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<document id="5200702F5173209CEB307FE683252301" ID-CLB-Dataset="24738" ID-DOI="10.11646/zootaxa.4702.1.14" ID-GBIF-Dataset="a6dcda0b-452e-43c3-9c84-a769ec073c42" ID-ISSN="1175-5326" ID-Zenodo-Dep="3562723" ID-ZooBank="F0FCDD0B-B356-41D0-8848-247BADCA73A2" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1575478845748" checkinUser="plazi" docAuthor="Koeda, Keita &amp; Ho, Hsuan-Ching" docDate="2019" docId="03C9DD2CFFA02E30FF1C2AB98A8DF974" docLanguage="en" docName="zootaxa.4702.1.14.pdf" docOrigin="Zootaxa 4702 (1)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Eustomias (Nominostomias) tritentaculatus Koeda &amp; Ho 2019, sp. nov." docType="treatment" docVersion="8" lastPageNumber="101" masterDocId="FFF0A554FFA72E37FF8B2828883BFE07" masterDocTitle="Review of the genus Eustomias (Stomiiformes: Stomiidae: Melanostomiinae) of Taiwan, with descriptions of three new species" masterLastPageNumber="106" masterPageNumber="94" pageNumber="101" updateTime="1698771538700" updateUser="ExternalLinkService">
<mods:mods id="11EF592CA1EB0ABA677AB3574AE5BF9B" xmlns:mods="http://www.loc.gov/mods/v3">
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<mods:title id="592C768B38EA5F90E42EE4735538D815">Review of the genus Eustomias (Stomiiformes: Stomiidae: Melanostomiinae) of Taiwan, with descriptions of three new species</mods:title>
</mods:titleInfo>
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<mods:namePart id="2047C2EB30E91692477273EA205C6EB1">Koeda, Keita</mods:namePart>
</mods:name>
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<mods:namePart id="592971AEB4A35B84F7EF476802BFD31A">Ho, Hsuan-Ching</mods:namePart>
</mods:name>
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<mods:title id="C092691F346DE845634C13DD7F6696AD">Zootaxa</mods:title>
</mods:titleInfo>
<mods:part id="9E1E074BA9E024CDC814B17BD31248F8">
<mods:date id="5E76236E215C36C09F1C4EAE6F90B5E1">2019</mods:date>
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<mods:number id="0EBE270CC8F11F92A21DFEBFF9F320E4">2019-12-04</mods:number>
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<mods:detail id="7A9B299ACB8E7907C528B60D2D5D968F" type="volume">
<mods:number id="FA49F7C2945485FCDB934C825CECD37F">4702</mods:number>
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<mods:classification id="A07A8B789A4EB16817693B5CFC279AC5">journal article</mods:classification>
<mods:identifier id="A07D5C848893DE518E8BBF2FBAB05F6F" type="CLB-Dataset">24738</mods:identifier>
<mods:identifier id="6F73C7F58C9A26C1D7C85DF4DA6471BA" type="DOI">10.11646/zootaxa.4702.1.14</mods:identifier>
<mods:identifier id="770B1CECD8277EBBE3EC732E0F0F7811" type="GBIF-Dataset">a6dcda0b-452e-43c3-9c84-a769ec073c42</mods:identifier>
<mods:identifier id="59710B36B482A7B4FDB812DB7E4D7F04" type="ISSN">1175-5326</mods:identifier>
<mods:identifier id="D26DF5B669F3AA39833492BF4BC7F631" type="Zenodo-Dep">3562723</mods:identifier>
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</mods:mods>
<treatment id="03C9DD2CFFA02E30FF1C2AB98A8DF974" ID-GBIF-Taxon="160618111" LSID="urn:lsid:plazi:treatment:03C9DD2CFFA02E30FF1C2AB98A8DF974" httpUri="http://treatment.plazi.org/id/03C9DD2CFFA02E30FF1C2AB98A8DF974" lastPageNumber="101" pageId="7" pageNumber="101">
<subSubSection id="C37A3FB1FFA02E30FF1C2AB98B30FCAC" box="[151,779,657,684]" pageId="7" pageNumber="101" type="nomenclature">
<paragraph id="8BDF6C3AFFA02E30FF1C2AB98B30FCAC" blockId="7.[151,779,657,719]" box="[151,779,657,684]" pageId="7" pageNumber="101">
<heading id="D097DB56FFA02E30FF1C2AB98B30FCAC" bold="true" box="[151,779,657,684]" fontSize="11" level="1" pageId="7" pageNumber="101" reason="1">
<emphasis id="B914B028FFA02E30FF1C2AB98B30FCAC" bold="true" box="[151,779,657,684]" pageId="7" pageNumber="101">
<taxonomicName id="4C6017B9FFA02E30FF1C2AB98A9DFCAB" authority="Koeda &amp; Ho, 2019" authorityName="Koeda &amp; Ho" authorityYear="2019" box="[151,678,657,684]" class="Actinopterygii" family="Stomiidae" genus="Eustomias" kingdom="Animalia" order="Stomiiformes" pageId="7" pageNumber="101" phylum="Chordata" rank="species" species="tritentaculatus" status="sp. nov." subGenus="Nominostomias">
<emphasis id="B914B028FFA02E30FF1C2AB9892DFCAC" bold="true" box="[151,278,657,683]" italics="true" pageId="7" pageNumber="101">Eustomias</emphasis>
(
<emphasis id="B914B028FFA02E30FEAC2AB989DFFCAC" bold="true" box="[295,484,657,683]" italics="true" pageId="7" pageNumber="101">Nominostomias</emphasis>
)
<emphasis id="B914B028FFA02E30FE7D2AB98A9DFCAB" bold="true" box="[502,678,657,684]" italics="true" pageId="7" pageNumber="101">tritentaculatus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A2270D53FFA02E30FD262AB98B30FCAC" box="[685,779,657,683]" pageId="7" pageNumber="101" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C37A3FB1FFA02E30FF1C2A9D8946FCC8" box="[151,381,693,719]" pageId="7" pageNumber="101" type="description">
<paragraph id="8BDF6C3AFFA02E30FF1C2A9D8946FCC8" blockId="7.[151,779,657,719]" box="[151,381,693,719]" pageId="7" pageNumber="101">
<figureCitation id="135B70BFFFA02E30FF1C2A9D88C3FCC8" box="[151,248,693,719]" captionStart="FIGURE 4" captionStartId="8.[151,250,1413,1438]" captionTargetBox="[151,1437,181,1375]" captionTargetId="figure@8.[151,1437,181,1388]" captionTargetPageId="8" captionText="FIGURE 4. Eustomias (Nominostomias) tritentaculatus sp. nov. from Taiwan, NMMB-P30396, holotype, 130.9 mm SL. Top: preserved specimen; middle: closeup illustration of distal part of barbel; bottom: diagram of dentition. Black and white color of teeth indicate fixed and movable, respectively." figureDoi="http://doi.org/10.5281/zenodo.3562731" httpUri="https://zenodo.org/record/3562731/files/figure.png" pageId="7" pageNumber="101">Figure 4</figureCitation>
;
<tableCitation id="C6E25981FFA02E30FE8F2A9D8946FCC8" box="[260,381,693,719]" captionStart-0="TABLE 1" captionStart-1="TABLE 2" captionStartId-0="3.[151,244,652,679]" captionStartId-1="10.[151,244,587,614]" captionTargetBox-0="[162,1947,694,1535]" captionTargetBox-1="[162,1970,633,1608]" captionTargetPageId-0="3" captionText-0="TABLE 1. Meristic data of the specimens of genus Eustomias from Taiwan." captionText-1="TABLE 2. Morphometric data of the specimens of genus Eustomias from Taiwan." httpUri-0="http://table.plazi.org/id/DF1F3CB2FFA42E34FF1C2AA48BD6FCA0" pageId="7" pageNumber="101" tableUuid-0="DF1F3CB2FFA42E34FF1C2AA48BD6FCA0">Tables 1–2</tableCitation>
</paragraph>
</subSubSection>
<subSubSection id="C37A3FB1FFA02E30FF1C2AD58C35FD58" pageId="7" pageNumber="101" type="materials_examined">
<paragraph id="8BDF6C3AFFA02E30FF1C2AD58DBBFD3C" blockId="7.[151,1437,764,1908]" pageId="7" pageNumber="101">
<materialsCitation id="3B086667FFA02E30FF1C2AD58DBBFD3C" ID-GBIF-Occurrence="2462934961" collectingDate="2005-03-16" collectionCode="NMMB-P" country="China" county="Dong-gang" latitude="22.65" location="Dong-gang" longLatPrecision="1259" longitude="120.4" municipality="Sea" pageId="7" pageNumber="101" specimenCount="1" typeStatus="holotype">
<emphasis id="B914B028FFA02E30FF1C2AD58930FD10" bold="true" box="[151,267,765,791]" pageId="7" pageNumber="101">
<typeStatus id="54DBD298FFA02E30FF1C2AD5893DFD10" box="[151,262,765,791]" pageId="7" pageNumber="101" type="holotype">Holotype</typeStatus>
.
</emphasis>
<collectionCode id="ED71F4FFFFA02E30FE9A2AD589BEFD10" box="[273,389,765,791]" pageId="7" pageNumber="101">NMMB-P</collectionCode>
<quantity id="4C98C1DFFFA02E30FE0E2AD58A7DFD10" box="[389,582,765,791]" metricMagnitude="4" metricUnit="m" metricValue="3.0396130899999996" pageId="7" pageNumber="101" unit="mm" value="3.03961309E7">30396, 130.9 mm</quantity>
SL, off
<location id="8EBF3AE1FFA02E30FD152AD58B27FD10" LSID="urn:lsid:plazi:treatment:03C9DD2CFFA02E30FF1C2AB98A8DF974:8EBF3AE1FFA02E30FD152AD58B27FD10" box="[670,796,765,791]" country="China" county="Dong-gang" latitude="22.65" longLatPrecision="1259" longitude="120.4" municipality="Sea" name="Dong-gang" pageId="7" pageNumber="101">Dong-gang</location>
(ca.
<geoCoordinate id="EE540AFDFFA02E30FCDA2AD48B8AFD10" box="[849,945,764,791]" degrees="22" direction="north" minutes="39" orientation="latitude" pageId="7" pageNumber="101" precision="925" value="22.65">22°39ʹN</geoCoordinate>
,
<geoCoordinate id="EE540AFDFFA02E30FC302AD48C1CFD10" box="[955,1063,764,791]" degrees="120" direction="east" minutes="24" orientation="longitude" pageId="7" pageNumber="101" precision="925" value="120.4">120°24ʹE</geoCoordinate>
),
<location id="8EBF3AE1FFA02E30FBB22AD58C9AFD10" LSID="urn:lsid:plazi:treatment:03C9DD2CFFA02E30FF1C2AB98A8DF974:8EBF3AE1FFA02E30FBB22AD58C9AFD10" box="[1081,1185,765,791]" country="China" county="Dong-gang" latitude="22.65" longLatPrecision="1259" longitude="120.4" municipality="Sea" name="Pingtung" pageId="7" pageNumber="101">Pingtung</location>
,
<location id="8EBF3AE1FFA02E30FB202AD58CD7FD10" LSID="urn:lsid:plazi:treatment:03C9DD2CFFA02E30FF1C2AB98A8DF974:8EBF3AE1FFA02E30FB202AD58CD7FD10" box="[1195,1260,765,791]" country="China" county="Dong-gang" latitude="22.65" longLatPrecision="1259" longitude="120.4" municipality="Sea" name="South" pageId="7" pageNumber="101">South</location>
<collectingCountry id="F3772CAAFFA02E30FB792AD58D0EFD10" box="[1266,1333,765,791]" name="China" pageId="7" pageNumber="101">China</collectingCountry>
<collectingMunicipality id="6BBBF640FFA02E30FAB02AD58D5DFD10" box="[1339,1382,765,791]" pageId="7" pageNumber="101">Sea</collectingMunicipality>
, Tai- wan,
<date id="FFDE4AFAFFA02E30FF5F2B098956FD3C" box="[212,365,800,827]" pageId="7" pageNumber="101" value="2005-03-16">
<collectingDate id="EF9AB312FFA02E30FF5F2B098956FD3C" box="[212,365,800,827]" pageId="7" pageNumber="101" value="2005-03-16">16 Mar. 2005</collectingDate>
</date>
, obtained at fish-landing ground at
<collectingCounty id="62BE14B6FFA02E30FD712B098B40FD3C" box="[762,891,801,827]" pageId="7" pageNumber="101">Dong-gang</collectingCounty>
, collected by commercial mid-water trawling.
</materialsCitation>
</paragraph>
<paragraph id="8BDF6C3AFFA02E30FF4C2B6D8C35FD58" blockId="7.[151,1437,764,1908]" box="[199,1038,837,863]" pageId="7" pageNumber="101">
<materialsCitation id="3B086667FFA02E30FF4C2B6D8C32FD58" ID-GBIF-Occurrence="2462934955" box="[199,1033,837,863]" collectingDate="2011-11-03" collectionCode="NMMB-P" pageId="7" pageNumber="101" specimenCount="2" typeStatus="paratype">
<emphasis id="B914B028FFA02E30FF4C2B6D8900FD58" bold="true" box="[199,315,837,863]" pageId="7" pageNumber="101">
<typeStatus id="54DBD298FFA02E30FF4C2B6D890DFD58" box="[199,310,837,863]" pageId="7" pageNumber="101" type="paratype">Paratype</typeStatus>
.
</emphasis>
<collectionCode id="ED71F4FFFFA02E30FEC92B6D898EFD58" box="[322,437,837,863]" pageId="7" pageNumber="101">NMMB-P</collectionCode>
<quantity id="4C98C1DFFFA02E30FE3D2B6D8A41FD58" box="[438,634,837,863]" metricMagnitude="4" metricUnit="m" metricValue="2.27601204" pageId="7" pageNumber="101" unit="mm" value="2.27601204E7">22760, 120.4 mm</quantity>
SL, same as
<typeStatus id="54DBD298FFA02E30FC852B6D8B48FD58" box="[782,883,837,863]" pageId="7" pageNumber="101" type="holotype">holotype</typeStatus>
,
<date id="FFDE4AFAFFA02E30FCF52B6D8C32FD58" box="[894,1033,837,863]" pageId="7" pageNumber="101" value="2011-11-03">
<collectingDate id="EF9AB312FFA02E30FCF52B6D8C32FD58" box="[894,1033,837,863]" pageId="7" pageNumber="101" value="2011-11-03">3 Nov. 2011</collectingDate>
</date>
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="C37A3FB1FFA02E30FF4C2B408D10FA14" pageId="7" pageNumber="101" type="diagnosis">
<paragraph id="8BDF6C3AFFA02E30FF4C2B408D10FA14" blockId="7.[151,1437,764,1908]" pageId="7" pageNumber="101">
<emphasis id="B914B028FFA02E30FF4C2B408979FD84" bold="true" box="[199,322,872,899]" pageId="7" pageNumber="101">Diagnosis.</emphasis>
A species of subgenus
<taxonomicName id="4C6017B9FFA02E30FDDB2B428AC5FD84" box="[592,766,874,899]" class="Actinopterygii" family="Stomiidae" genus="Eustomias" kingdom="Animalia" order="Stomiiformes" pageId="7" pageNumber="101" phylum="Chordata" rank="subGenus" subGenus="Nominostomias">
<emphasis id="B914B028FFA02E30FDDB2B428AC5FD84" box="[592,766,874,899]" italics="true" pageId="7" pageNumber="101">Nominostomias</emphasis>
</taxonomicName>
with a unique barbel structure: barbel moderately long, its length 65.3% SL; terminal bulb small, its length 1.2% SL; three terminal filaments, one of which thicker than others with few pigmentations near base, all filaments simple and lacking bulblets and secondary branches; longest filament length 4.1% SL; axis of stem weakly pigmented; a distinct ring-shaped black spot surrounding base of bulb. Some of the meristic characters are also distinct: anal-fin rays 35; IP 7; PV 34; VAV 19; OV 31; VAL 19.
</paragraph>
</subSubSection>
<subSubSection id="C37A3FB1FFA02E30FF4C2C348C27FAA4" pageId="7" pageNumber="101" type="description">
<paragraph id="8BDF6C3AFFA02E30FF4C2C348BFBFA30" blockId="7.[151,1437,764,1908]" box="[199,960,1052,1079]" pageId="7" pageNumber="101">
<emphasis id="B914B028FFA02E30FF4C2C348962FA30" bold="true" box="[199,345,1052,1079]" pageId="7" pageNumber="101">Description.</emphasis>
Counts and measurements are given in
<tableCitation id="C6E25981FFA02E30FC9C2C358B4EFA30" box="[791,885,1053,1079]" captionStart="TABLE 1" captionStartId="3.[151,244,652,679]" captionTargetBox="[162,1947,694,1535]" captionText="TABLE 1. Meristic data of the specimens of genus Eustomias from Taiwan." httpUri="http://table.plazi.org/id/DF1F3CB2FFA42E34FF1C2AA48BD6FCA0" pageId="7" pageNumber="101" tableUuid="DF1F3CB2FFA42E34FF1C2AA48BD6FCA0">Tables 1</tableCitation>
and
<tableCitation id="C6E25981FFA02E30FC202C358B87FA30" box="[939,956,1053,1079]" captionStart="TABLE 2" captionStartId="10.[151,244,587,614]" captionTargetBox="[162,1970,633,1608]" captionText="TABLE 2. Morphometric data of the specimens of genus Eustomias from Taiwan." pageId="7" pageNumber="101">2</tableCitation>
.
</paragraph>
<paragraph id="8BDF6C3AFFA02E30FF4C2C698C27FAA4" blockId="7.[151,1437,764,1908]" pageId="7" pageNumber="101">
Jaws not curved. Twelve teeth on premaxilla, anterior 2 teeth (and 6th in
<typeStatus id="54DBD298FFA02E30FC7B2C698C69FA5C" box="[1008,1106,1089,1115]" pageId="7" pageNumber="101" type="paratype">paratype</typeStatus>
) fixed; 8–10 strongly oblique comb-like teeth on maxilla; 15–18 teeth on dentary, all teeth (except for 2nd in
<typeStatus id="54DBD298FFA02E30FB822C4D8C56FA87" box="[1033,1133,1125,1152]" pageId="7" pageNumber="101" type="holotype">holotype</typeStatus>
) movable. Postorbital photophore rounded and small, its diameter as long as anteriormost upper jaw tooth.
</paragraph>
</subSubSection>
<subSubSection id="C37A3FB1FFA02E30FF4C2C858A2DFAEC" pageId="7" pageNumber="101" type="etymology">
<paragraph id="8BDF6C3AFFA02E30FF4C2C858A2DFAEC" blockId="7.[151,1437,764,1908]" pageId="7" pageNumber="101">
<emphasis id="B914B028FFA02E30FF4C2C858975FAC0" bold="true" box="[199,334,1197,1223]" pageId="7" pageNumber="101">Etymology.</emphasis>
The specific name of the new species,
<taxonomicName id="4C6017B9FFA02E30FC8D2C858B92FAC0" authorityName="Koeda &amp; Ho" authorityYear="2019" box="[774,937,1197,1223]" class="Actinopterygii" family="Stomiidae" genus="Eustomias" kingdom="Animalia" order="Stomiiformes" pageId="7" pageNumber="101" phylum="Chordata" rank="species" species="tritentaculatus">
<emphasis id="B914B028FFA02E30FC8D2C858B92FAC0" box="[774,937,1197,1223]" italics="true" pageId="7" pageNumber="101">tritentaculatus</emphasis>
</taxonomicName>
, in reference to the three terminal filaments, which is diagnostic of the species.
</paragraph>
</subSubSection>
<subSubSection id="C37A3FB1FFA02E30FF4C2CDC8D62FB34" pageId="7" pageNumber="101" type="distribution">
<paragraph id="8BDF6C3AFFA02E30FF4C2CDC8D62FB34" blockId="7.[151,1437,764,1908]" pageId="7" pageNumber="101">
<emphasis id="B914B028FFA02E30FF4C2CDC895AFB08" bold="true" box="[199,353,1268,1295]" pageId="7" pageNumber="101">Distribution.</emphasis>
<taxonomicName id="4C6017B9FFA02E30FEEC2CDE8ABEFB08" authority="Koeda &amp; Ho, 2019" authorityName="Koeda &amp; Ho" authorityYear="2019" box="[359,645,1269,1295]" class="Actinopterygii" family="Stomiidae" genus="Eustomias" kingdom="Animalia" order="Stomiiformes" pageId="7" pageNumber="101" phylum="Chordata" rank="species" species="tritentaculatus" status="sp. nov.">
<emphasis id="B914B028FFA02E30FEEC2CDE8ABEFB08" box="[359,645,1269,1295]" italics="true" pageId="7" pageNumber="101">Eustomias tritentaculatus</emphasis>
</taxonomicName>
<emphasis id="B914B028FFA02E30FD072CDD8ADFFB08" bold="true" box="[652,740,1269,1295]" pageId="7" pageNumber="101">
<taxonomicNameLabel id="A2270D53FFA02E30FD072CDD8ADFFB08" box="[652,740,1269,1295]" pageId="7" pageNumber="101" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
is currently only known from southwestern
<collectingCountry id="F3772CAAFFA02E30FB5B2CDD8D1DFB08" box="[1232,1318,1269,1295]" name="Taiwan" pageId="7" pageNumber="101">Taiwan</collectingCountry>
. The
<typeStatus id="54DBD298FFA02E30FAEA2CDD88FCFB34" pageId="7" pageNumber="101" type="holotype">holotype</typeStatus>
was collected in the upper
<quantity id="4C98C1DFFFA02E30FE712D318A7AFB34" box="[506,577,1305,1331]" metricMagnitude="2" metricUnit="m" metricValue="1.0" pageId="7" pageNumber="101" unit="m" value="100.0">100 m</quantity>
by a mid-water trawl targeting Sakura-shrimp (
<taxonomicName id="4C6017B9FFA02E30FBD32D328D72FB34" baseAuthorityName="Hansen" baseAuthorityYear="1922" box="[1112,1353,1305,1331]" class="Malacostraca" family="Sergestidae" genus="Lucensosergia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="7" pageNumber="101" phylum="Arthropoda" rank="species" species="lucens">
<emphasis id="B914B028FFA02E30FBD32D328D72FB34" box="[1112,1353,1305,1331]" italics="true" pageId="7" pageNumber="101">Lucensosergia lucens</emphasis>
</taxonomicName>
).
</paragraph>
</subSubSection>
<subSubSection id="C37A3FB1FFA02E30FF4C2D158A8DF974" pageId="7" pageNumber="101" type="discussion">
<paragraph id="8BDF6C3AFFA02E30FF4C2D158A8DF974" blockId="7.[151,1437,764,1908]" pageId="7" pageNumber="101">
<emphasis id="B914B028FFA02E30FF4C2D158900FB50" bold="true" box="[199,315,1341,1367]" pageId="7" pageNumber="101">Remarks.</emphasis>
The present specimens were identified as a species of the subgenus
<taxonomicName id="4C6017B9FFA02E30FBBB2D168CE5FB50" box="[1072,1246,1342,1367]" class="Actinopterygii" family="Stomiidae" genus="Eustomias" kingdom="Animalia" order="Stomiiformes" pageId="7" pageNumber="101" phylum="Chordata" rank="subGenus" subGenus="Nominostomias">
<emphasis id="B914B028FFA02E30FBBB2D168CE5FB50" box="[1072,1246,1342,1367]" italics="true" pageId="7" pageNumber="101">Nominostomias</emphasis>
</taxonomicName>
by the following combination of characters: a single terminal bulb on barbel; three terminal filaments of the bulb; no tapering branch on the barbel; external pigmentation of the barbel absent (axis pigmented); pectoral-fin rays 3; pelvic-fin rays 7; vertebrae 66–68; number of teeth on jaws high (
<bibRefCitation id="EFF111CBFFA02E30FD3F2D818BEBFBC4" author="Regan, C. E. &amp; Trewavas, E." box="[692,976,1449,1475]" pageId="7" pageNumber="101" pagination="1 - 143" refId="ref5991" refString="Regan, C. E. &amp; Trewavas, E. (1930) The fishes of the families Stomiatidae and Malacosteidae. Dana Report, 6, 1 - 143." type="journal article" year="1930">Regan &amp; Trewavas 1930</bibRefCitation>
;
<bibRefCitation id="EFF111CBFFA02E30FC572D818C9AFBC4" author="Gibbs Jr., R. H. &amp; Clarke, T. A. &amp; Gomon, J. R." box="[988,1185,1449,1475]" pageId="7" pageNumber="101" pagination="1 - 139" refId="ref5632" refString="Gibbs Jr., R. H., Clarke, T. A. &amp; Gomon, J. R. (1983) Taxonomy and distribution of the stomioid fish genus Eustomias (Melanostomiidae). I. Subgenus Nominostomias. Smithsonian Contributions to Zoology, 380, 1 - 139. https: // doi. org / 10.5479 / si. 00810282.380" type="journal article" year="1983">
Gibbs
<emphasis id="B914B028FFA02E30FBAD2D828C64FBC4" box="[1062,1119,1449,1475]" italics="true" pageId="7" pageNumber="101">et al.</emphasis>
1983
</bibRefCitation>
;
<bibRefCitation id="EFF111CBFFA02E30FB262D818D65FBC4" author="Prokofiev, A. M." box="[1197,1374,1449,1475]" pageId="7" pageNumber="101" pagination="2 - 9" refId="ref5894" refString="Prokofiev, A. M. (2018) Three new species of Eustomias (Melanostomiidae) from the southern subtropical waters with the description of a new subgenus. Voprosy Ikhtiologii, 58 (1), 2 - 9. https: // doi. org / 10.1134 / S 0032945218010113" type="journal article" year="2018">Prokofiev 2018</bibRefCitation>
; only typical characters listed).
<bibRefCitation id="EFF111CBFFA02E30FE332DE58AB6FBE0" author="Gibbs Jr., R. H. &amp; Clarke, T. A. &amp; Gomon, J. R." box="[440,653,1485,1511]" pageId="7" pageNumber="101" pagination="1 - 139" refId="ref5632" refString="Gibbs Jr., R. H., Clarke, T. A. &amp; Gomon, J. R. (1983) Taxonomy and distribution of the stomioid fish genus Eustomias (Melanostomiidae). I. Subgenus Nominostomias. Smithsonian Contributions to Zoology, 380, 1 - 139. https: // doi. org / 10.5479 / si. 00810282.380" type="journal article" year="1983">
Gibbs
<emphasis id="B914B028FFA02E30FD892DE68A00FBE0" box="[514,571,1485,1511]" italics="true" pageId="7" pageNumber="101">et al.</emphasis>
(1983)
</bibRefCitation>
reviewed this subgenus and recognized 38 valid species, 25 of which were described as new. They also separated this subgenus into five species groups on the basis of the structure of the barbels.
<taxonomicName id="4C6017B9FFA02E30FF7C2E3E8A2DF828" authority="Koeda &amp; Ho, 2019" authorityName="Koeda &amp; Ho" authorityYear="2019" box="[247,534,1557,1583]" class="Actinopterygii" family="Stomiidae" genus="Eustomias" kingdom="Animalia" order="Stomiiformes" pageId="7" pageNumber="101" phylum="Chordata" rank="species" species="tritentaculatus" status="sp. nov.">
<emphasis id="B914B028FFA02E30FF7C2E3E8A2DF828" box="[247,534,1557,1583]" italics="true" pageId="7" pageNumber="101">Eustomias tritentaculatus</emphasis>
</taxonomicName>
<emphasis id="B914B028FFA02E30FD952E3D8A42F828" bold="true" box="[542,633,1557,1583]" pageId="7" pageNumber="101">
<taxonomicNameLabel id="A2270D53FFA02E30FD952E3D8A42F828" box="[542,633,1557,1583]" pageId="7" pageNumber="101" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
has a structure of barbel with the species of their group IV (8 species), species of which have a single terminal bulb with one to three terminal filaments.
<taxonomicName id="4C6017B9FFA02E30FBAB2E128D06F854" authority="Koeda &amp; Ho, 2019" authorityName="Koeda &amp; Ho" authorityYear="2019" box="[1056,1341,1593,1619]" class="Actinopterygii" family="Stomiidae" genus="Eustomias" kingdom="Animalia" order="Stomiiformes" pageId="7" pageNumber="101" phylum="Chordata" rank="species" species="tritentaculatus" status="sp. nov.">
<emphasis id="B914B028FFA02E30FBAB2E128D06F854" box="[1056,1341,1593,1619]" italics="true" pageId="7" pageNumber="101">Eustomias tritentaculatus</emphasis>
</taxonomicName>
<emphasis id="B914B028FFA02E30FACF2E118DA7F854" bold="true" box="[1348,1436,1593,1619]" pageId="7" pageNumber="101">
<taxonomicNameLabel id="A2270D53FFA02E30FACF2E118DA7F854" box="[1348,1436,1593,1619]" pageId="7" pageNumber="101" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
is most similar to
<taxonomicName id="4C6017B9FFA02E30FED02E768C4CF870" authority="Gibbs, Clarke &amp; Gomon, 1983" authorityName="Gibbs, Clarke &amp; Gomon" authorityYear="1983" box="[347,1143,1629,1655]" class="Actinopterygii" family="Stomiidae" genus="Eustomias" kingdom="Animalia" order="Stomiiformes" pageId="7" pageNumber="101" phylum="Chordata" rank="species" species="cancriensis" subGenus="Nominostomias">
<emphasis id="B914B028FFA02E30FED02E7689EBF870" box="[347,464,1630,1655]" italics="true" pageId="7" pageNumber="101">Eustomias</emphasis>
(
<emphasis id="B914B028FFA02E30FE542E768AB6F870" box="[479,653,1630,1655]" italics="true" pageId="7" pageNumber="101">Nominostomias</emphasis>
)
<emphasis id="B914B028FFA02E30FD162E768B26F870" box="[669,797,1630,1655]" italics="true" pageId="7" pageNumber="101">cancriensis</emphasis>
<bibRefCitation id="EFF111CBFFA02E30FCA92E758C4CF870" author="Gibbs Jr., R. H. &amp; Clarke, T. A. &amp; Gomon, J. R." box="[802,1143,1629,1655]" pageId="7" pageNumber="101" pagination="1 - 139" refId="ref5632" refString="Gibbs Jr., R. H., Clarke, T. A. &amp; Gomon, J. R. (1983) Taxonomy and distribution of the stomioid fish genus Eustomias (Melanostomiidae). I. Subgenus Nominostomias. Smithsonian Contributions to Zoology, 380, 1 - 139. https: // doi. org / 10.5479 / si. 00810282.380" type="journal article" year="1983">Gibbs, Clarke &amp; Gomon, 1983</bibRefCitation>
</taxonomicName>
from the western and central-north Pacific oceans, but differs in having a longer barbel, its length 65.3% (vs. 47–53% SL in
<taxonomicName id="4C6017B9FFA02E30FB572EAA8D41F89C" authorityName="Gibbs, Clarke &amp; Gomon" authorityYear="1983" box="[1244,1402,1666,1691]" class="Actinopterygii" family="Stomiidae" genus="Eustomias" kingdom="Animalia" order="Stomiiformes" pageId="7" pageNumber="101" phylum="Chordata" rank="species" species="cancriensis">
<emphasis id="B914B028FFA02E30FB572EAA8D41F89C" box="[1244,1402,1666,1691]" italics="true" pageId="7" pageNumber="101">E. cancriensis</emphasis>
</taxonomicName>
), a distinct ring-shaped black spot at base of bulb (vs. no pigmentation at base of bulb), and fewer anal-fin rays counts 35 (vs. 39) (
<bibRefCitation id="EFF111CBFFA02E30FEAE2EE189C9F8E4" author="Gibbs Jr., R. H. &amp; Clarke, T. A. &amp; Gomon, J. R." box="[293,498,1737,1763]" pageId="7" pageNumber="101" pagination="1 - 139" refId="ref5632" refString="Gibbs Jr., R. H., Clarke, T. A. &amp; Gomon, J. R. (1983) Taxonomy and distribution of the stomioid fish genus Eustomias (Melanostomiidae). I. Subgenus Nominostomias. Smithsonian Contributions to Zoology, 380, 1 - 139. https: // doi. org / 10.5479 / si. 00810282.380" type="journal article" year="1983">
Gibbs
<emphasis id="B914B028FFA02E30FEF92EE28996F8E4" box="[370,429,1737,1763]" italics="true" pageId="7" pageNumber="101">et al.</emphasis>
1983
</bibRefCitation>
).
<taxonomicName id="4C6017B9FFA02E30FD8C2EE28B13F8E4" authority="Koeda &amp; Ho, 2019" authorityName="Koeda &amp; Ho" authorityYear="2019" box="[519,808,1737,1763]" class="Actinopterygii" family="Stomiidae" genus="Eustomias" kingdom="Animalia" order="Stomiiformes" pageId="7" pageNumber="101" phylum="Chordata" rank="species" species="tritentaculatus" status="sp. nov.">
<emphasis id="B914B028FFA02E30FD8C2EE28B13F8E4" box="[519,808,1737,1763]" italics="true" pageId="7" pageNumber="101">Eustomias tritentaculatus</emphasis>
</taxonomicName>
<emphasis id="B914B028FFA02E30FCBA2EE18BB6F8E4" bold="true" box="[817,909,1737,1763]" pageId="7" pageNumber="101">
<taxonomicNameLabel id="A2270D53FFA02E30FCBA2EE18BB6F8E4" box="[817,909,1737,1763]" pageId="7" pageNumber="101" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
is also similar to
<taxonomicName id="4C6017B9FFA02E30FBD52EE28A26F900" authority="Johnson &amp; Rosenblatt, 1971" authorityName="Johnson &amp; Rosenblatt" authorityYear="1971" class="Actinopterygii" family="Stomiidae" genus="Eustomias" kingdom="Animalia" order="Stomiiformes" pageId="7" pageNumber="101" phylum="Chordata" rank="species" species="gibbsi" subGenus="Nominostomias">
<emphasis id="B914B028FFA02E30FBD52EE28CE8F8E4" box="[1118,1235,1738,1763]" italics="true" pageId="7" pageNumber="101">Eustomias</emphasis>
(
<emphasis id="B914B028FFA02E30FB6E2EE28DA8F8E4" box="[1253,1427,1738,1763]" italics="true" pageId="7" pageNumber="101">Nominostomias</emphasis>
)
<emphasis id="B914B028FFA02E30FF1C2EC588E0F900" box="[151,219,1773,1799]" italics="true" pageId="7" pageNumber="101">gibbsi</emphasis>
Johnson &amp; Rosenblatt, 1971
</taxonomicName>
from the Pacific Ocean; both have a black spot at the base of the bulb. However,
<taxonomicName id="4C6017B9FFA02E30FF1C2F3A898DF92C" authority="Koeda &amp; Ho, 2019" authorityName="Koeda &amp; Ho" authorityYear="2019" box="[151,438,1809,1835]" class="Actinopterygii" family="Stomiidae" genus="Eustomias" kingdom="Animalia" order="Stomiiformes" pageId="7" pageNumber="101" phylum="Chordata" rank="species" species="tritentaculatus" status="sp. nov.">
<emphasis id="B914B028FFA02E30FF1C2F3A898DF92C" box="[151,438,1809,1835]" italics="true" pageId="7" pageNumber="101">Eustomias tritentaculatus</emphasis>
</taxonomicName>
<emphasis id="B914B028FFA02E30FE362F398A2DF92C" bold="true" box="[445,534,1809,1835]" pageId="7" pageNumber="101">
<taxonomicNameLabel id="A2270D53FFA02E30FE362F398A2DF92C" box="[445,534,1809,1835]" pageId="7" pageNumber="101" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
differs from the latter in having three terminal filaments (vs. single in
<taxonomicName id="4C6017B9FFA02E30FAA22F3A8DB0F92C" authorityName="Johnson &amp; Rosenblatt" authorityYear="1971" box="[1321,1419,1809,1835]" class="Actinopterygii" family="Stomiidae" genus="Eustomias" kingdom="Animalia" order="Stomiiformes" pageId="7" pageNumber="101" phylum="Chordata" rank="species" species="gibbsi">
<emphasis id="B914B028FFA02E30FAA22F3A8DB0F92C" box="[1321,1419,1809,1835]" italics="true" pageId="7" pageNumber="101">E. gibbsi</emphasis>
</taxonomicName>
), barbel length 65.3% (vs. 30–52% SL), and a longer terminal filament, its maximum length 4.1% SL (vs. 1.6–2.0 in the similar sized specimens) (
<bibRefCitation id="EFF111CBFFA02E30FE692F718A92F974" author="Gibbs Jr., R. H. &amp; Clarke, T. A. &amp; Gomon, J. R." box="[482,681,1881,1907]" pageId="7" pageNumber="101" pagination="1 - 139" refId="ref5632" refString="Gibbs Jr., R. H., Clarke, T. A. &amp; Gomon, J. R. (1983) Taxonomy and distribution of the stomioid fish genus Eustomias (Melanostomiidae). I. Subgenus Nominostomias. Smithsonian Contributions to Zoology, 380, 1 - 139. https: // doi. org / 10.5479 / si. 00810282.380" type="journal article" year="1983">
Gibbs
<emphasis id="B914B028FFA02E30FDA52F728A5DF974" box="[558,614,1881,1907]" italics="true" pageId="7" pageNumber="101">et al.</emphasis>
1983
</bibRefCitation>
).
</paragraph>
</subSubSection>
</treatment>
</document>
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