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<document id="5790240B84ACC391EC2AB1E8F70E6AC8" ID-DOI="10.5281/zenodo.276689" ID-GBIF-Dataset="b6781430-d288-4cb4-b178-db6192a61dde" ID-ISSN="1175-5326" ID-Zenodo-Dep="276689" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1460230557710" checkinUser="plazi" docAuthor="Enghoff, Henrik" docDate="2011" docId="03BC87EAFFB23140FF22F94F050CFDA9" docLanguage="en" docName="zt02753p041.pdf" docOrigin="Zootaxa 2753" docStyle="DocumentStyle:890A69B780ED73D6DB8551B71C8AC79E.4:Zootaxa.2009-2012.journal_article" docStyleId="890A69B780ED73D6DB8551B71C8AC79E" docStyleName="Zootaxa.2009-2012.journal_article" docStyleVersion="4" docTitle="Epibolus Cook 1897" docType="treatment" docVersion="9" lastPageNumber="33" masterDocId="FF85FF92FFAD3160FFB5FFF00761FFF9" masterDocTitle="East African giant millipedes of the tribe Pachybolini (Diplopoda, Spirobolida, Pachybolidae)" masterLastPageNumber="41" masterPageNumber="1" pageNumber="32" updateTime="1698238933756" updateUser="plazi">
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<mods:title id="2900AA3B11BF0518B1D8AEAC48CAB865">East African giant millipedes of the tribe Pachybolini (Diplopoda, Spirobolida, Pachybolidae)</mods:title>
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<mods:namePart id="F69454A1385CBFB7864DDB28CB635D31">Enghoff, Henrik</mods:namePart>
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<mods:date id="F7C53FE446FF973A6290FE719C877B54">2011</mods:date>
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<treatment id="03BC87EAFFB23140FF22F94F050CFDA9" ID-DOI="http://doi.org/10.5281/zenodo.5612674" ID-GBIF-Taxon="119387241" ID-Zenodo-Dep="5612674" LSID="urn:lsid:plazi:treatment:03BC87EAFFB23140FF22F94F050CFDA9" httpUri="http://treatment.plazi.org/id/03BC87EAFFB23140FF22F94F050CFDA9" lastPageId="32" lastPageNumber="33" pageId="31" pageNumber="32">
<subSubSection id="C30F6577FFB2317FFF22F94F068CF920" box="[151,493,1727,1753]" pageId="31" pageNumber="32" type="nomenclature">
<paragraph id="8BAA36FCFFB2317FFF22F94F068CF920" blockId="31.[151,493,1727,1753]" box="[151,493,1727,1753]" pageId="31" pageNumber="32">
<heading id="D0E28190FFB2317FFF22F94F068CF920" bold="true" box="[151,493,1727,1753]" fontSize="11" level="1" pageId="31" pageNumber="32" reason="1">
<emphasis id="B961EAEEFFB2317FFF22F94F068CF920" bold="true" box="[151,493,1727,1753]" pageId="31" pageNumber="32">
Genus
<taxonomicName id="4C154D7FFFB2317FFF5BF94F068CF920" authority="Cook, 1897" authorityName="Cook" authorityYear="1897" box="[238,493,1727,1753]" class="Actinopterygii" family="Labridae" genus="Epibolus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Perciformes" pageId="31" pageNumber="32" phylum="Chordata" rank="genus">
<emphasis id="B961EAEEFFB2317FFF5BF94F063BF920" bold="true" box="[238,346,1727,1753]" italics="true" pageId="31" pageNumber="32">Epibolus</emphasis>
<bibRefCitation id="EF844B0DFFB2317FFED4F94F068CF920" author="Cook" box="[353,493,1727,1753]" pageId="31" pageNumber="40" refString="Cook, O. F. (1897) New relatives of Spirobolus giganteus. Brandtia, 18, 73 - 75." type="journal article" year="1897">Cook, 1897</bibRefCitation>
</taxonomicName>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C30F6577FFB23140FF22F90C050CFDA9" lastPageId="32" lastPageNumber="33" pageId="31" pageNumber="32" type="materials_examined">
<paragraph id="8BAA36FCFFB2317FFF22F90C05D7F8E8" blockId="31.[151,694,1787,1809]" box="[151,694,1787,1809]" pageId="31" pageNumber="32">
<typeStatus id="54AE885EFFB2317FFF22F90C07A9F8E8" box="[151,200,1788,1809]" pageId="31" pageNumber="32">Type</typeStatus>
species:
<taxonomicName id="4C154D7FFFB2317FFE93F90B05D7F8E8" authority="Gerstacker, 1873" authorityName="Gerstacker" authorityYear="1873" box="[294,694,1787,1809]" class="Diplopoda" family="Spirobolidae" genus="Spirobolus" kingdom="Animalia" order="Spirobolida" pageId="31" pageNumber="32" phylum="Arthropoda" rank="species" species="pulchripes">
<emphasis id="B961EAEEFFB2317FFE93F90B0565F8E8" box="[294,516,1787,1809]" italics="true" pageId="31" pageNumber="32">Spirobolus pulchripes</emphasis>
Gerstäcker, 1873
</taxonomicName>
</paragraph>
<paragraph id="8BAA36FCFFB2317FFF22F8C604C1F896" blockId="31.[151,1157,1845,1904]" pageId="31" pageNumber="32">
<emphasis id="B961EAEEFFB2317FFF22F8C60600F8B7" bold="true" box="[151,353,1845,1870]" pageId="31" pageNumber="32">Included species.</emphasis>
<taxonomicName id="4C154D7FFFB2317FFEDDF8C705B6F8B7" authority="Gerstacker, 1873" authorityName="Gerstacker" authorityYear="1873" box="[360,727,1846,1870]" class="Diplopoda" family="Spirobolidae" genus="Spirobolus" kingdom="Animalia" order="Spirobolida" pageId="31" pageNumber="32" phylum="Arthropoda" rank="species" species="pulchripes">
<emphasis id="B961EAEEFFB2317FFEDDF8C7069CF8B7" box="[360,509,1846,1870]" italics="true" pageId="31" pageNumber="32">E. pulchripes</emphasis>
(Gerstäcker, 1873)
</taxonomicName>
;
<taxonomicName id="4C154D7FFFB2317FFD51F8C703E4F8B7" authority="Lawrence, 1967" authorityName="Lawrence" authorityYear="1967" box="[740,1157,1846,1870]" class="Actinopterygii" family="Labridae" genus="Epibolus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Perciformes" pageId="31" pageNumber="32" phylum="Chordata" rank="species" species="mossambicensis">
<emphasis id="B961EAEEFFB2317FFD51F8C704D6F8B7" box="[740,951,1846,1870]" italics="true" pageId="31" pageNumber="32">E. mossambicensis</emphasis>
(Lawrence, 1967)
</taxonomicName>
<emphasis id="B961EAEEFFB2317FFF70F8A6065EF896" bold="true" box="[197,319,1878,1903]" pageId="31" pageNumber="32">Diagnosis.</emphasis>
A genus of East African Pachybolini characterized by
</paragraph>
<paragraph id="8BAA36FCFFB2317FFF22F86A0537F848" blockId="31.[151,1181,1945,2004]" box="[151,598,1945,1969]" pageId="31" pageNumber="32">- no coxal lobes on male legs 6 and 7</paragraph>
<paragraph id="8BAA36FCFFB2317FFF22F84D03FCF82D" blockId="31.[151,1181,1945,2004]" box="[151,1181,1980,2004]" pageId="31" pageNumber="32">
- three (two to five) pairs of ventral macrosetae on male legs from pair 5 onward (
<figureCitation id="132E2A79FFB2317FFBFEF84C03EEF82D" box="[1099,1167,1980,2004]" captionStart="FIGURES 4 8" captionStartId="5.[151,264,1919,1942]" captionTargetBox="[157,1430,195,1890]" captionTargetId="figure@5.[151,1436,193,1898]" captionTargetPageId="5" captionText="FIGURES 4 8. Male legs, SEM. 4: Crurifarcimen vagans n. gen., n. sp., paratype, tarsus, lateral view. 5: C. vagans, paratype, tarsus, ventral view. 6: Epibolus pulchripes, tarsus, lateral view. 7: Hadrobolus crassicollis, holotype, ventral-lateral view. 8: C. vagans, paratype, surface structure of ventral pad. Scales 1 mm (5, 7), 0.1 mm (4, 6), 0.01 mm (8)." httpUri="https://zenodo.org/record/276692/files/figure.png" pageId="31" pageNumber="32">Fig. 6</figureCitation>
).
</paragraph>
<paragraph id="8BAA36FCFF8D3140FF22FF68044DFF56" blockId="32.[151,1290,151,384]" box="[151,812,151,175]" pageId="32" pageNumber="33">- a moveable lateral appendix on the posterior gonopods</paragraph>
<paragraph id="8BAA36FCFF8D3140FF22FF4B0429FF2B" blockId="32.[151,1290,151,384]" box="[151,840,186,210]" pageId="32" pageNumber="33">- striated membranous lamellae at tip of posterior gonopod</paragraph>
<paragraph id="8BAA36FCFF8D3140FF22FF2E0455FEEE" blockId="32.[151,1290,151,384]" pageId="32" pageNumber="33">- no mesal process, no free solenomerite, no distal complex of parallel ridges on posterior gonopod - no long meso-distal process on anterior gonopod coxite</paragraph>
<paragraph id="8BAA36FCFF8D3140FF22FED3047FFEC3" blockId="32.[151,1290,151,384]" box="[151,798,290,314]" pageId="32" pageNumber="33">- no overlapping lobes on anterior gonopod telopodites</paragraph>
<paragraph id="8BAA36FCFF8D3140FF22FEB6051EFE86" blockId="32.[151,1290,151,384]" pageId="32" pageNumber="33">
- apical margin of winglike expansions of second female coxae regularly rounded (
<figureCitation id="132E2A79FF8D3140FBE2FEB503C9FEA4" box="[1111,1192,325,349]" captionStart="FIGURES 11 12" captionStartId="7.[151,264,965,988]" captionTargetBox="[210,1380,180,949]" captionTargetId="figure@7.[209,1382,179,953]" captionTargetPageId="7" captionText="FIGURES 11 12. Female second pair of legs with one vulva attached, anterior view. 11: Crurifarcimen vagans, 12: Epibolus pulchripes. The arrows point to the distal margin of the lateral extensions of the coxosternum. Scales 1 cm. P. Pimvichai phot." httpUri="https://zenodo.org/record/276694/files/figure.png" pageId="32" pageNumber="33">Fig. 12</figureCitation>
) - an apical appendix on oral vulval valve
</paragraph>
<paragraph id="8BAA36FCFF8D3140FF72FE5D050CFDA9" blockId="32.[151,1436,429,592]" pageId="32" pageNumber="33">
<bibRefCitation id="EF844B0DFF8D3140FF72FE5D0615FE3C" author="Enghoff" box="[199,372,429,453]" pageId="32" pageNumber="40" refString="Enghoff, H. (1977) Revision of the East African millipede genus Epibolus Cook, 1897. With notes on the biology of E. pulchripes (Gerstacker) (Diplopoda, Spirobolida: Pachybolidae). Entomologica scandinavica, 8, 1 - 8." type="journal article" year="1977">Enghoff (1977)</bibRefCitation>
recognised three species of this genus: the widespread
<taxonomicName id="4C154D7FFF8D3140FC57FE5E0316FE3C" box="[994,1143,429,453]" class="Diplopoda" family="Spirobolidae" genus="Spirobolus" kingdom="Animalia" order="Spirobolida" pageId="32" pageNumber="33" phylum="Arthropoda" rank="species" species="pulchripes">
<emphasis id="B961EAEEFF8D3140FC57FE5E0316FE3C" box="[994,1143,429,453]" italics="true" pageId="32" pageNumber="33">E. pulchripes</emphasis>
</taxonomicName>
(
<collectingCountry id="F302766CFF8D3140FB33FE5D03B0FE3C" box="[1158,1233,429,453]" name="Kenya" pageId="32" pageNumber="33">Kenya</collectingCountry>
and
<collectingCountry id="F302766CFF8D3140FABDFE5D0210FE3C" box="[1288,1393,429,453]" name="Tanzania" pageId="32" pageNumber="33">Tanzania</collectingCountry>
),
<taxonomicName id="4C154D7FFF8D3140FA31FE5E0663FE11" class="Actinopterygii" family="Labridae" genus="Epibolus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Perciformes" pageId="32" pageNumber="33" phylum="Chordata" rank="species" species="bravensis">
<emphasis id="B961EAEEFF8D3140FA31FE5E0663FE11" italics="true" pageId="32" pageNumber="33">E. bravensis</emphasis>
</taxonomicName>
from
<collectingCountry id="F302766CFF8D3140FEFFFE3F06C7FE1E" box="[330,422,463,487]" name="Somalia" pageId="32" pageNumber="33">Somalia</collectingCountry>
and
<taxonomicName id="4C154D7FFF8D3140FE54FE2105D7FE11" box="[481,694,464,488]" class="Actinopterygii" family="Labridae" genus="Epibolus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Perciformes" pageId="32" pageNumber="33" phylum="Chordata" rank="species" species="mossambicensis">
<emphasis id="B961EAEEFF8D3140FE54FE2105D7FE11" box="[481,694,464,488]" italics="true" pageId="32" pageNumber="33">E. mossambicensis</emphasis>
</taxonomicName>
from
<collectingCountry id="F302766CFF8D3140FD4BFE3F04F4FE1E" box="[766,917,463,487]" name="Mozambique" pageId="32" pageNumber="33">Mozambique</collectingCountry>
. The morphological characters distinguishing the two “geographical satellite” species from
<taxonomicName id="4C154D7FFF8D3140FD1CFE040423FDF2" box="[681,834,499,523]" class="Diplopoda" family="Spirobolidae" genus="Spirobolus" kingdom="Animalia" order="Spirobolida" pageId="32" pageNumber="33" phylum="Arthropoda" rank="species" species="pulchripes">
<emphasis id="B961EAEEFF8D3140FD1CFE040423FDF2" box="[681,834,499,523]" italics="true" pageId="32" pageNumber="33">E. pulchripes</emphasis>
</taxonomicName>
were subtle, and in the case of
<taxonomicName id="4C154D7FFF8D3140FB0BFE04022AFDF2" box="[1214,1355,499,523]" class="Actinopterygii" family="Labridae" genus="Epibolus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Perciformes" pageId="32" pageNumber="33" phylum="Chordata" rank="species" species="bravensis">
<emphasis id="B961EAEEFF8D3140FB0BFE04022AFDF2" box="[1214,1355,499,523]" italics="true" pageId="32" pageNumber="33">E. bravensis</emphasis>
</taxonomicName>
drawn exclusively from the original description (Silvestri, 1897) because these characters could not be seen on the male
<typeStatus id="54AE885EFF8D3140FF22FDC7079CFDB6" box="[151,253,567,591]" pageId="32" pageNumber="33" type="holotype">holotype</typeStatus>
. See further under
<emphasis id="B961EAEEFF8D3140FE65FDC9050CFDA9" box="[464,621,568,592]" italics="true" pageId="32" pageNumber="33">
<taxonomicName id="4C154D7FFF8D3140FE65FDC90508FDA9" box="[464,617,568,592]" class="Diplopoda" family="Spirobolidae" genus="Spirobolus" kingdom="Animalia" order="Spirobolida" pageId="32" pageNumber="33" phylum="Arthropoda" rank="species" species="pulchripes">E. pulchripes</taxonomicName>
.
</emphasis>
</paragraph>
</subSubSection>
</treatment>
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