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<document id="CC28A3C78E7F531F1F06810BE51C23DB" ID="10.11646/zootaxa.4258.3.5SLASH10912" ID-CLB-Dataset="33142" ID-DOI="10.11646/zootaxa.4258.3.5" ID-GBIF-Dataset="240971f9-b053-4fd8-9612-ce7f71e9661a" ID-ISSN="1175-5326" ID-Zenodo-Dep="569773" ID-ZooBank="C8AB4963-75A9-49E1-883F-28740F91EC44" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1493366664018" checkinUser="plazi" docAuthor="Purchart, Luboš" docDate="2017" docId="03E55A0EFFBEF941FF583AAEFC0456B3" docLanguage="en" docName="zootaxa.4258.3.5SLASH10912.pdf" docOrigin="Zootaxa 4258 (3)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Adelostoma Purchart, 2017, sp. nov." docType="treatment" docVersion="7" lastPageNumber="285" masterDocId="FFDC2276FFBCF945FFCF3A39FF955436" masterDocTitle="Revision of the genus Adelostoma (Coleoptera: Tenebrionidae). Part 2: A new subgenus and species from Oman" masterLastPageNumber="286" masterPageNumber="281" pageNumber="283" updateTime="1698439146008" updateUser="ExternalLinkService">
<mods:mods id="12BE0E5A42191FC51960EB9F9C0DBDF8" xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo id="3425D39518BCB942E05879A5DC71EB09">
<mods:title id="37F254ABAA18925CC3206AED463FAEF5">Revision of the genus Adelostoma (Coleoptera: Tenebrionidae). Part 2: A new subgenus and species from Oman</mods:title>
</mods:titleInfo>
<mods:name id="D73B7C15F31A7FCCF8100CF64A74FBF3" type="personal">
<mods:role id="8E099604B9F2ED2BF6B45384A0DB2A49">
<mods:roleTerm id="D254554B92B0DDEECFBD7E2941A1315A">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="B5373083A58794452094A92BD1F1FF7C">Purchart, Luboš</mods:namePart>
</mods:name>
<mods:typeOfResource id="5747327BB5E758F768A9CB7274D9C351">text</mods:typeOfResource>
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<mods:titleInfo id="86E2962570787B955D4E5FC6877B3FBD">
<mods:title id="2442E4F3F016B85ADB22B963B86D6C8C">Zootaxa</mods:title>
</mods:titleInfo>
<mods:part id="2DFE86BA7C191EED721ADC83C856DDE0">
<mods:date id="5BF65B10E948A82170876BC3F227F925">2017</mods:date>
<mods:detail id="2E9A390A3EC2627F0E6BE00FD88A1A06" type="volume">
<mods:number id="98DF2CBBEB78D0E25DE2114FC8FA4BA6">4258</mods:number>
</mods:detail>
<mods:detail id="9DF56558379DAFE936D1EE68FF41C63A" type="issue">
<mods:number id="39EBE034A97D108B7635AB81570A6AE4">3</mods:number>
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<mods:start id="1B0C485FB7E4C9E1BFAD6E5F7E07C602">281</mods:start>
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<mods:classification id="83369C8164DF10A91272B3CD1B7C42BE">journal article</mods:classification>
<mods:identifier id="225687E126593882CB6B34B0A8836760" type="CLB-Dataset">33142</mods:identifier>
<mods:identifier id="DEF03C359A4F7F2C4837AA594594B64A" type="DOI">10.11646/zootaxa.4258.3.5</mods:identifier>
<mods:identifier id="CCB5F26253EA34A14C46E34782F3651D" type="GBIF-Dataset">240971f9-b053-4fd8-9612-ce7f71e9661a</mods:identifier>
<mods:identifier id="705D948B9AF1B212B594846F713695C9" type="ISSN">1175-5326</mods:identifier>
<mods:identifier id="DBA0CA4840C9A6F904276F22C9F3E650" type="Zenodo-Dep">569773</mods:identifier>
<mods:identifier id="5DCDDB6CABB6A787E61C2D0EA28EEB50" type="ZooBank">C8AB4963-75A9-49E1-883F-28740F91EC44</mods:identifier>
</mods:mods>
<treatment id="03E55A0EFFBEF941FF583AAEFC0456B3" ID-DOI="http://doi.org/10.5281/zenodo.6038407" ID-GBIF-Taxon="128810453" ID-Zenodo-Dep="6038407" LSID="urn:lsid:plazi:treatment:03E55A0EFFBEF941FF583AAEFC0456B3" httpUri="http://treatment.plazi.org/id/03E55A0EFFBEF941FF583AAEFC0456B3" lastPageId="4" lastPageNumber="285" pageId="2" pageNumber="283">
<subSubSection id="C356B893FFBEF947FF583AAEFE8754E5" pageId="2" pageNumber="283" type="nomenclature">
<paragraph id="8BF3EB18FFBEF947FF583AAEFC0F5484" blockId="2.[151,922,151,211]" box="[151,922,151,178]" pageId="2" pageNumber="283">
<heading id="D0BB5C74FFBEF947FF583AAEFC0F5484" bold="true" box="[151,922,151,178]" fontSize="11" level="1" pageId="2" pageNumber="283" reason="1">
<emphasis id="B938370AFFBEF947FF583AAEFC0F5484" bold="true" box="[151,922,151,178]" pageId="2" pageNumber="283">
<taxonomicName id="4C4C909BFFBEF947FF583AAEFEB05487" box="[151,293,151,177]" class="Insecta" family="Tenebrionidae" genus="Adelostoma" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="283" phylum="Arthropoda" rank="genus">
<emphasis id="B938370AFFBEF947FF583AAEFEB05487" bold="true" box="[151,293,151,177]" italics="true" pageId="2" pageNumber="283">Adelostoma</emphasis>
</taxonomicName>
(
<emphasis id="B938370AFFBEF947FEFA3AAEFE6F5487" bold="true" box="[309,506,151,177]" italics="true" pageId="2" pageNumber="283">Omandelostoma</emphasis>
subgen. nov.)
<taxonomicName id="4C4C909BFFBEF947FD623AAEFCA35487" box="[685,822,151,177]" class="Insecta" family="Tenebrionidae" genus="Adelostoma" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="283" phylum="Arthropoda" rank="species" species="muqalensis" status="sp. nov.">
<emphasis id="B938370AFFBEF947FD623AAEFCA35487" bold="true" box="[685,822,151,177]" italics="true" pageId="2" pageNumber="283">muqalensis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A20B8A71FFBEF947FCF23AA1FC0F5484" box="[829,922,152,178]" pageId="2" pageNumber="283" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
<paragraph id="8BF3EB18FFBEF947FF583A83FE8754E5" blockId="2.[151,922,151,211]" box="[151,274,186,211]" pageId="2" pageNumber="283">
(
<figureCitation id="1377F79DFFBEF947FF503A83FE9D54E5" box="[159,264,186,211]" captionStart="FIGURES 1 – 5" captionStartId="3.[151,264,1834,1857]" captionTargetBox="[156,1430,301,1813]" captionTargetId="figure@3.[156,1430,301,1813]" captionTargetPageId="3" captionText="FIGURES 1 – 5. Adelostoma muqalensis sp. nov., holotype. 1 — habitus, dorsal view; 2 — habitus lateral view; 3 — right antenna; 4 — head (white arrow indicates midlongitudinal carina on head); 5 — pronotum (white arrows indicate midlongitudinal carinae). Scale bars: Figs. 1, 2 = 2 mm; fig. 3 = 1 mm; figs. 4,5 = 0.5 mm." httpUri="https://zenodo.org/record/569774/files/figure.png" pageId="2" pageNumber="283">Figs. 1–5</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="C356B893FFBEF947FF583B3BFD64552C" box="[151,753,258,283]" pageId="2" pageNumber="283" type="materials_examined">
<paragraph id="8BF3EB18FFBEF947FF583B3BFD64552C" blockId="2.[151,1437,258,2011]" box="[151,753,258,283]" pageId="2" pageNumber="283">
<materialsCitation id="3B24E145FFBEF947FF583B3BFD64552C" ID-GBIF-Occurrence="1499315724" box="[151,753,258,283]" country="Oman" location="Wadi Bani Khalid" municipality="Muqal" pageId="2" pageNumber="283" specimenCount="1" typeStatus="holotype">
<emphasis id="B938370AFFBEF947FF583B3BFEA0552D" bold="true" box="[151,309,258,283]" pageId="2" pageNumber="283">
<typeStatus id="54F755BAFFBEF947FF583B3BFF47552D" box="[151,210,258,283]" pageId="2" pageNumber="283">Type</typeStatus>
locality.
</emphasis>
<collectingCountry id="F35BAB88FFBEF947FEF33B3AFE16552D" box="[316,387,259,283]" name="Oman" pageId="2" pageNumber="283">Oman</collectingCountry>
,
<location id="8E93BDC3FFBEF947FE413B3BFDCE552D" LSID="urn:lsid:plazi:treatment:03E55A0EFFBEF941FF583AAEFC0456B3:8E93BDC3FFBEF947FE413B3BFDCE552D" box="[398,603,258,283]" country="Oman" municipality="Muqal" name="Wadi Bani Khalid" pageId="2" pageNumber="283">Wadi Bani Khalid</location>
,
<collectingMunicipality id="6B977162FFBEF947FDA93B3BFD3A552D" box="[614,687,258,283]" pageId="2" pageNumber="283">Muqal</collectingMunicipality>
cave.
</materialsCitation>
</paragraph>
</subSubSection>
<subSubSection id="C356B893FFBEF947FF083B1FFB0D5555" pageId="2" pageNumber="283" type="discussion">
<paragraph id="8BF3EB18FFBEF947FF083B1FFB0D5555" blockId="2.[151,1437,258,2011]" pageId="2" pageNumber="283">
<materialsCitation id="3B24E145FFBEF947FF083B1FFB0D5555" ID-GBIF-Occurrence="1499315725" country="Oman" location="Ash Sharqiyah South Governorate" pageId="2" pageNumber="283" specimenCount="1" stateProvince="Ash Sharqiyah South Governorate">
<emphasis id="B938370AFFBEF947FF083B1FFEBC5509" bold="true" box="[199,297,294,319]" pageId="2" pageNumber="283">Remark</emphasis>
. The locality label mentions Munkal cave (“Munkal-Höhle”) which is an incorrect transcription of Muqal cave located in Wadi Bani Khalid in
<collectingRegion id="498825FAFFBEF947FD4C3B73FB925555" box="[643,1031,330,355]" country="Oman" name="Janub ash Sharqiyah" pageId="2" pageNumber="283">
<collectingRegion id="498825FAFFBEF947FD4C3B73FCBC5555" box="[643,809,330,355]" country="Saudi Arabia" name="Ash Sharqiyah" pageId="2" pageNumber="283">
Ash
<collectingRegion id="498825FAFFBEF947FD783B73FCBC5555" box="[695,809,330,355]" inferredBySuffix="true" pageId="2" pageNumber="283">Sharqiyah</collectingRegion>
</collectingRegion>
<collectingRegion id="498825FAFFBEF947FCFF3B73FB925555" box="[816,1031,330,355]" inferredBySuffix="true" pageId="2" pageNumber="283">South Governorate</collectingRegion>
</collectingRegion>
, west
<collectingCountry id="F35BAB88FFBEF947FB833B72FB015555" box="[1100,1172,331,355]" name="Oman" pageId="2" pageNumber="283">Oman</collectingCountry>
.
</materialsCitation>
</paragraph>
</subSubSection>
<subSubSection id="C356B893FFBEF947FF083B57FEA755F9" pageId="2" pageNumber="283" type="materials_examined">
<paragraph id="8BF3EB18FFBEF947FF083B57FEA755F9" blockId="2.[151,1437,258,2011]" pageId="2" pageNumber="283">
<materialsCitation id="3B24E145FFBEF947FF083B57FEA755F9" ID-GBIF-Occurrence="1499315726" collectionCode="NHMW" country="Oman" determinerName="L. Purchart" elevation="700" location="Bani Khalid" municipality="Munkal-Hohle" pageId="2" pageNumber="283" specimenCount="3" typeStatus="holotype">
<emphasis id="B938370AFFBEF947FF083B57FEED55B1" bold="true" box="[199,376,366,391]" pageId="2" pageNumber="283">
<typeStatus id="54F755BAFFBEF947FF083B57FE9755B1" box="[199,258,366,391]" pageId="2" pageNumber="283">Type</typeStatus>
material.
</emphasis>
<typeStatus id="54F755BAFFBEF947FE4E3B56FE6E55B3" box="[385,507,367,390]" pageId="2" pageNumber="283" type="holotype">HOLOTYPE</typeStatus>
[unsexed] (
<collectionCode id="ED5D73DDFFBEF947FD4A3B56FD7255B1" box="[645,743,367,391]" country="Austria" httpUri="http://grbio.org/cool/91g3-0mnw" name="Naturhistorisches Museum, Wien" pageId="2" pageNumber="283">NHMW</collectionCode>
), labelled:
<collectingCountry id="F35BAB88FFBEF947FCA83B56FC2A55B1" box="[871,959,367,391]" name="Oman" pageId="2" pageNumber="283">OMAN</collectingCountry>
, W,
<location id="8E93BDC3FFBEF947FC393B57FB1455B1" LSID="urn:lsid:plazi:treatment:03E55A0EFFBEF941FF583AAEFC0456B3:8E93BDC3FFBEF947FC393B57FB1455B1" box="[1014,1153,366,391]" country="Oman" municipality="Munkal-Hohle" name="Bani Khalid" pageId="2" pageNumber="283">Bani Khalid</location>
/
<collectingMunicipality id="6B977162FFBEF947FB533B57FAD455B1" box="[1180,1345,366,391]" pageId="2" pageNumber="283">Munkal-Höhle</collectingMunicipality>
/
<elevation id="00610C2BFFBEF947FA933B56FA0955B1" box="[1372,1436,367,391]" metricMagnitude="2" metricUnit="m" metricValue="7.0" pageId="2" pageNumber="283" unit="m" value="700.0">
<quantity id="4CB446FDFFBEF947FA933B56FA0955B1" box="[1372,1436,367,391]" metricMagnitude="2" metricUnit="m" metricValue="7.0" pageId="2" pageNumber="283" unit="m" value="700.0">700m</quantity>
</elevation>
a.s.l / 11.10.[20]06 [white, printed] //
<typeStatus id="54F755BAFFBEF947FD833BAAFD67559D" box="[588,754,403,427]" pageId="2" pageNumber="283" type="holotype">HOLOTYPUS</typeStatus>
/
<emphasis id="B938370AFFBEF947FCC23BAAFBBF559D" box="[781,1066,403,427]" italics="true" pageId="2" pageNumber="283">
<taxonomicName id="4C4C909BFFBEF947FCC23BAAFC04559D" box="[781,913,403,427]" class="Insecta" family="Tenebrionidae" genus="Adelostoma" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="283" phylum="Arthropoda" rank="genus">Adelostoma</taxonomicName>
/
<taxonomicName id="4C4C909BFFBEF947FC633BAAFBBF559D" box="[940,1066,403,427]" class="Insecta" family="Tenebrionidae" genus="Adelostoma" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="283" phylum="Arthropoda" rank="species" species="muqalensis" status="sp. nov.">muqalensis</taxonomicName>
</emphasis>
<taxonomicNameLabel id="A20B8A71FFBEF947FBFB3BAAFB18559C" box="[1076,1165,403,426]" pageId="2" pageNumber="283" rank="species">sp. nov.</taxonomicNameLabel>
/ det.
<determinerName id="7E2F07DCFFBEF947FB153BAAFACE559D" box="[1242,1371,402,427]" pageId="2" pageNumber="283">L. Purchart</determinerName>
2016 [red, printed].
</materialsCitation>
</paragraph>
</subSubSection>
<subSubSection id="C356B893FFBEF947FF083BE3FC9055C5" box="[199,773,474,499]" pageId="2" pageNumber="283" type="discussion">
<paragraph id="8BF3EB18FFBEF947FF083BE3FC9055C5" blockId="2.[151,1437,258,2011]" box="[199,773,474,499]" pageId="2" pageNumber="283">
<emphasis id="B938370AFFBEF947FF083BE3FEA555C5" bold="true" box="[199,304,474,499]" pageId="2" pageNumber="283">Remark.</emphasis>
Collector of the new species is unknown.
</paragraph>
</subSubSection>
<subSubSection id="C356B893FFBEF947FF083BC7FB035795" pageId="2" pageNumber="283" type="diagnosis">
<paragraph id="8BF3EB18FFBEF947FF083BC7FB035795" blockId="2.[151,1437,258,2011]" pageId="2" pageNumber="283">
<emphasis id="B938370AFFBEF947FF083BC7FE585621" bold="true" box="[199,461,510,535]" pageId="2" pageNumber="283">Differential diagnosis.</emphasis>
<taxonomicName id="4C4C909BFFBEF947FE1C3BC6FDFA5620" box="[467,623,510,534]" class="Insecta" family="Tenebrionidae" genus="Adelostoma" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="283" phylum="Arthropoda" rank="species" species="muqalensis" status="sp. nov.">
<emphasis id="B938370AFFBEF947FE1C3BC6FDFA5620" box="[467,623,510,534]" italics="true" pageId="2" pageNumber="283">A. muqalensis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A20B8A71FFBEF947FDB93BC7FD5A5621" box="[630,719,510,535]" pageId="2" pageNumber="283" rank="species">
<emphasis id="B938370AFFBEF947FDB93BC7FD5A5621" bold="true" box="[630,719,510,535]" pageId="2" pageNumber="283">sp. nov.</emphasis>
</taxonomicNameLabel>
possesses an epipleural carina which is only developed apically, a feature which is shared with all members of the subgenus
<emphasis id="B938370AFFBEF947FCFB381AFC79560D" box="[820,1004,547,571]" italics="true" pageId="2" pageNumber="283">Zarudnionymus.</emphasis>
However, it can be distinguished from those species mainly by antennomeres IV-IX being longer than wide and especially by strongly elongated antennomere III being three times longer than antennomere II (in
<emphasis id="B938370AFFBEF947FCB23852FBB856B5" box="[893,1069,619,643]" italics="true" pageId="2" pageNumber="283">Zarudnionymus</emphasis>
species antennomeres IV-IX are transverse and antennomere III is at maximum twice as long as antennomere II). Furthermore, the new species differs from the rest of its congeners by two midlongitudinal carinae on the disc of the pronotum which are developed only at the anterior fourth of pronotum, and by a large transverse impression in the middle of the pronotum (two pronotal carinae completely developed and no impression on pronotum present in the rest of the genus). Spaces between coarse and deep punctures on reflected portion of elytra are larger than puncture diameter in
<emphasis id="B938370AFFBEF947FF7C397AFEFF576D" box="[179,362,835,859]" italics="true" pageId="2" pageNumber="283">Omandelostoma</emphasis>
subgen. nov., while in all
<emphasis id="B938370AFFBEF947FD41397AFCAB576D" box="[654,830,835,859]" italics="true" pageId="2" pageNumber="283">Zarudnionymus</emphasis>
species the spaces are smaller than puncture diameter. Last but not least, the new species differs by absence of long erect or semi-erect hairs on dorsal body surface (members of the subgenus
<emphasis id="B938370AFFBEF947FE0B39B2FDE15795" box="[452,628,907,931]" italics="true" pageId="2" pageNumber="283">Zarudnionymus</emphasis>
are covered with long erect or semi-erect setae).
</paragraph>
</subSubSection>
<subSubSection id="C356B893FFBEF941FF083997FD2F55B1" lastPageId="4" lastPageNumber="285" pageId="2" pageNumber="283" type="description">
<paragraph id="8BF3EB18FFBEF947FF083997FC5C57DD" blockId="2.[151,1437,258,2011]" pageId="2" pageNumber="283">
<emphasis id="B938370AFFBEF947FF083997FECD57F0" bold="true" box="[199,344,942,967]" pageId="2" pageNumber="283">Description.</emphasis>
Body length
<quantity id="4CB446FDFFBEF947FE363997FDC157F0" box="[505,596,942,967]" metricMagnitude="-3" metricUnit="m" metricValue="7.6" pageId="2" pageNumber="283" unit="mm" value="7.6">7.6 mm</quantity>
, width
<quantity id="4CB446FDFFBEF947FD633996FC9D57F0" box="[684,776,943,967]" metricMagnitude="-3" metricUnit="m" metricValue="3.4" pageId="2" pageNumber="283" unit="mm" value="3.4">3.4 mm</quantity>
. Body narrow, elongate and glabrous. Dorsal and ventral sides of body pale brown (perhaps teneral specimen) and dull (
<figureCitation id="1377F79DFFBEF947FC9939EBFC2E57DD" box="[854,955,978,1003]" captionStart="FIGURES 1 – 5" captionStartId="3.[151,264,1834,1857]" captionTargetBox="[156,1430,301,1813]" captionTargetId="figure@3.[156,1430,301,1813]" captionTargetPageId="3" captionText="FIGURES 1 – 5. Adelostoma muqalensis sp. nov., holotype. 1 — habitus, dorsal view; 2 — habitus lateral view; 3 — right antenna; 4 — head (white arrow indicates midlongitudinal carina on head); 5 — pronotum (white arrows indicate midlongitudinal carinae). Scale bars: Figs. 1, 2 = 2 mm; fig. 3 = 1 mm; figs. 4,5 = 0.5 mm." httpUri="https://zenodo.org/record/569774/files/figure.png" pageId="2" pageNumber="283">Figs 1, 2</figureCitation>
).
</paragraph>
<paragraph id="8BF3EB18FFBEF947FF0839CFFA8651AD" blockId="2.[151,1437,258,2011]" pageId="2" pageNumber="283">
<emphasis id="B938370AFFBEF947FF0839CFFE915038" box="[199,260,1014,1038]" italics="true" pageId="2" pageNumber="283">Head</emphasis>
0.95 times as long as wide, 0.6 times as wide as pronotum (
<figureCitation id="1377F79DFFBEF947FC5239CFFC745039" box="[925,993,1014,1039]" captionStart="FIGURES 1 – 5" captionStartId="3.[151,264,1834,1857]" captionTargetBox="[156,1430,301,1813]" captionTargetId="figure@3.[156,1430,301,1813]" captionTargetPageId="3" captionText="FIGURES 1 – 5. Adelostoma muqalensis sp. nov., holotype. 1 — habitus, dorsal view; 2 — habitus lateral view; 3 — right antenna; 4 — head (white arrow indicates midlongitudinal carina on head); 5 — pronotum (white arrows indicate midlongitudinal carinae). Scale bars: Figs. 1, 2 = 2 mm; fig. 3 = 1 mm; figs. 4,5 = 0.5 mm." httpUri="https://zenodo.org/record/569774/files/figure.png" pageId="2" pageNumber="283">Fig. 4</figureCitation>
). Widest at genae. Shagreened, covered sparsely with short whitish hairs on genae and epistoma. Length of hairs approximately as long as two eye facets combined. Surface of head with net-like sculpturing composed of concave granules which are approximately 2–3 times larger than one eye facet. Genae strongly developed. Ocular carinae sharp, moderately high (in lateral view), parallel. Midlongitudinal carinae incomplete, short, starting before anterior part of eyes and reaching anterior margin of clypeus, approximately as high as ocular carinae (in lateral view). Depressed area between antennal sulcus and midlongitudinal carina large and deep. Clypeus asymmetrically cut out in the middle (left side deeper than right), with two small teeth in the middle (
<figureCitation id="1377F79DFFBEF947FD753ECBFC97513D" box="[698,770,1266,1291]" captionStart="FIGURES 1 – 5" captionStartId="3.[151,264,1834,1857]" captionTargetBox="[156,1430,301,1813]" captionTargetId="figure@3.[156,1430,301,1813]" captionTargetPageId="3" captionText="FIGURES 1 – 5. Adelostoma muqalensis sp. nov., holotype. 1 — habitus, dorsal view; 2 — habitus lateral view; 3 — right antenna; 4 — head (white arrow indicates midlongitudinal carina on head); 5 — pronotum (white arrows indicate midlongitudinal carinae). Scale bars: Figs. 1, 2 = 2 mm; fig. 3 = 1 mm; figs. 4,5 = 0.5 mm." httpUri="https://zenodo.org/record/569774/files/figure.png" pageId="2" pageNumber="283">Fig. 4</figureCitation>
). Antennae 1.66 times longer than width of head, covered with whitish hairs. Hairs approximately as long as three eye facets combined. All antennomeres longer than wide. Antennomere III strongly prolonged, approximately twice as long as antennomere IV and three times longer than antennomere II (
<figureCitation id="1377F79DFFBEF947FE9E3F67FE005141" box="[337,405,1374,1399]" captionStart="FIGURES 1 – 5" captionStartId="3.[151,264,1834,1857]" captionTargetBox="[156,1430,301,1813]" captionTargetId="figure@3.[156,1430,301,1813]" captionTargetPageId="3" captionText="FIGURES 1 – 5. Adelostoma muqalensis sp. nov., holotype. 1 — habitus, dorsal view; 2 — habitus lateral view; 3 — right antenna; 4 — head (white arrow indicates midlongitudinal carina on head); 5 — pronotum (white arrows indicate midlongitudinal carinae). Scale bars: Figs. 1, 2 = 2 mm; fig. 3 = 1 mm; figs. 4,5 = 0.5 mm." httpUri="https://zenodo.org/record/569774/files/figure.png" pageId="2" pageNumber="283">Fig. 3</figureCitation>
). Ultimate antennomere obliquely truncate and strongly widened towards apex, longer than wide, approximately one and a half times longer than penultimate antennomere. Antennal sulcus deep.
</paragraph>
<paragraph id="8BF3EB18FFBEF947FF083F9EFC75528D" blockId="2.[151,1437,258,2011]" pageId="2" pageNumber="283">
<emphasis id="B938370AFFBEF947FF083F9EFEA35188" box="[199,310,1447,1470]" italics="true" pageId="2" pageNumber="283">Pronotum</emphasis>
strongly transverse, 1.77 times wider than long, with rounded sides, strongly narrowing anteriorly, widest in posterior third (
<figureCitation id="1377F79DFFBEF947FE7B3FF3FE6C51D5" box="[436,505,1482,1507]" captionStart="FIGURES 1 – 5" captionStartId="3.[151,264,1834,1857]" captionTargetBox="[156,1430,301,1813]" captionTargetId="figure@3.[156,1430,301,1813]" captionTargetPageId="3" captionText="FIGURES 1 – 5. Adelostoma muqalensis sp. nov., holotype. 1 — habitus, dorsal view; 2 — habitus lateral view; 3 — right antenna; 4 — head (white arrow indicates midlongitudinal carina on head); 5 — pronotum (white arrows indicate midlongitudinal carinae). Scale bars: Figs. 1, 2 = 2 mm; fig. 3 = 1 mm; figs. 4,5 = 0.5 mm." httpUri="https://zenodo.org/record/569774/files/figure.png" pageId="2" pageNumber="283">Fig. 5</figureCitation>
). Glabrous, shagreened, without net-like surface, sparsely covered with very small granules which are smaller than one eye facet. Anterior fourth with two parallel, very short midlongitudinal carinae in the middle. Anterior margin of pronotum strongly emarginated. Anterior angles of pronotum more or less rounded.
<collectingRegion id="498825FAFFBEF947FEC83C0FFECC5279" box="[263,345,1590,1615]" country="Oman" name="Al Wusta" pageId="2" pageNumber="283">Central</collectingRegion>
part of pronotum with large and deep transverse impression. Lateral portions of pronotum foliaceous, distinctly bent upward. Lateral margins of pronotum slightly but more or less regularly dentate with very small teeth. Base of pronotum emarginated laterally and centrally, resulting in obtuse anterior angles and two large teeth, each situated between lateral and middle third of pronotal base.
</paragraph>
<paragraph id="8BF3EB18FFBEF946FF083CFFFEA154E3" blockId="2.[151,1437,258,2011]" lastBlockId="3.[151,1436,151,248]" lastPageId="3" lastPageNumber="284" pageId="2" pageNumber="283">
<emphasis id="B938370AFFBEF947FF083CFFFE9852E8" box="[199,269,1734,1758]" italics="true" pageId="2" pageNumber="283">Elytra</emphasis>
glabrous, shagreened, ovoid, with three longitudinal, strongly developed and relatively high keels on each elytron (
<figureCitation id="1377F79DFFBEF947FEFC3CD3FE0D5335" box="[307,408,1770,1795]" captionStart="FIGURES 1 – 5" captionStartId="3.[151,264,1834,1857]" captionTargetBox="[156,1430,301,1813]" captionTargetId="figure@3.[156,1430,301,1813]" captionTargetPageId="3" captionText="FIGURES 1 – 5. Adelostoma muqalensis sp. nov., holotype. 1 — habitus, dorsal view; 2 — habitus lateral view; 3 — right antenna; 4 — head (white arrow indicates midlongitudinal carina on head); 5 — pronotum (white arrows indicate midlongitudinal carinae). Scale bars: Figs. 1, 2 = 2 mm; fig. 3 = 1 mm; figs. 4,5 = 0.5 mm." httpUri="https://zenodo.org/record/569774/files/figure.png" pageId="2" pageNumber="283">Fig. 1, 2</figureCitation>
). Space between elytral suture and first (inner) keel irregularly deeply punctuate with 4–5 puncture interval. Space between first and second and second and third keel with 3–4 puncture intervals with punctures arranged more or less in rows. Surface between these punctures with very small tubercles smaller than one eye facet. First (inner, not elytral suture) and third (outer) keels connected anteriorly with basal margin of elytra. All three keels unconnected with each other or the apex of elytra. First carina beginning at level of metacoxae. Epipleural carina developed only apically. Reflected portion of elytra (sensu
<bibRefCitation id="EFDD96E9FFBEF947FB4D3DA7FA975381" author="Koch" box="[1154,1282,1950,1975]" pageId="2" pageNumber="283" refString="Koch, C. (1952) The Tenebrionidae of Southern Africa. VIII. Materials for a monographic study on Eurychorini (Coleoptera). Bulletin de la Societe Fouad 1 er d'Entomologie, 36, 1 - 125." type="journal article" year="1952">Koch 1952</bibRefCitation>
: 5, 7) at level of 1st and 2nd abdominal ventrites with five puncture intervals, with coarse and deep punctures arranged more or less irregularly, space between those punctures much bigger than their diameter. Apex of elytra moderately steep (lateral view).
</paragraph>
<paragraph id="8BF3EB18FFBFF946FF083AD8FBAD54CE" blockId="3.[151,1436,151,248]" box="[199,1080,223,248]" pageId="3" pageNumber="284">
<emphasis id="B938370AFFBFF946FF083AD8FF6954CE" box="[199,252,225,248]" italics="true" pageId="3" pageNumber="284">Legs</emphasis>
shagreened and covered with whittish setae, without any special features.
</paragraph>
<caption id="DF33BB90FFBFF946FF583D13FCC75348" httpUri="https://zenodo.org/record/569774/files/figure.png" pageId="3" pageNumber="284" targetBox="[156,1430,301,1813]" targetPageId="3">
<paragraph id="8BF3EB18FFBFF946FF583D13FCC75348" blockId="3.[151,1436,1834,1918]" pageId="3" pageNumber="284">
<emphasis id="B938370AFFBFF946FF583D13FEAC5376" bold="true" box="[151,313,1834,1857]" pageId="3" pageNumber="284">FIGURES 1–5.</emphasis>
<taxonomicName id="4C4C909BFFBFF946FEF03D12FDBB5376" box="[319,558,1835,1856]" class="Insecta" family="Tenebrionidae" genus="Adelostoma" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="284" phylum="Arthropoda" rank="species" species="muqalensis" status="sp. nov.">
<emphasis id="B938370AFFBFF946FEF03D12FDBB5376" box="[319,558,1835,1856]" italics="true" pageId="3" pageNumber="284">Adelostoma muqalensis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A20B8A71FFBFF946FDFB3D12FD115376" box="[564,644,1835,1856]" pageId="3" pageNumber="284" rank="species">
<emphasis id="B938370AFFBFF946FDFB3D12FD115376" bold="true" box="[564,644,1835,1856]" pageId="3" pageNumber="284">sp. nov.</emphasis>
</taxonomicNameLabel>
, holotype. 1—habitus, dorsal view; 2—habitus lateral view; 3—right antenna; 4—head (white arrow indicates midlongitudinal carina on head); 5—pronotum (white arrows indicate midlongitudinal carinae). Scale bars: Figs. 1,2 = 2mm; fig. 3=1mm; figs. 4,5=0.5mm.
</paragraph>
</caption>
<paragraph id="8BF3EB18FFB8F941FF083AA1FD2F55B1" blockId="4.[151,1437,151,645]" pageId="4" pageNumber="285">
<emphasis id="B938370AFFB8F941FF083AA1FECF5486" box="[199,346,152,176]" italics="true" pageId="4" pageNumber="285">Ventral side.</emphasis>
Mentum strongly transverse, subcordiform. Apical margin of buccal fissure lobes dentate, triangular (sensu
<bibRefCitation id="EFDD96E9FFB8F941FEAA3A85FE7F54E3" author="Koch" box="[357,490,188,213]" pageId="4" pageNumber="285" refString="Koch, C. (1952) The Tenebrionidae of Southern Africa. VIII. Materials for a monographic study on Eurychorini (Coleoptera). Bulletin de la Societe Fouad 1 er d'Entomologie, 36, 1 - 125." type="journal article" year="1952">Koch 1952</bibRefCitation>
: 15). Entire ventral side of head, pro-, meso- and metaventrite shagreened and sculptured with net-like pattern composed of concave granules. Ventral side of head covered with hairs similar to those on antennae. Prosternal process narrow, as broad as mesosternal and narrower than metasternal apophysis; subparallel, projecting behind procoxae, rounded apically (in dorsal view) and almost horizontal (in lateral view). Hypomeron and abdominal ventrites glabrous, shagreened, sparsely covered with fine granules similar to those on elytra. Each granule with recumbent, short setae.
</paragraph>
</subSubSection>
<subSubSection id="C356B893FFB8F941FF083BADFB69559B" box="[199,1276,404,429]" pageId="4" pageNumber="285" type="etymology">
<paragraph id="8BF3EB18FFB8F941FF083BADFB69559B" blockId="4.[151,1437,151,645]" box="[199,1276,404,429]" pageId="4" pageNumber="285">
<emphasis id="B938370AFFB8F941FF083BADFEDB559B" bold="true" box="[199,334,404,429]" pageId="4" pageNumber="285">Etymology.</emphasis>
The name “
<taxonomicName id="4C4C909BFFB8F941FE183BADFDC0559A" box="[471,597,404,428]" class="Insecta" family="Tenebrionidae" genus="Adelostoma" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="285" phylum="Arthropoda" rank="species" species="muqalensis">
<emphasis id="B938370AFFB8F941FE183BADFDC0559A" box="[471,597,404,428]" italics="true" pageId="4" pageNumber="285">muqalensis</emphasis>
</taxonomicName>
” refers to Muqal cave (
<collectingCountry id="F35BAB88FFB8F941FC913BADFC33559A" box="[862,934,404,428]" name="Oman" pageId="4" pageNumber="285">Oman</collectingCountry>
) where the species was found.
</paragraph>
</subSubSection>
<subSubSection id="C356B893FFB8F941FF083B8EFC375656" pageId="4" pageNumber="285" type="discussion">
<paragraph id="8BF3EB18FFB8F941FF083B8EFC375656" blockId="4.[151,1437,151,645]" pageId="4" pageNumber="285">
<emphasis id="B938370AFFB8F941FF083B8EFEA555E6" bold="true" box="[199,304,439,464]" pageId="4" pageNumber="285">Remark.</emphasis>
<emphasis id="B938370AFFB8F941FEF63B81FDBD55F9" box="[313,552,440,464]" italics="true" pageId="4" pageNumber="285">Symphochora simoni</emphasis>
(Fairmaire, 1894) and
<taxonomicName id="4C4C909BFFB8F941FCFB3B81FB6055E6" authority="Purchart, 2009" authorityName="Purchart" authorityYear="2009" box="[820,1269,439,464]" class="Insecta" family="Tenebrionidae" genus="Adelostoma" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="285" phylum="Arthropoda" rank="species" species="schawalleri">
<emphasis id="B938370AFFB8F941FCFB3B81FBD155E6" box="[820,1092,440,464]" italics="true" pageId="4" pageNumber="285">Adelostoma schawalleri</emphasis>
<bibRefCitation id="EFDD96E9FFB8F941FB823B8EFB6055E6" author="Purchart" box="[1101,1269,439,464]" pageId="4" pageNumber="285" refString="Purchart, L. (2009) Review of the genus Adelostoma Duponchel, 1827 (Coleoptera: Tenebrionidae: Adelostomini) from the Socotra Archipelago, with description of a new species. African Entomology, 17, 23 - 27." type="journal article" year="2009">Purchart, 2009</bibRefCitation>
</taxonomicName>
were the only members of the tribe Adelostomini that were considered as truly cavernicolous species so far (
<bibRefCitation id="EFDD96E9FFB8F941FB7D3BE5FAA755C3" author="Koch" box="[1202,1330,476,501]" pageId="4" pageNumber="285" refString="Koch, C. (1952) The Tenebrionidae of Southern Africa. VIII. Materials for a monographic study on Eurychorini (Coleoptera). Bulletin de la Societe Fouad 1 er d'Entomologie, 36, 1 - 125." type="journal article" year="1952">Koch 1952</bibRefCitation>
;
<bibRefCitation id="EFDD96E9FFB8F941FAF23BE5FF41562E" author="Purchart" pageId="4" pageNumber="285" refString="Purchart, L. (2009) Review of the genus Adelostoma Duponchel, 1827 (Coleoptera: Tenebrionidae: Adelostomini) from the Socotra Archipelago, with description of a new species. African Entomology, 17, 23 - 27." type="journal article" year="2009">Purchart 2009</bibRefCitation>
). According to the locality label and also due to the considerably elongated antennomeres, which is one of the typical features for cavernicolous insects,
<taxonomicName id="4C4C909BFFB8F941FD54381DFCD4560A" box="[667,833,548,572]" class="Insecta" family="Tenebrionidae" genus="Adelostoma" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="285" phylum="Arthropoda" rank="species" species="muqalensis" status="sp. nov.">
<emphasis id="B938370AFFB8F941FD54381DFCD4560A" box="[667,833,548,572]" italics="true" pageId="4" pageNumber="285">A. muqalensis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A20B8A71FFB8F941FC9E381DFC21560B" box="[849,948,548,573]" pageId="4" pageNumber="285" rank="species">
<emphasis id="B938370AFFB8F941FC9E381DFC21560B" bold="true" box="[849,948,548,573]" pageId="4" pageNumber="285">sp. nov.</emphasis>
</taxonomicNameLabel>
might be another member of the tribe Adelostomini which can be considered cavernicolous species as well.
</paragraph>
</subSubSection>
<subSubSection id="C356B893FFB8F941FF083855FC0456B3" box="[199,913,620,645]" pageId="4" pageNumber="285" type="distribution">
<paragraph id="8BF3EB18FFB8F941FF083855FC0456B3" blockId="4.[151,1437,151,645]" box="[199,913,620,645]" pageId="4" pageNumber="285">
<emphasis id="B938370AFFB8F941FF083855FEF556B3" bold="true" box="[199,352,620,645]" pageId="4" pageNumber="285">Distribution.</emphasis>
Oman—so far known from the
<typeStatus id="54F755BAFFB8F941FD083854FD6256B3" box="[711,759,621,645]" pageId="4" pageNumber="285">type</typeStatus>
locality only.
</paragraph>
</subSubSection>
</treatment>
</document>
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