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<document ID-DOI="10.25223/brad.n38.2020.a23" ID-GBIF-Dataset="69b0526e-7e3a-401d-8bea-b8a2541c1f36" ID-Zenodo-Dep="7871694" IM.bibliography_approvedBy="tatiana" IM.illustrations_approvedBy="tatiana" IM.materialsCitations_approvedBy="tatiana" IM.metadata_approvedBy="tatiana" IM.taxonomicNames_approvedBy="tatiana" IM.treatmentCitations_approvedBy="tatiana" IM.treatments_approvedBy="tatiana" checkinTime="1681981549586" checkinUser="hestersteyn" docAuthor="Thiede, Joachim, Campbell-Barker, Theo Peter, Hargreaves, Bruce J., Smith, Gideon F. &amp; Figueiredo, Estrela" docDate="2020" docId="03D2FB250D61163F7920F9C7FE02E23A" docLanguage="en" docName="Bradleya.38.247-253.pdf.imf" docOrigin="Bradleya 38" docSource="https://www.mendeley.com/catalogue/8fe3b13d-ffa9-34d8-a569-ada13d0ae5ad/" docTitle="Aloe lateritia var. lateritia Engl. var. lateritia" docType="treatment" docVersion="5" lastPageNumber="252" masterDocId="FFEB835D0D62163A794AFFD6FFA2E033" masterDocTitle="Notes on three aloes of Malawi: Aloe cam's, A. lateritia, and A. suffulta (Asphodelaceae subfam. Alooideae)" masterLastPageNumber="253" masterPageNumber="247" pageNumber="250" updateTime="1682614337913" updateUser="ExternalLinkService" zenodo-license-document="CLOSED">
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<mods:titleInfo>
<mods:title>Notes on three aloes of Malawi: Aloe cam's, A. lateritia, and A. suffulta (Asphodelaceae subfam. Alooideae)</mods:title>
</mods:titleInfo>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Thiede, Joachim</mods:namePart>
<mods:nameIdentifier type="ORCID">0000-0002-0130-8424</mods:nameIdentifier>
<mods:affiliation>Schenefelder Holt 3, 22589 Hamburg, Germany</mods:affiliation>
<mods:nameIdentifier type="email">joachim_thiede@gmx.de</mods:nameIdentifier>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Campbell-Barker, Theo Peter</mods:namePart>
<mods:affiliation>22 Faringdon Avenue, Lutterworth LE 17 4 DJ, England (email: theo. cb @ talktalk. net).</mods:affiliation>
<mods:nameIdentifier type="email">theo.cb@talktalk.net</mods:nameIdentifier>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Hargreaves, Bruce J.</mods:namePart>
<mods:affiliation>5817 Pryor Street, Bakersfield, CA 93308, USA</mods:affiliation>
<mods:nameIdentifier type="email">brucejhargreaves@hotmail.com</mods:nameIdentifier>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Smith, Gideon F.</mods:namePart>
<mods:nameIdentifier type="ORCID">0000-0002-5417-9208</mods:nameIdentifier>
<mods:affiliation>Department of Botany, P. O. Box 77000, Nelson Mandela University, Port Elizabeth, 6031 South Africa</mods:affiliation>
<mods:nameIdentifier type="email">smithgideon1@gmail.com</mods:nameIdentifier>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Figueiredo, Estrela</mods:namePart>
<mods:nameIdentifier type="ORCID">0000-0002-8511-8213</mods:nameIdentifier>
<mods:affiliation>Department of Botany, P. O. Box 77000, Nelson Mandela University, Port Elizabeth, 6031 South Africa</mods:affiliation>
<mods:nameIdentifier type="email">epnfigueiredo@gmail.com</mods:nameIdentifier>
</mods:name>
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<mods:title>Bradleya</mods:title>
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<mods:date>2020</mods:date>
<mods:detail type="pubDate">
<mods:number>2020-06-26</mods:number>
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<mods:detail type="volume">
<mods:number>38</mods:number>
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<mods:start>247</mods:start>
<mods:end>253</mods:end>
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<mods:location>
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<mods:identifier type="DOI">10.25223/brad.n38.2020.a23</mods:identifier>
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<treatment ID-DOI="http://doi.org/10.5281/zenodo.7871704" ID-Zenodo-Dep="7871704" LSID="urn:lsid:plazi:treatment:03D2FB250D61163F7920F9C7FE02E23A" httpUri="http://treatment.plazi.org/id/03D2FB250D61163F7920F9C7FE02E23A" lastPageId="5" lastPageNumber="252" pageId="3" pageNumber="250">
<subSubSection box="[106,497,1553,1576]" pageId="3" pageNumber="250" type="nomenclature">
<paragraph blockId="3.[106,666,1553,1744]" box="[106,497,1553,1576]" pageId="3" pageNumber="250">
<taxonomicName ID-CoL="5M5M6" authority="Engl. var. lateritia" box="[106,497,1553,1576]" class="Liliopsida" family="Asphodelaceae" genus="Aloe" kingdom="Plantae" order="Asparagales" pageId="3" pageNumber="250" phylum="Tracheophyta" rank="variety" species="lateritia" variety="lateritia">
<emphasis bold="true" box="[106,270,1553,1575]" italics="true" pageId="3" pageNumber="250">Aloe lateritia</emphasis>
Engl. var.
<emphasis bold="true" box="[393,497,1553,1575]" italics="true" pageId="3" pageNumber="250">lateritia</emphasis>
</taxonomicName>
</paragraph>
</subSubSection>
<subSubSection lastPageId="4" lastPageNumber="251" pageId="3" pageNumber="250" type="discussion">
<paragraph blockId="3.[106,666,1553,1744]" lastBlockId="3.[711,1270,994,1744]" pageId="3" pageNumber="250">
<taxonomicName authority="var. lateritia" box="[144,430,1582,1603]" class="Liliopsida" family="Asphodelaceae" genus="Aloe" kingdom="Plantae" order="Asparagales" pageId="3" pageNumber="250" phylum="Tracheophyta" rank="variety" species="lateritia" variety="lateritia">
<emphasis box="[144,286,1582,1603]" italics="true" pageId="3" pageNumber="250">Aloe lateritia</emphasis>
var.
<emphasis box="[341,430,1582,1603]" italics="true" pageId="3" pageNumber="250">lateritia</emphasis>
</taxonomicName>
, as presently circumscribed, is a maculate aloe documented by specimens from grasslands or rocky places in southern Kenya and Tanzania.
<taxonomicName authority="(Reynolds) S. Carter" authorityName="S. Carter" baseAuthorityName="Reynolds" class="Liliopsida" family="Asphodelaceae" genus="Aloe" kingdom="Plantae" order="Asparagales" pageId="3" pageNumber="250" phylum="Tracheophyta" rank="variety" species="lateritia" variety="graminicola">
<emphasis box="[466,613,1666,1687]" italics="true" pageId="3" pageNumber="250">Aloe lateritia</emphasis>
var.
<emphasis box="[106,242,1694,1715]" italics="true" pageId="3" pageNumber="250">graminicola</emphasis>
(Reynolds) S.Carter
</taxonomicName>
is known from Kenya only (
<bibRefCitation author="CARTER, S." box="[242,457,1722,1744]" editor="POLHILL, R. M. D." firstAuthor="Carter" journalOrPublisher="A. A. Balkema, Rotterdam" page="15 - 17" pageId="3" pageNumber="250" refId="ref4206" refString="CARTER, S. (1994). Aloaceae. In: POLHILL, R. M. D.), Flora of Tropical East Africa. A. A. Balkema, Rotterdam." title="Aloaceae" type="book" volumeTitle="Flora of Tropical East Africa" year="1994">Carter, 1994: 1517</bibRefCitation>
, pl. 1;
<bibRefCitation author="CARTER, S. &amp; LAVRANOS, J. J. &amp; NEWTON, L. E. &amp; WALKER, C. C." box="[528,870,994,1743]" etAl="et al." firstAuthor="Carter" journalOrPublisher="Kew Publishing, Royal Botanic Gardens, Kew / British Cactus &amp; Succulent Society" page="179 - 180" pageId="3" pageNumber="250" refId="ref4282" refString="CARTER, S., LAVRANOS, J. J., NEWTON, L. E. &amp; WALKER, C. C. (2011). Aloes: The definitive guide. Kew Publishing, Royal Botanic Gardens, Kew / British Cactus &amp; Succulent Society." title="Aloes: The definitive guide" type="book" year="2011">
Carter
<emphasis box="[606,652,1722,1743]" italics="true" pageId="3" pageNumber="250">et al</emphasis>
., 2011: 179180
</bibRefCitation>
). Alarmingly, for Malawi
<bibRefCitation author="MSEKANDIANA, G. &amp; MLANGENI, E." editor="GOLDING, J." firstAuthor="Msekandiana" journalOrPublisher="Southern African Botanical Diversity Network Report No" page="34" pageId="3" pageNumber="250" pagination="31 - 42" part="14" refId="ref4723" refString="MSEKANDIANA, G. &amp; MLANGENI, E. (2002). Malawi. In: GOLDING, J. (ED.), Southern African Plant Red Data Lists. Southern African Botanical Diversity Network Report No. 14: 31 - 42. SABO- NET, Pretoria." title="Malawi" type="book chapter" volumeTitle="Southern African Plant Red Data Lists" year="2002">Msekandiana &amp; Mlangeni (2002: 34)</bibRefCitation>
recorded
<taxonomicName authorityName=", Engler" authorityYear="1895" box="[1124,1270,1022,1043]" class="Liliopsida" family="Asphodelaceae" genus="Aloe" kingdom="Plantae" order="Asparagales" pageId="3" pageNumber="250" phylum="Tracheophyta" rank="species" species="lateritia">
<emphasis box="[1124,1270,1022,1043]" italics="true" pageId="3" pageNumber="250">Aloe lateritia</emphasis>
</taxonomicName>
as CR B1B2cD, i.e., Critically Endangered with a Small distribution and Decline or Fluctuation evidently based on severely fragmented locations or known to exist at one or fewer locations and continuing [to] decline in its area, extent and/or quality of habitat with fewer than 50 mature individuals.
</paragraph>
<paragraph blockId="3.[711,1270,994,1744]" pageId="3" pageNumber="250">
One of us (
<collectionCode box="[861,911,1246,1267]" pageId="3" pageNumber="250">BJH</collectionCode>
) found
<taxonomicName authority="var. lateritia" box="[991,1270,1246,1267]" class="Liliopsida" family="Asphodelaceae" genus="Aloe" kingdom="Plantae" order="Asparagales" pageId="3" pageNumber="250" phylum="Tracheophyta" rank="variety" species="lateritia" variety="lateritia">
<emphasis box="[991,1130,1246,1267]" italics="true" pageId="3" pageNumber="250">Aloe lateritia</emphasis>
var.
<emphasis box="[1183,1270,1246,1267]" italics="true" pageId="3" pageNumber="250">lateritia</emphasis>
</taxonomicName>
in northernmost Malawi in 1967 at Ipenza Village near Songwe River which forms the border between Malawi and Tanzania (in Chitipa District, Northern Province;
<figureCitation box="[825,923,1358,1380]" captionStart="Figure 1" captionStartId="1.[106,186,1283,1305]" captionTargetBox="[106,665,106,1268]" captionTargetId="figure-1255@1.[107,664,107,1267]" captionTargetPageId="1" captionText="Figure 1. Distribution of three Aloe species rare in Malawi: 1) A. canis (yellow dots, a = Senga Hills; b = inland from Domira Bay; c = Ntchisi Hills; Central Region); 2) A. lateritia var. lateritia (blue dot = near Ipenza, two records; Northern Region); and 3) A. suffulta (red dot = Mwabvi Wildlife Reserve; Southern Region). Placements of the dots are partly approximations. The map insert shows the position of Malawi (red) in Africa (map: Wikimedia Commons 2019; insert map: mapsof.net 2015)." figureDoi="http://doi.org/10.5281/zenodo.7871700" httpUri="https://zenodo.org/record/7871700/files/figure.png" pageId="3" pageNumber="250">Figure 1</figureCitation>
: 2), and revisited the locality together with Roger Royle in
<date box="[1021,1125,1386,1408]" pageId="3" pageNumber="250" value="1968-07">July 1968</date>
(
<bibRefCitation author="HARGREAVES, B. J." firstAuthor="Hargreaves" journalOrPublisher="Cactus and Succulent Journal (US)" page="92" pageId="3" pageNumber="250" pagination="86 - 104" part="Suppl. Vol. 1975" refId="ref4454" refString="HARGREAVES, B. J. (1975). Succulents of Chitipa - the muddy place. Cactus and Succulent Journal (US) Suppl. Vol. 1975: 86 - 104 [text reprinted in The Society of Malawi Journal 30 (1): 28 - 34, 1977)." title="Succulents of Chitipa - the muddy place" type="journal article" year="1975">Hargreaves, 1975: 92</bibRefCitation>
and
<figureCitation box="[893,1036,1414,1436]" captionStart-0="Figure 4" captionStart-1="Figure 5" captionStart-2="Figure 6" captionStart-3="Figure 7" captionStart-4="Figure 8" captionStartId-0="2.[726,806,1028,1050]" captionStartId-1="3.[106,185,851,873]" captionStartId-2="3.[711,791,791,813]" captionStartId-3="4.[121,201,938,960]" captionStartId-4="4.[726,805,473,495]" captionTargetBox-0="[718,1285,106,1015]" captionTargetBox-1="[106,665,106,840]" captionTargetBox-2="[711,1270,106,778]" captionTargetBox-3="[121,680,106,930]" captionTargetBox-4="[726,1285,106,461]" captionTargetId-0="figure-1878@2.[727,1284,107,1015]" captionTargetId-1="figure-2126@3.[107,664,107,839]" captionTargetId-2="figure-2279@3.[711,1269,106,777]" captionTargetId-3="figure-1553@4.[122,680,106,928]" captionTargetId-4="figure-1760@4.[727,1284,107,460]" captionTargetPageId-0="2" captionTargetPageId-1="3" captionTargetPageId-2="3" captionTargetPageId-3="4" captionTargetPageId-4="4" captionText-0="Figure 4. The specimen of Aloe canis that Ms Ruth Mthawanji received from Reverend Stewart Lane in the early-2000s photographed in Blantyre, Malawi, in 2015. Photograph: R. Mthawanji." captionText-1="Figure 5. Panicle of the specimen of Aloe canis that Ms Ruth Mthawanji received from Reverend Stewart Lane photographed in Blantyre, Malawi, in 2019. Photograph: R. Mthawanji." captionText-2="Figure 6. Aloe lateritia var. lateritia in habitat in northernmost Malawi at Ipenza Village in July 1968. The spotted leaves can reach a length of 45cm (18), but grow much longer in cultivation when irrigated throughout the year (Hargreaves, 1975: 92) (published earlier in Hargreaves, 1975: Figure 4). Photograph: Bruce J. Hargreaves." captionText-3="Figure 7. An inflorescence of Aloe lateritia var. lateritia at the locality noted in the caption of Figure 4, about 1.20m (4 feet) high (Hargreaves, 1975: 92) (published earlier in Hargreaves, 1975: Figure 5). Photograph: Bruce J. Hargreaves" captionText-4="Figure 8. Aloe suffulta in habitat in what is now the Mwabvi Wildlife Reserve (Malawi), growing on a river bank (rather likely of the Mwabvi River) in the dense undergrowth of bushes with the long, dry inflorescences literally woven through the supporting bushes and emerging through the top, ca. 1970. Drawing: Pastor Theo Peter Campbell-Barker." figureDoi-0="http://doi.org/10.5281/zenodo.7871706" figureDoi-1="http://doi.org/10.5281/zenodo.7871712" figureDoi-2="http://doi.org/10.5281/zenodo.7871708" figureDoi-3="http://doi.org/10.5281/zenodo.7871716" figureDoi-4="http://doi.org/10.5281/zenodo.7871714" httpUri-0="https://zenodo.org/record/7871706/files/figure.png" httpUri-1="https://zenodo.org/record/7871712/files/figure.png" httpUri-2="https://zenodo.org/record/7871708/files/figure.png" httpUri-3="https://zenodo.org/record/7871716/files/figure.png" httpUri-4="https://zenodo.org/record/7871714/files/figure.png" pageId="3" pageNumber="250">Figures 48</figureCitation>
, two of these are reproduced here as
<figureCitation box="[932,1038,1442,1464]" captionStart="Figure 6" captionStartId="3.[711,791,791,813]" captionTargetBox="[711,1270,106,778]" captionTargetId="figure-2279@3.[711,1269,106,777]" captionTargetPageId="3" captionText="Figure 6. Aloe lateritia var. lateritia in habitat in northernmost Malawi at Ipenza Village in July 1968. The spotted leaves can reach a length of 45cm (18), but grow much longer in cultivation when irrigated throughout the year (Hargreaves, 1975: 92) (published earlier in Hargreaves, 1975: Figure 4). Photograph: Bruce J. Hargreaves." figureDoi="http://doi.org/10.5281/zenodo.7871708" httpUri="https://zenodo.org/record/7871708/files/figure.png" pageId="3" pageNumber="250">Figures 6</figureCitation>
&amp;
<figureCitation box="[1074,1087,1442,1463]" captionStart="Figure 7" captionStartId="4.[121,201,938,960]" captionTargetBox="[121,680,106,930]" captionTargetId="figure-1553@4.[122,680,106,928]" captionTargetPageId="4" captionText="Figure 7. An inflorescence of Aloe lateritia var. lateritia at the locality noted in the caption of Figure 4, about 1.20m (4 feet) high (Hargreaves, 1975: 92) (published earlier in Hargreaves, 1975: Figure 5). Photograph: Bruce J. Hargreaves" figureDoi="http://doi.org/10.5281/zenodo.7871716" httpUri="https://zenodo.org/record/7871716/files/figure.png" pageId="3" pageNumber="250">7</figureCitation>
;
<bibRefCitation author="WHELLAN, J. A." firstAuthor="Whellan" journalOrPublisher="Excelsa" page="111" pageId="3" pageNumber="250" pagination="108 - 113" part="5" refId="ref5406" refString="WHELLAN, J. A. (1975). The Aloes of Malawi. Excelsa 5: 108 - 113." title="The Aloes of Malawi" type="journal article" year="1975">Whellan, 1975: 111</bibRefCitation>
). Philip Downs (†) discovered a further locality in
<date box="[741,922,1498,1520]" pageId="3" pageNumber="250" value="1991-09">September 1991</date>
along the track parallel to the Songwe River between the Ipenza and Chiwanga villages on the hills to the left (southern) side of the track (
<bibRefCitation author="THIEDE, J. &amp; HARGREAVES, B. J. &amp; DOWNS, P. E." box="[782,1035,1582,1604]" etAl="et al." firstAuthor="Thiede" journalOrPublisher="Cactus and Succulent Journal (US)" page="228" pageId="3" pageNumber="250" pagination="224 - 230" part="81" refId="ref5087" refString="THIEDE, J., HARGREAVES, B. J. &amp; DOWNS, P. E. (2009). A gorgeous gem. Sansevieria downsii at Njakwa Gorge, Malawi. Cactus and Succulent Journal (US) 81 (5): 224 - 230." title="A gorgeous gem. Sansevieria downsii at Njakwa Gorge, Malawi" type="journal article" year="2009">
Thiede
<emphasis box="[864,924,1582,1603]" italics="true" pageId="3" pageNumber="250">et al.,</emphasis>
2009: 228
</bibRefCitation>
;
<figureCitation box="[1048,1140,1582,1604]" captionStart="Figure 1" captionStartId="1.[106,186,1283,1305]" captionTargetBox="[106,665,106,1268]" captionTargetId="figure-1255@1.[107,664,107,1267]" captionTargetPageId="1" captionText="Figure 1. Distribution of three Aloe species rare in Malawi: 1) A. canis (yellow dots, a = Senga Hills; b = inland from Domira Bay; c = Ntchisi Hills; Central Region); 2) A. lateritia var. lateritia (blue dot = near Ipenza, two records; Northern Region); and 3) A. suffulta (red dot = Mwabvi Wildlife Reserve; Southern Region). Placements of the dots are partly approximations. The map insert shows the position of Malawi (red) in Africa (map: Wikimedia Commons 2019; insert map: mapsof.net 2015)." figureDoi="http://doi.org/10.5281/zenodo.7871700" httpUri="https://zenodo.org/record/7871700/files/figure.png" pageId="3" pageNumber="250">Figure 1</figureCitation>
: 2).
</paragraph>
<paragraph blockId="3.[711,1270,994,1744]" lastBlockId="4.[121,681,1103,1741]" lastPageId="4" lastPageNumber="251" pageId="3" pageNumber="250">
Specimens were not preserved. Since the two localities are imprecisely given as “beyond Chitipa” (
<bibRefCitation author="WHELLAN, J. A." box="[719,948,1666,1688]" firstAuthor="Whellan" journalOrPublisher="Excelsa" page="111" pageId="3" pageNumber="250" pagination="108 - 113" part="5" refId="ref5406" refString="WHELLAN, J. A. (1975). The Aloes of Malawi. Excelsa 5: 108 - 113." title="The Aloes of Malawi" type="journal article" year="1975">Whellan, 1975: 111</bibRefCitation>
), “near the Songwe River” (
<bibRefCitation author="LANE, S. S." box="[719,889,1694,1715]" firstAuthor="Lane" journalOrPublisher="Umdaus Press, Hatfield" page="27" pageId="3" pageNumber="250" refId="ref4656" refString="LANE, S. S. (2004). A field guide to the aloes of Malawi. Umdaus Press, Hatfield." title="A field guide to the aloes of Malawi" type="book" year="2004">Lane, 2004: 27</bibRefCitation>
), or “south of the Songwe River” (
<bibRefCitation author="KLOPPER, R. R. &amp; LANE, S. S. &amp; MSEKANDIANA-MKWAPATIRA, G. &amp; SMITH, G. F." box="[719,1012,1722,1744]" etAl="et al." firstAuthor="Klopper" journalOrPublisher="Bradleya" page="87" pageId="3" pageNumber="250" pagination="65 - 92" part="30" refId="ref4579" refString="KLOPPER, R. R., LANE, S. S., MSEKANDIANA-MKWAPATIRA, G. &amp; SMITH, G. F. (2012). The genus Aloe L. (Asphodelaceae: Alooideae) in Malawi. Bradleya 30: 65 - 92." title="The genus Aloe L. (Asphodelaceae: Alooideae) in Malawi" type="journal article" year="2012">
Klopper
<emphasis box="[824,887,1722,1743]" italics="true" pageId="3" pageNumber="250">et al.</emphasis>
, 2012: 87
</bibRefCitation>
), more exact locality information is provided here to enable the collection of preserved specimens for deposition in a herbarium. As preserved specimens were not prepared, the specimen-based treatments of
<emphasis box="[632,680,1187,1208]" italics="true" pageId="4" pageNumber="251">Aloe</emphasis>
for the
<emphasis box="[201,519,1215,1237]" italics="true" pageId="4" pageNumber="251">Flora of Tropical East Africa</emphasis>
(
<bibRefCitation author="CARTER, S." editor="POLHILL, R. M. D." firstAuthor="Carter" journalOrPublisher="A. A. Balkema, Rotterdam" page="15 - 17" pageId="4" pageNumber="251" refId="ref4206" refString="CARTER, S. (1994). Aloaceae. In: POLHILL, R. M. D.), Flora of Tropical East Africa. A. A. Balkema, Rotterdam." title="Aloaceae" type="book" volumeTitle="Flora of Tropical East Africa" year="1994">Carter, 1994: 1517</bibRefCitation>
) and
<emphasis box="[268,473,1243,1264]" italics="true" pageId="4" pageNumber="251">Flora zambesiaca</emphasis>
projects (
<bibRefCitation author="CARTER, S." editor="POPE, G. V." firstAuthor="Carter" journalOrPublisher="Royal Botanic Gardens, Kew." pageId="4" pageNumber="251" pagination="48 - 98" part="12" refId="ref4242" refString="CARTER, S. (2001). Aloaceae. In: POPE, G. V. (ED.), Flora zambesiaca 12 (3): 48 - 98. Royal Botanic Gardens, Kew." title="Aloaceae" type="journal article" volumeTitle="Flora zambesiaca" year="2001">Carter, 2001</bibRefCitation>
) omitted the record of
<taxonomicName authority="var. lateritia" class="Liliopsida" family="Asphodelaceae" genus="Aloe" kingdom="Plantae" order="Asparagales" pageId="4" pageNumber="251" phylum="Tracheophyta" rank="variety" species="lateritia" variety="lateritia">
<emphasis box="[468,622,1271,1292]" italics="true" pageId="4" pageNumber="251">Aloe lateritia</emphasis>
var.
<emphasis box="[121,209,1299,1320]" italics="true" pageId="4" pageNumber="251">lateritia</emphasis>
</taxonomicName>
from Malawi. In contrast, the Plants of the World Online database (
<bibRefCitation accessDate="2019-12-29" author="POWO" box="[415,565,1327,1349]" firstAuthor="POWO" journalOrPublisher="Facilitated by the Royal Botanic Gardens, Kew. Published on the Internet at" pageId="4" pageNumber="251" publicationUrl="http://www.plantsoftheworldonline.org" refId="ref4822" refString="POWO. (2019). Plants of the World Online. Facilitated by the Royal Botanic Gardens, Kew. Published on the Internet at http: // www. plantsoftheworld online. org. Retrieved 29 December 2019." title="Plants of the World Online" type="url" year="2019">
<collectionCode box="[415,493,1327,1348]" pageId="4" pageNumber="251">POWO</collectionCode>
, 2019
</bibRefCitation>
) lists “N. Malawi” as its native range.
</paragraph>
<caption ID-DOI="http://doi.org/10.5281/zenodo.7871714" ID-Zenodo-Dep="7871714" httpUri="https://zenodo.org/record/7871714/files/figure.png" pageId="4" pageNumber="251" startId="4.[726,805,473,495]" targetBox="[726,1285,106,461]" targetPageId="4" targetType="figure">
<paragraph blockId="4.[726,1285,473,654]" pageId="4" pageNumber="251">
<emphasis bold="true" box="[726,831,473,495]" pageId="4" pageNumber="251">Figure 8.</emphasis>
<taxonomicName box="[838,971,473,494]" class="Liliopsida" family="Asphodelaceae" genus="Aloe" kingdom="Plantae" order="Asparagales" pageId="4" pageNumber="251" phylum="Tracheophyta" rank="species" species="suffulta">
<emphasis box="[838,971,473,494]" italics="true" pageId="4" pageNumber="251">Aloe suffulta</emphasis>
</taxonomicName>
in habitat in what is now the Mwabvi Wildlife Reserve (Malawi), growing on a river bank (rather likely of the Mwabvi River) in the dense undergrowth of bushes with the long, dry inflorescences literally woven through the supporting bushes and emerging through the top,
<emphasis box="[1052,1075,606,627]" italics="true" pageId="4" pageNumber="251">ca</emphasis>
. 1970.
</paragraph>
<paragraph blockId="4.[726,1285,473,654]" box="[798,1285,632,654]" pageId="4" pageNumber="251">Drawing: Pastor Theo Peter Campbell-Barker.</paragraph>
</caption>
<caption ID-DOI="http://doi.org/10.5281/zenodo.7871716" ID-Zenodo-Dep="7871716" httpUri="https://zenodo.org/record/7871716/files/figure.png" pageId="4" pageNumber="251" startId="4.[121,201,938,960]" targetBox="[121,680,106,930]" targetPageId="4" targetType="figure">
<paragraph blockId="4.[121,681,938,1065]" pageId="4" pageNumber="251">
<emphasis bold="true" box="[121,235,938,960]" pageId="4" pageNumber="251">Figure 7.</emphasis>
An inflorescence of
<taxonomicName authority="var. lateritia" class="Liliopsida" family="Asphodelaceae" genus="Aloe" kingdom="Plantae" order="Asparagales" pageId="4" pageNumber="251" phylum="Tracheophyta" rank="variety" species="lateritia" variety="lateritia">
<emphasis box="[479,625,938,959]" italics="true" pageId="4" pageNumber="251">Aloe lateritia</emphasis>
var.
<emphasis box="[121,206,964,985]" italics="true" pageId="4" pageNumber="251">lateritia</emphasis>
</taxonomicName>
at the locality noted in the caption of Figure 4, about 1.20m (4 feet) high (
<bibRefCitation author="HARGREAVES, B. J." box="[442,672,990,1012]" firstAuthor="Hargreaves" journalOrPublisher="Cactus and Succulent Journal (US)" page="92" pageId="4" pageNumber="251" pagination="86 - 104" part="Suppl. Vol. 1975" refId="ref4454" refString="HARGREAVES, B. J. (1975). Succulents of Chitipa - the muddy place. Cactus and Succulent Journal (US) Suppl. Vol. 1975: 86 - 104 [text reprinted in The Society of Malawi Journal 30 (1): 28 - 34, 1977)." title="Succulents of Chitipa - the muddy place" type="journal article" year="1975">Hargreaves, 1975: 92</bibRefCitation>
) (published earlier in
<bibRefCitation author="HARGREAVES, B. J." box="[340,525,1017,1039]" firstAuthor="Hargreaves" journalOrPublisher="Cactus and Succulent Journal (US)" pageId="4" pageNumber="251" pagination="86 - 104" part="Suppl. Vol. 1975" refId="ref4454" refString="HARGREAVES, B. J. (1975). Succulents of Chitipa - the muddy place. Cactus and Succulent Journal (US) Suppl. Vol. 1975: 86 - 104 [text reprinted in The Society of Malawi Journal 30 (1): 28 - 34, 1977)." title="Succulents of Chitipa - the muddy place" type="journal article" year="1975">Hargreaves, 1975</bibRefCitation>
: Figure 5).
</paragraph>
<paragraph blockId="4.[121,681,938,1065]" box="[331,681,1043,1065]" pageId="4" pageNumber="251">Photograph: Bruce J. Hargreaves</paragraph>
</caption>
<caption ID-DOI="http://doi.org/10.5281/zenodo.7871718" ID-Zenodo-Dep="7871718" httpUri="https://zenodo.org/record/7871718/files/figure.png" pageId="4" pageNumber="251" startId="4.[726,806,1117,1139]" targetBox="[726,1285,675,1105]" targetPageId="4" targetType="figure">
<paragraph blockId="4.[726,1285,1117,1271]" pageId="4" pageNumber="251">
<emphasis bold="true" box="[726,835,1117,1139]" pageId="4" pageNumber="251">Figure 9.</emphasis>
<taxonomicName box="[845,983,1117,1138]" class="Liliopsida" family="Asphodelaceae" genus="Aloe" kingdom="Plantae" order="Asparagales" pageId="4" pageNumber="251" phylum="Tracheophyta" rank="species" species="suffulta">
<emphasis box="[845,983,1117,1138]" italics="true" pageId="4" pageNumber="251">Aloe suffulta</emphasis>
</taxonomicName>
: a young plant collected in what is now the Mwabvi Wildlife Reserve (Malawi). A living specimen was brought to Blantyre in
<emphasis box="[1197,1220,1171,1192]" italics="true" pageId="4" pageNumber="251">ca</emphasis>
. 1970 (published earlier in
<bibRefCitation author="SMITH, G. F. &amp; CROUCH, N. R." box="[943,1213,1197,1219]" firstAuthor="Smith" journalOrPublisher="Flowering Plants of Africa 57" page="21" pageId="4" pageNumber="251" pagination="16 - 22" part="t" refId="ref4982" refString="SMITH, G. F. &amp; CROUCH, N. R. (2001). Aloe suffulta. Flowering Plants of Africa 57: t. 2163, 16 - 22." title="Aloe suffulta" type="journal article" year="2001">Smith &amp; Crouch, 2001: 21</bibRefCitation>
and in
<bibRefCitation author="LANE, S. S." box="[726,879,1223,1245]" firstAuthor="Lane" journalOrPublisher="Umdaus Press, Hatfield" page="45" pageId="4" pageNumber="251" refId="ref4656" refString="LANE, S. S. (2004). A field guide to the aloes of Malawi. Umdaus Press, Hatfield." title="A field guide to the aloes of Malawi" type="book" year="2004">Lane, 2004: 45</bibRefCitation>
).
</paragraph>
<paragraph blockId="4.[726,1285,1117,1271]" box="[798,1285,1249,1271]" pageId="4" pageNumber="251">Drawing: Pastor Theo Peter Campbell-Barker.</paragraph>
</caption>
<paragraph blockId="4.[121,681,1103,1741]" pageId="4" pageNumber="251">
<bibRefCitation author="LANE, S. S." box="[159,339,1383,1405]" firstAuthor="Lane" journalOrPublisher="Umdaus Press, Hatfield" page="27" pageId="4" pageNumber="251" refId="ref4656" refString="LANE, S. S. (2004). A field guide to the aloes of Malawi. Umdaus Press, Hatfield." title="A field guide to the aloes of Malawi" type="book" year="2004">Lane (2004: 27)</bibRefCitation>
and
<bibRefCitation author="KLOPPER, R. R. &amp; LANE, S. S. &amp; MSEKANDIANA-MKWAPATIRA, G. &amp; SMITH, G. F." box="[400,680,1383,1405]" etAl="et al." firstAuthor="Klopper" journalOrPublisher="Bradleya" page="87" pageId="4" pageNumber="251" pagination="65 - 92" part="30" refId="ref4579" refString="KLOPPER, R. R., LANE, S. S., MSEKANDIANA-MKWAPATIRA, G. &amp; SMITH, G. F. (2012). The genus Aloe L. (Asphodelaceae: Alooideae) in Malawi. Bradleya 30: 65 - 92." title="The genus Aloe L. (Asphodelaceae: Alooideae) in Malawi" type="journal article" year="2012">
Klopper
<emphasis box="[499,550,1383,1405]" italics="true" pageId="4" pageNumber="251">et al</emphasis>
. (2012: 87)
</bibRefCitation>
suggest that the plants of
<taxonomicName authorityName=", Engler" authorityYear="1895" box="[429,555,1411,1433]" class="Liliopsida" family="Asphodelaceae" genus="Aloe" kingdom="Plantae" order="Asparagales" pageId="4" pageNumber="251" phylum="Tracheophyta" rank="species" species="lateritia">
<emphasis box="[429,555,1411,1433]" italics="true" pageId="4" pageNumber="251">A. lateritia</emphasis>
</taxonomicName>
that occur near Ipenza may have been brought to Malawi by humans travelling back and forth between Malawi and Tanzania. However, the photographs of
<bibRefCitation author="HARGREAVES, B. J." box="[121,330,1523,1545]" firstAuthor="Hargreaves" journalOrPublisher="Cactus and Succulent Journal (US)" pageId="4" pageNumber="251" pagination="86 - 104" part="Suppl. Vol. 1975" refId="ref4454" refString="HARGREAVES, B. J. (1975). Succulents of Chitipa - the muddy place. Cactus and Succulent Journal (US) Suppl. Vol. 1975: 86 - 104 [text reprinted in The Society of Malawi Journal 30 (1): 28 - 34, 1977)." title="Succulents of Chitipa - the muddy place" type="journal article" year="1975">Hargreaves (1975</bibRefCitation>
:
<figureCitation box="[353,453,1523,1545]" captionStart-0="Figure 4" captionStart-1="Figure 5" captionStart-2="Figure 6" captionStart-3="Figure 7" captionStart-4="Figure 8" captionStartId-0="2.[726,806,1028,1050]" captionStartId-1="3.[106,185,851,873]" captionStartId-2="3.[711,791,791,813]" captionStartId-3="4.[121,201,938,960]" captionStartId-4="4.[726,805,473,495]" captionTargetBox-0="[718,1285,106,1015]" captionTargetBox-1="[106,665,106,840]" captionTargetBox-2="[711,1270,106,778]" captionTargetBox-3="[121,680,106,930]" captionTargetBox-4="[726,1285,106,461]" captionTargetId-0="figure-1878@2.[727,1284,107,1015]" captionTargetId-1="figure-2126@3.[107,664,107,839]" captionTargetId-2="figure-2279@3.[711,1269,106,777]" captionTargetId-3="figure-1553@4.[122,680,106,928]" captionTargetId-4="figure-1760@4.[727,1284,107,460]" captionTargetPageId-0="2" captionTargetPageId-1="3" captionTargetPageId-2="3" captionTargetPageId-3="4" captionTargetPageId-4="4" captionText-0="Figure 4. The specimen of Aloe canis that Ms Ruth Mthawanji received from Reverend Stewart Lane in the early-2000s photographed in Blantyre, Malawi, in 2015. Photograph: R. Mthawanji." captionText-1="Figure 5. Panicle of the specimen of Aloe canis that Ms Ruth Mthawanji received from Reverend Stewart Lane photographed in Blantyre, Malawi, in 2019. Photograph: R. Mthawanji." captionText-2="Figure 6. Aloe lateritia var. lateritia in habitat in northernmost Malawi at Ipenza Village in July 1968. The spotted leaves can reach a length of 45cm (18), but grow much longer in cultivation when irrigated throughout the year (Hargreaves, 1975: 92) (published earlier in Hargreaves, 1975: Figure 4). Photograph: Bruce J. Hargreaves." captionText-3="Figure 7. An inflorescence of Aloe lateritia var. lateritia at the locality noted in the caption of Figure 4, about 1.20m (4 feet) high (Hargreaves, 1975: 92) (published earlier in Hargreaves, 1975: Figure 5). Photograph: Bruce J. Hargreaves" captionText-4="Figure 8. Aloe suffulta in habitat in what is now the Mwabvi Wildlife Reserve (Malawi), growing on a river bank (rather likely of the Mwabvi River) in the dense undergrowth of bushes with the long, dry inflorescences literally woven through the supporting bushes and emerging through the top, ca. 1970. Drawing: Pastor Theo Peter Campbell-Barker." figureDoi-0="http://doi.org/10.5281/zenodo.7871706" figureDoi-1="http://doi.org/10.5281/zenodo.7871712" figureDoi-2="http://doi.org/10.5281/zenodo.7871708" figureDoi-3="http://doi.org/10.5281/zenodo.7871716" figureDoi-4="http://doi.org/10.5281/zenodo.7871714" httpUri-0="https://zenodo.org/record/7871706/files/figure.png" httpUri-1="https://zenodo.org/record/7871712/files/figure.png" httpUri-2="https://zenodo.org/record/7871708/files/figure.png" httpUri-3="https://zenodo.org/record/7871716/files/figure.png" httpUri-4="https://zenodo.org/record/7871714/files/figure.png" pageId="4" pageNumber="251">Fig. 48</figureCitation>
, two of these are reproduced here as
<figureCitation box="[355,466,1551,1573]" captionStart="Figure 6" captionStartId="3.[711,791,791,813]" captionTargetBox="[711,1270,106,778]" captionTargetId="figure-2279@3.[711,1269,106,777]" captionTargetPageId="3" captionText="Figure 6. Aloe lateritia var. lateritia in habitat in northernmost Malawi at Ipenza Village in July 1968. The spotted leaves can reach a length of 45cm (18), but grow much longer in cultivation when irrigated throughout the year (Hargreaves, 1975: 92) (published earlier in Hargreaves, 1975: Figure 4). Photograph: Bruce J. Hargreaves." figureDoi="http://doi.org/10.5281/zenodo.7871708" httpUri="https://zenodo.org/record/7871708/files/figure.png" pageId="4" pageNumber="251">Figures 6</figureCitation>
&amp;
<figureCitation box="[508,521,1552,1573]" captionStart="Figure 7" captionStartId="4.[121,201,938,960]" captionTargetBox="[121,680,106,930]" captionTargetId="figure-1553@4.[122,680,106,928]" captionTargetPageId="4" captionText="Figure 7. An inflorescence of Aloe lateritia var. lateritia at the locality noted in the caption of Figure 4, about 1.20m (4 feet) high (Hargreaves, 1975: 92) (published earlier in Hargreaves, 1975: Figure 5). Photograph: Bruce J. Hargreaves" figureDoi="http://doi.org/10.5281/zenodo.7871716" httpUri="https://zenodo.org/record/7871716/files/figure.png" pageId="4" pageNumber="251">7</figureCitation>
) show plants apparently growing naturally amongst undisturbed grasses, and the locality found by Downs is beyond villages.
</paragraph>
<paragraph blockId="4.[121,681,1103,1741]" lastBlockId="4.[726,1286,1300,1742]" pageId="4" pageNumber="251">
In the protologue of the name
<taxonomicName authority=", Engler (1895: 140 - 141)" authorityName=", Engler" authorityPageNumber="140 - 141" authorityYear="1895" box="[523,674,1663,1684]" class="Liliopsida" family="Asphodelaceae" genus="Aloe" kingdom="Plantae" order="Asparagales" pageId="4" pageNumber="251" phylum="Tracheophyta" rank="species" species="lateritia">
<emphasis box="[523,674,1663,1684]" italics="true" pageId="4" pageNumber="251">Aloe lateritia</emphasis>
</taxonomicName>
,
<bibRefCitation author="ENGLER, A." bookContentInfo="Verzeichniss der bis jetzt aus Ost-Afrika bekannt gewordenen Pflanzen, mit 45 Tafein" box="[121,393,1691,1713]" firstAuthor="Engler" journalOrPublisher="D. Reimer, Berlin" page="140 - 141" pageId="4" pageNumber="251" part="Theil C" refId="ref4370" refString="ENGLER, A. (1895). Die Pflanzenwelt Ost-Afrikas und der Nachbargebiete. Theil C. Verzeichniss der bis jetzt aus Ost-Afrika bekannt gewordenen Pflanzen, mit 45 Tafein. D. Reimer, Berlin." title="Die Pflanzenwelt Ost-Afrikas und der Nachbargebiete" type="book" year="1895">Engler (1895: 140141)</bibRefCitation>
cited a single collection “([TANZANIA, Moshi distr.,] Rombo ― Volk. [-ens] n. 404) ― An sandigen und steinigen Abhängen, an Felsvorsprüngen um
<quantity box="[968,1116,1328,1350]" metricMagnitude="3" metricUnit="m" metricValue="1.45" metricValueMax="1.5" metricValueMin="1.4" pageId="4" pageNumber="251" unit="m" value="1450.0" valueMax="1500.0" valueMin="1400.0">1400―1500m</quantity>
”. [English: On sandy and stony slopes, on rock spurs around
<quantity metricMagnitude="3" metricUnit="m" metricValue="1.45" metricValueMax="1.5" metricValueMin="1.4" pageId="4" pageNumber="251" unit="m" value="1450.0" valueMax="1500.0" valueMin="1400.0">1400 1500m</quantity>
].
</paragraph>
</subSubSection>
<subSubSection lastPageId="5" lastPageNumber="252" pageId="4" pageNumber="251" type="materials_examined">
<paragraph blockId="4.[726,1286,1300,1742]" lastBlockId="5.[106,666,107,521]" lastPageId="5" lastPageNumber="252" pageId="4" pageNumber="251">
<materialsCitation collectionCode="B" collectorName="Volkens" location="Herb" municipality="Reynolds" pageId="4" pageNumber="251" specimenCode="B 10 0165570" specimenCount="1" typeStatus="holotype">
Two specimens are available for examination online. The holotype, cited by
<bibRefCitation author="REYNOLDS, G. W." firstAuthor="Reynolds" journalOrPublisher="The Trustees of the Aloes Book Fund, Mbabane" page="95" pageId="4" pageNumber="251" refId="ref4954" refString="REYNOLDS, G. W. (1966). The aloes of Tropical Africa and Madagascar. The Trustees of the Aloes Book Fund, Mbabane." title="The aloes of Tropical Africa and Madagascar" type="book" year="1966">
<collectingMunicipality box="[1109,1210,1412,1434]" pageId="4" pageNumber="251">Reynolds</collectingMunicipality>
(1966: 95
</bibRefCitation>
, as “type” and on p. 96 as “
<typeStatus box="[1083,1179,1440,1462]" pageId="4" pageNumber="251">holotype</typeStatus>
”) and by
<bibRefCitation author="CARTER, S." box="[726,906,1468,1490]" editor="POLHILL, R. M. D." firstAuthor="Carter" journalOrPublisher="A. A. Balkema, Rotterdam" page="15" pageId="4" pageNumber="251" refId="ref4206" refString="CARTER, S. (1994). Aloaceae. In: POLHILL, R. M. D.), Flora of Tropical East Africa. A. A. Balkema, Rotterdam." title="Aloaceae" type="book" volumeTitle="Flora of Tropical East Africa" year="1994">Carter (1994: 15</bibRefCitation>
, as “holo.”) is kept at Herb. B,
<emphasis italics="true" pageId="4" pageNumber="251">
<collectorName pageId="4" pageNumber="251">Volkens</collectorName>
404
</emphasis>
(
<collectionCode box="[840,857,1497,1518]" pageId="4" pageNumber="251">B</collectionCode>
barcode
<specimenCode box="[960,1115,1496,1518]" pageId="4" pageNumber="251">B 10 0165570</specimenCode>
, digital image! at https://herbarium.bgbm. org/object/B100165570); a photograph of this specimen is at
<location LSID="urn:lsid:plazi:treatment:03D2FB250D61163F7920F9C7FE02E23A:8EA41CE80D66163E7DE9F9C6FB7EE615" box="[1187,1244,1552,1574]" municipality="Reynolds" name="Herb" pageId="4" pageNumber="251">Herb</location>
. K (
<accessionNumber box="[734,873,1581,1602]" pageId="4" pageNumber="251">K000210270</accessionNumber>
, at https://apps.kew.org/ herbcat/detailsQuery.do?imageId=&amp;pageCode=1&amp;present- Page=1&amp;queryId=1&amp;sessionId=8
<collectionCode box="[1086,1102,1637,1658]" pageId="4" pageNumber="251">F</collectionCode>
9279
<collectionCode box="[1154,1172,1637,1658]" pageId="4" pageNumber="251">D</collectionCode>
2
<collectionCode box="[1186,1202,1637,1658]" pageId="4" pageNumber="251">F</collectionCode>
48866
<collectionCode box="[1267,1284,1637,1658]" pageId="4" pageNumber="251">A</collectionCode>
08
<collectionCode box="[752,786,1665,1686]" pageId="4" pageNumber="251">BE</collectionCode>
26
<collectionCode box="[813,830,1665,1686]" pageId="4" pageNumber="251">C</collectionCode>
47902
<collectionCode box="[895,929,1665,1686]" pageId="4" pageNumber="251">AC</collectionCode>
9
<collectionCode box="[943,979,1665,1686]" pageId="4" pageNumber="251">DC</collectionCode>
&amp;barcode=
<collectionCode box="[1097,1115,1665,1686]" pageId="4" pageNumber="251">K</collectionCode>
000210270).
</materialsCitation>
<materialsCitation collectionCode="BM" collectorName="Volkens" lastPageId="5" lastPageNumber="252" pageId="4" pageNumber="251" specimenCode="BM000911694" specimenCount="1" typeStatus="isotype">
An
<typeStatus box="[726,805,1692,1714]" pageId="4" pageNumber="251">isotype</typeStatus>
exists at Herb. BM
<emphasis box="[1038,1174,1692,1714]" italics="true" pageId="4" pageNumber="251">
<collectorName box="[1038,1125,1692,1713]" pageId="4" pageNumber="251">Volkens</collectorName>
404
</emphasis>
(
<collectionCode box="[1191,1231,1693,1714]" pageId="4" pageNumber="251">BM</collectionCode>
barcode
<specimenCode box="[817,980,1720,1742]" pageId="4" pageNumber="251">BM000911694</specimenCode>
, digital image! at https://plants.jstor. org/stable/10.5555/al.ap.specimen.bm000911694?searchUri=filter%3Dfree_text %26so%3Dps_group_by_genus_species%2Basc%26 Query%3
<collectionCode box="[213,273,192,213]" pageId="5" pageNumber="252">DBM</collectionCode>
000911694%26loggedin%3Dtrue).
</materialsCitation>
</paragraph>
</subSubSection>
<subSubSection pageId="5" pageNumber="252" type="discussion">
<paragraph blockId="5.[106,666,107,521]" pageId="5" pageNumber="252">
The
<typeStatus box="[154,248,219,241]" pageId="5" pageNumber="252">holotype</typeStatus>
, i.e., the Herb. B specimen, was dated as having been collected on
<date box="[488,665,247,269]" pageId="5" pageNumber="252" value="1893-06-13">13 June 1893</date>
[“
<date box="[124,205,275,297]" pageId="5" pageNumber="252" value="1993-06-13">13.6.93</date>
”], while the isotype, i.e., the Herb. BM specimen, carries the date “Juni [June] 1893”, with the exact day on which it was collected therefore omitted. However, since a black-and-white image of the more precisely dated
<typeStatus box="[401,494,387,409]" pageId="5" pageNumber="252">holotype</typeStatus>
is affixed to the BM specimen, albeit with some plant fragments removed, we interpret the BM specimen as an isotype and not as a different gathering (
<bibRefCitation author="TURLAND, N. J. &amp; WIERSEMA, J. H. &amp; BARRIE, F. R. &amp; GREUTER, W. &amp; HAWKSWORTH, D. L. &amp; HERENDEEN, P. S. &amp; KNAPP, S. &amp; KUSBER, W. - H. &amp; LI, D. - Z. &amp; MARHOLD, K. &amp; MAY, T. W. &amp; MCNEILL, J. &amp; MONRO, A. M. &amp; PRADO, J. &amp; PRICE, M. J. &amp; SMITH, G. F." etAl="et al." firstAuthor="Turland" journalOrPublisher="Koeltz Botanical Books, Glashutten" page="16" pageId="5" pageNumber="252" part="159" refId="ref5188" refString="TURLAND, N. J., WIERSEMA, J. H., BARRIE, F. R., GREUTER, W., HAWKSWORTH, D. L., HERENDEEN, P. S., KNAPP, S., KUSBER, W. - H., LI, D. - Z., MARHOLD, K., MAY, T. W., MCNEILL, J., MONRO, A. M., PRADO, J., PRICE, M. J. &amp; SMITH, G. F. DS.) (2018). International Code of Nomenclature for algae, fungi, and plants (Shenzhen Code) adopted by the Nineteenth International Botanical Congress Shenzhen, China, July 2017. Regnum Vegetabile 159. Koeltz Botanical Books, Glashutten." title="International Code of Nomenclature for algae, fungi, and plants (Shenzhen Code) adopted by the Nineteenth International Botanical Congress Shenzhen, China, July 2017" type="book" volumeTitle="Regnum Vegetabile" year="2018">
Turland
<emphasis box="[600,651,471,493]" italics="true" pageId="5" pageNumber="252">et al</emphasis>
., 2018: 16
</bibRefCitation>
, Art. 8.2 Footnote).
</paragraph>
</subSubSection>
</treatment>
</document>