892; Fig. 14).</td>
</tr>
<tr id="357C5A0CFFDA6E9B62CBFB68F8EFFB31" box="[147,1854,1255,1279]" gridrow="6" pageId="13" pageNumber="222">
<th id="76AD3370FFDA6E9B62CBFB68FD88FB31" box="[147,601,1255,1279]" gridcol="0" gridrow="6" pageId="13" pageNumber="222">
<taxonomicName id="4C4C23CDFFDA916962CBFB68FDC5FB31" authority="(Dujardin, 1839)" authorityName="Dujardin" authorityYear="1839" baseAuthorityName="Dujardin" baseAuthorityYear="1839" box="[147,532,1255,1279]" class="Polychaeta" family="Opheliidae" genus="Polyophthalmus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Opheliida" pageId="13" pageNumber="222" phylum="Annelida" rank="species" species="pictus">
<emphasis id="B938845CFFDA916962CBFB68FEBCFB31" box="[147,365,1255,1279]" italics="true" pageId="13" pageNumber="222">Polyophthalmus pictus</emphasis>
(Dujardin, 1839)
</taxonomicName>
</th>
<td id="76AD3370FFDA6E9B602EFB68FD2CFB31" box="[630,765,1255,1279]" gridcol="1" gridrow="6" pageId="13" pageNumber="222">8–10</td>
<td id="76AD3370FFDA6E9B6147FB68FC5CFB31" box="[799,909,1255,1279]" gridcol="2" gridrow="6" pageId="13" pageNumber="222">~29</td>
<td id="76AD3370FFDA6E9B6190FB68F8EFFB31" box="[968,1854,1255,1279]" gridcol="3" gridrow="6" pageId="13" pageNumber="222">Green bands in live individuals; drawings show a pair of dark spots per segment.</td>
</tr>
<tr id="357C5A0CFFDA6E9B62CBFAADF8EFFABB" box="[147,1854,1314,1397]" gridrow="7" pageId="13" pageNumber="222">
<th id="76AD3370FFDA6E9B62CBFAADFD88FABB" box="[147,601,1314,1397]" gridcol="0" gridrow="7" pageId="13" pageNumber="222">
<taxonomicName id="4C4C23CDFFDA916962CBFAADFEF9FA96" authority="Purschke, Ding &amp; Muller, 1995" authorityName="Purschke, Ding &amp; Muller" authorityYear="1995" class="Polychaeta" family="Opheliidae" genus="Polyophthalmus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Opheliida" pageId="13" pageNumber="222" phylum="Annelida" rank="species" species="qingdaoensis">
<emphasis id="B938845CFFDA916962CBFAADFE63FAF4" box="[147,434,1314,1338]" italics="true" pageId="13" pageNumber="222">Polyophthalmus qingdaoensis</emphasis>
Purschke, Ding &amp; Müller, 1995
</taxonomicName>
</th>
<td id="76AD3370FFDA6E9B602EFAADFD2CFABB" box="[630,765,1314,1397]" gridcol="1" gridrow="7" pageId="13" pageNumber="222">3.9–5</td>
<td id="76AD3370FFDA6E9B6147FAADFC5CFABB" box="[799,909,1314,1397]" gridcol="2" gridrow="7" pageId="13" pageNumber="222">26</td>
<td id="76AD3370FFDA6E9B6190FAADF8EFFABB" box="[968,1854,1314,1397]" gridcol="3" gridrow="7" pageId="13" pageNumber="222">Almost colorless and translucent (living material), yellow-white (alcohol), dorsal ribbon-like spot of red pigment present in most segments except posterior-most 2–3; about 10 dark pigmented structures inside brain including ocelli.</td>
</tr>
<tr id="357C5A0CFFDA6E9B62CBFA16F8EFFA7F" box="[147,1854,1433,1457]" gridrow="8" pageId="13" pageNumber="222">
<th id="76AD3370FFDA6E9B62CBFA16FD88FA7F" box="[147,601,1433,1457]" gridcol="0" gridrow="8" pageId="13" pageNumber="222">
<taxonomicName id="4C4C23CDFFDA916962CBFA16FDF1FA7F" authority="Kukenthal, 1887" authorityName="Kukenthal" authorityYear="1887" box="[147,544,1433,1457]" class="Polychaeta" family="Opheliidae" genus="Polyophthalmus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Opheliida" pageId="13" pageNumber="222" phylum="Annelida" rank="species" species="striatus">
<emphasis id="B938845CFFDA916962CBFA16FEADFA7F" box="[147,380,1433,1457]" italics="true" pageId="13" pageNumber="222">Polyophthalmus striatus</emphasis>
Kükenthal, 1887
</taxonomicName>
</th>
<td id="76AD3370FFDA6E9B602EFA16FD2CFA7F" box="[630,765,1433,1457]" gridcol="1" gridrow="8" pageId="13" pageNumber="222">12</td>
<td id="76AD3370FFDA6E9B6147FA16FC5CFA7F" box="[799,909,1433,1457]" gridcol="2" gridrow="8" pageId="13" pageNumber="222">27</td>
<td id="76AD3370FFDA6E9B6190FA16F8EFFA7F" box="[968,1854,1433,1457]" gridcol="3" gridrow="8" pageId="13" pageNumber="222">Distinct pigmentation forming brown rings in intersegmental regions</td>
</tr>
</table>
</paragraph>
<paragraph id="8BF3584EFFDA91696444FA4AF899FA13" blockId="13.[1564,1864,1477,1501]" box="[1564,1864,1477,1501]" pageId="13" pageNumber="222">
<emphasis id="B938845CFFDA91696444FA4AF899FA13" box="[1564,1864,1477,1501]" italics="true" pageId="13" pageNumber="222">……continued on the next page</emphasis>
</paragraph>
<caption id="DF3308C6FFD9916A62CBFD8DFEAEFDD9" box="[147,383,514,535]" pageId="14" pageNumber="223">
<paragraph id="8BF3584EFFD9916A62CBFD8DFEAEFDD9" blockId="14.[147,383,514,535]" box="[147,383,514,535]" pageId="14" pageNumber="223">TABLE 1. (Continued)</paragraph>
</caption>
<paragraph id="8BF3584EFFD9916A62CBFDB6F8C8F9A9" pageId="14" pageNumber="223">
<table id="F94CAAEEFFD96E9B62CBFDB6F899F952" box="[147,1864,569,1692]" gridcols="10" gridrows="9" pageId="14" pageNumber="223" rowsContinueFrom="13.[147,1854,798,1457]">
<tr id="357C5A0CFFD96E9B62CBFDB6F899FD4A" box="[147,1864,569,644]" gridrow="0" pageId="14" pageNumber="223">
<th id="76AD3370FFD96E9B62CBFDB6FE91FD4A" box="[147,320,569,644]" gridcol="0" gridrow="0" pageId="14" pageNumber="223">
<emphasis id="B938845CFFD9916A62CBFDB6FF0FFD80" bold="true" box="[147,222,569,590]" pageId="14" pageNumber="223">Species</emphasis>
</th>
<th id="76AD3370FFD96E9B633CFDB6FE3BFD4A" box="[356,490,569,644]" gridcol="1" gridrow="0" pageId="14" pageNumber="223">
<emphasis id="B938845CFFD9916A633CFDB6FE32FD80" bold="true" box="[356,483,569,590]" pageId="14" pageNumber="223">Prostomium</emphasis>
</th>
<th id="76AD3370FFD96E9B607BFDB6FD7FFD4A" box="[547,686,569,644]" gridcol="2" gridrow="0" pageId="14" pageNumber="223">
<emphasis id="B938845CFFD9916A607BFDB6FD9FFDA7" bold="true" box="[547,654,569,617]" pageId="14" pageNumber="223">Segmental eyes</emphasis>
</th>
<th id="76AD3370FFD96E9B6093FDB6FCF6FD4A" box="[715,807,569,644]" gridcol="3" gridrow="0" pageId="14" pageNumber="223">
<emphasis id="B938845CFFD9916A6093FDB6FCF2FD80" bold="true" box="[715,803,569,590]" pageId="14" pageNumber="223">Pharynx</emphasis>
</th>
<th id="76AD3370FFD96E9B6104FDB6FB90FD4A" box="[860,1089,569,644]" gridcol="4" gridrow="0" pageId="14" pageNumber="223">
<emphasis id="B938845CFFD9916A6104FDB6FBCDFD80" bold="true" box="[860,1052,569,590]" pageId="14" pageNumber="223">Parapodia/chaetae</emphasis>
</th>
<th id="76AD3370FFD96E9B663DFDB6FB17FD4A" box="[1125,1222,569,644]" gridcol="5" gridrow="0" pageId="14" pageNumber="223">
<emphasis id="B938845CFFD9916A663DFDB6FB17FD4A" bold="true" pageId="14" pageNumber="223">Shape/ length anal cone</emphasis>
</th>
<th id="76AD3370FFD96E9B66BEFDB6FABEFD4A" box="[1254,1391,569,644]" gridcol="6" gridrow="0" pageId="14" pageNumber="223">
<emphasis id="B938845CFFD9916A66BEFDB6FABBFDA7" bold="true" box="[1254,1386,569,617]" pageId="14" pageNumber="223">Marginal anal papillae</emphasis>
</th>
<th id="76AD3370FFD96E9B67D4FDB6F9C4FD4A" box="[1420,1557,569,644]" gridcol="7" gridrow="0" pageId="14" pageNumber="223">
<emphasis id="B938845CFFD9916A67D4FDB6FA0FFDA7" bold="true" pageId="14" pageNumber="223">Basal anal papillae</emphasis>
</th>
<th id="76AD3370FFD96E9B646AFDB6F969FD4A" box="[1586,1720,569,644]" gridcol="8" gridrow="0" pageId="14" pageNumber="223">
<emphasis id="B938845CFFD9916A646BFDB6F956FDA7" bold="true" box="[1587,1671,569,617]" pageId="14" pageNumber="223">Type Locality</emphasis>
</th>
<th id="76AD3370FFD96E9B6481FDB6F899FD4A" box="[1753,1864,569,644]" gridcol="9" gridrow="0" pageId="14" pageNumber="223">
<emphasis id="B938845CFFD9916A6481FDB6F891FD80" bold="true" box="[1753,1856,569,590]" pageId="14" pageNumber="223">Reference</emphasis>
</th>
</tr>
<tr id="357C5A0CFFD96E9B62CBFD05F899FC8F" box="[147,1864,650,833]" gridrow="1" pageId="14" pageNumber="223">
<th id="76AD3370FFD96E9B62CBFD05FE91FC8F" box="[147,320,650,833]" gridcol="0" gridrow="1" pageId="14" pageNumber="223">
<taxonomicName id="4C4C23CDFFD9916A62CBFD05FF38FD74" authority="Magalhães &amp; Rizzo &amp; Bailey-Brock, 2019" authorityName="Magalhães &amp; Rizzo &amp; Bailey-Brock" authorityYear="2019" box="[147,304,650,698]" class="Polychaeta" family="Opheliidae" genus="Polyophthalmus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Opheliida" pageId="14" pageNumber="223" phylum="Annelida" rank="species" species="mauliola" status="sp. nov.">
<emphasis id="B938845CFFD9916A62CBFD05FF38FD74" box="[147,304,650,698]" italics="true" pageId="14" pageNumber="223">Polyophthalmus mauliola</emphasis>
</taxonomicName>
<emphasis id="B938845CFFD9916A62B6FD29FEEDFD75" bold="true" box="[238,316,678,699]" pageId="14" pageNumber="223">
<taxonomicNameLabel id="A20B3927FFD9916A62B6FD29FEEDFD75" box="[238,316,678,699]" pageId="14" pageNumber="223" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
</th>
<td id="76AD3370FFD96E9B633CFD05FE3BFC8F" box="[356,490,650,833]" gridcol="1" gridrow="1" pageId="14" pageNumber="223">Broadly rounded</td>
<td id="76AD3370FFD96E9B607BFD05FD7FFC8F" box="[547,686,650,833]" gridcol="2" gridrow="1" pageId="14" pageNumber="223">Ch. 7–17 (smaller 7, 17)</td>
<td id="76AD3370FFD96E9B6093FD05FCF6FC8F" box="[715,807,650,833]" gridcol="3" gridrow="1" pageId="14" pageNumber="223">Oral tentacles not seen</td>
<td id="76AD3370FFD96E9B6104FD05FB90FC8F" box="[860,1089,650,833]" gridcol="4" gridrow="1" pageId="14" pageNumber="223">A short, rounded interramal papilla present throughout and more distinct on last 5– 7 chaetigers; a minute digitate ventral cirrus also present throughout</td>
<td id="76AD3370FFD96E9B663DFD05FB17FC8F" box="[1125,1222,650,833]" gridcol="5" gridrow="1" pageId="14" pageNumber="223">Short</td>
<td id="76AD3370FFD96E9B66BEFD05FABEFC8F" box="[1254,1391,650,833]" gridcol="6" gridrow="1" pageId="14" pageNumber="223">Up to four pairs of short, digitate</td>
<td id="76AD3370FFD96E9B67D4FD05F9C4FC8F" box="[1420,1557,650,833]" gridcol="7" gridrow="1" pageId="14" pageNumber="223">Conical, longer than marginal papillae</td>
<td id="76AD3370FFD96E9B646AFD05F969FC8F" box="[1586,1720,650,833]" gridcol="8" gridrow="1" pageId="14" pageNumber="223">Mamala Bay, Hawaii</td>
<td id="76AD3370FFD96E9B6481FD05F899FC8F" box="[1753,1864,650,833]" gridcol="9" gridrow="1" pageId="14" pageNumber="223">This study</td>
</tr>
<tr id="357C5A0CFFD96E9B62CBFCEEF899FC09" box="[147,1864,865,967]" gridrow="2" pageId="14" pageNumber="223">
<th id="76AD3370FFD96E9B62CBFCEEFE91FC09" box="[147,320,865,967]" gridcol="0" gridrow="2" pageId="14" pageNumber="223">
<taxonomicName id="4C4C23CDFFD9916A62CBFCEEFF12FC62" authority="Grube, 1869" authorityName="Grube" authorityYear="1869" class="Polychaeta" family="Opheliidae" genus="Polyophthalmus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Opheliida" pageId="14" pageNumber="223" phylum="Annelida" rank="species" species="australis">
<emphasis id="B938845CFFD9916A62CBFCEEFF39FC5E" box="[147,304,865,912]" italics="true" pageId="14" pageNumber="223">Polyophthalmus australis</emphasis>
Grube, 1869
</taxonomicName>
</th>
<td id="76AD3370FFD96E9B633CFCEEFE3BFC09" box="[356,490,865,967]" gridcol="1" gridrow="2" pageId="14" pageNumber="223">Broadly rounded</td>
<td id="76AD3370FFD96E9B607BFCEEFD7FFC09" box="[547,686,865,967]" gridcol="2" gridrow="2" pageId="14" pageNumber="223">
Ch. 7–17 (larger than 
<taxonomicName id="4C4C23CDFFD9916A60C0FCF4FD8CFC65" baseAuthorityName="Dujardin" baseAuthorityYear="1839" class="Polychaeta" family="Opheliidae" genus="Polyophthalmus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Opheliida" pageId="14" pageNumber="223" phylum="Annelida" rank="species" species="pictus">
<emphasis id="B938845CFFD9916A60C0FCF4FD8CFC65" italics="true" pageId="14" pageNumber="223">P. pictus</emphasis>
</taxonomicName>
and apparent lens)
</td>
<td id="76AD3370FFD96E9B6093FCEEFCF6FC09" box="[715,807,865,967]" gridcol="3" gridrow="2" pageId="14" pageNumber="223">Not described</td>
<td id="76AD3370FFD96E9B6104FCEEFB90FC09" box="[860,1089,865,967]" gridcol="4" gridrow="2" pageId="14" pageNumber="223">Not described</td>
<td id="76AD3370FFD96E9B663DFCEEFB17FC09" box="[1125,1222,865,967]" gridcol="5" gridrow="2" pageId="14" pageNumber="223">Not described</td>
<td id="76AD3370FFD96E9B66BEFCEEFABEFC09" box="[1254,1391,865,967]" gridcol="6" gridrow="2" pageId="14" pageNumber="223">Nine</td>
<td id="76AD3370FFD96E9B67D4FCEEF9C4FC09" box="[1420,1557,865,967]" gridcol="7" gridrow="2" pageId="14" pageNumber="223">Present, a pair, larger than marginal papillae</td>
<td id="76AD3370FFD96E9B646AFCEEF969FC09" box="[1586,1720,865,967]" gridcol="8" gridrow="2" pageId="14" pageNumber="223">Queensland, Australia</td>
<td id="76AD3370FFD96E9B6481FCEEF899FC09" box="[1753,1864,865,967]" gridcol="9" gridrow="2" pageId="14" pageNumber="223">Grube (1869)</td>
</tr>
<tr id="357C5A0CFFD96E9B62CBFC43F899FBD6" box="[147,1864,972,1048]" gridrow="3" pageId="14" pageNumber="223">
<th id="76AD3370FFD96E9B62CBFC43FE91FBD6" box="[147,320,972,1048]" gridcol="0" gridrow="3" pageId="14" pageNumber="223">
<taxonomicName id="4C4C23CDFFD9916A62CBFC43FEE2FBD6" authority="Kukenthal, 1887" authorityName="Kukenthal" authorityYear="1887" box="[147,307,972,1048]" class="Polychaeta" family="Opheliidae" genus="Polyophthalmus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Opheliida" pageId="14" pageNumber="223" phylum="Annelida" rank="species" species="ceylonensis">
<emphasis id="B938845CFFD9916A62CBFC43FED3FC32" box="[147,304,972,1020]" italics="true" pageId="14" pageNumber="223">Polyophthalmus ceylonensis</emphasis>
Kükenthal, 1887
</taxonomicName>
</th>
<td id="76AD3370FFD96E9B633CFC43FE3BFBD6" box="[356,490,972,1048]" gridcol="1" gridrow="3" pageId="14" pageNumber="223">?</td>
<td id="76AD3370FFD96E9B607BFC43FD7FFBD6" box="[547,686,972,1048]" gridcol="2" gridrow="3" pageId="14" pageNumber="223">Ch. 7–15 (same sized)</td>
<td id="76AD3370FFD96E9B6093FC43FCF6FBD6" box="[715,807,972,1048]" gridcol="3" gridrow="3" pageId="14" pageNumber="223">Not described</td>
<td id="76AD3370FFD96E9B6104FC43FB90FBD6" box="[860,1089,972,1048]" gridcol="4" gridrow="3" pageId="14" pageNumber="223">Chaetae short and thin</td>
<td id="76AD3370FFD96E9B663DFC43FB17FBD6" box="[1125,1222,972,1048]" gridcol="5" gridrow="3" pageId="14" pageNumber="223">Not described</td>
<td id="76AD3370FFD96E9B66BEFC43FABEFBD6" box="[1254,1391,972,1048]" gridcol="6" gridrow="3" pageId="14" pageNumber="223">?</td>
<td id="76AD3370FFD96E9B67D4FC43F9C4FBD6" box="[1420,1557,972,1048]" gridcol="7" gridrow="3" pageId="14" pageNumber="223">A pair, elongate</td>
<td id="76AD3370FFD96E9B646AFC43F969FBD6" box="[1586,1720,972,1048]" gridcol="8" gridrow="3" pageId="14" pageNumber="223">Sri lanka</td>
<td id="76AD3370FFD96E9B6481FC43F899FBD6" box="[1753,1864,972,1048]" gridcol="9" gridrow="3" pageId="14" pageNumber="223">Kükenthal (1887)</td>
</tr>
<tr id="357C5A0CFFD96E9B62CBFBB7F899FB4D" box="[147,1864,1080,1155]" gridrow="4" pageId="14" pageNumber="223">
<th id="76AD3370FFD96E9B62CBFBB7FE91FB4D" box="[147,320,1080,1155]" gridcol="0" gridrow="4" pageId="14" pageNumber="223">
<taxonomicName id="4C4C23CDFFD9916A62CBFBB7FEEEFB4D" authority="Michaelsen, 1892" authorityName="Michaelsen" authorityYear="1892" box="[147,319,1080,1155]" class="Polychaeta" family="Opheliidae" genus="Polyophthalmus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Opheliida" pageId="14" pageNumber="223" phylum="Annelida" rank="species" species="collaris">
<emphasis id="B938845CFFD9916A62CBFBB7FF0CFBA9" box="[147,304,1080,1127]" italics="true" pageId="14" pageNumber="223">Polyophthalmus collaris</emphasis>
Michaelsen, 1892
</taxonomicName>
</th>
<td id="76AD3370FFD96E9B633CFBB7FE3BFB4D" box="[356,490,1080,1155]" gridcol="1" gridrow="4" pageId="14" pageNumber="223">Broadly rounded</td>
<td id="76AD3370FFD96E9B607BFBB7FD7FFB4D" box="[547,686,1080,1155]" gridcol="2" gridrow="4" pageId="14" pageNumber="223">11 pairs</td>
<td id="76AD3370FFD96E9B6093FBB7FCF6FB4D" box="[715,807,1080,1155]" gridcol="3" gridrow="4" pageId="14" pageNumber="223">Not described</td>
<td id="76AD3370FFD96E9B6104FBB7FB90FB4D" box="[860,1089,1080,1155]" gridcol="4" gridrow="4" pageId="14" pageNumber="223">Chaetae short, not exceeding body width</td>
<td id="76AD3370FFD96E9B663DFBB7FB17FB4D" box="[1125,1222,1080,1155]" gridcol="5" gridrow="4" pageId="14" pageNumber="223">Not described</td>
<td id="76AD3370FFD96E9B66BEFBB7FABEFB4D" box="[1254,1391,1080,1155]" gridcol="6" gridrow="4" pageId="14" pageNumber="223">Not described</td>
<td id="76AD3370FFD96E9B67D4FBB7F9C4FB4D" box="[1420,1557,1080,1155]" gridcol="7" gridrow="4" pageId="14" pageNumber="223">Not described</td>
<td id="76AD3370FFD96E9B646AFBB7F969FB4D" box="[1586,1720,1080,1155]" gridcol="8" gridrow="4" pageId="14" pageNumber="223">Sri Lanka</td>
<td id="76AD3370FFD96E9B6481FBB7F899FB4D" box="[1753,1864,1080,1155]" gridcol="9" gridrow="4" pageId="14" pageNumber="223">Michaelsen (1892)</td>
</tr>
<tr id="357C5A0CFFD96E9B62CBFB2CF899FAC4" box="[147,1864,1187,1290]" gridrow="5" pageId="14" pageNumber="223">
<th id="76AD3370FFD96E9B62CBFB2CFE91FAC4" box="[147,320,1187,1290]" gridcol="0" gridrow="5" pageId="14" pageNumber="223">
<taxonomicName id="4C4C23CDFFD9916A62CBFB2CFEEEFB20" authority="Michaelsen, 1892" authorityName="Michaelsen" authorityYear="1892" box="[147,319,1187,1262]" class="Polychaeta" family="Opheliidae" genus="Polyophthalmus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Opheliida" pageId="14" pageNumber="223" phylum="Annelida" rank="species" species="longisetosus">
<emphasis id="B938845CFFD9916A62CBFB2CFEDBFB1D" box="[147,304,1187,1235]" italics="true" pageId="14" pageNumber="223">Polyophthalmus longisetosus</emphasis>
Michaelsen, 1892
</taxonomicName>
</th>
<td id="76AD3370FFD96E9B633CFB2CFE3BFAC4" box="[356,490,1187,1290]" gridcol="1" gridrow="5" pageId="14" pageNumber="223">Broadly rounded, eyes absent</td>
<td id="76AD3370FFD96E9B607BFB2CFD7FFAC4" box="[547,686,1187,1290]" gridcol="2" gridrow="5" pageId="14" pageNumber="223">Ch. 4–18? (smaller 4, 17, 18?)</td>
<td id="76AD3370FFD96E9B6093FB2CFCF6FAC4" box="[715,807,1187,1290]" gridcol="3" gridrow="5" pageId="14" pageNumber="223">Not described</td>
<td id="76AD3370FFD96E9B6104FB2CFB90FAC4" box="[860,1089,1187,1290]" gridcol="4" gridrow="5" pageId="14" pageNumber="223">Not described; chaetae of last 5 segments greatly enlarged and small interramal papilla</td>
<td id="76AD3370FFD96E9B663DFB2CFB17FAC4" box="[1125,1222,1187,1290]" gridcol="5" gridrow="5" pageId="14" pageNumber="223">Not described</td>
<td id="76AD3370FFD96E9B66BEFB2CFABEFAC4" box="[1254,1391,1187,1290]" gridcol="6" gridrow="5" pageId="14" pageNumber="223">Elongate, at least 3 pairs (Michaelsen 1892; Fig. 14)</td>
<td id="76AD3370FFD96E9B67D4FB2CF9C4FAC4" box="[1420,1557,1187,1290]" gridcol="7" gridrow="5" pageId="14" pageNumber="223">A pair may be present based (Michaelsen 1892; Fig. 14)</td>
<td id="76AD3370FFD96E9B646AFB2CF969FAC4" box="[1586,1720,1187,1290]" gridcol="8" gridrow="5" pageId="14" pageNumber="223">Sri Lanka</td>
<td id="76AD3370FFD96E9B6481FB2CF899FAC4" box="[1753,1864,1187,1290]" gridcol="9" gridrow="5" pageId="14" pageNumber="223">Michaelsen (1892)</td>
</tr>
<tr id="357C5A0CFFD96E9B62CBFAA6F899FA5E" box="[147,1864,1321,1424]" gridrow="6" pageId="14" pageNumber="223">
<th id="76AD3370FFD96E9B62CBFAA6FE91FA5E" box="[147,320,1321,1424]" gridcol="0" gridrow="6" pageId="14" pageNumber="223">
<taxonomicName id="4C4C23CDFFD9916A62CBFAA6FF1AFABB" authority="(Dujardin, 1839)" baseAuthorityName="Dujardin" baseAuthorityYear="1839" class="Polychaeta" family="Opheliidae" genus="Polyophthalmus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Opheliida" pageId="14" pageNumber="223" phylum="Annelida" rank="species" species="pictus">
<emphasis id="B938845CFFD9916A62CBFAA6FF1CFA97" box="[147,304,1321,1369]" italics="true" pageId="14" pageNumber="223">Polyophthalmus pictus</emphasis>
(Dujardin, 1839)
</taxonomicName>
</th>
<td id="76AD3370FFD96E9B633CFAA6FE3BFA5E" box="[356,490,1321,1424]" gridcol="1" gridrow="6" pageId="14" pageNumber="223">Broadly rounded</td>
<td id="76AD3370FFD96E9B607BFAA6FD7FFA5E" box="[547,686,1321,1424]" gridcol="2" gridrow="6" pageId="14" pageNumber="223">?</td>
<td id="76AD3370FFD96E9B6093FAA6FCF6FA5E" box="[715,807,1321,1424]" gridcol="3" gridrow="6" pageId="14" pageNumber="223">Not described</td>
<td id="76AD3370FFD96E9B6104FAA6FB90FA5E" box="[860,1089,1321,1424]" gridcol="4" gridrow="6" pageId="14" pageNumber="223">Not described</td>
<td id="76AD3370FFD96E9B663DFAA6FB17FA5E" box="[1125,1222,1321,1424]" gridcol="5" gridrow="6" pageId="14" pageNumber="223">Short</td>
<td id="76AD3370FFD96E9B66BEFAA6FABEFA5E" box="[1254,1391,1321,1424]" gridcol="6" gridrow="6" pageId="14" pageNumber="223">Uneven, at least five, dorsal ones longer</td>
<td id="76AD3370FFD96E9B67D4FAA6F9C4FA5E" box="[1420,1557,1321,1424]" gridcol="7" gridrow="6" pageId="14" pageNumber="223">?</td>
<td id="76AD3370FFD96E9B646AFAA6F969FA5E" box="[1586,1720,1321,1424]" gridcol="8" gridrow="6" pageId="14" pageNumber="223">France</td>
<td id="76AD3370FFD96E9B6481FAA6F899FA5E" box="[1753,1864,1321,1424]" gridcol="9" gridrow="6" pageId="14" pageNumber="223">Dujardin (1839)</td>
</tr>
<tr id="357C5A0CFFD96E9B62CBFA3FF899F9D8" box="[147,1864,1456,1558]" gridrow="7" pageId="14" pageNumber="223">
<th id="76AD3370FFD96E9B62CBFA3FFE91F9D8" box="[147,320,1456,1558]" gridcol="0" gridrow="7" pageId="14" pageNumber="223">
<taxonomicName id="4C4C23CDFFD9916A62CBFA3FFEC1F9D8" authority="Purschke, Ding &amp; Muller, 1995" authorityName="Purschke, Ding &amp; Muller" authorityYear="1995" class="Polychaeta" family="Opheliidae" genus="Polyophthalmus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Opheliida" pageId="14" pageNumber="223" phylum="Annelida" rank="species" species="qingdaoensis">
<emphasis id="B938845CFFD9916A62CBFA3FFEC3FA2E" box="[147,304,1456,1504]" italics="true" pageId="14" pageNumber="223">Polyophthalmus qingdaoensis</emphasis>
Purschke, Ding &amp; Müller, 1995
</taxonomicName>
</th>
<td id="76AD3370FFD96E9B633CFA3FFE3BF9D8" box="[356,490,1456,1558]" gridcol="1" gridrow="7" pageId="14" pageNumber="223">Broadly rounded</td>
<td id="76AD3370FFD96E9B607BFA3FFD7FF9D8" box="[547,686,1456,1558]" gridcol="2" gridrow="7" pageId="14" pageNumber="223">Ch. 7–18 (smaller 7, 18)</td>
<td id="76AD3370FFD96E9B6093FA3FFCF6F9D8" box="[715,807,1456,1558]" gridcol="3" gridrow="7" pageId="14" pageNumber="223">Not described</td>
<td id="76AD3370FFD96E9B6104FA3FFB90F9D8" box="[860,1089,1456,1558]" gridcol="4" gridrow="7" pageId="14" pageNumber="223">Small lobe, lacking dorsal and ventral cirri</td>
<td id="76AD3370FFD96E9B663DFA3FFB17F9D8" box="[1125,1222,1456,1558]" gridcol="5" gridrow="7" pageId="14" pageNumber="223">Short, dorsal notch</td>
<td id="76AD3370FFD96E9B66BEFA3FFABEF9D8" box="[1254,1391,1456,1558]" gridcol="6" gridrow="7" pageId="14" pageNumber="223">Digitiform, 3 pairs, dorsal pair shorter</td>
<td id="76AD3370FFD96E9B67D4FA3FF9C4F9D8" box="[1420,1557,1456,1558]" gridcol="7" gridrow="7" pageId="14" pageNumber="223">Absent</td>
<td id="76AD3370FFD96E9B646AFA3FF969F9D8" box="[1586,1720,1456,1558]" gridcol="8" gridrow="7" pageId="14" pageNumber="223">Qingdao Pier, China</td>
<td id="76AD3370FFD96E9B6481FA3FF899F9D8" box="[1753,1864,1456,1558]" gridcol="9" gridrow="7" pageId="14" pageNumber="223">
Purschke 
<emphasis id="B938845CFFD9916A656FFA3FF922FA2E" italics="true" pageId="14" pageNumber="223">et al.</emphasis>
(1995)
</td>
</tr>
<tr id="357C5A0CFFD96E9B62CBF9B9F899F952" box="[147,1864,1590,1692]" gridrow="8" pageId="14" pageNumber="223">
<th id="76AD3370FFD96E9B62CBF9B9FE91F952" box="[147,320,1590,1692]" gridcol="0" gridrow="8" pageId="14" pageNumber="223">
<taxonomicName id="4C4C23CDFFD9916A62CBF9B9FEE2F94C" authority="Kukenthal, 1887" authorityName="Kukenthal" authorityYear="1887" box="[147,307,1590,1666]" class="Polychaeta" family="Opheliidae" genus="Polyophthalmus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Opheliida" pageId="14" pageNumber="223" phylum="Annelida" rank="species" species="striatus">
<emphasis id="B938845CFFD9916A62CBF9B9FF0AF9A8" box="[147,304,1590,1638]" italics="true" pageId="14" pageNumber="223">Polyophthalmus striatus</emphasis>
Kükenthal, 1887
</taxonomicName>
</th>
<td id="76AD3370FFD96E9B633CF9B9FE3BF952" box="[356,490,1590,1692]" gridcol="1" gridrow="8" pageId="14" pageNumber="223">Broadly rounded</td>
<td id="76AD3370FFD96E9B607BF9B9FD7FF952" box="[547,686,1590,1692]" gridcol="2" gridrow="8" pageId="14" pageNumber="223">Ch. 5–16 (same sized)</td>
<td id="76AD3370FFD96E9B6093F9B9FCF6F952" box="[715,807,1590,1692]" gridcol="3" gridrow="8" pageId="14" pageNumber="223">Not described</td>
<td id="76AD3370FFD96E9B6104F9B9FB90F952" box="[860,1089,1590,1692]" gridcol="4" gridrow="8" pageId="14" pageNumber="223">Round lobe more distinct on posterior segments, lacking dorsal and ventral cirri</td>
<td id="76AD3370FFD96E9B663DF9B9FB17F952" box="[1125,1222,1590,1692]" gridcol="5" gridrow="8" pageId="14" pageNumber="223">Short, cylinder</td>
<td id="76AD3370FFD96E9B66BEF9B9FABEF952" box="[1254,1391,1590,1692]" gridcol="6" gridrow="8" pageId="14" pageNumber="223">Digitiform, 4 pairs</td>
<td id="76AD3370FFD96E9B67D4F9B9F9C4F952" box="[1420,1557,1590,1692]" gridcol="7" gridrow="8" pageId="14" pageNumber="223">Present, thicker than marginal papillae</td>
<td id="76AD3370FFD96E9B646AF9B9F969F952" box="[1586,1720,1590,1692]" gridcol="8" gridrow="8" pageId="14" pageNumber="223">Hong Kong</td>
<td id="76AD3370FFD96E9B6481F9B9F899F952" box="[1753,1864,1590,1692]" gridcol="9" gridrow="8" pageId="14" pageNumber="223">Kükenthal (1887)</td>
</tr>
</table>
</paragraph>
<paragraph id="8BF3584EFFD8916B629FFF18FA57FEAA" blockId="15.[151,1437,151,357]" pageId="15" pageNumber="224">
The structures of the anal cone are also helpful on separating species of this genus. 
<taxonomicName id="4C4C23CDFFD8916B6601FF18FAA4FF7E" authority="Magalhães &amp; Rizzo &amp; Bailey-Brock, 2019" authorityName="Magalhães &amp; Rizzo &amp; Bailey-Brock" authorityYear="2019" box="[1113,1397,151,176]" class="Polychaeta" family="Opheliidae" genus="Polyophthalmus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Opheliida" pageId="15" pageNumber="224" phylum="Annelida" rank="species" species="mauliola" status="sp. nov.">
<emphasis id="B938845CFFD8916B6601FF18FAA4FF7E" box="[1113,1397,151,176]" italics="true" pageId="15" pageNumber="224">Polyophthalmus mauliola</emphasis>
</taxonomicName>
<emphasis id="B938845CFFD8916B6723FF18FF16FF1B" bold="true" pageId="15" pageNumber="224">
<taxonomicNameLabel id="A20B3927FFD8916B6723FF18FF16FF1B" pageId="15" pageNumber="224" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
has a short anal cone with a pair of basal and elongate papillae clearly distinct from the eight short and digitate marginal papillae. 
<taxonomicName id="4C4C23CDFFD8916B633BFF50FDADFF36" authorityName="Grube" authorityYear="1869" box="[355,636,223,248]" class="Polychaeta" family="Opheliidae" genus="Polyophthalmus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Opheliida" pageId="15" pageNumber="224" phylum="Annelida" rank="species" species="australis">
<emphasis id="B938845CFFD8916B633BFF50FDADFF36" box="[355,636,223,248]" italics="true" pageId="15" pageNumber="224">Polyophthalmus australis</emphasis>
</taxonomicName>
and 
<taxonomicName id="4C4C23CDFFD8916B60E8FF6EFCCFFF36" authorityName="Kukenthal" authorityYear="1887" box="[688,798,225,248]" class="Polychaeta" family="Opheliidae" genus="Polyophthalmus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Opheliida" pageId="15" pageNumber="224" phylum="Annelida" rank="species" species="striatus">
<emphasis id="B938845CFFD8916B60E8FF6EFCCFFF36" box="[688,798,225,248]" italics="true" pageId="15" pageNumber="224">P. striatus</emphasis>
</taxonomicName>
were also described with a pair of basal papillae that were larger than eight (
<taxonomicName id="4C4C23CDFFD8916B6306FE8AFE1DFED2" authorityName="Kukenthal" authorityYear="1887" box="[350,460,261,284]" class="Polychaeta" family="Opheliidae" genus="Polyophthalmus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Opheliida" pageId="15" pageNumber="224" phylum="Annelida" rank="species" species="striatus">
<emphasis id="B938845CFFD8916B6306FE8AFE1DFED2" box="[350,460,261,284]" italics="true" pageId="15" pageNumber="224">P. striatus</emphasis>
</taxonomicName>
) or nine (
<taxonomicName id="4C4C23CDFFD8916B6066FE8AFD6BFED3" authorityName="Grube" authorityYear="1869" box="[574,698,260,285]" class="Polychaeta" family="Opheliidae" genus="Polyophthalmus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Opheliida" pageId="15" pageNumber="224" phylum="Annelida" rank="species" species="australis">
<emphasis id="B938845CFFD8916B6066FE8AFD6BFED3" box="[574,698,260,285]" italics="true" pageId="15" pageNumber="224">P. australis</emphasis>
</taxonomicName>
) marginal papillae; no illustrations were provided for 
<taxonomicName id="4C4C23CDFFD8916B6741FE8AFA44FED3" authorityName="Grube" authorityYear="1869" box="[1305,1429,260,285]" class="Polychaeta" family="Opheliidae" genus="Polyophthalmus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Opheliida" pageId="15" pageNumber="224" phylum="Annelida" rank="species" species="australis">
<emphasis id="B938845CFFD8916B6741FE8AFA44FED3" box="[1305,1429,260,285]" italics="true" pageId="15" pageNumber="224">P. australis</emphasis>
</taxonomicName>
. 
<taxonomicName id="4C4C23CDFFD8916B62CFFEA8FE44FE8E" baseAuthorityName="Dujardin" baseAuthorityYear="1839" box="[151,405,295,320]" class="Polychaeta" family="Opheliidae" genus="Polyophthalmus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Opheliida" pageId="15" pageNumber="224" phylum="Annelida" rank="species" species="pictus">
<emphasis id="B938845CFFD8916B62CFFEA8FE44FE8E" box="[151,405,295,320]" italics="true" pageId="15" pageNumber="224">Polyophthalmus pictus</emphasis>
</taxonomicName>
and 
<taxonomicName id="4C4C23CDFFD8916B6391FEA6FDA5FE8E" authorityName="Purschke, Ding &amp; Muller" authorityYear="1995" box="[457,628,295,320]" class="Polychaeta" family="Opheliidae" genus="Polyophthalmus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Opheliida" pageId="15" pageNumber="224" phylum="Annelida" rank="species" species="qingdaoensis">
<emphasis id="B938845CFFD8916B6391FEA6FDA5FE8E" box="[457,628,295,320]" italics="true" pageId="15" pageNumber="224">P. qingdaoensis</emphasis>
</taxonomicName>
lacked distinct basal anal papillae and both species have the dorsal pair of marginal papillae larger than ventral ones; 
<taxonomicName id="4C4C23CDFFD8916B6034FEC2FCC6FEAB" authorityName="Purschke, Ding &amp; Muller" authorityYear="1995" box="[620,791,332,357]" class="Polychaeta" family="Opheliidae" genus="Polyophthalmus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Opheliida" pageId="15" pageNumber="224" phylum="Annelida" rank="species" species="qingdaoensis">
<emphasis id="B938845CFFD8916B6034FEC2FCC6FEAB" box="[620,791,332,357]" italics="true" pageId="15" pageNumber="224">P. qingdaoensis</emphasis>
</taxonomicName>
was also described with a dorsal notch on the anal cone.
</paragraph>
</subSubSection>
<caption id="DF3308C6FFD8916B62CFF8D0FD78F87C" httpUri="https://zenodo.org/record/2624309/files/figure.png" pageId="15" pageNumber="224" startId="15.[151,250,1887,1909]" targetBox="[253,1294,443,1847]" targetPageId="15">
<paragraph id="8BF3584EFFD8916B62CFF8D0FD78F87C" blockId="15.[151,1436,1887,1970]" pageId="15" pageNumber="224">
<emphasis id="B938845CFFD8916B62CFF8D0FEC5F8BB" bold="true" box="[151,276,1887,1909]" pageId="15" pageNumber="224">FIGURE 8.</emphasis>
<taxonomicName id="4C4C23CDFFD8916B6343F8D0FDF0F8BB" authority="Magalhães &amp; Rizzo &amp; Bailey-Brock, 2019" authorityName="Magalhães &amp; Rizzo &amp; Bailey-Brock" authorityYear="2019" box="[283,545,1887,1909]" class="Polychaeta" family="Opheliidae" genus="Polyophthalmus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Opheliida" pageId="15" pageNumber="224" phylum="Annelida" rank="species" species="mauliola" status="sp. nov.">
<emphasis id="B938845CFFD8916B6343F8D0FDF0F8BB" box="[283,545,1887,1909]" italics="true" pageId="15" pageNumber="224">Polyophthalmus mauliola</emphasis>
</taxonomicName>
<emphasis id="B938845CFFD8916B6071F8D0FDAAF8BB" bold="true" box="[553,635,1887,1909]" pageId="15" pageNumber="224">
<taxonomicNameLabel id="A20B3927FFD8916B6071F8D0FDAAF8BB" box="[553,635,1887,1909]" pageId="15" pageNumber="224" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
from Mamala Bay, Hawaii: A, complete specimen in dorsal view; B, anterior end in lateral view; C, anterior end in dorsal view; D, parapodium of the posterior end; E, posterior end with pygidium in lateral view; F, posterior end with pygidium in dorsal view.
</paragraph>
</caption>
<subSubSection id="C3560BC5FFC79174629FFF18FA85FF36" pageId="16" pageNumber="225" type="etymology">
<paragraph id="8BF3584EFFC79174629FFF18FA85FF36" blockId="16.[151,1436,151,357]" pageId="16" pageNumber="225">
<emphasis id="B938845CFFC79174629FFF18FE9FFF7E" bold="true" box="[199,334,151,176]" pageId="16" pageNumber="225">Etymology.</emphasis>
The epithet of this species refers to the Hawaiian term 
<emphasis id="B938845CFFC7917461E2FF18FBFBFF7E" box="[954,1066,151,176]" italics="true" pageId="16" pageNumber="225">Mauli-ola</emphasis>
meaning breath of life, power of healing and also a God of health. 
<emphasis id="B938845CFFC791746052FF33FDA0FF1B" box="[522,625,188,213]" italics="true" pageId="16" pageNumber="225">Mauliola</emphasis>
was also the name of Sand Island, a small island near the 
<typeStatus id="54F7E6ECFFC7917466ACFF32FAF5FF1B" box="[1268,1316,189,213]" pageId="16" pageNumber="225">type</typeStatus>
locality of this species in Mamala Bay that was used as a quarantine station in 1869 (also known as Quarantine Island).
</paragraph>
</subSubSection>
<subSubSection id="C3560BC5FFC79174629FFE8BFE5EFEAB" pageId="16" pageNumber="225" type="distribution">
<paragraph id="8BF3584EFFC79174629FFE8BFE5EFEAB" blockId="16.[151,1436,151,357]" pageId="16" pageNumber="225">
<emphasis id="B938845CFFC79174629FFE8BFEB1FED3" bold="true" box="[199,352,260,285]" pageId="16" pageNumber="225">Distribution.</emphasis>
The 
<typeStatus id="54F7E6ECFFC7917463C6FE8AFE1FFED3" box="[414,462,261,285]" pageId="16" pageNumber="225">type</typeStatus>
locality is in Mamala Bay, south shore of Oahu, Hawaii. This species has also been collected in Waianae, Kaneohe Bay, Waikiki, Pearl Harbor (Oahu) and Halape (Big Island) from 
<quantity id="4CB4F5ABFFC79174668FFEA8FAEBFEF1" box="[1239,1338,295,320]" metricMagnitude="1" metricUnit="m" metricValue="4.15" metricValueMax="5.6" metricValueMin="2.7" pageId="16" pageNumber="225" unit="m" value="41.5" valueMax="56.0" valueMin="27.0">27–56 m</quantity>
depth in sand and coral rubble.
</paragraph>
</subSubSection>
</treatment>
</document>